taxonID	type	description	language	source
015D8E70FF9EFF92FDFF0054FADBBFF7.taxon	type_taxon	Type genus: Pseudocethera Villiers, 1963 (= Voconia Stål, 1866). Reduviinae Latreille, 1807: 126 – 127 (in part, new concept).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9EFF92FDFF0054FADBBFF7.taxon	diagnosis	Diagnosis Distinguished from other Reduviidae by the combination of the following characters: head and pronotum pubescent with interspersed macrosetae or setigerous tubercles; labial segment II straight, as long as remaining segments, and adpressed to head (Fig. 3 E – F); apex of the third visible labial segment bent and slender; antennal insertion near eye, medially in lateral view (Fig. 3 E – F); pedicel longer than scape; prosternum drawn into an anteriad-directed process (Fig. 6); absence of dorsal abdominal glands on terga IV – VI; ventral surface of all femora with anterior and posterior rows of spine-like protuberances with macrosetae at base of protuberance (Fig. 4 F, I); males with fossula spongiosa on fore and mid legs (except in V. motoensis sp. nov. where the the fossula spongiosa is absent on the midleg) (Fig. 4 G – H); females with fossula spongiosa on fore leg, but fossula spongiosa usually absent or vestigial on mid leg; tibiae laterally compressed (Fig. 4 H); three-segmented tarsi (Fig. 4 H).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9EFF92FDFF0054FADBBFF7.taxon	description	Redescription Usually macropterous, only micropterous morphs are known in three species and one species includes both micropterous and macropterous morphs. BODY LENGTH. 6.7 – 12.4 mm. BODY. Either robust (Fig. 8, e. g., V. motoensis sp. nov.) or slender (Fig. 8, e. g., V. minima sp. nov.) with variable color patterns. HEAD. Elongate or globose; pubescent with interspersed macrosetae or setigerous tubercles; interocular region with two glabrous lines; scape and pedicel with dense, short macrosetae; scape reaching or slightly surpassing head apex, 0.2 – 0.3 times length of pedicel; ocelli present in macropterous morphs and minute or absent in micropterous morphs; buccula with ventrolateral swelling; labial segment III drawn into bent, slender apex. THORAX. Prosternum in lateral view drawn into anteriad-directed process; scutellar spine long, either sub-horizontal or raised. HEMELYTRON. In macropterous individuals, Cu-An 1 cell triangular, shorter than M-Cu cell (Fig. 4 B) or Cu-An 1 cell similar in length to M-Cu cell, slender and tapered apically (Fig. 4 A). LEGS. Fossula spongiosa present on fore tibia, usually present on mid tibia of males; all femora with ventral anterior and posterior rows of spine-like protuberances with macrosetae at base of protuberance (Fig. 4 I); tibiae laterally compressed (Fig. 4 G – H). ABDOMEN. Posterolateral margin of dorsal laterotergites with (Fig. 5, e. g., Gerbelius sp. 2) or without single macroseta; anterior margin of terga carinulate; dorsal abdominal glands absent. Male PYGOPHORE. Posterior margin with a short median apical process, either straight or bent posteriorly. PARAMERES. Sinusoidal or rounded. AEDEAGUS. Basal plate extension long in relation to phallotheca; endosoma covered in spicules. Habitat and collecting method Pseudocetherinae have been collected using light traps, but records on habitat and collecting methods are scarce on specimen labels.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9EFF92FDFF0054FADBBFF7.taxon	distribution	Distribution (Figs 17 – 20) Pseudocetherinae are widespread, with species distributed in the Afrotropical, Neotropical, Oriental, and Australasian regions. The highest diversity occurs in Southeast Asia (Fig. 20) and the lowest in the Neotropics (Fig. 19).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	description	Figs 1, 2, 3 A, G, 4 A, 5, 11, 14, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	type_taxon	Type species: Gerbelius typicus Distant, 1903, by original designation.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	diagnosis	Revised diagnosis Distinguished from the other genus in the subfamily, Voconia, by the smooth or corrugated surface of the head and pronotum (Fig. 3 G); laterally flattened shape of the maxillary plates (Fig. 3 A); long, basally adjacent mandibular plates (Fig. 3 A); stout labium densely covered with long macrosetae on the morphologically ventral surface of labial segment I (Fig. 3 G); morphologically dorsal surface of labial segment II 0.8 – 0.9 times length of labial segment I (Fig. 3 G); and Cu-An 1 cell slender and tapering apically, similar in size to the M-Cu cell (Fig. 4 A).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	description	Redescription BODY LENGTH. 7.0 – 8.0 mm. BODY. Robust; only macropterous morphs known (Fig. 5). COLORATION. Variable color patterns; scutellum uniform or with contrasting yellow scutellar spine. INTEGUMENT AND VESTITURE. Head and pronotum: smooth to corrugated with dense pubescence and sparse, long macrosetae; antennifer with macroseta, without lateral protuberance (Fig. 3 A); labium with dense, long macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae. Hemelytron: corium with dense setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and three or fewer small spines. Abdomen: ventral surface pubescent with long macrosetae. HEAD (Fig. 3 A, G). Elongate, 1.2 – 1.3 times as long as wide; anteocular region about a third of head length and shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view globose and longer than eye; pedicel 1.1 – 1.3 times length of head width; mandibular plates in dorsal view long with basal half adjacent; maxillary plates in dorsal view enlarged laterally forming flattened projections, adjacent to clypeus; clypeus width in dorsal view narrower than maxillary plates; interocular glabrous markings V-shaped, joined medially at interocular sulcus; eye width in dorsal view narrower than synthlipsis, eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labium stout; second (first visible) labial segment in lateral view surpassing posteroventral eye margin, gradually widened apically; dorsal surface of third (second visible) labial segment convex, 0.8 – 0.9 times length of second segment; fourth (third visible) labial segment drawn into bent, slender apex. THORAX. Pronotal collar narrow medially with anterolateral angles short and flared laterally; anterior pronotal lobe lateral margins rounded; scutellum width 0.8 times length; scutellar spine subhorizontal; anteriad-directed process on prosternum with paramedial lobes; anterior margin of stridulitrum projected into sharp spine; proepimeron with obtuse protuberance on posteroventral margin. HEMELYTRON (Fig. 4 A). Cu-An 1 cell similar in size to M-Cu cell; Cu-An 1 cell slender, tapered apically. LEGS. Fossula spongiosa present on fore and mid legs in males, present on fore but absent or vestigial on mid legs of females. ABDOMEN. Anterior margin of terga weakly carinulate; terga II and III with paired, prominent, and longitudinal carinae, almost reaching posterior margin of tergum III. PYGOPHORE (Fig. 11). Transverse bridge rounded on posterior margin; lateral pygophore margin flat; paramere shape in dorsal view sinusoidal, apex tapered into squarely round tip. AEDEAGUS (Fig. 15). Endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite rounded in dorsal view; basal plate extension 4.4 – 4.8 times as long as wide.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	distribution	Distribution This genus currently comprises two described species from India and Southeast Asia, and we are aware of at least four undescribed species distributed across India, Laos, Thailand, and Vietnam. Gerbelius typicus is the most widespread among them.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF9DFF9DFD8406EDFE13B88F.taxon	discussion	Remarks Photographs of type specimens in their unit trays at the BMNH were used to evaluate species concepts for G. typicus and G. confluens. This genus previously had a third described species that is now transferred to Voconia, V. ornata. Molecular phylogenetics recovered a Gerbelius clade (Gerbelius sp. 3 + [Gerbelius sp. 1 + Gerbelius sp. 4]) sister to Voconia wegneri sp. nov. (Hwang & Weirauch, 2012). We also recovered a well-supported Gerbelius clade sister to Voconia (Figs 1 – 2). The pygophore of two specimens were dissected for this study (Figs 11, 14); however, the median apical process of G. typicus (AMNH _ IZC 00321200) broke off during dissection (Fig. 11). We decided not to describe new species of Gerbelius as part of this study. Since species of Gerbelius somewhat resemble other small-bodied assassin bugs currently classified as Reduviinae, we suspect that our specimen searches in various natural history collections were not comprehensive for this genus. However, this material should be included in a future revision of this genus.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF92FF9DFDA901C7FC1DB9D6.taxon	materials_examined	Type material Lectotype (present designation) MALAYSIA [“ SIAMESE MALAY STATES ”] • ♀; Terengganu, Bukit Besar; 1903; Annandale and Robinson leg.; USI: UCR _ ENT 00048383; BMNH (E) 1255221.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF92FF9DFDB9070EFA86BCBD.taxon	materials_examined	Type material Lectotype (present designation) MYANMAR [“ BIRMANIA ”] • ♀; Bhamo; Aug. 1885; L. Fea leg.; Distant coll. 1911 - 383; USI: UCR _ ENT 00048382; BMNH (E) 1255220. Paralectotype MYANMAR • 1 ♀; “ Palon (Pegu) ” [Palon]; Aug. – Sep. “ 87 ” [1887]; L. Fea leg.; Distant coll. 1911 - 383; BMNH. Additional material examined INDIA • 1 ♂; Kerala, Trivandrum, Poonmudi Range [Ponmudi]; [8.77 ° N, 77.12 ° E]; elev. “ 3000 ft ” [914 m]; Sep. 1971; T. R. S. Nathan leg.; dissected genitalia in vial; USI: AMNH _ IZC 00321200.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF92FF9EFD5805B8FB79BB54.taxon	materials_examined	Material examined LAOS • 1 ♀; Vientiane Prov., Houayang Nat. Park; 18.09815 ° N, 102.67535 ° E; elev. 180 m; 19 – 20 Jun. 2008; A. Solodovnikov and J. Pedersen leg.; collecting event: LAO 08 - 2 G; mostly secondary rainforest; DNA voucher specimen R _ CW 0705; USI: UCR _ ENT 00052189; UCR. THAILAND • 1 ♀; Chaiyaphum, Pa Hin Ngam; 15.63553 ° N, 101.3987 ° E; elev. 698 m; 11 – 18 Aug. 2006; Katae Sa-nog and Buakaw Adnafai leg.; collecting event: T 444; ecotone between mix deciduous / dry dipterocarp; DNA voucher specimen R _ CW 1185; USI: UCR _ ENT 00002541; UCR.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9EFD58028FFC7BB802.taxon	materials_examined	Material examined VIETNAM • 1 ♂; Gia Lai, 20 km N of Pleiku; [14.15 ° N, 107.95 ° E]; elev. 650 m; 9 May 1960; S. Quate leg.; dissected genitalia in vial; USI: UCR _ ENT 00073816; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9EFD58017AFD5DB90B.taxon	materials_examined	Material examined LAOS • 1 ♀; Vientiane Prov., Phou Khao Khouay; 18.33948 ° N, 102.80871 ° E; elev. 750 m; 26 – 31 May 2008; A. Solodovnikov and J. Pedersen leg.; collecting event: LAO 08 - 7 b; DNA voucher specimen R _ CW 0704; USI: UCR _ ENT 00052219; UCR.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9EFD580042FCC5BE13.taxon	materials_examined	Material examined THAILAND • 1 ♀; Phetchabun Prov., Nam Nao National Park; 16.728217 ° N, 101.563933 ° E; elev. 917 m; 6 – 13 Nov. 2006; Leng Janteab leg.; collecting event: T 1018; malaise trap; DNA voucher specimen R _ CW 0552; USI: UCR _ ENT 00052225; UCR.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	description	Figs 1 – 4, 6 – 14, 17 – 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	type_taxon	Type species: Voconia pallidipes Stål, 1866, by subsequent monotypy (Stål 1866 b: 165). Gender feminine.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	type_taxon	Type species: Microvarus conradti Jeannel, 1917, by original designation. syn. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	type_taxon	Type species: Kayanocoris wegneri Miller, 1954, by original designation. syn. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	type_taxon	Type species: Paragerbelius typicus Miller, 1958, by original designation. syn. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	type_taxon	Type species: Pseudocethera monodi Villiers, 1963, by original designation. syn. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	diagnosis	Diagnosis Distinguished from the other genus in the subfamily, Gerbelius, by the granulose integument of the head and pronotum (Fig. 6 A, D), ellipsoid shape of the maxillary plates (Fig. 6 B, blue), short non-adjacent mandibular plates (Fig. 6 B, green), gracile labium with short macrosetae on the morphologically ventral surface of labial segment I (Fig. 6 E), and the Cu-An 1 cell of the fore wing shorter than the M-Cu cell and triangular (Fig. 6 C).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	description	Redescription BODY LENGTH. 6.7 – 12.4 mm. BODY. Slender or robust; macropterous or micropterous (Figs 7 – 10). COLORATION. Thorax: scutellum uniform or with contrasting yellow scutellar spine. Hemelytron (abducted): corium with various combinations of yellow spots on the proximal, anteromedial, posteromedial, and distal areas, with medial yellow stripe, or uniformly yellow; membrane uniformly dark or with pale V-shaped marking along R and M veins. INTEGUMENT AND VESTITURE. Head and pronotum: granulose with pubescence and interspersed macrosetae or setigerous tubercles; interocular region with two glabrous lines; antennifer with macroseta, with or without lateral protuberance; morphologically ventral surface of labial segment I with short macrosetae or sparse long macrosetae. Thorax: anterolateral angles of pronotal collar armed with macrosetae or setigerous tubercles. Legs: posterior row of spine-like protuberances on all femora well developed, anterior row poorly developed on ventral surface of mid and hind femora. Abdomen: ventral surface pubescent with long macrosetae. STRUCTURE. Head: globose to elongate; mandibular plates in dorsal view short, not adjacent basally; maxillary plates in dorsal view elongate anteriorly, forming ellipsoid projections; pedicel 0.6 – 1.9 times length of head width; ventrolateral swelling of buccula with or without lateral protrusion that surpasses buccular margin; labium gracile; morphologically dorsal surface of labial segment II 0.2 – 0.7 times length of segment I; labial segment III drawn into bent, slender apex. Thorax: scutellum width 0.5 – 0.8 times length; scutellar spine subhorizontal or raised; proepimeron without or with acute protuberance on posteroventral margin. Hemelytron: Cu-An 1 cell triangular, smaller than M-Cu cell. Legs: fossula spongiosa present on fore and mid legs in males, either present on fore and mid legs or absent on mid legs of females.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	distribution	Distribution This genus currently comprises nine described species and 23 new species with a circumtropical distribution. The lowest known species diversity is in the Neotropics, and the highest species diversity for this genus is in Southeast Asia across India, Indonesia, Laos, Brunei, and the Philippines.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF91FF9BFD66074AFADBBF52.taxon	discussion	Remarks Originally a monotypic genus. It is here expanded to include numerous new species as well as several previously described species formerly placed in Reduviinae.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	description	urn: lsid: zoobank. org: act: 21 FEECCD- 754 F- 4 DF 8 - 815 B-E 18666 CE 9 CE 5 Figs 1 – 2, 7, 9, 11, 14, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	diagnosis	Diagnosis This species most closely resembles two other dark-brown Philippine species, V. minima sp. nov. and V. nyx sp. nov., with yellow posteromedial yellow spots on the abducted corium, but is differentiated by the yellow patches on the dorsal laterotergites, the faint yellow spot on the anteromedial portion of the abducted corium, the presence of small paramedial lobes on the anteriad-directed process of the prosternum, and the stridulitrum being elongated anteriorly into a small protuberance. It specifically differs from V. minima sp. nov. by the uniformly dark legs and short labial segment I that does not reach the posteroventral eye margin and from V. nyx sp. nov. by the uniformly dark pronotum.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	etymology	Etymology Named after Austin Baker, a former colleague at UCR who provided support and insightful comments on this revision.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂ (dissected pygophore and aedeagus in vial); Mindanao, Mts Or., Mt Pomalihi, Gingoog City, 21 km W of Gingoog City; [8.81 ° N, 124.92 ° E]; elev. 800 – 1000 m; 16 Oct. 1965; H. M. Torrevillas leg.; light trap; USI: UCR _ ENT 00073810; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 9.8 mm; macropterous. COLORATION. Head: dark brown with paired pale patches adjacent to lateral ocellar margin; labium nearly as dark as head. Thorax: coloration as head; scutellum uniformly dark brown. Hemelytron (abducted): clavus dark reddish-brown, distal half yellow; corium dark reddish-brown with anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale, remainder as membrane. Legs: dark brown. Abdomen: dark reddish-brown, almost black; dorsal laterotergites dark with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with macroseta, base without lateral protuberance; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large and two small spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.2 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins in dorsal view gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view converging; clypeus in dorsal view wider than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: pronotal collar narrow medially with anterolateral angles short and flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum deep, elongated longitudinally; scutellar spine long, raised; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate; terga II and III with paired prominent longitudinal carinae, reaching about two-thirds of tergum III. Pygophore (Fig. 11): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongue-shaped in dorsal view; basal plate extension about 4.3 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in the Philippines. Based on the record of a Miridae Hahn, 1831 specimen with the same collection event, Mt Pomalihi is presumably in Mt Balatukan Range Natural Park. The type locality is also shared with V. isosceles sp. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FF94FFA6FD690689FA89BCF4.taxon	discussion	Remarks The median apical process broke off during dissection (Fig. 11). It was presumably short as is the case in all Pseudocetherinae, but it was not possible to determine if it was upright or bent posteriorly.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	description	urn: lsid: zoobank. org: act: 32 D 3 A 427 - 888 D- 4 D 48 - B 0 C 3 - 5 EB 69 CB 43 B 1 A Figs 1, 2, 3 J, 4 C, K, 7, 9, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	diagnosis	Diagnosis Distinguished from other species of the genus Voconia by its long, slender body (about 10.8 mm), long head (about 1.7 times as long as wide), hind leg with four large spines in the posterior row, a large pale spot between R and M veins, and dorsal laterotergites I and II yellow, the remainder dark brown. This species is most similar to V. typica, but is differentiated by the yellow proximal third of the mid and hind femora as well as the anterior pronotal lobe, which is convex, strongly curved inward on the posterolateral margins.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ bracatus, - a, - um ’, meaning ‘ wearing trousers’. It refers to the yellow coloration of the proximal half of the femora.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	materials_examined	Type material Holotype PAPUA NEW GUINEA • ♂; Milne Bay Province, Waikaiuna, Normanby Island; [10.06 ° S, 150.97 ° E]; elev. 0 – 50 m; 14 Apr. 1956; L. J. Brass leg.; Fifth Archibold Exped. to New Guinea; USI: AMNH _ PBI 00170707; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 10.8 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin and paired pale patches adjacent to lateral ocellar margin; labial segments II and III lighter than head. Thorax: coloration as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark reddish-brown with distal half yellow; corium dark reddish-brown with anteromedial, posteromedial, and distal yellow spots; membrane dark, large pale spot between R and M veins; membranal veins R and M proximally pale, remainder as membrane. Legs (Fig. 4 K): dark brown with yellow mid and hind coxae, trochanter, and proximal third of femora. Abdomen (Fig. 4 C): dark reddishbrown; dorsal laterotergites I and II yellow. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among dense pubescence; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae; anterolateral angles of pronotal collar without macrosetae or setigerous tubercles. Legs (Fig. 4 K): posterior row of protuberances on mid and hind femora with four large spines on distal half. Abdomen: ventral surface pubescent with long macrosetae. STRUCTURE. Head: elongate, about 1.7 times as long as wide; anteocular region about one third of head length, subequal in length to postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel about 1.8 times length of head width; maxillary plates ellipsoidal, adjacent to and reaching apex of clypeus; apex of maxillary plates in dorsal view directed straight; clypeus in dorsal view slightly narrower than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view slightly bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II slightly curving apically, about 0.4 times length of segment I. Thorax: pronotal collar narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe subequal in length to posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum deep and elongated transversely; scutellar spine long and subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes (Fig. 3 J); anterior margin of stridulitrum not elongated into protuberance (Fig. 3 J); proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	distribution	Distribution (Fig. 18) This species is only known from the type locality of Normanby Island off of Papua New Guinea, where no other Pseudocetherinae species are known.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA9FFA0FD6005EFFE91B851.taxon	discussion	Remarks This species forms a well-supported clade with another species from Papua New Guinea, V. typica (Figs 1 – 2).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	description	urn: lsid: zoobank. org: act: AB 497 C 37 - 1 CBA- 4270 - AAE 0 - DDA 9 ADDCAB 25 Figs 1 – 2, 7, 9, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	diagnosis	Diagnosis Recognized by its Australasian distribution, small size (about 7.4 mm), globose head (about as long as wide), labial segment II swollen ventromedially, and coarsely granulose, tuberculate head and pronotum. Most closely resembles V. dolichocephala sp. nov. in size and hemelytral coloration but V. dolichocephala sp. nov. has a conspicuously elongate head.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	etymology	Etymology The specific epithet is the latinized Greek adjective ‘ brachycephalus, - a, - um ’, formed from the prefix ‘ brachy- ’ (meaning ‘ short’) and the noun ‘ cephale ’ (meaning ‘ head’), referring to the fact that this species has the shortest head among species of Voconia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	materials_examined	Type material Holotype PAPUA NEW GUINEA [“ N. GUINEA ”] • ♀; Madang Province, Erima; [5.42 ° S, 145.73 ° E]; 1896; L. Bíró leg.; “ Astrolabe B. ”; USI: AMNH _ PBI 00213935; HNHM. Material examined from literature AUSTRALIA • 1 ♀; Northern Territory, 2 km E of Saddle Creek; 15.95694 ° S, 129.56167 ° E; elev. 75 m; 27 Sep. 1983; E. L. Schlinger and M. E. Irwin leg.; INHS 914,917.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	description	Description Female (Figs 7, 9) BODY LENGTH. About 7.4 mm; macropterous. COLORATION. Head: dark brown, postocular region with dark patches adjacent to medial ocellar margin; labium lighter than head. Thorax: coloration as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium reddish-brown with anteromedial and posteromedial yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale, remainder as membrane. Legs: dark yellowishbrown. Abdomen: dark brown; dorsal laterotergites uniformly dark. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose, with dense, short setigerous tubercles interspersed among sparse pubescence; interocular region with two pairs of stout setigerous tubercles paramedially and one medially; antennifer with two adjacent lateral setigerous tubercles; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with short macrosetae on setigerous tubercles. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and three or fewer small spines. Abdomen: ventral surface pubescent with long macrosetae. STRUCTURE. Head: globose, about as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and shorter than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view wider than maxillary plates; clypeal apex bifid, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view gradually thickens distally, surpassing posteroventral eye margin; labial segment II swollen ventromedially; morphologically dorsal surface of labial segment II curves ventrally, about 0.3 times length of segment I. Thorax: pronotal collar narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long and raised; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa of fore leg present. Male Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	distribution	Distribution (Fig. 18) This species has a disjunct distribution. It is known from Erima in Papua New Guinea (type locality) and the Northern Territory of Australia (Swanson 2015: 268, as V. pallidipes).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFAFFFA2FDBB018CFAD4B9D6.taxon	discussion	Remarks There is little doubt that the specimen figured by Swanson (2015: 269, fig. 10) as V. pallidipes pertains to V. brachycephala sp. nov. Although the size of the specimen in concern is unknown, its coloration and the short, tuberculate head do not seem to differ from the condition seen in the new species. Since the specimen could not be re-examined, it is, however, not included in the type material of the new species.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	description	urn: lsid: zoobank. org: act: 79 E 22180 - F 528 - 4166 - 8 D 25 - 02 AD 3 BA 726 BE Figs 1 – 2, 7, 9, 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	diagnosis	Diagnosis Recognized from other African Voconia species by the unique coloration of the corium, abdomen, and legs which are entirely yellow. Additionally, the posterior row of spines on the mid and hind femora have three large and many small spines, and the antennifer lacks a lateral protuberance. Females possess a fossula spongiosa on the mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ chrysopterus, - a, - um ’, meaning ‘ golden-winged’, referring to the entirely yellow corium.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	materials_examined	Type material Holotype KENYA • ♀; Eastern Province, Mwingi, Nguni; 7 Apr. 2004; M. Snizek leg.; USI: UCR _ ENT 00001551; NHMW.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	description	Description Female (Figs 7, 9) BODY LENGTH. 10.5 mm; macropterous. COLORATION. Head: dark brown, maxillary plates lighter; postocular region dark brown with paired pale patches adjacent to lateral ocellar margin; labium lighter than head, yellow. Thorax: coloration as head, with posterior pronotal lobe lighter than anterior lobe; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus and corium uniformly yellow; membrane uniformly dark; membranal veins as remainder of membrane. Legs: yellow. Abdomen: ventral surface and dorsal laterotergites yellow. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with sparse pubescence and sparse, short macrosetae; interocular region with two pairs of macrosetae paramedially; antennifer without lateral projection; labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles; scutellar lateral carinae with short macrosetae. Hemelytron: corium with dense short setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and many small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.3 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel about 1.8 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view wider than maxillary plates, apex rounded, not narrowed; interocular glabrous markings curved anterolaterally, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis, reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin, setigerous tubercles along apical margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: with pronotal collar narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.5 times length of posterior pronotal lobe, lateral margins rounded, glabrous markings not depressed; median apodeme depression of pronotum shallow and elongated longitudinally; scutellar spine long and raised; anteriad-directed process of prosternum smoothly rounded without paramedial lobes; anterior margin of stridulitrum projecting into a small protuberance. Legs: fossula spongiosa present on fore and mid legs. Male Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	distribution	Distribution (Fig. 17) This species is only known from the type locality in Mwingi, Kenya. It is the easternmost Afrotropical species.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFADFFACFD8A070DFB1AB9FA.taxon	discussion	Remarks This is the only female in the subfamily with fossula spongiosa present on the mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	description	Figs 1 – 2, 7, 9, 11, 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	diagnosis	Diagnosis Distinguished from other species in Voconia by its Afrotropical distribution and stout body. This species most closely resembles V. motoensis and V. smithae sp. nov. It differs from V. motoensis by its distinct coloration: yellow scutellar spine, corium with a proximal and distal yellow spot, and contrasting yellow anterior spots on dark dorsal laterotergites. Despite the strong superficial resemblance to V. smithae sp. nov. due to its coloration and stoutness, closer examination reveals that V. conradti is distinguishable by the longer body (10.4 – 11.4 mm), dark dorsal laterotergites with strongly contrasting yellow anterior spots, finely granulose head, antennifer with short or absent laterally projecting setigerous tubercle, lack of paired interocular setigerous tubercles, maxillary plates with medial margins diverging, shorter anteocular region than postocular region (measured to where the posterior margin of the granulations meet the anterior margin of the smooth neck), buccula without lateral protrusion, and strongly bulging eyes that greatly surpass the postocular lateral margin in dorsal view and reach the ventral head margin in lateral view.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	materials_examined	Type material Lectotype (present designation) EQUATORIAL GUINEA • ♂; “ Fernando-Po ” [Bioko Island]; 1901; L. Conradt leg.; MNHN _ EH 24709. Additional material examined CAMEROON • 1 ♀; N Kolbisson; [3.87 ° N, 11.45 ° E]; 27 Jun. 1966; B. de Mire leg.; USI: AMNH _ PBI 00170693; AMNH • 1 ♂; same collection data as for preceding; 31 Mar. 1971; USI: AMNH _ PBI 00170694; AMNH. IVORY COAST • 1 ♂; Foro Foro; [7.98 ° N, 5.06 ° W]; 13 Oct. 1971; D. Duviard leg.; USI: AMNH _ PBI 00170691; AMNH • 1 ♂; same locality as for preceding; 25 – 28 Sep. 1973; R. Linnavuori leg.; dissected pygophore in vial (Fig. 11), aedeagus lost; USI: AMNH _ PBI 00170695; AMNH. NIGERIA • 1 ♂; W State, Ife; [7.47 ° N, 4.56 ° E]; 7 Jul. – 14 Aug. 1973; R. Linnavuori leg.; USI: AMNH _ PBI 00170692; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	description	Redescription Male (Figs 7, 9) BODY LENGTH. 10.4 – 11.4 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin and pale patches adjacent to lateral ocellar margin; labium lighter than head. Thorax: coloration as head; pronotum uniformly dark (Fig. 3 K) or posterior pronotal lobe lighter than anterior lobe; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish-brown with proximal, anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: completely yellow or yellow with distal half of femur dark brown. Abdomen: dark brown, dorsal laterotergites with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax (Fig. 3 K): anterolateral angles of pronotal collar without macrosetae or setigerous tubercles. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.1 – 1.4 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye or shorter, lateral margins subrectangular; pedicel about 1.3 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view with medial margins diverging; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view either wider or narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax (Fig. 3 K): pronotal collar in dorsal view narrow medially, with anterolateral angles short, projected forward; anterior pronotal lobe 0.5 – 0.6 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum slightly depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, raised; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate; terga II and III with paired prominent longitudinal carinae reaching about one-third of tergum III. Pygophore (Fig. 11): transverse bridge with triangular posterior margin; posterior region of ventral surface of pygophore in lateral view with slight swelling; median apical process in caudal view short; direction of median apical process in lateral view sharply bent posteriorly; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Female Mostly as male but differs from males by the following. Body length: about 11.2 mm. Head: elongate, about 1.4 times as long as wide; pedicel about 1.2 times length of head width; ocelli minute, occupying 0.15 times length of postocular region. Legs: fossula absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	distribution	Distribution (Fig. 17) This species is distributed in Western Africa. The type locality is from Bioko, an island off the west coast, north of Equatorial Guinea.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA3FFAFFDF700F0FE92B9BB.taxon	discussion	Remarks Originally described as Microvarus conradti based on several specimens which are to be treated as syntypes (Jeannel 1917). The type material of M. conradti is deposited at the MNHN under loan restrictions; habitus photographs of one of the syntypes (Fig. 7; USI: MNHN _ EH 24709) were used to evaluate this species concept and this specimen is designated as the lectotype. The synonymy of M. achteni proposed by Villiers (1948) is accepted here without re-examining the type material of Schouteden (1929). This species is transferred to Voconia based on the cladistic analysis conducted in this study. Based on the phylogeny, V. conradti and V. smithae sp. nov. form a well-supported clade (Figs 1 – 2).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	description	urn: lsid: zoobank. org: act: 13 ED 6 C 82 - C 06 A- 4 DE 4 - 92 E 3 - EAD 71 C 9 B 8 D 71 Figs 1 – 2, 7, 9, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	diagnosis	Diagnosis Distinguished from most species of Voconia by the semicircular yellow stripe encircling both ocelli, membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane, and proepimeron with protuberance on posteroventral margin. This species is most similar to two other Oriental species, V. decorata sp. nov. and V. lirophleps sp. nov. It is distinguishable from V. decorata sp. nov. by its distribution in Sumatra and the dark brown, almost black coloration of the body. Despite the holotypes of V. coronata sp. nov. and V. lirophleps sp. nov. being collected within about 106 km of each other, there are several differences, particularly in size. Among our larger series, conspecific Voconia males do not vary more than about ± 1 mm from each other, and V. coronata sp. nov. is about 2 mm longer than V. lirophleps sp. nov. It is distinguishable by the slightly more elongate head (about 1.3 times as long as wide), the pedicel is much longer (about 1.9 times length of head width), maxillary plates directed straight in dorsal view, postocular region is about as long as the eye, labial segment I is long, almost reaching posterior margin of head, wide pronotal collar, long anterior pronotal lobe relative to posterior pronotal lobe (posterior lobe about 1.8 times as long as anterior lobe), scutellum dark with brown apical spine, inconspicuous distal yellow spot on the corium, and the presence of three large spines and many small spines on the posterior row of spines of the hind leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ coronatus, - a, - um ’, meaning ‘ crowned’ or ‘ encircled’, referring to the yellow ring around the postocular region of the head.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	materials_examined	Type material Holotype INDONESIA • ♀; Sumatra, “ Central Atjeh ” [Aceh], Ketambe; [3.68 ° N, 97.66 ° E]; elev. 400 m; 26 – 27 Jan. 1974; E. W. Diehl leg.; USI: AMNH _ PBI 00170706; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 11.0 mm; macropterous. COLORATION. Head: dark brown; postocular region dark with a semicircular pale stripe encircling both ocelli; labium nearly as dark as head. Thorax: coloration as head; posterior pronotal lobe lighter than anterior lobe; scutellum dark with brown apical spine. Hemelytron (abducted): clavus dark with distal yellow stripe; corium dark with anteroproximal yellow stripe, posteromedial and small distal yellow spots; membrane uniformly dark; membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane. Legs: dark brown, femora darker. Abdomen: almost black, dorsal laterotergites dark with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and more than three small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.3 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.9 times length of head width; maxillary plates ellipsoidal, adjacent to and surpassing clypeal apex; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles short and flared laterally; anterior pronotal lobe about 0.6 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality of Ketambe in the northernmost province of Sumatra, Aceh.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA0FFA8FD9900B2FB83BD47.taxon	discussion	Remarks The phylogenetic placement of V. coronata sp. nov. is uncertain (Figs 1 – 2).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	description	urn: lsid: zoobank. org: act: 93 BE 46 E 2 - 210 B- 45 C 7 - 8 FD 6 - 2 C 38 EC 62 B 2 D 9 Figs 1 – 2, 3 B, 4 J, 7, 9, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	diagnosis	Diagnosis Distinguished from most other species of Voconia by the yellow ring around the postocular region, membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane, and proepimeron with protuberance on posteroventral margin. This species is most similar to two other Oriental species, V. coronata sp. nov. and V. lirophleps sp. nov. It is distinguishable from these species by its distribution in the Malaysian state of Sabah in Borneo, the light brown coloration of the body, the labium being lighter than the head, and the dark scutellum with a contrasting yellow apical spine.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ decoratus, - a, - um ’, meaning ‘ decorated, elegant’, referring to its light brown coloration and yellow markings on the hemelytron and dorsal laterotergites that gives it a flashier and more elegant look.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	materials_examined	Type material Holotype MALAYSIA [“ NORTH BORNEO ”] • ♀; Sabah, Tawau, Quoin Hill, Cocoa Res. Stn, [4.42 ° N, 118.02 ° E]; elev. 225 m; 16 Sep. 1962; K. J. Kuncheria leg.; USI: UCR _ ENT 00073809; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 10.0 mm; macropterous. COLORATION. Head (Fig. 3 B): light brown, yellowish; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head. Thorax: coloration as head; posterior margin of posterior pronotal lobe lighter; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium reddish-brown with anteroproximal yellow stripe, posteromedial and distal yellow spots; membrane light brown with pale V-shaped marking along R and M veins; membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane. Legs: yellowish-brown, distal half of femora darker. Abdomen: yellowish-brown, dorsal laterotergites with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head (Fig. 3 B) and pronotum: finely granulose with sparse, short macrosetae interspersed among sparse pubescence; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae. Hemelytron: corium with sparse, long setation. Legs (Fig. 4 J): posterior row of protuberances on mid and hind femora with three large spines on distal half and one small spine. Abdomen: ventral surface pubescent. STRUCTURE. Head (Fig. 3 B): elongate, 1.3 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.3 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view about as wide as maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality of Quoin Hill in the Malaysian state of Sabah on the island of Borneo.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA6FFAAFD670216FB82B9F5.taxon	discussion	Remarks The phylogenetic placement of V. decorata sp. nov. is uncertain (Figs 1 – 2).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	description	urn: lsid: zoobank. org: act: 7528828 E-ADAD- 49 A 4 - BA 42 - 98 BCE 32 BF 011 Figs 1 – 2, 6 – 7, 9, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	diagnosis	Diagnosis Distinguished from most other species of Voconia by its tuberculate and coarsely granulose head and pronotum. This species most closely resembles another Australian species, V. tuberculata sp. nov. These two species are recognized from other Australian species by the acute protuberance on the posteroventral margin of the proepimeron, long macrosetae of the setigerous tubercles, elongate head (1.4 – 1.6 times as long as wide), and yellow spots antero- and posteromedially on the abducted corium. Voconia dolichocephala sp. nov. differs from V. tuberculata sp. nov. by its overall dark brown, almost black coloration, dorsal laterotergites uniformly dark or with yellow spots posteriorly, and males are smaller than about 7.5 mm. Females slightly surpass the high-end range for female length of V. tuberculata sp. nov. (8.4 – 8.8 mm).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	etymology	Etymology The specific epithet is the latinized Greek adjective ‘ dolichocephalus, - a, - um ’, formed from the prefix ‘ dolicho- ’ (meaning ‘ long’) and the noun ‘ cephale ’ (meaning ‘ head’), referring to the long head of the type specimen, which is about a fifth of the body length.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	materials_examined	Type material Holotype AUSTRALIA • ♀; Northern Territory, Tindal; [14.52 ° S, 132.38 ° E]; 1 – 20 Dec. 1967; W. Vestjens leg.; at light; USI: AMNH _ PBI 00168747; ANIC. Paratypes AUSTRALIA • 1 ♀; Northern Territory, Kakadu NP, S Alligator River, Gungaree Rainforest; [12.65 ° S, 132.47 ° E]; 17 Dec. 1993; S. and J. Peck leg.; coll. S. and J Peck 93 - 131; at light; USI: AMNH _ PBI 00170709; deposited in AMNH • 1 ♀; same collection data as for holotype; USI: AMNH _ PBI 00168748; AMNH • 1 ♂; same collection data as for holotype; USI: AMNH _ PBI 00168745; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. 7.1 – 7.5 mm; macropterous. COLORATION. Head (Fig. 3 F): dark brown; postocular region uniformly dark with pale patches adjacent to lateral ocellar margin; labial segment I nearly as dark as head, segments II and III lighter than head. Thorax: coloration as head; scutellum uniformly dark brown. Hemelytron (abducted): clavus dark brown with distal yellow stripe; corium dark brown with anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: dark brown, tarsi yellow. Abdomen: dark brown; dorsal laterotergites uniformly dark or dark with yellow spots posteriorly. INTEGUMENT AND VESTITURE. Head (Fig. 3 F) and pronotum: coarsely granulose with sparse, long setigerous tubercles interspersed among sparse pubescence; interocular region with two pairs of stout setigerous tubercles paramedially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with setigerous tubercles with long macrosetae. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with four large spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head (Fig. 3 F): elongate, 1.5 – 1.6 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view as long as eye, lateral margins gently rounded; pedicel 1.3 – 1.5 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view with medial margins diverging; clypeus in dorsal view wider than maxillary plates; clypeal apex bifid, not narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, 0.3 – 0.4 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles long, flared laterally; anterior pronotal lobe 0.7 – 0.8 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate, terga II and III with deeper and longer carinules; terga II and III with paired prominent longitudinal carinae, reaching about two-thirds of tergum III. Female Mostly as male, distinguished from males by the following. Body length: 8.8 – 9.0 mm. Head: elongate, 1.5 – 1.6 times as long as wide; pedicel about 1.5 – 1.7 times length of head width. Legs: fossula spongiosa absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	distribution	Distribution (Fig. 18) This species is only known from the Northern Territory of Australia. The nearest species of Voconia is V. tuberculata sp. nov., known from about 100 km southeast of the three specimens collected in Tindal.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFA5FFB4FDBF00F0FCF8B835.taxon	discussion	Remarks Voconia dolichocephala sp. nov. is nested among the Australian clade. According to the phylogeny (Figs 1 – 2), it is the sister taxon to V. tuberculata sp. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	description	urn: lsid: zoobank. org: act: 22 FAB 401 - 5 B 52 - 4866 - 86 CD- 0 ED 8 B 980 A 3 A 5 Figs 1 – 2, 7, 9, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	diagnosis	Diagnosis Distinguished from most other species of Voconia by its tuberculate and coarsely granulose head and pronotum, lateral protrusion of the buccula, and dorsally projecting mandibular plates. This species most closely resembles three other Australian species, V. grandioculata sp. nov., V. pallidipes, and V. vittata sp. nov. This species, V. grandioculata sp. nov., and V. vittata sp. nov. have a similar yellow stripe across the corium; however, the stripe in V. fasciata sp. nov. is pale, almost translucent and the body is larger, total length is about 8.7 mm long. It is easily distinguished from V. pallidipes by its smaller size and coloration, i. e., the membranal veins are not yellow, the proximal half of the corium is not entirely yellow, and the legs are not completely yellow as described above.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ fasciatus, - a, - um ’, meaning ‘ banded’, referring to the pale yellow band that crosses the middle of the corium and scutellar spine.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	materials_examined	Type material Holotype AUSTRALIA • ♂; Queensland, Einasleigh River, 33 km W of Mt Surprise; [18.13 ° S, 143.99 ° E]; 10 – 11 Jun. 1993; M. and B. Baehr leg.; USI: AMNH _ PBI 00127097; ZSMC.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 8.7 mm; macropterous. COLORATION. Dark brown, maxillary plates lighter; postocular dark with pale patches adjacent to lateral ocellar margin; labium lighter than head. Thorax: coloration as head; posterior pronotal lobe with yellow posterolateral margins; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark brown with distal yellow stripe; corium dark brown with thick medial yellow stripe; membrane and membranal veins uniformly dark. Legs: femora dark brown, tibiae and tarsi yellowishbrown. Abdomen: ventral surface light brown medially, dark brown laterally; dorsal laterotergites I and II paler than remainder. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with dense, short setigerous tubercles interspersed among sparse pubescence; interocular region with setigerous tubercles medially and two pairs paramedially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with short macrosetae on setigerous tubercles. Hemelytron: corium with dense, long macrosetae. Legs: posterior row of protuberances on mid and hind femora with four spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head: globose, about as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.3 times length of head width; mandibular plates in anterior view projecting dorsolaterally; maxillary plates ellipsoidal, adjacent to and reaching apex of clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view wider than maxillary plates; clypeal apex bifid, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin, setigerous tubercles along apical margin; labial segment I in lateral view gradually thickens distally, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, about 0.3 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles short, flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum depressed near posterolateral margins; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into sharp spine; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	distribution	Distribution (Fig. 18) This species is only known from the type locality of Einasleigh River in Queensland, Australia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBBFFB5FD620130FBA3BF7B.taxon	discussion	Remarks Voconia fasciata sp. nov. is nested within the Australian clade (Figs 1 – 2). Though poorly supported, it forms the sister taxon to V. brachycephala sp. nov. and V. vittata sp. nov.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	description	urn: lsid: zoobank. org: act: 23167 BBE- 77 C 8 - 434 F- 9574 - B 9 F 59130 EAAB Figs 1 – 2, 4 E, 7, 9, 11, 14, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	diagnosis	Diagnosis Distinguished from most other species of Voconia by its tuberculate and coarsely granulose head and pronotum, the lateral protrusion of the buccula, and notably the large ocelli that occupy half the length of the postocular region. Due to the yellow stripe across the corium, this species resembles two other Australian species, V. fasciata sp. nov. and V. vittata sp. nov. However, the yellow stripe of this species is bright and opaque, unlike that of V. fasciata sp. nov. Additionally, the legs are entirely yellow, and the corium has a small proximal yellow spot. It is easily distinguished from V. pallidipes by the brown membranal veins, the proximal half of the corium being not entirely yellow, and its globose head.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ grandioculatus, - a, - um ’, formed from the prefix ‘ grandi- ’ (meaning ‘ large’) combined with the adjective ‘ oculatus, - a, - um ’ (meaning ‘ possessing eyes’), referring to the large ocelli of this species, occupying half the length of the postocular region.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	materials_examined	Type material Holotype AUSTRALIA • ♂ (dissected pygophore and aedeagus in vial); Queensland, 3 km NE of Mt Webb; [15.05 ° S, 145.15 ° E]; 30 Apr. – 3 May 1981; A. Calder leg.; at light; USI: AMNH _ PBI 00168749; ANIC. Paratype AUSTRALIA • 1 ♂; Western Australia, East York; USI: AMNH _ PBI 00213934; HNHM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. 9.2 – 9.7 mm; macropterous. COLORATION. Head: dark brown; postocular region with pale patches adjacent to lateral ocellar margin; labial segment I yellowish brown, segments II and III yellow. Thorax (Fig. 4 E): coloration as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark brown with distal yellow stripe; corium proximal half mostly yellow, distal half dark brown with yellow spot at distal apex; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: yellow. Abdomen: dark brown; dorsal laterotergites uniformly colored. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with dense, short setigerous tubercles interspersed among sparse pubescence; interocular region with setigerous tubercles medially and two pairs paramedially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with short macrosetae on setigerous tubercles. Hemelytron: corium with dense, long macrosetae. Legs: posterior row of protuberances on mid and hind femora with four spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: globose, about as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel 1.4 – 1.6 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view wider than maxillary plates; clypeal apex bifid, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ocelli large, occupying halflength of postocular region; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin, setigerous tubercles along apical margin; labial segment I in lateral view gradually thickens distally, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, flared laterally; anterior pronotal lobe 0.6 – 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum depressed near posterolateral margins; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised (Fig. 4 E); anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into sharp spine; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of tergum III weakly carinulate, remaining terga not carinulate; terga II and III with paired prominent longitudinal carinae, reaching about one-third of tergum III. Pygophore (Fig. 11): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view with slight swelling; median apical process short, sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex round, not sharply tapered or expanded. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongue-shaped in dorsal view; basal plate extension about 5.7 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	distribution	Distribution (Fig. 18) This species is known from Queensland and Western Australia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBAFFB7FD870672FD9FB92A.taxon	discussion	Remarks The phylogenetic placement of V. grandioculata sp. nov. within the Australian clade is uncertain (Figs 1 – 2). The abdomen of the holotype was extracted, but we did not get sufficient DNA in our NGS library for Illumina sequencing.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	description	urn: lsid: zoobank. org: act: A 6 B 29189 - 616 B- 4489 - BAE 0 - 4 A 1 F 5 F 055 DEA Figs 1 – 2, 7, 9, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	diagnosis	Diagnosis This species is most similar to other Southeast Asian Voconia species with a finely granulose head and pronotum and yellow spots on the anteromedial, posteromedial, and distal apex of the abducted corium. Voconia hemera sp. nov. is larger than V. minima sp. nov. (9.2 – 9.3 mm). It is distinguished from V. ornata, V. nyx sp. nov., and V. bakeri sp. nov. by its uniformly yellow legs and antenna as well as its uniformly brown scutellum and dorsal laterotergites.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	etymology	Etymology Named after the Greek primordial goddess of the day, Hemera. Refers to the yellow legs and antennae of this species, which differs from a similar yet darker species, V. nyx sp. nov., named after the goddess of night. A proper noun in apposition.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂; Laguna, Los Baños; [14.17 ° N, 121.18 ° E]; 1925; Pemberton leg.; USI: UCR _ ENT 00073812; BPBM. Paratype PHILIPPINES • 1 ♂; Luzon, Camarines Sur, Mt Isarog, Pili; [13.66 ° N, 123.37 ° E]; elev. 800 m; 25 Apr. 1965; H. M. Torrevillas leg.; light trap; USI: UCR _ ENT 00073813; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	description	Description Male (Figs 1 – 2) BODY LENGTH. 9.2 – 9.3 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head, brownish yellow. Thorax: as head; scutellum uniformly dark brown. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium reddish-brown with anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: yellowish-brown. Abdomen: dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large and two small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially, projected forward; anterior pronotal lobe about 0.8 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin, gently depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	distribution	Distribution This species is only known from the type locality on the Philippine island of Luzon.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB8FFB0FD600020FEECBC29.taxon	discussion	Remarks The holotype locality of Los Baños is about 10 km from the holotype locality of V. nyx sp. nov. at Mt Maquiling.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	description	Fig. 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	diagnosis	Revised diagnosis Distinguished from other species of Voconia by the following combination of characters: short body length (about 6.5 mm); coarsely granulose head and pronotum; maxillary plates are not adjacent to and far surpass clypeal apex; maxillary plates in lateral view are wide, about twice the width of scape; antennifer with long lateral spine; eyes do not reach ventral head margin in lateral view. It is recognized from micropterous individuals of V. schoutedeni and V. monodi by the toothed apex of the maxillary plates.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	materials_examined	Type material Holotype SENEGAL • ♂; “ Forêt classée de Tobor ” [Tobor Forest]; 21 Nov. 1961; “ Mission IFAN ” [Institut Fondamental d’Afrique Noire exped.]; “ sous un morceau de bois mort ” [under a piece of dead wood]; MNHN.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	description	Description Male BODY LENGTH. About 6.5 mm; micropterous. COLORATION. As described for V. monodi, except legs testaceous. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with setigerous tubercles; interocular region with two pairs of large setigerous tubercles paramedially. STRUCTURE. Head: scape not reaching head apex; antennifer with long lateral spine; postocular region in dorsal view as long as eye, lateral margins subrectangular; maxillary plates in lateral view about twice the width of scape and apex toothed; eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin; labial segment I in lateral view straight, surpassing posteroventral eye margin. Thorax: scutellar spine reduced, subhorizontal. Abdomen: anterior margin of terga carinulate; tergum II with paired prominent longitudinal carinae reaching posterior margin of segment. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	distribution	Distribution (Fig. 17) This species is known from Northwest Africa.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBFFFB1FDC30524FADBBFE7.taxon	discussion	Remarks The holotype was on loan and not available for study at the time we requested it. Consequently, this description is based on the interpretation of the short differential diagnosis and illustration of the head in lateral view provided by Villiers (1963 a). Due to the limited data that we were able to obtain, it was excluded from cladistic analyses, but we are confident that it likely forms a clade with V. schoutedeni and V. tridens sp. nov. because of the unique head morphology, microptery, and Afrotropical distribution.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	description	urn: lsid: zoobank. org: act: 826 EFBC 8 - FA 03 - 42 DA- 8 E 5 E-E 0 D 4 F 29 A 5 B 66 Figs 1 – 2, 7, 9, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	diagnosis	Diagnosis This species most closely resembles another Southeast Asian species, V. loki sp. nov., due to the finely granulose head and pronotum, yellow legs and antennae, dark pronotum with contrasting yellow posterior margin, and the proximal half of the corium being entirely yellow, distal half dark with small distal yellow spot. Voconia isosceles sp. nov. is slightly larger (about 8.8 mm long), the membranal veins forming the Cu-An 1 and M-Cu cells are paler than the remainder of the membrane, and the postocular region is dark with a semicircular pale stripe encircling both ocelli.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ isosceles, -, - ’, which has the same meaning as its equivalent geometric term used in English for a triangle having at least two sides of equal length (also known as a golden triangle). Refers to the yellow or golden isosceles triangle on the proximal half of the corium.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂; Mindanao, Gingoog City, Mts Or., Mt Pomalihi, 21 km W of Gingoog City; [8.81 ° N, 124.92 ° E]; elev. 800 – 1000 m; 16 – 18 Oct. 1965; H. M. Torrevillas leg.; light trap; USI: UCR _ ENT 00073814; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 8.8 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head, yellow. Thorax: as head; posterior pronotal lobe with posterior margin lighter, yellow; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium proximal half yellow, distal half reddish-brown with yellow spot at distal apex; membrane brown with pale V-shaped marking along R and M veins; membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane. Legs: yellow. Abdomen: dark brown ventrally; laterotergites light brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.4 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.3 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.6 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga prominently carinulate; terga II and III with paired prominent longitudinal carinae, almost reaching posterior margin of tergum III. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in the Philippines.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBEFFB3FD67051EFCD1B92A.taxon	discussion	Remarks Based on the record of a Miridae specimen from the same collection event as the holotype, Mt Pomalihi is presumably in Mt Balatukan Range Natural Park. The type locality is also shared with V. bakeri sp. nov. Though V. isosceles sp. nov. and V. loki sp. nov. closely resemble each other, we are treating them as separate species because they were not recovered as closely related taxa in our analyses (Figs 1 – 2). This hypothesis should be further tested with additional material.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	description	urn: lsid: zoobank. org: act: D 92 E 888 A- 8436 - 42 C 2 - 8316 - 881775941725 Figs 1 – 2, 7, 9, 12, 14, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	diagnosis	Diagnosis This species is distinguishable by the elongate Cu-An 1 cell (reaching at least half the length of M-Cu cell) and short labial segment I (not surpassing ventral eye margin in lateral view). This species most closely resembles three other Southeast Asian species, V. isosceles sp. nov., V. lasiosoma sp. nov., and V. loki sp. nov., due to the finely granulose head and pronotum, long setation of the body, semicircular pale stripe encircling both ocelli, and the dark pronotum with contrasting pale posterior margin. However, in V. laosensis sp. nov. this pale posterior margin occupies almost the entire posterior pronotal lobe.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	etymology	Etymology A proper noun with the Latin adjectival suffix ‘ - ensis ’ meaning ‘ belonging to’, referring to the country of the type locality, Laos. To be treated as an adjective.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	materials_examined	Type material Holotype LAOS • ♂ (dissected pygophore and aedeagus in vial); Khammouan Prov., Ban Khoun Ngeun; 18.11666 ° N, 104.48333 ° E; elev. 250 m; 4 – 16 Nov. 2000; E. Jendek and P. Pacholátko leg.; USI: UCR _ ENT 00001552; NHMW.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 9.2 mm; macropterous. COLORATION. Head: dark brown, maxillary plates and clypeus lighter; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head, yellow. Thorax: as head; posterior pronotal lobe lighter than anterior lobe; scutellum dark with brown apical spine. Hemelytron (abducted): clavus dark reddish-brown with proximal and distal yellow stripe; corium dark reddish-brown with anteroproximal yellow stripe, posteromedial and distal yellow spots; membrane uniformly dark; pale V-shaped marking along R and M veins. Legs: yellowish-brown. Abdomen: dark brown, laterotergites light brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, long macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large and one small spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.2 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and reaching apex of clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially, anterolateral angles short, flared laterally; anterior pronotal lobe about 0.5 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance. Hemelytron: Cu-An 1 cell elongate, reaching at least half the length of M-Cu cell. Legs: fossula spongiosa present on fore and mid legs. Pygophore (Fig. 12): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; short median apical process upright in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongueshaped in dorsal view; basal plate extension 4.4 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in Laos. This is the only known species of Voconia on mainland Southeast Asia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFBCFFBCFD990020FC7EBCBE.taxon	discussion	Remarks The pygophore was dissected from this pinned holotype and used to extract DNA, for which we acquired the lowest DNA concentration (0.124 ng / µL) and did not sequence.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	description	urn: lsid: zoobank. org: act: C 5495703 - AEE 2 - 49 FC- 8 B 08 - 099 ED 0 F 398 D 3 Figs 1 – 2, 7, 9, 12, 14, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	diagnosis	Diagnosis Recognized from other Voconia by the densely long setation on the pronotum, corium, legs, and abdomen. This species most closely resembles two other Southeast Asian species, V. isosceles sp. nov. and V. loki sp. nov. due to the finely granulose head and pronotum and dark pronotum with contrasting pale posterior margin. While V. loki sp. nov. is also hairy, V. lasiosoma sp. nov. is larger (about 9.4 mm) than these two species; legs, antennae, and corium are dark yellowish-brown rather than yellow; dark portions of corium and clavus are black rather than reddish-brown; Cu-An 1 cell stout (less than half the length of M-Cu cell); and proximal half of corium not entirely yellow (abducted corium with anteroproximal yellowish-brown stripe and posteromedial yellow spot).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	etymology	Etymology Derived from the Latinized Greek prefix ‘ lasio- ’ meaning ‘ hairy’ combined with the Latinized Greek noun ‘ - soma ’ meaning ‘ body’. Refers to the long macrosetae covering the body and hemelytra. A noun in apposition.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	materials_examined	Type material Holotype MALAYSIA • ♂ (dissected pygophore and aedeagus in vial); Sarawak, Lanjak Entimau Wildlife Sanctuary; [1.55 ° N, 110.36 ° E]; [elev. 3 m]; 9 – 10 Oct. 2017; W. Hwang et al. leg.; collecting event: SW 17 _ L _ 27; MV light trap in dipterocarp forest; DNA voucher R _ CW 5472; USI: UCR _ ENT 00127826; ZRC.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 9.4 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head, yellow. Thorax: anterior pronotal lobe darker than head and posterior pronotal lobe; posterior pronotal lobe with dark yellowish-brown gradient; scutellum dark with contrasting dark yellowish-brown apical spine. Hemelytron (abducted): clavus black with proximal dark yellowish-brown spot and distal yellow stripe; corium black with anteroproximal dark yellowish-brown stripe, posteromedial and distal yellow spots; membrane mostly dark with pale stripe along most of M vein; membranal veins R and M proximally pale. Legs: yellowishbrown. Abdomen: dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large and three small spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; length of pedicel subequal to head width; maxillary plates ellipsoidal, adjacent to and shorter than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially, with anterolateral angles short, flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum deep, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga prominently carinulate; terga II and III with paired prominent longitudinal carinae, reaching posterior margin of tergum III. Pygophore (Fig. 12): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; short median apical process, upright in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongueshaped in dorsal view; basal plate extension 5.6 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in Sarawak, Malaysia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB3FFBFFD9F05B5FD09B8EF.taxon	discussion	Remarks The holotype was preserved in ethanol when the abdomen was dissected for DNA extraction. We acquired sufficient DNA in our NGS library for Illumina sequencing for future phylogenomic studies, referred to as DNA voucher R _ CW 5472.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	description	urn: lsid: zoobank. org: act: 66547 E 0 B-D 2 F 8 - 46 E 0 - B 1 F 7 - CC 2 DD 12 BFC 2 A Figs 1 – 2, 4 B, 7, 9, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	diagnosis	Diagnosis Distinguished from most species of Voconia by the pale membranal veins. This species is most similar to two other Oriental species, V. decorata sp. nov. and V. coronata sp. nov. It is distinguishable from V. decorata sp. nov. by its distribution in Sumatra; the dark brown, almost black coloration of the body; and uniformly brown scutellum. V. lirophleps sp. nov. is about 2 mm shorter than V. coronata sp. nov. It is also distinguishable by the yellow spots adjacent to the lateral ocellar margin; slightly more globose head (about 1.2 times as long as wide); pedicel much shorter (about 1.1 times length of head width); maxillary plates converge in dorsal view, postocular region is shorter than eye; narrow pronotal collar; short anterior pronotal lobe relative to posterior pronotal lobe (about 2.2 times shorter than posterior lobe); conspicuous distal yellow spot on the corium; and only three large spines and two small spines on the posterior row of spines of the hind leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	etymology	Etymology Derived from the Latinized Greek prefix ‘ liro- ’ meaning ‘ pale’ combined with the Latinized Greek noun ‘ phleps ’ meaning ‘ vein’. Refers to the prominent pale coloration of the membranal veins. To be treated as a noun in apposition, ending not to be changed.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	materials_examined	Type material Holotype INDONESIA • ♂; Sumatra, Djulu, Rajeu, 250 km N of Medan; elev. 50 m; E. W. Diehl leg.; USI: AMNH _ PBI 00170704; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	description	Description Male (Figs 7, 9) BODY LENGTH. About 8.9 mm; macropterous. COLORATION. Head: dark brown; postocular region uniformly dark with pale patches adjacent to lateral ocellar margin; labium nearly as dark as head. Thorax: coloration as head; scutellum uniformly dark brown. Hemelytron (Fig. 4 B; abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddishbrown with anteroproximal yellow stripe, posteromedial and distal yellow spots; membrane uniformly dark; membranal veins forming Cu-An 1 and M-Cu cells paler than remainder of membrane. Legs: dark brown, femora darker. Abdomen (Fig. 4 B): dark brown; dorsal laterotergites dark with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large and three small spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.2 times as long as wide; anteocular region about one quarter of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.1 times length of head width; maxillary plates ellipsoidal, adjacent to and as long as clypeus; apices of maxillary plates in dorsal view converging; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.5 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in North Sumatra.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB0FFB8FD9F01E7FBA0BC67.taxon	discussion	Remarks Despite the holotypes of V. coronata sp. nov. and V. lirophleps sp. nov. being collected within about 106 km of each other, there are several differences, particularly in size. Among our larger series, conspecific Voconia males do not vary much in size, and V. lirophleps sp. nov. is about 2 mm shorter than V. coronata sp. nov.; thus, we are keeping them as separate species.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	description	urn: lsid: zoobank. org: act: BBC 757 E 0 - 8 E 3 F- 4 CCC- 9 B 5 F- 8 E 7 FA 5 FAB 7 A 2 Figs 1 – 2, 3 H, 8, 10, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	diagnosis	Diagnosis This species most closely resembles two other Southeast Asian species, V. isosceles sp. nov. and V. lasiosoma sp. nov., due to the finely granulose head and pronotum; legs and antennae yellow; pronotum dark with contrasting yellow posterior margin; and relatively denser and longer setation. Voconia loki sp. nov. is smaller (about 8.3 mm long) than these two species. It is also recognized from V. isosceles sp. nov. by the stout Cu-An 1 cell (less than half the length of the M-Cu cell), brown membranal veins that form the Cu-An 1 and M-Cu cells, and the postocular region brown having dark patches adjacent to the medial ocellar margin. Apart from its smaller size, this species is distinguished from V. lasiosoma sp. nov. by the almost entirely yellow proximal half of the corium, reddish-brown distal half of the corium instead of black, and yellow legs instead of a darker yellowish-brown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	etymology	Etymology Named after the cunning trickster from Norse mythology and from the Marvel Comics’ character, Loki, since this specimen deceived and tricked authors in a previous study (Hwang & Weirauch 2012) who misidentified it as “ Kayanocoris wegneri ” (V. wegneri comb. nov.). A proper noun in apposition.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	materials_examined	Type material Holotype BRUNEI • ♂; Belait District, 21.5 km N of Labi on Labi Rd, Karanga Forest; 4.58244 ° N, 114.50508 ° E; elev. 40 m; 28 Jun. 2010; C. Weirauch and W. Hwang leg.; collecting event: BR 10 _ L 18; hand collected; low-elevation dipterocarp forest; DNA voucher R _ CW 1590; USI: UCR _ ENT 00052216; UCR.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. About 8.3 mm; macropterous. COLORATION. Head (Fig. 3 H): light brown; clypeus yellow; postocular region with dark patches adjacent to medial ocellar margin; labium lighter than head, yellow. Thorax: pronotum darker than head; posterior pronotal lobe with posterior margin lighter; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; proximal half of corium mostly yellow with small black spot, distal half reddish-brown with distal yellow spot; membrane dark with proximally pale V-shaped marking along R and M veins; membranal veins R and M proximally pale, veins forming Cu-An 1 cell pale. Legs: yellow. Abdomen: dark brown; dorsal laterotergites yellow. INTEGUMENT AND VESTITURE. Head and pronotum (Fig. 3 H): finely granulose with dense, long macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with strong macroseta, base not protruded laterally; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large and three small spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head (Fig. 3 H): elongate, about 1.2 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view as long as eye, lateral margins gently rounded; pedicel about 0.9 times length of head width; maxillary plates ellipsoidal, adjacent to and shorter than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum deep, elongated longitudinally; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in Brunei on the island of Borneo.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB7FFBAFD79059DFAB8B8C8.taxon	discussion	Remarks The holotype is referred to as Kayanocoris wegneri R _ CW 1590 in Hwang & Weirauch’s (2012) molecular phylogeny, where it is recovered as the sister taxon to all other species of Gerbelius.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	description	urn: lsid: zoobank. org: act: 2 FAEA 443 - 4 F 17 - 4 ABE- 9 F 77 - C 28792 F 7 D 5 D 0 Figs 1 – 2, 3 I, 4 I, 8, 10, 12, 14, 19	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	diagnosis	Diagnosis One of two New World species of Voconia. Recognized by the coarsely granulose head and prothorax; wide anteocular region; morphologically ventral surface of labium with sparsely long macrosetae; buccula with lateral tubercles protruding past buccular margin; and yellow antero- and posteromedial spots on the abducted corium.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ mexicanus, - a, - um ’, meaning ‘ from or pertaining to Mexico’, referring to the country of the type locality.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	materials_examined	Type material Holotype MEXICO • ♂ (dissected pygophore and aedeagus in vial); Veracruz, 4 mi N of “ Cardale ” [José Cardel]; 30 Oct. 1982; G. Gordh leg.; DNA voucher R _ CW 2768; USI: UCR _ ENT 00004570; UCR. Paratypes HONDURAS • 1 ♀; El Paraíso, 13 km NW of El Zamorano; [14.15 ° N, 86.47 ° W]; 27 Jul. 1977; C. W. and L. B. O’Brien and Marshall leg.; USI: AMNH _ PBI 00170701; AMNH. MEXICO • 1 ♂; Oaxaca, Tehuantepec; [16.33 ° N, 95.23 ° W]; 11 Jun. 1964; J. C. and D. Pallister leg.; USI: AMNH _ PBI 00170702; AMNH. • 1 ♂; same collection data as for preceding; USI: AMNH _ PBI 00170703; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. 10.3 – 10.9 mm; macropterous. COLORATION. Head: uniformly dark brown; labium lighter than head, yellow brown. Thorax: as head; posterior pronotal lobe with posterior margin lighter; scutellum dark with brown apical spine. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish- brown with anteromedial and posteromedial yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R, M, and An 1 proximally pale. Legs: dark brown. Abdomen: dark brown, laterotergites light brown. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with sparse pubescence on setigerous tubercles; interocular region unarmed with macrosetae; antennifer with 1 – 3 short lateral setigerous tubercles; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles. Hemelytron: corium with dense, short setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and two to five small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.2 – 1.3 times as long as wide; anteocular region about one third of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel 1.4 – 1.9 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight or diverging; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin; labial segment I in lateral view straight, reaching or slightly shorter than posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.5 – 0.6 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and gently depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, subhorizontal or raised; anteriad-directed process of prosternum with paramedial lobes (Fig. 3 I); anterior margin of stridulitrum projected into small protuberance (Fig. 3 I); proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate; terga II and III with paired prominent longitudinal carinae, reaching about one-third of tergum III. Pygophore (Fig. 12): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view with large swelling; short median apical process sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on flat margin; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite rounded in dorsal view; basal plate extension about 4.3 times as long as wide. Female Distinguished from males by the following. Body length: about 12.2 mm. Head: elongate, about 1.2 times as long as wide; anteocular region shorter than postocular region (measured to anterior margin of neck); with sparse, short macrosetae on setigerous tubercles and sparse pubescence on setigerous tubercles; pedicel about 1.6 times length of head width; apices of maxillary plates in dorsal view directed straight; labium lighter than head, brown; labial segment I in lateral view surpassing posteroventral eye margin. Thorax: pronotal collar in dorsal view narrow, almost absent medially; anterolateral angles of pronotum long, projected forward. Hemelytron: corium dark with thick medial yellow stripe; with dense, long macrosetae. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and many small spines (Fig. 4 I); fossula spongiosa absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	distribution	Distribution (Fig. 19) This species is distributed in Southern Mexico and Honduras.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFB5FFC4FD9D0003FB50BB86.taxon	discussion	Remarks The female from Honduras is considerably different from the males from Mexico as the description states above. We keep them as a single species until this hypothesis can be further tested with additional material. Specimen voucher UCR _ ENT 00004570 was mis-labelled as collected “ 4 mi N of Cardale ”, though it should be “ 4 mi N of José Cardel ” as recorded by other collectors on the same trip.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	description	urn: lsid: zoobank. org: act: CFBDCF 33 - C 855 - 47 F 0 - 9945 - 98 BE 6 E 37 A 338 Figs 1 – 2, 8, 10, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	diagnosis	Diagnosis This species is recognized from other Philippine species such as V. bakeri sp. nov. and V. nyx sp. nov. by its short body length (about 8.5 mm long); dark tibiae with proximal and distal ends yellow; and abducted corium with yellow stripe occupying half of the anteroproximal margin and expanding medially to become adjacent to posteromedial yellow spot.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ minimus, - a, - um ’, meaning ‘ smallest’. Refers to the fact that this is among the smallest species of Voconia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂; Mindanao, Zamboanga del Sur, Lemesahan; [7.84 ° N, 123.30 ° E]; elev. 600 m; 7 Sep. 1958; H. E. Milliron leg.; tight trap; USI: UCR _ ENT 00073811; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. About 8.5 mm; macropterous. COLORATION. Head: dark brown; postocular region with pale patches adjacent to lateral ocellar margin; labium nearly as dark as head. Thorax: as head; scutellum dark, tip of apical spine contrasting yellow. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddishbrown with anteroproximal yellow stripe, posteromedial and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: dark brown, proximal and distal ends of tibiae and tarsi yellow. Abdomen: dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse, long setation. Legs: posterior row of protuberances on mid and hind femora with three large and two small spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.4 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view converging; clypeus in dorsal view about as wide as maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.6 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	distribution	Distribution (Fig. 8) This species is only known from the type locality on the Philippine island of Mindanao.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCBFFC5FD6102FDFEF4BE33.taxon	discussion	Remarks The carinules on the dorsal surface of the abdomen were not examined in order to prevent damage to the holotype.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	description	Fig. 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	diagnosis	Revised diagnosis Distinguished from other species of Voconia by the following combination of characters: short body length (about 7.0 mm long); coarsely granulose head and pronotum; maxillary plates not adjacent to and far surpass clypeal apex; maxillary plates in lateral view wide, about twice the width of the scape and not toothed apically; antennifer with long lateral spine. It is recognized from the micropterous V. schoutedeni by the paired, short, and not prominent longitudinal carinae on tergum II, not reaching the posterior margin of the segment; reliefs of the anterior lobe of the pronotum less strong, effaced in front; and indistinct ocelli.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	materials_examined	Type material Holotype GUINEA • ♀; “ Région du Mont Nimba ” [Mount Richard-Molard region], “ Camp du Gouan ” [Guinea camp]; 5 Apr. 1957; L. A. V. leg.; prairie; MNHN.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	description	Redescription Female BODY LENGTH. About 7.0 mm; micropterous. COLORATION. Head: yellowish brown. Thorax: Pronotum yellowish in front and on the sides, brown on the rest of the surface; scutellum brown. Micropterous wing: yellowish. Legs: brown with proximal and distal part of femur and tarsus testaceous. Abdomen: dark brown, dorsal laterotergites with anterior half yellow. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with setigerous tubercles; interocular region with two pairs of large setigerous tubercles paramedially; antennifer with long lateral spine. Legs: posterior row of protuberances on mid and hind femora with three or four spines. STRUCTURE. Head: scape not reaching head apex and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; eye width in dorsal view narrower than synthlipsis; ocelli absent; postocular region in dorsal view longer than eye, lateral margins subrectangular; labial segment I in lateral view straight, surpassing posteroventral eye margin; pronotal anterolateral angles long, flared laterally; scutellar spine reduced, slightly raised. Abdomen: anterior margin of terga carinulate; tergum II with paired, short, and not prominent longitudinal carinae. Male Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	distribution	Distribution (Fig. 17) This species is known from Northwest Africa.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCAFFC6FDF1072AFD43BEC7.taxon	discussion	Remarks Specimens were on loan and not available for this study. Consequently, this description and localities were inferred from the French description, illustrations, and key provided by Villiers (Villiers 1963 b, 1964). Villiers (1963 b) includes an additional specimen from Guinea that was not treated as type material that we could not examine and have limited information for. We therefore refrain from including it. A drawer image of two specimens from the BMNH in 2011 suggests that this species may be distributed in Sierra Leone (collection event: 11 May 1926) and Nigeria, but their identification is difficult to confirm without better images and without the holotype.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	description	Figs 1 – 2, 8, 10, 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	diagnosis	Revised diagnosis Distinguished from other species in Voconia by its finely granulose head; short labial segment I that does not reach posteroventral eye margin; anterior margin of buccula with dense, long macrosetae; and stout body. This species most closely resembles V. conradti and V. smithae sp. nov. in its stout body plan and Afrotropical distribution. It is easily recognized from these two species by the absent or vestigial fossula spongiosa on the midleg and its distinct coloration: uniformly brown scutellum, abducted corium with an anteromedial yellow spot, and uniformly brown dorsal laterotergites.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	materials_examined	Type material Holotype DEMOCRATIC REPUBLIC OF THE CONGO • ♀; Haut-Uélé, Moto; [2.90556 ° N, 29.415 ° E]; [elev. 898 m]; 1920; L. Burgeon leg.; USI: UCR _ ENT 00070525; RMCA ENT 000018004.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	description	Redescription Female (Figs 8, 10) BODY LENGTH. About 11.6 mm; macropterous. COLORATION. Head: dark brown; labium nearly as dark as head. Thorax: coloration as head; posterior pronotal lobe with posterior margin lighter; scutellum uniformly dark brown. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish-brown with anteromedial and posteromedial yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins proximally pale. Legs: yellowish-brown with distal half of femora dark brown. Abdomen: dark brown; dorsal laterotergites uniformly dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among dense pubescence; antennifer with macroseta, base not protruded; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae. Hemelytron: corium with sparse long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half, middle spine much longer. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.2 times as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.5 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis, eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.7 times length of segment I. Thorax: pronotal collar wide medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, raised; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore leg, absent or vestigial on mid leg. Male Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	distribution	Distribution (Fig. 17) Only known from Moto and Garamba National Park (Villiers 1964) in the Haut-Uélé province of the Democratic Republic of the Congo.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC9FFC7FE09063DFDEEBDA0.taxon	discussion	Remarks The carinules on the dorsal surface of the abdomen were not examined to prevent damage to the holotype. The synonymy of Microvarus inermiceps proposed by Villiers (1976: 171) was accepted without a reexamination of the type material.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	description	urn: lsid: zoobank. org: act: 374635 C 1 - 1613 - 483 F-A 653 - 17 F 63 F 0 C 9545 Figs 1 – 2, 4 F – H, 8, 10, 12, 14, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	diagnosis	Diagnosis This species is most similar to other Southeast Asian species of Voconia with a finely granulose head and pronotum and abducted corium with yellow spots on the posteromedial and distal apex. Despite the similarity in color, V. nyx sp. nov. is larger than V. minima sp. nov. (about 10.0 mm long) and has uniformly brown legs. It is distinguished from V. bakeri sp. nov. by its uniformly brown dorsal laterotergites, pale stripe along the posterior margin of the pronotum, and the abducted corium with an anteroproximal yellowish-brown stripe.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	etymology	Etymology Named after the Greek primordial goddess of the night, Nyx. Refers to the dark legs and antennae of this species, which differ from those of a similar yet lighter colored species, Voconia hemera sp. nov., named after the goddess of day. A proper noun in apposition.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂ (dissected pygophore and aedeagus in vial); Laguna, Mt Maquiling; [14.13 ° N, 121.20 ° E]; elev. 100 m; 28 Feb. 1949; A. E. Bigornia leg.; USI: AMNH _ PBI 00170705; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. About 10.0 mm; macropterous. COLORATION. Head: dark brown; postocular region with pale patches adjacent to lateral ocellar margin; labium lighter than head, brown. Thorax (Fig. 4 F): as head; posterior pronotal lobe with posterior margin lighter; scutellum uniformly dark brown. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish-brown with yellowish-brown stripe on anteroproximal margin, posteromedial and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M pale. Legs: dark reddish-brown. Abdomen: dark reddish-brown; dorsal laterotergites dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and three or fewer small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; pedicel about 1.2 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view about as wide as maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.6 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised (Fig. 4 F); anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs (Fig. 4 G – H): fossula spongiosa present on fore and mid legs. Pygophore (Fig. 12): transverse bridge with triangular posterior margin; posterior region of ventral surface of pygophore in lateral view with large swelling; short median apical process bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongue-shaped in dorsal view; basal plate extension 4.6 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	distribution	Distribution (Fig. 17) This species is only known from the type locality on the Philippine island of Luzon.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCFFFC1FD790216FEF2BE45.taxon	discussion	Remarks The holotype locality of Mt Maquiling is about 10 km from the holotype locality of V. hemera sp. nov. at Los Baños.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	description	Figs 1 – 2, 8, 10, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	diagnosis	Diagnosis Distinguished from other species of Voconia by the following combination of characters: slender and short body (~ 7.8 – 8.7 mm long); proepimeron with protuberance on posteroventral margin; legs yellow with distal half of femora brown; scutellum dark with contrasting yellow apical spine; and dark abducted corium with anteromedial, posteromedial, and distal yellow spots.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	materials_examined	Type material Lectotype (present designation) SRI LANKA • ♂; “ Ceylon ” [Sri Lanka]; E. E. Green leg.; Distant coll. 1911 – 383; USI: UCR _ ENT 00048381; BMNH. Additional material examined INDIA – Madhya Pradesh • 1 ♀; Jahalpur; elev. “ 1600 ft ” [488 m]; Oct. 1957; P. S. Nathan leg.; dissected abdomen in vial; USI: AMNH _ PBI 00170698; AMNH • 1 ♀; same collection data as for preceding; Susai Nathan leg.; dissected abdomen in vial; USI: AMNH _ PBI 00170697; AMNH. – Tamil Nadu • 1 ♀; Tranquebar, Madras; 30 Mar. 1969; K. Krishna leg.; USI: AMNH _ PBI 00170699; AMNH. – Uttar Pradesh • 1 ♂; Maldevta, Dehra Dun; elev. “ 3500 ft ” [1067 m]; 15 Apr. 1942; Jai K. Uniyal leg.; USI: AMNH _ PBI 00170696; AMNH. UNKNOWN COUNTRY • 1 ♂; “ 832 - 13 ”; USI: AMNH _ PBI 00213933; HNHM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. 8.0 – 9.0 mm; macropterous. COLORATION. Head: light brown; postocular region with pale patches adjacent to lateral ocellar margin; labium lighter than head, brown to yellow. Thorax: as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium reddishbrown with anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: yellow, distal half of femora brown. Abdomen: light brown; dorsal laterotergites dark brown with faint pale bands on posterior margins. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, short macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae. Hemelytron: corium with dense, long macrosetae. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and many small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel 1.3 – 1.4 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight or diverging; clypeus in dorsal view about as wide as maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin, setigerous tubercles along apical margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe 0.7 – 0.8 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum thin and deeply depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Mostly as male, distinguished from males by the following. Body length: 7.8 – 8.5 mm. Head: elongate, 1.3 – 1.4 times as long as wide; pedicel 1.3 – 1.5 times length of head width. Legs: fossula spongiosa absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	distribution	Distribution (Fig. 20) Distributed in India and Sri Lanka.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCEFFC3FDFB0780FD35B89C.taxon	discussion	Remarks Originally described as Gerbelius ornatus based on an unspecified number of specimens (syntypes) from Sri Lanka (Distant 1903 a). Cladistic analyses (Weirauch 2008; present study: Figs 1 – 2) confirm that this species is nested within Voconia, and hence it is hereby transferred to the latter genus. The abdomen of pinned specimen USI: AMNH _ PBI 00170697 was removed for DNA extraction, but insufficient DNA was acquired in the NGS library for Illumina sequencing. A single male pertaining to the type series (Fig. 8), here designated as the lectotype, was examined from a unit tray image at the BMNH, along with six other specimens for which we can only distinguish the locality for one specimen as Halwaldi, India and is not included in our distribution map nor material examined (Fig. 20). While the locality and date for the lectotype are not specified, we are confident that this specimen was studied by Distant. Kirby (1891) notes that the collector E. E. Green shared his collection of Hemiptera with the BMNH, most of which were collected at Pundaloya and could be inferred as the locality where no locality is mentioned. Less likely localities include Nitagala, Nawalapitya, Kandy, Colombo, or other.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	description	Figs 1 – 2, 3 C, 8, 10, 13 – 14, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	diagnosis	Diagnosis Distinguished from most other species of Voconia by the tuberculate and coarsely granulose head and pronotum, tuberculate lateral protrusion of the buccula, the anterolateral angles of the pronotal collar with many (more than two) setigerous tubercles, and the abducted corium with the proximal half entirely yellow and distal half dark with distal yellow spot. This species resembles another Australian species, V. grandioculata sp. nov., in coloration but is easily distinguished by its larger size (10.5 – 11.3 mm long), smaller eyes and ocelli (ocellus does not occupy half-length of postocular region), M-Cu cell partially or entirely yellow, membranal veins forming Cu-An 1 and M-Cu cells mostly or entirely yellow, and yellow clavus with a dark medial spot.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	materials_examined	Type material Lectotype (present designation) AUSTRALIA • 1 ♀; Queensland, Moreton Bay; Stevens leg.; NHRS-GULI 000007827. Additional material examined: AUSTRALIA – New South Wales • 1 ♀; Inverell; [29.78 ° S, 151.12 ° E]; Armstrong leg.; USI: AMNH _ PBI 00170708; AMNH. — Queensland • 1 ♂; Clermont; [22.82 ° S, 147.64 ° E]; Jul. 1928; K. K. Spence leg.; USI: AMNH _ PBI 00088008; AMS K 156495 • 1 ♂; Condamine River, 10 km WSW of Dalby; [27.21 ° S, 151.18 ° E]; 5 Nov. 1990; M. Baehr leg.; USI: AMNH _ PBI 00127094; ZSMC • 1 ♂; Mackenzie River Fitzroy Developmental Road; [22.94 ° S, 148.92 ° E]; 11 Nov. 1990; M. Müller leg.; USI: AMNH _ PBI 00127093; ZSMC.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. 10.5 – 11.3 mm; macropterous. COLORATION. Head (Fig. 3 C): dark brown; postocular region with pale patches adjacent to lateral ocellar margin; labium lighter than head, yellow. Thorax: as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus yellow with dark medial spot; corium proximal half yellow, distal half dark reddish-brown with yellow spot at distal apex; membrane dark with pale V-shaped marking along R and M veins, M-Cu cell entirely or partially yellow; membranal veins forming Cu-An 1 and M-Cu cells mostly or entirely yellow. Legs: yellow. Abdomen: light brown, laterotergites uniformly yellow. INTEGUMENT AND VESTITURE. Head (Fig. 3 C) and pronotum: coarsely granulose with dense, short setigerous tubercles interspersed among sparse pubescence; antennifer with long and 1 – 2 adjacent short lateral setigerous tubercles; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with short macrosetae on setigerous tubercles. Hemelytron: corium with short macrosetae. Legs: posterior row of protuberances on mid and hind femora with four or more large spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head (Fig. 3 C): elongate, 1.2 – 1.3 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel 1.4 – 1.6 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view about as wide as maxillary plates; clypeal apex bifid, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; ventral eye margin surpassed by setigerous tubercles in lateral view; ventrolateral swelling of buccula with large lateral setigerous tubercles that surpass buccular margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, flared laterally; anterior pronotal lobe about 0.8 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on anterior pronotal lobe depressed near posterolateral margins of pronotum; median apodeme depression of pronotum deep, elongated transversely; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate, tergum II without carinules laterally; terga II and III with paired prominent longitudinal carinae, reaching about one-third of tergum III. Pygophore (Fig. 13): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view with large swelling; short median apical process, bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on decline; parameres sinusoidal, apex tapered sharply. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite bifid in dorsal view; basal plate extension about 5.8 times as long as wide. Female As male. Body length: about 10.7 mm. Head: elongate, about 1.3 times as long as wide; pedicel about 1.5 times length of head width. Legs: fossula spongiosa absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	distribution	Distribution Only known from Eastern Australia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFCCFFCDFD8B01D7FE70BB7A.taxon	discussion	Remarks We were unable to physically examine the type material of Voconia pallidipes (photographs available on the NHRS website) and we thus based our redescription and character coding on a specimen deposited at the AMNH that was identified as V. pallidipes (USI: AMNH _ PBI 00170708) by P. Wygodzinsky and that matches the diagnostic features visible in the type photographs. The specimen in the NHRS photographs is the only specimen in the series and thus designated as the lectotype. Characters of the dorsal part of the abdomen (i. e., carinules and carinae), pygophore, and aedeagus are based on a single specimen (USI: AMNH _ PBI 00127093), to limit damage to specimens. This species is nested within the well supported Australian clade (Figs 1 – 2). The record and illustration of V. pallidipes by Swanson (2015: 268 – 269, fig. 10) is based on misidentification and pertains to V. brachycephala sp. nov. (see under the latter species).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	description	Figs 1 – 2, 3 D, 4 D, 8, 10, 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	diagnosis	Diagnosis Distinguished from other species of Voconia by the following combination of characters: coarsely granulose head and pronotum; maxillary plates not adjacent to and far surpassing clypeal apex; maxillary plates in lateral view wide, about twice the width of the scape and not toothed apically; antennifer with long lateral spine; eyes in lateral view do not reach ventral head margin; pedicel short, about 0.6 times length of head width. The macropterous morph is recognized from V. tridens sp. nov. by the almost entirely dark brown head and pronotum, the scutellar spine is slightly raised, and the hemelytra are blackish with pale coloration at the base and apex of the corium.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	materials_examined	Type material Holotype DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♂; “ P. N. G. ” [Garamba National Park], “ Aka- Gar / 4 ” [confluence of rivers Aka and Garamba south of Mt Bawezi, locality no. 4]; [3.88 ° N, 29.21 ° E]; 2 Feb. 1951; Miss. H. De Saeger and J. Verschuren leg.; No. 1195; USI: UCR _ ENT 00070524; RMCA- ENT- 000018003. Paratype DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♀; Garamba National Park; 1952; Miss. H. De Saeger leg.; No. 3283; MNHN. Material examined from original description DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♂; Garamba National Park; No. 1281; repository not specified in description.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	description	Redescription Male (Figs 8, 10) BODY LENGTH. About 6.7 mm, micropterous; about 6.7 mm, macropterous. COLORATION. Head (Fig. 3 D): yellow; labium as remainder of head (micropterous); almost entirely dark brown (macropterous). Thorax: as head; posterior pronotal lobe darker than anterior lobe; scutellum brown with darker lateral margins. Wing stumps: yellow with brown posterior margins. Hemelytra: blackish, slightly cleared up at the base and apex of corium. Legs: yellow, distal third of femora dark brown. Abdomen (Fig. 4 D): dark brown, dorsal laterotergites with anterior half yellow. INTEGUMENT AND VESTITURE. Head (Fig. 3 D) and pronotum: coarsely granulose with dense, short macrosetae on setigerous tubercles; interocular region with two pairs of large setigerous tubercles paramedially; antennifer with long lateral spine; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles; scutellar lateral carinae with pubescent tubercles. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and three or fewer small spines. Abdomen (Fig. 4 D): ventral surface pubescent. STRUCTURE. Head (Fig. 3 D): elongate, about 1.4 times as long as wide, narrower in macropterous morph; anteocular region two-fifths of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins subrectangular; scape not reaching apex of head, about 0.4 times length of pedicel; pedicel about 0.6 times length of head width; maxillary plates conical, not adjacent to or longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis in micropterous morph, eyes considerably larger in macropterous morph; eye not reaching ventral head margin in lateral view; ocelli minute, occupying about 0.16 times length of postocular region in micropterous morph, ocelli considerably larger in macropterous morph; ventrolateral swelling of buccula with acute protrusion surpassing buccular margin, small setigerous tubercles along margin of protrusion; labial segment I in lateral view straight, not reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles long, flared laterally; anterior pronotal lobe about 1.3 times length of posterior pronotal lobe, lateral margins rounded in micropterous morph; anterior pronotal lobe shorter than posterior pronotal lobe in macropterous morph; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine reduced and subhorizontal in micropterous morph; scutellum about 2 times as long as wide at base, with long slightly raised scutellar spine in macropterous morph; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen (Fig. 4 D): anterior margin of terga prominently carinulate, terga II and III with deeper and longer carinules; terga II with paired prominent longitudinal carinae reaching posterior margin of segment. Female As micropterous male. Body length: about 7.0 mm.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	distribution	Distribution Distributed in Garamba National Park in the Democratic Republic of the Congo.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC2FFC8FE1C0272FADDBE33.taxon	discussion	Remarks This description of V. schoutedeni is based on the micropterous male holotype (UCR _ ENT 00070524), which was the only type specimen available for examination. Any descriptive information of the micropterous female or macropterous form is taken directly from the original description by Villiers (1964). A lectotype was also designated for the macropterous morph, Pseudocethera schoutedeni forme alata Villiers, 1964. However, this infrasubspecific name is permanently unavailable under ICZN Article 45.6.3, therefore this specimen (referred to as No. 1281 by Villiers) has no type status. A drawer image of specimens at the BMNH in 2011 suggests that the macropterous morph may be distributed as far north as Sierra Leone, but this specimen and the macropterous paratype need to be properly examined.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	description	urn: lsid: zoobank. org: act: 6 E 950 EB 8 - E 03 F- 4 F 9 F-B 422 - BD 1 D 7 A 1 A 1594 Figs 12, 14	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	diagnosis	Diagnosis Distinguished from other species in Voconia by its Afrotropical distribution and stout body. This species most closely resembles V. motoensis and V. conradti. Voconia smithae sp. nov. differs from V. motoensis by its distinct coloration: yellow scutellar spine; abducted corium with a yellow spot at proximal and distal apex, and brown dorsal laterotergites with contrasting yellow anterior spots. Despite the strong superficial resemblance to V. conradti due its to coloration and stoutness, closer examination reveals that V. smithae sp. nov. is distinguishable by the following: shorter body (9.7 – 9.8 mm long); light brown dorsal laterotergites with weakly contrasting yellow anterior spots; antennifer with long, laterally projecting setigerous tubercle; paired interocular setigerous tubercles; maxillary plates globular, directed straight anteriorly; longer anteocular region than postocular region (measured to where the posterior margin of the granulations meet the anterior margin of the smooth neck); and eyes do not bulge strongly laterally in dorsal view and do not reach ventral head margin in lateral view.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	etymology	Etymology Named after one of the collectors of the holotype and colleague from the Weirauch lab, Samantha Smith.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	materials_examined	Type material Holotype GABON • ♂; Ogooué-Ivindo, Ipassa Reserve; 0.51167 ° N, 12.80306 ° E; elev. 516 m; 3 – 6 Feb. 2019; S. Smith, Y. Pacheco, S. Bybee, G. Svenson and G. Powell leg.; collecting event: GA 19 _ L 8 _ H; light trap; lowland forest; DNA voucher R _ CW 5801; USI: UCR _ ENT 00127891; UCR. Paratypes REPUBLIC OF THE CONGO • 1 ♂; Sibiti; [3.68 ° S, 13.35 ° E]; 25 Nov. 1963; Endrody-Younga leg.; Soil-Zoological Exped.; lamplight; USI: AMNH _ PB 00213936; HNHM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. 9.7 – 9.8 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin, pale patches adjacent to lateral ocellar margin; labium lighter than head. Thorax: coloration as head, posterior margin of posterior pronotal lobe lighter; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus brown with distal yellow stripe; corium dark or light brown with proximal, anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: yellowish-brown. Abdomen: yellowish-brown; dorsal laterotergites light brown with yellow spots anteriorly. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with sparse, short macrosetae interspersed among sparse pubescence; interocular region with two pairs of stout setigerous tubercles paramedially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and three or fewer small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, 1.3 – 1.4 times as long as wide; anteocular region about one third of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded and subrectangular; pedicel 1.4 – 1.5 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula with tuberculate protrusion that surpasses buccular margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: pronotal collar narrow medially with anterolateral angles short and flared laterally; anterior pronotal lobe about 0.6 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum slightly depressed; median apodeme depression of pronotum deep, circular; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate; terga II and III with paired prominent longitudinal carinae, reaching about one-third of tergum III. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	distribution	Distribution (Fig. 17) Only known from Congo and Gabon in Africa.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC7FFCAFD62072AFACFBBAA.taxon	discussion	Remarks The male paratype varies from the holotype in the following: head and pronotum finely granulose, postocular region subrectangular, buccula without lateral tuberculate protrusion. The holotype was preserved in ethanol when the mid and hind legs were dissected for DNA extraction but was not Illumina sequenced.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	description	urn: lsid: zoobank. org: act: 82 E 67 D 5 A-E 019 - 40 EE-A 019 - E 3 BCD 5 E 781 DA Figs 1 – 2, 8, 10, 13 – 14, 17	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	diagnosis	Diagnosis Distinguished from other species of Voconia by the following combination of characters: coarsely granulose head and pronotum; maxillary plates not adjacent to and far surpass clypeal apex; and antennifers with a long, lateral spine. This species is recognized from the macropterous morph of V. schoutedeni by the yellow head and pronotum; the maxillary plates in lateral view are not wide, about the width of the scape; scutellar spine is subhorizontal; and the hemelytra is not blackish, it is a light brown with pale coloration at the base and apex of the corium.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ tridens, -, - ’, meaning ‘ three-toothed, three-pronged’, derived from the prefix ‘ tri - ’ combined with the Latin noun ‘ dens ’, referring to the elongated and separated maxillary plates and clypeus that form three prongs.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	materials_examined	Type material Holotype SUDAN • ♂ (dissected pygophore and aedeagus in vial); Blue Nile, Abu Hashim-Galegu; [12.25 ° N, 34.69 ° E]; 23 – 24 Nov. 1962; R. Linnavuori leg.; USI: AMNH _ PBI 00170712; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. About 8.0 mm; macropterous. COLORATION. Head: yellow; postocular region with red patches adjacent to medial ocellar margin; labium as head. Thorax: uniformly yellow; scutellum yellow. Hemelytron (abducted): clavus uniformly yellow; corium proximal half mostly yellow, distal half brown with yellow spot at distal apex; membrane dark with pale V-shaped marking along R and M veins. Legs: yellow. Abdomen: yellow, dorsal laterotergites with a pale brown stripe on posterior margin. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with dense, short setigerous tubercles; interocular region with two pairs of stout setigerous tubercles paramedially; antennifer with long lateral spine; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles; scutellar lateral carinae with pubescent tubercles. Hemelytron: corium with short macrosetae. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region two-fifths of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins subrectangular; scape not reaching head apex; pedicel about 0.9 times length of head width; maxillary plates ellipsoidal, not adjacent to or longer than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with acute protrusion surpassing buccular margin, small setigerous tubercles along margin of protrusion; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles long, flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Pygophore (Fig. 13): transverse bridge with triangular posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; short median apical process sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres round, apex tapered into squarely rounded tip. Aedeagus: endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongueshaped in dorsal view; basal plate extension about 4.6 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	distribution	Distribution (Fig. 17) Only known from the type locality in Northeastern Africa, Sudan.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFC5FFD4FD6C02A0FCD1BF76.taxon	discussion	Remarks The antennifer spine on the right side of the head of the holotype is absent. We assume it was damaged since three species, V. schoutedeni, V. monodi, and V. ifana, have similar long spines that are paired. The pygophore of specimen AMNH _ PBI 00170712 was used for DNA extraction, but we did not get sufficient DNA in our NGS library for Illumina sequencing.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	description	urn: lsid: zoobank. org: act: F 5 E 8 C 8 C 5 - 6 BD 2 - 4544 - A 81 D- 0 F 31 F 846 EF 2 E Figs 1 – 2, 8, 10, 19	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	diagnosis	Diagnosis One of two New World species of Voconia. Recognized by the finely granulose head and prothorax; proepimeron with protuberance on posteroventral margin; pronotum dark with contrasting yellow posterior margin; corium dark with anteroproximal yellow stripe, posteromedial and distal white spots; labial segment I in lateral view not surpassing posteroventral eye margin.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	etymology	Etymology Named after the collecting locality of the holotype in Trinidad. A proper noun with the Latin adjectival suffix ‘ - ensis ’ meaning ‘ belonging to’. To be treated as an adjective.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	materials_examined	Type material Holotype TRINIDAD AND TOBAGO • ♀; Trinidad; J. W. Chapman leg.; USI: AMNH _ PBI 00170700; AMNH.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	description	Description Female (Figs 8, 10) BODY LENGTH. 10.5 mm; macropterous. COLORATION. Head: dark brown; maxillary plates light brown; postocular region with pale patches adjacent to lateral ocellar margin; labium lighter than head, yellow. Thorax: as head; posterior pronotal lobe with posterior margin lighter; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus reddish-brown with distal white stripe; corium reddish-brown with anteroproximal yellow stripe, posteromedial and distal white spots; membrane dark with white V-shaped marking along R and M veins; membranal veins R and M proximally white. Legs: yellow. Abdomen: dark brown, dorsal laterotergites brown medially, with black stripe adjacent to yellow lateral margin. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, long macrosetae interspersed among dense pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with sparse, long setation. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half and many small spines. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.4 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.3 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view wider than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, reaching posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.6 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe about 0.5 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum thin, not deeply depressed; median apodeme depression of pronotum deep, elongated transversely; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa of fore leg present, absent or vestigial on mid leg. Abdomen: anterior margin of terga prominently carinulate, posterior margin of terga III – VI weakly carinulate; terga II and III with paired prominent longitudinal carinae, reaching about two-thirds of tergum III; medial region between carinae on tergum II coarsely granulose.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDBFFD5FD81066DFCD4BCF5.taxon	distribution	Distribution (Fig. 19) This species is only known from the type locality, Trinidad.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	description	urn: lsid: zoobank. org: act: B 50 D 3 ECC- 82 C 1 - 431 B- 95 C 1 - 6 C 8326049678 Figs 12, 14, 17 – 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	diagnosis	Diagnosis Distinguished from most species of Voconia by its tuberculate and coarsely granulose head and pronotum. This species most closely resembles another Australian species, V. dolichocephala sp. nov. These two species are recognized from other Australian species by the acute protuberance on the posteroventral margin of the proepimeron, the long macrosetae of the setigerous tubercles, the elongate head (1.4 – 1.6 times as long as wide), and the yellow spots antero- and posteromedially on the abducted corium. Voconia tuberculata sp. nov. differs from V. dolichocephala sp. nov. by its overall light brown coloration and males are as long as about 7.9 mm. Females are smaller than the examined V. dolichocephala sp. nov. females (about 8.8 and 9.0 mm).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ tuberculatus, - a, - um ’, meaning ‘ warty’ or ‘ tuberculate’. Refers to the densely tuberculate surfaces of the body.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	materials_examined	Type material Holotype AUSTRALIA • ♂ (dissected pygophore and aedeagus in vial); Northern Territory, 17 km NE of Willeroo; [15.23 ° S, 131.67 ° E]; 8 – 9 Nov. 1984; M. and B. Baehr leg.; USI: AMNH _ PBI 00127096; ZSMC. Paratypes AUSTRALIA – Northern Territory • 1 ♂; same collection data as for holotype; USI: AMNH _ PBI 00127095; ZSMC • 1 adult sex unknown (abdomen missing); 30 km ESE of Victoria R. Downs; 16.19 ° S, 131.17 ° E; 18 Jun. 1969; M. Mendum leg.; USI: AMNH _ PBI 00168741; ANIC • 1 ♂; Elsey Ck, 19 km SSE of Mataranka; 15.05 ° S, 133.07 ° E; 15 Oct. 1972; Upton and Barrett leg.; USI: AMNH _ PBI 00168742; ANIC. — Queensland • 1 ♀; Burketown, site of old Broom Hotel: [17.85 ° S, 139.62 ° E]; 18 May 1931; T. G. Campbell leg.; light trap; USI: AMNH _ PBI 00168743; ANIC • 1 ♀; same collection data as for preceding; USI: AMNH _ PBI 00168744; ANIC • 1 ♂; Kunkulla Station, Gregory River; 25 May 1931; I. M. Mackerras leg.; USI: AMNH _ PBI 00168746; ANIC.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. 7.9 – 8.9 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin and pale patches adjacent to lateral ocellar margin; labium lighter than head. Thorax: coloration as head; scutellum uniformly brown. Hemelytron (abducted): clavus reddish-brown with distal yellow stripe; corium reddish-brown with anteromedial, posteromedial, and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: yellowish-brown. Abdomen: light brown, dorsal laterotergites I and II paler than remainder. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with sparse, long setigerous tubercles interspersed among sparse pubescence; interocular region with two pairs of stout setigerous tubercles paramedially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, short macrosetae. Thorax (Fig. 3 L): anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with setigerous tubercles with long macrosetae. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with four or more large spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head: elongate, 1.4 – 1.6 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel about 1.5 – 2.2 times length of head width; maxillary plates ellipsoidal, adjacent to and longer than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view about as wide as maxillary plates; clypeal apex bifid, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially or medially forming a V-shaped marking; interocular sulcus in dorsal view nearly straight; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, about 0.3 times length of segment I. Thorax (Fig. 3 L): pronotal collar in dorsal view wide medially with anterolateral angles long, flared laterally; anterior pronotal lobe 0.6 – 0.8 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum conspicuous, not depressed; median apodeme depression of pronotum shallow, elongated transversely; scutellar spine long, raised; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate, terga II and III with deeper and longer carinules; terga II and III with paired prominent longitudinal carinae, reaching about two-thirds of tergum III. Pygophore (Fig. 13): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view with large swelling; short median apical process, sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered sharply. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite serrated; basal plate extension about 5.7 times as long as wide. Female Mostly as male, distinguished from males by the following. Body length: 8.4 – 8.8 mm. Head: elongate, 1.5 times as long as wide; pedicel about 1.5 – 1.7 times length of head width. Legs: fossula spongiosa present on fore leg, absent or vestigial on mid leg.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	distribution	Distribution (Fig. 18) This species is distributed across Northern Territory and Queensland in Australia. The nearest Voconia species is V. dolichocephala sp. nov., found about 100 km northwest of specimen AMNH _ PBI 00168742.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDAFFD0FD8805EFFE18B8A8.taxon	discussion	Remarks The direction of the maxillary plates in dorsal view varies from nearly straight or parallel (e. g., AMNH _ PBI 00168741) to strongly diverging (e. g., AMNH _ PBI 00168746). Voconia tuberculata sp. nov. is nested within the Australian clade. According to our phylogeny (Figs 1 – 2), V. tuberculata sp. nov. was recovered as the sister taxon to V. dolichocephala sp. nov. The abdomen of specimen AMNH _ PBI 00127096 was used for DNA extraction, but we did not get sufficient DNA in our NGS library for Illumina sequencing.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	description	Figs 1 – 2, 8, 10, 13 – 14, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	diagnosis	Revised diagnosis This species is most similar to V. bracata sp. nov. They are distinguished from other species of the genus Voconia by the slender body, long head (1.7 times as long as wide), the ventral surface of the hind leg with four large spines in the posterior row, the large pale spot between the R and M veins, and the dorsal laterotergites I and II being yellow, the remainder dark brown. The following differentiate V. typica: largest species of Voconia (12.4 – 12.7 mm long); uniformly brown legs; anterior pronotal lobe is gently rounded, posterolateral margins nearly straight.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	materials_examined	Type material Holotype INDONESIA • ♂; “ North New Guinea ” [Papua], Jayapura Co., Cyclops Mts, Sabron camp; [2.50 ° N, 140.42 ° E]; elev. “ 2200 ft ” [671 m]; Jun. 1936; L. E. C. leg.; coll. B. M. 1936 - 271; USI: UCR _ ENT 00048384; NHMUK 013588138 (previously BMNH (E) 1255222). Paratype INDONESIA • 1 ♂; same collection data as for holotype; BMNH. Additional material examined: INDONESIA • 1 ♂; “ West New Guinea ” [Papua], Central Mountains, Archbold Lake; [3.41 ° S, 138.53 ° E]; elev. 760 m; 26 Nov. – 3 Dec. 1961; S. Quate and L. Quate leg.; dissected pygophore and aedeagus in vial; USI: UCR _ ENT 00073815; BPBM.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	description	Redescription Male (Figs 8, 10) BODY LENGTH. 12.4 – 12.7 mm; macropterous. COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin and pale patches adjacent to lateral ocellar margin; labium lighter than head, light brown. Thorax: as head; pronotum uniformly coloured; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium reddish-brown with anteromedial, posteromedial, and distal yellow spots; membrane dark with large pale spot between R and M veins; membranal veins R and M proximally pale, remainder as membrane. Legs: dark brown. Abdomen: dark brown, dorsal laterotergites I and II yellow. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with sparse, short macrosetae interspersed among dense pubescence; interocular region with two paired macrosetae; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles. Hemelytron: corium with sparse, long setation. Legs: posterior row of protuberances on mid and hind femora with four large spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head: elongate, about 1.7 times as long as wide; anteocular region about one third of head length, about as long as postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel about 1.8 times length of head width; maxillary plates ellipsoidal, adjacent to and reaching apex of clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view wider than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe subequal in length to posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum thin and deeply depressed; median apodeme depression of pronotum deep and elongated transversely; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga weakly carinulate, tergum II without carinules laterally; terga II and III with paired prominent longitudinal carinae, almost reaching posterior margin of tergum III. Pygophore (Fig. 13): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view with large swelling; short median apical process sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on decline; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (Fig. 14): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite bifid in dorsal view; basal plate extension about 7.9 times as long as wide. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	distribution	Distribution (Fig. 18) This species is distributed on the island of New Guinea, in Papua.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDFFFD3FDC001A3FC53B8B2.taxon	discussion	Remarks This description is based largely on UCR _ ENT 00073815, since the holotype (Fig. 8) and paratype remain at the BMNH. Images of the types in their unit trays at the BMNH were used to evaluate this species concept. The abdomen of specimen UCR _ ENT 00073815 was used for DNA extraction, but we did not get sufficient DNA in our NGS library for Illumina sequencing.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	description	urn: lsid: zoobank. org: act: 772 BEB 7 A- 1101 - 4 AD 1 - 91 C 0 - 8 EED 38 AE 4516 Figs 1 – 2, 8, 10, 12, 14, 18	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	diagnosis	Diagnosis Distinguished from other species of Voconia by the following combination of characters: short body length (about 7.1 mm long); globose head; tuberculate and coarsely granulose head and pronotum; interocular region with two setigerous tubercles medially; apex of clypeus bifid; corium dark with thick medial yellow stripe; and dark brown femora with contrasting yellowish-brown tibiae and tarsi. This species most closely resembles another Australian species, V. fasciata sp. nov., but V. vittata sp. nov. is about 1.6 mm shorter and the yellow medial stripe on the corium is opaque.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	etymology	Etymology The specific epithet is the Latin adjective ‘ vittatus, - a, - um ’, meaning ‘ banded’. Refers to the yellow band or color pattern that crosses the middle of the corium and scutellar spine.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	materials_examined	Type material Holotype AUSTRALIA • ♂; Queensland, Cape York Peninsula, Dulhunty Rd; 11.86666 ° S, 142.5 ° E; 22 Mar. 1992; G. Cassis leg.; at light; USI: AMNH _ PBI 00088406; AMS K 156496.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	description	Description Male (Figs 8, 10) BODY LENGTH. About 7.1 mm; macropterous. COLORATION. Head: uniformly dark brown; labium lighter than head, labial segment I brown, II and III yellowish-brown. Thorax: as head; scutellum dark with contrasting yellow apical spine. Hemelytron (abducted): clavus dark reddish-brown with distal yellow stripe; corium dark reddish-brown with thick medial yellow stripe; membrane dark with pale V-shaped marking along R and M veins; membranal veins An 1 and R proximally pale. Legs: femora dark brown, tibiae and tarsi yellowish-brown. Abdomen: yellowish-brown medially, dark reddish-brown laterally; dorsal laterotergites I and II paler than remainder. INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with dense, short setigerous tubercles interspersed among sparse pubescence; interocular region with two pairs of stout setigerous tubercles paramedially and two medially; antennifer with long lateral setigerous tubercle; morphologically ventral surface of labium with sparse, long macrosetae. Thorax: anterolateral angles of pronotal collar with setigerous tubercles; scutellar lateral carinae with short macrosetae on setigerous tubercles. Hemelytron: corium with dense, long macrosetae. Legs: posterior row of protuberances on mid and hind femora with three or four spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed. STRUCTURE. Head: globose, about as long as wide; anteocular region about one quarter of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view about as long as eye, lateral margins gently rounded; pedicel about 1.4 times length of head width; maxillary plates ellipsoidal, adjacent to and reaching apex of clypeus; apices of maxillary plates in dorsal view converging; clypeus in dorsal view wider than maxillary plates; clypeal apex bifid, narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view bent anteromedially, subtriangular; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II curves ventrally, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles short, flared laterally; anterior pronotal lobe about 0.9 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, raised; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	distribution	Distribution (Fig. 18) This species is only known from the type locality in Northern Australia.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFDCFFDDFD6801A8FC22BEC2.taxon	discussion	Remarks Nested within the well supported Australian clade (Figs 1 – 2).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	description	Figs 1 – 2, 3 E, 8, 10, 20	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	diagnosis	Revised diagnosis Distinguished from other species of Voconia by the minute ocelli, occupying 0.12 times length of postocular region; labial segment I swollen subapically on ventral surface; labial segment II swollen ventrobasally and narrowed apically; labial segment III very short (about 0.14 mm long), about 0.2 times length of segment II.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	materials_examined	Type material Holotype INDONESIA • ♂; E Borneo, Balikpapan, Mentawir River; [1.04 ° S, 116.75 ° E]; elev. 50 m; 14 Oct. 1950; A. M. R. Wegner leg.; USI: RMNH. INS 1091587; NBCN (previously RMNH).	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	description	Redescription Male (Figs 8, 10) BODY LENGTH. ~ 9.1 mm; macropterous. COLORATION. Head: dark brown with a semicircular pale stripe encircling both ocelli; labium lighter than head, brown. Thorax: as head; scutellum dark with brown apical spine. Hemelytron (abducted): clavus dark with distal yellow stripe; corium dark with anteroproximal yellow stripe, posteromedial and distal yellow spots; membrane dark with pale V-shaped marking along R and M veins; membranal veins R and M proximally pale. Legs: dark brown. Abdomen: dark brown; dorsal laterotergites dark brown. INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among dense pubescence; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles. Legs: posterior row of protuberances on mid and hind femora with four spines on distal half. Abdomen: ventral surface pubescent. STRUCTURE. Head (Fig. 3 E): elongate, about 1.5 times as long as wide; anteocular region about half of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins gently rounded; pedicel about 1.1 times length of head width; maxillary plates ellipsoidal, adjacent to and shorter than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view about as wide as maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ocelli minute, occupying 0.12 times length of postocular region; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I swollen subapically on ventral surface, surpassing posteroventral eye margin; labial segment II swollen ventrobasally, narrowed apically; morphologically dorsal surface of labial segment II curves ventrally, ~ 0.2 times length of segment I; labial segment III short, ~ 0.2 times length of segment II. Thorax: pronotal collar in dorsal view narrow medially with anterolateral angles short, projected forward; anterior pronotal lobe ~ 0.8 times length of posterior pronotal lobe; glabrous markings on pronotum thin and not deeply depressed; median apodeme depression of pronotum deep, elongated transversely; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore legs, unknown on mid legs. Female Unknown.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	distribution	Distribution (Fig. 20) This species is only known from the type locality in Eastern Borneo.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
015D8E70FFD2FFDEFDFC0638FC73BD37.taxon	discussion	Remarks The holotype remains at the NBCN under loan restrictions; habitus photographs were used to evaluate this species concept. Most closely resembles the two species from New Guinea, V. bracata sp. nov. and V. typica. These three species form a poorly supported clade in our analyses (Figs 1 – 2). This hypothesis should be further tested with additional material and molecular data.	en	Castillo, Stephanie, Rédei, Dávid, Weirauch, Christiane (2022): Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs. European Journal of Taxonomy 788 (1): 1-95, DOI: https://doi.org/10.5852/ejt.2022.788.1625, URL: http://dx.doi.org/10.5852/ejt.2022.788.1625
