taxonID	type	description	language	source
03AB87D4CE5AFF8CFF2A93FCE818FB82.taxon	description	(Figs 1, 4, 16 – 18, 29, 31)	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE5AFF8CFF2A93FCE818FB82.taxon	materials_examined	Holotype ♀: 200 _ 13.2 [collecting number] Switzerland, GR, S-charl, 1800 m, Hochstauden entlang Clemgia, 5.7.2013, leg. C. Germann (NMBE). Paratypes: (39 in total), all from Switzerland: 2 ♀ [GR], Bernina, coll. L. v. Heyden (SDEI). 1 ♀ [GR], St. Mtz. [St. Moritz, most likely collected 1862 after von Heyden (1863)], coll. L. v. Heyden (SDEI). 2 ♀ GR, Miralago, 18.7.1970, leg. P. Scherler (NMBE). 1 ♀ GR, Alp Grüm, 21.7.1970, leg. P. Scherler (NMBE). 1 ♀ GR, Val dal Fain, 23.7.1979, Mousses sol [sifted], leg. P. Scherler (NMBE). 1 ♀ GR, E Alp Trupchun, Kurzrasen, 22. 6. – 16.7.2000, 2100 m, 802.480 / 163.680 [barber traps], leg. B. Lüscher (NMBE). 1 ♀ GR, Alp Chaschauna, 2200 m, 800.380 / 161.760, 14. 7. – 4.8.2000 [barber traps], leg. B. Lüscher (NMBE). 1 ♀ GR, Alp Trupchun, Vegetationsmosaik, Seggen, Kurzweide, 2060 m, 802.360 / 163.920, 16. 7. – 5.8.2000 [barber traps], leg. B. Lüscher (NMBE). 2 ♀ GR, W Alp Trupchun, Vegetationsmosaik, Eisenhut / Seggen / Kurzweide, 2020 m, 802.040 / 164.100, 5. 8. – 1.9.2000 [barber traps], leg. B. Lüscher (NMBE). 2 ♀ GR, W Alp Trupchun, Vegetationsmosaik Eisenhut, Seggen, Kurzweide, 2020 m, 802.040 / 164.100, 31. 8. – 28.9.2000 [barber traps], leg. B. Lüscher (cCG, NMBE). 4 ♀ GR, W Alp Chaschauna, Vegetation mosaikartig, feucht, 2200 m, 800.280 / 162.030, 21.10.2000 – 19.6.2001 [barber traps], leg. B. Lüscher (NMBE). 1 ♀ GR, Sur, Alp Flix, 2145 m, 770.700 / 154.500, Gesiebe Rhododendron, Polsterpflanzen, Moos und unter Stein, 20.6.2002, leg. C. Germann (NMBE). 1 ♀ GR, Valle Poschiavo, Garbela, 2 km NW Brusio, 1000 m, 28.8.2004, leg. C. Germann (NMBE). 3 ♀ GR, Alp Flix, Salategnas, 1960 m, 24. 7. – 21.8.2005, leg. P. Muff (BNM, NMBE). 1 ♀ 021 _ 06.1 & 2 [collecting number] GR, Ftan, 812.730 / 186.400, 1900 m, Gesiebe Moos, S-Hang, 9.11.2006, leg. C. Germann (NMBE). 1 ♀ GR, Ardez, Chanoua, 811.850 / 184.850, 1600 m, 30.6.2007, Käscherfang, leg. M. Geiser (cMG). 4 ♀ GR, 4 km SW Zernez, Prazet, 1650 m, 46 ° 40 ‘ 12 " N, 10 ° 02 ‘ 51 " E, leg. C. Braunert (cCB). 1 ♀ GR, Ftan, Piz Clünas, 814.100 / 188.854, S-exp. Hang, 2560 m, Gesiebe Felssimse Flechten & Polsterpflanzen, 12.8.2011, leg. C. Germann (NMBE). 1 ♀ 199 _ 13.5 [collecting number] GR, Val Mora, DÖss Radond, 823.289 / 161.728, 2230 m, 28.6.2013, leg. C. Germann (NMBE). 1 ♀ 199 _ 13.1 [collecting number] GR, Valchava, Val Vau, Spi da Vau, 826 / 164, 1700 – 1800 m, 28.6.2013, leg. C. Germann (NML). 7 ♀ 200 _ 13.2 [collecting number] GR, S-charl, 1800 m, Hochstauden entlang Clemgia, 5.7.2013, leg. C. Germann (cCG, cHW, NMBE).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE5AFF8CFF2A93FCE818FB82.taxon	materials_examined	Italy: 1 ♀ Südtirol, oberhalb Trafoi, 1700 m, 10.8.1973, in Schlag abends gestreift, leg. M. Kahlen (TLMF). 1 ♀ [Trentino] Val Camonica, [locality handwritten, illegible, without date], leg. G. Krüger, coll. O. Leonhard (SDEI). 1 ♀ [Trentino] Mt. Pari, Judicaria [without date], coll. Reiss (TLMF). 1 ♀ [Trentino] P. Fugazza [Vallarsa], 24.6.1897, coll. Knabl (TLMF). 2 ♀ [Trentino] Piano della Fugazza, 30.7.09, Vallarsa Südtirol, Gesiebt aus Laub, leg. StÖcklein (NMBA). 1 ♀ Trentino, Rollepass, 1950 m, 4.6.1977, unter Stein, leg. M. Kahlen (TLMF). 1 ♀ Trentino, Cma. Pari, 1950 – 1990 m, 15.6.1979, Rasengesiebe, leg. M. Kahlen (TLMF). 1 ♀ Trentino, Cima Pari, Gipfel, 1950 – 1980 m, Grünerlenstreu, 6.7.1986, leg. M. Kahlen (TLMF). 1 ♀ Trentino, Monte Pari, Baitone di Gelos, 1450 m, 14.6.1984, leg. M. Kahlen (TLMF). 2 ♀ Trentino, Cima Pari, Gipfel, 1950 – 1990 m, Grünerlenstreu, 6.7.1986, leg. M. Kahlen (TLMF). 1 ♀ [Veneto] Vicenza, Altipiano d’Asiago, Cma. Undici, W-Hang, 2100 m, 9.8.1978, Rasengesiebe, leg. M. Kahlen (TLMF). 1 ♀ [Veneto] Campogrosso [without date], coll. Knabl (TLMF). 1 ♀ [Veneto] Vicenza, Altipiano d’Asiago, Dosso del Cuvolin, 2100 m, 30.5.1980, Alpenrosenmulm, leg. M. Kahlen (TLMF). 1 ♀ [Veneto] Alpi Lessini, Cima Carega, 2100 – 2200 m, 25.6.2007, leg. A. Szallies (cAS). Slovenia: 1 ♀ Julijske Alpe, M. Mojstrovka, N-Kar, 1800 m, 3.6.1980, Rhododendron, leg. Kahlen (TLMF). 1 ♀ Steiner Alpen, Logarska Dolina, 27.6.1988, leg. M. Egger (SDEI).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE5AFF8CFF2A93FCE818FB82.taxon	description	Size holotype: 4.6 mm Body colour: Auburn, head and rostrum dark brown. Head, rostrum and antennae: Head globular, eyes inversely drop-shaped, flat, not protruding from the heads outline. Frons between the eyes narrower than rostrum between the antennal insertion. Upper part of head and epistome up to insertion of antennae densely covered with narrow, oval and beige coppery shimmering scales. Rostrum a little longer than its widest part at the outline of the pterygia. Pterygia prominent, rostral apex glossy and sloped, V-shaped, where both tips are prolonged in form of little horns. Antennal scape strong, clubbed, thickest after its last third. First two segments of antennal funiculus longer than wide (L / B 1 st and 2 nd: 2.3), following five segments as long as wide, globular, club fusiform, twice as wide as last antennal segment. Pronotum: Index (L / B): 0.9. Transverse, laterally strongly rounded, widest in or just behind middle, constricted at front and hind margin. Surface tuberculate and microgranulate. Vestiture consisting of adherent round, beige coppery shimmering scales and raised, narrow and dark brown setulae, radially pointing to the middle of the pronotum. Elytra: Index (L / B): 1.3. Base slightly broader than pronotum, elytra oval, egg shaped, without humeral callus (apterous). Vestiture consisting of irregularly dispersed, beige to darker brown, coppery shimmering, adherent round scales. Uneven intervals (3, 5, 7) weakly elevated, regularly set with long and raised, clubbed setulae, up to 4 times longer than wide. Even intervals with somewhat shorter raised, clubbed setulae, best visible towards apex. Striae regularly punctuate with round scales inserting at their frontal margin. Legs: strong, femora unarmed, three visible tarsal segments (fourth minute), first twice as long as second, third twice the width of second, claw segment as long as second and third combined, claws simple. Spiculum ventrale, spermatheca and ovipositor: branches of the spiculum at base of plate angular and slightly diverging towards apex (Fig. 16). Spermatheca with well-developed long and bowed cornu, and short globular ramus and nodulus (Fig. 17). Ovipositor as in Fig. 18. Variation: Sizes vary (3.5 – 4.6 mm), proportions are rather constant, as is same with the contrasting vestiture. Body colour varies from auburn to dark brown. The branches of the spiculum's plate vary from diverging to parallel. Derivation of name: The new species is dedicated to Patrick Muff (Bern), passionate biologist, enthusiastic gymnasium teacher of future biologists and collector of the new species.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE5AFF8CFF2A93FCE818FB82.taxon	discussion	Otiorhynchus muffi sp. nov. is presently known from Grisons in the West to Slovenia in the East (Fig. 29) and was collected at altitudes from 1000 m (Garbela, Valle Poschiavo) up to 2560 m a. s. l. at Piz Clünas, lower Engadine (Fig. 31). The habitat varies from spruce tree forests (Larix decidua, Piceae abies) where O. muffi sp. nov. was beaten from tall forbs and Alnus incana, to moss and leaf litter and from low vegetation on rocks where the new species was sifted and extracted with the method illustrated in Germann (2014 a). On Alp Trupchun and Alp Flix (Grisons) both O. muffi sp. nov. and O. subcostatus were collected together in barber / pitfall-traps set in mosaic like growing plants as Aconitum and Carex in alpine meadows. Manfred Kahlen collected specimens of O. muffi sp. nov. by sifting leaf litter under Alnus and Rhododendron and from alpine meadows.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE5EFF80FF2B977CEE09FC86.taxon	description	Species of the subgenus Nihus are recognisable by the following traits (modified after Reitter 1913): i) sloped and glossy apex of rostrum, ii) tuberculate pronotum, iii) frons between the eyes not or hardly wider than the rostral dorsum between the insertions of the antennae, iv) comparably dense vestiture of body. Eunihus shares the traits mentioned for Nihus, except for the frons which is considerably wider than the rostral dorsum. Sizes vary, also due to polyploidy in the case of O. carinatopunctatus and supposedly also in O. subcostatus. A sexual dimorphism is present where males are narrower with stronger legs (O. azaleae, O. spaethi, O. hypocrita) or males are stouter and more globular (O. gredleri). Also genital structures vary to a certain extent, see as an example the form of the plate of the spiculum in the case of O. spaethi (Figs 20 – 21) or O. azaleae (Figs 22 – 23). On the other hand the shape of the spermatheca was found to be rather constant in all species examined and hence not useful for species discrimination. All examined specimens (except for the widespread and unambiguously recognisable species O. uncinatus and O. carinatopunctatus) are listed and their records are shown on the maps (Figs 29 – 30).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE55FF84FF4F9701EE12FC65.taxon	materials_examined	Type locality: Engadin Lectotype ♀: [Switzerland, GR], Engadin [handwritten] // O. subcostatus [handwritten] m. [mihi] // coll. Stierlin // Syntypus [red paper] // coll. DEI Müncheberg. Red label: Lectotype Otiorhynchus (Nihus) subcostatus Stierlin, 1866 des. C. Germann 2014 (SDEI).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE55FF84FF4F9701EE12FC65.taxon	materials_examined	Austria: 1 ♀ [North Tyrol] Brandenberg, Ht. Sonnwendjoch, SW-Hang, 1800 m, 7.7.1984, leg. Kahlen (TLFM). 1 ♀ Nordtirol, Stubaier Alpen, Sandestal, 1850 m, Zwergstrauchheide, Vaccinium - Streu, 7.6.2003, leg. M. Kahlen (TLMF). 1 ♀ Osttirol, Granatspitzgruppe, Steinertal, 2050 m, 21.5.1993, leg. M. Kahlen (TLMF). 1 ♀ Osttirol, Lienz Umgebung, Amlach, 16.7.1994, 1050 m, leg. A. Kofler (TLMF). France: Alpes-Maritimes, Vacherie du Boréon, 1680 m, 28.6.2010, leg. M. Kahlen (TLMF). 2 ♀ Hautes Alpes, Dévoluy, Tête de Lapras, 1700 m, 7.5.1999, leg. M. Kahlen (TLMF). 3 ♀ 113 _ 10.4 [collecting number] Haut Jura, Grand Crêt, 485.675 / 126.772, 1630 m, 4.6.2010, leg. C. Germann (cCG). 1 ♀ Haute-Savoie, Samoens, Pte. de Ressache, 1830 m, Gesiebe alpiner Rasen, 20.5.2002, leg. C. Germann (NMBE). 1 ♀ Germany: 1 ♀ [Bavaria] Berchtesgaden, Kiefernwald, 1100 m (cHW). 1 ♀ Bayern, Wimbach, 2.8.1986, leg. H. Winkelmann (cHW). 1 ♀ 034 _ 07.1 [collecting number] Bayern, Allgäuer Alpen, Oberstorf, 1490 m, 26.7.2007, leg. C. Germann (cCG). 1 ♀ Bayern, Oberbayern, Bayrischzell, 17.7.2012, leg. H. Mühle (NMBE). Italy: 1 ♀ [Friuli] FVG, Sella Nevea, 1300 – 1400 m, 9.10.2001, leg. Bayer & Winkelmann (cHW). 1 ♀ [Lombardia] Brescia, Passo di Croce Domini, 2300 m, 6.7.1980, leg. M. Kahlen (TLMF). 1 ♀ [Lombardia] Brescia, Monte Guglielmo, Dosso Pedalta, 1950 m, 2.11.1989, leg. M. Kahlen (TLFM). 2 ♀ Lombardia, Grigna settentrionale, 1874 m, Gesiebe und geklopft Rhododendron, Rhamnus, Salix, 7.7.2002, leg. C. Germann (cCG, NMBE). 1 ♀ Lombardia, Macugnaga [without date] (NMBE). 1 ♀ [Lombardia] Bergamo, Alpi Orobie, Averara, Passo di Verobbio, 2030 m, 13.7.2010, leg. M. Kahlen (TLMF). 1 ♀ [Piemonte] Cuneo, Terme di Valdieri, S Vallone di Ciriegia, 2150 m, 24.6.2008, leg. M. Kahlen (TLMF). 1 ♀ Südtirol, Seiser Alpe, W-Seite unt. Weiden gesiebt, 19.5.1973, leg. M. Kahlen (TLMF). 1 ♀ Südtirol, oberhalb Trafoi, 1700 m, 10.8.1973, leg. M. Kahlen (TLMF). 1 ♀ Südtirol, FranzenshÖhe, 2300 m, 14.8.1977, unt. Stein, leg. M. Kahlen (TLFM). 1 ♀ Trentino, Monte Baldo, Mt. Altissimo, S-Seite, 1900 m, Weidenlaub, 12.10.1997, leg. M. Kahlen (TLFM). 1 ♀ [Veneto] Verona, Mt. Baldo, Cma. Valdritta, Gipfel, 2200 m, 18.6.1983, nachts an Felsen, leg. M. Kahlen (TLFM). 1 ♀ [Veneto] Udine, Prealpi Giulie, Monte Matajur, N-Seite, 1630 m, 25.9.1998, leg. M. Kahlen (TLMF). 1 ♀ [Veneto] Treviso, Mt. Grappa, Val die Lebi, 1100 m, BF [Becherfalle] 15. 5. – 23.9.2006, leg. M. Kahlen (TLMF). Slovenia: 1 ♀ Kamniske Alpe, Okreselj, 1400 m, 27.6.1988, leg. M. Kahlen (TLMF). Switzerland: 1 ♀ BE, Gadmen, 6.8.1903, leg. A. Rätzer (NMBE). 1 ♀ BE, Gurnigel, 19.7.1959, leg. H. Schenk (NMBE). 1 ♀ BE, Bäderhorn, 16.9.1979, leg. P. Scherler (NMBE). 1 ♀ BE, Grischbachtal, 586 / 153, 1500 m, gekäschert, M. 6.1996, leg. C. Germann (cCG). 1 ♀ BE, Sustenpass, 1700 – 2000 m, gesiebt aus Moos und Bodenstreu, 14.8.2001, leg. C. Germann (cCG). 1 ♀ BE, Kiental, Dreispitz, 2000 m, gesiebt aus Moos, 5.10.2001, leg. C. Germann (cCG). 1 ♀ BE, Wimmis, 613.970 / 168.800, 630 m, Auenwald, 17.5.2006, leg. C. Germann (cCG). 1 ♀ BE, Kandersteg, Sunnbüel, 1960 m, unter Dryas - Polster, 18.7.2012, leg. P. Sonderegger (NMBE). 1 ♀ FR, Plaffeien, Kaisereggpass, 2100 m, 6. – 11. 2010, Fallen, leg. A. Szallies (cAS). 1 ♀ GL, Linthal, Ochsenstein, 28. 6. – 11.7.2008, leg. P. Marti (NMBE). 1 ♀ GL, Linthal, Ochsenstein, 19. 6. – 28.6.2008, leg. D. Gloor (NMBE). 1 ♀ GR, Tantermozza, 12.8.1920, Moos, leg. E. Handschin (BNM). 2 ♀ GR, Val Sinestra, 21.7.1971, leg. P. Scherler (NMBE). 1 ♀ GR, E Alp Trupchun, Vegetationsmosaik Seggen / Kurzweide, 22. 6. – 13.7.2000, 2140 m, leg. B. Lüscher (NMBE). 1 ♀ GR, E Alp Trupchun, 2140 m, 802.700 / 163.450, Barberfalle, 16. 6. – 23.7.2001, leg. B. Lüscher (NMBE). 2 ♀ GR, Passo del Bernina, 2100 – 2300 m, 799 / 143, gesiebt aus Bodenstreu u. Moos, 24.8.2001, leg. C. Germann (cCG). 1 ♀ GR, E Alp Trupchun, Vegetationsmosaik, Seggen, Kurzweide, 2140 m, 802.700 / 163.450, 23. 7. – 17.8.2001 [barber traps], leg. B. Lüscher. 1 ♀ GR, Alp Flix, Malpass, 2422 m, 771.457 / 154.324, Kalk und Silikat, Gesiebe Saxifraga caesia, 20.6.2002, leg. C. Germann (cCG). 2 ♀ 019 _ 06.8 [collecting number] GR, V. Poschiavo, Cavaione, Gesiebe Moos an Felsen im Wald (Larix), 1200 m, 6.10.2006, leg. C. Germann (cCG). 8 ♀ GR, Bergün, Preda oberhalb, 1830 m, Gesiebe Bodenstreu, Moos, 13.6.2008, leg. C. Germann (cCG, NMBE). 1 ♀ 124 _ 10.1 [collecting number] GR, Passo del Bernina, Ospizio Bernina, Giuf, 2290 m, Gesiebe Bodenstreu, 16.9.2010, leg. C. Germann (cCG). 4 ♀ GR, Corn dal Solcun, 2300 m, 24.7.2012, leg. M. Kahlen (TLMF). 1 ♀ OW, Mettental, 2000 m, Blockhalde, 15.11.2005, leg. A. Szallies (cAS). 2 ♀ TI, Generoso (B-Vista), 21.7.1956, Mousses sol [sifted], leg. P. Scherler (NMBE). 1 ♀ TI, Val Calanca, Alp Pianasc, 2100 m, 728 / 142, 24.6.1990, leg. W. Marggi (NMBE). 1 ♀ TI, Quinto, Val Piora, Mottone, 1860 m, 19. – 21.7.2010, leg. E. Knopp (MCSN). 2 ♀ 174 _ 12.3 [collecting number] UR, Isenthal, Brisen, 677.262 / 194.577, 2210 m, 14.6.2012, leg. C. Germann (cCG). 1 ♀ VD, Alpes vaudoises, 27.7.1912, leg. A. Mathey (NMBE). 1 ♀ VD, Jaman, 14.7.1955, leg. P. Scherler (NMBE). 2 ♀ VD, Famelon, Lapiaz, 17.8.1985, Mousses sol [sifted], leg. P. Scherler (NMBE). 1 ♀ 032 _ 07.2 [collecting number] VD, Col des Mosses, Lac Lioson, 1800 m, 29.6.2007, leg. C. Germann (cCG). 2 ♀ VS, Val Ferret, Ban Darray, 9.8.1957, leg. P. Scherler (NMBE). 1 ♀ VS, Grammont, 19.11.1978, Mousses sol [sifted], 19.11.1978, leg. P. Scherler (NMBE). 1 ♀ VS, Gemmipass, Daubensee, unter Stein, 2206 m, A. 7. 1997, leg. C. Germann (cCG). 1 ♀ VS, Leuk, 1700 m, 14. – 21.6.2005 [trap catches], leg. A. Pasche (NMBE). 2 ♀ VS, Chandolin, [without date], leg. A. Rätzer (NMBE). 1 ♀ VS, Val Ferret, [without date], leg. A. Rätzer (NMBE). 1 ♀ VS, Branche d‘en Haut, 1400 m, 16.7.2007, leg. C. Germann (NMBE). 1 ♀ VS, Val Ferret, La Peule, 2460 m, 17.7.2007, leg. C. Germann (cCG). 1 ♀ VS, Col de Balme, vers Tête de Balme, Kalkstein, 2300 m, Gesiebe Moos, Polsterpflanzen, 17.7.2012, leg. C. Germann (NMBE).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE55FF84FF4F9701EE12FC65.taxon	discussion	Remarks: A series of specimens of O. subcostatus from Alp Flix (Switzerland, Grisons) with the following data: 6 ♀ GR, Alp Flix, Salategnas, 1960 m, 24. 7. – 21.8.2005, leg. P. Muff (NMBE), and a single female specimen from Italy, Trentino with the data: Trentino, Gampenjoch, 6. 2004, leg. Grunwald (SDEI) are constantly bigger and more robust, which might indicate the presence of polyploidy, shown in the case of O. carinatopunctatus by Suomalainen (1969).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE56FF84FF2A975BE9CAF9DF.taxon	materials_examined	Type locality: Monte Pasubio Holotype ♀: [Italy] Pasubio Süd-Tirol // Dr. Spaeth // Gredleri Spaethi // azaleae v. Spaethi m // Holotypus Otiorrh. (sic!) azaleae v. spaethi Reitter 1913 // coll. Reitter (HNHM). Italy: 1 ♀ Trentino, Monte Pasubio, Palon, 2200 m, 23.7.1977, unter Stein, leg. M. Kahlen (TLMF). 1 ♂, 2 ♀ Trentino, Vallarsa, Campogrosso, Boale die Fondi, 1500 m, 26.6.1983, unt. Stein, leg. M. Kahlen (TLMF). 4 ♂, 3 ♀ Trentino, Mti. Lessini, Carega, Cma. Mosca SW-Seite, 1950 m, 7.7.1985, an Schneerand, leg. M. Kahlen (TLMF). 1 ♀ Trentino, Lessinische Alpen, Campogrosso, 1250 m, 2.7.1986, leg. L. Dieckmann (SDEI). 1 ♀ [Trentino] Mte. Lessini, Passo di Campogrosso, 1460 m, 1.8.1994, leg. L. Zerche (SDEI). 4 ♀ Trentino, Monti Lessini, Cima Carega, Gipfel, 2250 – 2259 m, 31.5.2003, Dryas - Rasen, leg. M. Kahlen (TLMF). 1 ♂ Trentino, Monti Lessini, Cima Carega, NO Gipfel, 2200 m, Rhododendron - Streu, N 45 ° 43 ’ 29 ” E 11 ° 07 ’ 01 ”, 31.5.2003, leg. L. Zerche (SDEI). 1 ♀ [Trentino / Veneto] Lessini, Cima Posta, 15.8.1972, leg. G. Osella (SDEI). 2 ♀ [Veneto] Aviano, Monte Cavallo, Tremoi, 1850 m, N 46 ° 06 ’ 55.2 ” E 12 ° 29 ’ 24.4 ”, 10.8.2013, leg. J. Kratky (cJK).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF9AFF2A9381E973FD0C.taxon	materials_examined	Type locality: Koralpe	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF9AFF2A9381E973FD0C.taxon	materials_examined	Austria: 1 ♀ Kärnten, Nockberge, WÖllaner Nock, E-Seite, 2100 m, Rasenstreu an Schneerand, 11.6.1998, leg. M. Kahlen (TLMF). 1 ♀ [Styria] Koralpe [without date], leg. E. Klimsch (MHNG). 1 ♂ Styria, Koralpe [without date], leg. V. Gspan (MHNG). 1 ♀ [Styria] Koralpe [without date], leg. Schuster (TLMF). 1 ♀ Koralpe [without date], leg. C. Mandl (TLMF). 2 ♂ Turnau, Styria sept. [without date and collector] (TLMF).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF9AFF2A9381E973FD0C.taxon	materials_examined	Austria: 1 ♀ Osttirol, Lienz-Umgebung, Deutsch Lavant, 4.5.1990, leg. A. Kofler (TLMF). Italy: 4 ♀ [Veneto] Udine, Mt. Sernio, Forc. Nuviernulis, S-Seite, 1700 m, 16.7.1988, Dryasrasen, leg. M. Kahlen (TLMF). 1 ♀ [Veneto] Udine, Mt. Sernio, Forcella Nuviernulis, 1732 m, 23.9.1998, unt. Saxifraga caesia, leg. M. Kahlen (TLMF).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF98FF2A91D1E825FBB8.taxon	materials_examined	Type locality: Cima Tombea	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF98FF2A91D1E825FBB8.taxon	materials_examined	Italy: 1 ♂, 1 ♀ [Lombardia] Brescia, Cima Tombea, S-Hang, 1900 – 1950 m, 12.7.1978, unter Stein, leg. M. Kahlen (TLMF). 1 ♂, 2 ♀ Südtirol, Mte. Pari, 20.6.1936, Rasen, leg. Pechlaner (TLFM). 1 ♀ [Trentino] Val Scaglia, Ledrosee [without date], coll. Reiss (TLFM). 2 ♂ Trentino, Cima Pari, Gipfel, 15.7.1978, Rasengesiebe, leg. M. Kahlen (TLMF). 1 ♀ [Trentino] Verona, Monte Baldo, Cima Telegrafo, 2000 – 2150 m, 13.7.1979, u. Stein, leg. M. Kahlen (TLMF). 1 ♂ [Trentino] Verona, Monte Baldo, Cima Valdritta, Gipfel, 2200 m, Nachts am Felsen, 18.6.1983, leg. M. Kahlen (TLMF). 1 ♀ Trentino, Judikarien, Mt. Tombea, unter Dryas - Polster, 1900 m, 4.7.1986, leg. L. Dieckmann (SDEI).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF98FF2A91D1E825FBB8.taxon	materials_examined	Switzerland: 3 ♀ GR, Ofenpass, 20.8.1969, leg. P. Scherler (NMBE). 3 ♀ GR, Ofenpass, 19.7.1973, Mousses sol [sifted] leg. P. Scherler (NMBE). 2 ♀ GR, Munt La Schera, 23.7.1977, leg. Dethier, coll. P. Scherler (NMBE). 2 ♀ GR, Munt La Schera, 17.9.1978, leg. P. Scherler (NMBE). 1 ♀ GR, E Alp Trupchun, Vegetationsmosaik, Seggen / Kurzweide, 2140 m, 16. 6. – 23.7.2001 [pitfall-traps], leg. B. Lüscher (cCG). 4 ♀ 038 _ 07.6 [collecting number} GR, Valbella, W Funtana da S-charl, Kalk, u. Steinen, 821.122 / 170.777, 2450 m, 14.9.2007, leg. C. Germann (cCG). 2 ♀ 149 _ 11.4 GR, Ftan, Piz Clünas, S-exp. Hang, 2500 – 2700 m, GS Pflanzenstreu Felsbänder, 10.8.2011, leg. C. Germann (cCG). 11 ♀ GR, Ftan, Piz Clünas, 814.100 / 188.854, S-exp. Hang, 2560 m, Gesiebe Felssimse Flechten & Polsterpflanzen, 12.8.2011, leg. C. Germann (cCG). 8 ♀ 199 _ 13.7 (collecting number) GR, Val Mora, unterh. Piz dal DÖss Radond, 823.323 / 162.634, 2500 m, GS Moos, Polsterpflanzen, Kalkstein, 29.6.2013, leg. C. Germann (cCG, NML).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE48FF98FF2A91D1E825FBB8.taxon	discussion	Remark: The type specimen (s) were not in the Daniel collections (ZSM), nor in Gredler’s collection (Franziskanergymnasium Bozen), and hence could unfortunately not be examined.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE4AFF9FFF2A9791E92CFBB8.taxon	materials_examined	Type locality: Tirol, FranzenshÖhe France: 1 ♀ Dauphiné, Col Lautaret, coll. J. Breit (NMBA). Italy: 1 ex. Lombardia, Tartano (SO): Valle di Lemma, 2100 m, 28.6.1993, leg. Perego, coll. Museo civico Storia Naturale di Morbegno (data taken from Pedroni 2013). 1 ♀ Piemonte, Coazze, fine VIII. 1898, leg. A. Dodero (NMBA). 1 ♂, 1 ♀ Piemonte, Ceresole Reale, VIII. 1900, leg. A. Dodero (NMBA). 1 ♂, 1 ♀ [Piemonte] Albergian, 8.7.1905, leg. J. Daniel (NMBA, MHNG). 1 ♀ [Piemonte] Alpe Cozie, Pragelato, 23.7.1922, leg. Botto (MHNG). 1 ♀ [Piemonte] Graje, Ceresole, 22.7.1935, leg. Tasso, Schatzmayer & Koch (NMBA). 1 ♀ [Piemonte] M. Cenisio, 1700 m, 25.7.1935, leg. Schatzmayer & Koch (NMBA). 2 ♀ [Piemonte] Cuneo, Monte Viso, oberhalb Piano de Re, N 44 ° 41 ’ 35 ” E 7 ° 05 ’ 41 ”, 2150 m, 9.7.2003, leg. C. Germann (cCG).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE4AFF9FFF2A9791E92CFBB8.taxon	discussion	Remark: The type specimen (s) were not in Rosenhauer's collection in the ZSM and hence could not be examined. The localities in the West (Alpes Maritimes) and those in the East (Trentino) are quite separate from each other, although recent records from Lombardia (Pedroni 2013) may explain this gap simply with presently missing records in-between.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE4DFF9FFF4F9791E831F981.taxon	materials_examined	Type locality: Alpes Maritimes France: 1 ♀ Alpes Maritimes, Castérino NW, Valmasque, Vastière des Dragons, 2000 m, 23.6.2009, Moos u. Rhododendron - Streu, leg. M. Kahlen (TLMF). 1 ♀ Alpes Maritimes, Vallée du Boréon, Talhang, Chalet Vidron SW, 1760 m, 26.6.2010, Nachts von Vaccinium myrtillus gestreift, leg. M. Kahlen (TLMF). 2 ♀ Alpes Maritimes, Camp Soubran, Le Boréon NW, 2300 m, 27.6.2010, Zwergstrauchheide, Rhododendron - Streu, leg. M. Kahlen (TLMF). Italy: 1 ♀ [Piemonte] Lago Brocan, Seealpen, leg. Dr. K. Singer (NMBA). 1 ♀ [Piemonte] Alpi Marittime, Val Meira, 1.6.1898, coll. StÖcklein (NMBA). 2 ♀ [Piemonte] Cuneo, Entraque SW, Piano del Praiet, 1800 m, 4.6.2009, Vaccinium - und Rhododendron - Streu, leg. M. Kahlen (TLMF).	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE4CFF9EFF2A965CE8BFF995.taxon	discussion	It was Roudier (1961) himself who already mentioned that his new species belongs to Nihus – by some authors regarded as synonym to Eunihus in the past. The type locality of O. levasseuri, Glacier du Canigou, is situated about 60 km E of the before mentioned locality.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
03AB87D4CE4CFF9EFF2B9381EB4DFCD8.taxon	discussion	Based on the key by Reitter (1913) and subsequently the one by Frieser (1981) to the taxa of Nihus, both O. azaleae and O. gredleri were mostly misinterpreted. One important detail, the shape of the eyes which is initially characterized by Daniel & Daniel (1898) in their description as " … stark gewÖlbte Augen ", followed by Reitter (1913) as " … selten sehr schwach vortretend v. Gredleri Dan. " and finally termed by Frieser (1981) as " … Augen wie bei azaleae … " (Whereas nothing is stated concerning the eyes in the case of azaleae …). This may have led to the repeated quotation of O. azaleae for Switzerland, initiated by the first record by Linder (1968) and followed by all subsequent authors (Germann & Lüscher 2007, Germann 2010 a, 2012, 2014 b, Magnano & Alonso-Zarazaga 2013). The present investigations, with a new key to this subgenus replacing those proposed by Reitter (1913) and Frieser (1981), showed that all Swiss specimens belong to O. gredleri, presently known from Italy (Trentino-Alto Adige) and Austria (Nordtirol). Contrary to Magnano & Alonso-Zarazaga (2013) no reliable records from Germany are presently known (P. Sprick, pers. comm.), although records from Austria (Ausserfern) are very close to Bavaria.	en	Germann, Christoph (2014): On the identity of Otiorhynchus (Nihus) subcostatus STIERLIN, 1866, description of Otiorhynchus muffi sp. nov., and an illustrated key to the species of Nihus REITTER, 1912 with taxonomic comments on both the subgenera Nihus and Eunihus REITTER, 1912 (Coleoptera, Curculionidae). Contributions to Natural History 25: 45-69, DOI: 10.5169/seals-787044
