taxonID	type	description	language	source
503B8E36676CB147FDA0A7AFE9555CB5.taxon	description	(Figures 1 (a, b), 2 (a – i), 3 (a – d); Table 1) Nomenclatural statement A life science identifier (LSID) number was obtained for the new genus: urn: lsid: zoobank. org: pub: 46218584 - 43 FE- 4 C 05 - 9537 - FFC 81 EACCFDB Synonymy ′ New genus E ̍, Cumberlidge et al. (2020): fig. 1.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E36676CB147FDA0A7AFE9555CB5.taxon	type_taxon	Type species Vichai cyanalepou sp. nov., by present designation, gender masculine.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E36676CB147FDA0A7AFE9555CB5.taxon	diagnosis	Diagnosis Carapace moderate height (CH / FW 1.0); front broad (CW / FW = 3.1), deflexed; surface with faint carinae in anterolateral and posterlateral regions, otherwise smooth; anterolateral margin between exorbital tooth, epibranchial tooth concave, granulated, lacking intermediate tooth; epibranchial tooth reduced to small granule, close to exorbital tooth, positioned in line with postorbital margin; lateral margin strongly convex, lined with granules, posterior end curving inward, not continuous with posterolateral margin; postfrontal crest faint, incomplete, not traversing entire carapace, epigastric crests faint, in advanced position, in line with postorbital margin, postorbital crests faint, ending before meeting epibranchial teeth; semicircular, cervical sulci deep, cervical sulcus ending before meeting postorbital crest (Figure 1 (a )). Suborbital, subhepatic regions of branchiostegite with faint granules, pterygostomial region smooth except for granules along epimeral (longitudinal) sulcus; vertical sulcus on branchiostegite curved, granular, running downward from base of epibranchial tooth to epimeral sulcus (Figure 1 (b), 2 (a )). Epistomial tooth triangular, deflexed, edges with faint granules, epistomial margins smooth (Figure 2 (a )); anterior lobe on terminal article of mandibular palp small (MPAL / MPTA = 0.4) (Figure 2 (i )). Ambulatory legs (P 2 – 5) stout, not elongated (P 2 – 5 / CW = 5.4) (Figure 1 (a); Table 1); cheliped ischium margins smooth, rounded (Figure 2 (d )); third maxilliped exopod with long flagellum (~ 0.5 × merus length) (Figure 2 (g, h )). G 1 TA medium length (G 1 TA / G 1 SA = 0.3), slim, basal two-thirds directed outward, distal third curving upward, tapering to blunt open tip; G 1 TA ventral side with faint sulcus between G 1 TA-G 1 SA junction (Figure 3 (a )); G 1 TA dorsal side with broad, trapezoid dorsal membrane (DM) at G 1 TA-G 1 SA junction; DM superior margin arrowhead-shaped, inferior margin U-shaped, lateral margin short, mesial margin elongated (Figure 3 (b )); G 1 SA lacking raised rounded shoulder on external margin near G 1 TA-G 1 SA junction (Figure 3 (a, b )); G 2 TA distinctly elongated, flagellum-like (G 2 TA / G 2 SA = 0.85) with pointed tip (Figure 3 (d )).	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E36676CB147FDA0A7AFE9555CB5.taxon	distribution	Distribution Endemic to Madagascar. Vichai gen. nov. is currently known only from Marojejy National Park in the Sava Region in northern Madagascar.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E36676CB147FDA0A7AFE9555CB5.taxon	etymology	Etymology The genus is named in honour of Vichai Srivaddhanaprabha, the owner of the English Premier League football club Leicester City (LCFC) from 2010 until his death in a tragic helicopter crash at the King Power Stadium in October 2018. Vichai was not only extremely mindful of the Leicester community and the LCFC supporters, but under his leadership the Football Club won the 2015 – 16 English Premier League title against all expectations, after starting the season as 5000 / 1 rank outsiders. The generic name, Vichai, is used as a Latin noun in nominative singular and treated as masculine.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E36676CB147FDA0A7AFE9555CB5.taxon	discussion	Remarks The genus Vichai is established for a new species, V. cyanalepou (see later). The recognition of the present new genus is based on phylogenetic and morphological evidence. The molecular phylogeny in Cumberlidge et al. (2020, fig. 1, ′ New genus E ̍) indicates that V. cyanalepou sp. nov. is part of a unique basal clade that is clearly genetically separate from all other Malagasy genera. Mophologically, Vichai gen. nov. can be distinguished from the three other genera found in the Marojejy National Park (i. e. Foza Reed and Cumberlidge, 2006, Hydrothelphusa A. Milne-Edwards, 1872, and Marojejy Cumberlidge, Boyko and Harvey, 2002) as follows. The G 1 TA of Vichai gen. nov. is medium in length (G 1 TA / G 1 SA = 0.3) and slim where the basal two-thirds are directed outward and the distal third curves upward (Figure 3 (a – c )) vs a G 1 TA that is short (G 1 TA / SA = 0.25), straight, and broadly conical in Foza (see Leever et al. 2022, fig. 7 g, h). The exorbital and epibranchial teeth in Vichai gen. nov. are separated by a shallow notch (Figure 1 (a )), whereas these teeth are separated by a deep cleft in Hydrothelphusa A. Milne-Edwards, 1872 (see Cumberlidge and Sternberg 2002, fig. 1 a – c; Cumberlidge et al. 2007, figs 1, 16). Finally, the ambulatory legs of Vichai gen. nov. are short and stout (ΣP 2 – 5 / CW = 5.4) (Figure 1 (a); Table 1) and its eyestalks and corneas are of normal length (Figures 1 (a), 2 (a )), whereas the ambulatory legs of Marojejy Cumberlidge et al. (2002) are elongated and slender (ΣP 2 – 5 / CW = 7.0) and its eyestalks are short, taper distally, and have a reduced cornea (see Cumberlidge et al. 2002, fig. 2 f). The medium-sized anterior lobe on the terminal article of the mandibular palp of Vichai gen. nov. (MPAL / MPTA = 0.4) (Figure 2 (i )) distinguishes it from three other Malagasy genera: Boreathelphusa (Cumberlidge, 2010), Madagapotamon Bott, 1955 and Skelosophusa Ng and Takeda (1994). In all three of these genera, the anterior lobe on the mandibular palp is noticeably small (MPAL / MPTA = 0.2), and ledge-like (see Cumberlidge and Sternberg 2002, fig. 4 h – l) rather than a broad rounded lobe as in Vichai gen. nov. (Figure 2 (i )). Vichai gen. nov. can be distinguished from the remaining Malagasy genera as follows. The postfrontal crest of Vichai gen. nov. is incomplete and faint and does not traverse the entire carapace (Figure 1 (a )), whereas the postfrontal crest is well defined and completely traverses the carapace in Agora (see Cumberlidge et al. 2020). The anterolateral regions of the carapace surface of Vichai gen. nov. are mostly smooth and only faintly carinated (Figure 1 (a )), vs anterolateral regions of the carapace that have fields of well-defined carinae in Crosnautes (see Cumberlidge et al. 2021, fig. 1 a, b). The G 1 TA of Vichai gen. nov. is slim, tapers evenly, and curves upward distally (Figure 3 (a, b )), vs a G 1 TA that is broadest at the midpoint and is straight along its length in Glabrithelphusa Meyer et al. (2014) (see Meyer et al. 2014, figs 1 a, 3 a). The lateral carapace margin of Vichai gen. nov. is lined with small granules (Figure (1 a )), vs a lateral margin that is lined with either small or medium-sized teeth in Malagasya Cumberlidge and Sternberg (2002) (see Cumberlidge and Sternberg 2002, fig. 1 e, f; Cumberlidge et al. 2020, fig. 10 a, c, e). Vichai gen. nov. can be also distinguished from Toamasina Leever et al. (2022), as follows. The margins of the cheliped ischium of Vichai gen. nov. are smooth and rounded (Figure 2 (d, e )), vs cheliped ischium margins that are lined with small teeth (see Leever et al. 2022, fig. 4 g, h); the exopod of the third maxilliped of Vichai gen. nov. has a mediumlength flagellum (~ 0.5 × merus length) (Figure 2 (g, h )), vs a third maxilliped exopod with an elongated flagellum (equal to the merus length) (see Leever et al. 2022, fig. 2 f). In addition, the ambulatory legs of Vichai gen. nov. (P 2 – 5) are short and stout (ΣP 2 – 5 / CW = 5.4) (Figure 1 (a); Table 1), vs ambulatory legs (P 2 – 5) that are medium length (ΣP 2 – 5 / CW = 6.4) in Toamasina (see Leever et al. 2022, table 3). Finally, Vichai gen. nov. can be distinguished from Vahatra Leever et al. (2022) by the characters of the chelipeds. The inner margin of the cheliped carpus of Vichai gen. nov. has a relatively small pointed distal tooth, a smaller acute proximal tooth, and granules between these teeth (Figure 2 (f )), vs a cheliped carpus inner margin with a relatively larger, pointed distal tooth and a significantly smaller, acute proximal tooth, with no granules between the teeth in Vahatra (see Leever et al. 2022, fig. 4 f). The cutting edge of the fixed finger (pollex of propodus) of the major (left) chela of Vichai gen. nov. has one large molar tooth flanked by small teeth proximally (Figure 2 (e )), whereas the fixed finger of the major (right) chela of Vahatra has four large molars proximally (see Leever et al. 2022, fig. 4 g, h). Further, the cutting edge of the movable finger (dactylus) of the major (left) chela of Vichai gen. nov. is lined with small teeth only and lacks large teeth (Figure 2 (e )), whereas the cutting edge of the movable finger of the major (right) chela of Vahatra has two large teeth proximally and one large tooth midway (see Leever et al. 2022, fig. 4 g, h).	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	description	(Figures 1 (a, b), 2 (a – i), 3 (a – d); Table 1) Nomenclatural statement A life science identifier (LSID) number was obtained for the new species: urn: lsid: zoobank. org: act: B 5985969 - B 677 - 47 E 4 - BE 59 - 37 C 1308 B 7 A 66	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	materials_examined	Material examined Type material. Holotype: Madagascar: adult J (CW 38.5, CL 27.7, CH 12.5, FW 12.4 mm), Sava Region, Marojejy National Park, 11.5 km SE of Doany (14.4267 ° S, 49.60833 ° E), 810 m ASL, in mostly undisturbed upper lowland forest, in tributary of Bemanivy River. Collected by hand during day on river bed, mostly rocky, with some open basins with sandy alluvium, coll. S. M. Goodman, 16 October 2001 (FMNH 7589) (GenBank MT 749742). Paratypes: Madagascar: subadult J (CW 28.4, CL 19.7, CH 11.0, FW 11.0 mm), same locality as for holotype, coll. V. Soarimalala and M. Raheriarisena, 8 February 2002 (FMNH 7579) (GenBank MT 749741); subadult ♀♀ (CW 35.5, CL 25.9, CH 11.4, FW 11.4 mm, CW 30.6, CL 21.3, CH 9.8, FW 9.8 mm, CW 27.5, CL 21.6, CH 9.3, FW 9.3 mm), same collection data as for holotype (FMNH 7589.2) (GenBank MT 749745); subadult J (CW 35.7, CL 25.6, CH 10.8, FW 10.8 mm), same details as holotype (FMNH 7589.3) (GenBank MT 749746); same locality as for holotype, coll. S. M. Goodman, 16 October 2001 (FMNH 7590) (GenBank MT 749744); subadult ♀ (CW 22.0, Cl 17.1 mm; CH 7.5, FW 7.5 mm), same locality as for holotype, coll. S. M. Goodman, 13 October 2001 (FMNH 7592) (GenBank MT 749743).	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	diagnosis	Diagnosis As for the genus.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	description	Description Based on holotype, adult J. Carapace outline transversely oval, moderate height (CH / FW 1.0) gently inflated, CH equal to FW; front broad (FW / CW = 0.3), deflexed covering part of antennulular fossae; dorsal surface with faint carinae in anterolateral, posterolateral regions, otherwise smooth. Anterolateral margin between exorbital, epibranchial teeth convex, granulated, lacking intermediate tooth; epibranchial tooth reduced to small granule, close to exorbital tooth, positioned in line with postorbital margin; lateral margin evenly curved outward, lined with granules, posterior end curving inward, not continuous with posterolateral margin; postfrontal crest faint, incomplete, not traversing entire carapace, ending before meeting epibranchial teeth, epigastric crests faint, in advanced position, in line with postorbital margin; postorbital crests, cardiac, urogastric sulci faint, semicircular sulcus deep, cervical sulcus faint, short, not meeting postorbital crest (Figure 1 (a )). Suborbital, subhepatic regions of branchiostegite with faint granules; pterygostomial region smooth except for granules along epimeral (longitudinal) sulcus; vertical sulcus on branchiostegite curved, granular, running downward from base of epibranchial tooth to epimeral sulcus (Figure 2 (a )). Epistomial tooth triangular, deflexed, edges smooth (Figure 2 (a )). Mandibular palp terminal article bilobed; anterior lobe on terminal article conspicuous, medium-sized (MPAL / MPTA = 0.4) (Figure 2 (i )). Exopod of third maxilliped reaching to lower half of merus, exopod with short flagellum (less than half merus length), ischium of third maxilliped with faint shallow vertical sulcus (Figure 2 (g, h )). Sternal suture S 1 / 2 short, faint; S 2 / 3 deep, completely traversing sternum; S 3 / 4 broad, U-shaped, deepest at edges, not meeting anterior margin of sternopleonal cavity; S 4 / 5 meeting pleon at level of telson / PL 6 suture; S 6 / 7 meeting midpoint of lateral margin of PL 6; episternal sulci S 4 / E 4, S 5 / E 5, S 6 / E 6, S 7 / E 7 absent, smooth (Figure 1 (b )). Left chela larger, proximal half of cutting edge of fixed finger (pollex of propodus) with one large molar flanked by small teeth, lined with small teeth distally; cutting edge of movable finger (dactylus) lined with small teeth, lacking large teeth (Figure 2 (c )); right chela slightly smaller than left chela, dentition same as left chela (Figure 2 (b )); lower margin of propodus of both chelae conspicuously indented medially (Figure 2 (b, c )). Inner margin of cheliped carpus flattened, broad, distal tooth on inner margin small, pointed; proximal tooth smaller, acute, small granules between teeth, and following proximal tooth (Figure 2 (f )). Inferior margins of cheliped merus both lined with small, rounded teeth, distal tooth largest; cheliped ischium margins smooth, rounded (Figure 2 (d, e )). Ambulatory legs P 2 – 5 short (ΣP 2 – 5 / CW = 5.4) (Figure 1 (a); Table 1). Pleon slim, lateral margins taper distally, widest at PL 3, narrowest at telson / PL 6 suture; PL 6 relatively broad; telson outline with straight lateral margins, apex rounded (Figure 1 (b )). G 1 TA medium length (G 1 TA / G 1 SA = 0.3), slim, basal two-thirds directed outward, distal onethird curving upward, tapering to blunt open tip; G 1 TA ventral side with faint sulcus between G 1 TA and G 1 SA (Figure 3 (a )); G 1 TA dorsal side with broad, trapezoid dorsal membrane (DM) at G 1 TA-G 1 SA junction; DM superior margin arrowhead-shaped, inferior margin U-shaped, lateral margin short, mesial margin elongated (Figure 3 (b )); G 1 SA lacking raised rounded shoulder on external margin near G 1 TA-G 1 SA junction (Figure 3 (a – b )); G 2 TA distinctly elongated, flagellum-like (G 2 TA / G 2 SA = 0.85) with pointed tip (Figure 3 (d )). Size Medium-sized species, largest known specimen CW 38.5 mm, pubertal moult between CWs 36 and 38 mm. Colour Unknown in life. Preserved specimens are uniformly pale brown.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	distribution	Distribution Vichai cyanalepou sp. nov. is only known from the type locality in the Marojejy National Park in the Sava Region of northern Madagascar.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	materials_examined	Type locality Madagascar, Sava Region, Marojejy National Park, 11.5 km SE of Doany, in tributary of Bemanivy River near junction with Bemanavy River, 810 m ASL (14.4267 ° S, 49.60833 ° E). Habitat and ecology The rainforests of Marojejy National Park in Sava Province are the northernmost part of this World Heritage Site, which comprises a chain of six national parks distributed in the highlands of eastern Madagascar stretching from Marojejy to as far south as the Anosyennes Mountains in the south-east of the island. This extensive World Heritage Site protects some of the last intact primary rainforest in Madagascar, and each park within it supports a rich biodiversity, including threatened species. The eastern mountains in Madagascar receive a high annual rainfall that supports humid rainforest at lower elevations (500 – 1200 m ASL), drier sclerophyllous mountain forest (1200 – 1600 m ASL), and montane forest on the highest slopes (1600 – 2064 m ASL). Marojejy National Park is a part of the Madagascar Eastern Highlands freshwater ecoregion (FEOW 581; Abell et al. 2008), and the new species was collected from under rocks on the bed of a tributary of the Bemanivy River in the mostly undisturbed upper reaches of the lowland forest belt of the park. Interestingly, V. cyanalepou sp. nov. was collected together with six subadult specimens of Hydrothelphusa madagascariensis (A. Milne-Edwards, 1872) (FMNH 7589.1, Genbank MT 749717), which is, therefore, syntopic with the new species.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	etymology	Etymology The species name ′ cyanalepou ̍ is derived from Greek and is a combination of two words ′ cyan ̍ for dark blue and ′ alepou ̍ for fox, a reference to the dark blue shirts that Leicester City play in, and the club mascot the fox. The specific epithet is used as a Latin noun in genitive singular and is treated as masculine.	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
503B8E366769B14AFE65A01AECAF5D28.taxon	discussion	Remarks Vichai cyanalepou sp. nov. was referred to as ′ New genus E ̍ in the phylogenetic tree of Cumberlidge et al. (2020, fig. 1). The seven specimens sequenced in that study (described here and listed under Material and methods above) are all closely related to each other (with short branch lengths). These specimens comprise a genetically separate lineage from the nearest sister clade consisting of specimens labelled ′ New genus C ̍, ′ New genus D ̍, ′ FMNH 13935 ̍ and ′ Foza raimundi ̍ in Cumberlidge et al. (2020, fig. 1). Subsequently, ′ New genus C ̍ was identified using molecular and morphological evidence as Crosnautes ranomafana Cumberlidge et al. (2021) (see Cumberlidge et al. 2021), ′ New genus D ̍ was identified as a second species of this genus, C. alainus Cumberlidge et al. (2021) (see Cumberlidge et al. 2021), and ′ Foza raimundi ̍ (FMNH 7438) and FMNH 13935 were both confirmed as Foza raimundi Reed and Cumberlidge (2006) (Cumberlidge et al. 2020, fig. 1). Vichai cyanalepou sp. nov. is the fourth genus and fourth species to be recorded from Marojejy National Park, the other taxa being Hydrothelphusa madagascariensis, Marojejy longimerus Cumberlidge et al. (2002) and Foza raimundi (see Cumberlidge and Sternberg 2002; Cumberlidge et al. 2004; Reed and Cumberlidge 2006). Two of these four taxa (Marojejy longimerus and Vichai cyanalepou sp. nov.) are endemic to Marojejy National Park. Earlier reports of Skelosophusa eumeces Ng and Takeda (1994), occurring in the Marojejy National Park by Cumberlidge et al. (2004) were misidentifications and are not supported by the present study, as those specimens (FMNH 7579, FMNH 7590, and FMNH 7592) are reidentified here as belonging to Vichai cyanalepou sp. nov. See above for comparisons of V. cyanalepou sp. nov. with Skelosophusa and Hydrothelphusa. Vichai cyanalepou sp. nov. can be distinguished from F. raimundi mainly by the G 1 TA which is glabrous on the dorsal side in V. cyanalepou sp. nov. (Figure 3 (b )), vs a G 1 TA that is heavily setose on the dorsal side in F. raimundi (Leever et al. 2022, fig. 7 (h )). Furthermore, a lack of setae on the superior part of the pterygostomial region and on the anterior subpleonal cavity in V. cyanalepou sp. nov. (Figure 2 (a )) distinguishes it from F. raimundi where both of these regions are distinctly setose (see Leever et al. 2022, fig. 7 (c )).	en	Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B., Leever, Ellen M. (2023): Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar. Journal of Natural History 57 (5 - 8): 463-474, DOI: 10.1080/00222933.2023.2192431, URL: http://dx.doi.org/10.1080/00222933.2023.2192431
