taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860366/files/figure.png	https://doi.org/10.5281/zenodo.7860366	FIGURE 3. Photographs of shell specimens of Pupa solidula (Linnaeus, 1758), ventral view. A. Specimen from Koumac, New Caledonia (MHNH IM-2013-86155). B. Specimen from Koumac, New Caledonia (MHNH IM-2013-86164). C. Neotype of Pupa solidula (Linnaeus, 1758) (MHNH IM-2013-86152). D. Specimen from Koumac, New Caledonia (MHNH IM-2013- 86163).	FIGURE 3. Photographs of shell specimens of Pupa solidula (Linnaeus, 1758), ventral view. A. Specimen from Koumac, New Caledonia (MHNH IM-2013-86155). B. Specimen from Koumac, New Caledonia (MHNH IM-2013-86164). C. Neotype of Pupa solidula (Linnaeus, 1758) (MHNH IM-2013-86152). D. Specimen from Koumac, New Caledonia (MHNH IM-2013- 86163).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8846678ED72F7B44A7F86B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF8446648ED72C8B422BFEAB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860382/files/figure.png	https://doi.org/10.5281/zenodo.7860382	FIGURE 9. Photographs of shell specimens of Pupa sulcata (Gmelin, 1791), ventral view. A–C. Syntypes of Tornatella glabra Reeve, 1842 (NHMUK 196956), ©NHMUK, photos by K. Webb. D. Specimen from Koumac, New Caledonia (MHNH IM-2013-86165). E. Specimen from Koumac, New Caledonia (MHNH IM-2013-86145). F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86147). G. Specimen from Koumac, New Caledonia (MHNH IM-2013-86146). H. Specimen from Koumac, New Caledonia (MHNH IM-2013-86160).	FIGURE 9. Photographs of shell specimens of Pupa sulcata (Gmelin, 1791), ventral view. A–C. Syntypes of Tornatella glabra Reeve, 1842 (NHMUK 196956), ©NHMUK, photos by K. Webb. D. Specimen from Koumac, New Caledonia (MHNH IM-2013-86165). E. Specimen from Koumac, New Caledonia (MHNH IM-2013-86145). F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86147). G. Specimen from Koumac, New Caledonia (MHNH IM-2013-86146). H. Specimen from Koumac, New Caledonia (MHNH IM-2013-86160).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF81467F8ED72AEC45EFFE1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860386/files/figure.png	https://doi.org/10.5281/zenodo.7860386	FIGURE 10. Photographs of shell specimens of Pupa nitidula (Lamarck, 1822), ventral view. A–E. Syntypes of Tornatella nitidula Lamarck, 1822 (MHNG Moll-51412), ©MHNG, photos by E. Tardy. F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86157). G. Specimen from Koumac, New Caledonia (MHNH IM-2013-86156).	FIGURE 10. Photographs of shell specimens of Pupa nitidula (Lamarck, 1822), ventral view. A–E. Syntypes of Tornatella nitidula Lamarck, 1822 (MHNG Moll-51412), ©MHNG, photos by E. Tardy. F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86157). G. Specimen from Koumac, New Caledonia (MHNH IM-2013-86156).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9D467D8ED72DE34082FEF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860390/files/figure.png	https://doi.org/10.5281/zenodo.7860390	FIGURE 11. Photographs of shell specimens of Pupa coccinata (Reeve, 1842), ventral view. A–C. Syntypes of Tornatella coccinata Reeve, 1842 (NHMUK 1998142), ©NHMUK, photos by K. Webb. D. Holotype of Pupa roseomaculata Iredale, 1936 (AM C.60702), ©AM. E. Syntype of Tornatella insculpta Reeve, 1842 (NHMUK 196957), ©NHMUK, photo by K. Webb. F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86186). G. Specimen from Koumac, New Caledonia (MHNH IM- 2013-86153). H. Specimen from Koumac, New Caledonia (MHNH IM-2013-86143).	FIGURE 11. Photographs of shell specimens of Pupa coccinata (Reeve, 1842), ventral view. A–C. Syntypes of Tornatella coccinata Reeve, 1842 (NHMUK 1998142), ©NHMUK, photos by K. Webb. D. Holotype of Pupa roseomaculata Iredale, 1936 (AM C.60702), ©AM. E. Syntype of Tornatella insculpta Reeve, 1842 (NHMUK 196957), ©NHMUK, photo by K. Webb. F. Specimen from Koumac, New Caledonia (MHNH IM-2013-86186). G. Specimen from Koumac, New Caledonia (MHNH IM- 2013-86153). H. Specimen from Koumac, New Caledonia (MHNH IM-2013-86143).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9F467C8ED72D8F4091FBCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860394/files/figure.png	https://doi.org/10.5281/zenodo.7860394	FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP.	FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9E46798ED728D340FCFB83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860398/files/figure.png	https://doi.org/10.5281/zenodo.7860398	FIGURE 13. Photographs of shell specimens of Pupa affinis (A.Adams, 1855). A. Lectotype of Solidula affinis A.Adams, 1855 from Port Essington, Australia (NHMUK 20020237c), ©NHMUK, photo by K. Webb. B. Paralectotype of Solidula affinis A. Adams, 1855 from Moreton Bay, Australia (NHMUK 20020236), ©NHMUK, photo by K. Webb. C. Paralectotype of Solidula affinis A. Adams, 1855 from Ticao Island, Philippines (NHMUK 20020238), ©NHMUK, photo by K. Webb. D–E. Syntypes of Tornatella fumata Reeve, 1865 (NHMUK 20020233), ©NHMUK, photos by K. Webb. F. Holotype of Buccinulus huttoni Kirk, 1882 (NMNZ M.000051), ©NMNZ. G–H. Syntypes of Actaeon cinereus R.B. Watson, 1884 (NHMUK 1887.2.9.2131), ©NHMUK, photos by K. Webb. I. Specimen from Koumac, New Caledonia (MNHN IM-2013-86166). J. Specimen from Koumac, New Caledonia (MNHN IM-2013-86144).	FIGURE 13. Photographs of shell specimens of Pupa affinis (A.Adams, 1855). A. Lectotype of Solidula affinis A.Adams, 1855 from Port Essington, Australia (NHMUK 20020237c), ©NHMUK, photo by K. Webb. B. Paralectotype of Solidula affinis A. Adams, 1855 from Moreton Bay, Australia (NHMUK 20020236), ©NHMUK, photo by K. Webb. C. Paralectotype of Solidula affinis A. Adams, 1855 from Ticao Island, Philippines (NHMUK 20020238), ©NHMUK, photo by K. Webb. D–E. Syntypes of Tornatella fumata Reeve, 1865 (NHMUK 20020233), ©NHMUK, photos by K. Webb. F. Holotype of Buccinulus huttoni Kirk, 1882 (NMNZ M.000051), ©NMNZ. G–H. Syntypes of Actaeon cinereus R.B. Watson, 1884 (NHMUK 1887.2.9.2131), ©NHMUK, photos by K. Webb. I. Specimen from Koumac, New Caledonia (MNHN IM-2013-86166). J. Specimen from Koumac, New Caledonia (MNHN IM-2013-86144).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9B46788ED7289B4062FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860400/files/figure.png	https://doi.org/10.5281/zenodo.7860400	FIGURE 14. Photographs of shell specimens of Pupa strigosa (Gould, 1859). A. Lectotype of Buccinulus strigosus Gould, 1859 (USNM 1311), ©USNM. B–C. Two syntypes of Buccinulus fraterculus Dunker, 1882 (NHMUK 1905.12.30.40), ©NHMUK, photos by K. Webb. D. Specimen from Tomioka Bay, Kumamoto Prefecture, Japan (LACM 82-23.12). E. Specimen from Namihaya-zaki, Wakayama Prefecture, Japan (LACM 82-19.23).	FIGURE 14. Photographs of shell specimens of Pupa strigosa (Gould, 1859). A. Lectotype of Buccinulus strigosus Gould, 1859 (USNM 1311), ©USNM. B–C. Two syntypes of Buccinulus fraterculus Dunker, 1882 (NHMUK 1905.12.30.40), ©NHMUK, photos by K. Webb. D. Specimen from Tomioka Bay, Kumamoto Prefecture, Japan (LACM 82-23.12). E. Specimen from Namihaya-zaki, Wakayama Prefecture, Japan (LACM 82-19.23).	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9B46788ED7289B4062FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9A46778ED72AEF4471FA53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860404/files/figure.png	https://doi.org/10.5281/zenodo.7860404	FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang.	FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9546758ED72A2B4290FDD7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860404/files/figure.png	https://doi.org/10.5281/zenodo.7860404	FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang.	FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9746758ED72EAF444FFA69.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860408/files/figure.png	https://doi.org/10.5281/zenodo.7860408	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9746748ED72A2E4393FE3F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860408/files/figure.png	https://doi.org/10.5281/zenodo.7860408	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860362/files/figure.png	https://doi.org/10.5281/zenodo.7860362	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860368/files/figure.png	https://doi.org/10.5281/zenodo.7860368	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860370/files/figure.png	https://doi.org/10.5281/zenodo.7860370	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860372/files/figure.png	https://doi.org/10.5281/zenodo.7860372	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860376/files/figure.png	https://doi.org/10.5281/zenodo.7860376	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860380/files/figure.png	https://doi.org/10.5281/zenodo.7860380	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
03C38798FF9646738ED72DC74231FC47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7860408/files/figure.png	https://doi.org/10.5281/zenodo.7860408	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.	2023-04-21	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.		Zenodo	biologists	Valdés, Ángel;Feliciano, Kendall;Malaquias, Manuel A. E.			
