taxonID	type	description	language	source
A6458955FFD96622D244FDEAFDF472DF.taxon	materials_examined	— Type (as given in protologue): “ In Hispania loco non notato (Herb. Pavon) ”. — Lectotype (designated by Sánchez-Mata et al. [1994]): Hispania, Pavon s. n. (Fig. 1 [G]).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD96622D244FDEAFDF472DF.taxon	discussion	REMARKS The lectotype shows two large flowering stems and a single sterile stem. The flowering stems are either not widened at the tip or only slightly widened. The widening observed on one of the stems appears to result from dessication or flattening of this stem. The leaves of the flowering stems are narrowly spatulate. As in the population that lives at high altitude, all the leaves of the sterile stems of the type are broadly spatulate. The capitula are very poorly preserved and the bracts were damaged during dissection by botanists. It is thus impossible to know how many external bracts were originally present. The morphological characteristics listed in Table 2 are essential to determine hybrid characters in Santolina ’ s populations, however the poorly preserved capitula did not allow a complete examination of Boissier’s original material. Therefore, it is not possible to determine if the lectotype of S. oblongifolia has hybrid characteristics or not. However, the morphological characteristics of this sample in the G-BOISS herbarium suggests that it was probably collected at an altitude below 1800 m in a mixed population including S. oblongifolia, S. rosmarinifolia subsp. rosmarinifolia and their putative hybrids. Rivas Goday (1955) considered that probably Pavon collected the original material of S. oblongifolia in locations close to Casatejada (Cáceres). Casatejada is located south of Central System, at not more than 300 m, outside the altitudinal range of this taxon. The present evaluation of the original material of S. oblongifolia suggests that the lectotype does not match with the current use of the name.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFDB6624D1DDFE4BFE8677E7.taxon	materials_examined	— Type (as given in protologue): “ In Hispania centrali regione montana: in monte Sierra de Gredos et in provinciae Avilensi prope Servanillos, Cutanda et Ysern! (1857). Floret Julio ”. — Lectotype (here designated): Spain, Sierra de Gredos, VI. 1855, Ysern s. n. (COI-WILLK 00035956; Fig. 2).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFDB6624D1DDFE4BFE8677E7.taxon	discussion	REMARKS De Candolle (1837) included S. canescens and S. viscosa into the circumscription of the genus Santolina L., and reduced S. pectinata Lag. as a variety of S. pectinata Benth. (nom. illegit.). The author retained the epithet “ pectinata ” for a variety α, whereas he conserved the type of S. pectinata Lag. for a variety β. He published var. β. heterophylla based on plants from the locus classicus of S. pectinata. He also cited S. pectinata Lag. as a synonym of this name, adding a short note as diagnosis “ foliis infimis et junioribus linearibus indivisis margine tuberculatis ”. Santolina pectinata Benth., non Lag. var. β. heterophylla DC., is an invalid name (ICBN: arts 53.1, 26.2; Mcneill et al. 2006). Furthermore, Willkomm & Cutanda in Willkomm (1859), and Willkomm in Willkomm & Lange (1865) ignored the variety β. heterophylla published by De Candolle (1837). Willkomm & Cutanda in Willkomm (1859) published a new taxon, S. heterophylla, based on a specimen from Sierra de Gredos. Willkomm in Willkomm & Lange (1865) reduced S. heterophylla Wilk. & Cut. to a variety of S. rosmarinifolia, but he omitted Cutanda as author of the epithet. He cited S. pectinata Lag. as a synonym of “ variat 1 ” of S. rosmarinifolia L. var. β heterophylla Willk., whereas Willkomm (1893) changed the original concept and accepted S. rosmarinifolia var. β heterophylla Willk., f. 1 Willk. as synonym of S. pectinata. He placed this variat in the type locality of S. pectinata, and in Sierra de la Sagra (COI-WILLK 00035964, MPU, P 00752621 from “ La Puebla de Don Fadrique et montem La Sagra ”). Willkomm misspelled the locality for the lectotype of S. heterophylla; the correct name is “ Serranillos ” (Cerveira Rodrigues Gomes 2000), and it will be used hereafter in this paper. Two collections of Isern were found. In the first one, labelled with the name “ S. oblongifolia ”, from Puerto del Pico (MA 126799), the date is only indicated as “ August ”. The other specimen (COI-WILLK 00035956), labelled by Willkomm with the names S. oblongifolia β ceratophylla and S. heterophylla was gathered by Isern in July 1855 in Sierra de Gredos. Presumably, Willkomm changed 1855 into 1857 during transcription of the original label. The lectotype (COI-WILLK 00035956; Fig. 2) has six flowering stems and various sterile stems. The sheet has two labels in the centre of the lower margin with the following notation in black ink: Label 1: “ Herbarium Willkommi [printed] // Santolina rosmarinifolia Mill. [underlined] var?. sericea [underlined] Wk. = S. heterophylla Wk. [underlined] et Cut. // Sierra de Gredos [Willkomm’s handwritten notation] // Legit [printed] Ysern // Julio 1855 [Willkomm’s handwritten notation] ”; Label 2: “ Santolina oblongifolia Boiss. [underlined] β ceratophylla [underlined] Wk. [Willkomm’s handwritten notation] ”. This specimen has two revision labels by D. Sánchez-Mata and S. Sardinero with the following notation in black ink: Label 1: “ Santolina oblongifolia Boiss var. ceratophylla Willk. in Willk. & Lange // Typus [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”; Label 2: “ Santolina oblongifolia Boiss. [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”. This specimen displays all the characteristics listed in the protologue (Appendix 1). The lectotype clearly shows intermediate characteristics between S. oblongifolia and S. rosmarinifolia subsp. rosmarinifolia. The relationship between S. heterophylla, S. rosmarinifolia subsp. rosmarinifolia and S. oblongifolia has been cited by Willkomm (1859) in the protologue: “ Specie proxima S. rosmarinifoliae Mill., a cujus formis numerosis foliis eximie heteromorphis planis sericeis statim distinguitur. Calathiorum florumque structura a S. rosmarinifolia non differre videtur, sed propter foliorum formam ab omnibus Santolinis valde discrepantem hanc stirpem ad illam speciem referre non audeo. S. oblongifolia Boiss. foliis planis sericeis pariter gaudens ramis sterilibus elongatis, foliis multo minus difformibus aliaque ratione formatis a nostra differre videtur […] ”. The sheet A 00251592 has two labels in the lower left margin: Label 1: “ HERBARIUM GRAELSIANUM // PLANTAE CASTELLANAE [printed] // Santolina heterophylla Wk. // in saxosis Sierra de Gredos et Puerto del Pico prope riventos // Julio [Graells’ handwritten notation] ”; and Label 2 (printed): “ HERBARIUM OF GEORGE CURLING JOAD. PRESENTED BY THE ROYAL GARDENS, KEW, 1882 ”. The labels do not show the collector and year. The notation of the labels does not match with the lectotype, however, this specimen is the counterpart of the type material of this name. Carrasco et al. (2002) discuss the value of the collections of the Real Colegio Alfonso XII historical herbarium of San Lorenzo del Escorial (Madrid, Spain). They argue that the Graells’ collection was used by Willkomm & Lange as part of the material examined for the Prodromus florae hispanicae. However, S. heterophylla was published several years before the publication of the species corology of the genus Santolina in Prodromus florae hispanicae (1865). There is not enough evidence to justify the choice of this material as original material of S. heterophylla.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFDD6624D191FB73FE5275ED.taxon	materials_examined	— Type (as given in protologue): “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ”. — Lectotype (here designated): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (COI-WILLK 00035955 [Fig. 3]; iso-, MANCH, MPU, P (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the right).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFDD662AD244FF46FEB975ED.taxon	materials_examined	— Type (as given in protologue): “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ”. — Lectotype (here designated): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (COI-WILLK 00035989 [Fig. 3]; isolecto-, A 00251664 (specimen to the left), HEID 702282 (specimen to the left), MPU (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the left]).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFDD662AD244FF46FEB975ED.taxon	discussion	REMARKS These names were published by Willkomm in Willkomm & Lange (1865) as S. oblongifolia α obtusifolia and S. oblongifolia β ceratophylla. The author did not mentioned the rank of “ α ” and “ β ” in the introductory chapter of Prodromus Florae Hispanicae. However, Willkomm (Willkomm & Lange 1865: 81) cited “ S. var. vulgaris S. rosmarinifoliae ”, indicating the rank of S. rosmarinifolia L. α vulgaris Boiss.; therefore the rank of “ α ” and “ β ” can clearly be established as variety. In addition, Lange in Willkomm & Lange (1880), cited the varietal status of both names. The varieties “ obtusifolia ” and “ ceratophylla ” are from the same locus classicus: “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ” (Willkomm in Willkomm & Lange 1865). Willkomm also mentioned “ Serranillos ” in the protologue of S. heterophylla. This indicates that Willkomm granted a varietal status to the intrapopulational variation. The specimens COI-WILLK 00035955 and COI-WILLK 00035989 of Willkomm’s herbarium are on the same sheet (Fig. 3). This sheet has two labels in the lower left-hand margin with the following notation in black ink: Label 1 (Bourgeau’s handwritten notation): “ E. Bourgeau – Pl. d’Espagne 1863 // 2540. Santolina oblongifolia Boiss. Diagn. / Pl. Or. Ser. 2, fasc. 3, p. 18 (Planta herbaria Paroniani, cujus patria nodium involucrate) // (J. Gay) // Rochers de la Sierra de Gredos au Puerto del Pico // 8 et 25 juill ”; Label 2 (Willkomm’s handwritten notation): “ Santolina oblongifolia Boiss. [underlined] // 1. var. α obtusifolia / 2. var. β ceratophylla ”. Voucher specimens from the locus classicus and with the number “ 2540 ” were found in A 00251664, G, HEID 702282, LY (2 sheets), MA 720935, MPU (3 sheets), P (3 sheets) and W (3 sheets), but they do not include a revision label of Willkomm with the epithets “ obtusifolia ” and “ ceratophylla ”. The collection to the left, COI-WILLK 00035955, comprises two samples from the same gathering, labelled with the name “ Santolina oblongifolia var. α obtusifolia ” (lectotype here designated; Fig. 3). One of them (sample to the right) has three flowering stems and a small sterile stem, and the other (sample to the left) is a robust sample with four flowering stems and several vegetative stems. The lectotype has two revision labels by Sánchez-Mata and Sardinero with the following notation in black ink: Label 1: “ Santolina oblongifolia Boiss. var. obtusifolia Willk. in Willk. & Lange // Lectotype [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”; Label 2: “ Santolina oblongifolia Boiss. [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”. Both samples have spatulate leaves, but the flowering stems are strongly thickened above and the capitula are big (Fig. 3). The literature does not show the publication of this lectotype by Sánchez-Mata & Sardinero. There is another specimen with the name “ Santolina oblongifolia var. α obtusifolia ” in W herbarium. The sheet (W 0026403; Fig. 4) has a different collection date (Sierra de Gredos, Leg. Bourgeau 3. VII. 1863, Cour. Willkomm), and the number “ 2540 ” is not shown. This specimen does not show intermediate characteristics between S. oblongifolia and S. rosmarinifolia subsp. rosmarinifolia. All the characteristics fit with those listed in the protologue (Appendix 1). Jordan & Fourreau (1869) published a new taxon (S. sericea) based on plants from the locus classicus of the varieties obtusifolia and ceratophylla. However, they did not cite it in the protologue. They also published a detailed diagnosis of S. sericea (Appendix 1). Furthermore, figure 321, table 240 (Jordan & Fourreau 1869) shows a robust sample of S. sericea. Two sheets are preserved in the Jordan’s herbarium (LY) with the epithets “ sericea ”. The records indicate that the sample Jordan used to illustrate this name was split into two units and mounted on two sheets. They were labelled as “ 1 ère feuille ” (Fig. 5) and “ 2 ème feuille ” (Fig. 6, specimen to the right). The sheet designated “ 1 ère feuille ” (Jordan’s handwritten notation in graphite in the lower left margin) has a sample, in the centre of the sheet, with three flowering stems, two fragments of flowering stems and three sterile stems. There is also a large, isolated sterile stem to the right of the sample (Fig. 5) [this sample is the portion to the right-hand of the original specimen]. A label in the central lower margin indicates: “ Santolina sericea [underlined] // Jord. et Fourr. // Lectotypus [underlined] // D. Rivera // 18 / 10 / 1984 [D. Rivera’s handwritten notation] ”. This lectotype was not published by D. Rivera. The sheet designated “ 2 ème feuille ” (Jordan’s handwritten notation in the central lower margin in graphite) has two samples (Fig. 6). The sample to the right is the portion to the left-hand-side of the original material. It has a decumbent branch with eight flowering stems. The sample to the left (representing a different gathering) has a small branch with three flowering stems. This sample shows clear intermediate characteristics (linear and narrowly spatulate leaves, peduncles strongly thickened above and umbilicate capitula) between S. oblongifolia and S. rosmarinifolia subsp. rosmarinifolia. The lectotype is the specimen from the sheet designated “ 1 ère feuille ” (Fig. 5 [sample to the left]). It has three flowering stems and three other short vegetative stems. Furthermore, Nyman (1879) cited the present Bourgeau’s collection as S. oblongifolia. Willkomm in Willkomm & Lage (1865), in the protologue of the variety ceratophylla, cited S. heterophylla as a synonym of this name. There are two possibilities regarding S. oblongifolia var. β ceratophylla: – Option 1: Willkomm created a new name, at varietal rank for S. heterophylla (even if he did not use the same final epithet). This theory is supported by the following evidences: 1) Willkomm & Cutanda in Willkomm (1859) in the protologue of S. heteroplylla wrote “ An varietas S. oblongifoliae? ”, indicating that the authors had doubts about the species status of S. heterophylla; 2) both names have the same locus classicus; 3) the single specimen cited in the protologue of S. heterophylla (COI-WILLK 0035956) bears the both names; 4) Willkomm (1865) cited S. heterophylla as synonym of the variety “ ceratophylla ”. If this is the case, the two names (S. heteroplylla and S. oblongifolia var. β ceratophylla) have the same lectotype (COI-WILLK 0035956, here designated; see ICBN: art. 7.3 and 7.4; McNeill et al. 2006), therefore the both names are homotypic synonyms. – Option 2: Willkomm created a new taxon at varietal rank for S. heterophylla. This theory is supported by the following evidences: 1) the author published a more accurate diagnosis with elements of the diagnosis of S. heterophylla; 2) he changed “ spathulato-linearibus ” (S. heterophylla) to “ cuneato-linearia ” (S. oblongifolia var. β ceratophylla); 3) the author included new elements in the diagnosis: “ folia superiora aut summa integerrima, omnia virentia ”; and 4) Willkomm studied more specimens and redefined the concept of S. heterophylla. If the option 2 is correct, the specimen to the right (Fig. 3; COI-WILLK 00035989) is the lectotype (here designated) of S. oblongifolia var. β ceratophylla. The sample has two flowering stems and a sterile stem. This specimen has two revision labels by Sánchez-Mata & Sardinero with the following notation in black ink (the labels are placed under one of the specimens of the variety “ obtusifolia ”, probably a handling error): Label 1: “ Santolina oblongifolia Boiss. var. ceratophylla Willk. in Willk. & Lange // Typus [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”; Label 2: “ Santolina oblongifolia Boiss. [handwritten] // Coimbra [printed] 1993 IX [handwritten] ”. However, the specimen COI-WILLK 00035956 (Fig. 2, see lectotypification of S. heterophylla) was also selected by Sánchez-Mata & Sardinero as typus of Santolina oblongifolia var. ceratophylla. The specimens display characteristics consistent with those listed in the protologue (Appendix 1) and show characteristics intermediate between S. oblongifolia and S. rosmarinifolia subsp. rosmarinifolia. AORG considers that option 2 is the most appropriate.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD3662AD3B6FF46FC0D70C2.taxon	materials_examined	— Type (as given in protologue): “ Hispania central. Sierra de Gredos, prope Bohoyo ”. — Lectotype (here designated): fig. 320, tab. 239: 12, 13 (Jordan & Fourreau 1869), cf. ICBN: arts 8.1 and 8 A. 1 (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD3662AD3B6FF46FC0D70C2.taxon	discussion	REMARKS Specimens from this locality were not found in the Lyon Herbarium (LY). Four specimens from the locus classicus were found in G, P (00752616, Fig. 8; 00752617, Fig. 9) and W (0026402, Fig. 7) herbaria. They are labelled as “ Sierra de Gredos, bords de la rivière près Bohoyo, à la base de la Sierra de Gredos, 9. VII. 1863, Bourgeau s. n. They do not show the epithet “ lobata ”. Probably, these materials were not examined by the authors of S. lobata. The figure 320 (Jordan & Fourreau 1869) has been selected as a lectotype of this name, in agreement with the articles 8.1 and 8 A. 1 of the ICBN (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD3662CD244FC4EFD507744.taxon	materials_examined	— Type (as given in protologue): “ Entre Baños de Montemayor y Garganta de Béjar; al pie de la Sierra, saliendo de la Garganta, abundante ”. — Neotype (here designated): Spain, Salamanca: Béjar, La Garganta, 40 ° 19 ’ 48 ’’ N, 5 ° 49 ’ 10 ’’ W, 1000 m, granites, A. O. Rivero-Guerra s. n. (SEV 249072).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD3662CD244FC4EFD507744.taxon	discussion	REMARKS Pau (1900) indicated that the plants from these localities show intermediate morphological characteristics between S. rosmarinifolia and the varieties “ obtusifolia ” and “ ceratophylla ”: “ Las hojas de los tallos floríferos, parecidos algo á los de la S. rosmarinifolia L., tan abundante en Castilla, son como los de la variedad ceratophylla Wk.; el color de todas la planta como en la obtusifolia Wk. ”. The type specimen of this name has been lost or destroyed and duplicates of the original material were not distributed. Consequently, no duplicate material is known, and unfortunately, no safe interpretation of this name can be provided by examination of the type specimen. No voucher specimen from this locality was found in more than 40 European Herbaria consulted. Therefore, in agreement with articles 9.6, 9.9 and 9.11 of the ICBN (McNeill et al. 2006), a neotype for S. oblongifolia Boiss. f. intermedia is designated.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ In Hispania loco non notato (Herb. Pavon) ”. — Lectotype (designated by Sánchez-Mata et al. [1994]): Hispania, Pavon s. n. (G).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ In Hispania centrali regione montana: in monte Sierra de Gredos et in province Avilensi prope Servanillos, Cutanda et Ysern! (1857). Floret Julio ”. — Lectotype (here designated): Spain, Sierra de Gredos VI. 1855, Ysern s. n. (COI-WILLK 00035956).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ”. — Lectotype (here designated): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (COI-WILLK 00035955; isolecto-, MANCH, MPU, P (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the right).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ Hispania central. Sierra de Gredos ad Puerto del Pico ”. — Lectotype (designated here): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (LY [Fig. 5, sample to the left]). Isolec- LY (Fig. 6, specimen to the right), cf. article 8.3 of the ICBN (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ Hispania central. Sierra de Gredos, prope Bohoyo ”. — Lectotype (here designated): fig. 320, tab. 239: 12, 13 (Jordan & Fourreau 1869), cf. articles 8.1 and 8 A. 1 of the ICBN (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFD76630D3E4FD6CFD5670DA.taxon	materials_examined	— Type (as given in protologue): “ Entre Baños de Montemayor y Garganta de Béjar; al pie de la Sierra, saliendo de la Garganta, abundante ”. — Neotype (designated here): Spain, Salamanca: Béjar, La Garganta, 40 ° 19 ’ 48 ’’ N, 5 ° 49 ’ 10 ’’ W, 1000 m, granites, A. O. Rivero-Guerra s. n. (SEV 249072).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ In Hispania loco non notato (Herb. Pavon) ”. — Lectotype (designated by Sánchez-Mata et al. [1994]): Hispania, Pavon s. n. (G).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ In Hispania centrali regione montana: in monte Sierra de Gredos et in province Avilensi prope Servanillos, Cutanda et Ysern! (1857). Floret Julio ”. — Lectotype (here designated): Spain, Sierra de Gredos VI. 1855, Ysern s. n. (COI-WILLK 00035956).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ”. — Lectotype (here designated): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (COI-WILLK 00035955; isolecto-, MANCH, MPU, P (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the right).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.! Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v. s.) ”. — Lectotype (here designated): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (COI-WILLK 00035989; isolecto-, A 00251664 (specimen to the left), HEID 702282 [specimen to the left], MPU (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the left).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ Hispania central. Sierra de Gredos, prope Bohoyo ”. — Lectotype (here designated): fig. 320, tab. 239: 12, 13 (Jordan & Fourreau 1869), cf. articles 8.1 and 8 A. 1 of the ICBN (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ Hispania central. Sierra de Gredos ad Puerto del Pico ”. — Lectotype (designated here): Spain, Rochers de la Sierra de Gredos au Puerto del Pico, 8 - 25. VII. 1863, Bourgeau 2540 (LY [Fig. 5, sample to the left]). Isolec- LY [Fig. 6, specimen to the right], cf. artticle 8.3 of the ICBN (McNeill et al. 2006).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFC96630D137FBF0FBED75ED.taxon	materials_examined	— Type (as given in protologue): “ Entre Baños de Montemayor y Garganta de Béjar; al pie de la Sierra, saliendo de la Garganta, abundante ”. — Neotype (designated here): Spain, Salamanca: Béjar, La Garganta, 40 ° 19 ’ 48 ’’ N, 5 ° 49 ’ 10 ’’ W, 1000 m, granites, A. O. Rivero-Guerra s. n. (SEV 249072).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFCB6632D13EFDCDFBD27085.taxon	description	(Fig. 10)	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFCB6632D13EFDCDFBD27085.taxon	diagnosis	A S. x oblongifolia Boiss., floriferis simplicibus caulibus 14.1 - 32.1 (47.5) cm longis (vs 16.1 - 54.1 cm longis), pedunculo 0.5 - 1.2 (2.2) mm, haud incrassato (vs (0.9) 1.1 - 2.0 mm, crasso), foliis spathulatis (vs spathulatis, anguste spathulatis vel linearis), (0) 1 - 8 (13) lobulis ellipticis (vs (0) 1 - 25 ellipticis vel linearibus), foliis basalibus fasciculatis cum 8 - 18 (52) lobis (vs (8) 14 - 80 (140) lobis), sterilibus caulibus 3.6 - 8.8 (21.1) cm in longitudine (vs 5.1 - 28.6 cm in longitudine), capitulo (5.5) 6.5 - 12.6 (14.08) × (2.7) 3.6 - 6.6 mm, haud umbilicato (vs (8.8) 10.3 - 16.6 × 4.4 - 6.7 (10.8) mm umbilicato), involucralibus trifariis bracteis (vs involucralibus 3 - 4 fariis bracteis) praecipue differt.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFCB6632D13EFDCDFBD27085.taxon	materials_examined	TYPUS. — Spain. Ávila, La Mira, 40 ° 15 ’ 59 ’’ N, 5 ° 10 ’ 28 ’’ W, 2200 m, granites, 20. VII. 1999, Rivero-Guerra s. n. (holo-, SEV 249071). PARATYPI. — Spain. Ávila, Canchal Negro, 40 ° 20 ’ 88 ’’ N, 5 ° 40 ’ 27 ’’ W, 2000 m, granites, 19. VII. 1999, Rivero-Guerra, s. n. (SEV 249072). — Salamanca, Béjar, La Garganta, 40 ° 19 ’ 48 ’’ N, 5 ° 49 ’ 10 ’’ W, 1000 m, granites, 10. VII. 1995, Rivero-Guerra s. n. (SEV 249073). — El Calvitero, 40 ° 17 ’ 16 ’’ N, 5 ° 44 ’ 18 ’’ W, 2360 m, granites, 19. VII. 1999, Rivero-Guerra s. n. (SEV 249074).	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
A6458955FFCB6632D13EFDCDFBD27085.taxon	description	DESCRIPTION Plant 35 - 70 × 18 - 60 cm, with 1 - 9 branches stock, glaucous, with indument sericeous. Flowering stem 14.1 - 32.1 (47.53) cm long, 0.5 - 1.2 (2.2) mm in diameter, simple, usually not solid. Peduncle 23.1 - 92.6 (142.4) mm long, not thickened above. Sterile stem 3.6 - 8.8 (21.1) cm long. Leaf 5.4 - 20.8 × 2.1 - 6.2 mm, spatulate, plane, with bi or trilobulate apex, usually pinnatisect or pinnatipatite, with (0) 1 - 8 (13) lobes of 0.4 - 4.1 (5.0) mm long; basal and fascicular leaves, usually pinnatifid or pinnatipartite. Lobes elliptical, along ⅓ upper, generally with obtuse-mucronate or acute-mucronate apex. Upper leaf 5.3 - 13.4 (15.8) × 0.8 - 2.1 (2.4) mm, narrowly elliptical, entire, with a beaked apex. Capitulum (5.5) 6.5 - 12.6 (14.1) × (2.7) 3.6 - 6.6 mm, usually campanulate, not umbilicate, with the flowers covering the capitulum. Receptacle 4.4 - 5.1 × 2.4 - 3.0 mm, hemispherical. Involucral bracts (1.7) 2.0 - 3.2 (3.7) × 0.7 - 1.6 (1.7) mm, in three rows, carinate, with an apical scarious appendage 0.2 - 0.8 × 0.2 - 2.5 (2.9) mm, generally decurrent along the upper ⅓ or ½; outer and middle bracts usually triangular or ovate, inner bracts usually ovate or elliptical. Interseminal bracts (1.8) 1.9 - 3.1 × 0.8 - 2.0 mm, usually elliptical or oblong, rarely obovate, with truncate or rounded apex, with erose or dentate margin. Corolla 3.5 - 5.1 mm long. Anthers 2.5 - 3.8 mm long. Style 3.0 - 5.1 mm long. Corolla tube 1.6 - 3.0 mm long. Corolla aperture 0.8 - 1.5 mm long. Corolla lobes 0.4 - 1.1 mm long. Achene 1.2 - 2.3 × 0.4 - 0.9 mm. Flowering from July to August.	en	Rivero-Guerra, Aixa O. (2012): Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss. Adansonia (3) 34 (1): 133-154, DOI: 10.5252/a2012n1a16, URL: http://dx.doi.org/10.5252/a2012n1a16
