taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C8746C4AFFE70A17FF1BF979FECBFE64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850651/files/figure.png	https://doi.org/10.5281/zenodo.7850651	FIGURE 5. An adult Desmognathus marmoratus from Laurel Creek (VA: Smyth). Note the indistinctiveness of the dorsal color pattern. Photo courtesy of T.W. Pierson (KSU).	FIGURE 5. An adult Desmognathus marmoratus from Laurel Creek (VA: Smyth). Note the indistinctiveness of the dorsal color pattern. Photo courtesy of T.W. Pierson (KSU).	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE70A17FF1BF979FECBFE64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850657/files/figure.png	https://doi.org/10.5281/zenodo.7850657	FIGURE 6. An adult Desmognathus marmoratus from Hickory Creek (NC: Henderson), from the isolated population (marmoratus G) in Hickory Nut Gorge. This phylogeographic sublineage of D. marmoratus contains significant genomic ancestry from D. intermedius and potentially other Pisgah-clade species (Pyron et al. 2022c). Photo courtesy of M.A. Seldes (UGA).	FIGURE 6. An adult Desmognathus marmoratus from Hickory Creek (NC: Henderson), from the isolated population (marmoratus G) in Hickory Nut Gorge. This phylogeographic sublineage of D. marmoratus contains significant genomic ancestry from D. intermedius and potentially other Pisgah-clade species (Pyron et al. 2022c). Photo courtesy of M.A. Seldes (UGA).	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE70A17FF1BF979FECBFE64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850645/files/figure.png	https://doi.org/10.5281/zenodo.7850645	FIGURE 2. Map of the Lower New River drainage (HUC8:05050004) in West Virginia (WV) showing surveyed localities from our recent fieldwork (2020–2022), highlighting sites containing Desmognathus kanawha, D. marmoratus, no aquatic Desmognathus, and likely candidates for future surveys. Each area was searched for at least 1 person/hour, netting the effluent of disturbed riffle zones adequate to detect Shovel-nosed Salamanders.The New (Kanawha) River is indicated in dark blue, with tributaries in gray. In addition to Piney Creek which we consider a likely site (?), numerous additional creeks and streams in the region remain to be evaluated. Watershed data are projected at 1:24,000 from the National Hydrography Dataset (https://www. usgs.gov/national-hydrography/national-hydrography-dataset), accessed 7 November 2022. Literature record (lit.) from Wooten and Rissler (2011).	FIGURE 2. Map of the Lower New River drainage (HUC8:05050004) in West Virginia (WV) showing surveyed localities from our recent fieldwork (2020–2022), highlighting sites containing Desmognathus kanawha, D. marmoratus, no aquatic Desmognathus, and likely candidates for future surveys. Each area was searched for at least 1 person/hour, netting the effluent of disturbed riffle zones adequate to detect Shovel-nosed Salamanders.The New (Kanawha) River is indicated in dark blue, with tributaries in gray. In addition to Piney Creek which we consider a likely site (?), numerous additional creeks and streams in the region remain to be evaluated. Watershed data are projected at 1:24,000 from the National Hydrography Dataset (https://www. usgs.gov/national-hydrography/national-hydrography-dataset), accessed 7 November 2022. Literature record (lit.) from Wooten and Rissler (2011).	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE70A17FF1BF979FECBFE64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850643/files/figure.png	https://doi.org/10.5281/zenodo.7850643	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE40A10FF1BFE62FCEFF9B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850643/files/figure.png	https://doi.org/10.5281/zenodo.7850643	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE40A10FF1BFE62FCEFF9B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850659/files/figure.png	https://doi.org/10.5281/zenodo.7850659	FIGURE 7. An adult Desmognathus aureatus from Pruitt Creek (GA: Lumpkin), near the type locality. Note the brighter yellow or golden lichenous dorsal blotches. Photo courtesy of MAS.	FIGURE 7. An adult Desmognathus aureatus from Pruitt Creek (GA: Lumpkin), near the type locality. Note the brighter yellow or golden lichenous dorsal blotches. Photo courtesy of MAS.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE30A12FF1BF953FDFBFA27.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850661/files/figure.png	https://doi.org/10.5281/zenodo.7850661	FIGURE 8. A large (75mm SVL) adult Desmognathus intermedius (RAP2814) from Wycle Fork (NC: Haywood) in life. Note the faded lichenous dorsal patches muddled into more indistinct blotches. Photo courtesy of MAS.	FIGURE 8. A large (75mm SVL) adult Desmognathus intermedius (RAP2814) from Wycle Fork (NC: Haywood) in life. Note the faded lichenous dorsal patches muddled into more indistinct blotches. Photo courtesy of MAS.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE30A12FF1BF953FDFBFA27.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850663/files/figure.png	https://doi.org/10.5281/zenodo.7850663	FIGURE 9. An adult Desmognathus intermedius from Little Santeetlah Creek (NC: Graham). Note the uniformly muddled dorsal blotches forming an indistinct pattern. Photo courtesy of MAS.	FIGURE 9. An adult Desmognathus intermedius from Little Santeetlah Creek (NC: Graham). Note the uniformly muddled dorsal blotches forming an indistinct pattern. Photo courtesy of MAS.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
C8746C4AFFE30A12FF1BF953FDFBFA27.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7850643/files/figure.png	https://doi.org/10.5281/zenodo.7850643	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	FIGURE 1. Map of genetic, specimen-vouchered, and literature localities for the three species of Shovel-nosed Salamanders, from our collections (RAP and DAB) and Martof (1956), Jackson (2005), Kozak et al. (2005), Wooten and Rissler (2011), Beamer and Lamb (2020), and Pyron et al. (2022c). Type localities are indicated with inverted triangles, and the site indicated in grey is a dubious locality reported by Wooten and Rissler (2011). Our extensive fieldwork at this and numerous other sites in the headwaters of the New (Kanawha) River yielded no specimens. State abbreviations are GA—Georgia, SC—South Carolina, NC—North Carolina, TN—Tennessee, VA—Virginia, and WV—West Virginia.	2023-04-20	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
