identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B07D7FC5250752D48742BC8001EFFC5C.text	B07D7FC5250752D48742BC8001EFFC5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus andrianjaka Rasoamanana & Fisher 2022	<div><p>Camponotus andrianjaka sp. nov.</p> <p>Figures 8D, 39B, 61A-C, 62</p> <p>Holotype worker.</p> <p>Madagascar, Province Fianarantsoa, Isalo III National Parc, 12.02 km SW Ranohira, -22.6158, 45.31084, 870 m, Bismarckia woodland, ex soil, 15-24 February 2010 (ARA), collection code: ARA0306, specimen code: CASENT0243690 (CAS). Paratype. one worker with same data as holotype but specimen coded as CASENT0243954 (CAS).</p> <p>Worker diagnosis.</p> <p>Camponotus andrianjaka can be easily distinguished from other species of the Camponotus ellioti group by the following character set: body bicolored, head and mesosoma reddish brown, gaster dark brown to black; anterior clypeal margin rounded, not produced; propodeum dorsum slightly concave at the level of metanotal suture; petiole nodiform and surmounted with standing, whitish hairs on its posterior margin.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 0.84 ± 0.07; 0.81-0.88). In full-face view, head subovate, usually somewhat longer than broad, posterior cephalic margin roundly convex; lateral margin of head straight and tapering to front (CWb/CL: 0.31 ± 0.01, 0.30-0.32). Eyes subelliptical, protruding, its border smoothly aligned to lateral margins of head (PoOC/CL: 0.08 ± 0.01, 0.08-0.09). Mandibles triangular with six teeth. Clypeus carinate, anterior clypeal margin with rounded triangular projection (ClyL/GPD: 0.24 ± 0.01, 0.23-0.25). Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL/CS: 0.36 ± 0.04, 0.35-0.38). In lateral view, mesosoma without anterolateral margination, promesonotum convex, mesopropodeal suture shallowly impressed, propodeum rather angulate with slightly concave basal and declivitous surface (MW/ML: 0.19 ± 0.02, 0.18-0.21, MPH/ML: 0.15 ± 0.01, 0.14-0.15). In lateral view, petiole nodiform, higher than long, its short anterior face rounding gradually to the convex posterior faces. Head, especially anterior portion, finely reticulate-areolate, gaster and mesosoma finely imbricate. Mandibles finely reticulate-punctate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; three to four pairs on each side of lateral corner of propodeum; four pairs transversely arranged on petiolar posteromargin; ground pubescence delicate. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remainder of the segment dark brown to black.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head as long as broad, with rather straight posterior and lateral borders (CS: 1.15 ± 0.05, 1.12-1.20, CWb/CL: 0.84 ± 0.01, 0.84-0.85). Eyes circular, placed dorsally next to lateral borders (PoOC/CL: 0.26 ± 0.03, 0.23-0.27). Anterior clypeal margin forms a rounded rectangular lobe (ClyL/GPD: 0.89 ± 0.05, 0.83-0.92). Antennal scape short, just reaching the occipital border (SL/CS: 0.70 ± 0.02, 0.69-0.72). Mesosoma the same as minor worker.</p> <p>Distribution and biology.</p> <p>Camponotus andrianjaka is a very distinctive species that appears endemic to Bismarckia woodland, shrubland, Uapaca woodland, and savannah woodland in the southern part of Madagascar (Fig. 70D). It can be found also on eucalyptus plantations and grassland. Altitude: 1300-1987 m.</p> <p>Discussion.</p> <p>At first sight, Camponotus andrianjaka may be confused with another new species, Camponotus MG132, subgenus Camponotus Myrmonesites, but can be separated easily with mesonotum that is gradually rounded in dorsal view; petiole higher than long, its dorsolateral margin surmounted with white hairs; presence of ground pubescence; and gaster all black. In addition, C. MG132 has been collected from rainforest and montane rainforest and absent from woodland.</p> <p>Etymology.</p> <p>This new species is named after the collector Andrianjaka Ravelomanana.</p> <p>Additional material examined.</p> <p>Province Fianarantsoa: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Ampandravelo</a> III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 10.72 km NE Ranohira, -22.53944, 45.51497, 869 m, shrubland (ARA) (CAS); Ampangabe III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 21.26 km W Itremo, -20.6125, 46.60883, 1412 m, savannah woodland (ARA) (CAS); Ampangabe VI <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 21.16 km W Itremo, -20.61444, 46.6104, 1379 m, shrubland (ARA) (CAS); Ampangabe VII <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 21.2 km W Itremo, -20.61417, 46.60989, 1420 m, shrubland (ARA) (CAS); Ampotoampoto II National Parc, 8.07 km NW Ilakaka, -22.62806, 45.18843, 919 m, savannah woodland (ARA) (ATOL UCDC); Ampotoampoto III National Parc, 7.91 km NW Ilakaka, -22.62944, 45.189, 919 m, savannah woodland (ARA) (ATOL voucher CAS); Antohatsahomby III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 22.79 km NW Itremo, -20.54806, 46.58599, 1499 m, Uapaca woodland (ARA) (CAS); Antohatsahomby V Non Protected Area, 22.63 km NW Itremo, -20.56722, 46.57923, 1726 m, Uapaca woodland (ARA) (CAS); Isalo III National Parc, 12.02 km SW Ranohira, -22.61583, 45.31084, 870 m, Bismarckia woodland (ARA) (CAS); Isalo IV National Parc, 12 km SW Ranohira, -22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (ATOL USNM); Isalo IV National Parc, 12 km SW Ranohira, -22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (CAS); Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, -20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (ATOL CAS).</p> </div>	http://treatment.plazi.org/id/B07D7FC5250752D48742BC8001EFFC5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
E5B75706343E523FBFA2FCE6EDDE3A5B.text	E5B75706343E523FBFA2FCE6EDDE3A5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus antsaraingy Rasoamanana & Fisher 2022	<div><p>Camponotus antsaraingy sp. nov.</p> <p>Figures 4 A-D, 13B, 19A, 51</p> <p>Holotype worker.</p> <p>Madagascar, Province Antsiranana, Antsaraingy, -12.90362, 49.65921, 66 m, littoral forest, ground nest, 21 October 2013 (BLF), collection code: BLF32314, specimen code: CASENT0371032 (CAS). Paratypes. Two workers with same data as holotype but collection code: BLF32341, specimen code: CASENT0371026 and CASENT0840640 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument entirely black, coarsely sculptured; gaster densely covered with enlarged white hairs; anterior clypeal margin with a rectangular lobe, broadly rounded medially; antennal scape flattened longitudinally; appressed, pointed-oval, whitish hairs present next to the antennal insertion; dorsal face of propodeum distinctly shorter than declivitous face; only three first gastral tergites visible dorsally.</p> <p>Description of minor worker.</p> <p>Large-sized species. Absolute cephalic size (CS: 2.14 ± 0.38, 2.00-2.33). In full-face view, head elongate (CWb/CL: 0.34 ± 0.03; 0.32-0.35); posterior margin of head straight, lateral margin of head even straighter, feebly converging toward base of mandible. Eyes elliptical, sublateral, and placed next to the vertex (PoOC/CL: 0.08 ± 0.02; 0.07-0.09). Mandibles triangular with six teeth. Clypeus strongly carinate throughout its length, with produced, angular, anterior margin, lateral corner angulate and median portion a rounded triangle (ClyL/GPD: 0.25 ± 0.05, 0.22-0.28). Antennal scape longitudinally flattened, its base scarcely enlarged (SL/CS: 0.44 ± 0.04, 0.42-0.45). In lateral view, mesosoma short and high, its dorsum smoothly arcuate, metanotal suture obsolete. Humeral angle rounded. Dorsal and declivitous face of propodeum difficult to distinguish and forming a rounded angle (MW/ML: 0.27 ± 0.02; 0.26-0.28; MPH/ML: 0.21 ± 0.02, 0.20-0.22). In lateral view, petiole squamiform with sharp edge, anterior face short and converging to the highest point of the petiole, posterior face straight. Ventral face of femora marginate laterally, the outer margin of tibia with long, whitish setae. Head and mesosoma coarsely reticulate-punctate, legs and gastral tergite finely imbricate. Mandible finely foveolate with sparse punctures. Whitish, suberect hairs present on occipital portion, to the anterior margin of eyes, and on posteroventral head surface, scarcely distributed on mesosoma dorsum but much closer on declivity; decumbent, spatulate, whitish hair on the ventral section of gena, below the antennal socket, on the posterolateral margin of clypeus, promesonotum dorsum, petiolar node, and gastral tergite; suberect and decumbent whitish hairs present on gastral segment. Entire body blackish except the mandible testaceous to reddish.</p> <p>Description of major worker.</p> <p>With characteristics of minor workers, except: head more cordate (CS: 3.32 ± 0.21, 2.94-3.55) (CWb/CL: 0.96 ± 0.01, 0.95-0.99); lateral margins slightly convex and tapering anteriorly. Eyes elliptical, smaller compared to head size (EL/CS: 0.20 ± 0.01; 0.20-0.21), placed dorsally next to posterior margin of frontal carina. Antennal scape short and not exceeding posterior margin of head (SL/CS: 0.81 ± 0.04; 0.76, 0.88); mandible more robust. Mesosoma and pilosity same as minor worker.</p> <p>Distribution and biology.</p> <p>This new species is only known from Antsaraingy, a littoral forest, from 66 to 90 m in elevation, located in the northern portion of Madagascar (Fig. 70A). Nests were found underground and inside termite mounds; four colonies were collected to represent this species.</p> <p>Discussion.</p> <p>Despite its large size, Camponotus antsaraingy may be confused with Camponotus mita and Camponotus voeltzkowii because in these three species the propodeal face of the propodeum is more reduced than the declivitous face. In C. antsaraingy the antennal segments are entirely flattened while in C. mita only the basal half is flattened; C. voeltzkowii has circular antennal segments. The pubescence on the gastral segment consists of thick, decumbent hair of the same type as the pilosity in C. antsaraingy, with short and filiform hairs on the other two species.</p> <p>Etymology.</p> <p>The species epithet is in reference to the type locality.</p> <p>Additional material examined.</p> <p>Province Antsiranana: Antsaraingy, -12.90665, 49.6606, 90 m, littoral forest, BLF (CAS).</p></div> 	http://treatment.plazi.org/id/E5B75706343E523FBFA2FCE6EDDE3A5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
B1BA149DAF575670859EC43BF1DA7E54.text	B1BA149DAF575670859EC43BF1DA7E54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus chrislaini Rasoamanana & Fisher 2022	<div><p>Camponotus chrislaini sp. nov.</p> <p>Figures 2D, 7C-D, 41D, 59</p> <p>Holotype worker.</p> <p>Madagascar, Province Antsiranana, Forêt d’Andavakoera, 21.4 km 75° ENE Ambilobe; 4.6 km 356° N Betsiaka, -13.11833, 49.23, 425 m, rainforest, ex rotten log, 16 December 2003 (BLF), collection code: BLF10296, specimen code: CASENT0498999 (CAS). Paratypes. Nine workers with the same data as holotype, CASENT0499000, CASENT0499001, CASENT0499002 (three workers in each pin) (CAS).</p> <p>Worker diagnosis.</p> <p>Camponotus chrislaini can be easily distinguished from the other members of the Camponotus efitra group by its mostly smooth and shiny sculpture; head more or less globose in full-face view; sigmoid frontal carina, and wavy propodeum dorsum.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 0.95 ± 0.17; 0.87-1.02). In full-face view, head ovoid (CWb/CL: 0.34 ± 0.03; 0.32-0.35); posterior margin rounded, lateral margin of head convex. Eyes circular, protruding, placed midway from the occipital corner (PoOC/CL: 0.12 ± 0.03; 0.11-0.13). Mandibles triangular with five teeth. Clypeus transversally trapezoidal, its anterior border broadly rounded, not angular. With head in full-face view, anterior clypeal margin rounded, not projecting (ClyL/GPD: 0.26 ± 0.03, 0.25-0.29). Antennal scape flattened and lobulated at the base, long, surpassing the occiput by the length of three basal funiculi (SL/CS: 0.46 ± 0.03; 0.45-0.48). In lateral view, anterodorsal corner of pronotum tuberculate, pronotal suture distinct, mesonotal suture distinctly impressed so that the anterior portion of propodeum is raised to a blunt edge in profile, and suture forms a concavity (MW/ML: 0.22 ± 0.02, 0.21-0.23; MPH/ML: 0.17 ± 0.02, 0.17-0.18). In lateral view, petiolar node nodiform, higher than long. Entire body, including mandible, smooth and shiny. One pair of standing hairs on angle formed by mesonotal dorsum and propodeum, head with scattered, subdecumbent, filiform brown hairs, scape with decumbent setae. Ant brownish; mandible, clypeus, and pronotum light brown; mid- and forecoxae, apical femora, basal tibiae, and apical portion of third and fourth tergite whitish.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head quadrate with straight margins (CS: 1.67 ± 0.06, 1.57-1.74; CWb/CL: 0.92 ± 0.03, 0.90-0.96). Eyes circular, closed to the lateral margin (PoOC/CL: 0.29 ± 0.02, 0.27-0.31). In profile, clypeus distinctly truncate (ClyL/GPD: 0.92 ± 0.03, 0.86-0.94), its anterior margin with a short concavity in full-face view. Antennal scape same as minor but short, just reaching the occipital border (SL/CS: 0.77 ± 0.04, 0.74-0.83). Promesonotum forms a unique dome followed by an impressed mesonotal suture, dorsum of propodeum slightly concave near the propodeal angle. Anterior portion of head with widely spaced large punctures, a single puncture in the center of the vertex. Filiform brownish hair present on head and mesosoma dorsum, more abundant on malar area.</p> <p>Distribution and biology.</p> <p>Camponotus chrislaini is restricted to the northern portion of Madagascar (Fig. 70B). This newly identified species prefers dry (tropical dry forest, dry forest, disturbed dry forest) as well as wet (montane forest and rainforest) habitats at moderate altitudes (below 1000 m). It was primarily collected from rotten logs, sifted litter, inside trees, or on branches above ground.</p> <p>Discussion.</p> <p>The new species is easily distinguishable from the Camponotus Mayria species, but morphologically is most similar to the unidentified species Camponotus MG098 from subgenus Camponotus Myrmonesites. The main character used to distinguish them is the form of the petiolar scale, as in the former the petiole is much thinner and its dorsal face forms a straight surface, while in the latter the petiole is thickened with the posterior face concave in dorsal view. Camponotus MG098 is also covered with plentiful setae which are scarce and widely distributed in C. chrislaini. The presence of C. chrislaini and C. MG098 in six localities (Sakaramy, Binara, Antsahabe, Analamerana-Ankavanana, Galoko, and Mont Kalabenono) suggests that they are reproductively isolated.</p> <p>Additional material examined.</p> <p>Province Antsiranana: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.16167&amp;materialsCitation.latitude=-13.79861" title="Search Plazi for locations around (long 48.16167/lat -13.79861)">Foret de Binara</a> , 7.5 km 230° SW Daraina, -13.255, 49.61667, 375 m, tropical dry forest (B.L. Fisher) (CAS); Forêt d’ Antsahabe, 11.4 km 275° W Daraina, -13.21167, 49.55667, 550 m, tropical dry forest (B.L. Fisher) (CAS); Forêt d’ Andavakoera, 21.4 km 75° ENE Ambilobe; 4.6 km 356° N Betsiaka, -13.11833, 49.23, 425 m, rainforest (B.L. Fisher) (CAS); Res. Analamerana, 16.7 km 123° Anivorano-Nord, -12.80467, 49.37383, 225 m, tropical dry forest (B.L. Fisher) (CAS); Parc National Montagne d’Ambre, Fozalanana, -12.46637, 49.22157, 475m, dry forest (BLF) (CAS); Res. Ankarana, 7 km SE Matsaborimanga, -12.9, 49.11667, 150 m, tropical dry forest (PSW) (PSWC); Galoko chain, Mont Galoko, -13.58487, 48.71818, 520 m, rainforest (BLF) (CAS); Galoko chain, Mont Galoko, -13.5888, 48.72864, 980 m, montane forest (BLF) (CAS); Galoko chain, Mont Galoko, -13.58745, 48.71419, 38 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, -13.63999, 48.67374, 498 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, -13.64179, 48.67282, 643 m, rainforest (BLF) (CAS); Galoko chain, Mont Kalabenono, -13.64609, 48.67732, 937 m, rainforest (BLF) (CAS); Ampasindava, Andranomatavy Forest, -13.6648, 47.98702, 275 m, disturbed dry forest (BLF) (CAS); Reserve Spéciale d’Ambre, 3.5 km 235° SW Sakaramy, -12.46889, 49.24217, 325 m, tropical dry forest (FGAT) (CAS); Reserve Spéciale de l’Ankarana, 22.9 km 224° SW Anivorano Nord, -12.90889, 49.10983, 80 m, tropical dry forest (FGAT) (CAS); Ampasindava, Forêt d’Ambilanivy, 3.9 km 181° S Ambaliha, -13.79861, 48.16167, 600 m, rainforest (FGAT) (CAS).</p> </div>	http://treatment.plazi.org/id/B1BA149DAF575670859EC43BF1DA7E54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
200598EEA3F45D8D821BACDB7A0AB8D2.text	200598EEA3F45D8D821BACDB7A0AB8D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus claveri Rasoamanana & Fisher 2022	<div><p>Camponotus claveri sp. nov.</p> <p>Figures 44B, 67</p> <p>Holotype worker.</p> <p>Madagascar, Province Fianarantsoa, Forêt d’ Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest, Malaise trap, 22-26 January 2003 (BLF), collection code: BLF07152, specimen code: CASENT0490618 (CAS). Paratypes. Three workers: one worker with same data as holotype but specimen collected from Malaise trap and with collection code: BLF07155, specimen code: CASENT0049849; two workers from Madagascar, Province Fianarantsoa, Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, -20.73528, 47.08382, 1486 m, Uapaca woodland, 03-07 February 2010 (ARA), collection code: ARA0492, specimen code: CASENT0168788 and CASENT0168787 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument shiny black; anterior clypeal margin with short, subtriangular lobe; erect hairs arranged in a transverse row on gastral segment, sparsely arranged on mesosoma dorsum; dorsum of mesosoma with promesonotal and mesometanotal sutures marked but not impressed; petiole squamiform.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 0.89 ± 0.08, 0.84-0.93). In full-face view, head as long as wide, posterior margin rounded, lateral margin straight and subparallel (CWb/CL: 0.33 ± 0.02, 0.32-0.34). Eye circular, protruding, its border smoothly aligned to lateral margins of head (PoOC/CL: 0.10 ± 0.02, 0.08-0.10). Mandibles triangular with six teeth. Clypeus carinate, its anterior margin produced medially into a subtriangular lobe (ClyL/GPD: 0.23 ± 0.07, 0.20-0.26). Antennal scape long, surpassing the occipital corner by the length of two-and-a-half basal funiculi (SL/CS: 0.39 ± 0.03, 0.37-0.40). In lateral view, mesosoma without anterolateral margination, promesonotum not very convex, mesopropodeal suture impressed, propodeum angular with the base fully twice as long as the declivity, which is straight (MW/ML: 0.18 ± 0.01, 0.18-0.19; MPH/ML: 0.14 ± 0.01, 0.14-0.15). Petiole cuneate in profile, its anterior face convex, and its posterior face flattened along entire broad border. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins on petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remaining segment dark brown to black.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head as long as broad, with rather convex posterior borders and subparallel lateral sides (CS: 1.18 ± 0.09, 1.12-1.24, CWb/CL: 0.86 ± 0.01, 0.85-0.87). Eyes circular, placed dorsally next to lateral borders (PoOC/CL: 0.26 ± 0.00, 0.25-0.26). Anterior clypeal margin forms a rounded rectangular lobe (ClyL/GPD: 0.88 ± 0.05, 0.85-0.91). Antennal scape short, just reaching the occipital border (SL/CS: 0.75 ± 0.04, 0.73-0.78). Mesosoma with the same form as minor worker. Sculpture on head coarsely reticulate-punctate anteriorly and tends to be striolate posteriorly.</p> <p>Distribution and biology.</p> <p>Camponotus claveri occupies two different habitats: montane rainforest and open habitats such as Uapaca woodland, eucalyptus plantation, shrubland, and deciduous dry forest, plus urban areas (Fig. 72A). Their nests are found in soil, under stones, and on the ground. It has been collected at elevations of 830-1987 m.</p> <p>Discussion.</p> <p>Camponotus claveri is similar to other species of the Camponotus repens group, but differs in having a thinner petiolar node with sharp dorsal edge, and a rounded triangular anterior clypeal margin. Morphometric analysis combined C. claveri and C. maintilany in the same cluster, meaning that they are the same size but have different qualitative morphological traits. Their distributions are quite distant but sympatric at Alasora, situated in the central highlands.</p> <p>Etymology.</p> <p>This species is named after the collector Claver Marotafika Randrianandrasana.</p> <p>Additional material examined.</p> <p>Province Antananarivo: Alasora, -18.96245, 47.58925, 1434 m, eucalyptus plantation (BLF) (CAS); Antaponimanadala III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 6.55 km E Manalalondo, -19.25583, 47.17751, 1987 m, savanna grassland (ARA) (CAS); Ilafy, -18.85415, 47.56575, 1385 m, urban/garden (BLF) (CAS). Province Fianarantsoa: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Amoron'i Mania Region</a> , <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">District of Ambositra</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Italaviana Uapaca forest</a>, 135 km SE of Antsirabe, -20.17333, 47.086, 1359 m, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Uapaca forest</a> (MG) (CAS); Ampandravelo II <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 10.78 km NE Ranohira, -22.53917, 45.51548, 87 3m, shrubland (ARA) (CAS); Antapia III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">Non Protected Area</a>, 26.43 km SW Ambositra, -20.72, 47.08785, 1494 m, Uapaca woodland (ARA) (CAS); Forêt d’ Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest (FGAT) (CAS); Mampiarika III Non Protected Area, 28.93 km SW Ambositra, -20.73583, 47.08399, 1487 m, Uapaca woodland (ARA) (CAS); Mampiarika IV Non Protected Area, 27.98 km SW Ambositra, -20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (CAS).</p> </div>	http://treatment.plazi.org/id/200598EEA3F45D8D821BACDB7A0AB8D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
1EDEEF999D9954D88F8FEDDBEF505189.text	1EDEEF999D9954D88F8FEDDBEF505189.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus darwinii Forel 1886	<div><p>Camponotus darwinii Forel</p> <p>Figures 5A, 16B, 17B, 52</p> <p>Camponotus darwinii Forel, 1886: 179. Lectotype minor worker, present designation, Centre de Madagascar [Ambatomanjaka, Miarinarivo, -18.766947, 46.869107, 1343 m] (Hildebrandt), AntWeb CASENT0101391 (MNHG). Paralectotypes: one alate queen and five major workers, same data as lectotype, but specimen coded respectively as CASENT0101385 (queen) andCASENT0101386 (MNHG), CASENT0104426, CASENT0104427, FOCOL2468 and FOCOL2469 (ZMHB) (examined).</p> <p>Camponotus (Myrmobrachys) darwinii Forel, 1914: 270; in Camponotus (Myrmepomis): Emery, 1920: 258; in Camponotus (Myrmopiromis): Wheeler, 1922: 1051; Emery, 1925: 128; Bolton, 1995: 95, 131.</p> <p>Camponotus darwinii var. rubropilosus Forel, 1891: 44. Lectotype major worker, present designation, Madagascar, Antananarivo (Camboué), AntWeb CASENT0101392 (MHNG). Paralectotypes: three major workers same data as the lectotype CASENT0101393 (MHNG), CASENT0104628 (ZMHB), CASENT0101193 (NHMB).</p> <p>Camponotus (Myrmopiromis) darwinii var. rubropilosus Wheeler, 1922: 1051; Emery, 1925: 128; Bolton, 1995: 121. Syn. nov.</p> <p>Camponotus darwinii r. rubropilosus var. robustior Forel, 1905: 165. Lectotype minor worker, present designation, Toliara, Menabe, Beronono; Mahabo, -21.0, 45.0, 339 m (Woltzkow), AntWeb CASENT0104629 (MNHG). Paralectotypes: two major workers, same data as lectotype, but specimen coded respectively as CASENT0101195andCASENT0101380 (MHNG).</p> <p>Camponotus (Myrmopiromis) darwinii r. rubropilosus var. robustior Wheeler, 1922: 1051; Emery, 1925: 128; Bolton, 1995: 121. Syn. nov.</p> <p>Worker diagnosis.</p> <p>Body integument matte black, finely reticulate-punctate; mesosoma and/or gastral dorsum with dense, long, and yellowish pilosity and pubescence; anterior clypeal margin with short, rounded, rectangular lobe; petiole squamiform.</p> <p>Description of minor worker.</p> <p>Medium size. Absolute cephalic size (CS: 1.45 ± 0.28; 1.22-1.63). In full-face view, head more quadrate (CWb/CL: 0.37 ± 0.03, 0.35-0.41); posterior margin of head almost flat, lateral margin of head slightly convex and tapering to front. Eyes circular, placed midway from the occipital corner (PoOC/CL: 0.08 ± 0.02; 0.06-0.09). Mandibles triangular with six teeth. With head in full-face view, anterior clypeal margin with a short, rounded, rectangular lobe; less convex in lateral view, not medially marginate (ClyL/GPD: 0.25 ± 0.04, 0.23-0.28). Antennal scape circular and long, surpassing the occiput by the length of two basal funiculi (SL/CS: 0.40 ± 0.04; 0.37-0.43). In lateral view, anterodorsal corner of pronotum rounded, mesosomal dorsal outline feebly and rather evenly arcuate above, dorsal face of propodeum generally longer than declivitous face (MW/ML: 0.21 ± 0.02; 0.19-0.23; MPD/ML: 0.24 ± 0.03; 0.20-0.26). In lateral view, petiolar node cuneate, anterodosum straight basally and inclined dorsally to reach the flattened posterodorsal margin. Tibia tubular, not prismatic, its extensor profile with suberect setae. Head dorsum is matte, finely reticulate-punctate with sparse punctures on malar area; mesosoma striate in lateral view; in dorsal view, petiole, mesosoma, and appendages less shiny, finely reticulate-striolate except the abdominal sternite, which is more matte. Mandible finely reticulate-punctate with sparse oblique punctures. Hairs yellowish white, orange, or tawny colored, erect hairs usually abundant, at least on gastral tergite, often on vertex, occipital portion, and thorax dorsum; petiolar node with 4 pairs of hairs on its dorsum. Shiny black, with mandible and basitarsus reddish to brownish.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head much wider posteriorly (CS: 2.14 ± 0.26, 1.70-2.47; CWb/CL: 1.04 ± 0.06, 0.94-1.20); lateral margins convex and converging towards base of mandible. Eyes semicircular, placed dorsolaterally midway from the occipital margin (PoOC/CL: 0.22 ± 0.01, 0.19-0.24). Clypeus much more quadrate in full-face view (ClyL/GPD: 0.72 ± 0.09, 0.48-0.81), its anterior margin slightly concave medially. Antennal scape short, just surpassing the occipital border (SL/CS: 0.83 ± 0.08, 0.75-0.99). Dorsal outline of mesosoma forms a continuous line so that the basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely reticulate above, punctate to finely areolate below.</p> <p>Distribution and biology.</p> <p>Camponotus darwinii is a famous species which is distributed widely across central of Madagascar, with a range extending from Ankazobe to Mount Papango, PN Befotaka Midongy (Fig. 58A). It has been collected from a wide variety of different habitats, including rainforest, montane rainforest, Uapaca forest, and Uapaca woodland as well as disturbed habitats such as urban and garden areas and cultivated land (Tavy). The species has also been found nesting inside Melia azedarach and Phellolophium madagascariense. Habitat elevations ranged from 1069-2150 m.</p> <p>Discussion.</p> <p>Camponotus darwinii should be separable from most similar species in this group by the combination of the following characters: presence of dense hairs on mesosoma and/or gastral dorsum, mandible and malar area distinctly reddish in color, propodeal and declivitous faces unequal in length. This species is much closer to C. ursus but differs in the density of mesosomal hairs and the aspect of the petiole. In addition, C. darwinii is much more robust than C. ursus.</p> <p>Additional material examined.</p> <p>Province Antananarivo: Analamanga Region, District of Ankazobe, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.53333&amp;materialsCitation.latitude=-18.91667" title="Search Plazi for locations around (long 47.53333/lat -18.91667)">Ambohitantely</a>, 46 km NE of Ankazobe, -18.198, 47.2815, 701 m, Forêt sclerophylle (MG) (CAS); Ambatolaona, -18.928, 47.88283, 1382 m, urban/garden (BLF) (CAS); Kaloy, -18.58998, 47.65102, 1423 m, disturbed montane rainforest, (BLF) (CAS); Ilafy, -18.85415, 47.56575, 1385 m, urban/garden (BLF) (CAS); Forêt de galerie, Telomirahavavy, 23.4 km NNE Ankazobe, -18.12167, 47.2062 7, 1520 m, disturbed gallery montane forest (BLF) (CAS); Stn. Forestiѐre Manjakatompo, -19.35, 47.31667, 2000 m, montane rainforest (Mark Pigeon) (CAS); 25 km NNE Ankazobe, -18.1, 47.18333, 1500 m, rainforest (PSW) (PSWC); Stn. Forestiѐre Manjakatompo, -19.35, 47.31667, 1600 m, montane rainforest (PSW) (PSWC); -18.91667, 47.53333, 1350 m, park/garden (PSW) (PSWC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.53333&amp;materialsCitation.latitude=-18.91667" title="Search Plazi for locations around (long 47.53333/lat -18.91667)">Parc de Tsimbazaza</a>; Antananarivo renivohitra, -18.93, 47.52611, 1275 m (D.M. Olson) (PSWC); -18.91667, 47.53333, 1300 m, park/garden (PSW) (PSWC); 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, -18.47333, 47.96, 1300 m, montane rainforest (FGAT) (CAS); Parc Botanique et <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.91167&amp;materialsCitation.latitude=-18.93233" title="Search Plazi for locations around (long 47.91167/lat -18.93233)">Zoologique de Tsimbazaza</a>, -18.93233, 47.91167, 1300 m (D.H. Kavanaugh) (CAS); Angavokely, -18.93333, 47.75 (B. Pettersson) (PSWC).</p> <p>Province Fianarantsoa: 29 km SSW Ambositra, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.165&amp;materialsCitation.latitude=-20.77667" title="Search Plazi for locations around (long 47.165/lat -20.77667)">Ankazomivady</a>, -20.77667, 47.165, 1700 m, disturbed montane rainforest (B.L. Fisher) (CAS); Forêt d’ Atsirakambiaty, 7.6 km 285° WNW <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.56333&amp;materialsCitation.latitude=-20.59333" title="Search Plazi for locations around (long 46.56333/lat -20.59333)">Itremo</a>, -20.59333, 46.56333, 1550 m, montane rainforest (FGAT)(CAS); 2 km W Andrambovato, along river Tatamaly, -21.51167, 47.41, 1075 m, cultivated land (tavy), recent 1-2 year-old tavy (BLF) (CAS); stream area, 900 m E of Isalo National Park Interpretive Center, -22.62667, 45.35817, 750 m, open area near stream (R. Harin’Hala) (CAS); Parc National d’Andringitra, Plateau d’Andohariana, 39.8 km 204° <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.36667&amp;materialsCitation.latitude=-21.23333" title="Search Plazi for locations around (long 47.36667/lat -21.23333)">Ambalavao</a>, -22.18767, 46.90083, 2150 m, rubicole thicket at base of cliff (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.36667&amp;materialsCitation.latitude=-21.23333" title="Search Plazi for locations around (long 47.36667/lat -21.23333)">Vohiparara</a>, -21.23333, 47.36667 (A. Pauly) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.36667&amp;materialsCitation.latitude=-21.23333" title="Search Plazi for locations around (long 47.36667/lat -21.23333)">Miandritsara Forest</a>, 40 km S of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.17567&amp;materialsCitation.latitude=-20.79267" title="Search Plazi for locations around (long 47.17567/lat -20.79267)">Ambositra</a>, -20.79267, 47.17567, 822 m, low altitude rainforest (MG) (CAS); dry wash, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.35817&amp;materialsCitation.latitude=-22.62667" title="Search Plazi for locations around (long 45.35817/lat -22.62667)">1 km E of Isalo National Park Interpretive Center</a>, -22.62667, 45.35817, 885 m, dry wash (R. Harin’Hala) (CAS); Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.9575&amp;materialsCitation.latitude=-23.84083" title="Search Plazi for locations around (long 46.9575/lat -23.84083)">Mount Papango</a>, -23.84083, 46.9575, 1250 m, montane rainforest (BLF) (CAS); Parc naturel communautaire, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.1791&amp;materialsCitation.latitude=-20.79751" title="Search Plazi for locations around (long 47.1791/lat -20.79751)">29.3 km SW Ambositra</a>, -20.79751, 47.1791, 1782 m, disturbed montane rainforest (BLF) (CAS); Parc naturel communautaire, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.18362&amp;materialsCitation.latitude=-20.775" title="Search Plazi for locations around (long 47.18362/lat -20.775)">26.8 km SW Ambositra</a>, -20.775, 47.18362, 1755 m, disturbed montane rainforest (BLF) (CAS); Parc naturel communautaire, 28.5 km SW Ambositra, -20.78414, 47.16699, 1780 m, disturbed montane rainforest along road side (BLF) (CAS); Mampiarika IV <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.16699&amp;materialsCitation.latitude=-20.78414" title="Search Plazi for locations around (long 47.16699/lat -20.78414)">Non Protected Area</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08382&amp;materialsCitation.latitude=-20.73528" title="Search Plazi for locations around (long 47.08382/lat -20.73528)">27.98 km SW Ambositra</a>, -20.73528, 47.08382, 1486 m, Uapaca woodland (ARA) (CAS); R.S. Ivohibe, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.955&amp;materialsCitation.latitude=-22.49667" title="Search Plazi for locations around (long 46.955/lat -22.49667)">6.5 km ESE Ivohibe</a>, -22.49667, 46.955, 1575 m, montane rainforest (B.L. Fisher (Sylvain)) (CAS); P. N. Andringitra, Forêt Ravaro 12.5 km SW Antanifotsy, -22.21167, 46.845, 1650 m, montane rainforest (S. Razafimandimby) (CAS); R.S. Ivohibe 8.0 km E Ivohibe, -22.48333, 46.96833, 1200 m, montane rainforest (B.L. Fisher (Sylvain)) (CAS); 28 km. SSW <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.16833&amp;materialsCitation.latitude=-20.775" title="Search Plazi for locations around (long 47.16833/lat -20.775)">Ambositra</a>, Ankazomivady, -20.775, 47.16833, 1670 m, montane rainforest edge (B.L. Fisher) (CAS); 38 km S Ambalavao, Res. Andringitra, -22.2, 46.96667, 1680 m, montane rainforest (B.L. Fisher) (CAS); 40 km S Ambalavao, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.96667&amp;materialsCitation.latitude=-22.21667" title="Search Plazi for locations around (long 46.96667/lat -22.21667)">Res. Andringitra</a>, -22.21667, 46.96667, 1275 m, montane rainforest (B.L. Fisher) (CAS); Res. Andringitra; 8.5 km SE Antanitotsy, -22.16667, 46.96667, 1990 m, montane rainforest (Sylvian) (CAS); 8.0 km NE Ivohibe, -22.42167, 46.89833, 1200 m, montane rainforest (B.L. Fisher (Sylvain)) (CAS); Antapia I <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.89833&amp;materialsCitation.latitude=-22.42167" title="Search Plazi for locations around (long 46.89833/lat -22.42167)">Non Protected Area</a>, 26.43 km SW Ambositra, -20.71972, 47.08685, 1495 m, Uapaca woodland (ARA) (CAS); Antohatsahomby I <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.08685&amp;materialsCitation.latitude=-20.71972" title="Search Plazi for locations around (long 47.08685/lat -20.71972)">Non Protected Area</a>, 22.77 km NW Ambatofinandrahana, -20.55056, 46.58562, 1550 m, Uapaca woodland (ARA) (CAS); Amoron’i Mania Region, District of Ambositra, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.58562&amp;materialsCitation.latitude=-20.55056" title="Search Plazi for locations around (long 46.58562/lat -20.55056)">Italaviana Uapaca forest</a>, 135 Km SE of Antsirabe, -20.17333, 47.086, 1359 m, Uapaca forest, (MG) (CAS); Antapia IV <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.086&amp;materialsCitation.latitude=-20.17333" title="Search Plazi for locations around (long 47.086/lat -20.17333)">Non Protected Area</a>, 26.42 km SW Ambositra, -20.71917, 47.0868, 1494 m, Uapaca woodland (ARA) (CAS); Antohatsahomby V <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.0868&amp;materialsCitation.latitude=-20.71917" title="Search Plazi for locations around (long 47.0868/lat -20.71917)">Non Protected Area</a>, 22.63 km NW Itremo, -20.56722, 46.57923, 1726 m, Uapaca woodland (ARA) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.57923&amp;materialsCitation.latitude=-20.56722" title="Search Plazi for locations around (long 46.57923/lat -20.56722)">Manombo Special Reserve</a>, 32 km SE of Farafangana, -23.02183, 47.72, 36 m, lowland rainforest (MG) (CAS); Fitovavy Fitovinany Region, District of Ifanadiana, 12 km W of Ranomafana, -21.25083, 47.40717, 1127 m, forest edge, open area (MG) (CAS); Antohatsahomby IV <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.40717&amp;materialsCitation.latitude=-21.25083" title="Search Plazi for locations around (long 47.40717/lat -21.25083)">Non Protected Area</a>, 22.67 km NW Itremo, -20.56306, 46.58097, 1708 m, Uapaca woodland (ARA) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.58097&amp;materialsCitation.latitude=-20.56306" title="Search Plazi for locations around (long 46.58097/lat -20.56306)">Ambinanindranomena Non Protected Area</a>, 39.45 km SE Ambalavao, -21.95386, 47.29427, 1069 m, montane rainforest (ARA) (CAS); Hte Mananjary; Ambinanindrano; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.633335&amp;materialsCitation.latitude=-20.733334" title="Search Plazi for locations around (long 47.633335/lat -20.733334)">Ambositra</a>, -20.733334, 47.633335, 657 m (R. Catala) (MNHN).</p> <p>Province Mahajanga: Sofia Region, District of Sofia, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.73417&amp;materialsCitation.latitude=-15.157" title="Search Plazi for locations around (long 47.73417/lat -15.157)">Anjiamangirana</a> 45 km S Antsohihy, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.73417&amp;materialsCitation.latitude=-15.157" title="Search Plazi for locations around (long 47.73417/lat -15.157)">Analagnambe Galery forest</a>, 5 km W Anjiamangirana, -15.157, 47.73417, 97 m, low degraded dry forest (MG) (CAS). Province Toamasina: Ambanizana, Parc National Masoala, -15.57222, 50.00694, 1020 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CAS); Moramanga, -18.948902, 48.224426, 914 m (Decarpentries) (NHMB); 6.9 km NE Ambanizana, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.635&amp;materialsCitation.latitude=-15.17833" title="Search Plazi for locations around (long 49.635/lat -15.17833)">Ambohitsitondroina</a>, -15.58506, 50.00952, 825 m, rainforest (B.L. Fisher) (CAS); Montagne d’Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, -15.17833, 49.635, 1100 m, montane rainforest (FGAT) (CAS). Province Toliara: [S.W. Madagascar]; Beronono; Mahabo, -21, 45, 339 m (<a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Voltzkow</a>) (NHMB); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Androy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">District of Tsihombe</a>, 74 km S of Tsihombe, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Cap Ste Marie Reserve</a>, -25.58767, 45.163, 36 m, spiny bush (MG) (CAS); 13 km NW Enkara, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Res Andohahela</a>, -24.55, 46.8, 1300 m, montane rainforest (B.L. Fisher) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">District of Tulear</a> II, Mikea decidious dry forest 3 km N Andranomavo village, -22.90367, 43.4755, 30 m, deciduous dry forest (MG) (CAS); Miary, -23.311972, 43.740917, 55 m (F. Geay) (MNHN); Ambovombe, -25.1775, 46.09283, 25 m (R. Decary) (MNHN); near road, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.73117&amp;materialsCitation.latitude=-22.8405" title="Search Plazi for locations around (long 44.73117/lat -22.8405)">Zombitse National Park</a>, -22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin’Hala) (CAS).</p> </div>	http://treatment.plazi.org/id/1EDEEF999D9954D88F8FEDDBEF505189	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
56A450F48CB95F468775CE68A76E8CDF.text	56A450F48CB95F468775CE68A76E8CDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus ellioti Forel 1891	<div><p>Camponotus ellioti Forel</p> <p>Figures 2A, 8A, 38B, 63</p> <p>Camponotus ellioti Forel, 1891: 37, 73. Holotype minor worker, Madagascar, collection Henri de Saussure. AntWeb CASENT0101382 (MHNG) (examined).</p> <p>Camponotus (Myrmepomis) ellioti Forel, 1912: 92; Emery, 1920: 258.</p> <p>Camponotus (Myrmopiromis) ellioti Wheeler, 1922: 1052; Emery, 1925: 128; Bolton, 1995: 97, 131.</p> <p>Camponotus ellioti var. relucens Santschi, 1911: 133. Lectotype two workers, present designation, Madagascar, Toliara Province, Region du Sud-Est Fort-Dauphin (Ch. Alluaudi), AntWeb CASENT0101196 (minor); CASENT0101197 (major) (NHMB).</p> <p>Camponotus (Myrmopiromis) ellioti var. relucens Wheeler, 1922: 1052; Emery, 1925: 129; Bolton, 1995: 120. Syn. nov.</p> <p>Diagnosis.</p> <p>Head and mesosoma reddish brown to dark brown, gaster always dark colored; anterior clypeal margin of clypeus with short, rectangular lobe; pronotal humeri tuberculate, dorsal outline of mesosoma arcuate; gastral tergite with short, decumbent, ochreous setae.</p> <p>Description of minor worker.</p> <p>Large-sized species. Absolute cephalic size (CS: 1.85 ± 0.55, 1.45-2.42). In full-face view, head trapezoidal, wider than length of pronotum in dorsal view; occipital margin slightly convex medially, and straight subparallel lateral margin of head (CWb/CL: 0.36 ± 0.03, 0.33-0.39). Eyes elliptical, positioned at the midline of head (PoOC/CL: 0.08 ± 0.01, 0.07-0.09). Mandibles short and triangular with six teeth. Clypeus trapeziform, less convex in profile, sharply carinate along its entire length, with a short, rectangular projection feebly sinuate in the middle (ClyL/GPD: 0.12 ± 0.01, 0.11-0.13). Frontal carina less sinuate and divergent. Antennal scape long and surpassing the occipital corner (SL/CS:0.41 ± 0.07;0.34-0.46). In lateral view, dorsal outline of mesosoma forms a smooth curve, anterodorsal corner of pronotum distinctly shouldered, propodeum rounded, its surface convex and not marginate laterally, declivitous face triangular in posterior view (MW/ML: 0.23 ± 0.03, 0.21-0.25; MPH/ML: 0.18 ± 0.03, 0.16-0.20). Petiolar node squamiform in profile, wider than long, anterodorsal face distinctly flat and inclined forward, posterodorsal face flat, node summit arcuate. First abdominal tergite with vertical anterior face. Entire body (frontal area, dorsal face of mesosoma, scape, and legs) entirely, finely, and densely reticulate-punctate and matte; lateral face of petiole and abdomen transversally striolate. Mandible finely reticulate-punctate and subopaque, with superimposed, abundant punctuation arranged transversally. Ground pilosity yellowish, short, sparse, and dilute on tibia and scape. Short white to yellowish standing hair present on entire dorsum, sparsely distributed on occipital region, regularly present on petiolar dorsum and onto declivity, absent on scape and tibia. Dorsum of gastral segment with appressed ochreous setae and whitish, suberect hairs arranged serially parallel and longitudinally. Head and mesosoma dark brown to reddish brown; mandible, basal half of scape, basitarsus, and gastral segment reddish.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head as long as broad, with rather straight posterior and lateral borders (CS: 4.13 ± 0.16, 3.90-4.33; CWb/CL: 1.04 ± 0.02, 1.01-1.07). Eyes circular, placed dorsally next to lateral borders (PoOC/CL: 0.28 ± 0.01, 0.26-0.29). Anterior clypeal margin forms a rectangular lobe (ClyL/GPD: 0.87 ± 0.05, 0.79-0.92). Antennal scape short, not reaching the occipital border (SL/CS: 0.61 ± 0.02, 0.58-0.65). Promesonotal suture strongly marked.</p> <p>Distribution and biology.</p> <p>Camponotus ellioti occurs in the dry forest regions of the southeast of Madagascar in habitats such as bush, coastal scrub, and coastal spiny bush on sandy soil, Euphorbia forest, spiny bush and thicket, gallery forest, and riparian scrub (Fig. 70C). It has been collected from sifted litter and Malaise traps. The altitudes of these localities range from 5 to 230 meters.</p> <p>Discussion.</p> <p>Camponotus ellioti is a distinctive species recognizable by the submargined anterolateral corner of the pronotum and the gaster covering of thick, blunt hairs the color of rust. In the traditional classification, C. ellioti is considered a member of the C. darwinii species group based on the density and color of the gastral pilosity and the sculpture pattern; however, we propose color as a character difference between these two groups.</p> <p>Additional material examined.</p> <p>Province Toliara: [Région du Sud Andrahomana, Museum Paris, Ch. Alluaud 1901]; Analapatsa I; Taolagnaro, -25.1833333, 46.6666667, 31 m, (NHMB); [Toliara S.W. Madagascar]; Beronono; Mahabo, -21, 45, 339 m (Voeltzkow) (MHNG); 3.5km 236° SW Marovato, -25.55389, 45.25583, 230 m, spiny forest/thicket (FGAT) (CAS); 5 km N Ampotaka, Malaise on trail in Vitambany gallery forest, -24.65033, 43.96317, 86 m, gallery forest (MG) (CAS); 7.0 km 156° SSE Lavanono, -25.47111, 44.9885, 50 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Ambohimahavelona village</a> 33 km NE of Toliara, Andoharano, dry forest, -23.44083, 43.89967, 46 m, dry forest (MG) (CAS); Andranovato, 5 km SE of Manombo, -22.81806, 43.50217, 18 m, Euphorbia forest (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Androy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">District of Tsihombe</a>, 74 km S of Tsihombe, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Cap Sainte Marie Reserve</a>, -25.58767, 45.163, 36 m, spiny bush (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Anosy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">District of Amboasary</a>, 58 km SW of Fort Dauphin, 8 km NW of Amboasary, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Berenty Special Reserve</a>, -25.00667, 46.30333, 85 m, gallery forest (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Atsimo-Andrefana Region</a>, Antsokay Arboretum, -23.41491, 43.75499, 13 m, spiny forest/thicket (B.L. Fisher, F.A. Esteves et al.) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Atsimo-Andrefana Region</a>, Famata 02, -23.45314, 43.76448, 20 m, coastal spiny bush on sandy soil (B.L. Fisher, F.A. Esteves et al.) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Atsimo-Andrefana Region</a>, Famata04, -23.53922, 43.77935, 20 m, riparian scrub (B.L. Fisher, F.A. Esteves et al.) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Foret de Petriky</a> , 12.5 km W 272° Tolagnaro, -25.06167, 46.87, 10 m, littoral rainforest (B.L. Fisher) (CAS); Libanona beach, Tolagnaro, -25.03883, 46.996, 20 m, coastal scrub (BLF) (CAS); Mahafaly Plateau, 6.2 km 74° ENE Itampolo, -24.65361, 43.99667, 80 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Mikea Forest</a>, spiny forest, -22.91333, 43.48222, 37 m, spiny forest (R. Harin’Hala) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Parc National de Tsimanampetsotsa</a>, 6.7 km 130° SE Efoetse, 23.0 km 175° S Beheloka, -24.10056, 43.76, 25 m, spiny forest/thicket (FishFGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Parc National de Tsimanampetsotsa</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Foret de Bemanateza</a> , 20.7 km 81° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Efoetse</a>, 23.0 km 131° SE Beheloka, -23.99222, 43.88067, 90 m, spiny forest/thicket (FiFGAT) (CAS); Ranobe, -23.04457, 43.61532, 20 m, spiny forest/thicket (Frontier Wilderness Project) (CAS); Réserve Privé Berenty, Forêt d’Anjapolo, 21.4 km 325° NW Amboasary, -24.92972, 46.20967, 65 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">ReserveSpeciale de Cap Sainte Marie</a> , 12.3 km 262° W Marovato, -25.58167, 45.16833, 200 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">ReserveSpeciale de Cap Sainte Marie</a> , 14.9 km 261° W Marovato, -25.59444, 45.14683, 160 m, spiny forest/thicket (FGAT) (CAS); Tolagnaro (= <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.14683&amp;materialsCitation.latitude=-25.59444" title="Search Plazi for locations around (long 45.14683/lat -25.59444)">Fort Dauphin</a>), -25.03333, 47, 5 m, littoral vegetation (B.L. Fisher) (CAS); Tolagnaro (= Ft. Dauphin), -25.04611, 47, 30 m, garden, hotel (D.M. Olson) (PSWC); Toliara, -23.3575, 43.669, 20 m, urban/garden (BLF) (CAS); Tsimanampetsotsa National Park, Mitoho Forest, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.75233&amp;materialsCitation.latitude=-24.0485" title="Search Plazi for locations around (long 43.75233/lat -24.0485)">Malaise</a> across trail at escarpment base, -24.0485, 43.75233, 120 m, dense dry forest and transitional forest (MG) (CAS); Tsimelahy - Parcel II, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.75233&amp;materialsCitation.latitude=-24.0485" title="Search Plazi for locations around (long 43.75233/lat -24.0485)">Andohahela National Park</a>, transitional forest, Toliara Province, -24.93683, 46.62667, 180 m, transition forest (M.E. Irwin, F.D. Parker, R. Harin’Hala) (CAS).</p> </div>	http://treatment.plazi.org/id/56A450F48CB95F468775CE68A76E8CDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
620DC68E0ECC53C5BE04D2935A560375.text	620DC68E0ECC53C5BE04D2935A560375.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus ihazofotsy Rasoamanana & Fisher 2022	<div><p>Camponotus ihazofotsy sp. nov.</p> <p>Figures 46B, 68</p> <p>Holotype worker.</p> <p>Madagascar, Province Toliara, Ihazofotsy - Parcel III, Andohahela National Park, -24.83483, 46.48683, 80 m, tropical dry forest, transition between spiny and dry deciduous forests, Malaise traps, 26 January- 03 March 2003 (M.E. Irwin, F.D. Parker, R. Harin’Hala), collection code: MA-02-21-08, specimen code: CASENT0062675 (CAS). Paratypes. Two workers with same data as holotype but collection code: MA-02-21-01, specimen codes: CASENT0062267 and CASENT0062269 (CAS)</p> <p>Worker diagnosis.</p> <p>Integument shiny black; standing pilosity sparse elsewhere; clypeus with short, rounded lobe; mesonotal suture impressed; petiole nodiform; gaster longer than mesosoma.</p> <p>Note.</p> <p>Habitat: eucalyptus plantation, grassland, savanna grassland, Uapaca forest, Uapaca woodland, urban/garden. Altitude: 1300-1987 m. Province: Antananarivo. Microhabitat: sifted litter (leaf mold, rotten wood), under stones, on low vegetation, ex soil, ground forager(s), ground nest. Method: MW 20 sample transect, 10 m; SD 20 digging sample transect, 10 m; 20 sardine and ground tuna fish bait; Malaise, 2 traps.</p> <p>Description of minor worker.</p> <p>Small-sized species. Absolute cephalic size (CS: 0.78 ± 0.12; 0.75-0.81). In full-face view, head ovate, usually somewhat longer than broad, lateral margin of head straight and tapering to front (CWb/CL: 0.34 ± 0.01, 0.34-0.35). Eye subelliptical, protruding, its border smoothly aligned to lateral margins of head (PoOC/CL: 0.09 ± 0.03, 0.08-0.10). Mandibles triangular with six teeth. Anterior clypeal margin not produced, entire, broadly rounded. Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL/CS: 0.35 ± 0.03, 0.34-0.36). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeum dorsum is raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW/ML: 0.18 ± 0.01, 0.17-0.18; MPH/ML: 0.10 ± 0.01, 0.10-0.10). In lateral view, petiole cuneate, its anterior face convex, its posterior face more flattened. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins of petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown, and the remaining segment dark brown to black.</p> <p>Distribution and biology.</p> <p>Camponotus ihazofotsy is known from the dry forest of Tsimanapetsotsa and Andohaela NP (Fig. 72B). We assume that this species is arboreal because all samples were collected from Malaise traps.</p> <p>Discussion.</p> <p>Camponotus ihazofotsy closely resembles C. tsimelahy, although morphological examination reveals a number of distinguishing characters. The most interesting character is the form and length of the petiolar scale; in C. ihazofotsy the petiole is cuboidal (anterior and posterior face about the same height), while in C. tsimelahy the posterior face is higher than the anterior face. In addition, these two species are sympatric at one site, Ihazofotsy. Molecular phylogenetic analysis provides strong evidence that they represent separate lineages (unpublished data).</p> <p>Etymology.</p> <p>The species epithet refers to the type locality.</p> <p>Additional material examined.</p> <p>Province Toliara: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.75233&amp;materialsCitation.latitude=-24.0485" title="Search Plazi for locations around (long 43.75233/lat -24.0485)">Tsimanampetsotsa National Park</a>, Mitoho Forest, Malaise across trail at escarpment base, -24.0485, 43.75233, 120 m, dense dry forest and transition forest (MG) (CAS).</p> </div>	http://treatment.plazi.org/id/620DC68E0ECC53C5BE04D2935A560375	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8AE00EA93EBC587ABDC2AB6DE530B564.text	8AE00EA93EBC587ABDC2AB6DE530B564.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus ivadia Rasoamanana & Fisher 2022	<div><p>Camponotus ivadia sp. nov.</p> <p>Figures 9D, 50C, D, 65</p> <p>Holotype worker.</p> <p>Madagascar, Province Antsiranana, Forêt d’ Ampondrabe, 26.3 km 10° NNE Daraina, tropical dry forest, on low vegetation, -12.97, 49.7, 175 m, 04 November 2009 (BLF) collection code: BLF10105, specimen code: CASENT0498634 (CAS). Paratype. One worker with same data as holotype but specimen coded as CASENT0498635 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument dark brown, apparently shiny; hairs on mesosoma and petiole blunt-tipped; fine, pointed hairs on head and gastral dorsum; anterior clypeal margin with straight rectangular lobe; dorsal outline of mesosoma nearly evenly arcuate; petiole higher than long.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 1.21 ± 0.19, 1.13-1.30). In full-face view, head almost quadrate, with feebly convex posterior and lateral border (CWb/CL: 0.35 ± 0.03, 0.33-0.36). Eyes elliptical, placed next to the lateral margins of head (PoOC/CL: 0.10 ± 0.01, 0.10-0.11). Mandibles triangular with six teeth. Anterior clypeal margin angularly produced, with straight lateral border and rounded anterolateral corner. Antennal scape long, surpassing the occipital corner by the length of two basal funiculi (SL/CS: 0.39 ± 0.02, 0.38-0.40). In lateral view, pronotum with short anterolateral margination, mesosoma moderately convex, propodeum angular in profile, with unequal basal and declivitous faces, each straight in profile and meeting at a rounded obtuse angle (MW/ML: 0.21 ± 0.03, 0.19-0.22; MPH/ML: 0.16 ± 0.01, 0.16-0.17). In lateral view, petiole rather narrow, its anterior face convex and its posterior face flattened. Body finely imbricate. Hairs whitish; enlarged, whitish hairs with blunt tip on mesonotum, propodeum, and petiolar dorsum; filiform, pointed, grayish hair on head, pronotum, and gastral segment; pubescence white, short, and more abundant across entire dorsum. Head, mesosoma and gaster dark brown, femora, tibia, and funiculi brown, scape and basitarsi slightly lighter.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head as long as wide, posterior borders medially concave and subparallel, lateral sides diverging apically (CS: 1.93; CWb/CL: 0.93). Eyes circular, placed dorsally in the middle of the lateral borders (PoOC/CL: 0.28). Anterior clypeal margin forms a short, rounded, rectangular lobe, slightly concave medially (ClyL/GPD: 0.84). Antennal scape short, just reaching the occipital border (SL/CS: 0.69). Sculpture, pilosity, and color as in minor worker.</p> <p>Distribution and biology.</p> <p>Camponotus ivadia has thus far been collected only in the Forêt d’ Ampondrabe within Ankarana Reserve, a region with tropical dry forest located in the northern portion of Madagascar (Fig. 70F). All specimens were collected by beating vegetation.</p> <p>Discussion.</p> <p>This species is easily recognized within the Camponotus madagascarensis group as one of few back and shiny species with white standing pubescence.</p> <p>Etymology.</p> <p>The species epithet Camponotus ivadia is derived from the low vegetation microhabitat it prefers.</p> <p>Additional material examined.</p> <p>Province Antsiranana: Rés. Ankarana, 7 km SE Matsaborimanga, -12.9, 49.11667, 150 m, tropical dry forest (PSW) (PSWC); Forêt d’ Ampondrabe, 26.3 km 10° NNE Daraina, -12.97, 49.7, 175 m, tropical dry forest (B.L. Fisher) (CAS).</p> </div>	http://treatment.plazi.org/id/8AE00EA93EBC587ABDC2AB6DE530B564	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
8F479C2D6B0958D4845A5D9316B32BA8.text	8F479C2D6B0958D4845A5D9316B32BA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus jjacquia Rasoamanana & Fisher 2022	<div><p>Camponotus jjacquia sp. nov.</p> <p>Figures 10C, D, 45A, 66</p> <p>Holotype worker.</p> <p>Madagascar, Province Toliara, Parc National de Tsimanampetsotsa, Forêt de Bemanateza, 20.7 km 81°E Efoetse, 23.0 km 131°SE Beheloka, -23.99222, 43.88067, 90 m, spiny forest/thicket, under stone, 22-26 March 2002 (BLF), collection code: BLF06310, specimen code: CASENT0445276 (CAS). Paratypes. Four workers with the same data as holotype CASENT0445284, CASENT0445285, CASENT0445287, CASENT0445288 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument black; entire body with plentiful, erect setae; anterior clypeal margin produced into a rounded triangular lobe; dorsal outline of mesosoma slightly impressed at the level of metanotum; petiole long and low.</p> <p>Description of minor worker.</p> <p>Medium-sized. Absolute cephalic size (CS: 0.99 ± 0.14; 0.91-1.09). In full-face view, head almost quadrate (CWb/CL: 0.32 ± 0.02; 0.31-0.34); posterior margin evenly straight, lateral margin of head anterior to eyes slightly convex. Eyes circular, placed above midline of head (PoOC/CL: 0.10 ± 0.02; 0.09-0.12). Mandibles triangular with six teeth. Clypeus carinate, its anterior border angularly produced in the middle (ClyL/GPD: 0.20 ± 0.06, 0.16-0.23). Antennal scape long, surpassing the occiput by the length of one basal funiculus (SL/CS: 0.34 ± 0.02; 0.32-0.34). In lateral view, anterodorsal corner of pronotum rounded, mesonotum dorsum inclined toward nearly straight propodeum face (MW/ML: 0.19 ± 0.02, 0.18-0.20; MPH/ML: 0.13 ± 0.02, 0.12-0.14). In lateral view, petiolar node longer than wide, with flat summit, its anterior face leaning forward. Ants finely foveolate throughout, apparently shiny. Entire body, including mandibles and appendages, covered with plentiful, erect, whitish setae, uneven in length, some long. Head, mesosoma, and gaster dark brown to black; antennae, mandibles, tibiae, and tarsi reddish brown.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head much wider posteriorly (CS: 1.68 ± 0.07, 1.61-1.76; CWb/CL: 0.91 ± 0.02, 0.89-0.92); lateral margins convex and converging towards base of mandible. Eyes circular placed dorsolaterally midway from the occipital margin (PoOC/CL: 0.30 ± 0.00, 0.29-0.30). Clypeus with a short rectangular lobe with rounded lateral angle (ClyL/GPD: 0.85 ± 0.03, 0.81-0.88). Antennal scape short, not reaching the occipital margin (SL/CS: 0.57 ± 0.02, 0.56-0.59). Dorsal outline of mesosoma interrupted by sutures, metanotal suture forms a canal. Head dorsum and mesosoma coarsely reticulate-punctate, petiole finely striolate, gaster imbricate and shiny.</p> <p>Distribution and biology.</p> <p>The known elevational range of Camponotus jjacquia extends from 43 to 923 m. Specimens have been collected on tree trunks, on low vegetation, in leaf litter, and under stones in Bismarckia woodland, deciduous forest, gallery forest, savannah woodland, spiny forest, and tropical dry forest (Fig. 72C). This species nests in the soil.</p> <p>Discussion.</p> <p>The unique petiolar shape, the density of its pilosity, and the shape of its metanotal dorsum make C. jjacquia a very distinctive species.</p> <p>Etymology.</p> <p>The new species is dedicated to Jean Jacques Rafanomezantsoa, who has been a Malagasy ant collector for more than twenty years.</p> <p>Additional material examined.</p> <p>Province Fianarantsoa: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Ampotoampoto</a> III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">National Park</a>, 7.91 km NW Ilakaka, -22.62944, 45.189, 919 m, savannah woodland (ARA) (CAS); Isalo IV National Park, 12 km SW Ranohira, -22.61472, 45.31304, 867 m, Bismarckia woodland (ARA) (CAS); Ampotoampoto IV National Park, 7.83 km NW Ilakaka, -22.62944, 45.1912, 923 m, savanna woodland (ARA) (CAS); Province Mahajanga: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Ampijoroa National Park</a>, 160 km N Maevatanana, deciduous forest, -16.31944, 46.81333, 43 m, deciduous forest (MG) (CAS). Province Toliara: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Anosy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">District of Amboasary</a>, 58 km SW of Fort Dauphin, 8 km NW of Amboasary, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Berenty Special Reserve</a>, -25.00667, 46.30333, 85 m, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Galery forest</a> (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Anosy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">District of Fort-Dauphin</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Andohaela National Park</a> Parcelle II, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Tsimelahy</a>, 42 km W of Fort-Dauphin, -24.93683, 46.62667, 176 m, transition forest (Mike Irwin, Frank Parker, Rin’ha) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Anosy Region</a>, Parc National d’Andohahela, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Foret de Manatalinjo</a> , -24.82505, 46.57811, 90 m, spiny forest/thicket (B.L. Fisher, F.A. Esteves et al.) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">District of Betioky</a>; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Beza Mahafaly Special reserve</a>, Parcelle Belle vue 7 km W of Research Station, -23.68983, 44.5755, 177 m, spiny forest (Rin’ha) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">District of Betioky</a>, 30 km E Betioky, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Beza Mahafaly Special Reserve</a> (around <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Research Station</a>), -23.6865, 44.591, 165 m, gallery dry deciduous forest (MG); Beza-Mahafaly, 27 km E Betioky, -23.65,4 4.63333, 135 m, tropical dry forest (B.L. Fisher) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Foret de Mahavelo</a> , <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Isantoria River</a>, -24.75833, 46.15717, 110 m, spiny forest/thicket (FGAT) (CAS); Parc National d’Andohahela, Forêt d’ Ambohibory, 1.7 km 61° ENE Tsimelahy, 36.1 km 308° NW Tolagnaro, -24.93, 46.6455, 300 m, tropical dry forest (FGAT) (CAS); Parc National d’Andohahela, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Foret de Manatalinjo</a> , 33.6 km 63° ENE Amboasary, 7.6 km 99° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Hazofotsy</a>, -24.81694, 46.61, 150 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Parc National de Kirindy Mite</a>, 16.3 km 127° SE Belo sur mer, -20.79528, 44.147, 80 m, tropical dry forest (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Parc National de Tsimanampetsotsa</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Foret de Bemanateza</a> , 20.7 km 81° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Efoetse</a>, 23.0 km 131° SE Beheloka, -23.99222, 43.88067, 90 m, spiny forest/thicket (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Beza Mahafaly Reserve</a>, near research station, Parcel I, -23.6865, 44.591, 165 m, dry deciduous forest (R. Harin’Hala) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Beza Mahafaly Reserve</a>, near Bellevue, Parcel II, -23.68983, 44.5755, 180 m, spiny forest (R. Harin’Hala) (CAS); Andohahela National Park, Tsimelahy - Parcel II, transitional forest, -24.93683, 46.62667, 180 m, transitional forest (M.E. Irwin, F.D. Parker, R. Harin’Hala) (CAS).</p> </div>	http://treatment.plazi.org/id/8F479C2D6B0958D4845A5D9316B32BA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
45336E9577B351B38C648D581ABF175A.text	45336E9577B351B38C648D581ABF175A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus maintilany Rasoamanana & Fisher 2022	<div><p>Camponotus maintilany sp. nov.</p> <p>Figures 37B, 39A, 64</p> <p>Holotype worker.</p> <p>Madagascar, Province Antananarivo, Ankalalahana, -19.00659, 47.1122, 1375 m, Uapaca woodland, ground forager(s), 29 March 2011 (BLF), collection code: BLF26368, specimen code: CASENT0245156 (CAS). Paratypes. Two workers with same data as holotype but collection code: BLF26364, specimen code: CASENT0245172; collection code: BLF26385, specimen code: CASENT0245187 (CAS).</p> <p>Worker diagnosis.</p> <p>Ants bicolored: head and mesosoma reddish brown, gaster dark brown to black; pronotum and mesonotum form a unique dome followed by an impressed metanotal suture, propodeum, and petiole angle with standing, whitish, spatulate hairs, head and gastral dorsum with sparse, fine, brown hairs; petiole squamiform.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 0.84 ± 0.05, 0.81-0.86). In full-face view, head distinctly longer than broad, with convex occipital margin and subparallel lateral sides (CWb/CL: 0.33 ± 0.02, 0.32-0.34). Eyes elliptical and break the lateral outlines of head (PoOC/CL: 0.09 ± 0.01, 0.08-0.10). Mandibles triangular with six short, blunt teeth. Clypeus convex, its anterior border produced into rounded lobe (ClyL/GPD: 0.24 ± 0.01, 0.22-0.25). Antennal scape long, surpassing the occipital margin (SL/CS: 0.40 ± 0.03, 0.39-0.43). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeum dorsum is raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW/ML: 0.20 ± 0.01, 0.19-0.20; MPH/ML: 0.14 ± 0.03, 0.12-0.15). In lateral view, petiole cuneate, its anterior face convex, its posterior face more flattened with entire border broad. Body finely imbricate. Hairs grayish, present only on vertex, one pair next to the pronotal suture, one pair on the middle of mesonotum dorsum, sparse on gastral tergite; enlarged, whitish hairs present on propodeal corner (two or three pairs) and on posteromargin of petiole; pubescence whitish and more dilute on gaster tergites. Head, mesosoma, and appendages reddish to brownish yellow; first gastral segment lighter colored than the remaining dark brown segments.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head apparently rectangular, with straight to feebly convex posterior margin and parallel lateral sides. (CS: 1.13 ± 0.05, 1.09-1.18; CWb/CL: 0.90 ± 0.02, 0.88-0.91). Eyes circular, placed dorsally next to the lateral margin (PoOC/CL: 0.26 ± 0.01, 0.25-0.27). Anterior clypeal margin forms a short, rounded lobe (ClyL/GPD: 0.83 ± 0.00, 0.83-0.83). Antennal scape short, just reaching the occipital border (SL/CS: 0.77 ± 0.03, 0.74-0.79). Dorsal outline of mesosoma not very convex and interrupted but pronotal and mesopropodeal suture present, the latter weakly impressed.</p> <p>Distribution and biology.</p> <p>Camponotus maintilany occurs from 1300-1987 m in elevation and inhabits a montane environment in the central highlands, where it prefers mostly Uapaca forest or woodland, but is also found in savanna grassland and eucalyptus plantations (Fig. 70E). This new species has been collected by digging in soil, sifting leaf litter, and baiting with tuna fish or sardines. In addition, a few workers were found in Malaise traps.</p> <p>Discussion.</p> <p>Camponotus maintilany is most readily distinguished from other members of the Camponotus ellioti species group by a deeper metanotal groove and shinier sculpture. It may be confused with C. andrianjaka but can be separated by its impressed metanotal suture and truncate petiolar node.</p> <p>Etymology.</p> <p>This species is derived from the coloration of the mesosoma and gaster. “Mainty” means black and “ilany” means second part.</p> <p>Additional material examined.</p> <p>Province Antananarivo: Alasora, -18.96245, 47.58925, 1434 m, grassland (BLF) (CAS); Ambatolaona, -18.928, 47.88283, 1382 m, urban/garden (BLF) (CAS); Ambohidrabiby, -18.7746, 47.60908, 1362 m, eucalyptus plantation (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Analamanga Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">District of Arivonimamo</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Andebadeba Uapaca forest</a>, 4 km from <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Arivonimamo</a>, -19.00806, 47.14861, 1300 m, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Uapaca forest</a> (MG) (CAS); Andohony I <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Non Protected Area</a>, 22.62 km SW Antsirabe, -20.06784, 46.99068, 1451 m, Savannah grassland (ARA) (CAS); Andohony II <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Non Protected Area</a>, 22.71 km SW Antsirabe, -20.06904, 46.99199, 1398 m, savanna grassland (ARA) (CAS); Ankalalahana, -19.00711, 47.1337, 1350 m, Uapaca woodland (BLF) (CAS); Ankalalahana, -19.00659, 47.1122, 1375 m, Uapaca woodland (BLF) Ankalalahana, -19.00659, 47.1122, 1375 m, Uapaca woodland (BLF) (CAS); Ankalalahana, -19.00716, 47.1124, 1370 m, Uapaca woodland (BLF) (CAS); Ankalalahana, -19.0531, 47.11304, 1314 m, Uapaca woodland (H.E. Marti et al.) (CAS); Ankalalahana, -18.99666, 47.1118, 1353 m, Uapaca woodland (H.E. Marti et al.) (CAS); Antaponimanadala III <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Non Protected Area</a>, 6.55 km E Manalalondo, -19.25583, 47.17751, 1987 m, savanna grassland (ARA) (CAS); Iharanandriana, -19.15823, 47.49702, 1513 m, Uapaca woodland (BLF) (CAS), Kaloy, -18.59568, 47.65333, 1338 m, grassland (BLF) (CAS); Navoatra I <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.12232&amp;materialsCitation.latitude=-18.98194" title="Search Plazi for locations around (long 47.12232/lat -18.98194)">Non Protected Area</a>, 7.64 km NW Arivonimamo, -18.97806, 47.11929, 1373 m, Uapaca woodland (ARA) (CAS); Navoatra III Non Protected Area, 7.39 km NW Arivonimamo, -18.98028, 47.12071, 1321 m, Uapaca woodland (ARA) (CAS); Navoatra IV Non Protected Area, 7.15 km NW Arivonimamo, -18.98194, 47.12232, 1323 m, Uapaca woodland (ARA) (CAS).</p> </div>	http://treatment.plazi.org/id/45336E9577B351B38C648D581ABF175A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
561C14F81F155920AD6C6310C917395C.text	561C14F81F155920AD6C6310C917395C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus norvigi Rasoamanana & Fisher 2022	<div><p>Camponotus norvigi sp. nov.</p> <p>Figures 15A, 53</p> <p>Holotype worker.</p> <p>Madagascar, Province Mahajanga, Region Sofia, Bemanevika, -14.337, 48.58874, 1606 m, montane rainforest, ex dead twig above ground, 14 January 2015 (BLF), collection code: BLF35614, specimen code: CASENT0923254 (CAS). Paratypes. Three workers with the same data as holotype: collection code: BLF35657, specimen code: CASENT0724324; collection code: BLF35614, specimen code: CASENT0724356; collection code: BLF35615, specimen code: CASENT0724357 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument black; head, mesosoma, and gastral dorsum clothed with dense, whitish hairs; anterior margin of clypeus with short rectangular lobe; dorsal outline of mesosoma almost flat; dorsum and declivitous faces of propodeum unequal in length and form an obtuse angle.</p> <p>Description minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 1.45 ± 0.24, 1.31-1.66). In full-face view, head more quadrate (CWb/CL: 0.36 ± 0.02; 0.35-0.38), posterior margin of head and lateral margin weakly convex. Eyes circular, placed laterally close to lateral head margin (PoOC/CL: 0.08 ± 0.01; 0.07-0.09). Mandibles triangular with six teeth. Clypeus convex, its anterior portion projected into a rectangular lobe. Antennal scape circular and long (SL/CS: 0.40 ± 0.03; 0.37-0.42). In lateral view, mesosoma low and long, its dorsum smoothly arcuate, metanotal suture obsolete. Humeral angle rounded. Propodeal dorsum about the same length as declivitous face (MPH/ML: 0.18 ± 0.02; 0.18-0.20). In lateral view, petiole squamiform with blunt edge, anterior face short and converging to the highest point of the petiole, posterior face straight. Ventral face of femora marginate laterally, the outer margin of tibia with long, whitish setae. Head and mesosoma coarsely reticulate-punctate, legs and gastral tergite finely imbricate. Mandible finely foveolate with sparse punctures. Pilosity and pubescence whitish, the former sparse and suberect, especially on occipital portion, frontal area, and mesosoma, though most conspicuous on the thoracic dorsum. On the gaster, the suberect hairs are long and abundant, overlapping with appressed hairs, pubescence inconspicuous and short. Petiolar node fringed with one row of whitish hair from the level of petiolar spiracle. Head including scape, mesosoma, and gaster dark brown to black, legs brownish, less dark than mesosoma.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head much wider posteriorly (CS: 1.98 ± 0.30, 1.78-2.32; CWb/CL: 0.99 ± 0.01, 0.98-1.00); lateral margins convex and converging towards base of mandible. Eyes semicircular, placed dorsolaterally midway from the occipital margin (PoOC/CL: 0.23 ± 0.01, 0.22-0.23). Clypeus much more quadrate in full-face view (ClyL/GPD: 0.72 ± 0.02, 0.69-0.73), its anterior margin slightly concave medially. Antennal scape short, just surpassing the occipital border (SL/CS: 0.90 ± 0.14, 0.75-1.00). Dorsal outline of mesosoma forms a continuous line so that the basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely reticulate above, punctate to finely areolate below.</p> <p>Distribution and biology.</p> <p>Camponotus norvigi has been collected primarily by Malaise trap and beating low vegetation, and rarely from yellow pan traps. No colonies have been collected. It is distributed mainly in montane forest, rainforest, and high-altitude rainforest in southern, western, and central Madagascar; in tropical dry forest in northern Madagascar, e.g., Res Tsingy de Bemaraha, and in gallery forest and open areas on the western portion of the island (Fig. 58B). It occurs at elevations of 20-1606 meters. This species is sympatric with C. darwinii at the edge of PN Isalo and Ranomafana, at Ambohitantely (Forêt sclerophylle).</p> <p>Discussion.</p> <p>Camponotus norvigi stands out within the species group. The color and disposition of the erect hairs covering its entire dorsum separate C. norvigi from C. darwinii. The latter species and C. ursus both have dense pubescence on their mesosoma or gastral dorsum, a character absent in C. norvigi. At first glance, C. norvigi appears closely related to the C. madagascarensis group because of its whitish pilosity, but the two differ in the structure of the mesosoma, which is low and long in the C. madagascarensis group and short and high in the C. darwinii group.</p> <p>Etymology.</p> <p>This species is named in honor of K. Norvig for support in training young scientists in Madagascar.</p> <p>Additional material examined.</p> <p>Province Antananarivo: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Analamanga Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">District of Ankazobe</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ambohitantely</a>, 46 km NE of Ankazobe, -18.198, 47.2815, 701 m, Forêt sclerophylle (MG) (CAS); Reserve Speciale d’Ambohitantely, -18.22444, 47.2774, 1490 m, montane forest (B.L. Fisher et al.) (CAS). Province Antsiranana: 9.2 km WSW Befingotra, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Res. Anjanaharibe-Sud</a>, -14.75, 49.46667, 1200 m, montane rainforest (B.L. Fisher) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Parc National de Marojejy</a>, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, -14.44333, 49.74333, 1325 m, montane rainforest (B.L. Fisher) (CAS); Parc National Montagne d’Ambre [Petit Lac road], -12.533333, 49.166668, 1125m, rainforest (R. Harin’Hala) (CAS). Province Fianarantsoa: 2 km NE Anjoma-Ramartina, -19.63333, 45.96667, 750 m, grassland (PSW) (CAS); 2 km W Andrambovato, along <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">river Tatamaly</a>, -21.51167, 47.41, 1075 m, cultivated land (tavy), recent 1-2 year-old tavy (BLF) (CAS); Ambalavao, -21.83267, 46.93867, 1020 m, urban/garden (BLF) (CAS); Belle Vue trail, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana National Park</a>, -21.2665, 47.42017, 1020 m, mixed tropical forest (R. Harin’Hala) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Fitovavy Fitovinany Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">District of Ifanadiana Belle</a> vue area, 1200 m S of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana National Park</a> entrance, -21.2665, 47.42017, 1018 m, rainforest (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Fitovavy Fitovinany Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">District of Ifanadiana</a>, 12 km W of Ranomafana, -21.25083, 47.40717, 1127 m, forest edge, open area, (MG) (CAS); JIRAMA water works near river, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana National Park</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Fianarantsoa Prov.</a>, -21.2485, 47.45217, 690m, open area near stream (R. Harin’Hala) (CAS); radio tower, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana National Park</a>, -21.25833, 47.40717, 1130 m, forest edge, mixed tropical forest, open area (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana</a>, -21.25, 47.36667 (A. Pauly) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ranomafana</a>, nr. Ifanadiana, -21.26667, 47.45, 610 m, open forest/shrubland (PSW) (CAS); stream area, 900 m E of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Isalo National Park Interpretive Center</a>, -22.62667, 45.35817, 750 m, open area near stream (R. Harin’Hala) (CAS); Vohiparara broken bridge, -21.22617, 47.36983, 1110 m, high altitude rainforest (R. Harin’Hala) (CAS). Province Mahajanga: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Parc National de Namoroka</a>, 16.9 km 317° NW Vilanandro, -16.40667, 45.31, 100 m, tropical dry forest (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Parc National Tsingy de Bemaraha</a>, 2.5 km 62° ENE Bekopaka, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Ankidrodroa River</a>, -19.13222, 44.81467, 100 m, tropical dry forest on Tsingy (FGAT) (CAS); Region Sofia, Bemanevika, -14.337, 48.58874, 1606 m, montane rainforest (BLF) (CAS). Province Toamasina: Ankerana, -18.40636, 48.80254, 1108 m, montane forest (BLF) (CAS); Corridor Forestier Analamay-Mantadia, Tsaravoniana, -18.76465, 48.41938, 1039 m, rainforest (BLF) (CAS); Toamasina Forêt Ambatovy, 14.3 km 57° Moramanga, -18.85083, 48.32, 1075 m, montane rainforest (B.L. Fisher) (CAS); Manakambahiny Atsinanana, -17.75, 48.71667, primary forest (A. Pauly) (CAS); P.N. Mantadia, -18.79167, 48.42667, 895m, rainforest (H.J. Ratsirarson) (CAS); Torotorofotsy, -18.77048, 48.43043, 1005 m, montane rainforest (B.L. Fisher et al.) (CAS). Province Toliara: 13 km NW Enakara, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Res. Andohahela</a>, -24.55, 46.8, 1180 m, montane rainforest (B.L. Fisher) (CAS); 2.7 km WNW 302° Ste. Luce, -24.77167, 47.17167, 20 m, littoral rainforest (B.L. Fisher) (CAS); 4 km N Isaka-Ivondro, -24.76667, 46.86667, 180 m, roadside (PSW) (CAS); 6 km ESE Imonty, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.40667&amp;materialsCitation.latitude=-18.26667" title="Search Plazi for locations around (long 45.40667/lat -18.26667)">Res. Andohahela</a>, -24.85, 46.75, 1200 m, rainforest (PSW) (CAS); Makay Mts., -21.227, 45.33222, 475 m, gallery forest on sandy soil (BLF) (CAS); Manatantely, 8.5 km NW Tolagnaro, -24.9875, 46.92617, 85 m, rainforest (BLF) (CAS); Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, -24.7585, 46.85367, 275 m, rainforest (BLF) (CAS); Parc National d’Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, -24.76389, 46.75167, 900 m, montane rainforest (FGAT) (CAS); Res. Andohahela, 6 km SSW Eminiminy, -24.73333, 46.8, 330 m, rainforest (PSW) (CAS); Reserve Speciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, -18.26667, 45.40667, 1050 m, montane rainforest (FGAT) (CAS).</p> </div>	http://treatment.plazi.org/id/561C14F81F155920AD6C6310C917395C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
643A9FA7C8CA5C60BD48BB038098503A.text	643A9FA7C8CA5C60BD48BB038098503A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus nossibeensis Andre 1887	<div><p>Camponotus nossibeensis Andre</p> <p>Figures 5C, 13A, 14A, 54</p> <p>Camponotus nossibeensis André, 1887: 281. Lectotype major worker, present designation, Madagascar, Province Antsiranana, Nossi-be, -13.291667, 48.258335, 146 m (Forel), AntWeb CASENT0101425 (MNHN). Paralectotype minor worker, same data as lectotype but specimen coded as CASENT0101373 (MHNG).</p> <p>Camponotus (Myrmobrachys) nossibeensis Forel, 1912: 91; 1914: 270.</p> <p>Camponotus (Myrmepomis) nossibeensis Emery, 1920: 258.</p> <p>Camponotus (Myrmopiromis) nossibeensis Wheeler, 1922: 1052; Emery, 1925: 129; Bolton, 1995: 114, 131.</p> <p>Worker diagnosis.</p> <p>Integument matte black; body a rust brown color, relatively dense on entire dorsum; anterior margin of clypeus with short rectangular lobe; dorsal outline of mesosoma weakly arcuate; declivitous face of propodeum longer than its dorsum.</p> <p>Description of minor worker.</p> <p>Large-sized species. Absolute cephalic size (CS: 2.24 0.46; 1.92-2.70). In full-face view, head slightly longer than broad, narrower in front than behind, with nearly straight posterior and lateral margins. (CWb/CL: 0.37 ± 0.03; 0.35-0.42). Eyes more elongate, elliptical, not breaking outlines of head (PoOC/CL: 0.08 ± 0.02; 0.06-0.09). Mandibles triangular withsix teeth. Clypeus carinate, anterior clypeal margin entire, projecting into a rectangular lobe with an obtuse anterolateral angle (ClyL/GPD: 0.28 ± 0.05, 0.24-0.32). Antennal scape long, surpassing the occiput by the length of one funiculus (SL/CS: 0.39 ± 0.05; 0.33-0.41). In lateral view, mesosoma short, robust, and depressed; its dorsal outline arcuate; promesonotal suture large, distinctly shiny, and glabrous; humeral angle distinctly rounded; metanotal suture obsolete; propodeal dorsum slightly convex in profile and distinctly shorter than the declivitous face into which it passes via a rounded obtuse angle (MW/ML: 0.22 ± 0.03; 0.20-0.24; MPH/ML: 0.21 ± 0.05; 0.18-0.24). In lateral view, petiolar node squamiform, anterior face confused with its dorsal face, node summit truncate, rounded triangle in dorsal view. Head, mesosoma, and gastral tergite finely and densely punctate. Mandible finely and longitudinally striate with sparse punctuation. Hairs golden yellow, erect to suberect, abundant, bending forward at its midlength on occipital region and mesosomal dorsum, but more abundant on mesonotum and propodeum dorsum; sparse, stiff, pointed, and a different length on gastral tergites. Petiole along its border with a fringe of golden, yellowish hairs not the same length. Body entirely black and a little shiny; masticatory margin reddish to dark brown.</p> <p>Description of major worker.</p> <p>With characteristics of minor worker, except: head much wider posteriorly (CS: 3.05 ± 0.41, 2.35-3.67; CWb/CL: 1.02 ± 0.04, 0.96-1.08), lateral margin almost straight and tapering anteriorly to the mandibular insertion. Eyes elliptical, moderately sized compared to the head (EL/CS: 0.20 ± 0.01, 0.1--0.23), placed dorsally next to the vertex (PoOC/CL: 0.23 ± 0.02, 0.18-0.27). Clypeus more rectangular (ClyL/GPD: 0.84 ± 0.05; 0.73-0.91). Antennal scape just surpassing the occipital margin (SL/CS: 0.77 ± 0.08; 0.68-0.91). Mandible strongly built with five large teeth, apical tooth sharp and long, and remainder decreasing in size to the basal margin. Mesosoma, pilosity, and pubescence same as minor worker. Head shiny black; anterior portion of head, including frontal area, finely alveolate punctate.</p> <p>Distribution and biology.</p> <p>Camponotus nossibeensis is an endemic species mostly known from the northern part of Madagascar, from Nosy Faly to Ambanja. This species seems very adaple because it has been collected from both humid and tropical dry forest, at altitudes 7-780 m (Fig. 58C). The data indicate that individual workers forage on the ground and on lower vegetation, while nests are mostly found in dead twigs above ground, rotten logs, dead tree stumps, and underground.</p> <p>Discussion.</p> <p>The combination of large size, big eyes, vertical declivitous face of propodeum, dense pilosity on mesosoma and gaster, and alveolate sculpture make C. nossibeensis a very distinctive species.</p> <p>Additional material examined.</p> <p>Province Antsiranana: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Foret Ambato</a> , 26.6 km 33° <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Ambanja</a>, -13.4645, 48.55167, 150 m, rainforest (B.L. Fisher) (CAS); Nosy faly, Tafiambotry, 35.3 km N Ambanja, -13.3654, 48.48775, 7 m, littoral rainforest (BLF) (CAS); Ankobahoba, 32.3 km N Ambanja, -13.39166, 48.48249, 41 m, disturbed littoral rainforest (BLF) (CAS); R.S. Manongarivo, 10.8 km 229° SW Antanambao, -13.96167, 48.43333, 400 m, rainforest (CAS); R.S. Manongarivo, 12.8 km 228° SW Antanambao, -13.97667, 48.42333, 780 m, rainforest(CAS); R.S. Manongarivo, 20.4 km 219° SW Antanambao, -14.04667, 48.40167, 1860 m, montane rainforest (CAS); R.S. Manongarivo 17.3 km 218° SW Antanambao, -14.02167, 48.41833, 1580 m, montane rainforest (B.L. Fisher) (CAS); Nosy Faly, -13.36435, 48.49137, 40 m, open secondary vegetation (BLF) (CAS); Nosy Faly, -13.3624, 48.49101, 15 m, open secondary vegetation (BLF) (CAS); Sahamalaza Peninsula, Forêt d’Anabohazo, 21.6 km 247° WSW Maromandia, -14.30889, 47.91433, 120 m, tropical dry forest (FGAT) (CAS); Nosy Be, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Reserve Naturelle Integrale de Lokobe</a> , 6.3 km 112° ESE Hellville, -13.41933, 48.33117, 30 m, rainforest (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Sahamalaza National Park</a>, -14.16361, 47.70277, 192 m, subhumid forest (MG) (CAS); Nosy Faly, -13.36255, 48.49104, 19 m, open secondary vegetation (BLF) (CAS). Province Mahajanga: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Parc National Tsingy de Bemaraha</a>, 3.4 km 93° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Bekopaka</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Tombeau Vazimba</a>, -19.14194, 44.828, 50 m, tropical dry forest (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Parc National Tsingy de Bemaraha</a>, 10.6 km ESE 123° Antsalova, -18.70944, 44.71817, 150 m, tropical dry forest on Tsingy (FGAT) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Parc National Tsingy de Bemaraha</a>, 2.5 km 62° ENE Bekopaka, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Ankidrodroa River</a>, -19.13222, 44.81467, 100 m, tropical dry forest on Tsingy (FGAT) (CAS); Réserve Spéciale Marotandrano, Marotandrano 48.3 km S Mandritsara, -16.28322, 48.81443, 865 m, transitional humid forest (BLF) (CAS); Ampamakiambato, 45 km SW Ambanja, -13.97545, 48.15929, 145 m, disturbed forest, in tsingy (BLF) (CAS); Manerinerina, 76.6 km N Antsohihy, -14.10744, 48.11046, 247 m, disturbed forest (BLF) (CAS); Réserve forestière Beanka, 50.7 km E Maintirano, -17.88021, 44.46877, 140 m, tropical dry forest on tsingy (BLF) (CAS); Réserve forestière Beanka, 54.3 km E Maintirano, -18.06009, 44.54086, 262 m, tropical dry forest on tsingy (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Melaky Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">District of Besalampy</a>, Marofototra dry forest, 17 km W of Besalampy, -16.72167, 44.42367, 51 m, dry wash in the dry forest (MG) (CAS); Bemeraha, 9 km E. Antsalova, -18.65, 44.71667 (D. C. Lees) (PSWC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Foret de Tsimembo</a> , 8.7 km 336° NNW Soatana, -19.02139, 44.44067, 20 m, tropical dry forest (FGAT) (CAS). Province Toliara: Belalanda, -23.29833, 43.64648, 5 m, spiny forest/thicket, wetlands (Frontier Wilderness Project) (CAS); near road, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Zombitse National Park</a>, -22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin’Hala) (CAS); near ANGAP office, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Zombitse National Park</a>, -22.8865, 44.69217, 840 m, deciduous spiny forest (R. Harin’Hala) (CAS); 15 km E Sakaraha, -22.9, 44.68333, 760 m, tropical dry forest (PSW) (PSWC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Makay Mts.</a>, -21.30997, 45.12946, 590 m, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Dry forest</a> on sandy soil (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Makay Mts.</a>, -21.227, 45.33222, 475 m, gallery forest on sandy soil (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38333&amp;materialsCitation.latitude=-22.48333" title="Search Plazi for locations around (long 45.38333/lat -22.48333)">Makay Mts.</a>, -21.21013, 45.35462, 540 m, gallery forest with bamboo (BLF) (CAS); Parc National de Zombitse, 17.7 km 98° E Sakaraha, -22.88833, 44.70167, 760 m, tropical dry forest (FGAT) (CAS); 9 km NNW Ranohira, P.N. Isalo, -22.48333, 45.38333, 800 m, tropical dry forest (PSW) (PSWC).</p> </div>	http://treatment.plazi.org/id/643A9FA7C8CA5C60BD48BB038098503A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
90859E1B4168588A97CB8325A074D9A9.text	90859E1B4168588A97CB8325A074D9A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus radovae Forel 1886	<div><p>Camponotus radovae Forel, 1886</p> <p>Figures 14B, 16D, 18C, D, 55</p> <p>Camponotus radovae Forel, 1886: civ. Lectotype minor worker, present designation, Madagascar, Province Antananarivo (Camboué), AntWeb CASENT0101363 (MHNG). Paralectotypes two workers with the same data as lectotype but specimen coded as CASENT0101362, CASENT0101364 (MHNG).</p> <p>Camponotus (Myrmobrachys) radovae Forel, 1912: 91.</p> <p>Camponotus (Myrmepomis) radovae Emery, 1920: 258.</p> <p>Camponotus (Myrmopiromis) radovae Wheeler, 1922: 1052; Emery, 1925: 129; Bolton, 1995: 119, 131.</p> <p>Camponotus radovae-darwinii Forel, 1891: 46. Lectotype minor worker, present designation, Madagascar, Province Antananarivo (Camboué), AntWeb CASENT0101367 (MHNG). Paralectotype one worker with the same data as lectotype but specimen coded as CASENT0101115 (MHNG).</p> <p>Camponotus (Myrmopiromis) radovae-darwinii Wheeler, 1922: 1053; Emery, 1925: 129; Bolton, 1995: 119. Syn, nov.</p> <p>Worker diagnosis.</p> <p>Integument matte black with sparse, yellowish brown hairs arranged in a series of transverse rows, pubescence reduced to absent on mesosomal dorsum. Anterior clypeal margin with a distinct produced rectangular lobe. Petiole cuneate in profile.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 1.59 ± 0.22, 1.43-1.77). In full-face view, head longer than broad, narrower in front than in back, with feebly convex occipital and lateral sides (CWb/CL: 0.37 ± 0.03; 0.35-0.40). Eyes circular, placed much closer to the lateral sides (PoOC/CL: 0.08 ± 0.01, 0.07-0.09). Mandibles triangular with six teeth. With head in full-face view, clypeus not carinate, its anterior margin produced, forming a short rectangular lobe with a sharp, lateral angle. Antennal scape circular, long, surpassing the occiput by the length of one basal funiculus (SL/CS: 0.39 ± 0.03, 0.37-0.41). In lateral view, anterodorsal corner of pronotum broadly rounded, mesosomal dorsal almost straight, propodeal dorsum shorter than declivity, both straight in profile and meeting at a blunt angle (MW/ML: 0.21 ± 0.02, 0.20-0.23; MPD/ML: 0.24 ± 0.02, 0.22-0.26). Promesonotal and metanotal sutures form bold, polished lines. In lateral view, petiolar node cuneate, with a short, vertical, anterior face which reaches toward the flattened posterior faces. Head and mesosoma finely alveolate, gaster more finely strigulate. Mandible finely foveate with sparse punctures. Pilosity of dorsal head and body consists of fine and pointed brownish yellow hairs, long and suberect on frontal area and mesosomal dorsum; mostly short and sparse on gastral tergites; petiolar dorsum edge with 4 to 5 pairs of brownish yellow hairs. Body entirely matte black, apical portion of mandible, basitarsi, and funiculus reddish brown.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head somewhat cordate, with nearly straight posterior margin and slightly convex lateral sides (CS: 2.76 ± 0.12, 2.59-2.85; CWb/CL: 1.05 ± 0.02, 1.02-1.07). Eyes elliptical, placed dorsally midway from the occipital margin (PoOC/CL: 0.24 ± 0.01, 0.23-0.25). Clypeus much more quadrate in full-face view with truncate anterior margin (ClyL/GPD: 0.76 ± 0.13, 0.57-0.86). Antennal scape short, just surpassing the occipital border (SL/CS: 0.74 ± 0.03, 0.72-0.79). Dorsal outline of mesosoma interrupted by pronotal and mesonotal sutures, basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely punctate, frontal area and vertex with deep punctures.</p> <p>Distribution and biology.</p> <p>Camponotus radovae is mainly distributed in the southern part of Madagascar in Zombitse National Park (Fig. 58D). This species is encountered in native forest habitats such as deciduous dry forest, tropical dry forests, gallery forests, and spiny forests and thickets. It was sampled mostly from Malaise traps and by beating lower vegetation, on the ground, and twice on leaf litter at altitudes 20-840 m.</p> <p>Discussion.</p> <p>Within the Camponotus darwinii group, Camponotus radovae cannot be confused with C. darwinii or C. themistocles since the latter two differ by size and pilosity color. However, C. radovae has reduced pubescence and is smaller in size.</p> <p>Additional material examined.</p> <p>Province Toliara: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">District of Sakaraha</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Zombitse National Park</a>; 900 m N from ANGAP <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Entrance</a> office, -22.8405, 44.73117, 823 m, spiny deciduous forest (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">District of Toliara</a> II, Mikea decidious dry forest 3 km N Andranomavo village, -22.90367, 43.4755, 30 m, Deciduous dry forest (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Atsimo Andrefana Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">District of Toliara</a> II, Mikea spiny forest 8 km N Andranomavo village, -22.91333, 43.39883, 37 m, spiny forest (MG) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Mikea Forest</a>, deciduous dry forest, -22.90367, 43.4755, 30 m, deciduous dry forest (R. Harin’Hala) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Mikea Forest</a>, spiny forest, -22.91333, 43.48222, 37 m, spiny forest (R. Harin’Hala) (CAS); near ANGAP office, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Zombitse National Park</a>, -22.8865, 44.69217, 840 m, deciduous spiny forest (R. Harin’Hala) (CAS); near road, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Zombitse National Park</a>, -22.8405, 44.73117, 825 m, spiny deciduous forest (R. Harin’Hala) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Parc National de Zombitse</a>, 17.7 km 98° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.6096&amp;materialsCitation.latitude=-23.03832" title="Search Plazi for locations around (long 43.6096/lat -23.03832)">Sakaraha</a>, -22.88833, 44.70167, 760 m, tropical dry forest (FGAT) (CAS); Parc National de Zombitse, 19.8 km 84° E Sakaraha, -22.84333, 44.71, 770 m, tropical dry forest (FGAT) (CAS); Ranobe, -23.03975, 43.6109, 30 m, spiny forest/thicket (Frontier Project) (CAS); Ranobe, -23.03957, 43.61032, 30 m, spiny forest/thicket (Frontier Project) (CAS); Ranobe, -23.03957, 43.61017, 30 m, spiny forest/thicket (Frontier Project) (CAS); Ranobe, -23.03883, 43.60982, 30 m, gallery forest (Frontier Wilderness Project) (CAS); Ranobe, -23.03952, 43.61015, 20 m, gallery forest (Frontier Wilderness Project) (CAS); Ranobe, -23.03832, 43.6096, 20 m, gallery forest (Frontier Wilderness Project) (CAS).</p> </div>	http://treatment.plazi.org/id/90859E1B4168588A97CB8325A074D9A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
02D9DC15904D5058BCD8DF2156D26ACC.text	02D9DC15904D5058BCD8DF2156D26ACC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus themistocles (Rasoamanana & Fisher 2022) Forel 2022	<div><p>Camponotus themistocles Forel stat. nov.</p> <p>Figures 5D, 15B, 16C, 18A, B, 56</p> <p>Camponotus darwinii themistocles Forel, 1910: 456. Holotype minor worker, Madagascar, Fort Dauphin S. E. Madagascar, AntWeb CASENT0101381 (MHNG).</p> <p>Camponotus (Myrmobrachys) darwinii themistocles Forel, 1914: 271.</p> <p>Camponotus (Myrmepomis) darwinii themistocles Emery, 1920: 258. Stat.nov.</p> <p>Camponotus darwinii Subspecies of Camponotus darwinii Santschi, 1911: 133.</p> <p>Camponotus Myrmopiromis Subspecies of Camponotus (Myrmopiromis) Wheeler, 1922: 1051; Emery, 1925: 128.</p> <p>Worker diagnosis.</p> <p>Integument dark brown with rust or red-brown suberect pilosity. Anterior clypeal margin with rectangular projection. Petiole scale thin with sharp border.</p> <p>Description of minor worker.</p> <p>Medium-sized. Absolute cephalic size (CS: 1.82 ± 0.37; 1.53-1.98). In full-face view, head slightly longer than broad, a little narrower in front than in back, with straight occipital and feebly convex lateral margins (CWb/CL: 0.36 ± 0.03; 0.34-0.38). Eyes circular, placed much closer to the lateral sides (PoOC/CL: 0.07 ± 0.01; 0.07-0.08). Mandibles triangular with sixteeth. With head in full-face view, anterior clypeal margin produced, entire, and angularly projecting in the middle. Antennal scape circular, long, surpassing the occiput by the lengths of two basal funiculi (SL/CS: 0.41 ± 0.05; 0.38-0.44). In lateral view, anterodorsal corner of pronotum broadly rounded, mesosomal dorsum moderately arcuate, propodeum dorsum a little shorter than the declivity, each straight in profile and meeting at an obtuse angle (MW/ML: 0.21 ± 0.02, 0.20-0.22; MPD/ML: 0.24 ± 0.03; 0.22-0.26). In lateral view, petiolar node wedge-shaped, with feebly convex anterior and flattened posterior faces. Ants entirely reticulate-punctate. Mandible finely striolate with sparse punctures. Pilosity and pubescence brownish yellow, the former long and curved forward especially on vertex, frontal area, and mesosomal dorsum; mostly short and sparse on gastral tergites; petiolar dorsum edge with four or five pairs of brownish yellow hairs. Body color matte black, apical portion of mandible reddish.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head somewhat cordate, with nearly straight posterior margin and slightly convex lateral sides (CS: 2.68 ± 0.36, 2.26-3.15; CWb/CL: 1.05 ± 0.04, 0.99-1.11). Eyes elliptical, placed dorsally midway from the occipital margin (PoOC/CL: 0.23 ± 0.01, 0.22-0.25). Clypeus much more quadrate in full-face view with truncate anterior margin (ClyL/GPD: 0.82 ± 0.07, 0.73-0.93). Antennal scape short, just surpassing the occipital border (SL/CS: 0.75 ± 0.03, 0.71-0.79). Dorsal outline of mesosoma interrupted by pronotal and mesonotal sutures, basal portion of propodeum forms an obtuse angle with the declivity. Lateral portion of head finely punctate, frontal area and vertex with deep punctures.</p> <p>Distribution and biology.</p> <p>Camponotus themistocles has been collected at three localities in southern Madagascar (Fig. 58E): two are littoral forest and one is gallery forest. All sites are at low altitudes from 10 to 30 meters. Camponotus themistocles is a ground nester; most collections have been made from sifted litter, rotten logs, and dead twigs above the ground.</p> <p>Discussion.</p> <p>Camponotus themistocles differs from C. radovae by its larger size and the sculpture of the lateral sides o f its mesosoma, which is finely reticulate-punctate in the former and widely striolate in the later. In addition, in lateral view, the petiole scale of C. themistocles has an acute summit but is blunt in C. radovae. Pubescence is long and abundant in C. themistocles, and short and dilute in C. radovae. Morphometrical analysis reveals that C. themistocles and C. ellioti are the same size, but differ in integument coloration and pilosity pattern on the gastral tergite.</p> <p>Additional material examined.</p> <p>Province Toliara: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.61012&amp;materialsCitation.latitude=-23.04085" title="Search Plazi for locations around (long 43.61012/lat -23.04085)">Foret de Petriky</a> , 12.5 km W 272° <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.61012&amp;materialsCitation.latitude=-23.04085" title="Search Plazi for locations around (long 43.61012/lat -23.04085)">Tolagnaro</a>, -25.06167, 46.87, 10 m, littoral rainforest (B.L. Fisher) (CAS); Forêt Mandena 8.5 km N Tolagnaro, -24.95267, 47.0025, 20 m, littoral rainforest (BLF) (CAS); Mandena, 8.4 km NNE 30° Tolagnaro, -24.95167, 47.00167, 20 m, littoral rainforest (B.L. Fisher) (CAS); Ranobe, -23.04085, 43.61012, 30 m, gallery forest (Frontier Wilderness Project) (CAS).</p> </div>	http://treatment.plazi.org/id/02D9DC15904D5058BCD8DF2156D26ACC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
5C1193616D2C5AF3B854A8A0A4C56EDB.text	5C1193616D2C5AF3B854A8A0A4C56EDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus tsimelahy Rasoamanana & Fisher 2022	<div><p>Camponotus tsimelahy sp. nov.</p> <p>Figures 43B, 45B, 46A, 69</p> <p>Holotype worker.</p> <p>Madagascar, Province Toliara, Parc National d 'Andohahela, Forêt de Manatalinjo, 33.6 km 63° ENE Amboasary, 7.6 km 99° E Hazofotsy, -24.81694, 46.61, 150 m, spiny forest/thicket, 12-16 January 2002 (BLF), collection code: BLF04814, specimen code: CASENT0446651 (CAS). Paratypes. One worker with same data as holotype but specimen code CASENT0446549; 3 workers from Parc National d’ Andohahela, Forêt d’ Ambohibory, 1.7 km 61° ENE Tsimelahy, 36.1 km 308° NW Tolagnaro, -24.93, 46.6455, tropical dry forest, 12-16 January 2002 (BLF), collection code: BLF04918, specimen code: CASENT0444910 (CAS).</p> <p>Note.</p> <p>Habitat: eucalyptus plantation, grassland, savanna grassland, Uapaca forest, Uapaca woodland, urban/garden. Altitude: 1300-1987 m. Province: Antananarivo. Microhabitat: sifted litter (leaf mold, rotten wood), under stones, on low vegetation, ex soil, ground forager(s), ground nest. Method: MW 20 sample transect, 10 m; SD 20 digging sample transect, 10 m; 20 sardine and ground tuna fish baits; Malaise, 2 traps.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 0.79 ± 0.04; 0.77-0.81). In full-face view, head ovate, usually somewhat longer than broad, lateral margin of head straight and tapering to front (CWb/CL: 0.29 ± 0.02, 0.28-0.29). Eyes subelliptical, protruding, their borders smoothly aligned to lateral margins of head (PoOC/CL: 0.09 ± 0.00, 0.08-0.09). Mandibles triangular with six teeth. Anterior clypeal margin not produced, entire, broadly rounded. Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL/CS: 0.41 ± 0.03, 0.40-0.42). In lateral view, mesosoma without anterolateral margination, promesonotum distinctly convex, mesopropodeal suture deeply impressed so that propodeal dorsum raised to the level of mesopropodeal suture, propodeum angular in profile, with unequal basal and declivitous faces, the former horizontal, the latter straight to evenly concave (MW/ML: 0.19 ± 0.02, 0.17-0.20; MPH/ML: 0.13 ± 0.01, 0.13-0.14). In lateral view, petiole cuneate with convex anterior face and flat dorsal face. Body finely imbricate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; two pairs on each side of lateral corner of propodeum; three pairs on lateral margins of petiole. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remaining segment dark brown to black.</p> <p>Distribution and biology.</p> <p>This new species seems to prefer deciduous forest habitats such as tropical dry forest, gallery forest, spiny forest or thicket, and transitional forest, but is much rarer in humid habitats at lower altitudes (Fig. 72D).</p> <p>Discussion.</p> <p>Camponotus tsimelahy may be confused with C. repens and C. ihazofotsy; all three species share the same sculpturation, form of petiole, and type of suberect setae. However, in profile, the dorsal outline of the mesosoma presents a slightly concave metanotal groove. See the discussion section under C. ihazofotsy.</p> <p>Etymology.</p> <p>The species epithet is in reference to the type locality.</p> <p>Additional material examined.</p> <p>Province Toliara: 18.4 km N Amboasary, -24.87367, 46.39767, 85 m, spiny forest/thicket (BLF) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.39767&amp;materialsCitation.latitude=-24.87367" title="Search Plazi for locations around (long 46.39767/lat -24.87367)">Anosy Region</a>, District of Amboasary, Andohaela National Park Parcelle III, Ihazofotsy, 32 km NE Amboasary, -24.83083, 46.53617, 58 m, dry forest, spiny forest (Mike Irwin, Frank Parker, Rin’ha) (CAS); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.53617&amp;materialsCitation.latitude=-24.83083" title="Search Plazi for locations around (long 46.53617/lat -24.83083)">Anosy Region</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.53617&amp;materialsCitation.latitude=-24.83083" title="Search Plazi for locations around (long 46.53617/lat -24.83083)">District of Fort-Dauphin</a>, Andohaela National Park Parcelle II, Tsimelahy, 42 km W of Fort-Dauphin, -24.93683, 46.62667, 176 m, transitional forest (Mike Irwin, Frank Parker, Rin’ha) (CAS); Ifaty 22 km N, -23.18333, 43.61667, 30 m,beach dunes (M.E. Irwin and E.I. Schlinger) (CAS); Ihazofotsy - Parcel III, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.48683&amp;materialsCitation.latitude=-24.83483" title="Search Plazi for locations around (long 46.48683/lat -24.83483)">Andohahela National Park</a>, transitional forest, -24.83483, 46.48683, 80 m, transition between spiny and dry deciduous forests (M.E. Irwin, F.D. Parker, R. Harin’Hala) (CAS); Parc National d’Andohahela, Forêt d’ Ambohibory, 1.7 km 61° ENE Tsimelahy, 36.1 km 308° NW Tolagnaro, -24.93, 46.6455, 300 m, tropical dry forest (FGAT) (CAS); Parc National d’Andohahela, Forêt de Manatalinjo, 33.6 km 63° ENE Amboasary, 7.6 km 99° E <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.62667&amp;materialsCitation.latitude=-24.93683" title="Search Plazi for locations around (long 46.62667/lat -24.93683)">Hazofotsy</a>, -24.81694, 46.61, 150 m, spiny forest/thicket (FGAT) (CAS) Tsimelahy - Parcel II, Andohahela National Park, transitional forest, -24.93683, 46.62667, 180 m, transitional forest (M.E. Irwin, F.D. Parker, R. Harin’Hala) (CAS).</p> </div>	http://treatment.plazi.org/id/5C1193616D2C5AF3B854A8A0A4C56EDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
CC6DAF11264F5C95B1453B2E816ECD40.text	CC6DAF11264F5C95B1453B2E816ECD40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus ursus Forel 1886	<div><p>Camponotus ursus Forel</p> <p>Figures 2C, 16A, 17A, 57</p> <p>Camponotus ursus Forel, 1886: ci. Lectotype minor worker, present designation, Madagascar (Forel), AntWeb CASENT0101374 (MHNG) [examined]. Paralectotype, one minor worker same data as lectotype but specimen coded as CASENT0101375 (MHNG).</p> <p>Camponotus (Myrmobrachys) ursus Forel, 1912: 91; 1914: 271.</p> <p>Camponotus (Myrmepomis) ursus Emery, 1920: 258.</p> <p>Camponotus (Myrmopiromis) ursus Wheeler, 1922: 1053; Emery, 1925: 129; Bolton, 1995: 128, 131.</p> <p>Worker diagnosis.</p> <p>Integument shiny black; anterior margin of clypeus with short, rectangular lobe; dorsum of mesosoma with dense, decumbent, golden-yellow setae; gastral dorsum with widely distributed, short, subdecumbent setae.</p> <p>Description of minor worker.</p> <p>Medium-sized species. Absolute cephalic size (CS: 1.54 ± 0.19; 1.43-1.64). In full-face view, head somewhat longer than broad, narrower in front than behind, with slightly convex lateral and posterior sides (CWb/CL: 0.37 ± 0.02; 0.36-0.38). Eyes elliptical, sublateral at midpoint from the lateral sides (PoOC/CL: 0.07 ± 0.02; 0.06-0.08). Mandibles triangular with six teeth. Clypeus not carinate, produced into a short, rectangular lobe. Antennal scape long, surpassing the occiput by the length of one basal funiculus. (SL/CS: 0.38 ± 0.02; 0.37-0.39). In lateral view, dorsal contour of mesosoma smoothly convex, humeral angle broadly rounded; in dorsal view, mesonotal suture distinct but not impressed, mesonotal suture obsolete so that mesonotum and propodeum are fused together, propodeum with convex base and sloping declivity (MPH/ML: 0.18 ± 0.02; 0.16-0.19). Petiole narrow, cuneate in profile, with short anterior face tapering dorsally to the flattened posterior face, its border rather sharp, produced upwards as a blunt angle in the middle. Head dorsum finely reticulate-punctate with shallow, sparse punctures; occipital region transversally striate-reticulate; lateral face of mesosoma, declivitous face, and petiolar face finely strigulate; legs finely reticulate; dorsum of mesosoma finely reticulate-striate-punctate with sparse excavation from which one suberect setae arises. Dorsum of gastral tergite finely striate-reticulate transversally, with sparse, small punctures. Mandible finely rugose with sparse, large punctures. Hairs golden yellow, the former abundant, long, and bending forward on entire mesosoma dorsum; suberect, short, and sparsely distributed on gastral tergites; the latter short and conspicuous on the abdominal segment, five pairs of erect hairs present on vertex. Body shiny black; scape, two basal funiculi, mandible, tarsi, and tibiae reddish.</p> <p>Description of major worker.</p> <p>Characteristics of minor workers, except: head as broad as long, with occipital and lateral margin almost straight (CS: 2.14 ± 0.15, 2.00-2.39; CWb/CL: 0.89 ± 0.07, 0.84-1.03). Eyes circular, placed dorsally next to the vertex (PoOC/CL: 0.23 ± 0.01, 0.21-0.24). Anterior clypeal margin forms a short, rounded lobe (ClyL/GPD: 1.03 ± 0.41, 0.73-1.78). Antennal scape short, just reaching the occipital border (SL/CS: 0.82 ± 0.03, 0.78-0.86). Dorsal outline of mesosoma almost flat, propodeum dorsum naked and the same length as the sloping declivity.</p> <p>Distribution and biology.</p> <p>Camponotus ursus is found in two different habitats: primary forest in the eastern portion and urban/garden areas in the central highlands of Madagascar (Fig. 58F). It is found foraging on low vegetation or inside branches above the ground. It occurs at altitudes above 1,200 meters.</p> <p>Discussion.</p> <p>Camponotus ursus is recognizable within the C. darwinii species group on the basis of its distinct mesosomal pilosity. In addition, it is the only species with reddish brown basitarsi. However, C. ursus is unlikely be confused with C Camponotus darwinii for several reasons. First, the body of the latter is much larger. Second, the head and gastral segment of C. ursus are covered with fine, short, sparse, and yellowish setae, giving the ant a glossy appearance that is completely different than that of C. darwinii.</p> <p>Additional material examined.</p> <p>Province Antananarivo: Ambatomanjaka; Miarinarivo, -18.766947, 46.869107, 1343 m (MHNG); Ankazobe, -18.31617, 47.11583, 1241 m (BLF) (CAS). Province Toamasina: Manakambahiny Atsinanana, -17.75, 48.71667 (A. Pauly) (CAS).</p></div> 	http://treatment.plazi.org/id/CC6DAF11264F5C95B1453B2E816ECD40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
30EEC29A33AB5533BC303620A258F4FA.text	30EEC29A33AB5533BC303620A258F4FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camponotus zoro Rasoamanana & Fisher 2022	<div><p>Camponotus zoro sp. nov.</p> <p>Figures 11A, B, 33C, D, 72</p> <p>Holotype worker.</p> <p>Madagascar, Province Fianarantsoa, Forêt d’ Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest, Malaise trap, 22-26 January 2003 (BLF), collection code: BLF07155, specimen code: CASENT0049853 (CAS). Paratype. One worker. Madagascar, Province Antananarivo, 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, -18.47333, 47.96, 1300 m, montane rainforest, pitfall trap, 5-13 December 2000 (BLF), collection code: BLF02370, specimen code: CASENT0409150 (CAS).</p> <p>Worker diagnosis.</p> <p>Integument black, entirely but finely sculptured, appendages dark brown; scape circular; anterior margin of clypeus rounded, not projected; promesonotal suture impressed, dorsolateral margin of propodeal margin.</p> <p>Description of minor worker.</p> <p>Medium-sized. Absolute cephalic size (CS: 1.02 ± 0.28; 0.92-1.14). In full-face view, head subquadrate (CWb/CL: 0.39 ± 0.01, 0.39-0.40), posterior margin of head broadly convex, lateral margin of head slightly convex. Eyes sub-elliptical, placed laterally, closer to the posterior margin (PoOC/CL: 0.09 ± 0.01; 0.08-0.09). Mandibles triangular with five teeth. Anterior clypeal margin rounded and not produced anteriorly (ClyL/GPD: 0.24 ± 0.01). Antennal scape virtually circular, surpassing posterior cephalic margin by the length of two basal flagella (SL/CS: 0.34 ± 0.04; 0.33-0.36). In lateral view, metanotal groove deeply impressed; pronotum, mesonotum, and propodeum dorsum marginated laterally; mesonotum dorsum inclined posteriorly; propodeum dorsum slightly concave, its lateral carina forming a blunt tubercle with the lateral margin of the declivity (MPD/ML: 0.25 ± 0.02; 0.24-0.25). In lateral view, petiolar node strongly convex anterodorsally and flat posteriorly, node summit convex. Head finely areolate and mesosoma feebly areolate-costulate, gastral tergite finely imbricate. Mandible glabrous. Vertex with three pairs of filiform erect setae; mesonotum and propodeum dorsum with single pair of whitish setae; gastral tergite with short, whitish setae aligned transversally on the middle length of each segment; petiolar node with two or three pairs of whitish setae on its dorsum. Body uniformly dark blackish brown to black, appendages brownish, mandible light brown.</p> <p>Description of major worker.</p> <p>With characteristics exactly the same as minor worker, but head more trapezoidal (CS: 1.74 ± 0.04, 1.71-1.77).</p> <p>Distribution and biology.</p> <p>Camponotus zoro has been collected by beating low vegetation, sifting leaf litter, and baiting with honey and tuna fish. One worker has been found in Malaise traps from deciduous forest in southern Madagascar (Fig. 72E). In addition, this species has been collected from montane rainforest and Uapaca woodland.</p> <p>Discussion.</p> <p>Camponotus zoro is characterized by a shallow but distinct metanotal groove, and should not be confused with any other species because of the margination of its mesosoma.</p> <p>Etymology.</p> <p>The new species is named after the structure of the mesosoma; " Camponotus zoro "means angle.</p> <p>Additional material examined.</p> <p>Province Antananarivo: Navoatra I <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.26427&amp;materialsCitation.latitude=-16.7755" title="Search Plazi for locations around (long 49.26427/lat -16.7755)">Non Protected Area</a>, 7.64 km NW Arivonimamo, -18.97806, 47.11929, 1373 m, Uapaca woodland (ARA) (CAS); Ankalalahana, -19.0531, 47.11304, 1314 m, Uapaca woodland (H.E. Marti et al.) (CAS); 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, -18.47333, 47.96, 1300 m, montane rainforest (FGAT) (CAS); Reg. Analamanga, St. Forestiѐre Mandraka, -18.9183, 47.91687, 1285 m, montane rainforest (B.L. Fisher, F.A. Esteves et al.) (CAS). Province Fianarantsoa: Forêt d’ Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest (FGAT) (CAS). Province Toamasina: Réserve Spéciale Ambatovaky, Sandrangato river, -16.7755, 49.26427, 430 m, rainforest (BLF) (CAS).</p> </div>	http://treatment.plazi.org/id/30EEC29A33AB5533BC303620A258F4FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasoamanana, Nicole;Fisher, Brian L.	Rasoamanana, Nicole, Fisher, Brian L. (2022): A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137-231, DOI: http://dx.doi.org/10.3897/zookeys.1081.71872, URL: http://dx.doi.org/10.3897/zookeys.1081.71872
