taxonID	type	description	language	source
B07D7FC5250752D48742BC8001EFFC5C.taxon	description	Figures 8 D, 39 B, 61 A-C, 62	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 0.84 + / - 0.07; 0.81 - 0.88). In full-face view, head subovate, usually somewhat longer than broad, posterior cephalic margin roundly convex; lateral margin of head straight and tapering to front (CWb / CL: 0.31 + / - 0.01, 0.30 - 0.32). Eyes subelliptical, protruding, its border smoothly aligned to lateral margins of head (PoOC / CL: 0.08 + / - 0.01, 0.08 - 0.09). Mandibles triangular with six teeth. Clypeus carinate, anterior clypeal margin with rounded triangular projection (ClyL / GPD: 0.24 + / - 0.01, 0.23 - 0.25). Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL / CS: 0.36 + / - 0.04, 0.35 - 0.38). In lateral view, mesosoma without anterolateral margination, promesonotum convex, mesopropodeal suture shallowly impressed, propodeum rather angulate with slightly concave basal and declivitous surface (MW / ML: 0.19 + / - 0.02, 0.18 - 0.21, MPH / ML: 0.15 + / - 0.01, 0.14 - 0.15). In lateral view, petiole nodiform, higher than long, its short anterior face rounding gradually to the convex posterior faces. Head, especially anterior portion, finely reticulate-areolate, gaster and mesosoma finely imbricate. Mandibles finely reticulate-punctate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; three to four pairs on each side of lateral corner of propodeum; four pairs transversely arranged on petiolar posteromargin; ground pubescence delicate. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remainder of the segment dark brown to black.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	description	Description of major worker. Characteristics of minor workers, except: head as long as broad, with rather straight posterior and lateral borders (CS: 1.15 + / - 0.05, 1.12 - 1.20, CWb / CL: 0.84 + / - 0.01, 0.84 - 0.85). Eyes circular, placed dorsally next to lateral borders (PoOC / CL: 0.26 + / - 0.03, 0.23 - 0.27). Anterior clypeal margin forms a rounded rectangular lobe (ClyL / GPD: 0.89 + / - 0.05, 0.83 - 0.92). Antennal scape short, just reaching the occipital border (SL / CS: 0.70 + / - 0.02, 0.69 - 0.72). Mesosoma the same as minor worker.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	distribution	Distribution and biology. Camponotus andrianjaka is a very distinctive species that appears endemic to Bismarckia woodland, shrubland, Uapaca woodland, and savannah woodland in the southern part of Madagascar (Fig. 70 D). It can be found also on eucalyptus plantations and grassland. Altitude: 1300 - 1987 m.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	discussion	Discussion. At first sight, Camponotus andrianjaka may be confused with another new species, Camponotus MG 132, subgenus Camponotus Myrmonesites, but can be separated easily with mesonotum that is gradually rounded in dorsal view; petiole higher than long, its dorsolateral margin surmounted with white hairs; presence of ground pubescence; and gaster all black. In addition, C. MG 132 has been collected from rainforest and montane rainforest and absent from woodland.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	etymology	Etymology. This new species is named after the collector Andrianjaka Ravelomanana.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B07D7FC5250752D48742BC8001EFFC5C.taxon	materials_examined	Additional material examined. Province Fianarantsoa: Ampandravelo III Non Protected Area, 10.72 km NE Ranohira, - 22.53944, 45.51497, 869 m, shrubland (ARA) (CAS); Ampangabe III Non Protected Area, 21.26 km W Itremo, - 20.6125, 46.60883, 1412 m, savannah woodland (ARA) (CAS); Ampangabe VI Non Protected Area, 21.16 km W Itremo, - 20.61444, 46.6104, 1379 m, shrubland (ARA) (CAS); Ampangabe VII Non Protected Area, 21.2 km W Itremo, - 20.61417, 46.60989, 1420 m, shrubland (ARA) (CAS); Ampotoampoto II National Parc, 8.07 km NW Ilakaka, - 22.62806, 45.18843, 919 m, savannah woodland (ARA) (ATOL UCDC); Ampotoampoto III National Parc, 7.91 km NW Ilakaka, - 22.62944, 45.189, 919 m, savannah woodland (ARA) (ATOL voucher CAS); Antohatsahomby III Non Protected Area, 22.79 km NW Itremo, - 20.54806, 46.58599, 1499 m, Uapaca woodland (ARA) (CAS); Antohatsahomby V Non Protected Area, 22.63 km NW Itremo, - 20.56722, 46.57923, 1726 m, Uapaca woodland (ARA) (CAS); Isalo III National Parc, 12.02 km SW Ranohira, - 22.61583, 45.31084, 870 m, Bismarckia woodland (ARA) (CAS); Isalo IV National Parc, 12 km SW Ranohira, - 22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (ATOL USNM); Isalo IV National Parc, 12 km SW Ranohira, - 22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (CAS); Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, - 20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (ATOL CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	description	Figures 4 A-D, 13 B, 19 A, 51	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	description	Description of minor worker. Large-sized species. Absolute cephalic size (CS: 2.14 + / - 0.38, 2.00 - 2.33). In full-face view, head elongate (CWb / CL: 0.34 + / - 0.03; 0.32 - 0.35); posterior margin of head straight, lateral margin of head even straighter, feebly converging toward base of mandible. Eyes elliptical, sublateral, and placed next to the vertex (PoOC / CL: 0.08 + / - 0.02; 0.07 - 0.09). Mandibles triangular with six teeth. Clypeus strongly carinate throughout its length, with produced, angular, anterior margin, lateral corner angulate and median portion a rounded triangle (ClyL / GPD: 0.25 + / - 0.05, 0.22 - 0.28). Antennal scape longitudinally flattened, its base scarcely enlarged (SL / CS: 0.44 + / - 0.04, 0.42 - 0.45). In lateral view, mesosoma short and high, its dorsum smoothly arcuate, metanotal suture obsolete. Humeral angle rounded. Dorsal and declivitous face of propodeum difficult to distinguish and forming a rounded angle (MW / ML: 0.27 + / - 0.02; 0.26 - 0.28; MPH / ML: 0.21 + / - 0.02, 0.20 - 0.22). In lateral view, petiole squamiform with sharp edge, anterior face short and converging to the highest point of the petiole, posterior face straight. Ventral face of femora marginate laterally, the outer margin of tibia with long, whitish setae. Head and mesosoma coarsely reticulate-punctate, legs and gastral tergite finely imbricate. Mandible finely foveolate with sparse punctures. Whitish, suberect hairs present on occipital portion, to the anterior margin of eyes, and on posteroventral head surface, scarcely distributed on mesosoma dorsum but much closer on declivity; decumbent, spatulate, whitish hair on the ventral section of gena, below the antennal socket, on the posterolateral margin of clypeus, promesonotum dorsum, petiolar node, and gastral tergite; suberect and decumbent whitish hairs present on gastral segment. Entire body blackish except the mandible testaceous to reddish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	description	Description of major worker. With characteristics of minor workers, except: head more cordate (CS: 3.32 + / - 0.21, 2.94 - 3.55) (CWb / CL: 0.96 + / - 0.01, 0.95 - 0.99); lateral margins slightly convex and tapering anteriorly. Eyes elliptical, smaller compared to head size (EL / CS: 0.20 + / - 0.01; 0.20 - 0.21), placed dorsally next to posterior margin of frontal carina. Antennal scape short and not exceeding posterior margin of head (SL / CS: 0.81 + / - 0.04; 0.76, 0.88); mandible more robust. Mesosoma and pilosity same as minor worker.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	distribution	Distribution and biology. This new species is only known from Antsaraingy, a littoral forest, from 66 to 90 m in elevation, located in the northern portion of Madagascar (Fig. 70 A). Nests were found underground and inside termite mounds; four colonies were collected to represent this species.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	discussion	Discussion. Despite its large size, Camponotus antsaraingy may be confused with Camponotus mita and Camponotus voeltzkowii because in these three species the propodeal face of the propodeum is more reduced than the declivitous face. In C. antsaraingy the antennal segments are entirely flattened while in C. mita only the basal half is flattened; C. voeltzkowii has circular antennal segments. The pubescence on the gastral segment consists of thick, decumbent hair of the same type as the pilosity in C. antsaraingy, with short and filiform hairs on the other two species.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	etymology	Etymology. The species epithet is in reference to the type locality.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	materials_examined	Additional material examined. Province Antsiranana: Antsaraingy, - 12.90665, 49.6606, 90 m, littoral forest, BLF (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	description	Figures 2 D, 7 C-D, 41 D, 59	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 0.95 + / - 0.17; 0.87 - 1.02). In full-face view, head ovoid (CWb / CL: 0.34 + / - 0.03; 0.32 - 0.35); posterior margin rounded, lateral margin of head convex. Eyes circular, protruding, placed midway from the occipital corner (PoOC / CL: 0.12 + / - 0.03; 0.11 - 0.13). Mandibles triangular with five teeth. Clypeus transversally trapezoidal, its anterior border broadly rounded, not angular. With head in full-face view, anterior clypeal margin rounded, not projecting (ClyL / GPD: 0.26 + / - 0.03, 0.25 - 0.29). Antennal scape flattened and lobulated at the base, long, surpassing the occiput by the length of three basal funiculi (SL / CS: 0.46 + / - 0.03; 0.45 - 0.48). In lateral view, anterodorsal corner of pronotum tuberculate, pronotal suture distinct, mesonotal suture distinctly impressed so that the anterior portion of propodeum is raised to a blunt edge in profile, and suture forms a concavity (MW / ML: 0.22 + / - 0.02, 0.21 - 0.23; MPH / ML: 0.17 + / - 0.02, 0.17 - 0.18). In lateral view, petiolar node nodiform, higher than long. Entire body, including mandible, smooth and shiny. One pair of standing hairs on angle formed by mesonotal dorsum and propodeum, head with scattered, subdecumbent, filiform brown hairs, scape with decumbent setae. Ant brownish; mandible, clypeus, and pronotum light brown; mid- and forecoxae, apical femora, basal tibiae, and apical portion of third and fourth tergite whitish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	description	Description of major worker. Characteristics of minor workers, except: head quadrate with straight margins (CS: 1.67 + / - 0.06, 1.57 - 1.74; CWb / CL: 0.92 + / - 0.03, 0.90 - 0.96). Eyes circular, closed to the lateral margin (PoOC / CL: 0.29 + / - 0.02, 0.27 - 0.31). In profile, clypeus distinctly truncate (ClyL / GPD: 0.92 + / - 0.03, 0.86 - 0.94), its anterior margin with a short concavity in full-face view. Antennal scape same as minor but short, just reaching the occipital border (SL / CS: 0.77 + / - 0.04, 0.74 - 0.83). Promesonotum forms a unique dome followed by an impressed mesonotal suture, dorsum of propodeum slightly concave near the propodeal angle. Anterior portion of head with widely spaced large punctures, a single puncture in the center of the vertex. Filiform brownish hair present on head and mesosoma dorsum, more abundant on malar area.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	distribution	Distribution and biology. Camponotus chrislaini is restricted to the northern portion of Madagascar (Fig. 70 B). This newly identified species prefers dry (tropical dry forest, dry forest, disturbed dry forest) as well as wet (montane forest and rainforest) habitats at moderate altitudes (below 1000 m). It was primarily collected from rotten logs, sifted litter, inside trees, or on branches above ground.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	discussion	Discussion. The new species is easily distinguishable from the Camponotus Mayria species, but morphologically is most similar to the unidentified species Camponotus MG 098 from subgenus Camponotus Myrmonesites. The main character used to distinguish them is the form of the petiolar scale, as in the former the petiole is much thinner and its dorsal face forms a straight surface, while in the latter the petiole is thickened with the posterior face concave in dorsal view. Camponotus MG 098 is also covered with plentiful setae which are scarce and widely distributed in C. chrislaini. The presence of C. chrislaini and C. MG 098 in six localities (Sakaramy, Binara, Antsahabe, Analamerana-Ankavanana, Galoko, and Mont Kalabenono) suggests that they are reproductively isolated.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.taxon	materials_examined	Additional material examined. Province Antsiranana: Foret de Binara, 7.5 km 230 ° SW Daraina, - 13.255, 49.61667, 375 m, tropical dry forest (B. L. Fisher) (CAS); Foret d' Antsahabe, 11.4 km 275 ° W Daraina, - 13.21167, 49.55667, 550 m, tropical dry forest (B. L. Fisher) (CAS); Foret d' Andavakoera, 21.4 km 75 ° ENE Ambilobe; 4.6 km 356 ° N Betsiaka, - 13.11833, 49.23, 425 m, rainforest (B. L. Fisher) (CAS); Res. Analamerana, 16.7 km 123 ° Anivorano-Nord, - 12.80467, 49.37383, 225 m, tropical dry forest (B. L. Fisher) (CAS); Parc National Montagne d'Ambre, Fozalanana, - 12.46637, 49.22157, 475 m, dry forest (BLF) (CAS); Res. Ankarana, 7 km SE Matsaborimanga, - 12.9, 49.11667, 150 m, tropical dry forest (PSW) (PSWC); Galoko chain, Mont Galoko, - 13.58487, 48.71818, 520 m, rainforest (BLF) (CAS); Galoko chain, Mont Galoko, - 13.5888, 48.72864, 980 m, montane forest (BLF) (CAS); Galoko chain, Mont Galoko, - 13.58745, 48.71419, 38 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, - 13.63999, 48.67374, 498 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, - 13.64179, 48.67282, 643 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, - 13.64609, 48.67732, 937 m, rainforest (BLF) (CAS); Ampasindava, Andranomatavy Forest, - 13.6648, 47.98702, 275 m, disturbed dry forest (BLF) (CAS); Reserve Speciale d'Ambre, 3.5 km 235 ° SW Sakaramy, - 12.46889, 49.24217, 325 m, tropical dry forest (FGAT) (CAS); Reserve Speciale de l'Ankarana, 22.9 km 224 ° SW Anivorano Nord, - 12.90889, 49.10983, 80 m, tropical dry forest (FGAT) (CAS); Ampasindava, Foret d'Ambilanivy, 3.9 km 181 ° S Ambaliha, - 13.79861, 48.16167, 600 m, rainforest (FGAT) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	description	Figures 44 B, 67	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 0.89 + / - 0.08, 0.84 - 0.93). In full-face view, head as long as wide, posterior margin rounded, lateral margin straight and subparallel (CWb / CL: 0.33 + / - 0.02, 0.32 - 0.34). Eye circular, protruding, its border smoothly aligned to lateral margins of head (PoOC / CL: 0.10 + / - 0.02, 0.08 - 0.10). Mandibles triangular with six teeth. Clypeus carinate, its anterior margin produced medially into a subtriangular lobe (ClyL / GPD: 0.23 + / - 0.07, 0.20 - 0.26). Antennal scape long, surpassing the occipital corner by the length of two-and-a-half basal funiculi (SL / CS: 0.39 + / - 0.03, 0.37 - 0.40). In lateral view, mesosoma without anterolateral margination, promesonotum not very convex, mesopropodeal suture impressed, propodeum angular with the base fully twice as long as the declivity, which is straight (MW / ML: 0.18 + / - 0.01, 0.18 - 0.19; MPH / ML: 0.14 + / - 0.01, 0.14 - 0.15). Petiole cuneate in profile, its anterior face convex, and its posterior face flattened along entire broad border. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins on petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remaining segment dark brown to black.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	description	Description of major worker. Characteristics of minor workers, except: head as long as broad, with rather convex posterior borders and subparallel lateral sides (CS: 1.18 + / - 0.09, 1.12 - 1.24, CWb / CL: 0.86 + / - 0.01, 0.85 - 0.87). Eyes circular, placed dorsally next to lateral borders (PoOC / CL: 0.26 + / - 0.00, 0.25 - 0.26). Anterior clypeal margin forms a rounded rectangular lobe (ClyL / GPD: 0.88 + / - 0.05, 0.85 - 0.91). Antennal scape short, just reaching the occipital border (SL / CS: 0.75 + / - 0.04, 0.73 - 0.78). Mesosoma with the same form as minor worker. Sculpture on head coarsely reticulate-punctate anteriorly and tends to be striolate posteriorly.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	distribution	Distribution and biology. Camponotus claveri occupies two different habitats: montane rainforest and open habitats such as Uapaca woodland, eucalyptus plantation, shrubland, and deciduous dry forest, plus urban areas (Fig. 72 A). Their nests are found in soil, under stones, and on the ground. It has been collected at elevations of 830 - 1987 m.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	discussion	Discussion. Camponotus claveri is similar to other species of the Camponotus repens group, but differs in having a thinner petiolar node with sharp dorsal edge, and a rounded triangular anterior clypeal margin. Morphometric analysis combined C. claveri and C. maintilany in the same cluster, meaning that they are the same size but have different qualitative morphological traits. Their distributions are quite distant but sympatric at Alasora, situated in the central highlands.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	etymology	Etymology. This species is named after the collector Claver Marotafika Randrianandrasana.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.taxon	materials_examined	Additional material examined. Province Antananarivo: Alasora, - 18.96245, 47.58925, 1434 m, eucalyptus plantation (BLF) (CAS); Antaponimanadala III Non Protected Area, 6.55 km E Manalalondo, - 19.25583, 47.17751, 1987 m, savanna grassland (ARA) (CAS); Ilafy, - 18.85415, 47.56575, 1385 m, urban / garden (BLF) (CAS). Province Fianarantsoa: Amoron'i Mania Region, District of Ambositra, Italaviana Uapaca forest, 135 km SE of Antsirabe, - 20.17333, 47.086, 1359 m, Uapaca forest (MG) (CAS); Ampandravelo II Non Protected Area, 10.78 km NE Ranohira, - 22.53917, 45.51548, 87 3 m, shrubland (ARA) (CAS); Antapia III Non Protected Area, 26.43 km SW Ambositra, - 20.72, 47.08785, 1494 m, Uapaca woodland (ARA) (CAS); Foret d' Atsirakambiaty, 7.6 km 285 ° WNW Itremo, - 20.59333, 46.56333, 1550 m, montane rainforest (FGAT) (CAS); Mampiarika III Non Protected Area, 28.93 km SW Ambositra, - 20.73583, 47.08399, 1487 m, Uapaca woodland (ARA) (CAS); Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, - 20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	description	Figures 5 A, 16 B, 17 B, 52	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	description	Description of minor worker. Medium size. Absolute cephalic size (CS: 1.45 + / - 0.28; 1.22 - 1.63). In full-face view, head more quadrate (CWb / CL: 0.37 + / - 0.03, 0.35 - 0.41); posterior margin of head almost flat, lateral margin of head slightly convex and tapering to front. Eyes circular, placed midway from the occipital corner (PoOC / CL: 0.08 + / - 0.02; 0.06 - 0.09). Mandibles triangular with six teeth. With head in full-face view, anterior clypeal margin with a short, rounded, rectangular lobe; less convex in lateral view, not medially marginate (ClyL / GPD: 0.25 + / - 0.04, 0.23 - 0.28). Antennal scape circular and long, surpassing the occiput by the length of two basal funiculi (SL / CS: 0.40 + / - 0.04; 0.37 - 0.43). In lateral view, anterodorsal corner of pronotum rounded, mesosomal dorsal outline feebly and rather evenly arcuate above, dorsal face of propodeum generally longer than declivitous face (MW / ML: 0.21 + / - 0.02; 0.19 - 0.23; MPD / ML: 0.24 + / - 0.03; 0.20 - 0.26). In lateral view, petiolar node cuneate, anterodosum straight basally and inclined dorsally to reach the flattened posterodorsal margin. Tibia tubular, not prismatic, its extensor profile with suberect setae. Head dorsum is matte, finely reticulate-punctate with sparse punctures on malar area; mesosoma striate in lateral view; in dorsal view, petiole, mesosoma, and appendages less shiny, finely reticulate-striolate except the abdominal sternite, which is more matte. Mandible finely reticulate-punctate with sparse oblique punctures. Hairs yellowish white, orange, or tawny colored, erect hairs usually abundant, at least on gastral tergite, often on vertex, occipital portion, and thorax dorsum; petiolar node with 4 pairs of hairs on its dorsum. Shiny black, with mandible and basitarsus reddish to brownish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	description	Description of major worker. Characteristics of minor workers, except: head much wider posteriorly (CS: 2.14 + / - 0.26, 1.70 - 2.47; CWb / CL: 1.04 + / - 0.06, 0.94 - 1.20); lateral margins convex and converging towards base of mandible. Eyes semicircular, placed dorsolaterally midway from the occipital margin (PoOC / CL: 0.22 + / - 0.01, 0.19 - 0.24). Clypeus much more quadrate in full-face view (ClyL / GPD: 0.72 + / - 0.09, 0.48 - 0.81), its anterior margin slightly concave medially. Antennal scape short, just surpassing the occipital border (SL / CS: 0.83 + / - 0.08, 0.75 - 0.99). Dorsal outline of mesosoma forms a continuous line so that the basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely reticulate above, punctate to finely areolate below.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	distribution	Distribution and biology. Camponotus darwinii is a famous species which is distributed widely across central of Madagascar, with a range extending from Ankazobe to Mount Papango, PN Befotaka Midongy (Fig. 58 A). It has been collected from a wide variety of different habitats, including rainforest, montane rainforest, Uapaca forest, and Uapaca woodland as well as disturbed habitats such as urban and garden areas and cultivated land (Tavy). The species has also been found nesting inside Melia azedarach and Phellolophium madagascariense. Habitat elevations ranged from 1069 - 2150 m.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	discussion	Discussion. Camponotus darwinii should be separable from most similar species in this group by the combination of the following characters: presence of dense hairs on mesosoma and / or gastral dorsum, mandible and malar area distinctly reddish in color, propodeal and declivitous faces unequal in length. This species is much closer to C. ursus but differs in the density of mesosomal hairs and the aspect of the petiole. In addition, C. darwinii is much more robust than C. ursus.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.taxon	materials_examined	Additional material examined. Province Antananarivo: Analamanga Region, District of Ankazobe, Ambohitantely, 46 km NE of Ankazobe, - 18.198, 47.2815, 701 m, Foret sclerophylle (MG) (CAS); Ambatolaona, - 18.928, 47.88283, 1382 m, urban / garden (BLF) (CAS); Kaloy, - 18.58998, 47.65102, 1423 m, disturbed montane rainforest, (BLF) (CAS); Ilafy, - 18.85415, 47.56575, 1385 m, urban / garden (BLF) (CAS); Foret de galerie, Telomirahavavy, 23.4 km NNE Ankazobe, - 18.12167, 47.2062 7, 1520 m, disturbed gallery montane forest (BLF) (CAS); Stn. Forestiѐre Manjakatompo, - 19.35, 47.31667, 2000 m, montane rainforest (Mark Pigeon) (CAS); 25 km NNE Ankazobe, - 18.1, 47.18333, 1500 m, rainforest (PSW) (PSWC); Stn. Forestiѐre Manjakatompo, - 19.35, 47.31667, 1600 m, montane rainforest (PSW) (PSWC); - 18.91667, 47.53333, 1350 m, park / garden (PSW) (PSWC); Parc de Tsimbazaza; Antananarivo renivohitra, - 18.93, 47.52611, 1275 m (D. M. Olson) (PSWC); - 18.91667, 47.53333, 1300 m, park / garden (PSW) (PSWC); 3 km 41 ° NE Andranomay, 11.5 km 147 ° SSE Anjozorobe, - 18.47333, 47.96, 1300 m, montane rainforest (FGAT) (CAS); Parc Botanique et Zoologique de Tsimbazaza, - 18.93233, 47.91167, 1300 m (D. H. Kavanaugh) (CAS); Angavokely, - 18.93333, 47.75 (B. Pettersson) (PSWC). Province Fianarantsoa: 29 km SSW Ambositra, Ankazomivady, - 20.77667, 47.165, 1700 m, disturbed montane rainforest (B. L. Fisher) (CAS); Foret d' Atsirakambiaty, 7.6 km 285 ° WNW Itremo, - 20.59333, 46.56333, 1550 m, montane rainforest (FGAT) (CAS); 2 km W Andrambovato, along river Tatamaly, - 21.51167, 47.41, 1075 m, cultivated land (tavy), recent 1 - 2 year-old tavy (BLF) (CAS); stream area, 900 m E of Isalo National Park Interpretive Center, - 22.62667, 45.35817, 750 m, open area near stream (R. Harin'Hala) (CAS); Parc National d'Andringitra, Plateau d'Andohariana, 39.8 km 204 ° Ambalavao, - 22.18767, 46.90083, 2150 m, rubicole thicket at base of cliff (BLF) (CAS); Vohiparara, - 21.23333, 47.36667 (A. Pauly) (CAS); Miandritsara Forest, 40 km S of Ambositra, - 20.79267, 47.17567, 822 m, low altitude rainforest (MG) (CAS); dry wash, 1 km E of Isalo National Park Interpretive Center, - 22.62667, 45.35817, 885 m, dry wash (R. Harin'Hala) (CAS); Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango, - 23.84083, 46.9575, 1250 m, montane rainforest (BLF) (CAS); Parc naturel communautaire, 29.3 km SW Ambositra, - 20.79751, 47.1791, 1782 m, disturbed montane rainforest (BLF) (CAS); Parc naturel communautaire, 26.8 km SW Ambositra, - 20.775, 47.18362, 1755 m, disturbed montane rainforest (BLF) (CAS); Parc naturel communautaire, 28.5 km SW Ambositra, - 20.78414, 47.16699, 1780 m, disturbed montane rainforest along road side (BLF) (CAS); Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, - 20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (CAS); R. S. Ivohibe, 6.5 km ESE Ivohibe, - 22.49667, 46.955, 1575 m, montane rainforest (B. L. Fisher (Sylvain )) (CAS); P. N. Andringitra, Foret Ravaro 12.5 km SW Antanifotsy, - 22.21167, 46.845, 1650 m, montane rainforest (S. Razafimandimby) (CAS); R. S. Ivohibe 8.0 km E Ivohibe, - 22.48333, 46.96833, 1200 m, montane rainforest (B. L. Fisher (Sylvain )) (CAS); 28 km. SSW Ambositra, Ankazomivady, - 20.775, 47.16833, 1670 m, montane rainforest edge (B. L. Fisher) (CAS); 38 km S Ambalavao, Res. Andringitra, - 22.2, 46.96667, 1680 m, montane rainforest (B. L. Fisher) (CAS); 40 km S Ambalavao, Res. Andringitra, - 22.21667, 46.96667, 1275 m, montane rainforest (B. L. Fisher) (CAS); Res. Andringitra; 8.5 km SE Antanitotsy, - 22.16667, 46.96667, 1990 m, montane rainforest (Sylvian) (CAS); 8.0 km NE Ivohibe, - 22.42167, 46.89833, 1200 m, montane rainforest (B. L. Fisher (Sylvain )) (CAS); Antapia I Non Protected Area, 26.43 km SW Ambositra, - 20.71972, 47.08685, 1495 m, Uapaca woodland (ARA) (CAS); Antohatsahomby I Non Protected Area, 22.77 km NW Ambatofinandrahana, - 20.55056, 46.58562, 1550 m, Uapaca woodland (ARA) (CAS); Amoron'i Mania Region, District of Ambositra, Italaviana Uapaca forest, 135 Km SE of Antsirabe, - 20.17333, 47.086, 1359 m, Uapaca forest, (MG) (CAS); Antapia IV Non Protected Area, 26.42 km SW Ambositra, - 20.71917, 47.0868, 1494 m, Uapaca woodland (ARA) (CAS); Antohatsahomby V Non Protected Area, 22.63 km NW Itremo, - 20.56722, 46.57923, 1726 m, Uapaca woodland (ARA) (CAS); Manombo Special Reserve, 32 km SE of Farafangana, - 23.02183, 47.72, 36 m, lowland rainforest (MG) (CAS); Fitovavy Fitovinany Region, District of Ifanadiana, 12 km W of Ranomafana, - 21.25083, 47.40717, 1127 m, forest edge, open area (MG) (CAS); Antohatsahomby IV Non Protected Area, 22.67 km NW Itremo, - 20.56306, 46.58097, 1708 m, Uapaca woodland (ARA) (CAS); Ambinanindranomena Non Protected Area, 39.45 km SE Ambalavao, - 21.95386, 47.29427, 1069 m, montane rainforest (ARA) (CAS); Hte Mananjary; Ambinanindrano; Ambositra, - 20.733334, 47.633335, 657 m (R. Catala) (MNHN). Province Mahajanga: Sofia Region, District of Sofia, Anjiamangirana 45 km S Antsohihy, Analagnambe Galery forest, 5 km W Anjiamangirana, - 15.157, 47.73417, 97 m, low degraded dry forest (MG) (CAS). Province Toamasina: Ambanizana, Parc National Masoala, - 15.57222, 50.00694, 1020 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CAS); Moramanga, - 18.948902, 48.224426, 914 m (Decarpentries) (NHMB); 6.9 km NE Ambanizana, Ambohitsitondroina, - 15.58506, 50.00952, 825 m, rainforest (B. L. Fisher) (CAS); Montagne d'Anjanaharibe, 19.5 km 27 ° NNE Ambinanitelo, - 15.17833, 49.635, 1100 m, montane rainforest (FGAT) (CAS). Province Toliara: [S. W. Madagascar]; Beronono; Mahabo, - 21, 45, 339 m (Voltzkow) (NHMB); Androy Region, District of Tsihombe, 74 km S of Tsihombe, Cap Ste Marie Reserve, - 25.58767, 45.163, 36 m, spiny bush (MG) (CAS); 13 km NW Enkara, Res Andohahela, - 24.55, 46.8, 1300 m, montane rainforest (B. L. Fisher) (CAS); Atsimo Andrefana Region, District of Tulear II, Mikea decidious dry forest 3 km N Andranomavo village, - 22.90367, 43.4755, 30 m, deciduous dry forest (MG) (CAS); Miary, - 23.311972, 43.740917, 55 m (F. Geay) (MNHN); Ambovombe, - 25.1775, 46.09283, 25 m (R. Decary) (MNHN); near road, Zombitse National Park, - 22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin'Hala) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	description	Figures 2 A, 8 A, 38 B, 63	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	diagnosis	Diagnosis. Head and mesosoma reddish brown to dark brown, gaster always dark colored; anterior clypeal margin of clypeus with short, rectangular lobe; pronotal humeri tuberculate, dorsal outline of mesosoma arcuate; gastral tergite with short, decumbent, ochreous setae.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	description	Description of minor worker. Large-sized species. Absolute cephalic size (CS: 1.85 + / - 0.55, 1.45 - 2.42). In full-face view, head trapezoidal, wider than length of pronotum in dorsal view; occipital margin slightly convex medially, and straight subparallel lateral margin of head (CWb / CL: 0.36 + / - 0.03, 0.33 - 0.39). Eyes elliptical, positioned at the midline of head (PoOC / CL: 0.08 + / - 0.01, 0.07 - 0.09). Mandibles short and triangular with six teeth. Clypeus trapeziform, less convex in profile, sharply carinate along its entire length, with a short, rectangular projection feebly sinuate in the middle (ClyL / GPD: 0.12 + / - 0.01, 0.11 - 0.13). Frontal carina less sinuate and divergent. Antennal scape long and surpassing the occipital corner (SL / CS: 0.41 + / - 0.07; 0.34 - 0.46). In lateral view, dorsal outline of mesosoma forms a smooth curve, anterodorsal corner of pronotum distinctly shouldered, propodeum rounded, its surface convex and not marginate laterally, declivitous face triangular in posterior view (MW / ML: 0.23 + / - 0.03, 0.21 - 0.25; MPH / ML: 0.18 + / - 0.03, 0.16 - 0.20). Petiolar node squamiform in profile, wider than long, anterodorsal face distinctly flat and inclined forward, posterodorsal face flat, node summit arcuate. First abdominal tergite with vertical anterior face. Entire body (frontal area, dorsal face of mesosoma, scape, and legs) entirely, finely, and densely reticulate-punctate and matte; lateral face of petiole and abdomen transversally striolate. Mandible finely reticulate-punctate and subopaque, with superimposed, abundant punctuation arranged transversally. Ground pilosity yellowish, short, sparse, and dilute on tibia and scape. Short white to yellowish standing hair present on entire dorsum, sparsely distributed on occipital region, regularly present on petiolar dorsum and onto declivity, absent on scape and tibia. Dorsum of gastral segment with appressed ochreous setae and whitish, suberect hairs arranged serially parallel and longitudinally. Head and mesosoma dark brown to reddish brown; mandible, basal half of scape, basitarsus, and gastral segment reddish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	description	Description of major worker. Characteristics of minor workers, except: head as long as broad, with rather straight posterior and lateral borders (CS: 4.13 + / - 0.16, 3.90 - 4.33; CWb / CL: 1.04 + / - 0.02, 1.01 - 1.07). Eyes circular, placed dorsally next to lateral borders (PoOC / CL: 0.28 + / - 0.01, 0.26 - 0.29). Anterior clypeal margin forms a rectangular lobe (ClyL / GPD: 0.87 + / - 0.05, 0.79 - 0.92). Antennal scape short, not reaching the occipital border (SL / CS: 0.61 + / - 0.02, 0.58 - 0.65). Promesonotal suture strongly marked.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	distribution	Distribution and biology. Camponotus ellioti occurs in the dry forest regions of the southeast of Madagascar in habitats such as bush, coastal scrub, and coastal spiny bush on sandy soil, Euphorbia forest, spiny bush and thicket, gallery forest, and riparian scrub (Fig. 70 C). It has been collected from sifted litter and Malaise traps. The altitudes of these localities range from 5 to 230 meters.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	discussion	Discussion. Camponotus ellioti is a distinctive species recognizable by the submargined anterolateral corner of the pronotum and the gaster covering of thick, blunt hairs the color of rust. In the traditional classification, C. ellioti is considered a member of the C. darwinii species group based on the density and color of the gastral pilosity and the sculpture pattern; however, we propose color as a character difference between these two groups.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.taxon	materials_examined	Additional material examined. Province Toliara: [Region du Sud Andrahomana, Museum Paris, Ch. Alluaud 1901]; Analapatsa I; Taolagnaro, - 25.1833333, 46.6666667, 31 m, (NHMB); [Toliara S. W. Madagascar]; Beronono; Mahabo, - 21, 45, 339 m (Voeltzkow) (MHNG); 3.5 km 236 ° SW Marovato, - 25.55389, 45.25583, 230 m, spiny forest / thicket (FGAT) (CAS); 5 km N Ampotaka, Malaise on trail in Vitambany gallery forest, - 24.65033, 43.96317, 86 m, gallery forest (MG) (CAS); 7.0 km 156 ° SSE Lavanono, - 25.47111, 44.9885, 50 m, spiny forest / thicket (FGAT) (CAS); Ambohimahavelona village 33 km NE of Toliara, Andoharano, dry forest, - 23.44083, 43.89967, 46 m, dry forest (MG) (CAS); Andranovato, 5 km SE of Manombo, - 22.81806, 43.50217, 18 m, Euphorbia forest (CAS); Androy Region, District of Tsihombe, 74 km S of Tsihombe, Cap Sainte Marie Reserve, - 25.58767, 45.163, 36 m, spiny bush (MG) (CAS); Anosy Region, District of Amboasary, 58 km SW of Fort Dauphin, 8 km NW of Amboasary, Berenty Special Reserve, - 25.00667, 46.30333, 85 m, gallery forest (MG) (CAS); Atsimo-Andrefana Region, Antsokay Arboretum, - 23.41491, 43.75499, 13 m, spiny forest / thicket (B. L. Fisher, F. A. Esteves et al.) (CAS); Atsimo-Andrefana Region, Famata 02, - 23.45314, 43.76448, 20 m, coastal spiny bush on sandy soil (B. L. Fisher, F. A. Esteves et al.) (CAS); Atsimo-Andrefana Region, Famata 04, - 23.53922, 43.77935, 20 m, riparian scrub (B. L. Fisher, F. A. Esteves et al.) (CAS); Foret de Petriky, 12.5 km W 272 ° Tolagnaro, - 25.06167, 46.87, 10 m, littoral rainforest (B. L. Fisher) (CAS); Libanona beach, Tolagnaro, - 25.03883, 46.996, 20 m, coastal scrub (BLF) (CAS); Mahafaly Plateau, 6.2 km 74 ° ENE Itampolo, - 24.65361, 43.99667, 80 m, spiny forest / thicket (FGAT) (CAS); Mikea Forest, spiny forest, - 22.91333, 43.48222, 37 m, spiny forest (R. Harin'Hala) (CAS); Parc National de Tsimanampetsotsa, 6.7 km 130 ° SE Efoetse, 23.0 km 175 ° S Beheloka, - 24.10056, 43.76, 25 m, spiny forest / thicket (FishFGAT) (CAS); Parc National de Tsimanampetsotsa, Foret de Bemanateza, 20.7 km 81 ° E Efoetse, 23.0 km 131 ° SE Beheloka, - 23.99222, 43.88067, 90 m, spiny forest / thicket (FiFGAT) (CAS); Ranobe, - 23.04457, 43.61532, 20 m, spiny forest / thicket (Frontier Wilderness Project) (CAS); Reserve Prive Berenty, Foret d'Anjapolo, 21.4 km 325 ° NW Amboasary, - 24.92972, 46.20967, 65 m, spiny forest / thicket (FGAT) (CAS); Reserve Speciale de Cap Sainte Marie, 12.3 km 262 ° W Marovato, - 25.58167, 45.16833, 200 m, spiny forest / thicket (FGAT) (CAS); Reserve Speciale de Cap Sainte Marie, 14.9 km 261 ° W Marovato, - 25.59444, 45.14683, 160 m, spiny forest / thicket (FGAT) (CAS); Tolagnaro (= Fort Dauphin), - 25.03333, 47, 5 m, littoral vegetation (B. L. Fisher) (CAS); Tolagnaro (= Ft. Dauphin), - 25.04611, 47, 30 m, garden, hotel (D. M. Olson) (PSWC); Toliara, - 23.3575, 43.669, 20 m, urban / garden (BLF) (CAS); Tsimanampetsotsa National Park, Mitoho Forest, Malaise across trail at escarpment base, - 24.0485, 43.75233, 120 m, dense dry forest and transitional forest (MG) (CAS); Tsimelahy - Parcel II, Andohahela National Park, transitional forest, Toliara Province, - 24.93683, 46.62667, 180 m, transition forest (M. E. Irwin, F. D. Parker, R. Harin'Hala) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	description	Figures 46 B, 68	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	description	Description of minor worker. Small-sized species. Absolute cephalic size (CS: 0.78 + / - 0.12; 0.75 - 0.81). In full-face view, head ovate, usually somewhat longer than broad, lateral margin of head straight and tapering to front (CWb / CL: 0.34 + / - 0.01, 0.34 - 0.35). Eye subelliptical, protruding, its border smoothly aligned to lateral margins of head (PoOC / CL: 0.09 + / - 0.03, 0.08 - 0.10). Mandibles triangular with six teeth. Anterior clypeal margin not produced, entire, broadly rounded. Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL / CS: 0.35 + / - 0.03, 0.34 - 0.36). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeum dorsum is raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW / ML: 0.18 + / - 0.01, 0.17 - 0.18; MPH / ML: 0.10 + / - 0.01, 0.10 - 0.10). In lateral view, petiole cuneate, its anterior face convex, its posterior face more flattened. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins of petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown, and the remaining segment dark brown to black.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	distribution	Distribution and biology. Camponotus ihazofotsy is known from the dry forest of Tsimanapetsotsa and Andohaela NP (Fig. 72 B). We assume that this species is arboreal because all samples were collected from Malaise traps.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	discussion	Discussion. Camponotus ihazofotsy closely resembles C. tsimelahy, although morphological examination reveals a number of distinguishing characters. The most interesting character is the form and length of the petiolar scale; in C. ihazofotsy the petiole is cuboidal (anterior and posterior face about the same height), while in C. tsimelahy the posterior face is higher than the anterior face. In addition, these two species are sympatric at one site, Ihazofotsy. Molecular phylogenetic analysis provides strong evidence that they represent separate lineages (unpublished data).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	etymology	Etymology. The species epithet refers to the type locality.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.taxon	materials_examined	Additional material examined. Province Toliara: Tsimanampetsotsa National Park, Mitoho Forest, Malaise across trail at escarpment base, - 24.0485, 43.75233, 120 m, dense dry forest and transition forest (MG) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	description	Figures 9 D, 50 C, D, 65	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 1.21 + / - 0.19, 1.13 - 1.30). In full-face view, head almost quadrate, with feebly convex posterior and lateral border (CWb / CL: 0.35 + / - 0.03, 0.33 - 0.36). Eyes elliptical, placed next to the lateral margins of head (PoOC / CL: 0.10 + / - 0.01, 0.10 - 0.11). Mandibles triangular with six teeth. Anterior clypeal margin angularly produced, with straight lateral border and rounded anterolateral corner. Antennal scape long, surpassing the occipital corner by the length of two basal funiculi (SL / CS: 0.39 + / - 0.02, 0.38 - 0.40). In lateral view, pronotum with short anterolateral margination, mesosoma moderately convex, propodeum angular in profile, with unequal basal and declivitous faces, each straight in profile and meeting at a rounded obtuse angle (MW / ML: 0.21 + / - 0.03, 0.19 - 0.22; MPH / ML: 0.16 + / - 0.01, 0.16 - 0.17). In lateral view, petiole rather narrow, its anterior face convex and its posterior face flattened. Body finely imbricate. Hairs whitish; enlarged, whitish hairs with blunt tip on mesonotum, propodeum, and petiolar dorsum; filiform, pointed, grayish hair on head, pronotum, and gastral segment; pubescence white, short, and more abundant across entire dorsum. Head, mesosoma and gaster dark brown, femora, tibia, and funiculi brown, scape and basitarsi slightly lighter.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	description	Description of major worker. Characteristics of minor workers, except: head as long as wide, posterior borders medially concave and subparallel, lateral sides diverging apically (CS: 1.93; CWb / CL: 0.93). Eyes circular, placed dorsally in the middle of the lateral borders (PoOC / CL: 0.28). Anterior clypeal margin forms a short, rounded, rectangular lobe, slightly concave medially (ClyL / GPD: 0.84). Antennal scape short, just reaching the occipital border (SL / CS: 0.69). Sculpture, pilosity, and color as in minor worker.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	distribution	Distribution and biology. Camponotus ivadia has thus far been collected only in the Foret d' Ampondrabe within Ankarana Reserve, a region with tropical dry forest located in the northern portion of Madagascar (Fig. 70 F). All specimens were collected by beating vegetation.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	discussion	Discussion. This species is easily recognized within the Camponotus madagascarensis group as one of few back and shiny species with white standing pubescence.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	etymology	Etymology. The species epithet Camponotus ivadia is derived from the low vegetation microhabitat it prefers.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.taxon	materials_examined	Additional material examined. Province Antsiranana: Res. Ankarana, 7 km SE Matsaborimanga, - 12.9, 49.11667, 150 m, tropical dry forest (PSW) (PSWC); Foret d' Ampondrabe, 26.3 km 10 ° NNE Daraina, - 12.97, 49.7, 175 m, tropical dry forest (B. L. Fisher) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	description	Figures 10 C, D, 45 A, 66	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	description	Description of minor worker. Medium-sized. Absolute cephalic size (CS: 0.99 + / - 0.14; 0.91 - 1.09). In full-face view, head almost quadrate (CWb / CL: 0.32 + / - 0.02; 0.31 - 0.34); posterior margin evenly straight, lateral margin of head anterior to eyes slightly convex. Eyes circular, placed above midline of head (PoOC / CL: 0.10 + / - 0.02; 0.09 - 0.12). Mandibles triangular with six teeth. Clypeus carinate, its anterior border angularly produced in the middle (ClyL / GPD: 0.20 + / - 0.06, 0.16 - 0.23). Antennal scape long, surpassing the occiput by the length of one basal funiculus (SL / CS: 0.34 + / - 0.02; 0.32 - 0.34). In lateral view, anterodorsal corner of pronotum rounded, mesonotum dorsum inclined toward nearly straight propodeum face (MW / ML: 0.19 + / - 0.02, 0.18 - 0.20; MPH / ML: 0.13 + / - 0.02, 0.12 - 0.14). In lateral view, petiolar node longer than wide, with flat summit, its anterior face leaning forward. Ants finely foveolate throughout, apparently shiny. Entire body, including mandibles and appendages, covered with plentiful, erect, whitish setae, uneven in length, some long. Head, mesosoma, and gaster dark brown to black; antennae, mandibles, tibiae, and tarsi reddish brown.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	description	Description of major worker. Characteristics of minor workers, except: head much wider posteriorly (CS: 1.68 + / - 0.07, 1.61 - 1.76; CWb / CL: 0.91 + / - 0.02, 0.89 - 0.92); lateral margins convex and converging towards base of mandible. Eyes circular placed dorsolaterally midway from the occipital margin (PoOC / CL: 0.30 + / - 0.00, 0.29 - 0.30). Clypeus with a short rectangular lobe with rounded lateral angle (ClyL / GPD: 0.85 + / - 0.03, 0.81 - 0.88). Antennal scape short, not reaching the occipital margin (SL / CS: 0.57 + / - 0.02, 0.56 - 0.59). Dorsal outline of mesosoma interrupted by sutures, metanotal suture forms a canal. Head dorsum and mesosoma coarsely reticulate-punctate, petiole finely striolate, gaster imbricate and shiny.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	distribution	Distribution and biology. The known elevational range of Camponotus jjacquia extends from 43 to 923 m. Specimens have been collected on tree trunks, on low vegetation, in leaf litter, and under stones in Bismarckia woodland, deciduous forest, gallery forest, savannah woodland, spiny forest, and tropical dry forest (Fig. 72 C). This species nests in the soil.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	discussion	Discussion. The unique petiolar shape, the density of its pilosity, and the shape of its metanotal dorsum make C. jjacquia a very distinctive species.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	etymology	Etymology. The new species is dedicated to Jean Jacques Rafanomezantsoa, who has been a Malagasy ant collector for more than twenty years.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.taxon	materials_examined	Additional material examined. Province Fianarantsoa: Ampotoampoto III National Park, 7.91 km NW Ilakaka, - 22.62944, 45.189, 919 m, savannah woodland (ARA) (CAS); Isalo IV National Park, 12 km SW Ranohira, - 22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (CAS); Ampotoampoto IV National Park, 7.83 km NW Ilakaka, - 22.62944, 45.1912, 923 m, savanna woodland (ARA) (CAS); Province Mahajanga: Ampijoroa National Park, 160 km N Maevatanana, deciduous forest, - 16.31944, 46.81333, 43 m, deciduous forest (MG) (CAS). Province Toliara: Anosy Region, District of Amboasary, 58 km SW of Fort Dauphin, 8 km NW of Amboasary, Berenty Special Reserve, - 25.00667, 46.30333, 85 m, Galery forest (MG) (CAS); Anosy Region, District of Fort-Dauphin, Andohaela National Park Parcelle II, Tsimelahy, 42 km W of Fort-Dauphin, - 24.93683, 46.62667, 176 m, transition forest (Mike Irwin, Frank Parker, Rin'ha) (CAS); Anosy Region, Parc National d'Andohahela, Foret de Manatalinjo, - 24.82505, 46.57811, 90 m, spiny forest / thicket (B. L. Fisher, F. A. Esteves et al.) (CAS); Atsimo Andrefana Region, District of Betioky; Beza Mahafaly Special reserve, Parcelle Belle vue 7 km W of Research Station, - 23.68983, 44.5755, 177 m, spiny forest (Rin'ha) (CAS); Atsimo Andrefana Region, District of Betioky, 30 km E Betioky, Beza Mahafaly Special Reserve (around Research Station), - 23.6865, 44.591, 165 m, gallery dry deciduous forest (MG); Beza-Mahafaly, 27 km E Betioky, - 23.65,4 4.63333, 135 m, tropical dry forest (B. L. Fisher) (CAS); Foret de Mahavelo, Isantoria River, - 24.75833, 46.15717, 110 m, spiny forest / thicket (FGAT) (CAS); Parc National d'Andohahela, Foret d' Ambohibory, 1.7 km 61 ° ENE Tsimelahy, 36.1 km 308 ° NW Tolagnaro, - 24.93, 46.6455, 300 m, tropical dry forest (FGAT) (CAS); Parc National d'Andohahela, Foret de Manatalinjo, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, - 24.81694, 46.61, 150 m, spiny forest / thicket (FGAT) (CAS); Parc National de Kirindy Mite, 16.3 km 127 ° SE Belo sur mer, - 20.79528, 44.147, 80 m, tropical dry forest (FGAT) (CAS); Parc National de Tsimanampetsotsa, Foret de Bemanateza, 20.7 km 81 ° E Efoetse, 23.0 km 131 ° SE Beheloka, - 23.99222, 43.88067, 90 m, spiny forest / thicket (FGAT) (CAS); Beza Mahafaly Reserve, near research station, Parcel I, - 23.6865, 44.591, 165 m, dry deciduous forest (R. Harin'Hala) (CAS); Beza Mahafaly Reserve, near Bellevue, Parcel II, - 23.68983, 44.5755, 180 m, spiny forest (R. Harin'Hala) (CAS); Andohahela National Park, Tsimelahy - Parcel II, transitional forest, - 24.93683, 46.62667, 180 m, transitional forest (M. E. Irwin, F. D. Parker, R. Harin'Hala) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	description	Figures 37 B, 39 A, 64	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 0.84 + / - 0.05, 0.81 - 0.86). In full-face view, head distinctly longer than broad, with convex occipital margin and subparallel lateral sides (CWb / CL: 0.33 + / - 0.02, 0.32 - 0.34). Eyes elliptical and break the lateral outlines of head (PoOC / CL: 0.09 + / - 0.01, 0.08 - 0.10). Mandibles triangular with six short, blunt teeth. Clypeus convex, its anterior border produced into rounded lobe (ClyL / GPD: 0.24 + / - 0.01, 0.22 - 0.25). Antennal scape long, surpassing the occipital margin (SL / CS: 0.40 + / - 0.03, 0.39 - 0.43). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeum dorsum is raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW / ML: 0.20 + / - 0.01, 0.19 - 0.20; MPH / ML: 0.14 + / - 0.03, 0.12 - 0.15). In lateral view, petiole cuneate, its anterior face convex, its posterior face more flattened with entire border broad. Body finely imbricate. Hairs grayish, present only on vertex, one pair next to the pronotal suture, one pair on the middle of mesonotum dorsum, sparse on gastral tergite; enlarged, whitish hairs present on propodeal corner (two or three pairs) and on posteromargin of petiole; pubescence whitish and more dilute on gaster tergites. Head, mesosoma, and appendages reddish to brownish yellow; first gastral segment lighter colored than the remaining dark brown segments.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	description	Description of major worker. Characteristics of minor workers, except: head apparently rectangular, with straight to feebly convex posterior margin and parallel lateral sides. (CS: 1.13 + / - 0.05, 1.09 - 1.18; CWb / CL: 0.90 + / - 0.02, 0.88 - 0.91). Eyes circular, placed dorsally next to the lateral margin (PoOC / CL: 0.26 + / - 0.01, 0.25 - 0.27). Anterior clypeal margin forms a short, rounded lobe (ClyL / GPD: 0.83 + / - 0.00, 0.83 - 0.83). Antennal scape short, just reaching the occipital border (SL / CS: 0.77 + / - 0.03, 0.74 - 0.79). Dorsal outline of mesosoma not very convex and interrupted but pronotal and mesopropodeal suture present, the latter weakly impressed.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	distribution	Distribution and biology. Camponotus maintilany occurs from 1300 - 1987 m in elevation and inhabits a montane environment in the central highlands, where it prefers mostly Uapaca forest or woodland, but is also found in savanna grassland and eucalyptus plantations (Fig. 70 E). This new species has been collected by digging in soil, sifting leaf litter, and baiting with tuna fish or sardines. In addition, a few workers were found in Malaise traps.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	discussion	Discussion. Camponotus maintilany is most readily distinguished from other members of the Camponotus ellioti species group by a deeper metanotal groove and shinier sculpture. It may be confused with C. andrianjaka but can be separated by its impressed metanotal suture and truncate petiolar node.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	etymology	Etymology. This species is derived from the coloration of the mesosoma and gaster. " Mainty " means black and " ilany " means second part.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.taxon	materials_examined	Additional material examined. Province Antananarivo: Alasora, - 18.96245, 47.58925, 1434 m, grassland (BLF) (CAS); Ambatolaona, - 18.928, 47.88283, 1382 m, urban / garden (BLF) (CAS); Ambohidrabiby, - 18.7746, 47.60908, 1362 m, eucalyptus plantation (BLF) (CAS); Analamanga Region, District of Arivonimamo, Andebadeba Uapaca forest, 4 km from Arivonimamo, - 19.00806, 47.14861, 1300 m, Uapaca forest (MG) (CAS); Andohony I Non Protected Area, 22.62 km SW Antsirabe, - 20.06784, 46.99068, 1451 m, Savannah grassland (ARA) (CAS); Andohony II Non Protected Area, 22.71 km SW Antsirabe, - 20.06904, 46.99199, 1398 m, savanna grassland (ARA) (CAS); Ankalalahana, - 19.00711, 47.1337, 1350 m, Uapaca woodland (BLF) (CAS); Ankalalahana, - 19.00659, 47.1122, 1375 m, Uapaca woodland (BLF) Ankalalahana, - 19.00659, 47.1122, 1375 m, Uapaca woodland (BLF) (CAS); Ankalalahana, - 19.00716, 47.1124, 1370 m, Uapaca woodland (BLF) (CAS); Ankalalahana, - 19.0531, 47.11304, 1314 m, Uapaca woodland (H. E. Marti et al.) (CAS); Ankalalahana, - 18.99666, 47.1118, 1353 m, Uapaca woodland (H. E. Marti et al.) (CAS); Antaponimanadala III Non Protected Area, 6.55 km E Manalalondo, - 19.25583, 47.17751, 1987 m, savanna grassland (ARA) (CAS); Iharanandriana, - 19.15823, 47.49702, 1513 m, Uapaca woodland (BLF) (CAS), Kaloy, - 18.59568, 47.65333, 1338 m, grassland (BLF) (CAS); Navoatra I Non Protected Area, 7.64 km NW Arivonimamo, - 18.97806, 47.11929, 1373 m, Uapaca woodland (ARA) (CAS); Navoatra III Non Protected Area, 7.39 km NW Arivonimamo, - 18.98028, 47.12071, 1321 m, Uapaca woodland (ARA) (CAS); Navoatra IV Non Protected Area, 7.15 km NW Arivonimamo, - 18.98194, 47.12232, 1323 m, Uapaca woodland (ARA) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	description	Figures 15 A, 53	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	description	Description minor worker. Medium-sized species. Absolute cephalic size (CS: 1.45 + / - 0.24, 1.31 - 1.66). In full-face view, head more quadrate (CWb / CL: 0.36 + / - 0.02; 0.35 - 0.38), posterior margin of head and lateral margin weakly convex. Eyes circular, placed laterally close to lateral head margin (PoOC / CL: 0.08 + / - 0.01; 0.07 - 0.09). Mandibles triangular with six teeth. Clypeus convex, its anterior portion projected into a rectangular lobe. Antennal scape circular and long (SL / CS: 0.40 + / - 0.03; 0.37 - 0.42). In lateral view, mesosoma low and long, its dorsum smoothly arcuate, metanotal suture obsolete. Humeral angle rounded. Propodeal dorsum about the same length as declivitous face (MPH / ML: 0.18 + / - 0.02; 0.18 - 0.20). In lateral view, petiole squamiform with blunt edge, anterior face short and converging to the highest point of the petiole, posterior face straight. Ventral face of femora marginate laterally, the outer margin of tibia with long, whitish setae. Head and mesosoma coarsely reticulate-punctate, legs and gastral tergite finely imbricate. Mandible finely foveolate with sparse punctures. Pilosity and pubescence whitish, the former sparse and suberect, especially on occipital portion, frontal area, and mesosoma, though most conspicuous on the thoracic dorsum. On the gaster, the suberect hairs are long and abundant, overlapping with appressed hairs, pubescence inconspicuous and short. Petiolar node fringed with one row of whitish hair from the level of petiolar spiracle. Head including scape, mesosoma, and gaster dark brown to black, legs brownish, less dark than mesosoma.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	description	Description of major worker. Characteristics of minor workers, except: head much wider posteriorly (CS: 1.98 + / - 0.30, 1.78 - 2.32; CWb / CL: 0.99 + / - 0.01, 0.98 - 1.00); lateral margins convex and converging towards base of mandible. Eyes semicircular, placed dorsolaterally midway from the occipital margin (PoOC / CL: 0.23 + / - 0.01, 0.22 - 0.23). Clypeus much more quadrate in full-face view (ClyL / GPD: 0.72 + / - 0.02, 0.69 - 0.73), its anterior margin slightly concave medially. Antennal scape short, just surpassing the occipital border (SL / CS: 0.90 + / - 0.14, 0.75 - 1.00). Dorsal outline of mesosoma forms a continuous line so that the basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely reticulate above, punctate to finely areolate below.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	distribution	Distribution and biology. Camponotus norvigi has been collected primarily by Malaise trap and beating low vegetation, and rarely from yellow pan traps. No colonies have been collected. It is distributed mainly in montane forest, rainforest, and high-altitude rainforest in southern, western, and central Madagascar; in tropical dry forest in northern Madagascar, e. g., Res Tsingy de Bemaraha, and in gallery forest and open areas on the western portion of the island (Fig. 58 B). It occurs at elevations of 20 - 1606 meters. This species is sympatric with C. darwinii at the edge of PN Isalo and Ranomafana, at Ambohitantely (Foret sclerophylle).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	discussion	Discussion. Camponotus norvigi stands out within the species group. The color and disposition of the erect hairs covering its entire dorsum separate C. norvigi from C. darwinii. The latter species and C. ursus both have dense pubescence on their mesosoma or gastral dorsum, a character absent in C. norvigi. At first glance, C. norvigi appears closely related to the C. madagascarensis group because of its whitish pilosity, but the two differ in the structure of the mesosoma, which is low and long in the C. madagascarensis group and short and high in the C. darwinii group.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	etymology	Etymology. This species is named in honor of K. Norvig for support in training young scientists in Madagascar.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.taxon	materials_examined	Additional material examined. Province Antananarivo: Analamanga Region, District of Ankazobe, Ambohitantely, 46 km NE of Ankazobe, - 18.198, 47.2815, 701 m, Foret sclerophylle (MG) (CAS); Reserve Speciale d'Ambohitantely, - 18.22444, 47.2774, 1490 m, montane forest (B. L. Fisher et al.) (CAS). Province Antsiranana: 9.2 km WSW Befingotra, Res. Anjanaharibe-Sud, - 14.75, 49.46667, 1200 m, montane rainforest (B. L. Fisher) (CAS); Parc National de Marojejy, Antranohofa, 26.6 km 31 ° NNE Andapa, 10.7 km 318 ° NW Manantenina, - 14.44333, 49.74333, 1325 m, montane rainforest (B. L. Fisher) (CAS); Parc National Montagne d'Ambre [Petit Lac road], - 12.533333, 49.166668, 1125 m, rainforest (R. Harin'Hala) (CAS). Province Fianarantsoa: 2 km NE Anjoma-Ramartina, - 19.63333, 45.96667, 750 m, grassland (PSW) (CAS); 2 km W Andrambovato, along river Tatamaly, - 21.51167, 47.41, 1075 m, cultivated land (tavy), recent 1 - 2 year-old tavy (BLF) (CAS); Ambalavao, - 21.83267, 46.93867, 1020 m, urban / garden (BLF) (CAS); Belle Vue trail, Ranomafana National Park, - 21.2665, 47.42017, 1020 m, mixed tropical forest (R. Harin'Hala) (CAS); Fitovavy Fitovinany Region, District of Ifanadiana Belle vue area, 1200 m S of Ranomafana National Park entrance, - 21.2665, 47.42017, 1018 m, rainforest (MG) (CAS); Fitovavy Fitovinany Region, District of Ifanadiana, 12 km W of Ranomafana, - 21.25083, 47.40717, 1127 m, forest edge, open area, (MG) (CAS); JIRAMA water works near river, Ranomafana National Park, Fianarantsoa Prov., - 21.2485, 47.45217, 690 m, open area near stream (R. Harin'Hala) (CAS); radio tower, Ranomafana National Park, - 21.25833, 47.40717, 1130 m, forest edge, mixed tropical forest, open area (MG) (CAS); Ranomafana, - 21.25, 47.36667 (A. Pauly) (CAS); Ranomafana, nr. Ifanadiana, - 21.26667, 47.45, 610 m, open forest / shrubland (PSW) (CAS); stream area, 900 m E of Isalo National Park Interpretive Center, - 22.62667, 45.35817, 750 m, open area near stream (R. Harin'Hala) (CAS); Vohiparara broken bridge, - 21.22617, 47.36983, 1110 m, high altitude rainforest (R. Harin'Hala) (CAS). Province Mahajanga: Parc National de Namoroka, 16.9 km 317 ° NW Vilanandro, - 16.40667, 45.31, 100 m, tropical dry forest (FGAT) (CAS); Parc National Tsingy de Bemaraha, 2.5 km 62 ° ENE Bekopaka, Ankidrodroa River, - 19.13222, 44.81467, 100 m, tropical dry forest on Tsingy (FGAT) (CAS); Region Sofia, Bemanevika, - 14.337, 48.58874, 1606 m, montane rainforest (BLF) (CAS). Province Toamasina: Ankerana, - 18.40636, 48.80254, 1108 m, montane forest (BLF) (CAS); Corridor Forestier Analamay-Mantadia, Tsaravoniana, - 18.76465, 48.41938, 1039 m, rainforest (BLF) (CAS); Toamasina Foret Ambatovy, 14.3 km 57 ° Moramanga, - 18.85083, 48.32, 1075 m, montane rainforest (B. L. Fisher) (CAS); Manakambahiny Atsinanana, - 17.75, 48.71667, primary forest (A. Pauly) (CAS); P. N. Mantadia, - 18.79167, 48.42667, 895 m, rainforest (H. J. Ratsirarson) (CAS); Torotorofotsy, - 18.77048, 48.43043, 1005 m, montane rainforest (B. L. Fisher et al.) (CAS). Province Toliara: 13 km NW Enakara, Res. Andohahela, - 24.55, 46.8, 1180 m, montane rainforest (B. L. Fisher) (CAS); 2.7 km WNW 302 ° Ste. Luce, - 24.77167, 47.17167, 20 m, littoral rainforest (B. L. Fisher) (CAS); 4 km N Isaka-Ivondro, - 24.76667, 46.86667, 180 m, roadside (PSW) (CAS); 6 km ESE Imonty, Res. Andohahela, - 24.85, 46.75, 1200 m, rainforest (PSW) (CAS); Makay Mts., - 21.227, 45.33222, 475 m, gallery forest on sandy soil (BLF) (CAS); Manatantely, 8.5 km NW Tolagnaro, - 24.9875, 46.92617, 85 m, rainforest (BLF) (CAS); Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, - 24.7585, 46.85367, 275 m, rainforest (BLF) (CAS); Parc National d'Andohahela, Col du Sedro, 3.8 km 113 ° ESE Mahamavo, 37.6 km 341 ° NNW Tolagnaro, - 24.76389, 46.75167, 900 m, montane rainforest (FGAT) (CAS); Res. Andohahela, 6 km SSW Eminiminy, - 24.73333, 46.8, 330 m, rainforest (PSW) (CAS); Reserve Speciale d'Ambohijanahary, Foret d'Ankazotsihitafototra, 35.2 km 312 ° NW Ambaravaranala, - 18.26667, 45.40667, 1050 m, montane rainforest (FGAT) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	description	Figures 5 C, 13 A, 14 A, 54	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	description	Description of minor worker. Large-sized species. Absolute cephalic size (CS: 2.24 0.46; 1.92 - 2.70). In full-face view, head slightly longer than broad, narrower in front than behind, with nearly straight posterior and lateral margins. (CWb / CL: 0.37 + / - 0.03; 0.35 - 0.42). Eyes more elongate, elliptical, not breaking outlines of head (PoOC / CL: 0.08 + / - 0.02; 0.06 - 0.09). Mandibles triangular withsix teeth. Clypeus carinate, anterior clypeal margin entire, projecting into a rectangular lobe with an obtuse anterolateral angle (ClyL / GPD: 0.28 + / - 0.05, 0.24 - 0.32). Antennal scape long, surpassing the occiput by the length of one funiculus (SL / CS: 0.39 + / - 0.05; 0.33 - 0.41). In lateral view, mesosoma short, robust, and depressed; its dorsal outline arcuate; promesonotal suture large, distinctly shiny, and glabrous; humeral angle distinctly rounded; metanotal suture obsolete; propodeal dorsum slightly convex in profile and distinctly shorter than the declivitous face into which it passes via a rounded obtuse angle (MW / ML: 0.22 + / - 0.03; 0.20 - 0.24; MPH / ML: 0.21 + / - 0.05; 0.18 - 0.24). In lateral view, petiolar node squamiform, anterior face confused with its dorsal face, node summit truncate, rounded triangle in dorsal view. Head, mesosoma, and gastral tergite finely and densely punctate. Mandible finely and longitudinally striate with sparse punctuation. Hairs golden yellow, erect to suberect, abundant, bending forward at its midlength on occipital region and mesosomal dorsum, but more abundant on mesonotum and propodeum dorsum; sparse, stiff, pointed, and a different length on gastral tergites. Petiole along its border with a fringe of golden, yellowish hairs not the same length. Body entirely black and a little shiny; masticatory margin reddish to dark brown.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	description	Description of major worker. With characteristics of minor worker, except: head much wider posteriorly (CS: 3.05 + / - 0.41, 2.35 - 3.67; CWb / CL: 1.02 + / - 0.04, 0.96 - 1.08), lateral margin almost straight and tapering anteriorly to the mandibular insertion. Eyes elliptical, moderately sized compared to the head (EL / CS: 0.20 + / - 0.01, 0.1 -- 0.23), placed dorsally next to the vertex (PoOC / CL: 0.23 + / - 0.02, 0.18 - 0.27). Clypeus more rectangular (ClyL / GPD: 0.84 + / - 0.05; 0.73 - 0.91). Antennal scape just surpassing the occipital margin (SL / CS: 0.77 + / - 0.08; 0.68 - 0.91). Mandible strongly built with five large teeth, apical tooth sharp and long, and remainder decreasing in size to the basal margin. Mesosoma, pilosity, and pubescence same as minor worker. Head shiny black; anterior portion of head, including frontal area, finely alveolate punctate.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	distribution	Distribution and biology. Camponotus nossibeensis is an endemic species mostly known from the northern part of Madagascar, from Nosy Faly to Ambanja. This species seems very adaple because it has been collected from both humid and tropical dry forest, at altitudes 7 - 780 m (Fig. 58 C). The data indicate that individual workers forage on the ground and on lower vegetation, while nests are mostly found in dead twigs above ground, rotten logs, dead tree stumps, and underground.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	discussion	Discussion. The combination of large size, big eyes, vertical declivitous face of propodeum, dense pilosity on mesosoma and gaster, and alveolate sculpture make C. nossibeensis a very distinctive species.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.taxon	materials_examined	Additional material examined. Province Antsiranana: Foret Ambato, 26.6 km 33 ° Ambanja, - 13.4645, 48.55167, 150 m, rainforest (B. L. Fisher) (CAS); Nosy faly, Tafiambotry, 35.3 km N Ambanja, - 13.3654, 48.48775, 7 m, littoral rainforest (BLF) (CAS); Ankobahoba, 32.3 km N Ambanja, - 13.39166, 48.48249, 41 m, disturbed littoral rainforest (BLF) (CAS); R. S. Manongarivo, 10.8 km 229 ° SW Antanambao, - 13.96167, 48.43333, 400 m, rainforest (CAS); R. S. Manongarivo, 12.8 km 228 ° SW Antanambao, - 13.97667, 48.42333, 780 m, rainforest (CAS); R. S. Manongarivo, 20.4 km 219 ° SW Antanambao, - 14.04667, 48.40167, 1860 m, montane rainforest (CAS); R. S. Manongarivo 17.3 km 218 ° SW Antanambao, - 14.02167, 48.41833, 1580 m, montane rainforest (B. L. Fisher) (CAS); Nosy Faly, - 13.36435, 48.49137, 40 m, open secondary vegetation (BLF) (CAS); Nosy Faly, - 13.3624, 48.49101, 15 m, open secondary vegetation (BLF) (CAS); Sahamalaza Peninsula, Foret d'Anabohazo, 21.6 km 247 ° WSW Maromandia, - 14.30889, 47.91433, 120 m, tropical dry forest (FGAT) (CAS); Nosy Be, Reserve Naturelle Integrale de Lokobe, 6.3 km 112 ° ESE Hellville, - 13.41933, 48.33117, 30 m, rainforest (FGAT) (CAS); Sahamalaza National Park, - 14.16361, 47.70277, 192 m, subhumid forest (MG) (CAS); Nosy Faly, - 13.36255, 48.49104, 19 m, open secondary vegetation (BLF) (CAS). Province Mahajanga: Parc National Tsingy de Bemaraha, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, - 19.14194, 44.828, 50 m, tropical dry forest (FGAT) (CAS); Parc National Tsingy de Bemaraha, 10.6 km ESE 123 ° Antsalova, - 18.70944, 44.71817, 150 m, tropical dry forest on Tsingy (FGAT) (CAS); Parc National Tsingy de Bemaraha, 2.5 km 62 ° ENE Bekopaka, Ankidrodroa River, - 19.13222, 44.81467, 100 m, tropical dry forest on Tsingy (FGAT) (CAS); Reserve Speciale Marotandrano, Marotandrano 48.3 km S Mandritsara, - 16.28322, 48.81443, 865 m, transitional humid forest (BLF) (CAS); Ampamakiambato, 45 km SW Ambanja, - 13.97545, 48.15929, 145 m, disturbed forest, in tsingy (BLF) (CAS); Manerinerina, 76.6 km N Antsohihy, - 14.10744, 48.11046, 247 m, disturbed forest (BLF) (CAS); Reserve forestiere Beanka, 50.7 km E Maintirano, - 17.88021, 44.46877, 140 m, tropical dry forest on tsingy (BLF) (CAS); Reserve forestiere Beanka, 54.3 km E Maintirano, - 18.06009, 44.54086, 262 m, tropical dry forest on tsingy (BLF) (CAS); Melaky Region, District of Besalampy, Marofototra dry forest, 17 km W of Besalampy, - 16.72167, 44.42367, 51 m, dry wash in the dry forest (MG) (CAS); Bemeraha, 9 km E. Antsalova, - 18.65, 44.71667 (D. C. Lees) (PSWC); Foret de Tsimembo, 8.7 km 336 ° NNW Soatana, - 19.02139, 44.44067, 20 m, tropical dry forest (FGAT) (CAS). Province Toliara: Belalanda, - 23.29833, 43.64648, 5 m, spiny forest / thicket, wetlands (Frontier Wilderness Project) (CAS); near road, Zombitse National Park, - 22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin'Hala) (CAS); near ANGAP office, Zombitse National Park, - 22.8865, 44.69217, 840 m, deciduous spiny forest (R. Harin'Hala) (CAS); 15 km E Sakaraha, - 22.9, 44.68333, 760 m, tropical dry forest (PSW) (PSWC); Makay Mts., - 21.30997, 45.12946, 590 m, Dry forest on sandy soil (BLF) (CAS); Makay Mts., - 21.227, 45.33222, 475 m, gallery forest on sandy soil (BLF) (CAS); Makay Mts., - 21.21013, 45.35462, 540 m, gallery forest with bamboo (BLF) (CAS); Parc National de Zombitse, 17.7 km 98 ° E Sakaraha, - 22.88833, 44.70167, 760 m, tropical dry forest (FGAT) (CAS); 9 km NNW Ranohira, P. N. Isalo, - 22.48333, 45.38333, 800 m, tropical dry forest (PSW) (PSWC).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	description	Figures 14 B, 16 D, 18 C, D, 55	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 1.59 + / - 0.22, 1.43 - 1.77). In full-face view, head longer than broad, narrower in front than in back, with feebly convex occipital and lateral sides (CWb / CL: 0.37 + / - 0.03; 0.35 - 0.40). Eyes circular, placed much closer to the lateral sides (PoOC / CL: 0.08 + / - 0.01, 0.07 - 0.09). Mandibles triangular with six teeth. With head in full-face view, clypeus not carinate, its anterior margin produced, forming a short rectangular lobe with a sharp, lateral angle. Antennal scape circular, long, surpassing the occiput by the length of one basal funiculus (SL / CS: 0.39 + / - 0.03, 0.37 - 0.41). In lateral view, anterodorsal corner of pronotum broadly rounded, mesosomal dorsal almost straight, propodeal dorsum shorter than declivity, both straight in profile and meeting at a blunt angle (MW / ML: 0.21 + / - 0.02, 0.20 - 0.23; MPD / ML: 0.24 + / - 0.02, 0.22 - 0.26). Promesonotal and metanotal sutures form bold, polished lines. In lateral view, petiolar node cuneate, with a short, vertical, anterior face which reaches toward the flattened posterior faces. Head and mesosoma finely alveolate, gaster more finely strigulate. Mandible finely foveate with sparse punctures. Pilosity of dorsal head and body consists of fine and pointed brownish yellow hairs, long and suberect on frontal area and mesosomal dorsum; mostly short and sparse on gastral tergites; petiolar dorsum edge with 4 to 5 pairs of brownish yellow hairs. Body entirely matte black, apical portion of mandible, basitarsi, and funiculus reddish brown.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	description	Description of major worker. Characteristics of minor workers, except: head somewhat cordate, with nearly straight posterior margin and slightly convex lateral sides (CS: 2.76 + / - 0.12, 2.59 - 2.85; CWb / CL: 1.05 + / - 0.02, 1.02 - 1.07). Eyes elliptical, placed dorsally midway from the occipital margin (PoOC / CL: 0.24 + / - 0.01, 0.23 - 0.25). Clypeus much more quadrate in full-face view with truncate anterior margin (ClyL / GPD: 0.76 + / - 0.13, 0.57 - 0.86). Antennal scape short, just surpassing the occipital border (SL / CS: 0.74 + / - 0.03, 0.72 - 0.79). Dorsal outline of mesosoma interrupted by pronotal and mesonotal sutures, basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely punctate, frontal area and vertex with deep punctures.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	distribution	Distribution and biology. Camponotus radovae is mainly distributed in the southern part of Madagascar in Zombitse National Park (Fig. 58 D). This species is encountered in native forest habitats such as deciduous dry forest, tropical dry forests, gallery forests, and spiny forests and thickets. It was sampled mostly from Malaise traps and by beating lower vegetation, on the ground, and twice on leaf litter at altitudes 20 - 840 m.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	discussion	Discussion. Within the Camponotus darwinii group, Camponotus radovae cannot be confused with C. darwinii or C. themistocles since the latter two differ by size and pilosity color. However, C. radovae has reduced pubescence and is smaller in size.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.taxon	materials_examined	Additional material examined. Province Toliara: Atsimo Andrefana Region, District of Sakaraha, Zombitse National Park; 900 m N from ANGAP Entrance office, - 22.8405, 44.73117, 823 m, spiny deciduous forest (MG) (CAS); Atsimo Andrefana Region, District of Toliara II, Mikea decidious dry forest 3 km N Andranomavo village, - 22.90367, 43.4755, 30 m, Deciduous dry forest (MG) (CAS); Atsimo Andrefana Region, District of Toliara II, Mikea spiny forest 8 km N Andranomavo village, - 22.91333, 43.39883, 37 m, spiny forest (MG) (CAS); Mikea Forest, deciduous dry forest, - 22.90367, 43.4755, 30 m, deciduous dry forest (R. Harin'Hala) (CAS); Mikea Forest, spiny forest, - 22.91333, 43.48222, 37 m, spiny forest (R. Harin'Hala) (CAS); near ANGAP office, Zombitse National Park, - 22.8865, 44.69217, 840 m, deciduous spiny forest (R. Harin'Hala) (CAS); near road, Zombitse National Park, - 22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin'Hala) (CAS); Parc National de Zombitse, 17.7 km 98 ° E Sakaraha, - 22.88833, 44.70167, 760 m, tropical dry forest (FGAT) (CAS); Parc National de Zombitse, 19.8 km 84 ° E Sakaraha, - 22.84333, 44.71, 770 m, tropical dry forest (FGAT) (CAS); Ranobe, - 23.03975, 43.6109, 30 m, spiny forest / thicket (Frontier Project) (CAS); Ranobe, - 23.03957, 43.61032, 30 m, spiny forest / thicket (Frontier Project) (CAS); Ranobe, - 23.03957, 43.61017, 30 m, spiny forest / thicket (Frontier Project) (CAS); Ranobe, - 23.03883, 43.60982, 30 m, gallery forest (Frontier Wilderness Project) (CAS); Ranobe, - 23.03952, 43.61015, 20 m, gallery forest (Frontier Wilderness Project) (CAS); Ranobe, - 23.03832, 43.6096, 20 m, gallery forest (Frontier Wilderness Project) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	description	Figures 5 D, 15 B, 16 C, 18 A, B, 56	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	description	Description of minor worker. Medium-sized. Absolute cephalic size (CS: 1.82 + / - 0.37; 1.53 - 1.98). In full-face view, head slightly longer than broad, a little narrower in front than in back, with straight occipital and feebly convex lateral margins (CWb / CL: 0.36 + / - 0.03; 0.34 - 0.38). Eyes circular, placed much closer to the lateral sides (PoOC / CL: 0.07 + / - 0.01; 0.07 - 0.08). Mandibles triangular with sixteeth. With head in full-face view, anterior clypeal margin produced, entire, and angularly projecting in the middle. Antennal scape circular, long, surpassing the occiput by the lengths of two basal funiculi (SL / CS: 0.41 + / - 0.05; 0.38 - 0.44). In lateral view, anterodorsal corner of pronotum broadly rounded, mesosomal dorsum moderately arcuate, propodeum dorsum a little shorter than the declivity, each straight in profile and meeting at an obtuse angle (MW / ML: 0.21 + / - 0.02, 0.20 - 0.22; MPD / ML: 0.24 + / - 0.03; 0.22 - 0.26). In lateral view, petiolar node wedge-shaped, with feebly convex anterior and flattened posterior faces. Ants entirely reticulate-punctate. Mandible finely striolate with sparse punctures. Pilosity and pubescence brownish yellow, the former long and curved forward especially on vertex, frontal area, and mesosomal dorsum; mostly short and sparse on gastral tergites; petiolar dorsum edge with four or five pairs of brownish yellow hairs. Body color matte black, apical portion of mandible reddish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	description	Description of major worker. Characteristics of minor workers, except: head somewhat cordate, with nearly straight posterior margin and slightly convex lateral sides (CS: 2.68 + / - 0.36, 2.26 - 3.15; CWb / CL: 1.05 + / - 0.04, 0.99 - 1.11). Eyes elliptical, placed dorsally midway from the occipital margin (PoOC / CL: 0.23 + / - 0.01, 0.22 - 0.25). Clypeus much more quadrate in full-face view with truncate anterior margin (ClyL / GPD: 0.82 + / - 0.07, 0.73 - 0.93). Antennal scape short, just surpassing the occipital border (SL / CS: 0.75 + / - 0.03, 0.71 - 0.79). Dorsal outline of mesosoma interrupted by pronotal and mesonotal sutures, basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely punctate, frontal area and vertex with deep punctures.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	distribution	Distribution and biology. Camponotus themistocles has been collected at three localities in southern Madagascar (Fig. 58 E): two are littoral forest and one is gallery forest. All sites are at low altitudes from 10 to 30 meters. Camponotus themistocles is a ground nester; most collections have been made from sifted litter, rotten logs, and dead twigs above the ground.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	discussion	Discussion. Camponotus themistocles differs from C. radovae by its larger size and the sculpture of the lateral sides o f its mesosoma, which is finely reticulate-punctate in the former and widely striolate in the later. In addition, in lateral view, the petiole scale of C. themistocles has an acute summit but is blunt in C. radovae. Pubescence is long and abundant in C. themistocles, and short and dilute in C. radovae. Morphometrical analysis reveals that C. themistocles and C. ellioti are the same size, but differ in integument coloration and pilosity pattern on the gastral tergite.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.taxon	materials_examined	Additional material examined. Province Toliara: Foret de Petriky, 12.5 km W 272 ° Tolagnaro, - 25.06167, 46.87, 10 m, littoral rainforest (B. L. Fisher) (CAS); Foret Mandena 8.5 km N Tolagnaro, - 24.95267, 47.0025, 20 m, littoral rainforest (BLF) (CAS); Mandena, 8.4 km NNE 30 ° Tolagnaro, - 24.95167, 47.00167, 20 m, littoral rainforest (B. L. Fisher) (CAS); Ranobe, - 23.04085, 43.61012, 30 m, gallery forest (Frontier Wilderness Project) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	description	Figures 43 B, 45 B, 46 A, 69	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 0.79 + / - 0.04; 0.77 - 0.81). In full-face view, head ovate, usually somewhat longer than broad, lateral margin of head straight and tapering to front (CWb / CL: 0.29 + / - 0.02, 0.28 - 0.29). Eyes subelliptical, protruding, their borders smoothly aligned to lateral margins of head (PoOC / CL: 0.09 + / - 0.00, 0.08 - 0.09). Mandibles triangular with six teeth. Anterior clypeal margin not produced, entire, broadly rounded. Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL / CS: 0.41 + / - 0.03, 0.40 - 0.42). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeal dorsum raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW / ML: 0.19 + / - 0.02, 0.17 - 0.20; MPH / ML: 0.13 + / - 0.01, 0.13 - 0.14). In lateral view, petiole cuneate with convex anterior face and flat dorsal face. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins of petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remaining segment dark brown to black.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	distribution	Distribution and biology. This new species seems to prefer deciduous forest habitats such as tropical dry forest, gallery forest, spiny forest or thicket, and transitional forest, but is much rarer in humid habitats at lower altitudes (Fig. 72 D).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	discussion	Discussion. Camponotus tsimelahy may be confused with C. repens and C. ihazofotsy; all three species share the same sculpturation, form of petiole, and type of suberect setae. However, in profile, the dorsal outline of the mesosoma presents a slightly concave metanotal groove. See the discussion section under C. ihazofotsy.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	etymology	Etymology. The species epithet is in reference to the type locality.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	materials_examined	Additional material examined. Province Toliara: 18.4 km N Amboasary, - 24.87367, 46.39767, 85 m, spiny forest / thicket (BLF) (CAS); Anosy Region, District of Amboasary, Andohaela National Park Parcelle III, Ihazofotsy, 32 km NE Amboasary, - 24.83083, 46.53617, 58 m, dry forest, spiny forest (Mike Irwin, Frank Parker, Rin'ha) (CAS); Anosy Region, District of Fort-Dauphin, Andohaela National Park Parcelle II, Tsimelahy, 42 km W of Fort-Dauphin, - 24.93683, 46.62667, 176 m, transitional forest (Mike Irwin, Frank Parker, Rin'ha) (CAS); Ifaty 22 km N, - 23.18333, 43.61667, 30 m, beach dunes (M. E. Irwin and E. I. Schlinger) (CAS); Ihazofotsy - Parcel III, Andohahela National Park, transitional forest, - 24.83483, 46.48683, 80 m, transition between spiny and dry deciduous forests (M. E. Irwin, F. D. Parker, R. Harin'Hala) (CAS); Parc National d'Andohahela, Foret d' Ambohibory, 1.7 km 61 ° ENE Tsimelahy, 36.1 km 308 ° NW Tolagnaro, - 24.93, 46.6455, 300 m, tropical dry forest (FGAT) (CAS); Parc National d'Andohahela, Foret de Manatalinjo, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, - 24.81694, 46.61, 150 m, spiny forest / thicket (FGAT) (CAS) Tsimelahy - Parcel II, Andohahela National Park, transitional forest, - 24.93683, 46.62667, 180 m, transitional forest (M. E. Irwin, F. D. Parker, R. Harin'Hala) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	description	Figures 2 C, 16 A, 17 A, 57	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	description	Description of minor worker. Medium-sized species. Absolute cephalic size (CS: 1.54 + / - 0.19; 1.43 - 1.64). In full-face view, head somewhat longer than broad, narrower in front than behind, with slightly convex lateral and posterior sides (CWb / CL: 0.37 + / - 0.02; 0.36 - 0.38). Eyes elliptical, sublateral at midpoint from the lateral sides (PoOC / CL: 0.07 + / - 0.02; 0.06 - 0.08). Mandibles triangular with six teeth. Clypeus not carinate, produced into a short, rectangular lobe. Antennal scape long, surpassing the occiput by the length of one basal funiculus. (SL / CS: 0.38 + / - 0.02; 0.37 - 0.39). In lateral view, dorsal contour of mesosoma smoothly convex, humeral angle broadly rounded; in dorsal view, mesonotal suture distinct but not impressed, mesonotal suture obsolete so that mesonotum and propodeum are fused together, propodeum with convex base and sloping declivity (MPH / ML: 0.18 + / - 0.02; 0.16 - 0.19). Petiole narrow, cuneate in profile, with short anterior face tapering dorsally to the flattened posterior face, its border rather sharp, produced upwards as a blunt angle in the middle. Head dorsum finely reticulate-punctate with shallow, sparse punctures; occipital region transversally striate-reticulate; lateral face of mesosoma, declivitous face, and petiolar face finely strigulate; legs finely reticulate; dorsum of mesosoma finely reticulate-striate-punctate with sparse excavation from which one suberect setae arises. Dorsum of gastral tergite finely striate-reticulate transversally, with sparse, small punctures. Mandible finely rugose with sparse, large punctures. Hairs golden yellow, the former abundant, long, and bending forward on entire mesosoma dorsum; suberect, short, and sparsely distributed on gastral tergites; the latter short and conspicuous on the abdominal segment, five pairs of erect hairs present on vertex. Body shiny black; scape, two basal funiculi, mandible, tarsi, and tibiae reddish.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	description	Description of major worker. Characteristics of minor workers, except: head as broad as long, with occipital and lateral margin almost straight (CS: 2.14 + / - 0.15, 2.00 - 2.39; CWb / CL: 0.89 + / - 0.07, 0.84 - 1.03). Eyes circular, placed dorsally next to the vertex (PoOC / CL: 0.23 + / - 0.01, 0.21 - 0.24). Anterior clypeal margin forms a short, rounded lobe (ClyL / GPD: 1.03 + / - 0.41, 0.73 - 1.78). Antennal scape short, just reaching the occipital border (SL / CS: 0.82 + / - 0.03, 0.78 - 0.86). Dorsal outline of mesosoma almost flat, propodeum dorsum naked and the same length as the sloping declivity.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	distribution	Distribution and biology. Camponotus ursus is found in two different habitats: primary forest in the eastern portion and urban / garden areas in the central highlands of Madagascar (Fig. 58 F). It is found foraging on low vegetation or inside branches above the ground. It occurs at altitudes above 1,200 meters.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	discussion	Discussion. Camponotus ursus is recognizable within the C. darwinii species group on the basis of its distinct mesosomal pilosity. In addition, it is the only species with reddish brown basitarsi. However, C. ursus is unlikely be confused with C Camponotus darwinii for several reasons. First, the body of the latter is much larger. Second, the head and gastral segment of C. ursus are covered with fine, short, sparse, and yellowish setae, giving the ant a glossy appearance that is completely different than that of C. darwinii.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.taxon	materials_examined	Additional material examined. Province Antananarivo: Ambatomanjaka; Miarinarivo, - 18.766947, 46.869107, 1343 m (MHNG); Ankazobe, - 18.31617, 47.11583, 1241 m (BLF) (CAS). Province Toamasina: Manakambahiny Atsinanana, - 17.75, 48.71667 (A. Pauly) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	description	Figures 11 A, B, 33 C, D, 72	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	description	Description of minor worker. Medium-sized. Absolute cephalic size (CS: 1.02 + / - 0.28; 0.92 - 1.14). In full-face view, head subquadrate (CWb / CL: 0.39 + / - 0.01, 0.39 - 0.40), posterior margin of head broadly convex, lateral margin of head slightly convex. Eyes sub-elliptical, placed laterally, closer to the posterior margin (PoOC / CL: 0.09 + / - 0.01; 0.08 - 0.09). Mandibles triangular with five teeth. Anterior clypeal margin rounded and not produced anteriorly (ClyL / GPD: 0.24 + / - 0.01). Antennal scape virtually circular, surpassing posterior cephalic margin by the length of two basal flagella (SL / CS: 0.34 + / - 0.04; 0.33 - 0.36). In lateral view, metanotal groove deeply impressed; pronotum, mesonotum, and propodeum dorsum marginated laterally; mesonotum dorsum inclined posteriorly; propodeum dorsum slightly concave, its lateral carina forming a blunt tubercle with the lateral margin of the declivity (MPD / ML: 0.25 + / - 0.02; 0.24 - 0.25). In lateral view, petiolar node strongly convex anterodorsally and flat posteriorly, node summit convex. Head finely areolate and mesosoma feebly areolate-costulate, gastral tergite finely imbricate. Mandible glabrous. Vertex with three pairs of filiform erect setae; mesonotum and propodeum dorsum with single pair of whitish setae; gastral tergite with short, whitish setae aligned transversally on the middle length of each segment; petiolar node with two or three pairs of whitish setae on its dorsum. Body uniformly dark blackish brown to black, appendages brownish, mandible light brown.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	description	Description of major worker. With characteristics exactly the same as minor worker, but head more trapezoidal (CS: 1.74 + / - 0.04, 1.71 - 1.77).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	distribution	Distribution and biology. Camponotus zoro has been collected by beating low vegetation, sifting leaf litter, and baiting with honey and tuna fish. One worker has been found in Malaise traps from deciduous forest in southern Madagascar (Fig. 72 E). In addition, this species has been collected from montane rainforest and Uapaca woodland.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	discussion	Discussion. Camponotus zoro is characterized by a shallow but distinct metanotal groove, and should not be confused with any other species because of the margination of its mesosoma.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	etymology	Etymology. The new species is named after the structure of the mesosoma; " Camponotus zoro " means angle.	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.taxon	materials_examined	Additional material examined. Province Antananarivo: Navoatra I Non Protected Area, 7.64 km NW Arivonimamo, - 18.97806, 47.11929, 1373 m, Uapaca woodland (ARA) (CAS); Ankalalahana, - 19.0531, 47.11304, 1314 m, Uapaca woodland (H. E. Marti et al.) (CAS); 3 km 41 ° NE Andranomay, 11.5 km 147 ° SSE Anjozorobe, - 18.47333, 47.96, 1300 m, montane rainforest (FGAT) (CAS); Reg. Analamanga, St. Forestiѐre Mandraka, - 18.9183, 47.91687, 1285 m, montane rainforest (B. L. Fisher, F. A. Esteves et al.) (CAS). Province Fianarantsoa: Foret d' Atsirakambiaty, 7.6 km 285 ° WNW Itremo, - 20.59333, 46.56333, 1550 m, montane rainforest (FGAT) (CAS). Province Toamasina: Reserve Speciale Ambatovaky, Sandrangato river, - 16.7755, 49.26427, 430 m, rainforest (BLF) (CAS).	en	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
