taxonID	type	description	language	source
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	description	Figs 2, 3, 4	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	materials_examined	Type. CAMEROON • [Littoral] Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 22 Dec. 2022; fr.; B. Sonke & M. Simo 7127; holotype: YA; isotypes: BR, BRLU, E, K, MA [MA 959269], MO, P.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	diagnosis	Diagnosis. Haec species a Drypete verrucosa Pierre ex Hutch. stipulis 1.9 - 2.2 mm longis anguste triangularibus caducis, petiolis (2.6 -) 3 - 4.6 (- 5.1) mm longis confertim subtiliter rugosis, laminis foliorum (6.5 -) 7 - 11.1 (- 13.7) cm longis ad basim plerumque obliquis aliquando asymmetricis subter pubescentibus imprimis in nervis, inflorescentiis in truncum et ramorum principalium positis constantibus ex 1 - 7 (- 18) floribus, pedicello florum masculinorum (9.2 -) 14.7 - 20 mm longo, sepalis florum masculinorum 4.7 - 4.9 mm longis, disco florum masculinorum 3.1 - 3.3 mm diametro aliquantum exili sparsim pubescenti, staminibus 19 - 20, filamento 1.5 - 1.6 mm longo, anthera 2.2 - 2.4 mm longa anguste ellipsoidea sparsim pubescenti, pedicello florum femininorum 5.5 - 6.2 mm longo, sepalis florum femininorum 2.6 - 3.2 mm longis, disco florum femininorum 2.1 - 2.4 mm diametro glabro, fructibus (18 -) 20 - 29 (- 31) x 17 - 23 (- 25) mm late ovoideis, exocarpio 1 - 3 mm lato phymatodeo brunneo diluto luteolo tum aurantiacescenti vel rubescenti differt.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	description	Description. Tree up to 20 m tall, with plagiotropic sinuous branches; trunk up to 20 cm in diameter, fluted; bark greenish-brown, smooth, rough and flaking off when old, slash light brown, later dark brown; young branchlets subterete, slightly sulcate, dark-coloured when dry, sparingly and shortly pubescent, glabrescent, trichomes 0.1 - 0.3 mm; terminal buds scaly, scales 1.8 - 2.8 x 0.7 - 2 mm, ovate, firm, slightly keeled, shortly pubescent outside with trichomes 0.1 - 0.3 mm, glabrous inside, ciliate, cilia 0.1 - 0.3 (- 0.5) mm. Leaves: stipules 1.9 - 2.2 x 0.5 - 0.9 mm, narrowly triangular, sparsely and shortly pubescent outside, glabrescent, trichomes 0.2 - 0.3 mm, glabrous inside, sparingly ciliate near the base, cilia 0.2 - 0.4 mm, caducous; petiole (2.6 -) 3 - 4.6 (- 5.1) mm long, (0.9 -) 1.1 - 1.6 (- 1.8) mm in diameter, finely transversely wrinkled, sparsely and shortly pubescent, glabrescent, old ones slightly blistered, trichomes 0.1 - 0.4 mm; blade (6.5 -) 7 - 11.1 (- 13.7) x (2.8 -) 3 - 5.8 (- 6.4) cm, elliptic, papery to subcoriaceous, base acute, sometimes slightly obtuse, oblique, sometimes asymmetrical, rounded on one side, basal sides usually meeting the petiole at the same point, margin crenulate-serrulate for most of its length, crenulae / teeth 0.1 - 0.6 mm, acute, frequently blunt, flat to slightly recurved near the blade base, shortly acuminate, apex 4 - 8 mm, abaxial surface between the nerves glabrous; midrib smooth, slightly longitudinally wrinkled when dry, glabrous; first order lateral veins 5 - 8 pairs, ascending, more or less regularly spaced, slightly depressed above, prominent beneath, courses curved and anastomosing well within the margin, oriented at (55 -) 60 - 73 (- 77) ° to the midrib, glabrous; second order venation loosely reticulate, hardly or not raised above, slightly so beneath. Male inflorescence borne on small cushion-like excrescences of the trunk and main branches (categories I and II), in clusters of (1 -) 3 - 7 flowers; bracts 0.7 - 0.8 x 0.5 - 0.9 mm, ovate to suborbicular, minutely pubescent outside, glabrous inside, minutely ciliate, trichomes and cilia 0.1 - 0.2 mm, barely distinguishable. Male flowers pedicellate, yellowish green or brownish green; pedicel (9.2 -) 14.7 - 20 mm long, 0.5 - 0.9 mm in diameter, slender, very sparingly and shortly pubescent, trichomes 0.1 - 0.2 mm; sepals 5 - 6, 4.7 - 4.9 x 4.2 - 5.1 mm, widely ovate, acute, cucullate, imbricate, the outer often smaller than the inner, glabrous outside and inside, margin minutely ciliate for most of its length, cilia 0.1 - 0.2 mm; stamens 19 - 20, one-whorled, surrounding the disc, more or less enveloped by its marginal lobes, filaments 1.5 - 1.6 mm, white, anthers 2.2 - 2.4 mm long, 0.6 - 0.8 mm in diameter, narrowly ellipsoid, sub-basifixed, introrse, pale yellow, sparsely pubescent, trichomes 0.1 - 0.2 mm; disc ca 0.2 mm high, 3.1 - 3.3 mm in diameter, rather flat, thin, rugose, sparsely and minutely pubescent, trichomes to 0.1 mm, pistillode to 0.4 mm, laminate, 3 - branched. Female inflorescence borne on small cushion-like excrescences of the trunk and main branches (categories I and II), in clusters of 1 - 7 (- 18) flowers; bracts 0.7 - 1.1 x 0.5 - 0.6 mm, otherwise similar to the male ones. Female flowers pedicellate, yellowish green or brownish green, sometimes pinkish; pedicel (5.5 -) 12 - 26 mm long, 0.5 - 0.9 mm in diameter, more or less robust, very sparingly and shortly pubescent, trichomes 0.1 - 0.2 mm; sepals 5 - 6, 2.6 - 3.2 x 2.5 - 3.4 mm, otherwise similar to the male ones; disc 0.2 - 0.3 mm high, 2.1 - 2.4 mm in diameter, concave, cupulate, fleshy, glabrous; style 1, sessile, unbranched; stigmas 3, obdeltoid, stigmatic surface ca 0.5 mm long, 1.1 - 1.2 mm wide; ovary 1.3 - 1.4 mm long, 2 - 2.2 mm in diameter, subglobose, 3 - celled, densely and minutely pubescent, trichomes 0.1 - 0.2 mm. Fruits (18 -) 20 - 29 (- 31) mm long, 17 - 23 (- 25) mm in diameter, widely ovoid, with caducous sepals and stigmas, exocarp warty, yellowish brown turning orange or red when ripe, shortly pubescent, trichomes 0.1 - 0.4 mm, mesocarp 1 - 3 mm thick when dried, hard, endocarp 0.8 - 1.3 mm thick, bony, 3 (- 4) - seeded; seeds ca 16.8 mm long, ca 17.2 mm in diameter; fruiting pedicel (15 -) 21 - 39 (- 42) mm long, 1.7 - 3.6 mm in diameter, very sparingly and shortly pubescent, trichomes 0.1 - 0.2 (- 0.4) mm.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	distribution	Distribution and chorology. Cameroon (Centre, East, Littoral, and West Regions) and eastern Nigeria (Adamawa) (Fig. 2). Lower Guinea subcentre of endemism (White 1979).	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	etymology	Etymology. According to the wishes of Bonaventure Sonke, the collector of the type specimen, we have named this new species after the botanist Tariq Stevart (b. 1974, Coordinator, West and Central Africa Program, Missouri Botanical Garden, Africa & Madagascar Department), in recognition of his helpful collaboration to our research on Drypetes and especially of his constant support to the second and third authors.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
5D78EE1D68B75DC7B65F6C2B91DFF08A.taxon	materials_examined	Additional material examined. CAMEROON - Centre • Goura; [4 ° 33 ' N, 11 ° 24 ' E]; Nov. 1938; fr.; H. Jacques-Felix 2383; P [P 04765168, P 04765171, P 04765172, P 04765173] • Goura, entre Bafia et Ntui; [4 ° 33 ' N, 11 ° 24 ' E]; fl. ♂; B. Mpom 520; P [P 04707491], YA [YA 0029435] • Mindou; 5 ° 10 ' N, 11 ° 35 ' E; 20 Jan. 2020; fr.; B. Sonke & M. Simo 6398; YA • Mpem et Djim, Malabo, plot Mpem 001, tag 245; 5 ° 01 ' N, 11 ° 39 ' E; 14 Feb. 2019; fr.; IRD plot 2354 (B. Sonke & al.); BRLU, YA • Parc National du Mpem et Djim; 5 ° 08 ' N, 11 ° 32 ' E; 12 Dec. 2022; fr.; F. Nzoyeuem & H. Leblanc 158; BR, BRLU, MA, MO, YA • Parc National du Mpem et Djim, Kounoungou; 5 ° 02 ' N, 11 ° 37 ' E; veg.; 24 Mar. 2023; B. Sonke & al. 7267; BR, BRLU, MO, YA • Parc National du Mpem et Djim, Kounoungou; 5 ° 01 ' N, 11 ° 36 ' E; fr.; 27 Mar. 2023; B. Sonke & al. 7303; BR, BRLU, MO, P, YA • Vers Biakoa a 117 km de Ntui; [4 ° 33 ' N, 11 ° 28 ' E]; 18 Nov. 1969; fr.; B. Mpom 535; P [P 04707489, P 04707490], YA [YA 0029434] • Versant septentrional des monts Meiki (983 m) au Sud de Ndo (25 km NNE d'Esse); [4 ° 19 ' N, 11 ° 58 ' E]; 9 Nov. 1969; fl. ♀, fr.; R. Letouzey 9543; BR [BR 0000015776754], COI, P [P 04707446], YA, WAG [WAG. 1564736]. - East • 60 km south of Yokadouma, 30 km after Ngato, 15 km after river, ALPICAM ' base de vie', then on forestry road starting 4 km before Masea village; 3 ° 09 ' N, 14 ° 44 ' E; 4 Mar. 2019; fr.; T. L. P. Couvreur & al. 1194; YA. - Littoral • Bilangue; 4 ° 10 ' N, 11 ° 00 ' E; 29 Apr. 2022; fr.; B. Sonke & al. 6975; BR, BRLU [BRLU 0032163], K, MO, P, WAG, YA • Bindamongo, km 106.5 sur route Yaounde-Bafia, a 15 km WNW de Ntui; [4 ° 31 ' N, 11 ° 30 ' E]; 18 Nov. 1969; fl. ♂; R. Letouzey 9568; BR [BR 0000015776730], P [P 04707231], YA [YA 0029469] • Bindamongo, km 106,5 sur route Yaounde-Bafia, a 15 km WNW de Ntui; [4 ° 31 ' N, 11 ° 30 ' E]; 18 Nov. 1969; fr.; R. Letouzey 9568 bis, BR [BR 0000015776747], P [P 04707227], YA [YA 0029462] • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 18 Oct. 2022; fl. ♂; B. Sonke 7097; BR, BRLU, M, MA [MA 959273], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 18 Oct. 2022; fl. ♀; B. Sonke 7096; BR, BRLU, M, MA [MA 959272], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 01 ' E; 18 Oct. 2022; fl. ♀; B. Sonke 7095; BR, BRLU, M, MA [MA 959277], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 18 Oct. 2022; fr.; B. Sonke 7094; BR, BRLU, E, MA [MA 959271], P n. v., YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 13 Oct. 2022; fr.; B. Sonke & M. Libalah 7074; BR, BRLU, MA [MA 959276], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 13 Oct. 2022; fr.; B. Sonke & M. Libalah 7072; BR, BRLU, MA [MA 959275], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 13 Oct. 2022; fr.; B. Sonke & M. Libalah 7071; BR, BRLU, M, MA [MA 959274], MO, P, YA • Binoum; 4 ° 12 ' N, 11 ° 02 ' E; 22 Dec. 2022, fr.; B. Sonke & M. Simo 7126; BR, BRLU, E, K, MA [MA 959270], MO, P, YA • Binoum, sur une coline; 4 ° 12 ' N, 11 ° 02 ' E; 15 Oct. 2022; fl. ♂; F. Nzoyeuem & al. 149; BRLU, YA • Kikot, en Aval du pont sur la Sanaga; 4 ° 09 ' N, 11 ° 00 ' E; 10 Nov. 2022; fl. ♀; TRCam plot 618 (F. Nzoyeuem & al.); BRLU, YA. - West • Pres Mankare, 45 km SE Foumban; [5 ° 24 ' N, 11 ° 10 ' E]; 23 Oct. 1974; fl. ♀, fr.; R. Letouzey 12989; BRLU [BRLU 0000164], MO [MO- 5594595], P [P 04707473], YA [YA 0029404]. NIGERIA - Adamawa • Gangumi, on the second mile line from the base line; [7 ° 56 ' N, 11 ° 11 ' E]; 11 Dec. 1954; fr.; FHI 28890 (M. G. Latilo & B. O. Daramola); FHO [FHO 00018584], K [2 sheets].	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	description	Figs 1, 2	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	materials_examined	Type. GABON • [Estuaire] Environs de Libreville, Sibang, [0 ° 25 ' N, 9 ° 30 ' E]; 23 Oct. 1902; fl. ♀, fr.; T. J. Klaine 2382; lectotype (designated here): P [P 04777265]; isolectotypes: BM, BR [BR 0000006238162, BR 0000006238490, BR 0000006238827, BR 0000006239152], K [K 000406384, K 000406385], P [P 04777262, P 04777263, P 04777264, P 04777266]; syntypes: GABON • [Estuaire] Environs de Libreville; 19 Feb. 1902; fl. ♀, fr.; T. J. Klaine 2482; BM, K, P [P 04777258, P 04777259, P 04777260, P 04777261] • Environs de Libreville; 26 Dec. 1901; fl. ♀, fr.; T. J. Klaine 2589; P [P 04777257, P 04777267]).	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	description	Description. Tree up to 25 m tall, with plagiotropic branches; trunk up to 16 cm in diameter, fluted or slightly fluted at the base; bark greenish-brown, smooth, rough and flaky when old; young branchlets subterete, slightly sulcate, shortly pubescent, trichomes 0.1 - 0.3 (- 0.6) mm; terminal buds scaly, scales 2.5 - 2.6 x 0.7 - 1 mm, ovate, firm, slightly keeled, shortly pubescent outside with trichomes 0.1 - 0.3 mm, glabrous inside, ciliate, cilia 0.1 - 0.3 mm. Leaves: stipules (3.7 -) 5.9 - 8 (- 12.5) x 0.2 - 0.5 mm, linear, laciniate, often with a single lacinia of 1.5 - 4.1 mm near the base, sometimes with 1 - 2 pairs of lateral laciniae, shortly pubescent outside, glabrous inside, ciliate, trichomes 0.2 - 0.5 mm, cilia 0.2 - 0.4 mm, subpersistent; petiole (4.5 -) 5.2 - 6.7 (- 9) mm long, (1.4 -) 1.7 - 2 mm in diameter, smooth or hardly wrinkled when dried, densely and shortly pubescent, glabrescent when old, trichomes 0.1 - 0.2 (- 0.3) mm; blade (9.3 -) 11 - 16.6 (- 17.7) x (3.4 -) 5.4 - 7.6 (- 8.8) cm, elliptic to broadly elliptic, papery to subcoriaceous, glossy and dark-medium green above, dull pale green beneath, base slightly obtuse to obtuse, markedly asymmetrical, rounded on one side, less often oblique, basal sides meeting the petiole up to 0.4 mm apart along the petiole length, margin crenulate-serrulate for most of its length, teeth to 1.3 (- 2.4) mm, acute, frequently blunt, flat to slightly recurved near the blade base, shortly acuminate, apex 10 - 20 mm, abaxial surface between the nerves sparingly pubescent, trichomes 0.1 - 0.4 mm; midrib smooth, slightly longitudinally wrinkled when dry, shortly and densely pubescent, trichomes 0.1 - 0.5 mm; first order lateral veins 6 - 8 pairs, ascending, more or less regularly spaced, slightly depressed above, prominent beneath, curved and anastomosing well within the margin, oriented at (49 -) 63 - 74 (- 79) ° to the midrib, minutely pubescent, trichomes 0.1 - 0.2 mm; second order venation loosely reticulate, hardly or not raised above, slightly so beneath. Male inflorescence borne on more or less horizontal cushion-like excrescences of the trunk (category I), in many-flowered clusters, often in groups of 30 or more flowers; bracts 0.6 - 0.9 x 0.7 - 0.8 mm, ovate to suborbicular, minutely pubescent outside, glabrous inside, minutely ciliate, trichomes and cilia 0.1 - 0.2 mm, barely distinguishable with a x 10 handlens. Male flowers long pedicellate, pinkish; pedicel 22 - 35 (- 40) mm long, 0.3 - 0.7 mm in diameter, slender, very sparingly and shortly pubescent, trichomes 0.1 - 0.2 mm; sepals (4 -) 5, 4.9 - 5.3 x 4.1 - 5.5 mm, widely ovate to suborbicular, cucullate, imbricate, the inner often smaller than the outer, glabrous outside and inside, margin margin minutely ciliate for most of its length, cilia 0.1 - 0.2 mm; stamens 14 - 16, one-whorled, surrounding the disc, more or less enveloped by its marginal lobes, filaments 4.4 - 4.5 (- 9.8) mm, white, anthers 1.2 - 1.5 (- 2) mm long, 0.9 - 1 (- 1.3) mm in diameter, ellipsoid, cream-coloured, subbasifixed to dorsifixed, introrse, pale yellow, glabrous; disc (0.3 -) 0.8 - 0.9 mm high, (2.7 -) 4.3 - 4.7 mm in diameter, convex, thick, very rugose, cream-coloured, densely and shortly pubescent, trichomes 0.1 - 0.3 mm, pistillode absent. Female inflorescence borne on cushion-like excrescences of the trunk (category I), in clusters of 10 - 35 flowers; bracts (0.6 -) 0.9 - 1.1 x 0.7 - 0.9 mm, otherwise similar to the male ones. Female flowers pedicellate; pedicel (6.3 -) 7.9 - 13.2 (- 20) mm long, 0.9 - 1.4 mm in diameter, robust, sparsely and minutely pubescent, glabrescent, trichomes 0.1 - 0.2 mm; sepals 5, 5 - 5.7 x (4 -) 6 - 6.3 mm, otherwise similar to the male ones; disc 0.9 - 1 mm high, (2.5 -) 4.2 - 4.4 mm in diameter, concave, cupulate, fleshy, shortly pubescent, more densely towards the margin, trichomes 0.2 - 0.3 mm; style 1, to 0.5 mm, unbranched; stigmas 3, obdeltoid, stigmatic surface 1.9 - 2.5 mm long, 1.3 - 3 (- 3.4) mm wide; ovary 2.4 - 3.4 mm long, 4.1 - 4.7 mm in diameter, subglobose, 3 (- 4) - celled, densely and shortly pubescent, trichomes 0.2 - 0.5 mm. Fruits (27 -) 29.5 - 34 (- 38.1) mm long, (15 -) 23 - 29 mm in diameter, ellipsoid, sepals, style, and stigmas caducous, exocarp uneven, yellowish brown turning yellowish orange when ripe, shortly pubescent, trichomes 0.1 - 0.3 mm, mesocarp 2 - 5 mm thick when dried, hard, endocarp 1 - 2 mm thick, bony, 1 - 3 - seeded; seeds 17.1 - 21 mm long, 8 - 9 mm in diameter; fruiting pedicel 21 - 33 mm long, 1.2 - 2.7 mm in diameter, sparsely and shortly pubescent, trichomes 0.1 - 0.3 mm.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	distribution	Distribution and chorology. Endemic to Gabon, mostly distributed in the northwest: Estuaire, Moyen-Ogooue, Ngounie, and Woleu-Ntem (Fig. 2). Lower Guinea subcentre of endemism (White 1979).	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	etymology	Etymology. The specific epithet comes from the Latin adjective verrucosus, which means warty, rough, or rugged. Hutchinson (1912) did not specify the reason for the epithet of this species name in the protologue, though it seems reasonable to assume that he referred to the rough, uneven surface of the fruit.	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
84FC3075E3F153D89D2BCC4550747D4C.taxon	materials_examined	Additional material examined. GABON - Estuaire • Concession Sud Estuaire, riviere Remboue; 0 ° 07 ' S, 9 ° 52 ' E; 19 Jun. 2021; fl. ♂; J. Klein & al. 564; BR n. v., BRLU, P n. v. • Mondah-Liby; [0 ° 46 ' N, 9 ° 38 ' E]; 30 Sep. 1952; veg.; F. Bernard 889; LBV [LBV 0030789] image, NHN [NHN 4557230] n. v. • National Park Akanda, Ngounzi debarcadere; 0 ° 34 ' N, 9 ° 41 ' E; 11 Dec. 2012; fr.; E. L. A. N. Simons & al. 1001; BR n. v., LBV n. v., MO [MO- 6561895] • Ndombo oil-concession area of CONOCO, ca 4 km SW of No Ayong; 0 ° 37 ' N, 9 ° 39 ' E; 22 Feb. 1991; fr.; J. M. Reitsma 3683; LBV [LBV 0030851] image • Remboue; [0 ° 02 ' N, 9 ° 50 ' E]; 30 Jul. 1951; veg.; Sabire & S. Etasse 340; LBV image. - Moyen-Ogooue • Au Nord-est du lac Azingo; 0 ° 19 ' S, 10 ° 03 ' E; 4 Jun. 2014; veg.; A. Boupoya & al. 944; BR [BR 0000016171442], BRLU n. v. • Au sud de Lambarene, entre 5 et 10 km depuis la ville, entre l'Ogooue et la route de Fougamou; 0 ° 55 ' S, 10 ° 21 ' E; 15 Apr. 2015; fr.; E. Bidault & al. 1940; BR [BR 0000016172166], BRLU [BRLU 0002958] n. v., P [P 00854948] • Lambarene, along the Ogooue stream; 0 ° 53 ' S, 10 ° 09 ' E; 15 Oct. 2012; fl. ♂; B. Sonke & al. 6041; BR [BR 0000024385589], BRLU, MO [MO- 6729952] • Route menant au lac Azingo, a 35 km au nord-ouest de Lambarene; 0 ° 27 ' S, 10 ° 05 ' E; 26 Oct. 2014; fl. ♂; E. Bidault & al. 1861; MO [MO- 6681694] • Zone de Mabounie, a 45 km au sud-ouest de Lambarene, rive nord de la riviere Ngounie; 0 ° 45 ' S, 10 ° 36 ' E; 4 Feb. 2013; fr.; E. Bidault & al. 1054; BRLU [BRLU 0000151], LBV image, MO [MO- 6729953], NHN [NHN 1380674] n. v., P [P 00854579] • Zone de Mabounie, a environ 45 km au sud-est de Lambarene, rive ouest de la Ngounie, plot Mabou 009; 0 ° 49 ' S, 10 ° 29 ' E; 14 Oct. 2012; veg.; IRD plot 332 (N. Barbier & al.); BRLU • Zone de Mabounie, a environ 45 km au sud-est de Lambarene, rive ouest de la Ngounie, plot Mabou 009; 0 ° 49 ' S, 10 ° 29 ' E; 14 Oct. 2012; veg.; IRD plot 331 (N. Barbier & al.); BRLU. - Ngounie • Foret au Nord de Lambarenekili, a environ 5 km au nord de la riviere Niambo-Kamba; [1 ° 26 ' S, 10 ° 17 ' E]; 15 Aug. 2008; veg.; G. Dauby & al. 1375; BRLU • Sindara; 1 ° 02 ' S, 10 ° 39 ' E; 16 Jul. 1979; fl. ♂; G. M. P. C. Le Testu 2291; B [B 10 0366532], BM, BR [BR 0000015019523, BR 0000015019547], K, LISC, P [P 04777254, P 04777255, P 04777256]. - Woleu-Ntem • [La] Lara; [0 ° 21 ' N, 11 ° 28 ' E]; 27 Oct. 1933; fl. ♂; G. M. P. C. Le Testu 9349; K, BM, BR [BR 0000015019554], MO [MO- 5594599], P [P 04707826], WAG [WAG. 1564868] • [La] Lara; [0 ° 21 ' N, 11 ° 28 ' E]; 27 Oct. 1933; fl. ♀; G. M. P. C. Le Testu 9347; K, BM, BR [BR 0000015777539], P [P 04707828, P 04707829, P 04707831], WAG [WAG. 1564872, WAG. 1564873] • Region entre Ogooue et Cameroon; 19 May 1936; veg.; G. M. P. C. Le Testu s. n.; P [P 04707827, P 04707830].	en	Quintanar, Alejandro, Sonke, Bonaventure, Simo-Droissart, Murielle, Barbera, Patricia, Libalah, Moses, Harris, David J. (2023): A matter of warts: a taxonomic treatment for Drypetes verrucosa (Putranjivaceae, Malpighiales) and a new cauliflorous species from Cameroon and Nigeria, D. stevartii. Plant Ecology and Evolution 156 (2): 160-173, DOI: http://dx.doi.org/10.5091/plecevo.102004, URL: http://dx.doi.org/10.5091/plecevo.102004
