taxonID	type	description	language	source
6C38C875FFB55B54FDE9FCFEE707FC35.taxon	description	Smetana (2017 a) defined this group by the position of the large lateral puncture of the pronotum, which is not touching the marginal bead, and the lack of micropunctures on the disc of the head and pronotum.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	description	Fig. 1 A	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	diagnosis	Diagnosis Indoquedius malaisei can be distinguished from other members of the Sikkimensis group in the mainland Oriental region by the combination of: antennomeres 6 – 11 milky yellow; elytra reddish and paler than head and pronotum.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	materials_examined	Material examined Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 2000 m; 12 – 17 / 6.34; Malaise [printed label] / HOLOTYPUS [red, handwritten label] / TYPUS Quedius malaisei, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) Malaisei n. sp., det. Scheerpeltz, 1944 [white printed label] / 6625 E 91 [printed blue label] / Malaisei Scheerpeltz [large, printed folded label] / NHRS - JLKB 000021047 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	description	Redescription The species was recently redescribed and diagnosed by Smetana (2017 a) under the name I. nonparallelus. For additional details, see Zhao & Zhou (2010). A dorsal habitus is provided in Fig. 1 A.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	distribution	Distribution Myanmar (Kachin), China (Yunnan).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB55B55FDFBFC15E18FFD8B.taxon	discussion	Remarks Although Scheerpeltz stated the holotype of I. malaisei was a female, it is in fact a male whose aedeagus corresponds perfectly to that of I. nonparallelus syn. nov., which becomes a junior synonym. The type locality of I. nonparallelus (“ Gonggashan County, Yunnan ” = Gongshan Derung and Nu Autonomous County) is north of Kambaiti Pass and near the Chinese border with Myanmar. The holotype of I. nonparallelus was not examined but it was examined by Smetana (2017 a) and illustrated in the original description with detail sufficient to make the above synonymy.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B55FE27FDC9E6B8FA0F.taxon	description	Fig. 1 B	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B55FE27FDC9E6B8FA0F.taxon	diagnosis	Diagnosis Indoquedius micantiventris can be distinguished from other members of the Sikkimensis group by the combination of: antennomeres 1 – 3 dark, 4 – 11 testaceous and becoming paler toward apex (but not milky-yellow); legs entirely dark except for vaguely paler tarsi; elytra dark, non-metallic.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B55FE27FDC9E6B8FA0F.taxon	materials_examined	Material examined Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 7000 ft.; 1 / 5 1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius micantiventris, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) micantiventris nov. spec., det. Scheerpeltz, 1944 [white handwritten label] / 6641 E 91 [printed blue label] / micantiventris Scheerpeltz [large, printed folded label] / NHRS-JLKB 000021048 ”; NHRS. Paratype MYANMAR – Kachin State • ♀; “ [female symbol on card] / N. E. Burma, Kambaiti; 7000 ft.; 1 / 5 1934; R. Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll. Scheerpeltz [printed blue label] / ALLOTYPUS [red handwritten label] / TYPUS Quedius micantiventris, O. Scheerpeltz [dark red-pink label] / micantiventris [pink card] ”; NHW.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B53FDC9FA35E72BFD5F.taxon	materials_examined	Material examined Paratype NEPAL – Bagmati Province • ♀; “ Nepal (Prov. Bagmati), Malemchi; 2800 m; 17. IV. 81; Löbl and Smetana / Paratype Indoquedius baliyo A. Smetana 1986 [yellow printed label] / PARATYPE CNC No. 20724 [yellow printed label] / CNC 978889 [identifier] ”; CNC. Non-type material MYANMAR – Kachin State • 1 ♂; Three River Junction; 26 ° 23 ′ 12.6 ″ N, 98 ° 39 ′ 36.3 ″ E; 3 Oct. 2010; M. Langer, S. Naumann and S. Loffler leg.; under wood / stones; ASC. NEPAL • 1 ♂; Manaslu Mts, E slope of Ngadi Khola valley; 28 ° 22 ′ N, 84 ° 29 ′ E; 2000 – 2300 m; 14 – 16 May 2005; J. Schmidt leg.; CNC • 2 ♂♂; same collection data as for preceding; ASC.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B53FDC9FA35E72BFD5F.taxon	description	Redescription Apart from Scheerpeltz (1965), the species has been described and diagnosed by Smetana (1988, 2017 a) under the name I. baliyo. A dorsal habitus of the holotype of I. micantiventris is given in Fig. 1 B.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B53FDC9FA35E72BFD5F.taxon	distribution	Distribution Nepal, India (West Bengal), Myanmar (Kachin), and China (Tibet / Xizang, Yunnan, Sichuan) (Smetana 2017 a).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB45B53FDC9FA35E72BFD5F.taxon	discussion	Remarks The type series of Indoquedius micantiventris includes one male and one female. The male was dissected to reveal an aedeagus identical to the common, widespread species I. baliyo. Therefore I. baliyo syn. nov., becomes a junior synonym of I. micantiventris. Both specimens of the type series of I. micantiventris have slightly darkened antennomeres 4 – 11 (especially 4 – 5) compared to all other specimens of I. baliyo seen. However, no other differences were observed and the darkening is likely an artefact of preservation. The recently collected, non-type specimen examined from Myanmar (~ 120 km northeast of Kambaiti) has entirely normal coloration.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B53FF67FD08E06FFCF9.taxon	description	Smetana (2017 a) defined this group by the position of the large lateral puncture of the pronotum, which is touching the marginal bead.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	description	Figs 1 C, 4 A – B, 7 A	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	diagnosis	Diagnosis This species is unique among all Indoquedius described from the mainland Oriental region for the combination of: sparse, irregular elytral punctation (Fig. 4 A); short elytra with strongly transverse scutellum (Fig. 1 C); and extremely finely and sparsely punctate abdominal tergites III – VI, IV – VI with the apical half to two-thirds impunctate (Fig. 4 B). Widespread and possibly sympatric I. filicornis (Eppelsheim) is somewhat similar in the short elytra but the elytral punctures are coarser and denser, the scutellum is densely and coarsely punctate and the abdominal punctures are much coarser. Although it has been reported that I. filicornis always lacks a palisade seam on abdominal tergite VII (Smetana 2017 a), it was present on one dark female specimen (ASC) from Yunnan, China, determined by A. Smetana and otherwise consistent with the species.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♀; “ N. E. Burma, Kambaiti; 7000 ft.; 3 – 7 / 5.1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius dispersepunctatus, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) dispersepunctatus nov. spec., det. Scheerpeltz [printed label] / 6476 E 91 [blue printed label] / NHRS-JLKB 000073667 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	description	Redescription Body entirely dark, maxillary and labial palpi testaceous, antennae testaceous without any obviously darkened segments, legs yellowish-brown with dark brown middle and hind femora. Head distinctly transverse; eyes very large and convex; tempora almost non-existent, with three parocular punctures along inner margin of eye; posterior frontal puncture slightly anteriad of hind margin of eye, puncture touching border; temporal puncture absent; nuchal ridge indistinct at middle of neck; antennomere 3 about 1.3 × as long as 2, antennomeres elongate and symmetrical, becoming shorter toward apical antennomere, antennomere 10 about 1.3 × as long as wide. Pronotum distinctly transverse, slightly convergent anteriad, with broadly rounded posterior margin, dorsal rows each with three punctures, sublateral rows absent, large lateral puncture touching marginal bead. Scutellum transverse, with seven moderately coarse punctures, most of scutellum surface impunctate; elytra short, slightly broader than pronotum, suture markedly shorter than pronotum at midline, at sides, slightly longer than pronotum at midline, punctures moderately coarse, sparsely and irregularly distributed, separated by at least one puncture diameter but most by several times that amount; wings fully developed; abdominal tergite VII with palisade fringe; abdominal punctures extremely fine and sparse; tergite III with only a short transverse row of basal punctures; tergites VI – VI with posterior ½ to ⅔ impunctate; segments with extremely fine and dense microsculpture of transverse waves. Male Unknown. Female Tergite X strongly narrowed to deeply and narrowly emarginate apex (Fig. 7 A).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	distribution	Distribution Known only from the type locality in Myanmar (Kachin).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB25B50FE3DFC5BE176FB00.taxon	discussion	Remarks The type material includes only a single female holotype. The holotype was dissected and female tergite X is illustrated in Fig. 7 A. Based on external morphology alone, I. dispersepunctatus does not have any synonyms and can be easily recognized.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB15B51FDDBFB31E0D7FD59.taxon	description	Figs 1 D – E, 7 B	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB15B51FDDBFB31E0D7FD59.taxon	diagnosis	Diagnosis Among members of the Bipunctatus group, Indoquedius parallelicollis can be distinguished by the darkened femora and tibiae (at least outer face) on all legs, the overall pale antennae without any markedly darkened segments, and the moderately coarse and sparse punctation of the elytra. Sympatric Indoquedius bicornutus Zhao & Zhou, 2010 is very similar but can be distinguished by the pale tibiae, coarser and denser elytral punctation. Smetana (2017 a) had stated that the elytra of I. parallelicollis (as I. bicoloris) were either entirely red or entirely dark but among the paratype specimens examined was one specimen with dark elytra and a faint basolateral red spot, similar to what can occur in I. bicornutus. The paratype of I. parallelicollis represents another transitional color pattern with less obvious pale areas of the elytra basally and posteriorly (Fig. 1 D). The abdominal coloration is also slightly more variable than previously stated (Smetana 2017 a), with the apices of tergites narrowly paler in some specimens, while they are broadly paler in I. bicornutus.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB15B51FDDBFB31E0D7FD59.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 2000 m; 4 / 6.1934; Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius parallelicollis, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) parallelicollis nov. spec., det. Scheerpeltz 1944 [printed label] / 6664 E 91 [printed blue label] / parallelicollis Scheerpeltz [large, printed, folded label] / NHRS-JLKB 000021050 ”; NHRS. Paratype MYANMAR – Kachin State • 1 ♀; “ N. E. Burma, Kambaiti; 2000 m; 4 / 6.1934; Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll. Scheerpeltz [printed blue label] / ALLOTYPUS [red handwritten label] / TYPUS Quedius parallelicollis, O. Scheerpeltz [dark red-pink label] / parallelicollis [pink card] ”; NHW.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B51FE53FD00E1B9FB87.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♀; “ N. E. Burma, Kambaiti; 7000 ft.; 28 / 5.1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius sanguinipennis, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) sanguinipennis nov. spec., det. Scheerpeltz [printed label] / 6706 E 91 [printed blue label] / NHRS-JLKB 000073670 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B5EFDD9FBC2E064FD24.taxon	materials_examined	Type material Paratypes CHINA – Yunnan Province • 1 ♂; “ Indoquedius bicoloris A. Smetana 2013 [printed yellow label] ”; Pu’er Pref., Ailao Shan, 37 km NW Jingdong; 24 ° 45 ′ 12 ″ N, 100 ° 41 ′ 24.5 ″ E; alt. 2300 m; 13. Oct. 2009; D. Wrase leg.; devastated forest remnant, litter / moss / grass roots sifted; ASC • 1 ♂; “ Indoquedius bicoloris A. Smetana 2013 [printed yellow label] ”; Dai Bai Nat. Aut. Pref., Diancang Shan, 3 km W Dali old town, pine forest at Cloud Road, right upper chair-lift station; 25 ° 41.1 ′ N, 100 ° 6.8 ′ E; alt. 2650 – 2750 m; 1 Sep. 2003; D. Wrase leg.; needleleaf litter; ASC • 1 ♀; “ Indoquedius bicoloris A. Smetana 2013 [printed yellow label] ”; E slope N Gaoligongshan; 27 ° 45 ′ 27.1 ″ N, 98 ° 35 ′ 34.5 ″ E; 2 Jun. 2010; alt. 2600 m; V. Grebennikov; sifting; ASC.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B5EFDD9FBC2E064FD24.taxon	description	Redescription The species has been recently redescribed and diagnosed in detail by Smetana (2017 a) under the name I. bicoloris. This description is supplemented here only by the comments above and Figs 1 D – E, 7 B.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B5EFDD9FBC2E064FD24.taxon	distribution	Distribution This widespread, potentially common species is known from medium elevations in northern Myanmar, and Yunnan and Sichuan, China (Smetana 2017 a, 2017 b).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B5EFDD9FBC2E064FD24.taxon	biology_ecology	Bionomics Indoquedius parallelicollis appears to be tolerant to human disturbance and has been collected in a haystack and plant debris in a ruderal habitat (Smetana 2014, 2017 a; Yan et al. 2017). Other specimens have been collected in a mixed forest by sifting litter, twigs and roots (Smetana 2015)	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB05B5EFDD9FBC2E064FD24.taxon	discussion	Remarks The type series of I. parallelicollis includes one male and one female (Fig. 1 D). The male was dissected to reveal an aedeagus identical to the color polymorphic species I. bicoloris. The current distribution of I. bicoloris includes the narrow part of Yunnan, China, west of the Salween River (Smetana 2014) and not far from Kambaiti Pass in Myanmar. Therefore I. bicoloris syn. nov., becomes a junior synonym of I. parallelicollis. The female holotype of I. sanguinipennis (Fig. 1 E) corresponds perfectly (coloration of legs and abdomen, elytral punctation) with a female paratype (ASC) of the red elytra morph of I. bicoloris. Female tergite X was also consistent with that of I. bicoloris and the paratype of I. parallelicollis, including the deep emargination (Fig. 7 B). Therefore, along with I. bicoloris, I. sanguinipennis syn. nov., becomes a junior synonym of I. paralellicollis.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	description	Figs 1 F, 4 C – D, 7 C	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	diagnosis	Diagnosis Among members of the Bipunctatus group, I. recticollis can be easily recognized by the combination of: impunctate middle areas and posterior half to two-thirds of abdominal tergites III – VI (Fig. 4 D); the sparse but regularly spaced elytral punctures (Fig. 4 C); and the darkened middle and hind femora (Fig. 1 F). Indoquedius recticollis is somewhat similar to I. dispersepunctatus in the sparsely punctate abdomen and darkened middle and hind femora but can easily be distinguished by the darkened first antennomere, much finer and regularly spaced elytral punctures, the longer elytra, the coarser abdominal punctures and the pale apices of the abdominal tergites. These two species may be related as they share a similar shape of female tergite X (Fig. 7 A, C).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♀; “ N. E. Burma, Kambaiti; 2000 m; 4 / 6.1934; Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius recticollis, O. Scheerpeltz [dark red-pink label] / Quedius (Indoquedius) recticollis nov. spec., det. Scheerpeltz [printed label] / 6692 E 91 [blue printed label] / NHRS-JLKB 000073669 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	description	Redescription Similar to Indoquedius dispersepunctatus and differing only in the following: antennomere 1 darkened on dorsal surface; abdominal tergites broadly paler apically; head with two parocular punctures; temples slightly smaller; antennomere 3 slightly longer than 2, about 1.4 ×; antennomere 10 distinctly broader at apex, shape more obviously triangular; pronotum slightly transverse, more strongly convergent anteriad; scutellum longer, more typical of the genus, densely punctate with only outer margins impunctate; elytra longer at both suture and at sides, at sides distinctly longer than pronotum at midline; elytral punctures much finer and less strongly impressed, regularly distributed though sparse, separated by more than one puncture diameter; abdominal punctation coarser, tergite III with sparse punctures across disc, middle of tergites impunctate or nearly so on tergites III – VI; female tergite X slightly less strongly narrowed to apex, which is broader overall and more shallowly and broadly emarginate (Fig. 7 C).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	distribution	Distribution Known only from the type locality in Myanmar (Kachin).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBF5B5FFDC3FD1CE373FD8E.taxon	discussion	Remarks The type material contains only the female holotype, which was dissected. Based on external morphology, this species does not appear to have any synonyms and can be recognized based on external morphology alone.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5FFF67FD86E707FC5B.taxon	discussion	Members of this large, monophyletic group can be recognized by the combination of: genal puncture present; basal puncture doubled; elytra with evenly distributed punctures or with at least asetose punctures on disc among serial rows of setose punctures. This lineage currently consists of Anthosaurus Smetana, 2015, Pseudorientis Watanabe, 1970, Korgella Özdikmen, 2005, Quedius (Velleius) (Fabricius, 1787) and the large polyphyletic subgenus Quedius (Microsaurus) Stephens, 1829, inside which all the aforementioned taxa are nested. To avoid treating all members of the Microsaurus lineage as a single, morphologically heterogeneous genus, and to avoid a major priority issue (Velleius Leach, 1819 has priority over Microsaurus Dejean, 1833), the major subclades of this lineage should be ranked as genera, pending comprehensive morphological study (Brunke et al. 2021). One of these taxa is described below.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	description	urn: lsid: zoobank. org: act: 6947 E 75 F- 7462 - 4642 - A 8 D 4 - 2 AA 3 EA 25 E 22 C Figs 2 A – B, 4 E – F, 5 A – B, 6 A – G, 7 D	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	type_taxon	Type species Quedius ruficeps Scheerpeltz, 1965.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	diagnosis	Diagnosis Within the Microsaurus lineage, Malaisdius gen. nov. can be recognized by the following: elytral disc distinctive, uneven and with deeply impressed, foveate and setose punctures (Fig. 4 F); abdominal tergites III – VI with distinct, coarsely punctate basal impressions, disc otherwise impunctate or nearly so, punctures elongate (Fig. 5 A – B). In the latter two features, species of Malaisdius gen. nov. superficially resemble some members of cyrtoquediine genus Bolitogyrus Chevrolat, 1842 though asetose punctures are always present in Bolitogyrus and the forebody is entirely without microsculpture.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	etymology	Etymology This genus is dedicated to the Swedish explorer and entomologist René Malaise, who collected a diversity of rove beetles at Kambaiti Pass, Kachin, Myanmar. Malaise is also famous for the invention of the insect trap that bears his name, which has revolutionized our knowledge of insect biodiversity and was first used on a significant scale in Kambaiti (Sjöberg 2014).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	description	Description Medium-sized, glossy beetles, with bright coloration. With the character states of Quediini (sensu Brunke et al. 2021) and the following: antennomere 3 longer than 2, without dense setation or tomentose pubescence; outer antennomeres (6 – 10 or 8 – 10) weakly transverse and slightly asymmetrical; antennae inserted far from eyes, about 2.0 × the width of antennal sclerite; head with eyes large, strongly convex, protruding from lateral head margin, temples distinct, about half the length of eyes; eyes subparallel, inner margin forming acute angle with supra-antennal ridge; basal puncture doubled, interocular punctures absent, genal puncture present but extremely small, rudimentary; frons not well developed anterolaterad of antennal insertions; labrum notched medially creating two lobes; apical maxillary and labial palpomeres fusiform and glabrous; infraorbital ridge complete to base of mandibles; gular sutures convergent towards neck, narrowly spaced posteriad; mandibles with small dorsolateral groove; right mandible with two teeth, distal tooth on lower plane compared to proximal; pronotum shield-shaped, slightly explanate; marginal punctures touching marginal bead, dorsal rows with three punctures, sublateral rows with only one puncture, therefore not extended posteriad of large lateral puncture (two female specimens with puncture-like elements (without sockets) giving superficial appearance of 2 – 3 punctures in the sublateral row), with single large lateral puncture; hypomeron strongly inflexed, hypomeron not visible; basisternum with pair of small, pale macrosetae, with well-developed longitudinal carina; scutellum impunctate; elytron with subbasal ridge complete, forming scutellar collar; row of humeral spines present but short (2 – 4 spines); disc of elytra without microsculpture between the punctures, surface uneven, punctures irregularly and sparsely distributed, or in very loose rows, punctures bearing setae and situated in broad depressions; foretibia without lateral spines but with apical spur; metatibia with only two slender spines on lateral face; metatarsomeres with disc setose; prototergal glands likely absent, with only vague, shallow impression lacking marginal setae; abdominal tergites III – VI with distinct basal depressions bearing elongate punctures, remainder of disc impunctate or nearly so; abdominal sternite III transverse basal line acutely and sharply produced posteriad at middle; males without modifications on sternite VII; aedeagus asymmetrical in known species, with well-developed paramere, paramere without peg setae; aedeagus without large copulatory sclerites in internal sac.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	distribution	Distribution Malaisdius gen. nov. is known from the Himalaya of Nepal and the mountains of northern Myanmar.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	biology_ecology	Bionomics Nothing is currently known about the natural history of Malaisdius gen. nov.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBE5B5CFDAFFBE7E707F883.taxon	discussion	Remarks According to phylogenomic analyses (Brunke et al. 2021), the sister group of Malaisdius gen. nov. is the more generalized Masasatoi group (see below), currently included within polyphyletic Quedius (Microsaurus) but will be treated as a separate genus in a future study.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	description	Figs 2 A, 5 A, 6 A – E, 7 D	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	diagnosis	Diagnosis Within the genus, M. ruficeps comb. nov. is easily distinguished from M. smetanai gen. et sp. nov. by the strikingly bicolored body (Fig. 2 A).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 7000 ft.; 12 / 5 193; R. Malaise [printed label] / Holotypus [handwritten red label] / TYPUS Quedius ruficeps, O. Scheerpeltz [typed dark red-pink label] / Quedius (Microsaurus) ruficeps nov. spec., det. Scheerpeltz, 1974 [white label] / 6693 E 91 [typed blue label] / NHRS-JLKB 000021051 [identifier] ”; NHRS. Paratypes MYANMAR – Kachin State • 1 ♀; “ [card with female symbol] / N. E. Burma, Kambaiti; 2000 m; 24 / 5.1934; Malaise [typed label] / Schwedisch Indian-Burma Expedition 1934 [typed label] / ex. coll. Scheerpeltz [typed blue label] / Allotypus [handwritten red label] / TYPUS Quedius ruficeps, O. Scheerpeltz [typed dark red-pink label] / ruficeps Schp [handwritten pink card]; NHW • 1 ♀; “ [card with female symbol] / N. E. Burma, Kambaiti; 7000 ft.; 28 / 5 1934; R. Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [typed label] / ex. coll. Scheerpeltz [typed blue label] / COTYPUS Quedius ruficeps O. Scheerpeltz [pink label] ”; NHW • 1 ♂; “ N. E. Burma, Kambaiti; 2000 m; 25 / 5.1934; Malaise [typed label] / COTYPUS Quedius ruficeps O. Scheerpeltz [pink label] / 6694 E 91 [typed blue label] / NHRS-JLKB 000021052 [identifier] / ruficeps Scheer [large folded label] ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	description	Redescription Head and pronotum pale orange-red, sharply contrasting with dark, vaguely bluish elytra and abdomen, scutellum very dark red, abdomen with iridescent reflection, antennomeres 1 – 4 pale, yellow, antennomeres 5 – 11 darker, brown to dark brown, becoming darker toward apex of antenna, legs entirely pale, yellow, palpi entirely pale, yellow. Body length (minus abdomen): 3.8 – 4.3 mm. Head slightly wider than long (1.12 – 1.15), with posterior angles absent; neck varying from moderately wide to narrow, creating an impression of a more elongate head; eyes large and clearly protruding from lateral head outline, tempora shorter than eyes (0.4 – 0.6); temporal puncture much closer to eye margin than hind margin of head; surface of head with extremely fine and superficial microsculpture of dense transverse waves. Antennae long, all segments longer than wide except 8 – 10, which are weakly transverse and slightly asymmetrical. Pronotum strongly convex, about as wide as long to distinctly wider than long (1.20), slightly explanate, punctures arranged as in generic description, microsculpture similar to that on head. Scutellum impunctate. Elytra at base slightly broader than pronotum, at middle and sides distinctly longer than pronotum at midline; disc without microsculpture, punctures sparsely and irregularly spaced, deeply impressed and foveate. Wings fully developed. Abdomen with tergite VII with palisade fringe; punctation as in generic description; tergites with extremely fine microsculpture of transverse waves. Male Sternite VIII with distinct but shallow emargination that is about 3.0 × as wide as long; sternite IX with long, asymmetrical basal part, most of disc setose, narrowing to asymmetrically rounded and entire apex; tergite X weakly narrowed to truncate and loosely crenulate apex, projections bearing setae; median lobe and paramere asymmetrical; median lobe in ventral view with apical third delimited by slight constriction, narrowed to rounded acute apex, with one lateral and one median fin-like tooth (Fig. 6 A – B); median lobe in lateral view with narrowed apical part distad of emargination, this part with a tooth corresponding to the lateral tooth in ventral view, median tooth clearly visible (Fig. 6 C); in the alternate lateral view, median lobe with apical part appearing broader and median tooth less visible (Fig. 6 D); paramere broad and subparallel over much of its length, narrowing into a finger-like lobe, with a small lateral bulge, apex with four pairs of small setae, peg setae absent (Fig. 6 E). Female Tergite X (Fig. 7 D) nearly oval in shape, narrowed to moderately wide and slightly acuminate apex, disc with distinct oval impression, apex with pair of very long setae.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	distribution	Distribution Known only from the type locality in Myanmar (Kachin).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBC5B5AFE32FEFEE0FDFC05.taxon	discussion	Remarks One of the female paratypes is slightly paler than the other individuals, the antennae appear narrower, with antennomeres 7 – 10 weakly transverse, and the elytra appear vaguely narrower. However, this is considered to be intraspecific variation.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	description	urn: lsid: zoobank. org: act: 2868 C 1 D 2 - F 3 E 2 - 42 CE-B 4 DF- 48 F 4 B 5 D 4705 F Figs 2 B, 4 E – F, 5 B, 6 F – G	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	diagnosis	Diagnosis Malaisdius smetanai gen. et sp. nov. is easily recognized by the overall pale coloration of the body (Fig 2 B). The only other species of the genus is strikingly bicolored. Additionally, the bicolored hind femur and the elytra with loose rows of punctures will distinguish this species from M. ruficeps comb. nov.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	etymology	Etymology This species is dedicated to the late Dr Aleš Smetana, who was working with me on an early morphological concept of Malaisdius gen. nov. (and who created the name) several years before his passing in 2021. Aleš spent his career uncovering the great diversity of Quediini and without this enormous contribution, a generic revision of the tribe would not be possible.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	materials_examined	Type material Holotype NEPAL – Gandaki Province • ♂; “ E-Nepal, Arun Valley, Deurali; ca. 2100 mNN; 27 ° 30 ′ N, 87 ° 16 ′ E; 10. V. 2014; J. Schmidt leg. [typed label] / collection NATURKUNDE MUSEUM ERFURT [typed yellow label] / AJB 0001337 [identifier] / Quediini phylogeny DNA Voucher Specimen, SeqDB: AKH- 07, Specimen AJB 0001337 [blue label] / Malaisdius smetanai Brunke, des Brunke 2023 [red label] ”; NME.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	description	Description Similar to M. ruficeps and differing only by the following: body (minus abdomen) 4.2 mm, paler overall, not sharply bicolored, head, centre of elytra, and abdominal tergites dark orange-brown to dark brown, pronotum, lateral parts of elytra and paratergites paler, brownish-orange; antennomeres 1 – 5 paler, orange-brown, 6 – 10 darker, brown; hind femora with dark apical band, hind tibia with very base darker; antennomeres 6 – 10 weakly transverse and slightly asymmetrical; pronotum less convex; elytral punctures arranged in loose rows, impressions even broader; abdominal tergites with punctures even more strongly impressed, slightly more numerous but still mostly restricted to basal impressions. Male Sternite VIII with emargination slightly wider; male sternite IX with basal part more strongly narrowed and hooked at the end, with distinct expansion opposite to this, apex slightly more rounded; male tergite X shorter, more strongly narrowed to apex, which is more rounded and without large setose crenulatons; median lobe in ventral view slightly broader, gradually narrowed to apical part, apex slightly broader, lateral tooth smaller and facing opposite direction (Fig. 6 F) [median tooth and entire lateral view not visible due to previous mounting of genitalia in Canada balsam]; paramere narrower than median lobe in ventral view, slightly convergent apicad over much of its length, finger-like lobe hooked and on opposite side of paramere, one side of apex crenulate, crenulations bearing pairs of setae (Fig. 6 G). Female Unknown.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBB5B5BFD9BFC3CE151FC35.taxon	distribution	Distribution Known only from the type locality in eastern Nepal.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFBA5B58FDE4FBD3E3ABFECF.taxon	discussion	Remarks This diverse subgenus is in great need of generic revision as it is rendered polyphyletic by several genera and subgenera of Quediini, including Malaisdius (see further comments above under “ Microsaurus lineage ”). Currently, it is not possible to give a meaningful diagnosis of Quedius (Microsaurus). The species groups listed below are redefined from Smetana (2017 a), likely monophyletic and correspond to clades in Brunke et al. (2021). They will be treated in much greater detail in a future publication.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	description	Figs 2 C, 5 C – E, 7 E	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	diagnosis	Diagnosis Within the Apicicornis group, Q. rutilipennis can be recognized by a combination of: the sublateral rows of the pronotum reaching posteriad of the large lateral puncture; and the sparsely punctate to impunctate disc of tergites III – IV (Fig. 5 E).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 7000 ft.; 9 / 6.1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius rutilipennis, O. Scheerpeltz [dark red-pink label] / Quedius (Microsaurus) rutilipennis nov. spec., det. Scheerpeltz 1944 [printed label] / 6703 E 91 [printed blue label] / NHRS-JLKB 000021053 ”; NHRS. Paratypes MYANMAR – Kachin State • 1 ♀; “ N. E. Burma, Kambaiti; 2000 m; 7 – 9 / 6.1934; Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll. Scheerpeltz [printed blue label] / ALLOTYPUS [red handwritten label] / TYPUS Quedius rutilipennis, O. Scheerpeltz [dark red-pink label] / rutilipennis [orange card] ”; NHW • 1 ♀; “ [female symbol on card] / N. E. Burma, Kambaiti; 7000 ft.; 9 / 6.1934; R. Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll. Scheerpeltz [printed blue label] / Cotypus Quedius rutilipennis O. Scheerpeltz [pink label] ”; NMW.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	description	Redescription This species was recently described (as Q. cornutus) by Cai et al. (2015) with sufficient detail. That description is supplemented here with a complete habitus illustration (Fig. 2 C) and some additional images of the antennae, forebody and abdomen (Fig. 5 C – E). The female was previously undescribed and so tergite X is illustrated in Fig. 7 E.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	distribution	Distribution The species is known from Myanmar (Kachin) and an adjacent area of Yunnan, China.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB95B58FE0CFE72E0A1F8D1.taxon	discussion	Remarks The type series includes the male holotype and two female paratypes. The male was dissected to reveal an aedeagus identical to Q. cornutus Cai, Yan-Peng, Zong-Yi Zhao & Hong-Zhang Zhou 2015. The type locality of Q. cornutus is located in the narrow part of Yunnan west of the Salween River (Cai et al. 2015) and not far from the type locality of Q. rutilipennis in Myanmar. Therefore Q. cornutus syn. nov., becomes a junior synonym of Q. rutilipennis.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B59FF67FEFEE731FB9E.taxon	discussion	Quedius masasatoi Smetana, 2007, from China (Fujian, Yunnan), Laos and Vietnam, was previously included in the Apicicornis group of Smetana (2017 a), likely out of convenience as it is rather different in habitus. In a similar way, Smetana (2007) placed Q. masatakai Smetana, 2007, from Laos and Vietnam (Smetana & Brunke 2021), in the Ripicola group even though it was rather different in habitus. Recently, these two species were shown to be sister groups within ‘ Clade E’ of the Microsaurus lineage, and together were sister to a species here described as Malaisdius smetanai gen. et sp. nov. (Brunke et al. 2021). Quedius masasatoi and Q. masatakai are here included in a newly delineated group, which also includes Q. piceolineatus Scheerpeltz, 1965 from the Malaise material collected in northern Myanmar. A new genus will need to be described for this group but additional morphological study is required in order to provide a more robust diagnosis. The Masasataoi group is provisionally diagnosed within the Microsaurus lineage by the following combination of characters: head with eyes large, clearly longer than temples but temples well-developed (Fig. 2 D), eyes protruding from lateral head margin, posterior frontal puncture situated at about hind margin of eye, without additional punctures between posterior frontal puncture and basal punctures; antennomeres 6 – 10 flattened as typical for Quediini, not inflated; pronotum slightly explanate, marginal punctures not removed from marginal bead, anterior angles with marginal bead partly hidden in dorsal view, dorsal row with second and third punctures rather close together (about one puncture diameter or less) and separated from the first by a long gap (Fig. 5 F), at least on one side; sublateral row not extended posteriad of large lateral puncture, without additional punctures between dorsal and sublateral rows (Fig. 5 F); scutellum impunctate but sometimes with micropunctures or scratch-like elements, elytra sparsely but evenly punctate, punctures setose (Fig. 5 F); abdominal tergites III or III – IV with median impunctate area (Fig. 2 D); mesotarsomeres 2 – 4 short, triangular.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	description	Figs 2 D, 6 H – J	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	diagnosis	Diagnosis Within the Masasatoi group, Q. piceolineatus can be recognized by the pale yellow-orange body and medial darkening on the sutural area of the elytra (Fig. 2 D). It is most similar to Q. masasatoi but the latter species is darker, bicolored reddish to orange-yellow and dark-brown (Fig. 5 F), with darkened parts of the abdominal tergites. Additionally, the median lobe in ventral view and paramere of Q. masasatoi (Fig. 6 K, M) are distinctly more slender than those of Q. piceolineatus (Fig. 6 H, J) and the apex of the median lobe in lateral view is broader in Q. masasatoi (Fig. 6 L). Quedius masatakai is entirely dark bodied and easily distinguished from pale Q. piceolineatus.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 2000 m; 14 / 5.1934; Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius piceolineatus, O. Scheerpeltz [dark red-pink label] / Quedius (Microsaurus) piceolineatus nov. spec., det. Scheerpeltz [printed label] / 6673 E 91 [blue printed label] / piceolineatus Scheerpeltz [large, printed, folded label] / NHRS-JLKB 000021049 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	description	Redescription Rather similar to Q. masasatoi (description in Smetana 2007; supplemented by Smetana 2012 a) and differing in the following: paler areas of the body in general lighter, brownish-yellow, abdominal tergites entirely pale; antennomeres 7 – 10 slightly less transverse but still wider than long; tergites III – IV with median impunctate area; male tergite X very similar but with slightly and minutely emarginate apex; male sternite IX similar but with expanded middle part more symmetrical and elongate basal part shorter and less elongate; median lobe in ventral view shorter and broader, pair of median teeth slightly farther apart (Fig. 6 H), median lobe in lateral view more strongly narrowed at apex (Fig. 6 I); paramere shorter and more strongly narrowed to apex, which is more rounded, setae on underside in slightly different arrangement (Fig. 6 J). Female Unknown.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	distribution	Distribution Known only from the type locality in Myanmar (Kachin).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	biology_ecology	Bionomics Nothing is known about the natural history of this species but the holotype has two deutonymphs of a uropodine mite attached to the pronotum (Fig. 2 D).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFB85B47FE16FBA2E1A0FCF9.taxon	discussion	Remarks Based on the rather similar male genitalia, slightly emarginate tergite VIII and overall habitus (Fig. 6 H – M), Q. piceolineatus is probably the sister species of Q. masasatoi.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	description	Figs 2 E – F, 7 F	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	diagnosis	Diagnosis Within the Kiangsiensis group, Quedius impressithorax may be distinguished by a combination of: small eyes, temples about 1.5 × as long as eyes (Fig. 2 F); head slightly elongate, with posterior frontal puncture about three puncture diameters from eye margin (Fig. 2 F); body entirely dark (Fig. 2 E).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♀; “ N. E. Burma, Kambaiti; 7000 ft.; 1 / 5.1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius impressithorax, O. Scheerpeltz [dark red-pink label] / Quedius (Microsaurus) impressithorax nov. spec., det. Scheerpeltz [printed label] / 6590 E 91 [blue printed label] / impressithorax Scheerpeltz [large, printed, folded label] / JLKB 000021054 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	description	Redescription Head, elytra, abdomen, antennae and legs dark brown, pronotum and apices of abdominal tergites slightly paler, dark reddish brown; apical tarsomeres of all legs paler, yellow-brown; abdomen vaguely iridescent. Head slightly longer than wide, with posterior angles indistinct; eyes small and only slightly protruding from lateral head outline, tempora longer than eyes, about 1.5 ×; without interocular punctures or additional punctures near the eye margin; posterior frontal puncture somewhat close to posterior margin of eye, about three puncture diameters, situated closer to margin of eye than to hind margin of head; temporal puncture slightly closer to posterior margin of head than margin of eye; surface of head with fine and dense microsculpture of transverse waves, becoming meshed at middle of disc, with distinct micropunctures moderately densely distributed. Antennae somewhat bulky, antennomere 4 only slightly transverse, 5 – 10 distinctly transverse, symmetrical, last segment shorter than previous two. Pronotum distinctly transverse, much wide than long, distinctly explanate laterally; dorsal rows with three punctures, sublateral rows with two punctures, not extended posteriad of large lateral puncture; microsculpture similar to that on head. Scutellum entirely and moderately densely punctate. Elytra at base distinctly narrower than pronotum, at both middle and sides, clearly longer than pronotum; punctures rather fine and dense, many punctures touching laterally; pubescence pale, yellow-brown; disc without microsculpture. Wing fully developed. Abdomen with tergite VII with palisade fringe; punctation sparser than that of elytra, punctures clearly separated by their diameter or more; tergite III with very small sparsely punctate area at middle; tergites with extremely fine microsculpture of transverse waves. Male Unknown. Female Tergite X elongate triangular, strongly narrowed to acutely projected apex, sides of apex sinuate, disc with median darkening from about posterior two-thirds to apex (Fig. 7 F).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	distribution	Distribution Known only from the type locality in Myanmar (Kachin).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA65B44FE1BFB22E033FA5B.taxon	discussion	Remarks The posterior frontal puncture is located at a distance from the eye that is similar to Q. kubani Smetana, 1996 and Q. kuatunensis Smetana, 2013 but in these species the head is clearly transverse, the eyes are larger and the apical antennomeres are asymmetrical. Quedius impressithorax keys to Q. albiorix Smetana, 2012 (Sichuan, China) in Smetana (2017 a) but is quite different in habitus, with entirely dark antennae, a longer head and a distinctly less distant posterior frontal puncture (4 – 5 puncture diameters in Q. albiorix). Quedius impressithorax is here provisionally treated as a valid species until males are found at or near the type locality.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA55B45FD9DFA0CE38EFECF.taxon	description	In its current composition, subgenus Quedius (Raphirus) is highly polyphyletic and it is not possible to provide a useful diagnosis. Currently, it is more practical to work with species groups until a generic revision of the Raphirus lineage is completed.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA55B45FD9DFA0CE38EFECF.taxon	discussion	The Muscicola group of Quedius (Raphirus) as defined here is equivalent to ‘ Clade W’ recovered by the phylogenomic analyses of Brunke et al. (2021). Its members can be recognized by the following combination of characters: head with single basal puncture; eyes large and occupying most of the lateral side of head; antennomere 2 shorter than to about as long as 3; scutellum punctate. Recently, the mainland Chinese species of this group (= part of subclade W 1 of Brunke et al. 2021) were treated together with Q. microsauroides Smetana, 2008 (Smetana 2017 a) as the Muscicola group. However, Q. microsauroides probably belongs to Clade X of Brunke et al. (2021) as it has an impunctate scutellum and much different habitus.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B45FE0AFE72E1B8FBB7.taxon	description	Fig. 3 A	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B45FE0AFE72E1B8FBB7.taxon	diagnosis	Diagnosis Among members of the Muscicola group in the Oriental region, Quedius semilaeviventris may be easily distinguished by a combination of: forebody very dark; pronotum narrow, vaguely longer than wide, strongly narrowed anteriad; elytra with sparse punctation; legs entirely pale; abdominal tergites with large impunctate areas at middle; tergite VII with palisade fringe. A dorsal habitus is given in Fig. 3 A.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B45FE0AFE72E1B8FBB7.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♂; “ N. E. Burma, Kambaiti; 7000 ft.; 12 / 4.1934; R. Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius semilaeviventris, O. Scheerpeltz [dark red-pink label] / Quedius (Raphirus) semilaeviventris nov. spec., det. Scheerpeltz 1944 [printed label] / 6711 E 91 [printed blue label] / NHRS-JLKB 000073671 ”; NHRS.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B45FDFCFB93E32BFA68.taxon	materials_examined	Type material Holotype NEPAL – Province Number 1 • ♂; “ Nepal, Khand-bari District / For. NE Kuwapani; 2500 m; 28. III. 82; A. & Z. Smetana leg. / HOLOTYPE Quedius sundar A. Smetana 1986 [red handwritten label] ”; ASC. Paratypes NEPAL – Province Number 1 • 2 ♂♂; “ Nepal, Khand-bari District / For. above Ahele; 2300 m; 26. III. 1982; A. & Z. Smetana leg. / PARATYPE Quedius sundar A. Smetana 1986 [yellow handwritten label] ”; ASC.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B40FDF3F9FEE085FEE0.taxon	materials_examined	Type material Holotype CHINA – Yunnan Province • ♂; “ CHINA (Yunnan) Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong; 24 ° 5 l′ 22 ″ N, 98 ° 45 ′ 36 ″ E; 2100 – 2200 m; (prim. decid. forest, brook bank, litter, moss sifted); 31. V. 2007; D. W. Wrase leg.; [14 B]; / HOLOTYPE Quedius hecato A. Smetana 2010 [red label] ”; ASC.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B40FDF3F9FEE085FEE0.taxon	description	Redescription The species has been redescribed and diagnosed by Smetana (2017 a) under the name Quedius hecato and described by Smetana (1988) under Q. sundar. These descriptions are only supplemented by the diagnosis above and a dorsal habitus image (Fig. 3 A).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B40FDF3F9FEE085FEE0.taxon	distribution	Distribution This species is now known from Nepal, Myanmar (Kachin) and China (Yunnan).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA45B40FDF3F9FEE085FEE0.taxon	discussion	Remarks The type material of Q. semilaeviventris contains only the male holotype, which was dissected. The aedeagus and external morphology closely matched that of Q. hecato Smetana. The paramere of the type of Q. semilaeviventris is slightly more constricted basally than in the type of Q. hecato but this is considered to be intraspecific variation. As might be expected, the ‘ dark pigmented aedeagus’ of Q. hecato (Smetana 2012 b) is not typical of the species as the aedeagus is normally pigmented in the type of Q. semilaeviventris. The type locality of Q. hecato is Gaoligong Shan, Yunnan, China, which is rather close to Kambaiti Pass in Myanmar. In its description (Smetana 2012 b), Quedius hecato from China was unfortunately never compared to Himalayan Q. sundar. Using only the available illustrations of the aedeagus (Smetana 1988, 2012 b), it would appear that the paramere in Q. hecato was slightly shorter than the median lobe, while it was about the same length in Q. sundar. The male holotype of Q. sundar, previously undissected and therefore not mounted into Canada balsam, was dissected and the aedeagus was examined in both ventral and lateral view. The paramere of the holotype is distinctly shorter than the median lobe and, in general, intermediate between the holotypes of Q. semilaeviventris and Q. hecato, and the available paratype of Q. sundar with the paramere still attached to the median lobe. The arrangement of the two rows of peg setae varies between well organized and highly irregular. The shape of the median lobe in lateral view, where available, was identical across all examined specimens. Therefore both Q. sundar and Q. hecato become junior synonyms of Q. semilaeviventris.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B41FDD9FEFEE0D8FD67.taxon	description	Fig. 3 B	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B41FDD9FEFEE0D8FD67.taxon	diagnosis	Diagnosis Among members of the Muscicola group in the Oriental region, Quedius muscicola may be distinguished by a combination of: pronotum no more than moderately narrowed anteriad; elytra with fine and dense punctation; legs entirely pale; abdominal tergites with sparsely punctate to impunctate areas at middle of apical half; tergite VII with palisade fringe; median lobe in ventral view with arcuate, broadly rounded apex. A dorsal habitus is given in Fig. 3 B.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B4EFE2AFCE3E02EFD00.taxon	materials_examined	Type material Holotype MYANMAR – Kachin State • ♀; “ N. E. Burma, Kambaiti; 2000 m; 4 / 6.1934; Malaise [printed label] / HOLOTYPUS [red handwritten label] / TYPUS Quedius kambaitiensis, O. Scheerpeltz [dark red-pink label] / Quedius (Raphirus) kambaitiensis nov. spec., det. Scheerpeltz 1944 [printed label] / 6599 E 91 / [printed blue label] / NHRS-JLKB 000073668 ”; NHRS. Paratype MYANMAR – Kachin State • 1 ♂; “ [male symbol on card] / N. E. Burma, Kambaiti; 2000 m; 4 / 6.1934; Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll. Scheerpeltz [printed blue label] / COTYPUS Quedius kambaitiensis O. Scheerpeltz [pink label] / kambaitiensis [orange card] / Quedius muscicola Cam., det. A. Smetana 2018 ”; NHW. Non-type material CHINA – Yunnan Province • 1 ♂; Nukiang Lisu, Autonomous Prefecture, Gongshan County, Gaoligong Shan, above ranger station; 27 ° 47.65 ′ N, 98 ° 35.41 ′ E; alt. 2000 m; 19 Jun. 2005; A. Smetana leg.; ASC. NEPAL – Province Number 1 • 1 ♂; East Nepal, Kosi, Chinchilla, “ s / Ahale ”; alt. 2200 m; 4 Apr. 1984; Löbl & Smetana leg.; ASC • 1 ♂; Khandbari District, forest northeast of Kuwapani; alt. 2400 m; 5 Apr. 1984; Smetana & Löbl leg.; ASC.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B4EFE2AFCE3E02EFD00.taxon	description	Redescription Quedius muscicola has been redescribed and diagnosed by Smetana (2017 a). This redescription is only supplemented by the above diagnosis and illustration of the dorsal habitus (Fig. 3 B).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B4EFE2AFCE3E02EFD00.taxon	distribution	Distribution This widespread species is known from India (West Bengal: Darjeeling Distr., Himachal Pradesh, Uttar Pradesh), Nepal, Myanmar, and China (Gansu, Guizhou, Hubei, Shaanxi, Sichuan, Yunnan).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B4EFE2AFCE3E02EFD00.taxon	biology_ecology	Bionomics Quedius muscicola has been collected from forests, field edges and other anthropogenic habitats in moist leaf litter, moss and grassy vegetation, rotting wood and bark, compost and flood debris (Smetana 2017 a).	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
6C38C875FFA05B4EFE2AFCE3E02EFD00.taxon	discussion	Remarks The male paratype of Q. kambaitiensis was dissected to reveal an aedeagus entirely consistent with that of widespread and variable species Q. muscicola. Therefore Q. kambaitiensis syn. nov., becomes a synonym of Q. muscicola. The determination was originally made by A. Smetana in 2018 but it is here confirmed after a comparison with similar species of the group occurring in both the Himalayan region and mainland China. Both the holotype and paratype are externally consistent with Q. muscicola and can be recognized as such by the features given in the diagnosis above. Quedius muscicola is a rather variable species, especially outside of mainland China (Smetana 2017 a), so it is not unexpected that Scheerpeltz (1965) also reported Q. muscicola from Kambaiti, while describing Q. kambaitiensis in the same paper.	en	Brunke, Adam James (2023): Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar. European Journal of Taxonomy 864: 117-145, DOI: https://doi.org/10.5852/ejt.2023.864.2093
