taxonID	type	description	language	source
8E8D13BF83915762B876AB0A8897FB78.taxon	description	Fig. 9	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
8E8D13BF83915762B876AB0A8897FB78.taxon	diagnosis	Diagnosis. This species corresponds to the mitochondrial lineage NE 2 as defined herein. It is assigned to the subgenus Guibemantis Pandanusicola of the genus Guibemantis based on presence of intercalary elements between ultimate and penultimate phalanges of fingers and toes (verified by external examination), small body size, moderate to weakly expressed webbing between toes, connected lateral metatarsalia, the presence of both inner and outer metatarsal tubercles, femoral glands in males, absence of nuptial pads, small body size (SVL 25.1 - 26.0 mm in males; female size unknown), and molecular phylogenetic relationships. Within Pandanusicola, the new species is distinguished from all species except G. liber, G. razandry, G. razoky, and G. tasifotsy by femoral glands type 1 (vs. type 2) as defined by Glaw et al. (2000), thus possessing a large number of small gland granules in a relatively diffuse field covering most of the thigh ventrally, and by its probable breeding in open swamps (vs. phytotelmic breeding in Pandanus leaf axils). It can be distinguished from G. tasifotsy by its different brownish color pattern lacking a green dorsal and lateral coloration with series of distinct white blotches along the lower flanks. The new species differs from all G. liber lineages occurring in the Northern Central East and Southern Central East of Madagascar by its high DNA divergence> 5 % in the mitochondrial 16 S gene, and probably by a somewhat smaller snoutvent length. Guibemantis razoky (see above) has a larger body size (26.5 - 33.9 mm in males, vs 25.1 - 26.0 mm in males of G. fotsitenda). Guibemantis razandry (see above) is the closest relative of G. fotsitenda sp. nov., and no obvious morphological differences between these two species are known, despite their clear divergence in mitochondrial and nuclear-encoded DNA in near-sympatry. The new species differs from G. liber, G. razandry, and G. razoky by 23, 23, and 50 diagnostic positions in the analyzed fragment of the cytochrome b gene, respectively (see Appendix 2 for a list of diagnostic sites).	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
8E8D13BF83915762B876AB0A8897FB78.taxon	description	Description of holotype. Adult male in good state of preservation (Fig. 9). Pieces of muscle tissue removed from both left and right thigh for molecular analysis. SVL 25.5 mm. For full morphometric measurements see Table 1. Body relatively slender; head slightly longer than wide, wider than body; snout slightly pointed in dorsal and lateral views, rounded in ventral view; nostrils much nearer to tip of snout than to eye and pointed anterolaterally; canthus rostralis relatively distinct, slightly concave; loreal region concave; tympanum distinct, relatively small, its diameter 60 % of eye diameter; distinct supratympanic fold; tongue ovoid, distinctly bifid posteriorly; vomerine teeth as one weakly expressed rounded aggregation posterolateral of each choana; choanae small, rounded. Forelimbs slender; subarticular tubercles distinct and single; central metacarpal tubercle large and rounded, outer metacarpal tubercle smaller and oval; a small but indistinct prepollex (which could also be considered as an inner metacarpal tubercle) at base of first finger. Fingers without webbing; relative finger length I <II <IV <III; finger discs distinctly enlarged; nuptial pads absent. Outer toe and finger discs darker than inner toe and finger discs. Hind limbs long and slender; when adpressed along body, tibiotarsal articulation reaches center of eye; lateral metatarsalia connected by tissue; inner metatarsal tubercle distinct, larger than outer; outer metatarsal tubercle distinct; webbing formula of the foot 1 (traces), 2 i (traces), 2 e (1), 3 i (2.25), 3 e (1.25), 4 i (2.5), 4 e (2.75), 5 (1); relative toe length I <II <V <III <IV. Skin dorsally smooth; ventral skin smooth on throat and chest, slightly granular on belly. Femoral glands not intact due to tissue excision. After sixteen years in preservative (70 % EtOH; Fig. 9), dorsal background coloration light brownish with irregular beige mottling, and several scattered dark brown spots. Area above eyes dark brown, a thin interocular band with a small beige medial interruption. A dark brown rostral stripe is present. Tympanic area dark brown. Several dark brown crossbands present on hindlimbs. Ventral side without dark color elements; belly of a faded beige, throat bright white. Coloration in life not recorded.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
8E8D13BF83915762B876AB0A8897FB78.taxon	distribution	Distribution. The species is reliably known from two sites in northern Madagascar: (1) the type locality Marojejy (Camp Mantella, at low elevation), and (2) a site between Andrakata and Andapa also located at rather low elevation. Furthermore, individuals from (3) Ambodivoangy at the north-eastern edge of the Makira Reserve, at ca. 30 m a. s. l., are provisionally assigned to this species based on evidence from nuclear genes, despite their assignment to G. razandry based on mitochondrial DNA (see Discussion below). This seems to be a species specialized to habitat at low elevations (known from near sea level to ca. 480 m a. s. l.).	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
8E8D13BF83915762B876AB0A8897FB78.taxon	etymology	Etymology. The name is derived from the Malagasy words fotsy meaning white, and tenda meaning throat, referring to the white throat (vocal sac) typical for this and other species of the G. liber complex. The name is used as a noun in apposition to the genus name.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
CC73535B35E85741ADF7D0BB12B425C4.taxon	description	Figs 9, 11	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
CC73535B35E85741ADF7D0BB12B425C4.taxon	diagnosis	Diagnosis. This species corresponds to the mitochondrial lineage NE 1 as defined herein, and to the candidate species Guibemantis sp. Ca 21 according to Perl et al. (2014). It is assigned to the subgenus Guibemantis Pandanusicola of the genus Guibemantis based on presence of intercalary elements between ultimate and penultimate phalanges of fingers and toes (verified by external examination), small body size, moderate to weakly expressed webbing between toes, connected lateral metatarsalia, the presence of both inner and outer metatarsal tubercles, femoral glands in males, absence of nuptial pads, small body size (SVL 24.4 - 25.9 mm in reliably sexed males and 22.8 mm in one female), and molecular phylogenetic relationships. Within Pandanusicola, the new species is distinguished from all species except G. liber and G. tasifotsy by femoral glands type 1 (vs. type 2) as defined by Glaw et al. (2000), thus possessing many small gland granules in a relatively diffuse field covering most of the thigh ventrally, and by its probable breeding in open swamps (vs. phytotelmic breeding in Pandanus leaf axils). It can be distinguished from G. tasifotsy by its different brownish color pattern lacking a green dorsal and lateral coloration with a series of distinct white blotches along the lower flanks and its strongly different advertisement call, consisting of a pulsatile note with numerous pulses being largely fused (vs. a trill-like note containing 3 - 7 distinctly separated pulses). The new species differs from all G. liber lineages occurring in the Northern and Southern Central East of Madagascar by its high DNA divergence, with> 5 % uncorrected pairwise distance in the mitochondrial 16 S gene and 20 diagnostic positions in the analyzed fragment of the cytochrome b gene (see Appendix 2 for a list of diagnostic sites), as well as probably by a somewhat smaller snout-vent length. For a distinction from the other two new species described herein, see below.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
CC73535B35E85741ADF7D0BB12B425C4.taxon	description	Description of holotype. Adult male in excellent state of preservation (Fig. 9). A small piece of muscle tissue from right thigh removed for molecular analysis. SVL 25.6 mm. For full morphometric measurements see Table 1. Body relatively slender; head slightly longer than wide, wider than body; snout rounded in dorsal, ventral, and lateral views; nostrils much nearer to tip of snout than to eye and pointed anterolaterally; canthus rostralis distinct, slightly concave; loreal region concave; tympanum distinct, relatively small, its diameter 69 % of eye diameter; distinct supratympanic fold; tongue ovoid, distinctly bifid posteriorly; vomerine teeth as one weakly expressed rounded aggregation posterolateral of each choana; choanae small, rounded. Forelimbs slender; subarticular tubercles distinct and single; central metacarpal tubercle large and rounded, outer metacarpal tubercle smaller and oval; a small but indistinct prepollex (which could also be considered as an inner metacarpal tubercle) at base of first finger. Fingers without webbing; relative finger length I <II <IV <III; finger discs distinctly enlarged; nuptial pads absent. Outer toe and finger discs darker than inner toe and finger discs. Hind limbs long and slender; when adpressed along body, tibiotarsal articulation reaches beyond eye; lateral metatarsalia connected by tissue; inner metatarsal tubercle distinct, larger than outer; outer metatarsal tubercle distinct; webbing formula of foot 1 (traces), 2 i (traces), 2 e (1), 3 i (2.5), 3 e (1.5), 4 i (2.75), 4 e (3), 5 (1); relative toe length I <II <III = V <IV. Skin dorsally smooth; ventral skin smooth on throat and chest, slightly granular on belly. Femoral glands relatively distinct from external view, consisting of large number of small gland granules in a relatively diffuse field covering most of thigh ventrally, thus of type 1 as defined by Glaw et al. (2000). In life gland granules distinctly recognizable as small greenish-yellowish units, at least 170 in one gland (Fig. 11). After sixteen years in preservative (70 % EtOH; Fig. 9), dorsal background coloration light brownish with two prominent dark brownish dorsolateral bands extending posteriorly from eye orbits to hips. Rostral stripe dark brownish. Dorsally numerous dark reddish brown irregular spots present, particularly between orbits and at middorsum. Forelimbs have irregular and partially interrupted dark, brownish bands and spots extending from shoulders to fingers. Outer finger discs reddishbrown. Dorsal surface of thigh with broad dark blotches and interrupted bands. These darkish patterns extend to shanks and continue as single blotches and spots on feet and toes. Like finger discs, some outer toe discs dark reddish brown, noticeably different than adjacent tissue. Based on photographs of holotype in life (Fig. 11), body coloration was as follows: on dorsum reddish brown, on limbs and laterally greenish gray. Dorsolaterally a thin yellowish line on each side. Irregular-sized dark blackish brown spots and dots middorsally and laterally, but particularly on both fore- and hindlimbs. Supratympanic fold and rostral stripe blackish brown. Paired, thin white dorsal lines in life (not visible in preservative) (Fig. 11). Background color of ventral surface whitish to greenish, chest and throat bright white. Posterior and lateral parts of abdomen semi-transparent. Femoral gland granules yellow. Iris (whitish in preservative) apparently glossy golden in life.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
CC73535B35E85741ADF7D0BB12B425C4.taxon	distribution	Distribution. The species is known from various sites in northern Madagascar, all at mid- to high-elevation: (1) the type locality Marojejy (Camp Simpona, at mid-elevation), (2) Bemanevika, (3) the southern slope of the Tsaratanana Massif, (4) the western slope of Makira Reserve, and (5) Anjanaharibe-Sud Reserve, based on specimen CRH 1693 (KU 347374) included in the FrogCap analysis (Fig. 8). At Makira (west), Bemanevika and Tsaratanana, the species occurs syntopically with G. razoky sp. nov. (described below). The species is known from elevations between 1093 and 2021 m a. s. l.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
CC73535B35E85741ADF7D0BB12B425C4.taxon	etymology	Etymology. The name is derived from the Malagasy word Guibemantis razandry meaning smaller (younger) sibling, and makes reference to the fact that this species is the smaller-sized relative of the syntopic larger-sized species of the G. liber complex described in the following. The name is used as a noun in apposition to the genus name.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
2603B965B02351009DF29C540AB6969A.taxon	description	Figs 9, 12, 13	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
2603B965B02351009DF29C540AB6969A.taxon	diagnosis	Diagnosis. This species corresponds to the mitochondrial lineages NOR + NCENTR as defined herein. It is assigned to the subgenus Guibemantis Pandanusicola of the genus Guibemantis based on presence of intercalary elements between ultimate and penultimate phalanges of fingers and toes (verified by external examination), moderate to weakly expressed webbing between toes, connected lateral metatarsalia, the presence of both inner and outer metatarsal tubercles, femoral glands in males, absence of nuptial pads, moderately small body size (SVL 26.5 - 33.9 mm in males and 29.8 - 32.8 mm in females), and molecular phylogenetic relationships. Within Pandanusicola, the new species is distinguished from all species except G. liber, G. razandry, and G. tasifotsy by femoral glands type 1 (vs. type 2) as defined by Glaw et al. (2000), thus possessing many small gland granules in a relatively diffuse field covering most of the thigh ventrally, and by its probable breeding in open swamps (vs. phytotelmic breeding in Pandanus leaf axils). It can be distinguished from G. tasifotsy by its different brownish color pattern lacking a green dorsal and lateral coloration with series of distinct white blotches along the lower flanks, and its different advertisement call, namely a short click-like note of 20 - 117 ms duration and 2598 - 3010 Hz dominant frequency (vs. a longer trill-like note of 147 - 516 ms duration and higher dominant frequency; Lehtinen et al. 2012). The new species differs from all G. liber lineages occurring in the Northern and Southern Central East of Madagascar by its high DNA divergence> 2.4 % in the mitochondrial 16 S gene, by a larger SVL, and differences in the advertisement call. It differs from G. razandry (described above) by larger SVL, different advertisement call, and a molecular 16 S divergence> 5.5 %. It also differs from G. liber and G. razandry by 3 and 41 diagnostic positions in the analyzed fragment of the cytochrome b gene, respectively (see Appendix 2 for a list of diagnostic sites). For a distinction from the third new species described herein, see below.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
2603B965B02351009DF29C540AB6969A.taxon	description	Description of holotype. Adult male in good state of preservation (Fig. 9). A small piece of muscle tissue from right thigh removed for molecular analysis. Ventral skin cut open and bladder removed for parasite examination. SVL 28.0 mm. For full morphometric measurements see Table 1. Body relatively slender; head slightly longer than wide, wider than body; snout rounded in dorsal, ventral, and lateral views; nostrils much nearer to tip of snout than to eye, slightly protuberant and pointed anterolaterally; canthus rostralis distinct, straight; loreal region straight; tympanum distinct, relatively small, its diameter 70 % of eye diameter; distinct supratympanic fold; tongue ovoid, strongly bifid posteriorly; posterior tongue extensions slightly serrated; vomerine teeth as one weakly expressed rounded aggregation posterolateral of each choana; choanae small, rounded. Forelimbs slender; subarticular tubercles distinct and single; central metacarpal tubercle large and rounded, outer metacarpal tubercle smaller and oval; a small but indistinct prepollex (which could also be considered as an inner metacarpal tubercle) at base of first finger. Fingers without webbing; relative finger length I <II <IV <III; finger discs distinctly enlarged; nuptial pads absent. Outer toe and finger discs darker than inner toe and finger discs. Hind limbs long and slender; when adpressed along body, the tibiotarsal articulation reaches beyond the eye; lateral metatarsalia connected by tissue; inner metatarsal tubercle distinct, larger than outer; outer metatarsal tubercle distinct; webbing formula of the foot 1 (traces), 2 i (traces), 2 e (1), 3 i (2), 3 e (1), 4 i (2.75), 4 e (3), 5 (1.25); relative toe length I <II <III = V <IV. Skin dorsally smooth; ventral skin as far as recognizable smooth on throat, chest, slightly granular on belly. Femoral glands recognizable from external view but not very distinct, also in life (Fig. 12) not very prominent and of same color as surrounding shank, possibly because the specimen was collected outside of the reproductive season. Glands consisting of many small gland granules in a relatively diffuse field covering most of thigh ventrally, thus of type 1 as defined by Glaw et al. (2000). After eleven years in preservative (70 % EtOH; Fig. 9), dorsal background coloration grayish brown with two prominent blackish dorsolateral discontinuous bands consisting of densely arranged blotches and extending posteriorly from eye orbits to hips. Rostral stripe dark brownish. Dorsally, from between orbits a dense field of brown blotches, running over most of dorsum and becoming more scattered on posterior dorsum. Interrupted and indistinct whitish middorsal line. Forelimbs with irregular and partially interrupted dark, brownish bands and spots extending from shoulders to fingers. Outer finger discs darkish brown. Ventrally, throat and forelimbs largely unpigmented (yellowish-brownish in preservative), chest with some fine dark brown dotting, belly more pigmented with dense pattern of brown dots that leave out some larger unpigmented markings, and the hindlimbs dark largely brown. Dorsal surface of thighs and shanks dark reddishbrown with indistinct broad dark blotches and interrupted bands. Like finger discs, some outer toe discs dark reddish brown, of noticeably different color than adjacent tissue. Based on photographs of the holotype specimen (Fig. 12), body coloration in life was pinkish brown. Dorsally and on both forelimbs and hindlimbs, with a chocolate brown pattern, consisting of irregular-sized blotches and spots. Also, supratympanic fold and rostral stripe chocolate brown. Dorsally, from an imaginary line between the orbits to the axilla with a dense field of brown blotches. A yellowish interrupted and indistinct middorsal line. Ventrally, background color pinkish-whitish, but chest, throat and the anterior part of belly bright white. Posteriorly and laterally belly semi-transparent. Ventrolaterally, small white spots in anterior part, larger bright yellow spots posteriorly. First finger and toe intensely yellow. Numerous femoral gland granules visible, but not highlighted in color. Iris (whitish in preservative) copper metallic in life.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
2603B965B02351009DF29C540AB6969A.taxon	distribution	Distribution. Since many Pandanusicola species appear to be phenotypically similar and are therefore often taxonomically confused, we restrict our assessment of distribution to populations for which molecular data are available. According to our data, G. razoky appears to be a regional endemic of northern Madagascar, and is so far reliably known from six localities (not taking into account the imprecise site " Bealanana region ": (1) Bemanevika, the type locality; (2) Ampotsidy; (3) Andranonafrindra forest; (3) Tsaratanana; (4) Makira; (5) Montagne d'Ambre (where a genetically distinct mitochondrial lineage occurs). These localities are from elevations between 1044 and 1466 m a. s. l. Furthermore, genetic samples assigned to this species based on mitochondrial DNA exist from the low-elevation site Ambodiriana (about 50 m a. s. l.) but no voucher specimens from this site are available, and this record thus requires confirmation.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
2603B965B02351009DF29C540AB6969A.taxon	etymology	Etymology. The name is derived from the Malagasy word Guibemantis razoky meaning larger (elder) sibling, and refers to the fact that this species is the larger-sized relative of the syntopic G. razandry. The name is used as a noun in apposition to the genus name.	en	Koppetsch, Thore, Pabijan, Maciej, Hutter, Carl R., Koehler, Joern, Gehring, Philip-Sebastian, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Scherz, Mark D., Vieites, David R., Glaw, Frank, Vences, Miguel (2023): An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397-432, DOI: http://dx.doi.org/10.3897/vz.73.e94063, URL: http://dx.doi.org/10.3897/vz.73.e94063
