taxonID	type	description	language	source
BF18F633A937FFD22B9CC25BFAB4FC90.taxon	materials_examined	Type species. Acmaea mitra Rathke, 1833, present day NE Pacific (subsequent designation by Dall 1871: p. 238).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A937FFD22B9CC288FC39F810.taxon	description	Figs 19 A – C	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A937FFD22B9CC288FC39F810.taxon	materials_examined	Diagnosis. Broadly oval Acmaea shell with height less than 20 % of length and a pointed apex situated 25 – 35 % of shell length from anterior margin. Shell surface smooth or with scattered weak radial striation. Derivation of name. Refers to the Danish island Sealand on which it is found. Type material. Holotype MGUH 33077 from Skeldervig at Stevns Klint, and paratypes MGUH 33078 from Bøgeskoven and MGUH 33079 from the cliff just north of Holtug Bay, both localities at Stevns Klint. All three specimens belong to the uppermost part of the Maastrichtian Tor Formation. Additional material. At least one and probably two specimens from Rødvig. Furthermore a specimen with shell, ØSM. 10042 - 199 - a with counterpart in ØSM. 10042 - 191 is found in the collection of Østsjaellands Museum. This is also from Rødvig. Type stratum and type locality. Upper part of the upper Maastrichtian Tor Formation at Skeldervig, Stevns Klint, Denmark. Occurrence. Uppermost Tor Formation at Stevns Klint. Description. Shell low, height corresponding to less than 20 % of length, broadly oval with cone-shaped apex situated 25 – 35 % of shell length from anterior margin. Surface of shell smooth or with scattered radial striation interrupted by moderately coarse, irregular growth lines. Interior surface with indistinct wide muscle imprint, narrowing in front of apex; widest part of imprint on posterior part of shell approximately 17 % of shell length. Measurements. Largest shell, holotype MGUH 33077, 6.0 mm long and 5.4 mm wide. Remarks. Acmaea selandica n. sp. differs from the Danian A. simplex (Briart & Cornet, 1887) from Belgium by the relatively lower shell with its slightly more anterior apex and from the Belgian A. hannonica (Vincent, 1930) of the late Maastrichtian, the Palaeocene Acmaea sp. 1 and Acmaea sp. 2 described by Kollmann & Peel (1983) from Greenland and A. rigida Kaunhowen, 1898 from Maastricht by the smoother surface. This species is distinguished from A. laevigata Binkhorst, 1861 by the very low shell and from Acmaea sp. 3 sensu Kollmann & Peel (1983) from the Palaeocene of West Greenland by the more pointed and slightly curved aperture. It is distinguished from the Maastrichtian A. ciplyana (Ryckholt, 1852) of Ciply, Belgium, by the lack of the very fine and dense striae covering the surface of the latter and from the Palaeocene A. poulseni Ravn, 1939 of Denmark by the less convex and smoother shell. Acmaea selandica n. sp. is rather close to Acmaea? sp. sensu Kollmann & Peel (1983) from which it mainly appears to differ by the strong effacement of growth-lines.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A937FFD22B9CC373FDF0FD43.taxon	materials_examined	Superfamily Lottioidea Gray, 1840 a	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A939FFDC2B9CC36EFF60FC2A.taxon	description	Figs 19 D – E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A939FFDC2B9CC36EFF60FC2A.taxon	materials_examined	Material. This taxon is represented by a single wax-cast, MGUH 33080, in the collections of the Geological Museum of Copenhagen, while the original appears lost. Occurrence. Upper part of the Maastrichtian Tor Formation slightly north of Kulsti Rende at Stevns Klint. Description. Large shell weakly convex, with central, apparently flattened apex and an elongated oval outline. Aperture width 77 % of length, shell height approximately 20 % of length. Surface bearing fairly strong growth lines and very fine, regular, but slightly grainy radial striation. Measurements. Shell reaching at least 18.8 mm in length. Remarks. The apex may be secondarily deformed on the wax cast, and the assignment to genus is thus tentative.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A939FFDC2B9CC5BEFAEDFAAC.taxon	materials_examined	Type species. Cocculina rathbuni Dall, 1882, Recent from Martinique (subsequent designation by Dall 1908: p. 340).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A939FFDC2B9CC4ECFD05F850.taxon	description	Figs 19 F – G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A939FFDC2B9CC4ECFD05F850.taxon	materials_examined	Material. MGUH 33081, a fragmentary mould of outer surface. Occurrence. Bay at Boesdal at Stevns Klint, where it was collected from the lower Danian Cerithium Limestone Member of the Rødvig Formation. Description. Protoconch narrow, smooth, not immersed in posterior slope of teleoconch. Teleoconch short ovate in outline, fairly high with nearly straight to slightly concave posterior slope and gently convex anterior slope; lateral flanks nearly straight. No sculpture except for some faint growth lines. Measurements. Height of shell approximately 3.3 mm, width at least 5.3 mm. Remarks. This species closely resembles Cocculina s. l. sp. sensu Kiel (2006) from the Albian of Madagascar with its smooth shell, but may differ in having a relatively higher and less elongated shell with a slightly smoother surface. From all other species of Cocculina it differs in its smooth protoconch and teleoconch, though specimens with nearly smooth shell belonging to the Miocene Cocculina dittmeri (Anderson, 1964) from Northern to Central Europe have been observed (see Janssen 1984).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC1ACFD70FEA0.taxon	materials_examined	Superfamily Symmetrocapuloidea Wenz, 1938	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC4ACFEC9F9BF.taxon	materials_examined	Type species. Emarginula conica Lamarck, 1801 accepted as Emarginula fissura (Linnaeus, 1758) (type by monotypy). Recent, European seas.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC310FDECFB93.taxon	description	Fig. 19 H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC310FDECFB93.taxon	materials_examined	Material. Specimen MGUH 33082, representing a fragmentary mould. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Shell low cap-shaped, apex located close to steep posterior margin. Protoconch 0.7 mm wide, obtusely conical to dome-shaped with at least 1.5 smooth whorls. Protoconch located at apex approximately 1 / 5 shell length from posterior margin, forming wide, somewhat bulbous beak over short, steep posterior margin. Adult shell thin, sub-circular, smooth, the only sculpture represented by weak, irregular growth lines. Measurements. MGUH 33082 is 2.6 mm long and 0.8 mm high. The width is unknown but seemingly nearly as great as the length. Remarks. This taxon differs from the typical Symmetrocapulus by the small size, lack of shell sculpture and the anterior location of the shell apex, and is consequently only tentatively assigned this genus. It may represent the youngest known species of this genus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC0F4FAE7FDF5.taxon	materials_examined	Type species. Patella rugosa Sowerby, 1816, Jurassic deposits in Great Britain (original designation by Dacqué 1934).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDF2B9CC58CFD36FAC0.taxon	materials_examined	Superfamily Fissurelloidea Fleming, 1822	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDE2B9CC7ECFDA7FAEC.taxon	description	Figs 19 I – K 1867 Emarginula coralliorum Lundgren: 19 – 20, pl. 1, figs 5 a – b. 1902 Emarginula coralliorum Lundgren — Ravn: 213, pl. 1, fig. 1. 1903 Emarginula coralliorum Lundgren — Ravn: 394 – 395, 416 – 17. 1940 Emarginula sp. I — Rosenkrantz: 512. 1940 Emarginula sp. II — Rosenkrantz: 512.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93AFFDE2B9CC7ECFDA7FAEC.taxon	materials_examined	Diagnosis. Species of Emarginula characterised by a strongly recurved apex typically situated close to posterior margin, a raised selenizone forming distinct keel with bordering flanges, coarsely cancellate sculpture and by approximately 28 primary radial riblets intercalated with the same number of secondary riblets. Interstices simple pit-like, at least sometimes divided into a pair by a microlirule. Material. The Højerup Member at Stevns Klint is represented by MGUH 33083, ØSM. 10042 - 189, ØSM. 10042 - 334 - b – c, together with 16 specimens with informal numbers (SH. 9, SS. 62. A – B, SR. 730, SR. 733. A – B, SR. 738, SR. 744, SR. 806, SR. 908. A – B, SR. 922, SR. 940, SR. 992. B, SR. 1002, SR. 1061. A – B, SR. 1062, SR. 1075. C and SR. 1118) and eight specimens lacking numbers in the collections of the Natural History Museum of Denmark. The Cerithium Limestone Member is represented by specimen MGUH 33084. Type stratum and type locality. The holotype LO 180 T (Lundgren 1867: fig. 5 a) stored at the Department of Geology, Lund University, Sweden, was collected from the Danian limestone at Limhamn, Sweden. Occurrence. In Denmark E. coralliora ranges from the Højerup Mb. of the upper Maastrichtian Tor Formation at Stevns Klint and up into the lower Danian Stevns Klint Fm. and middle Danian Faxe Formation at Faxe (Ravn 1902). Based on the museum collections and newly collected material, it appears to be extremely rare in the Rødvig Formation (only one specimen). Outside Denmark it is known from the Danian of Belgium, France, Germany, Sweden and Ukraine and from the Thanetian of the Carpathians (Pacaud 2004). Description. Shell patelliform with elongate-oval base. Relative height strongly variable, but typically corresponding to between 40 and 55 % of length. Apex posterior, strongly recurved, situated close to or above posterior margin. Anterior slope broadly convex. Posterior slope concave to nearly straight below apex. Sides of shell nearly straight. Protoconch minute, more or less overhung by succeeding shell. Selenizone raised, forming distinct keel with bordering flanges. Slit short and narrow, length in dorsal view approximating 15 % of anterior slope. Sculpture coarsely cancellate with approximately 28 primary radial riblets intercalated with equal number of weak secondary riblets. Raised concentric ridges reaching around half the thickness of primary radial riblets, but usually markedly stronger than secondary radials. Interstices simple pit-like rectangular, at least sometimes divided into a pair by a microlirule. Margin weakly crenulated by rib extensions. Measurements. The largest specimen from the Danish Cretaceous-Palaeogene boundary is approximately 13 mm long, 7.4 mm wide and 5.7 mm high. It was collected by S. B. Andersen (05.06.1970) from the top of the Højerup Member of the upper Maastrichtian Tor Formation at Holtug, Stevns Klint. Remarks. As already noted by Ravn (1902) the specimens from the Cretaceous-Palaeogene boundary differ from the typical E. coralliora from slightly younger strata by a generally relatively lower shell, though the variation inside the two stratigraphic populations is considerable. Except for this one feature there appear to be no differences. The size also appears unchanged.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93BFFDE2B9CC41EFBA2FA4C.taxon	materials_examined	Type species. Cranopsis pelex A. Adams, 1860 (type by monotypy). Recent, off Japan.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93BFFD92B9CC4B8FD82FCBC.taxon	description	Figs 19 L – Q 2014 Fissurellinid gen. et sp. indet.; Hansen & Surlyk: 339, fig. 5: 18, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93BFFD92B9CC4B8FD82FCBC.taxon	materials_examined	Diagnosis. Shell high with slightly anteriorly turned apex situated at highest point, one third from anterior margin. Trema long, reaching from just behind anterior margin, three quarters the distance to apex, bordered by strong and clearly convex flanks. Sculpture coarsely cancellate with 16 to 19 primary radial ribs. Derivation of name. Refers to its Cretaceous age. Type material. The holotype MGUH 33085 and paratype MGUH 33086 are preserved as internal and external moulds. The paratype is from Rødvig, Stevns Klint. Additional material. The material consists of, in addition to the types, an internal mould in sample SR. 986. A from the uppermost 15 centimetres of the Højerup Member at Rødvig, and an external and internal mould, ØSM. 10042 - 257 - a and b, housed at Østsjaellands Museum, also from the lithified top of the Højerup Member at Rødvig, Stevns Klint. Type stratum and type locality. Lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Knøsen, Stevns Klint. Occurrence. Upper 35 centimetres of the Maastrichtian Højerup Member at Rødvig and somewhere from the upper part of the Tor Formation, and here most likely the top of the Højerup Member at Knøsen, Stevns Klint. Description. Shell high patelliform with bilateral symmetry. Protoconch turned slightly anterior, situated at top of shell approximately one third of shell length from anterior margin. Shell profile slightly concave anteriorly, becoming weakly convex behind apex. Flanks strong, weakly convex, nearly merging at anterior margin. Fissure long, narrow, pyriform, reaching from apex to slightly behind anterior margin with the widest part located half-way; interior side with well-developed, wide septum, reaching three quarters the distance from anterior margin to apex. Shell surface with sculpture of strong radial ribs cancellated by slightly weaker collabral ribs. The 16 to 19 primary radial ribs are intercalated by weaker secondary and tertiary ribs. External sculpture clearly seen on internal side. Measurements. The largest specimen, holotype MGUH 33085, is 3.7 mm long, 3.0 mm wide and 2.5 mm high. Remarks. Cranopsis cretacea n. sp. is somewhat variable with regard to its relative height, which may be distinctly less to slightly more than the width. The genus Cranopsis A. Adams is mostly represented by extant species, with a few fossil taxa known from Neogene deposits (Wenz 1938; Knight et al. 1960). Cranopsis cretacea n. sp. thus becomes the by far oldest known representative from this genus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93DFFD82B9CC52DFB70F8E2.taxon	description	Figs 20 C – I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93DFFD82B9CC52DFB70F8E2.taxon	materials_examined	Material. The Højerup Member is represented by MGUH 33088, MGUH 33089, MGUH 33090 and an external mould without number in the collection of the Natural History Museum. The Cerithium Limestone Member is represented by MGUH 33091. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig and the basal 10 centimetres of the Danian Cerithium Limestone Member at Højerup Church, corresponding to the foraminiferal zone Pα. Description. Protoconch at least 0.3 mm in diameter, consisting of two smooth but angular whorls separated by distinct suture line. Selenizone peripheral, situated just above abapical suture. Teleoconch whorl more flattened with no angulation at wide selenizone. Adapical whorl surface covered by no or a few weak spiral ribs and fine, sharp, densely spaced prosocyrt transverse threads curving backwards towards selenizone; transverse threads numbering between 40 and 60 per whorl. Base of whorl typically covered by faint, very fine, densely spaced transverse threads and fewer faint spiral threads. Number of transverse threads approximately twice that of the transverse ribs on the adapical whorl side. Selenizone bordered by wide ridges. Measurements. The largest specimen, MGUH 33088 is 1.3 mm in diameter and consists of three whorls. Remarks. This taxon differs from Anatoma hedegaardi (Bandel, 1998) and A. rosenkrantzi Schnetler et al., 2001 from the Danish middle Danian in a slightly more step-like shell, the typically marked increase in the number of transverse threads from adapical to abapical side of whorl and in the slightly wider selenizone.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93DFFD82B9CC2FCFBFDFBEE.taxon	materials_examined	Type species. Scissurella crispata Fleming, 1828 (type by monotypy). Recent, North Atlantic.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93DFFDA2B9CC619FB2EFD12.taxon	description	Fig. 20 J	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93DFFDA2B9CC619FB2EFD12.taxon	materials_examined	Material. Specimen MGUH 33092, representing a very fragmentary external mould. Occurrence. Lithified top of the Maastrichtian Højerup Member, Tor Formation, at Rødvig, Stevns Klint. Description. Adapical part of spire whorl weakly convex, nearly horizontal. Base moderately convex. Sculpture on spire whorls consisting of close to 60 sigmoid transverse threads per whorl, threads continuing onto base, becoming slightly opisthocyrt to sigmoid. Transverse threads crossed by weaker, more closely spaced spiral threads. Remarks. The fragmentary state of the specimens makes generic classification rather difficult, but it is here tentatively assigned to Anatoma. Anatoma? sp. B resembles the contemporaneous Anatoma sp. A described above, but differs by the weakly sigmoid transverse threads and the closely spaced spiral threads.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93FFFDA2B9CC285FEEDFBA0.taxon	materials_examined	Type species. Sigaretus carinatus Münster, 1841 (type by original designation). From the Triassic St. Cassian Formation of the Italian Alps.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93FFFA52B9CC5C6FCFAFCD8.taxon	description	Figs 20 K – M 2014 Temnotropis sp. Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A93FFFA52B9CC5C6FCFAFCD8.taxon	materials_examined	Diagnosis. Temnotropis with selenizone located immediately above abapical suture. Last whorl well rounded with convex base. Sculpture fine with granulated spire whorls and basal spiral ribs. Derivation of name. Refers to the famous Stevns Klint coastal exposure at which it has been found. Type material. The holotype MGUH 33093 is an internal and external mould, while paratype MGUH 33094 is a well preserved external and internal mould from the lithified top of the Højerup Member at Rødvig, Stevns Klint. Additional material. ØSM. 10042 - 244 - a. Additionally eight informally numbered and fragmentary specimens preserved as external and internal moulds are found in the collections of the Natural History Museum of Denmark (SR. 754, SR. 781, SR. 786, SR. 787. D and F, SR. 905. A – B, SR. 910, SR. 934. A, SR. 1045. A – C). Type stratum and type locality. Lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Holtug, Stevns Klint. Occurrence. Uppermost part of the Maastrichtian Tor Formation at Rødvig and Holtug Quarry, Stevns. Description. Shell dextral, umboniiform, with 2 ½ fairly quickly widening whorls. Spire low, flatly rounded with shallow suture lines between whorls. Protoconch consisting of around 1 ¼ whorls, seemingly smooth. Teleoconch whorls wide, weakly convex adapically, becoming increasingly more convex laterally; midwhorl with slight angle on which upper side a narrow, slightly raised and gently convex selenizone is found; abapical whorl ramp short and straight, distinguished from lower midwhorl by strong angle. Selenizone partly covered by adapical wall of succeeding whorl. Base convex, with central narrow umbilicus. Aperture oblique, rounded. Teleoconch sculpture on first half whorl consisting of faint transverse threads just inside selenizone; following whorls with beaded fine spiral threads, the beading formed by the crossing of even finer and denser collabral threads describing a moderately opisthocyrt course from adapical suture and out to selenizone. Number of spiral threads above selenizone approximately 27 on last whorl. Base covered by evenly spaced, fine, unbeaded spiral threads spaced much farther from each other than on apical side of whorl; number of spiral threads at least 16 on second whorl, increasing with growth. Growth lines nearly effaced, slightly opisthocyrt while strongly opisthocline. Measurements. Largest specimen 3.4 mm wide and approximately 1.8 mm high. Remarks. The latest Maastrichtian age of the Danish species makes it the youngest of its genus described so far, although a few as yet undescribed specimens have been found in mid-Danian limestone deposits from Denmark as documented by Lauridsen & Schnetler (2014). They appear morphologically very similar to the present species and could possibly be conspecific. The stratigraphically and morphologically closest Cretaceous relative is the Spanish Temnotropis frydai Kiel & Bandel, 2000 from the Campanian. Temnotropis stevniensis n. sp. differs from the latter by the more inflated whorls carrying a finer, denser sculpture with distinctly beaded spiral threads on apical whorl surface and by the convex base covered by at least 16 spiral threads on second whorl. Both Temnotropis frydai and T. stevniensis n. sp. seems with regard to the general shape of the shell, the low selenizone and the presence of a distinct umbilicus more similar to the extant genus Sasakiconcha Geiger, 2006 from the Western Pacific than to the Triassic type of Temnotropis. On the other hand both species seem to lack the foramen formed by converging margins of the slit, and the umbilical wall on at least T. frydai appears to connect with the previous whorl. No preserved apertures and umbilical troughs have so far been uncovered for the Danish taxon, although the mid-Danian taxon do seem to have a rather wide umbilicus (see Lauridsen & Schnetler 2014, Fig. 5). Better material may thus suggest a reassignment of the species to Sasakiconcha or related genera.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A940FFA52B9CC550FE86FB74.taxon	materials_examined	Type species. Pleurotomaria amoena Eudes-Deslongchamps, 1849 (type by original designation). Bajocian (Middle Jurassic) of France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A940FFA42B9CC5A7FC5AFBD4.taxon	description	Figs 20 N – P 1820 Trochilites niloticiformis Schlotheim: 156 – 157. 1902 Pleurotomaria niloticiformis v. Schlotheim sp. [partim] — Ravn: 215 (not Pl. 1, figs 3 and 4). 1903 Pleurotomaria niloticiformis v. Schlotheim sp. [partim] — Ravn: 380 (other species included). 1933 Pleurotomaria niloticiformis (Schlotheim) — Ravn: 25, Pl. 1, fig. 10. 2014 Leptomaria niloticiformis (von Schlotheim 1829) — Lauridsen & Schnetler: 36 – 37, fig. 3 A – D.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A940FFA42B9CC5A7FC5AFBD4.taxon	materials_examined	Diagnosis. Very large, relatively high-spired Leptomaria with nearly straight sides dominated by fairly smooth spiral ribs on later whorls. Umbilicus wide (20 – 25 % of whorl diameter) on internal moulds. Selenizone located approximately 1 / 3 of the distance between the whorl sutures from the adapical suture (emended from Pacaud (2004) and Schlotheim (1820 )). Material. A large specimen MGUH 33095 from the Korsnaeb Mb. infilling at the base of the Cerithium Limestone Mb. at Højerup, and a smaller specimen, ØSM. 10042 - 73 from a Thalassinoides burrow in the Cerithium Limestone at Rødvig. A further specimen with a diameter of 89.5 mm and a height of more than 68 mm has been examined from the private collection of Claus Heinberg. That one came from the Thalassinoides burrows cutting through the Cerithium Limestone Member. Type stratum and type locality. Middle Danian Coral Limestone from Faxe Quarry, Denmark. Occurrence. This species occurs in the lower Danian Cerithium Limestone Member of the Rødvig Fm. at Stevns Klint, where the shells are found in Thalassinoides burrows containing material from the succeeding Stevns Klint Formation. The specimens, however, belong to a now eroded upper layer of the Cerithium Limestone Member as evidenced by their internal moulds consisting of Cerithium Limestone (pers. comm. Claus Heinberg Clausen, Roskilde University, Denmark). It is furthermore reported from the middle Danian Faxe Formation and upper Stevns Klint Formation at Faxe, Denmark (Pacaud 2004; Ravn 1902). Description of Cerithium Limestone material. Shell cyrtoconoid with a spire angle of between 70 and 80 ˚; umbilicus narrow or completely closed, but taking up between 20 and 25 % of total width on internal moulds; whorls with moderately distinct shoulder, resulting in a somewhat stepped shell outline. Whorl surface adapically and abapically of shoulder nearly straight to weakly convex, but may become slightly concave. Shell periphery with moderately pronounced, sharply rounded keel. Abaxial part of whorl fairly weakly convex. Selenizone narrow, taking up approximately 7 % of whorl height measured from suture to suture; selenizone generally indistinct, located immediately above shoulder about 1 / 3 of distance between keel and adapical suture from adapical suture. Selenizone replaced by open slit around 2 / 3 of a whorl from aperture. Aperture wide, subtriangular with flattened outer wall. Teleoconch sculpture above keel consisting of 10 weak spiral threads abapically of selenizone. Adapical part of whorl with about four weak spiral threads on adapical half and a more or less smooth abapical half adjoining selenizone. Growth lines very weak, mainly seen on shell base, but may produce a slightly grainy sculpture on the very early whorls. Measurements. The most complete specimen examined from the Cerithium Limestone Member has a diameter of 89.5 mm and a height in excess of 68 mm (apex missing), but specimens of more than 10 cm in height are known from the middle Danien coral limestone of the type locality. Remarks. This species seem first to have moved out into the Danish Basin shortly before the marked sea level drop, which terminated the muddy Cerithium Limestone environment, and has thus not been found in situ in the Cerithium Limestone but only as redeposited shells in Thalassinoides burrows reaching down from the top of the formation. The material from the Cerithium Limestone Member may deviate slightly from the typical form from the middle Danian limestone at Faxe Quarry, Denmark, by a slightly more high-spired shape (spire angle below 80 ˚) and perhaps relatively finer spiral threads.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A941FFA72B9CC5C4FCE9FD80.taxon	description	Figs 20 Q, 21 A – B 2004 Leptomaria meyeri n. sp. Pacaud: 620 – 623, figs 22, 23 B.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A941FFA72B9CC5C4FCE9FD80.taxon	materials_examined	Diagnosis. Cyrtoconoid; umbilicus narrow or closed; whorls moderately to strongly sigmoid with relatively sharply rounded keel; rather indistinct selenizone located on most convex part of whorl nearly one third of distance between sutures from adapical suture; teleoconch sculpture of well-developed spiral ribs cancellated by weaker collabral threads. Material. MGUH 33097 and two internal and external moulds without numbers are found in the collections of the Natural History Museum of Denmark. Type stratum and type locality. The holotype MNHN R 63509 stored at Muséum national d’Histoire naturelle, Paris, collections du domaine Sciences de la Terre, was found in Danian deposits at Montainville, France. Occurrence. Leptomaria meyeri Pacaud occurs in the lower Danian Cerithium Limestone Member of the Rødvig Fm. at Stevns Klint, where the shells are found in the limestone and in Thalassinoides burrows containing material from the succeeding Stevns Klint Formation. The specimen from a burrow, however, belongs to a now eroded upper layer of the Cerithium Limestone Member as evidenced by its internal mould consisting of Cerithium Limestone (pers. comm. Claus Heinberg Clausen, Roskilde University, Denmark). Additionally it is known from the middle Danian Faxe Formation, Denmark, and from Danian deposits at Montainville and Bray-et-Lü in France (Pacaud 2004). Description of Cerithium Limestone material. Shell cyrtoconoid with a spire angle of approximately 80 ˚; whorls sigmoid in outline with convex adapical part, changing to distinctly concave on central to abapical part outside selenizone; abaxial part of whorl fairly weakly convex. Last whorl with moderately pronounced, sharply rounded keel. Selenizone narrow, generally very indistinct, located on most convex part of whorl between 1 / 4 and 1 / 3 the distance between sutures from adapical suture. Selenizone replaced by open slit on last whorl. Umbilicus on internal moulds taking up between 1 / 5 and ¼ of whorl diameter. Teleoconch sculpture above keel consisting of moderately coarse spiral threads of varying strength, cancellated by fine collabral threads, giving the surface a slightly granular appearance. Number of spiral threads increases rapidly on first whorls, but keep around 3 spirals adapically of selenizone. Abapical part of whorl has up to at least 13 spirals between selenizone and abapical suture. Sculpture below keel consisting of weak spiral threads. Measurements. MGUH 33097 is approximately 14.5 mm wide and 12.5 mm high. Remarks. Leptomaria meyeri Pacaud is distinguished from the Danian L. penultima (d’Orbigny, 1850 a) from northern Europe, which was redescribed by Pacaud (2004), in the generally more step-like outline of the spire and in the generally more pronounced teleoconch sculpture. It differs furthermore from the large L. niloticiformis (Schlotheim, 1820) treated above in the smaller size and the more adapical location of the selenizone. In contrast to what was postulated by Pacaud (2004) no difference may be found between the two species in the relative width of the umbilicus, at least on internal moulds found in Denmark.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A942FFA72B9CC3F8FADFFCF2.taxon	materials_examined	Type species. Trochus reticulatus Sowerby, 1821 (type by original designation). Kimmeridgian (Late Jurassic) of England.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A942FFA62B9CC22EFCEDFF18.taxon	description	Figs 21 C – H 2014 Bathrotomaria sp. Hansen & Surlyk: 335, fig. 5: 20, tab. 2.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A942FFA62B9CC22EFCEDFF18.taxon	materials_examined	Diagnosis. Species of Bathrotomaria characterized by a strong, thin peripheral keel, a low spire with sharp, well separated threads and by the fine growth lines changing to transverse ribbing at transition to wide umbilicus. Derivation of name. In honour of conservator Sten Lennart Jakobsen at the Geological Museum of Copenhagen, Denmark, who has been extremely helpful throughout this project. Type material. Holotype MGUH 33098 is a nearly complete internal and external mould but for extensive early diagenetic dissolution on the apical side. Paratypes MGUH 33099 and MGUH 33100 from the lithified top of the Tor Formation (probably lower Højerup Member) at Holtug, Stevns, represent very fragmentary external moulds. Type stratum and type locality. Slightly lithified upper ten centimetres of the Maastrichtian Sigerslev Member, Tor Formation, at Kulsti Rende, Stevns Klint. Occurrence. Top Sigerslev Member to basal Højerup Member of the Maastrichtian Tor Formation at Holtug and Kulsti Rende, Stevns Klint. Description. Protoconch not known. Teleoconch broadly conical, nearly twice as wide as high. Whorls sigmoid in outline with wide, only slightly inclined adapical ramp divided by weak angle from concave face. Selenizone located at mid-whorl on angle between adapical ramp and face of whorl, terminating in exhalant slit a quarter of a whorl from aperture, the width of which takes up nearly 20 % of whorl width. Periphery with very broad, attenuated keel delimiting strongly convex base. Transition from base to wide umbilicus gradual; width of umbilicus taking up 20 % of shell width. Aperture subrectangular, width taking up 44 % of shell width on holotype, corresponding to 142 % of aperture height. Whorl sculpture consisting of fine spiral threads crossed by deeply opisthocyrt collabral threads with V-shaped sinus at selenizone; small tubercles occur at connecting points between spiral and collabral threads. Ramp carrying around four spiral ribs, while six to seven are observed on whorl face. Two additional but stronger ribs border the selenizone. Base with fine opisthocyrt growth lines on abaxial part, changing to low transverse ribs at transition to umbilicus. Spiral threads may occur. Measurements. Holotype MGUH 33098, 11.1 mm high and 20.9 mm wide. Remarks. Bathrotomaria jakobseni n. sp. differs from the Late Cretaceous B. praepyrenaica Astre, 1935 from southwestern France by the distinctly lower and less step-like spire. It is separated from the Senonian B. regalis (Roemer, 1841) and B. perspectiva (Mantell, 1822) from Germany and Great Britain, respectively, by the strong, thin keel and the transverse ribbing at the transition to the umbilicus. Bathrotomaria jakobseni n. sp. strongly re- sembles the Early Albian Pleurotomaria shenleyensis Cox, 1960 b, but differs in the main generic character, namely the location of the selenizone at the ramp angle and not well below it. Nevertheless the strong resemblance suggests that there may be problems with the present generic division.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA12B9CC2E4FE2EFA7B.taxon	description	Fig. 21 I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA12B9CC2E4FE2EFA7B.taxon	materials_examined	Material. Specimen MGUH 33101, a natural mould of parts of the last whorl. Occurrence. The specimen comes from the lithified top of the Maastrichtian Højerup Member, Tor Formation, in Holtug Quarry. Description. Last whorl convex with at least four sharp, beaded spiral ribs interconnected by oblique, discontinuous and sharply narrow transverse cords. Beading on keel rib spiny, becoming less pronounced on base ribs and adapical rib. Beading on adapical spiral rib much finer and more densely spaced than following ribs. Measurements. The whorl fragment is 3.65 mm wide. Remarks. The sculpture and whorl profile is very similar to that of the extant Bathybembix bairdii (Dall, 1889), which is why the present taxon is tentatively assigned here in spite of a current oldest generic record from the late Eocene (see Hickman & McLean 1990). Better material, though, may still lead to a reassignment to Solariella or perhaps Pareuchelus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA12B9CC3B6FBB1FCB5.taxon	materials_examined	Type species. Bembix aeola Watson, 1879 (type by original designation). Recent, Japan.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA12B9CC77CFBFCF96D.taxon	materials_examined	Type species. Monodonta tinei Calcara, 1839 (type by monotypy). Recent, the Mediterranean.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA02B9CC7ADFA34FE14.taxon	description	Figs 21 J – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A944FFA02B9CC7ADFA34FE14.taxon	materials_examined	Material. MGUH 33102 is an external mould. Occurrence. Upper metre of the lower Danian ‘ dead layer’ in Vokslev Quarry, Northern Jutland. Description. Protoconch planispiral to slightly heterostrophic, submerged, nearly 0.4 mm in width, with 1 ¼ moderately convex whorls, appearing smooth. Transition to teleoconch marked by appearance of transverse ribs. Teleoconch with strongly convex, low whorls separated by fairly deep suture located slightly below periphery of preceding whorl. Base convex. Aperture rounded with concave columella. No columellar folds or denticles present on preserved part. Teleoconch sculpture dominated by sharp and strong prosocline and prosocyrt transverse ribs, numbering 22 to 23 per whorl on initial whorls. Transverse ribs crossed by five or six weak spiral ribs, becoming stronger on later whorls. Connecting points marked by tubercles. Measurements. MGUH 33102 1.9 mm high and 1.3 mm wide, consisting of parts of 3 ½ teleoconch whorls in addition to protoconch. Remarks. Danilia sp. is easily distinguished from Danilia kosslerae Kiel & Bandel, 2001 b from the Campanian of Spain by the convex whorls and the nearly planispiral protoconch. It has a passing resemblance to the middle Danian Danilia quadricordata Ravn, 1933 from Denmark, but differs by the more convex whorls and finer spirals.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A945FFA02B9CC376FEBAFD55.taxon	materials_examined	Type species. Chilodonta clathrata Étallon, 1859 (type by subsequent designation of Bayan 1874: p. 335). Late Jurassic, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A945FFA32B9CC3B6FDBFFF19.taxon	description	Figs 21 L – Q 1861 Turbo rudis Binkhorst: 49, pl. III, fig. 8 a – b. 1861 Turbo detritus — Binkhorst: 45 – 46, pl. III, fig. 10 a – b. 1861 Turbo bidentatus — Binkhorst: 46, pl. III, fig. 9. 1898 Trochus (Craspedotus) rudis Binkhorst — Kaunhowen: 38 – 39, pl. II, figs 1 – 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A945FFA32B9CC3B6FDBFFF19.taxon	materials_examined	Diagnosis. Low conical; whorls weakly convex, keeled; aperture with three strong denticles on outer lip, a parietal denticle and strong columellar fold; whorl sculpture relatively fine, adapically consisting of well developed, sharp transverse ribs cancellated by weaker spiral ribs and abapically with transverse ribs or just fine and densely spaced collabral threads cancellated by coarser spiral ribs. Material. The Danish material consists of MGUH 33103 to MGUH 33106, the informally numbered SR. 728, SR. 756, SR. 924. A, SR. 954, SR. 1069. A and SR. 1071. A – B, together with five internal and external moulds lacking numbers in the old collections of the Natural History Museum of Denmark. Type stratum and type locality. The holotype figured by Binkhorst (1861) on plate III: 8 a – b was collected from the upper Maastrichtian stratotype at Limburg, the Netherlands. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Stevns Klint (Rødvig, Skeldervig, Mandehoved, Holtug Quarry and Kulsti Rende). A single specimen coming from the lower Danian Cerithium Limestone Member at Stevns Klint has been examined in the collections of C. Heinberg, Denmark (Fig. 21 Q), but it may be reworked from the Maastrichtian deposits. Outside of Denmark it is known from the Maastrichtian of Poland and the Netherlands (Abdel-Gawad 1986). Description. Shell small, low conical with a spire angle of between 65 ˚ and 75 ˚ and up to around five whorls. Protoconch nearly planispiral, consisting of approximately one smooth whorl with a small initial whorl. Protoconch width around 0.25 mm. Teleoconch sharply rounded at midwhorl, keel separating weakly convex adapical and abapical side; suture distinct, located just below keel of preceding whorl with the two whorls meeting at an angle of approximately 90 ˚. Shell narrowly umbilicate in smaller specimens, becoming covered by parietal callus in larger shells. Aperture with a fairly strong columellar fold or elongated denticle on lower columella, a strong parietal ridge or denticle and three strong denticles on the outer lip distributed with one above keel and two below. Teleoconch sculpture adapical to keel consisting of strong, sharp, densely spaced transverse ribs with 5 to 8 weaker spiral ribs (increases with size), sometimes intercalated with secondary ribs. Abapical side with spiral ribs cancellated by fine, densely spaced collabral threads. Measurements. The most complete Danish specimen MGUH 33103 is 6.0 mm wide and 5.8 mm high with about 5 1 / 3 teleoconch whorls in addition to one protoconch whorl. Remarks. The type material of Chilodonta rudis (Binkhorst, 1861) from the Netherlands appears in general to be characterized by coarser transverse sculpture on the base of the last whorl than is typical for the Danish material, but otherwise they seem identical.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A946FFA32B9CC0D9FF57FDF3.taxon	materials_examined	Type species. Trochus binodosus Münster, 1841 (type by original designation). Late Triassic St. Cassian Formation, Italian Alps.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A946FFA32B9CC32CFC47F9AB.taxon	description	Figs 21 R – T	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A946FFA32B9CC32CFC47F9AB.taxon	materials_examined	Diagnosis. Spire whorls rounded, not angular. Lamellar to cord-like transverse ribs present within half a whorl from the protoconch. Later whorls also with sharp spiral ribs, forming strong reticulate pattern with transverse ribs. Derivation of name. In reference to the delicate ribbing of the teleoconch. Type material. Holotype MGUH 33108 is an external mould of the spire. Paratype MGUH 33107 is an external mould of the transition from periphery to base. The paratype comes from the same locality and stratigraphic level as the holotype. Additional material. The informally numbered SR. 809, SR. 993. A – B and D, SR. 1116, and a mould lacking number, all from the collections of the Natural History Museum of Denmark. Type stratum and type locality. Lithified top of the Maastrichtian Højerup Member at Rødvig, Stevns Klint. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Protoconch low heterostrophic with approximately 1 ¼ smooth and strongly convex whorls and deep suture. Height measuring nearly 0.2 mm, while width reaches 0.4 mm. Transition to teleoconch sharp. Teleoconch turbiniform with three strongly convex whorls and deep suture located just adapically of strongly angular transition to flattened base of preceding whorl. Aperture not known. First teleoconch whorl covered by 14 lamellar and slightly prosocline transverse ribs, the rib number increasing slowly on later whorls. Transverse ribs crossed by two to four sharply defined spiral cords, forming spiny tubercles at connecting points. Angle between whorl side and base of last whorl carrying yet another strong spiral cord. Whorl base with at least one strongly developed spiral rib and no transverse ribs. Measurements. Holotype MGUH 33108 composed of just over four whorls and measuring 2.3 mm in width and over 2.0 mm in height. Remarks. Eucycloscala gracilicostatus n. sp. resembles the middle Danian Eucycloscala ultima Ravn, 1933 from Denmark but for the strongly developed spiral cords on the later whorls. It differs from the Campanian Eucycloscala cretacea Kiel & Bandel, 2001 b from Spain by the strongly rounded, non-angular spire whorls, a first teleoconch whorl not flattened around the protoconch, and by the sharp ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A946FFA32B9CC63FFD86F80E.taxon	materials_examined	Type species. Phorcus margaritaceus Risso, 1826 accepted as Phorcus richardi (Payraudeau, 1826) (type by subsequent designation). Recent, the Mediterranean.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC1D4FDE7FD80.taxon	description	Figs 21 U, 22 A	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC1D4FDE7FD80.taxon	materials_examined	Material. Specimen MGUH 33109, an external mould of the spire. Occurrence. Lithified top of the Maastrichtian Tor Formation at Holtug Quarry. Description. Protoconch smooth, weakly convex with flattened whorls. Transition to teleoconch gradual. Teleoconch with low cyrtoconoid spire and weakly convex whorls separated by distinct suture. Whorls in cross-section broadly rounded without keel; smooth. Aperture and abapical side of whorl not known. Measurements. Specimen MGUH 33109 is 3.1 mm high. Remarks. Phorcus sp. differs from the middle Danian Phorcus carinatus (Ravn, 1933) from Denmark by the lack of a carination on the whorls and a relatively higher teleoconch shell. It differs from the two Paleocene species of Phorcus from Greenland, presented under the names Osilinus sp. 1 and 2 by Kollmann & Peel (1983), by the higher shell and lack of spiral striation.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC726FBA6F924.taxon	materials_examined	Type species. Solariella maculata Wood, 1842 (type by monotypy). Pliocene, England.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC287FC5BFA34.taxon	description	Figs 22 B – C	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC287FC5BFA34.taxon	materials_examined	Material. Specimen MGUH 33110 is an external mould of the apex. Occurrence. The lower Danian ‘ dead layer’ at Vokslev Quarry, Northern Jutland. Description. Low-spired shell with apex relatively close to posterior margin. Protoconch moderately highspired with 1 2 / 3 smooth, convex whorls and a moderately wide initial chamber. The width is 0.3 mm. Transition to teleoconch marked by slight lessening in whorl convexity and a beginning of growth lines. Teleoconch whorls convex, relatively high, with well marked suture line. Whorl angle close to 110 ˚. Whorl width increasing moderately fast. Transition to base seemingly sharp. Teleoconch surface smooth with weak, prosocyrt growth lines. Measurements. MGUH 33110 is 1.4 mm wide and approximately 0.4 mm high, consisting of 1 ½ teleoconch whorls and protoconch. Remarks. The taxon is clearly distinguished from the Danish middle Danian Stomatella sp. sensu Lauridsen & Schnetler (2014) by the marked suture line and convex, well defined whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFA22B9CC3B0FF52FC9A.taxon	materials_examined	Type species. Stomatella auricula Lamarck, 1816 (subsequent designation by Anton 1838). Recent, central and western Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFAC2B9CC643FCE3FC0E.taxon	description	Figs 22 D – F 2014 Solariella sp. Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A947FFAC2B9CC643FCE3FC0E.taxon	materials_examined	Material. MGUH 33111, MGUH 33112, ØSM. 10042 - 272 - a and an external mould of spire without number in the collections of the Natural History Museum of Denmark. Occurrence. Top Højerup Member of the Maastrichtian Tor Formation at Rødvig, Højerup, Holtug Quarry and north of Harvig Stige at Stevns Klint, Denmark. Description. Protoconch smooth with deep sutures and a moderately high spire with two or three whorls. Teleoconch whorls broadly convex, angular with strong and sharp spiral cords, three of which form a wide obsolete keel; collabral threads or ridges very faint on older whorls, becoming more pronounced on later whorls. Upper suture situated between second and third keel rib of preceding whorl. Base and aperture unknown. Measurements. Largest shell approximately 6.5 mm wide and 4.8 mm high. Remarks. This taxon differs from S. ladrierei (Briart & Cornet, 1887) and S. multilineata (Briart & Cornet, 1887) from the Danian of Belgium by the more angular whorls with their coarser spiral cords and by the presence of distinct collabral ribs. It is distinguished from S. vincenti Cossmann, 1915 by the larger number of spiral ribs and less stepped spire and from S. briarti (Vincent in Cossmann 1915), also from the Danian of Belgium, by the more keeled outline of the whorl with its stronger collabral threads.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A949FFAC2B9CC5CAFA9AFA97.taxon	materials_examined	Type species. Helix serpuloides Montagu, 1808 (subsequent designation by Gray 1847: p. 152). Recent, Great Britain. Remarks. Three Campanian species S. wareni, S. suturata and S. torallensis from Northern Spain were described and tentatively assigned to this genus by Kiel & Bandel (2002 a). They clearly do not belong here as their sutures are too little impressed, separating rather abutting whorls and they have a rather strongly reinforced aperture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A949FFAC2B9CC488FBECF818.taxon	description	Figs 22 G – I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A949FFAC2B9CC488FBECF818.taxon	materials_examined	Material. MGUH 33113, MGUH 33114, ØSM. 10042 - 194 and a mould in the informally numbered sample SR. 1101 deposited at the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Rødvig and Holtug Quarry, Stevns Klint. Description. Shell low spired and umbilical, consisting of at least four thin-shelled and loosely coiled whorls separated by deep suture. Transition between protoconch and teleoconch obscure. Whorls smooth, highly convex with weak peripheral keel on first whorl fading away on last; whorl expansion initially slow, becoming pronounced on last whorl. Base strongly convex with wide and deep umbilicus, width approximating 1 / 5 shell width. Aperture circular in outline, height corresponding to around 4 / 5 of shell height. Measurements. MGUH 33114 is 10.2 mm wide and at least 6.8 mm high with two preserved teleoconch whorls. Remarks. Skenea sp. looks quite a lot like the extant type species Skenea serpuloides (Montagu) except for the smooth whorls and lower spire. It differs from the Middle Palaeocene Circulus sp. of Kollmann & Peel (1983: fig. 38) from Greenland by the smooth and more strongly expanding teleoconch whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAF2B9CC073FBA6FC97.taxon	description	Figs 22 J – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAF2B9CC073FBA6FC97.taxon	materials_examined	Material. Specimen MGUH 33115 is an incomplete external mould. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Holtug Quarry, Stevns Klint. Description. Shell very low trochiform with flattened spire. Nearly planispiral protoconch 0.55 mm wide, consisting of approximately two smooth and only slowly widening flat whorls; whorls separated by deep sutures. Transition to teleoconch not visible. Teleoconch with around two fairly rapidly widening, adapically flattened whorls. Deep suture situated just above sharply rounded periphery of preceding whorl. Base weakly convex. Aperture and umbilical area unknown. Teleoconch sculpture consisting of a single distinct adapical furrow. Measurements. Specimen MGUH 33115 is 1.6 mm high and at least 2.4 mm wide, consisting of 3 3 / 4 whorls. Remarks. This taxon differs from the middle Danian Leucorhynchia marginata Ravn, 1933 from Denmark by a slightly higher shell and lower protoconch, but seems otherwise quite similar.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAF2B9CC1D7FB86FF15.taxon	materials_examined	Type species. Leucorhynchia caledonica Crosse, 1867 (type by monotypy). Recent, Japan.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAF2B9CC543FE0BFB6A.taxon	materials_examined	Type species. Tectus pagodalis Montfort, 1810 accepted as Tectus mauritianus (Gmelin, 1790) (type by original designation). Recent, South Africa.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAE2B9CC591FC68FDDC.taxon	description	Figs 22 L – N 2014 Tectus sp. Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94AFFAE2B9CC591FC68FDDC.taxon	materials_examined	Diagnosis. High-spired with weakly convex whorls. Columella nearly straight, oriented slightly oblique to axial line; columellar fold nearly effaced. Transverse ribs short and strong, numbering around 10 per whorl. Spirals finely tuberculated, present on base as well as spire whorls. Derivation of name. Refers to the generally worn and rather ugly appearance of the specimens at hand. Type material. Holotype MGUH 33116 is an external mould with protoconch. Paratype ØSM. 10042 - 327 - a and b is an external and internal mould of the lower teleoconch whorls from the same stratigraphic level north of Kulsti Rende, Stevns Klint. Additional material. Two weathered moulds (ØSM. 10042 - 182 - a – b, ØSM. 10042 - 253 - a – b). Type stratum and type locality. Lithified top of the Højerup Member at Rødvig, Stevns Klint. Occurrence. Lithified top of the Maastrichtian Højerup Member at Stevns Klint. Description. Protoconch obtusely conical to dome-shaped paucispiral with close to two smooth whorls separated by shallow suture. Protoconch width approximating 0.6 mm. Teleoconch trochiform with weakly convex whorls and shallow suture. Whorl height corresponding to nearly half the width of early whorls, decreasing to one third on later whorls. Base flattened with no umbilicus. Aperture subrectangular with short and weakly convex columellar lip, oriented slightly oblique to axial line. Columellar fold nearly completely effaced. Teleoconch sculpture dominated by around 10 strong, bulbous transverse ribs per whorl on abapical whorl half. Spire whorls additionally covered by fairly coarse and irregular spiral threads, which are heavily tuberculated due to cancellation by slightly sigmoid and strongly prosocline growth lines. Base finely tuberculated due to interference between strongly prosocline and opisthocyrt growth lines and finer, more densely spaced spiral threads. Measurements. Holotype MGUH 33116 is 2.0 mm high and 1.6 mm wide, consisting of protoconch and about 3 ½ teleoconch whorls. Paratype ØSM. 10042 - 327 - a has a width of at least 20.3 mm. Remarks. Tectus? indecorus n. sp. to some degree resembles the Campanian Tectus sensuyi (Vidal, 1921) from Torallola in Spain, but is easily distinguished by the fewer transverse ribs and the presence of spiral sculpture on the base. Of the two species encountered in the early Danian limestone of Belgium it is most similar to Tectus carinadentatus (Briart & Cornet, 1887), but differs by the absence of a strong columellar fold, the higher spire and by the coarser and sparser transverse sculpture. The relatively high whorls and that the transverse whorls appear better developed on the adapical whorl surface than at the keel may suggest that the species do not belong in this genus, though a few recent species do feature somewhat similar characteristics.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94BFFAE2B9CC646FB5BF884.taxon	materials_examined	Type species. Turbo squamulosus Lamarck, 1804 (type by original designation). Eocene, Paris Basin, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94BFFAE2B9CC224FBDDFC26.taxon	materials_examined	Type species. Turbo sanguineaum Linnaeus, 1758 (type by monotypy). Coast of Catalane, Spain.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94BFFAE2B9CC55AFD9BF9CA.taxon	description	Figs 22 O – P	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94BFFAE2B9CC55AFD9BF9CA.taxon	materials_examined	Material. Specimen MGUH 33117 is a very fragmentary external and internal mould of the final 2 ½ whorls. Occurrence. The lower Danian Cerithium Limestone Member at Skeldervig, Stevns Klint. Description. Teleoconch fairly low conical; whorls highly convex, nearly round in cross-section, separated by shallow suture. Aperture round. Teleoconch sculpture consisting of numerous weak and fine spiral bands, more or less equally developed on spire whorls as on base. A beaded spiral band numbering perhaps 50 weak beads on last whorl runs adjacent to adapical suture. Weaker beading may be seen on several more adapical spiral bands. Measurements. Tube diameter of last whorl 6.3 mm. Whorl radius reaches at least 6.4 mm. Remarks. The Danish taxon resembles somewhat the Palaeocene Homalopoma sp. 1 of Kollmann & Peel (1983: p. 30 – 31) from Greenland, but differs by the weak subsutural constriction of the whorls and the weakly beaded adapical spiral bands. It differs from the Campanian Homalopoma minimum (Quintero & Revilla, 1966) from Spain by the finer spiral bands.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94CFFA92B9CC298FAC3FBD1.taxon	materials_examined	Type species. Turbo cruentata Mühlfeld, 1824 (subsequent designation by Woodring (1928: p. 422). Recent, West Indies.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94CFFA92B9CC1D4FCFCFCD8.taxon	description	Fig. 22 Q	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94CFFA92B9CC1D4FCFCFCD8.taxon	materials_examined	Material. MGUH 33118 and a very fragmentary external mould of the last whorl stored in the old collections at the Natural History Museum of Denmark. Occurrence. Lithified top of the Højerup Member of the upper Maastrichtian Tor Formation at Rødvig and Højerup Church, Stevns Klint. Description. Shell moderately bulbous. Last whorl convex with evenly rounded periphery and weakly convex base; umbilicus deep. Growth lines prosocyrt and prosocline. Sculpture consisting of approximately 25 fine and closely spaced, somewhat flattened spiral ribs, changing from relatively high and rounded, widely spaced adapical ones over closely spaced peripheral ribs to wider and lower abapical ones. Base with unevenly distributed, coarse growth ridges. Measurements. Whorl diameter of MGUH 33118 is 2.8 mm. Remarks. The specimens resemble the Danian C. firketi (Briart & Cornet, 1887) from Belgium as illustrated by Glibert (1973: pl. 1: 22), but has a more rounded periphery; relatively wider spacing of the adapical spiral ribs and prosocyrt abapical growth lines. It differs from the Maastrichtian C. (C.) granulatus (Kaunhowen, 1898) from Belgium by the smooth adapical spiral ribs. It differs from the Late Cretaceous C. (C.) americanus Wade, 1926, C. (C.) nudus Sohl, 1960 and C. (C.) paucispirilus Sohl, 1964 b from the USA by the higher number of spiral ribs, which lack beading adapically, and by the narrower umbilicus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94CFFA82B9CC5C9FDFFFE84.taxon	description	Figs 22 R – U, 23 A	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94CFFA82B9CC5C9FDFFFE84.taxon	materials_examined	Diagnosis. Shell relatively high; planispirally coiled protoconch featuring convex whorls; teleoconch with three to four strong, beaded spiral cords above whorl periphery. Derivation of name. In honour of Sten Lennart Jakobsen, conservator at the Geological Museum of Copenhagen, who has been a tremendous help to this project. Type material. Holotype MGUH 33119 is a natural external and internal mould of the spire. Paratype MGUH 33120 is an external mould of the base and aperture from Holtug Quarry, possible from one of the Korsnaeb Member infilled Thalassinoides burrows in the Cerithium Limestone Member, but the gastropod is filled out with material from the Cerithium Limestone. Type stratum and type locality. Lower Danian Cerithium Limestone Member at Holtug Quarry, Stevns Klint. Occurrence. Lower Danian Cerithium Limestone Member of the Rødvig Formation at Holtug Quarry, Stevns Klint. Description. Shell relatively high with planispirally coiled protoconch. Protoconch 1.6 mm wide, consisting of 2 smooth or weakly transversely striated and slightly depressed whorls. Teleoconch turbiniform with deeply impressed sutures and strongly convex whorls; a weak keel may be present; base of shell weakly flattened with deep, but narrow umbilicus. Aperture circular with thick and smooth, flattened lip and low varix. Teleoconch surface heavily sculptured with eight to ten beaded or spiny spiral cords, three to four strong cords on adapical side, one at the periphery and the rest, which generally are much finer and more closely spaced, on the base. Spiny tubercles hollow in growth direction; restricted to primary three spiral ribs. Spiral cords on base decreasing in strength towards umbilicus, which is bounded by a single more strongly developed beaded spiral rib. Cords crossed by very fine and densely spaced transverse striae. Measurements. Holotype MGUH 33119 is 9.2 mm high and 8.6 mm wide. Remarks. Arene jakobseni n. sp. differs from the Campanian A. mcleani Kiel & Bandel, 2002 a from Spain and the Maastrichtian A. truncatospaera Sohl, 1998 from Jamaica in the sparser and spinier sculpture and the more strongly convex whorls. It closely resembles the middle Danian Arene depressa (Ravn, 1933) from Denmark, but differs by the distinctly larger size, the coarser and sparser teleoconch sculpture and possible by a narrower umbilicus, though this may relate to ontogeny.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94DFFA82B9CC3DDFAC6FD0E.taxon	materials_examined	Type species. Ataphrus crassus Gabb, 1869 (type by original designation). Late Cretaceous, Martinez in California, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94DFFA82B9CC272FBA5F853.taxon	description	Figs 23 B – E 2014 Ataphrus (Ataphrus) sp. Hansen & Surlyk: 339, fig. 6: 6, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94DFFA82B9CC272FBA5F853.taxon	materials_examined	Diagnosis. Shell with convex spire, flattened top, no umbilicus and a relatively strong spiral sculpture. Derivation of name. In reference to the type locality Knøsen at Stevns Klint. Type material. Holotype MGUH 33121 is an external and internal mould. Paratype MGUH 33122 is an external and part of internal mould from Stevns Klint, while paratype MGUH 33123 is an external mould from Holtug, Stevns Klint. Additional material. ØSM. 10042 - 253 - c – d, the informally catalogued specimens SR. 753. A – B, SR. 757, SR. 785. A – B, SR. 971, SR. 981, SR. 983, SR. 1018, SR. 1071. A, SR. 1108. A, and a mould without number placed in the old collections of the Natural History Museum of Denmark. Type stratum and type locality. The lithified top of the upper Maastrichtian Højerup Member at Knøsen, Stevns Klint. Occurrence. Uppermost Maastrichtian at Stevns Klint in Denmark, where it has been collected from the lithified top of the Tor Formation at Holtug Quarry, Knøsen, Rødvig and in the cove at Boesdal. Description. Shell globosely turbiniform. Protoconch flattened paucispiral to planispiral, approximately 0.6 mm wide, consisting of two whorls; surface nearly smooth but for faint spiral ribs, which slowly increases in number and distinctiveness. Transition to teleoconch gradual. Teleoconch whorls evenly rounded with very faint keel. Base slightly flattened. Last whorl distinctly downturned with strong suture passing down below keel of preceding whorl. Surface covered by weak collabral threads and moderately weak spiral threads, which become slightly stronger around keel. Measurements. Largest specimen, holotype MGUH 33121, is 4.4 mm wide and 4.5 mm high. Remarks. Ataphrus (Ataphrus) knoesensis n. sp. differs from most species within this subgenus in the presence of spiral sculpture. It closely resembles the contemporaneous Ataphrus (Ataphrus) compactus Gabb, 1869 from California, which has been refigured by Stewart (1927), but the abapical turn of the last whorl appears more pronounced, the base is not as convex with a sharper transition to the whorl flank, and the spiral threads are coarser and slightly stronger, especially around the keel. This species is distinguished from Ataphrus sp. sensu Kollmann & Peel (1983) from the Palaeocene of Greenland by the weaker keel, slightly stronger whorl inflation and a distinctly downturned last whorl. It is distinguished from the Jurassic A. (A.) marschmidti Gründel & Kaim (2006) from Poland in the lack of an umbilicus, stronger spiral threads and more convex spire.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAB2B9CC34CFDFDFD9E.taxon	materials_examined	Type species. Cerithium adansonii Bruguière, 1792 accepted as Cerithium nodulosum Bruguière, 1789 (subsequent designation by ICZN 1978). Recent, Ellice Islands.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAB2B9CC383FE1BF9AA.taxon	description	Figs 23 F – H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAB2B9CC383FE1BF9AA.taxon	materials_examined	Material. MGUH 33124, MGUH 33125, MGUH 33126, and two fragmentary specimens without number located in the old collections of the Natural History Museum of Denmark. Occurrence. It ranges from the Lithified top of the Maastrichtian Tor Formation and up into the lower Danian Korsnaeb Member infilled Thalassinoides burrows in the Cerithium Limestone at Stevns Klint. Description. Protoconch unknown. Teleoconch whorls flat, separated by narrow suture located immediately abapical to keel ridge. Wide and low varices scattered unevenly on spire, the last occurring approximately opposite aperture on last whorl. Keel carrying two sharply developed ridges, adapical one only visible on last whorl. Base flattened. Aperture relatively small, strongly rounded adaxially against only slightly convex columella; inner lip well developed with strong parietal ridge forming continuation of abapical keel ridge. Columellar fold absent. Teleoconch sculpture on younger whorls consisting of weak transverse ribs crossed by spiral lirae, forming up to five weak, beaded spiral ribs. Adult teleoconch whorls nearly smooth but for fine spiral lirae and a weak spiral band of 40 to 50 tubercles per whorl adjacent to adapical suture. Growth lines slightly opisthocyrt, forming very fine and dense pattern on whorl surface, on adapical keel ridge developing into very fine and dense beading. Base covered by spiral ribs of unequal strength. Measurements. Largest specimen, MGUH 33124, 10.5 mm in diameter with a last whorl height approximating 13 mm. Remarks. Because of the flattened and weakly sculptured whorls this taxon is not a Cerithium sensu stricto, but belongs most likely to one of the many subgenera erected for this genus. Its affinity with the genus is clearly seen from the aperture with its distinctly rounded outline, strong parietal lip, nearly centrally placed columella, well developed parietal ridge and rather weakly opisthocyrt growth lines. It is distinguished from the genus Metacerithium Cossmann, 1906 by the presence of varices, the strong parietal ridge and the rather weakly forwardly reaching sinus of the basal growth lines.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAB2B9CC7D6FBF1F914.taxon	materials_examined	Type species. Murex aluco Linnaeus, 1758 (type by original designation). Recent, Philippines.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAA2B9CC604FF47FE31.taxon	description	Fig. 23 I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94EFFAA2B9CC604FF47FE31.taxon	materials_examined	Material. Specimen MGUH 33127 is an external mould of a teleoconch. Occurrence. Lithified top of the Maastrichtian Højerup Member at Holtug Quarry, Stevns Klint. Description. Teleoconch whorls flattened, dextrally coiled, nearly twice as wide as high and separated by fairly deep and V-shaped suture. Transition to flattened whorl base sharp, angular. Teleoconch sculpture consisting of three narrow spiral ribs bordered by wide and shallow spiral furrows; central furrow deepest. Very weak and strongly opisthocline transverse ribs may be discerned between two abapical spiral ribs. Measurements. MGUH 33127 is 4.0 mm high and 1.6 mm wide, consisting of 5 teleoconch whorls. Remarks. Except for the dextrally coiled whorls and more pronounced suture the specimen resembles Epetrium s. l. skeldervigensis n. sp. described below, and it is therefore possible that it should be assigned to the triphorids instead.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFAA2B9CC77EFB96F94D.taxon	materials_examined	Type species. Melania lineolata Gray in Griffith & Pidgeon, 1833 accepted as Cerithium obtusum Lamarck, 1822 (subsequent designation by Pilsbry & Harbison (1933: p. 115). Recent, Indian Ocean and Indonesia.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFAA2B9CC369FBD0FD63.taxon	materials_examined	Type species. Cerithium striatum Bruguière, 1792 (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFAA2B9CC39EFDE3FA7A.taxon	description	Figs 23 J – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFAA2B9CC39EFDE3FA7A.taxon	materials_examined	Material. Specimen MGUH 33128 is an external mould of the teleoconch. Occurrence. Lower Danian Cerithium Limestone Member at Skeldervig, Stevns Klint. Description. Whorls weakly convex, separated by shallow but well marked suture. Whorl height corresponding to less than half the width. Whorl periphery located on lower third of whorl. Transition to convex base gradual. Columella fairly short, weakly concave with no columellar folds. Inner lip rather strong, well defined, smooth. Outer lip thick, flat and sharply cut, inside smooth. Growth lines opisthocline, nearly straight. Teleoconch sculpture weak, dominated on early whorls by approximately 40 to nearly 60 low and sharp-ridged, weakly opisthocline transverse ribs. Later whorls nearly smooth but for a single spiral thread located approximately a third of the whorl height from adapical suture. Measurements. MGUH 33128 measures 2.5 mm in width, and consists of the final two whorls. Remarks. The available specimen is a juvenile, meaning that the aperture has not been fully developed with the characteristic oblique siphonal canal. The generic assignment is consequently only tentative with the possibility that the taxon may turn out to belong somewhere else within the Cerithiidae. Striovertagus? sp. is closely related to the middle Danian Trypanaxis? faxensis Ravn, 1933 of Denmark, from which it differs by the nearly absent spiral sculpture and a well developed transverse ribbing. Neither taxon belongs to Trypanaxis Cossmann, 1889. It resembles the Danian Striovertagus punctifibrum (Briart & Cornet, 1873) from Belgium, but the whorls are more convex and the transverse sculpture more pronounced.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFB42B9CC64DFAEDFB26.taxon	description	Figs 23 L – O	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A94FFFB42B9CC64DFAEDFB26.taxon	materials_examined	Material. The two external teleoconch moulds MGUH 33129 and MGUH 33130. Occurrence. Lithified top of the Maastrichtian Højerup Member north of Holtug Quarry, Stevns Klint and from the upper metre of the lower Danian ‘ dead layer’ at Vokslev Quarry, Northern Jutland. Description. Protoconch unknown. Shell high turriform with weakly convex whorls separated by shallow but well marked suture. Whorls twice as wide as high. Periphery located slightly above fairly sharp transition to moderately flat base. Columella fairly short, terminating in rather deep and oblique, narrow siphonal canal. Columellar folds lacking. Growth lines opisthocline, nearly straight. Teleoconch sculpture on early whorls dominated by around 20 to later 40 low and sharp-ridged, weakly opisthocline transverse ribs. Transverse ribs crossed by seven to nine narrow and weak spiral ribs, abapical most demarcating whorl base. Adapical rib beaded and stronger than succeeding ones, demarcated abapically by relatively wide spiral furrow crossing transverse ribs. Later whorls characterized by fine spiral ribs and furrows fading out centrally on whorl. Base smooth but for weak spiral ribs abaxially. Measurements. MGUH 33130 measures 3.65 mm in width and 10.1 mm in height, consisting of at least eight teleoconch whorls. The original height was most probably around 11.0 mm. Remarks. The taxon has an uncharacteristically deep and narrow siphonal canal, but otherwise agrees well with the diagnostic features of the Miocene to Recent genus Cerithidea. It is therefore tentatively assigned here.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A951FFB42B9CC412FA2BFA59.taxon	materials_examined	Type species. Cerithium subspinosum Deshayes in Leymiere 1842 (type by original designation). Early Cretaceous, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A951FFB72B9CC740FE30FAED.taxon	description	Figs 23 P – S 1885 Scalaria crassilabris Koenen: 66, pl. 3, fig. 2. 1939 Eucycloscala? crassilabris (v. Koenen) — Ravn: 58, pl. 2, fig. 1 a – c. 1971 Scalina crassilabris (v. Koenen) — Rosenkrantz: 100.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A951FFB72B9CC740FE30FAED.taxon	materials_examined	Diagnosis. Protoconch with only three whorls, the last half whorl carrying spiral ribs. Teleoconch with around 12 transverse ribs on first whorl, increasing rapidly on later whorls. Aperture with very weakly developed siphonal canal. Type material. The type material should include more than 30 specimens out of which one formed the basis of the drawings presented by von Koenen (1885: pl. 3: 2 a – d) and got a museum number. This specimen, MGUH 865, is here selected as the lectotype. MGUH 865 is a nearly complete specimen only lacking the first whorl of the protoconch and the aperture (Fig. 23 R). Examined material. A single external and internal mould, MGUH 33131, from the Cerithium Limestone Member at Holtug, Stevns; a specimen ØSM. 10042 - 61 - a from the Korsnaeb Member at Korsnaeb, Stevns Klint; an informally catalogued mould SH. 320 from the basal Cerithium Limestone at Højerup Church, Stevns Klint; the lectotype MGUH 865 from the Lellinge Greensand at Vestre Gasvaerk, and the complete specimen, MGUH 3751, figured by Ravn (1939: pl. 2: 1 a – c) from the Lellinge Greensand at Sundkrogen, Copenhagen. Type stratum and type locality. Black clay unit from the Lower Selandian Lellinge Greensand at Vestre Gasvaerk in Copenhagen. Occurrence. Lower Danian Cerithium Limestone Member of the Rødvig Formation and up into the Lower Selandian Lellinge Greensand of eastern Denmark. Description. Protoconch conical, consisting of three convex whorls, first 2 ½ whorls smooth with keel just above abapical suture. Last quarter whorl with three spiral ribs, one on the keel and two adapically of this. Transition to teleoconch fairly sharp. Teleoconch whorls convex, evenly rounded, weakly bicarinate with nearly continuous transition to base. Aperture subcircular with strong varix at outer lip and well developed callus and straight and sharp columellar fold partly hiding narrow umbilicus. No teeth or columellar folds present. Siphonal canal only weakly developed. Teleoconch sculpture initially consisting of three spiral ribs and approximately 12 coarse, slightly opisthocyrt transverse ribs, increasing in number to around 20 on fourth whorl and about 35 on fifth whorl. Sinus of transverse ribs located between two keels. Number of spiral ribs increasing gradually with ontogenetic growth. Low and coarse tubercles are formed at crossing points between transverse ribs and three to four primary spiral ribs, one or two of them located at adapical suture, while last two are found at keels. Base nearly smooth or covered by weak spiral ribs. Measurements. Specimen MGUH 33131 from the Cerithium Limestone Member measures at least 9.7 mm in height and 5.0 mm in width, consisting of six teleoconch whorls. Remarks. The early Danian specimen may differ from the type in having slightly less pronounced sculpture, by a higher number of transverse ribs on later whorls and in having a nearly smooth base. This kind of variation is however common in a normal population of living cerithiid gastropods (Kase 1984) and the specimen is here regarded as conspecific with the Selandian Cirsocerithium crassilabris. C. crassilabris resembles somewhat the Early Cretaceous type species C. subspinosum Deshayes from France and the Mid Cretaceous C. collignoni Kiel, 2006 from Madagascar, but differs by the higher number of transverse ribs, by the weaker siphonal canal, by the lower number of protoconch whorls and by the absence of denticles in the aperture. The aperture and protoconch compares well with that of C. kulickii (Schröder, 1995) from the Early Cretaceous of Poland (see Kaim 2004: fig. 25 c), but the shell is easily distinguished by the protoconch only having three whorls, the last of which carries three and not two spiral ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A952FFB72B9CC4E5FE9CF9E9.taxon	materials_examined	Genus indet.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB62B9CC45EFB4BFA8C.taxon	materials_examined	Type species. Turritella mesal Deshayes, 1843 (type by original designation). Recent, southwestern Europe.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB62B9CC0D9FB87FE12.taxon	materials_examined	Type species. Turbo terebra Linnaeus, 1758 (type by monotypy). Recent, Western Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB62B9CC30EFF56FB9A.taxon	description	Figs 23 V – W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB62B9CC30EFF56FB9A.taxon	materials_examined	Material. MGUH 33134 and a fragmentary teleoconch mould lacking number, which is stored in old collections at the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Tor Formation at Stevns Klint. Description. Whorls flattened, rectangular in cross-section with sharp peripheral transition to relatively flat base. Periphery of preceding whorl reaching slightly farther out than top of succeeding one, divided by fairly deep suture. Lower half of whorl with wide and shallow spiral furrow, separated from abapical suture by broad spiral band taking up just over a quarter of the whorl height. Aperture widened with the parietal lip slightly raised. Whorl surface covered by weak spiral lirae and indistinct, sharply sigmoid growth lines. Measurements. The most complete specimen, MGUH 33134, measures 17.2 mm in diameter and has a last whorl height of 17.8 mm. Remarks. The material is too fragmentary for a confident identification, but the taxon shows a close resemblance to the contemporaneous Turritella (Haustator) plana Binkhorst, 1861 from Belgium to which it is tentatively assigned.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB12B9CC48CFD32FD2D.taxon	description	Figs 23 X – Y 2014 Mesalia sp. Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A953FFB12B9CC48CFD32FD2D.taxon	materials_examined	Diagnosis. Protoconch broad and low with four whorls. Teleoconch spire relatively low with weakly beaded spiral ribs and a smooth, flattened base. Derivation of name. danica (Latin) = Danish. Type material. Holotype MGUH 33135 is a well preserved external mould of the protoconch and teleoconch spire. Paratype MGUH 33136 is an external mould of the protoconch from the same stratigraphic level at Højerup, Stevns Klint. Additional material. Two fragmentary external moulds from the uncatalogued old collections at the Natural History Museum of Denmark, and a complete external mould (ØSM. 10042 - 335 - a and b) from the collections at Østsjaellands Museum. Type stratum and type locality. Lithified top of the Højerup Member of the upper Maastrichtian Tor Formation at Holtug Quarry, Stevns Klint. Occurrence. Lithified top of the Tor Formation at Højerup Church, Mandehoved and Holtug, Stevns Klint. Description. Protoconch low conical, consisting of four smooth whorls with a very tiny initial chamber. Height and width equal, measuring between 0.5 and 0.6 mm. Teleoconch whorls sharply convex with flattened upper shoulder and keel-like periphery. Shoulder carrying three to four secondary spiral ribs, followed by three primary spiral ribs intercalated by two secondary ones around the keel. Spiral ribs crossed by strong opisthocyrt growth lines forming weak, diffuse tubercles at connecting points. Broad growth line sinus located just above keel. Base flattened, smooth. Aperture rounded subrectangular. Measurements. Shell height of MGUH 33135 larger than 4.2 mm and width larger than 2.3 mm. Remarks. Mesalia danica n. sp. differs from other species of this genus by the broad and low protoconch with its four whorls; the relatively compact teleoconch shell and the weakly beaded spiral ribs. It shows some resemblance to the slightly younger Mesalia instabilis (Briart & Cornet, 1873) from the Danian of Belgium, but is separated by the more compact shell with a higher number of spiral ribs and a smooth, flattened base. Mesalia danica n. sp. differs from the Danian M. virginiae Stilwell et al., 2004 from Antarctica and the Middle Palaeocene Mesalia sp. 1 and Mesalia sp. 2 sensu Kollmann & Peel (1983) from West Greenland by the convex whorls with weak tuberculation. It is distinguished from the Maastrichtian M. (Woodsalia) jovisammonis (Quaas, 1902) from Libya by the more compact shell with its tuberculated spiral ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A954FFB12B9CC26DFA26F995.taxon	description	Fig. 23 Z	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A954FFB12B9CC26DFA26F995.taxon	materials_examined	Material. MGUH 33137 and the two informally catalogued and fragmentary moulds SR. 918. A and SR. 1041 stored at the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Shell high turriform, slender. Protoconch only poorly preserved, conical, consisting of up to two or three whorls. Transition to teleoconch not discerned. Teleoconch whorls strongly convex, more than twice as wide as high with impressed suture. Transition to narrow and flattened, slightly thickened base marked by spiral cord or rib. Narrow umbilical opening present. Aperture seemingly rounded ovoid. Teleoconch sculpture on early whorls consisting of three strong spiral ribs, adapical two strongest, situated around whorl periphery. Last whorl carrying an additional spiral rib between adapical three and the spiral rib delimiting the base; distance between ribs decreasing abapically on whorl. Spiral ribs increasing in number on later whorls. Measurements. Specimen MGUH 33137 is 1.0 mm high and 0.6 mm wide, consisting of the initial four or five whorls. Remarks. The taxon resembles Mesalia danica n. sp. described above from the same stratigraphic level, but differs by its smaller size, slender spire, convex and relatively higher whorls and narrower base. It is distinguished from the Danian Mesalia marthae (Briart & Cornet, 1873) from Belgium by the slightly less densely spaced spiral ribs; a narrow, but smooth base and possibly by a relatively higher whorl. It does not seem to have the siphonal canal seen on the Danish middle Danian Mesalia? sp. 1 sensu Lauridsen & Schnetler (2014: p. 50), which in any case appears more similar to the cerithiid genus Fastigiella Reeve, 1848 than to any members of the family Turritellidae.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A954FFB12B9CC65DFAD2F88E.taxon	materials_examined	Type species. Serpula anguina Linnaeus, 1758 (subsequent designation by Cossmann 1912: p. 146). Recent, Indian Ocean.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A955FFB02B9CC555FB59FB97.taxon	materials_examined	Type species. Cerithium giganteum Lamarck, 1804 (subsequent designation by Sacco 1895). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A955FFB02B9CC1D4FD13FCD5.taxon	description	Figs 23 AE, 24 A 2014 Tenagodus sp. Hansen & Surlyk: 339, fig. 6: 3, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A955FFB02B9CC1D4FD13FCD5.taxon	materials_examined	Material. MGUH 33138, MGUH 33139, and 15 external and internal moulds inofficially catalogued by the sample numbers: SH. 8. A, SR. 729, SR. 750. A – B, SR. 787. B and D, SR. 915. A – B, SR. 931. A – B, SR. 943, SR. 964, SR. 985, SR. 988, SR. 989, SR. 1029. A, SR. 1073. A – B, SR. 1075. A – B and SR. 1094. A – B. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig, Skeldervig, Højerup Church and north of Holtug Quarry, Stevns. Description. Whorls free, loosely coiled, tubular with narrow, even-sided slit on adapical side, slit width 0.9 mm. Teleoconch surface smooth or covered by relatively fine and weak spiral ribs, ribs becoming weaker on adapical whorl side. Cracks and growth ridges may be present, but unevenly distributed. Measurements. Large, largest specimen consisting of two whorls, 28 mm high and with an aperture width of 5.6 mm. Remarks. The preserved part of this species does not reveal any diagnostic features except for the generic characters, but shows a close resemblance to other species of Tenagodus with no spines such as the extant T. obtusus (Schumacher, 1817) and the Scandinavian middle Danian Tenagodus ornatus (Lundgren, 1867). In fact better material may show it to be conspecific with the latter.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A955FFB32B9CC58BFDF8FBF0.taxon	description	Figs 24 B – D 1867 Cerithium selandicum Lundgren: 17, pl. 1, fig. 2. 1902 Cerithium pseudotelescopium n. sp. — Ravn: 219 – 220, pl. 1, figs 13 – 14. 1902 Cerithium selandicum Lundgren — Ravn: 220 – 221, pl. 1, figs 15 – 16. 1933 Cerithium (Campanile?) pseudotelescopium Ravn — Ravn: 45, pl. 4, fig. 8 a – b. 1933 Metacerithium? selandicum Lundgren — Ravn: 46.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A955FFB32B9CC58BFDF8FBF0.taxon	materials_examined	Diagnosis. Columella with single strong fold. Teleoconch sculpture on early whorls consisting of two or three distinct spiral rows of tubercles. Suture bordered by two rows, while last row runs on adapical half of whorl, connected by weak transverse rib to adapical tubercle row. Later whorls with fine spiral ribs. Material from the boundary layers. MGUH 33140, MGUH 33141 and ØSM. 10042 - 129. The old collections at the Natural History Museum of Denmark includes another five uncatalogued moulds. Type stratum and type locality. The middle Danian Faxekalk at Limhamn, southern Sweden. Occurrence. From the lower Danian Cerithium Limestone Member and up to the middle Danian Faxe Limestone in Sweden and Denmark. At Stevns Klint it is found in the Cerithium Limestone Member and succeeding Korsnaeb Member. A specimen has been found by C. Heinberg in the contemporaneous ‘ dead layer’ at Vokslev, Northern Jutland. Description of Cerithium Limestone material. Protoconch not known. Teleoconch spire high, slender with low, flat whorls and sharp keel, whorl height corresponding to one third the width. Suture enhanced by spiral ribs on early growth stages, becoming thin and shallow on later growth stages. Last whorls more inflated with keel close to abapical suture. Base nearly perpendicular to whorl side, flat to slightly concave, becoming more convex on last whorl. Aperture subrectangular with short columella carrying single sharp fold centrally, bordered abapically by deep canal. Teleoconch sculpture variable, on early growth stages consisting of two primary beaded spiral ribs, the typically strongest at adapical suture, the other on keel at abapical suture. Primary spiral ribs separated by several weaker secondary ribs, the adapical one being the strongest; secondary ribs increasing in number on later whorls while becoming fainter. Tubercles, reflecting slightly opisthocyrt growth lines, generally merging into transverse ribs on adapical part of whorl, fading out abapically, often merging with tubercles on abapical primary spiral rib; number of adapical tubercles approximately 15 in a half whorl on second last whorl of MGUH 33141 and 18 in abapical row. Transverse ribs decreasing in strength and length with increasing whorl number. Base with single primary smooth spiral rib located close to keel at transition to flat part of base and axially succeeded by faint secondary spiral ribs. Sculpture on later growth stages changing to one of moderately fine, unbeaded spiral ribs only, the exact rib-configuration varying from specimen to specimen, but tending towards a fainter ribbing centrally on whorl. Base with weak spiral ribs. Measurements. Most complete specimen, MGUH 33140, 32.7 mm high and 11.4 mm wide, consisting of 12 teleoconch whorls. Remarks. The taxon is variable with regard to shell sculpture, varying between forms with only spiral ribs to others with two or three stronger spiral rows of tubercles, the adapical two connected by more or less weak transverse ribs. The transverse sculpture is typically most prominent on early teleoconch whorls, fading out on later whorls. The wide variability has led to the erection of the subjective junior synonym Campanile pseudotelescopium (Ravn, 1902). It was thought to differ from C. selandicum (Lundgren, 1867) by having only one subsutural tubercle row and by the whorls being generally slightly higher and with a more abapical location of the columellar fold. An examination of the material stored at the Geological Museum of Copenhagen together with new material figured by Damholt et al. (2010) do not support such a division. The taxon belongs to Campanile and not Metacerithium due to the well developed columellar fold and the strongly flattened base on early whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A956FFB32B9CC774FA92F81F.taxon	description	Fig. 24 G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A956FFB32B9CC774FA92F81F.taxon	materials_examined	Material. Specimen MGUH 33143 is an external whorl mould. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Holtug Quarry, Stevns. Description. Teleoconch whorls flattened, slightly sigmoid in outline with sharp, pronounced abapical keel and weakly convex base. Suture deep. Columella with sharp, nearly horizontal fold. Teleoconch sculpture fairly coarse, consisting of spiral ribs and coarse tubercles on keel and finer beading on spiral rib 1 / 3 of whorl height from adapical suture; weak, slightly prosocline transverse ribs reaching adapical from spiral beads to suture. Measurements. Whorl height 3.4 mm. Remarks. MGUH 33143 differs from the two taxa described above by the deep suture and the prosocline transverse adapical ribs. The strong differences suggest that the taxon should be regarded as a separate species.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A956FFB32B9CC528FBD6FA04.taxon	description	Figs 24 E – F	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A956FFB32B9CC528FBD6FA04.taxon	materials_examined	Material. Specimen MGUH 33142 is a mould of the last whorl. Occurrence. Lithified top of the Maastrichtian Højerup Member at Holtug Quarry, Stevns Klint. Description. The mature whorl carries ten or more moderately strong spiral ribs. Growth lines slightly opisthocyrt, nearly straight. Base carrying one moderately strong spiral rib close to keel, followed by a number of faint secondary ribs. Measurements. MGUH 33142 is a 2.1 mm high whorl. Remarks. The specimen could easily turn out to belong to Campanile selandicum (Lundgren, 1867) described above, but until better material has been uncovered it is placed under open nomenclature.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A958FFBD2B9CC3FEFBD8FCED.taxon	materials_examined	Type species. Cerithium trimonile Michelin, 1838 (type by original designation). Albian, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A958FFBC2B9CC22CFCFDFBB8.taxon	description	Figs 24 H – K 1902 Cerithium balticum Forchhammer Ravn: 221 – 222, pl. I, figs 17 – 18. 1902 Cerithium Sartorii J. Müller? — Ravn: 222, pl. I, fig. 19. 1923 Cerithium balticum Forchhammer — Jessen & Ødum: 44.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A958FFBC2B9CC22CFCFDFBB8.taxon	materials_examined	Diagnosis. Metacerithium with strongly transverse canal, broadly rounded periphery and base and three rows of relatively fine and sparsely distributed tubercles of semiequal strength. Type stratum and type locality. Among the material used by Ravn (1902) for his description of this species only the two figured (Ravn 1902: pl. I: 17 – 18) have been designated as types. They have the museum numbers MGUH 91 and MGUH 92, respectively. Both syntypes originate from the Cerithium Limestone Member at Stevns Klint. Ravn never selected a holotype for this species. Due to its better preserved spire and aperture specimen MGUH 91 is here selected as lectotype. Material. MGUH 91, MGUH 92, MGUH 93, MGUH 33144, MGUH 33145, ØSM. 10042 - 40, ØSM. 10042 - 91 - a, ØSM. 10042 - 183 - a – b and ØSM. 10042 - 374 - a. Another 133 specimens are catalogued by the unofficial sample numbers: SH. 221, SH. 228, SH. 257 (2 specimens), SH. 279, SH. 281. A and C, SH. 281. B – D, SH. 281. C – D, SH. 286, SH. 288 (2 specimens), SH. 289, SH. 290, SH. 294. A, SH. 297. A, SH. 297. A – B, SH. 334. A, SH. 342. A (3 specimens), SH. 351. A, SH. 352. A, SH. 360, SH. 372, SH. 373. A – B, SH. 380, SH. 381. B – C, SH. 385. B, SH. 388, SH. 390, SH. 392, SH. 394 (2 specimens), SH. 396, SH. 398, SH. 399, SH. 403. A – B, SH. 407. A – B, SH. 411 (2 specimens), SH. 412, SH. 413, SH. 414. B, SH. 415. A – B, SH. 417, SH. 418. A – B, SH. 419. A – B, SH. 419. B, SH. 428, SH. 436. A – B, SH. 451 (4 specimens), SH. 452 (2 specimens), SH. 458. A, SH. 460, SH. 463, SH. 470, SH. 471, SH. 473 (2 specimens), SH. 483. C, SH. 484. B, SH. 517, SS. 169, SR. 143, SR. 149, SR. 152. A – B (2 specimens), SR. 152. A, SR. 210, SR. 248, SR. 271, SR, 277. C, SR. 286, SR. 287, SR. 288, SR. 289, SR. 295, SR. 299 (2 specimens), SR. 312, SR. 315, SR. 320, SR. 358, SR. 362, SR. 373, SR. 374 (2 specimens), SR. 377, SR. 387, SR. 402, SR. 406, SR. 441, SR. 446, SR. 480, SR. 501, SR. 508, SR. 510, SR. 516, SR. 527, SR. 557. B, SR. 562, SR. 563, SR. 566, SR. 568, SR. 581. A – B, SR. 581. B, SR. 583. A – B, SR. 583. B, SR. 610, SR. 617. B, SR. 623, SR. 632, SR. 638, SR. 639 (2 specimens), SR. 641, SR. 642. A – B (2 specimens), SR. 646, SR. 656, SR. 683, SR. 684, SR. 686, SR. 698. A – B, SO. 109, SO. 186 (2 specimens), SO. 187 and SO. 188. A further 87 external and internal moulds from the Cerithium Limestone Member at Stevns Klint and four external moulds from Bundgaard near Rebild just south of the Limfjord, Jutland are found in the uncatalogued old collections at the Natural History Museum of Denmark. Occurrence. Metacerithium balticum ranges from the lower part of the lower Danian Parvularugoglobigerina eugubina Partial Foraminiferal Range Zone (Pα) and up into the upper part of the Parasubbotina pseudobulloides Partial Foraminiferal Range Subzone (P 1 a) at Stevns Klint on Sealand and at least in some part of the informal ‘ dead layer’ at Bundgaard, Dania and Vokslev quarries in the Limfjord area of northern Jutland. Stevns Klint: Rødvig (20 – 72 cm above the Cretaceous), Skeldervig, Harvig, Klintekongens Hule, Højerup (10 – 37 cm above the Cretaceous), North of Kirkevig, Kirkebraade, South of the Lighthouse, Barmhjertighedshuse Mandehoved, Holtug (44 – 64 cm above the Cretaceous) and North of Kulsti. Description. Protoconch nearly 0.4 mm high and 0.4 mm wide, low conical with 2 ½ smooth and strongly convex whorls. Embryonic part moderately small. Transition to teleoconch marked by sinusigera riblet followed by strong transverse ribs and a slight median keel. Teleoconch turriculate to slightly coeloconoid with up to at least 12 flattened whorls; initial whorls may appear convex due to transverse ribbing; whorls wide and low, the H / W ratio corresponding to between 0.35 and 0.40 on the later whorls. Suture deep and narrow. Last whorl featuring slight angle at transition to base; base flattened. Aperture rounded rectangular; columella slightly concave with no folds, terminated by nearly transverse canal directed parallel to lower margin of outer lip. Outer lip with three denticles somewhat removed from aperture, a weak one located on adapical part and two stronger ones situated on the flattened part close to the siphonal canal. A fourth strong denticle is located on the parietal lip closer to the aperture. Sculpture on first four to six whorls composed of 10 or more varix-like opisthocyrt ribs, which then begin to split up into two distinct spiral rows of tubercles, the abapical row located at periphery just above abapical suture, while adapical row runs approximately midway between the two sutures. The number of transverse ribs and tubercles increases slowly by about one per whorl. After the first four or five whorls a third adapical row of finer and slightly more densely spaced tubercles appears on a band or spiral rib at the adapical suture. The spiral rib increases in strength abapically, while the tubercle rows become less pronounced. Whorl surface covered by very fine spiral striae and forwardly concave growth lines with sinus between tubercle rows 2 and 3. Measurements. Lectotype MGUH 91 is 11.4 mm wide and more than 25 mm high with 10 preserved teleoconch whorls, thereby representing one of the largest specimens known of this species. Remarks. The name Cerithium balticum was used by Forchhammer (1861: p. 781) in his description of the Danian deposits of Denmark, but the name was first formalized forty years later when Ravn (1902) described the Danish gastropod fauna from Cretaceous and Danian deposits.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A959FFBC2B9CC5E0FC91F9E5.taxon	materials_examined	Type species. Dzikella trammeri Kaim, 2004 (type by original designation). Early Cretaceous, Poland. Remarks. Kiel (2006: p. 459) noted a close resemblance between this genus and Metacerithium treated above and suggested that they are if not congeneric then at least closely related. The taxon described below shows similarities to both genera, but is tentatively assigned Dzikella based on its close resemblance to D. chuzikensis Kaim, Beisel & Kurushin, 2004. The fragmentary nature of the new material, though, does not allow for a better understanding of the phylogenetic relationship between the two genera, and the status of Dzikella as a distinct genus or junior subjective synonym of Metacerithium can thus not be solved here.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A959FFBF2B9CC71AFE5CFEA1.taxon	description	Fig. 24 L	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A959FFBF2B9CC71AFE5CFEA1.taxon	materials_examined	Material. Specimen MGUH 33146 is a fragmentary teleoconch mould. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation north of Kulsti Rende. Description. Teleoconch broadly coeloconoid with weakly convex, nearly flattened whorls with a strongly angular abapical keel and separated by somewhat grooved sutures. Whorl height approximating 2 / 5 the width. Base weakly convex. Columella smooth with no columellar fold. Whorls subrectangular in cross section. Early teleoconch sculpture consisting of 12 to 18 coarse transverse ribs and fine spiral lirae. Transverse ribs reaching from adapical two spiral lirae and abapically to keel. Later teleoconch whorls smooth except for two weak adapical spiral lirae and two similarly weak abapical spiral lirae located slightly adapically of keel. Growth lines strongly prosocline, slightly prosocyrt. Measurements. Specimen MGUH 33146 is 7.2 mm high and 6.0 mm wide, consisting of 6 ½ teleoconch whorls. Remarks. The Danish taxon resembles the late Maastrichtian D. chukinensis Kaim, Beisel & Kurushin, 2004 from Western Sibiria, but differs by the concave sided spire, the more flattened whorls with the abapically located keel and by a less convex base.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95AFFBF2B9CC3C6FEDEF8C5.taxon	description	Figs 24 M – Q	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95AFFBF2B9CC3C6FEDEF8C5.taxon	materials_examined	Diagnosis. The shell is high-spired with distinctly concave base, a deeply grooved suture, a whorl periphery located well above abapical suture, a very narrow umbilical opening and with pitted spiral grooves. Derivation of name. In honour of J. P. J. Ravn, who greatly contributed to our knowledge of the gastropod faunas from the Cretaceous and especially Palaeocene deposits of Denmark. Type material. The holotype MGUH 33147 is a nearly complete external mould. Paratype MGUH 33150 is an external mould of the protoconch from Stevns Klint. Paratype MGUH 33149 is a mould with replaced shell from Holtug. Paratype MGUH 33148 is an internal mould from Holtug. All paratypes come from the Cerithium Limestone Member. Additional material. Three specimens are found in the informally catalogued samples SH. 278, SH. 325. A and C, and SR. 525. Another seven external and internal moulds have been studied from the uncatalogued old collections at the Natural History Museum of Denmark. Type stratum and type locality. The Cerithium Limestone at Stevns Klint. Occurrence. The lower Danian Cerithium Limestone Member at Stevns Klint. Description. Protoconch conical, with seemingly around 2 ½ moderately convex and smooth whorls. Height and width approximately 0.6 mm. Teleoconch coeloconoid with a moderately high spire and a globose last whorl. Spire whorls moderately convex with periphery located just below 1 / 3 the whorl height from the abapical suture. Suture initially shallow, on later whorls becoming deeply grooved with nearly axially oriented sides and bordered by a sharp abaxial shoulder. Base evenly rounded. Umbilical opening narrow. Aperture semicircular with width corresponding to about 70 % of height; aperture height approximating 60 % of shell height. Callus on inner lip thin, not reaching outside aperture opening. Growth lines fine, prosocline, crossed by weak, but more or less unevenly distributed pitted spiral grooves. Measurements. Holotype MGUH 33147 is 14.0 mm high and 9.9 mm wide, consisting of at least seven whorls. Largest specimen paratype MGUH 33148 is 13.2 mm wide and in excess of 15.4 mm in height. Remarks. Amaurellina ravni n. sp. differs from the Danian Amaurellina sp. and “ aff. Amaurellina ” sp. sensu Kollmann & Peel (1983) from Greenland by the distinctly higher spire and more deeply incised suture. It is distinguished from the Danian Amaurellina julei (Briart & Cornet, 1873) by the relatively larger last whorl, a concavesided spire and a narrower umbilical opening. It has some resemblance to Amaurellina guerrera Perrilliat, Vega & Corona, 2000 from the Maastrichtian of Mexico, but differs by the more convex spire whorls and the presence of faint spiral grooves.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95AFFBF2B9CC0D9FC76FDDB.taxon	materials_examined	Genus Amaurellina Bayle in Fischer (1884) Type species. Natica spirata Lamarck, 1804 (type by monotypy). Eocene, Europe.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFB92B9CC437FDE7FF18.taxon	description	Figs 24 S – W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFB92B9CC437FDE7FF18.taxon	materials_examined	Material. Three nearly complete internal moulds (MGUH 33152, MGUH 33153 and ØSM. 10042 - 311) with parts of external mould preserved. Occurrence. Lithified top of the Maastrichtian Højerup Member north of Kulsti Rende and the middle part of the lower Danian Cerithium Limestone Member near Knøsen, Stevns Klint. Specimen MGUH 33152 is indicated to have come from the lithified top of the upper Maastrichtian Højerup Member, but the matrix would rather suggest the lower Danian Korsnaeb Member, probably from one of the Danian Thalassinoides burrows into the Cretaceous strata. Description. Shell thin, cap-shaped with protoconch located close to posterior margin. Highest point of shell situated 2 / 3 of shell length from anterior margin or farther back. Interior with thin process or plate reaching from just behind apex and halfway to highest point of shell or, in specimen MGUH 33152, to around midline of shell. A possible muscle scar describes a wide semicircle halfway between apex and anterior margin. Outer surface characterized by irregular growth increments and very fine and dense radial striae and coarser, short radial ribs unevenly distributed across the surface. Measurements. Specimen MGUH 33152 is 8.0 mm long, 8.5 mm wide and nearly 4.0 mm high. Remarks. The Danish specimen strongly resembles the British Albian type species of this genus, but may differ by a more horizontally directed inner process and a less steep anterior margin. These characters may on the other hand just as well reflect environmental influences on the growth. The age difference may be the strongest support for regarding this as a separate species.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFBE2B9CC581FB03FACA.taxon	materials_examined	Type species. Galericulus altus Seeley, 1861 (type by monotypy). Early Cretaceous, United Kingdom.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFBE2B9CC2B9FD17FBB5.taxon	materials_examined	Superfamily Calyptraeoidea Lamarck, 1809	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFBE2B9CC1D4FAFCFEDE.taxon	materials_examined	Genus Trochocerithium Cossmann & Sacco in Sacco (1897) Type species. Trochus turritus Bonelli in Bellardi & Michelotti (1841) (type by original designation). Miocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFBE2B9CC0DDFE94FC41.taxon	description	Fig. 24 R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95BFFBE2B9CC0DDFE94FC41.taxon	materials_examined	Material. Specimen MGUH 33151 is an external mould of the spire. Occurrence. Lower Danian Cerithium Limestone Member just south of Højerup Church, Stevns. Description. Protoconch unknown. Teleoconch broad-spired, conical, with flat sides; whorls low, height corresponding to just over 35 % of width. Whorl sculpture dominated by up to around ten weak, beaded spiral ribs and a slightly more pronounced abapical beaded spiral rib located just adjacent to abapical suture. Nearly effaced prosocline and slightly opisthocyrt striae cross the spiral ribs, resulting in a dense beading of spiral ribs. Abapical spiral rib carrying nearly 90 tubercles on sixth preserved whorl. Measurements. Specimen MGUH 33151 measures 13.3 mm in height and 9.0 mm in width and has 8 whorls. Remarks. The Danish specimen is easily distinguished from the Danian Trochocerithium chapuisi (Briart & Cornet, 1873) from Belgium in the extensive spiral and transverse ribbing of the whorls and the distinct tuberculation of the primary abapical spiral rib. It may be conspecific with the Cretaceous Cerithium heberti Geinitz, 1875 from Germany	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95CFFB92B9CC38BFB7BF987.taxon	description	Figs 24 X, 25 A – G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95CFFB92B9CC38BFB7BF987.taxon	materials_examined	Material. MGUH 33154, MGUH 33155, MGUH 33156, ØSM. 10042 - 88 - e and ØSM. 10042 - 104 - b – c. An additional 47 specimens are found in the informally catalogued samples SH. 104. A – B, SH. 140. A, SH. 142 (2 specimens), SH. 157. A – B, SH. 183. B, SH. 198, SH. 199, SH. 200, SH. 201, SH. 202, SH. 226, SH. 228, SH. 232, SH. 249, SH. 250, SH. 274. B (3 specimens), SH. 277, SH. 281. A and E, SH. 285. A, SH. 311, SH. 312, SH. 367, SH. 406. A – B, SH. 448, SH. 466, SH. 467, SH. 518. A, SR. 297, SR. 303. A – B, SR. 312, SR. 365. B, SR. 380, SR. 381, SR. 402, SR. 572. A – B, SR. 573, SR. 589. A – C, SR. 623, SR. 657. A, SR. 668. B, SR. 688, SO. 129, SO. 135. C and SO. 192. Three specimens collected by A. Rosenkrantz at Skeldervig (13.07.1944 and August 1944) and Holtug Vig (August 1924) are located in the uncatalogued old collections at the Natural History Museum of Denmark. Occurrence. The lower Danian Cerithium Limestone at Stevns Klint and contemporaneous ‘ dead layer’ at Vokslev in Northern Jutland. Description. Protoconch planispiral or very weakly heterostrophic at apex with close to 2 ½ flattened whorls; width approximately 0.85 mm. Embryonic shell 0.45 mm in width, corresponding to the initial ¾ of a whorl. Teleoconch thick-shelled, globose with short spire, rounded whorls and impressed suture. Umbilicus moderately wide and deep with no separating ridge from the shell base. Aperture semicircular with straight, thickened inner lip ending in sharply angular abapical corner; height corresponding to around ¾ the shell height. Central columellar callus extension more or less completely absent. Parietal callus strong and wide, rectangular; the height corresponding to nearly half the aperture height. Growth lines simple, straight and prosocline, turned slightly adaxially at suture. Some specimens have a subsutural furrow on at least the final protoconch whorl and initial teleoconch whorl. Measurements. Largest specimen, MGUH 33154, 7.8 mm high and 7.9 mm wide, consisting of four whorls. Remarks. The taxon varies somewhat in the convexity of the spire whorls and the relative spire height, sometimes making it resemble small specimens of Polinices sp. A described below. It is, though, recognizable on strength of the generally deep suture; the lower, more stepped spire and by the rectangular parietal callus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95CFFB92B9CC368FA88FD62.taxon	materials_examined	Type species. Nerita vitellus Linnaeus, 1758 (subsequent designation by Anton 1838: p. 31). Recent, Indo-Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95CFFB82B9CC7E4FBCAFE30.taxon	description	Figs 25 H – I 2014 Natica sp. Hansen & Surlyk: 339, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95CFFB82B9CC7E4FBCAFE30.taxon	materials_examined	Material. MGUH 33157 and ØSM. 10042 - 329 - b – c. An additional four fragmentary moulds with the unofficial sample numbers SH. 17, SR. 749. B, SR. 1090. B – C and SR. 1118 are deposited at the Natural History Museum of Denmark. Occurrence. Top upper Maastrichtian Højerup Member, Tor Formation at Rødvig, Højerup and Holtug, Stevns Klint. Description. Protoconch low, consisting of two smooth and slowly widening whorls. Transition to teleoconch marked by distinct growth stop. Teleoconch seemingly sub-globose with fast expanding and in profile evenly rounded whorls lacking adapical ramp. Whorls abutting. Measurements. Most complete specimen, MGUH 33157, is at least 2.3 mm wide, consisting of slightly over three whorls. Remarks. The taxon resembles Natica sp. A described above, but differs by a more rounded aperture and the absence of a shoulder ramp. As the material is very limited and fragmentary, completely lacking a preserved aperture, it is not possible to completely verify whether the two taxa are conspecific or not.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95DFFB82B9CC24AFB78FA4A.taxon	description	Figs 25 J – N	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95DFFB82B9CC24AFB78FA4A.taxon	materials_examined	Material. MGUH 33158 and MGUH 33159. 39 specimens are found in the unofficially catalogued samples: SH. 103. A, SH. 104. B, SH. 106. A – B, SH. 140. B, SH. 142, SH. 158, SH. 174, SH. 188. A – C, SH. 204, SH. 205, SH. 218, SH. 285. A – B, SH. 288, SH. 290 (2 specimens), SH. 303. A, SH. 324, SH. 410. A – B, SH. 470, SR. 364, SR. 389. A – B, SR. 404, SR. 477, SR. 493, SR. 501, SR. 523, SR. 574, SR. 575, SR. 576. A – B, SR. 588. A – B, SR. 609, SR. 616, SR. 641, SR. 643. A – B, SR. 644, SR. 650, SR. 651, SR. 652 and SR. 668. A – B. A final 14 fragmentary external and internal moulds without numbers are located in the uncatalogued old collections at the Natural History Museum of Denmark. Occurrence. Lower Danian Cerithium Limestone Member and basal Korsnaeb Member at Stevns Klint. It furthermore occurs in the lower Danian ‘ dead layer’ at Vokslev, Northern Jutland. Description. Protoconch low-spired, but poorly preserved on available material. Teleoconch ovoid turbiniform and moderately low-spired with slightly adpressed and moderately convex whorls separated by distinct suture. Aperture semicircular and distinctly oblique to axial line. Umbilical opening narrow, partly covered by moderately thick, subtriangular callus. Surface smooth except for more or less straight, prosocline growth lines. Measurements. Largest specimen 8.9 mm wide, but most specimens reaches sizes of only a few millimetres. Remarks. The shape of the shell varies somewhat from forms with a strong ear-shape and fairly flattened spire whorls to those with a more stepped outline somewhat resembling Natica sp. A described above.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95DFFB82B9CC4A0FBB8F819.taxon	description	Figs 25 O – P 2014 Polinices? sp. Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95DFFB82B9CC4A0FBB8F819.taxon	materials_examined	Material. MGUH 33160, a specimen from the unofficially catalogued sample SR. 741. C, and a last specimen without number deposited in the old collections of the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig and Holtug Quarry, Stevns Klint. Description. Protoconch low-spired, on specimen MGUH 33160 2.1 mm wide and about 1.6 mm high, consisting of around 2 ¾ whorls. Whorl height only slowly increasing. Teleoconch fairly high-spired with relatively weakly convex whorls, separated by deep suture. Whorl height corresponding to half the width, increasing rapidly abapically. Growth lines prosocline, nearly straight. Remarks. The fragmentary preservation of the specimens prevents a sound generic assignment, but the overall form strongly resembles that of Polinices. It is distinguished from Polinices? sp. A described above by the elongated spire and apparently also by a slightly higher whorl number for the protoconch.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95DFFB82B9CC312FBD7FD5F.taxon	materials_examined	Type species. Polinices albus Montfort, 1810 (type by original designation). Recent, Indo-Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFFBA2B9CC241FB25FA85.taxon	description	Figs 25 Q – R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFFBA2B9CC241FB25FA85.taxon	materials_examined	Material. Specimen MGUH 33161 is an external mould of the protoconch and two first teleoconch whorls. Occurrence. The lower Danian Cerithium Limestone Member, 20 to 25 cm above the base of the Rødvig Formation at Højerup Church, Stevns Klint. Description. Protoconch conical, just over 0.4 mm high and 0.45 mm wide, consisting of 3 ½ smooth, weakly convex whorls. Transition to teleoconch relatively sharp. Teleoconch whorls weakly convex, twice as wide as high with periphery located immediately adapically of deeply impressed linear suture. Teleoconch whorls carrying strong and closely spaced, nearly straight transverse ribs, number of ribs 20 or just below on first teleoconch whorl. Measurements. Specimen MGUH 33161 is 1.15 mm high, consisting of five whorls with a 0.6 mm wide penultimate whorl. Remarks. The material does not allow for a closer taxonomical identification, but the general protoconch outline and the teleoconch with its well defined transverse ribs on weakly convex whorls separated by deeply impressed sutures supports a placement within this genus. It differs from the Cretaceous Zebinella selandica n. sp. described below by a more conical protoconch and the slightly coarser and straighter transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFFBA2B9CC326FAABFD24.taxon	materials_examined	Type species. Rissoina inca d’Orbigny, 1840 (type by monotypy). Recent, off the west coast of South America.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFFBA2B9CC4E6FAD8F9E4.taxon	materials_examined	Type species. Helix decussata Montagu, 1803 (subsequent designation by Martens 1878). Recent, West Indies and Forida.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFF852B9CC701FC13FCBC.taxon	description	Figs 25 S – U	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A95FFF852B9CC701FC13FCBC.taxon	materials_examined	Diagnosis. Shell slender with rounded apex, moderately convex whorls and well defined transverse ribs terminating at the whorl periphery. Transverse ribs numbering around 20 on first teleoconch whorl. Suture line well impressed. Derivation of name. Refers to the Danish island Sealand from where it derives. Type material. Holotype MGUH 33162 is a nearly complete external mould. Additional material. The collections at the Natural History Museum of Denmark includes 11 moulds with the unofficial sample numbers SH. 12, SR. 730, SR. 745, SR. 746. A, SR. 748. B – C, SR. 1043. A, SR. 1047, SR. 1063, SR. 1089. A, SR. 1093 and SR. 1126. Type stratum and type locality. Lithified top of the Højerup Member of the upper Maastrichtian Tor Formation at Rødvig, Stevns Klint. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Rødvig and Højerup, Stevns. Description. Shell small to medium-sized, slender with an elongated conical spire. Protoconch obtusely conical, 0.55 mm high and 0.55 mm wide, consisting of approximately 3 ¾ smooth, convex whorls. Transition to teleoconch moderately sharp. Teleoconch with weakly convex whorls more than twice as wide as high and divided by moderately deeply impressed linear suture located immediately abapically of preceding whorl periphery. Last whorl corresponding to less than half the shell height. Base convex. Aperture slightly flaring with thickened outer lip. Abapical part of aperture and columella not known. Teleoconch sculpture consisting of closely spaced, fine and slightly opisthocline, weakly opisthocyrt transverse ribs reaching from adapical suture to whorl periphery; number of ribs just surpassing 20 on first teleoconch whorl increasing to about 40 on third whorl. Base covered by weak spiral lirae, around nine on third whorl. Measurements. Holotype MGUH 33162 0.8 mm wide and 2.1 mm high, consisting of protoconch and 3 ½ teleoconch whorls. Remarks. Zebinella selandica n. sp. differs from the two Selandian taxa figured by Kollmann & Peel (1983) as Zebinella sp. 1 and Z. sp. 2 from Greenland by the teleoconch sculpture with the transverse ribs terminating at the periphery and by the more rounded outline of the spire top. It differs from the Danian Z. nuda Briart & Cornet, 1887 and Z. tenuicancellata Briart & Cornet, 1887 from Belgium by the presence of distinct teleoconch sculpture and well impressed suture line (see Cossmann 1924; Glibert 1973). The lack of both of these characters may indicate that the two Belgian species most likely do not belong to this subgenus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A960FF852B9CC2B4FDC8FB97.taxon	materials_examined	Type species. Alvania europea Risso, 1826 accepted as Turbo cimex Linnaeus, 1758 (subsequent designation by Nevill 1885). Holocene to Recent, Mediterranean Sea.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A960FF852B9CC58BFC87F837.taxon	description	Figs 25 V, 26 A – B	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A960FF852B9CC58BFC87F837.taxon	materials_examined	Material. MGUH 33163, MGUH 33164 and an external mould in the unofficially catalogued sample SR. 1015 deposited at the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Højerup Member, Tor Formation, at Rødvig and Højerup Church, Stevns Klint. Description. Spire fairly low and wide. Protoconch low spired with smooth and relatively flattened whorls. Transition to teleoconch apparently rather gradual. Teleoconch whorls moderately convex with periphery slightly adapical of transition to base. Suture narrow and moderately shallow. Transition to strongly convex base rather gradual. Aperture rounded abapically with strongly developed and smooth varix. Teleoconch sculpture consisting of around 10 strong and sharp-ridged, slightly opisthocyrt and opisthocline transverse ribs per half whorl; ribs fading out abapically at transition to base. Transverse ribs weak on first teleoconch whorl, crossed by weak spiral threads. Adapical suture bordered by beaded collar formed by stronger spiral rib delineated abapically by spiral furrow. Transition to weakly spirally ribbed base marked by another stronger spiral rib. Base covered by up to six or seven very weak spiral ribs, crossed adapically by nearly effaced continuations of transverse ribs. Measurements. Most complete specimen, MGUH 33163, measures 1.6 mm in height and 1.3 mm in width, consisting of three whorls but lacking top of protoconch. Remarks. Alvania sp. A resembles the Late Cretaceous Alvania costata (Wade, 1926) and Alvania tallahatchiensis (Sohl, 1960) from the U. S. A., but is readily recognized by the spiral rib bordering the base, the restricted spiral sculpture and the beaded spiral rib adapically on the whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC1D4FC1BFCF4.taxon	description	Fig. 26 C	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC1D4FC1BFCF4.taxon	materials_examined	Material. MGUH 33165 and an external mould in the uncatalogued collections of the Natural History Museum of Denmark. Occurrence. The lower Danian ‘ dead layer’ in Vokslev Quarry, Northern Jutland. Description. Protoconch obtusely conical with 2 ¼ to 2 ½ convex whorls. Whorls seemingly smooth, but could have been weakly postulate. Protoconch height between 0.3 and 0.35 mm, while the width reaches 0.55 mm. Shell high trochiform with weakly convex whorls separated by fairly distinct suture. Transition to base at periphery moderately sharp. Aperture not preserved. Teleoconch sculpture dominated by about 22 to 25 opisthocyrt transverse ribs per whorl. Ribs reaching down to periphery before fading out. Transverse ribs crossed by generally weaker spiral ribs; adapical one much stronger than succeeding ones, carrying small beads representing adapical part of transverse ribs. Spiral ribs increasing in strength at and just abapically of periphery with up to five moderately distinct spirals on adapical part of base. Measurements. MGUH 33165 is 0.8 mm wide, consisting of the last protoconch whorl and the succeeding teleoconch whorl. Remarks. Alvania sp. B resembles the late Maastrichtian A. sp. A described above, but differs by the stronger spiral sculpture and the more opisthocline nature of the transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC6D2FB5CF818.taxon	materials_examined	Type species. Rissoa proxima Forbes & Hanley, 1850 (type by monotypy). Recent, Spanish coast of Catalane.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC2E6FA2FFBE4.taxon	materials_examined	Type species. Rissoina semiglabrata A. Adams, 1854 (subsequent designation by Rehder 1980: p. 27). Recent, Indo-Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC501FC6DF929.taxon	description	Fig. 26 D	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A961FF842B9CC501FC6DF929.taxon	materials_examined	Material. Specimen MGUH 33166 is an external mould. Occurrence. The specimen was found in the Cerithium Limestone Member, 52 – 62 cm above the K-Pg boundary at Rødvig, Stevns Klint. Description. Spire blunt with smooth and flatly dome-shaped protoconch consisting of 1 ½ convex whorls; transition to teleoconch rather gradual. Teleoconch with fairly high whorls, the first being rather weakly convex, the following becoming more convex with increasingly steeply inclined flanks. Suture distinct but not deep. Aperture narrow, obliquely lenticular with a weakly developed anterior apertural channel; aperture height approximating 40 % of shell height, the width corresponding to just over 60 % of height. Measurements. Minute gastropod, MGUH 33166 is 2.0 mm high and 0.9 mm wide and consists of approximately 5 ½ whorls, protoconch included. Remarks. Zebina sp. is distinguished from most species belonging to this genus by the unusually well developed anterior apertural channel and strongly rounded base. It has a passing resemblance to the Early Cretaceous Zebina dziki Kaim, 2004 from Poland, but differs by the thinner shell, the rounded base and the narrow aperture. It differs from the mid Danian Zebina sp. 1 and Zebina sp. 2 sensu Lauridsen & Schnetler (2014) from Denmark by the relatively broader apex and narrow aperture with developed channel.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A963FF862B9CC3FEFE4CF902.taxon	description	Figs 26 E – H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A963FF862B9CC3FEFE4CF902.taxon	materials_examined	Diagnosis. Shell slender with slowly expanding and relatively flat whorls; apex blunt with low, obtusely conical protoconch and first teleoconch whorl; aperture distinctly lenticular. Teleoconch with very weak spiral striation. Derivation of name. Refers to the type locality Rødvig at the southern end of Stevns Klint, Denmark. Type material. Holotype MGUH 33167 is an external and internal mould from 10 to 20 cm below the top of the Cerithium Limestone Member at Rødvig. Paratype MGUH 33168 is an external and internal mould from the same locality and stratigraphic horizon as the holotype. Additional material. ØSM. 10042 - 108 and 16 external and internal moulds in the inofficially catalogued samples SH. 140. A – B, SH. 274. B, SH. 416, SH. 456. A – B, SH. 479, SS. 211. A – B, SR. 152. A, SR. 275, SR. 395, SR. 518, SR. 591, SR. 620. A, SR. 628, SR. 630 and SR. 696. B – C are known from the Cerithium Limestone Member of Stevns Klint. A specimen in sample DN. 28. A – B comes from the contemporaneous ‘ dead layer’ in Dania Quarry, Northern Jutland. Additionally a single specimen, ØSM. 10042 - 70, from the hardground about two metres above the base of the Korsnaeb Member at Stevns Klint was examined. Type stratum and type locality. Uppermost part of the lower Danian Cerithium Limestone Member at Rødvig, Stevns Klint. Occurrence. Ranges from 30 cm above the Cretaceous to the top of the lower Danian Cerithium Limestone Member at Rødvig and up to the hardground in the lower part of the succeeding Korsnaeb Member at Stevns Klint. It is furthermore found in the lower Danian ‘ dead layer’ at Dania quarry, Northern Jutland. Description. Spire blunt with smooth and obtusely conical protoconch consisting of 1 ½ convex whorls; transition to teleoconch rather gradual. Teleoconch with very finely spirally striated and fairly high whorls, the first rather weakly convex, the following becoming more convex with increasingly steeply inclined flanks. Growth lines weak, slightly opisthocline. Suture distinct but not deep. Measurements. Minute gastropod, the largest specimen from sample SR. 628 being just over 2 mm high and 0.95 mm wide and consisting of approximately 6 ½ whorls, protoconch included. Remarks. Ceratia rodvigensis n. sp. is distinguished from most species belonging to this genus by the low, obtusely conical apex, the relatively flat whorls and the fairly slow whorl expansion. This species may superficially resemble the Palaeocene Ceratia sp. 2 and C. sp. 3 sensu Kollmann & Peel (1983) from Greenland but for the blunt apex and the less deep suture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A963FF812B9CC67EFDB1FEA0.taxon	description	Figs 26 I – J	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A963FF812B9CC67EFDB1FEA0.taxon	materials_examined	Material. MGUH 33169 and MGUH 33170, both fragmentary moulds. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig, Stevns Klint. Description. Minute conical shell with smooth, weakly convex whorls and narrow suture. Transition to base marked by increased convexity. Aperture fairly small, ovoid, seemingly without columellar fold. Measurements. MGUH 33169 is 0.85 mm wide and 1.3 mm high, consisting of three whorls. Apex hidden. Remarks. The taxon resembles Ceratia rodvigensis n. sp. described above, but differs by the less convex whorls and a more ovoid aperture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF812B9CC39EFC5CFA96.taxon	description	Fig. 26 K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF812B9CC39EFC5CFA96.taxon	materials_examined	Material. Specimen MGUH 33171 is an external mould of parts of the last two whorls. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Holtug Quarry, Stevns. Description. Whorls relatively high and moderately convex, the height corresponding to half the width. Suture very narrow. Second last whorl with two moderately strong central rows of tubercles and an abapical tuberculated spiral rib situated just above suture. The stronger rows or ribs intercalated by two narrow and weak spiral threads. Last whorl sculpture consisting of two weak adapical rows of tubercles, succeeded abapically by two strong and sharp peripheral ribs carrying tubercles. Base of whorl covered by at least eight strong to weaker spiral rows of tubercles, five located above the canal. The peripheral two ridges and the slightly less developed adapical rib or row of tubercles on the base diverge at transition to apertual wing. Measurements. Preserved diameter of last whorl 8.1 mm. Remarks. The specimen somewhat resembles the contemporaneous A. baptos Elder & Saul, 1996 from the North American Pacific Slope, but differs by the finer and slightly less developed spiral ribs and in the slightly lower location of the two strong peripheral spiral ribs on the last whorl. It is distinguished from the Late Cretaceous A. securifera (Forbes, 1846) from Cameroon as redescribed by Kiel & Bandel (2002 b) in the weaker sculpture and the presence of two nearly equally strong peripheral spiral ribs on the last whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF812B9CC368FB06FD62.taxon	materials_examined	Type species. Anchura abrupta Conrad, 1860 (type by monotypy). Late Cretaceous, Mississippi, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF812B9CC742FB06F98F.taxon	materials_examined	Type species. Cassis corallina Quenstedt, 1858 (type by original designation). Late Jurassic, Germany.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF802B9CC7FCFCEEFC0C.taxon	description	Figs 26 L – M 1861 Pyrula tuberculosa Binkhorst: 8, pl. III, fig. 5 a – b. 1861 Pyrula? plicata — Binkhorst: 68, pl. Va. 3, fig. Aa – b. 1898 Columbellaria tuberculosa (Binkhorst) — Kaunhowen: 79 – 80, pl. IX, figs 7 – 8.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A964FF802B9CC7FCFCEEFC0C.taxon	materials_examined	Diagnosis. Low-spired and bulbous with unkeeled whorls. Sculpture of nearly smooth and sharp spiral ribs. Transverse ribs completely lacking. Material. The present Danish material consists of specimen ØSM. 10042 - 202 - a – c and an uncatalogued external and internal apical mould from Holtug quarry (A. Rosenkrantz, 1 — 6 / 8 1940), which is deposited in the old collections at the Natural History Museum of Denmark. Type stratum and type locality. The holotype figured by Binkhorst (1861: pl. III: 5 a – b) comes from the upper Maastrichtian limestone at Limburg, Belgium. Occurrence. In Denmark Columbellaria tuberculosa is found in the lithified top of the Maastrichtian Højerup Member at Rødvig and Holtug Quarry, Stevns Klint. Outside Denmark it is known from the Early Senonian of West Germany, the late Senonian of the Polish Lvov region and the Maastrichtian of Limburg (Belgium), Kunrade (Holland) and Middle Vistula Valley of Poland (Abdel-Gawad 1986). Description of Danish material. Protoconch with three smooth and moderately convex, slowly expanding whorls. Initial two whorls nearly planispiral. Protoconch (of ØSM. 10042 - 202 - b) 1.55 mm in width and about 0.8 mm in height. Teleoconch low-spired with moderately convex spire whorls. Shell width corresponding to 80 % of the height. Last whorl bulbous, constituting approximately 90 % of the shell height. Base evenly rounded. Aperture narrowly elongated, height corresponding to around 70 % of shell height. Inner lip only partly preserved, wide, with at least six teeth. Teleoconch sculpture consisting of seven narrow and sharp spiral ribs of nearly equal strength on spire whorls. Last whorl with at least 16 near equal spiral ribs and more or less completely lacking tuberculation throughout. Growth lines nearly orthogonal to slightly opisthocline. Measurements. Specimen ØSM. 10042 - 202 is a largely complete shell measuring about 5.9 mm in height and 5.2 mm in width. Remarks. The rare occurrence in the Danish deposits strongly suggests that Columbariella tuberculosa (Binkhorst, 1861) did not belong in this deep water muddy environment, but rather to a more shallow water firm-ground environment like that found in Holland, Belgium and Poland.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A965FF802B9CC678FE9CF873.taxon	materials_examined	Genus indet.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A965FF802B9CC460FBF3F94C.taxon	description	Fig. 26 N	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A965FF802B9CC460FBF3F94C.taxon	materials_examined	Material. MGUH 33172 and a fragmentary internal mould in the same sample. Occurrence. The lower Danian ‘ dead layer’ in Dania Quarry, Northern Jutland. Description. Shell large, strongly rounded in outline with a low spire and spire whorls. The most complete specimen MGUH 33172 shows signs of an internal raised band just abapical of periphery. Aperture moderately narrow with no sign of teeth or varices on outer lip or parietal region. Remarks. The material is very fragmentary and does not allow for a confident generic assignment, but it shows a strong resemblance to Tylostoma, and here especially the middle Danian T. ampullariaeforme Ravn, 1902 from Denmark. It differs from this species by having a lower spire and a narrower aperture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A965FF802B9CC5C4FAFEFB06.taxon	materials_examined	Type species. Tylostoma torrubiae Sharpe, 1849 (subsequent designation by White 1880: p. 142). Cretaceous, Portugal.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A966FF832B9CC394FF52FC78.taxon	materials_examined	Genus Sassia Bellardi, 1872 Type species. Triton appenninica Sasso, 1827 (subsequent designation by Cossmann 1903: p. 93). Miocene to Pliocene, Europe.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A966FF822B9CC2A3FECAFBB8.taxon	description	Figs 26 Q – U 1902 Tritonium fenestratum n. sp. Ravn: 227 – 228, pl. II, figs 7 – 8. 1933 Tritonium (Sassia) faxense n. nom. — Ravn: 58 – 59, pl. V fig. 16 a – b, pl. VI fig. 7 a – b.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A966FF822B9CC2A3FECAFBB8.taxon	materials_examined	Diagnosis. Shell relatively short, width typically around half the height. Protoconch small with 3 ½ to 3 ¾ whorls; last whorls covered by fine and closely spaced, prosocline collabral threads cancellated by a few, generally three, spiral threads. Teleoconch aperture taking up approximately half of shell height; columella only weakly concave. Teleoconch with strong reticulate pattern of sharp transverse and spiral ribs. Material. In addition to the Faxe Quarry material stored at the Natural History Museum of Denmark, 70 moulds from the Cerithium Limestone, one from the basal Korsnaeb Member and 11 from the lithified top of the Højerup Member at Stevns Klint have been examined. These include specimens MGUH 33173, MGUH 33174, MGUH 33175, MGUH 33176, MGUH 33177, ØSM. 10042 - 184 - a – b, and the 26 informally catalogued specimens from samples SH. 389, SR. 568, SR. 759. A – B, DN. 22, SH. 108, SH. 185. A – B, SH. 281. A and C, SH. 284. A – B, SH. 295. A – B, SH. 302. A – B, SH. 387, SH. 447, SH. 458. B, SH. 516, SR. 303. A – B, SR. 393. A – B, SR. 394, SR. 482, SR. 491, SR. 529, SR. 572. B, SR. 613, SR. 668. A – B, SR. 911. A – B and SR. 924. B. The rest are uncatalogued specimens deposited in the collections of the Natural History Museum of Denmark. Type stratum and type locality. No holotype or lectotype was selected by Ravn (1902, 1933), but the type material was collected from the middle Danian Faxe Formation of Faxe Quarry. Occurrence. Known from the lithified top of the Maastrichtian Højerup Member and the lowermost Danian Cerithium Limestone and Korsnaeb members at Stevns Klint, and from the middle Danian limestones of Faxe Quarry, Denmark. In Northern Jutland it has been found in the lower Danian ‘ dead layer’ at Vokslev. Description. Shell fusiform with tall, conical spire, shell width approximating half the height. Protoconch low conical, small, width typically around 1.3 to 1.7 mm, with 3 ½ to 3 ¾ convex whorls and low apex. Suture distinct. Last two whorls covered by fine and closely spaced, prosocline collabral threads (about 60 per whorl) cancellated by typically three narrow spiral cords. Transition to teleoconch abrupt, marked by coarsening of sculpture. Teleoconch somewhat variable regarding height and width, but with low to moderately convex and evenly rounded whorls. Suture adpressed. Varices broad and strongly raised, recurring approximately every 240 ˚ on teleooconch. Aperture, including siphonal canal, taking up between 45 and 53 % of shell height. Outer lip formed by strong varix with marked ridge running along inner side adapically, changing to outer margin abapically at transition to base of last whorl. Adapical transition to moderately wide callus smooth. Transition of outer lip to siphonal canal rather smooth with no sharp angle. Callus moderately thin, wide adapically, narrowing at transition to weakly concave columella, disappearing completely at short and left wards deflected deep siphonal canal. Number and strength of teeth on inner and outer lips quite variable. Inner lip typically carrying six teeth on columella and an additional six but weaker teeth on parietal lip. No folds. Outer lip carrying three teeth on abapical part and up to an additional three on adapical half. Teleoconch sculpture of varying strength, strongly reticulate, consisting of sharp transverse and spiral ribs narrower than interspaces, and with tubercles at intersections. Each whorl carrying around 17 sharp transverse ribs, ribs slightly sigmoid with a prosocline, opisthocyrt adapical part. Primary spiral cords numbering 3 – 4 on spire whorls and about 10 on last whorl. Up to several fine spiral threads may be added between primary spirals on later whorls, most numereous on base. Measurements. Specimens from the Cerithium Limestone Member may reach a height of at least 15.1 mm and a width of 10.7 mm, but most are less than 10 mm high. Remarks. This species is somewhat variable with regard to protoconch size and the strength of the teleoconch sculpture, something which is seen for both the mid Danian and the latest Maastrichtian to earliest Danian material. The fine collabral threads on the protoconch are generally not preserved, but if present they number around 30 to 60 per whorl. There is also some variation in the coarseness of the teleoconch sculpture and in the number of columellar and parietal teeth, where some specimens carry weak teeth on the transition between columella and the parietal region, while others are smooth in this area. This, though, is clearly intraspecific variation as a similar large intraspecific variation is found among recent species of Sassia. Ravn (1902) originally named his new species Tritonium fenestratum, but later changed the name to Tritonium (Sassia) faxense Ravn, 1933 as Tritonium fenestratum Vincent, 1878 from the Eocene of Belgium had priority on the original name.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A967FF822B9CC470FB77FA7A.taxon	materials_examined	Type species. Patella cornucopia Lamarck, 1803 b (subsequent designation by Gray 1847). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A967FF822B9CC4A1FAEBF830.taxon	description	Figs 26 V – X	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A967FF822B9CC4A1FAEBF830.taxon	materials_examined	Material. MGUH 33178 and MGUH 33179, both internal moulds; the latter with fragments of shell preserved. Occurrence. The lower Danian ‘ dead layer’ in Dania Quarry, Northern Jutland. Description. Shell fairly thin, limpet-shaped, with variable height; apex above or behind rear margin. Margin irregular, probably conforming to substrate surface, not crenulated within. Shell surface, if preserved, smooth. Interior with very weak anteriorly opened horseshoe shaped muscle scar. Specimen MGUH 33179 additionally with weak axial ridge anteriorly, reaching 2 / 3 of the distance from apex out towards anterior margin, dividing a left and right attachment area. Measurements. Largest specimen, MGUH 33178, 23.8 mm long at base, 19.5 mm wide and 16.0 mm high. Remarks. The taxon may be conspecific with the Danian (?) Patella nov. sp. of Kaunhowen (1898: p. 15, pl. I: 3 – 4) from Holland. It appears to have been more thin-shelled than Hipponix dunkerianus Bosquet, 1848 as illustrated by Binkhorst (1861: p. 58, pl. IV: 12 a – c) from the upper Maastrichtian of Limburg, the Netherlands.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A968FF8D2B9CC0F4FB2DFDD6.taxon	materials_examined	Type species. Turbo subulatus Donovan, 1804 accepted as Eulima glabra (da Costa, 1778) (subsequent designation by the International Commission on Zoological Nomenclature 1993). Recent, Weymouth in England.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A968FF8D2B9CC1D4FB99FE84.taxon	materials_examined	Remarks. This family is extremely difficult to differentiate taxonomically based upon shell morphology as pointed out by Bouchet & Warén (1986). In most cases the higher classification may not be solved without knowledge of the soft parts and / or colouration of the shell, both features not usually preserved in the fossil record. As a consequence the generic assignment of the following species may never be more than tentative.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A968FF8C2B9CC3CBFCAEFEA0.taxon	description	Figs 26 Y – Z 1933 Eulima (Polygyreulima) danica n. sp. Ravn: 41, pl. IV, figs 1 a – b, 2 a – b.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A968FF8C2B9CC3CBFCAEFEA0.taxon	materials_examined	Diagnosis. Shell relatively high, straight, with highly flattened whorls separated by nearly invisible sutures. Transition to base moderately convex. Aperture relatively high and narrow with very sharp upper angle and gently sigmoid inner side. Columellar lip distinct with evenly rounded outer margin. Outer lip gently sigmoid. Material. Material examined herein consists of holotype MGUH 3148 and paratype MGUH 3149 from the Danian ‘ Naesekalk’, specimens MGUH 33180, MGUH 33181, ØSM. 10042 - 313 - a and b, three specimens from the informally catalogued samples SR. 575, SH. 141. A – B and SH. 169. B, and five uncatalogued moulds from the lower Danian Cerithium Limestone. The latter specimens are deposited in the old collections at the Natural History Museum of Denmark. Type stratum and type locality. The middle Danian ‘ Naesekalk’ at Faxe Quarry, Sealand, Denmark. Occurrence. Eulima? danica is known from an unspecified level within the lower Danian Cerithium Limestone Member at Skeldervig and from the lower 10 cm of the Cerithium Limestone Member at Højerup Church, Stevns Klint. The type material belongs to the middle Danian ‘ Naesekalk’ in Faxe Quarry, Denmark. In addition a single specimen, ØSM. 10042 - 313 - a + b, is known from the ‘ Krabbelaget’, a hardground located two metres up in the Korsnaeb Member. A further specimen in the collections of C. Heinberg comes from the lower Danian ‘ dead layer’ at Vokslev, Northern Jutland. Description. Slender shell generally nearly straight, awl-shaped conical with a height corresponding to around 3.7 times the width. Protoconch consisting of 1 3 / 4 smooth and moderately convex whorls. Protoconch differing only slightly from teleoconch by a distinctly stronger whorl convexity. Teleoconch whorls relatively high and flat with a smooth surface except for the weak growth lines. A few moderately weak incremental scars may be present, while a false suture is absent or at most very faintly developed slightly below the shallow suture. Aperture high and narrow, the height corresponding to nearly 2.6 times the width. Outer lip straight in frontal view, in lateral view slightly prosocline, strongly projecting with strongest projection below mid-line. Base of aperture slightly pointed. Parietal callus thin but distinct. Umbilicus absent. Measurements. Shell of most complete specimen from the Cerithium Limestone, MGUH 33180, 9.2 mm high and 2.5 mm wide with an aperture measuring 3.1 mm in height and 1.2 mm in width and consisting of 7 ½ teleoconch whorls. Last whorl 4.8 mm in height. Remarks. Eulima? danica could also belong to the closely related genus Melanella Bowdich, 1822, but differs by the relative height of the aperture corresponding to nearly 2.6 times the width, whereas Melanella typically is characterized by an aperture H / W ratio of less than 2.4 (Bouchet & Waren 1986) and by a more convex outer lip profile (pers. comm. A. Warén, 2009). Another possible genus is Polygyreulima Sacco, 1892, which generally is characterized by a relatively slender spire like the present species, but the very straight outer lip profile would rather suggest an assignment to Eulima (pers. comm. A. Warén, 2009).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A969FF8C2B9CC0D8FD74FC60.taxon	description	Fig. 26 AE	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A969FF8C2B9CC0D8FD74FC60.taxon	materials_examined	Material. Specimen MGUH 33182 consists of an internal and external mould of the lower half of the shell. Occurrence. The lithified top of the Højerup Member of the upper Maastrichtian Tor Formation at Højerup Church, Stevns Klint. Description. Protoconch unknown. Teleoconch whorls flat, separated by indistinct suture; whorl height corresponding to 2 / 3 the width. Aperture moderately high and narrow tear-shaped, the H / W ratio corresponding to 2.4 or just above. Inner aperture margin sigmoid with nearly straight columella. Measurements. MGUH 33182 is 2.85 mm high and 1.25 mm wide, consisting of three teleoconch whorls. Remarks. The taxon strongly resembles Eulima? danica (Ravn, 1933) treated above, but differs through a relatively wider shell and aperture and a slightly more curved outline of the inner aperture margin. It resembles the Late Cretaceous Eulima gracilistylis Sohl, 1964 a and E. persimplicata (Wade, 1926) from Mississippi, USA, but differs in the narrower aperture and a more slender outline.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A969FF8C2B9CC298FBE0FBD1.taxon	materials_examined	Type species. Melanella dufresnei Bowdich, 1822 (type by monotypy). Recent, Indo-Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A969FF8C2B9CC5CEFC77F8E1.taxon	description	Figs 27 A – B	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A969FF8C2B9CC5CEFC77F8E1.taxon	materials_examined	Material. A single external mould, MGUH 33183. Occurrence. Lower Danian Cerithium Limestone 5 – 600 metres south of Harvig Stige, Stevns Klint. Description. Shell short, broadly conical with the height corresponding to two times the width. Shell wall thick. Protoconch homeostrophic, consisting of around two convex whorls. Teleoconch whorls flat, smooth, separated by shallow sutures. Incremental scars not strongly developed. Aperture small and low, the height corresponding to 38 % of shell height on measured specimen and 1.7 times the aperture width. Outer lip slightly curved in frontal view, in lateral view strongly prosocline, describing a gently convex curve with no central projection. Inner lip curved with no parietal callus. Measurements. Measured specimen 2.6 mm high and just over 1.2 mm wide. Aperture height 1.0 mm and width 0.6 mm. The shell consists of 6 ½ to 7 whorls, protoconch included. Remarks. The relatively low and wide shell makes this more likely to represent a Melanella than either Eulima or Polygyreulima, both of which it also closely resembles. The relatively low spire, the flattened whorls with fairly strongly prosocline growth lines and the relatively wide aperture makes this species stand out from most related species. It shows some resemblance to the much younger Melanella (Polygyreulima) montilysensis Bałuk, 1995 from the Middle Miocene of Poland, but differs in the relatively higher aperture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8F2B9CC779FA25F972.taxon	materials_examined	Type species. Serpulorbis polyphragma Sasso, 1827 (subsequent designation by Keen 1961: p. 191). Recent, Mediterranean.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8F2B9CC1D4FAE0FF15.taxon	materials_examined	Type species. Vermetus adansoni Daudin, 1800 (subsequent designation by Cossmann 1912: p. 134). Recent, Senegal.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8F2B9CC004FDFEFA01.taxon	description	Figs 27 C – E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8F2B9CC004FDFEFA01.taxon	materials_examined	Material. MGUH 33184, MGUH 33185, MGUH 33186, ØSM. 10042 - 134 - a, ØSM. 10042 - 187, ØSM. 10042 - 188, ØSM. 10042 - 386, ØSM. 10042 - 388 - b and ØSM. 10042 - 393. The collection of the Natural History Museum of Denmark furthermore contains several specimens lacking numbers. These may be grouped into 14 very fragmentary external teleoconch moulds from the upper Maastrichtian Højerup Member (samples: SH. 4, SR. 773, SR. 914, SR. 921, SR. 924. A, SR. 1039, SR. 1067. A – B, SR. 1069. A – B, SR. 1086. A – B, SR. 1091. B, SR. 1092. A – B, SR. 1105, SR. 1107, SR. 1114. A) and 52 fragmentary moulds from the Lower Danian Cerithium Limestone Member (informal sample numbers: SH. 80. B, SH. 84, SH. 149. B – C, SH. 169. A, SH. 174, SH. 191, SH. 234, SH. 254, SH. 277, SH. 297. A – B, SH. 302. A, SH. 415. B, SH. 454, SH. 455, SH. 458. A – B, SH. 468. A, B, SH. 483. A – B, SH. 483. B – C, SR. 143, SR. 146, SR. 189, SR. 211. A – B, SR. 250, SR. 253, SR. 281 (2 specimens), SR. 285, SR. 291, SR. 295, SR. 312, SR. 331, SR. 372, SR. 383, SR. 384, SR. 389. A – B, SR. 400, SR. 404. A – B, SR. 450, SR. 458, SR. 586. A – B, SR. 597, SR. 616 (2 specimens), SR. 622, SR. 632, SR. 642. A – B, SR. 648, SO. 90. A – B, SO. 101, SO. 176, and two without number from the old collections at the Natural History Museum of Denmark). Occurrence. Lithified top of the upper Maastrichtian Højerup Member at Rødvig and Højerup Church. Low- er Danian Cerithium Limestone Member at Rødvig, Højerup Church, Skeldervig and Holtug, Stevns. The lower Danian ‘ dead layer’ at Vokslev, Northern Jutland. The lower hardground (‘ krabbelaget’) in the Korsnaeb Member at Korsnaeb, Stevns Klint. The best exposed protoconch possibly comes from the infilling of Korsnaeb Member into a burrow in the lower middle part of the Cerithium Limestone Member at Højerup Church. Description. Protoconch 0.65 to 0.9 mm high and 0.5 to 0.55 mm wide and regularly coiled trochispiral with 3 ½ to 4 whorls. Surface without sculpture. Teleoconch uncoiled, irregular with few unevenly distributed and moderately sharp spiral ribs and fairly strong, irregularly distributed growth increments. Older specimens may have a denser spiral sculpture with more rounded ribs. Aperture tube-shaped. Measurements. Largest specimen from Holtug with a largest whorl diameter of 6.6 mm and aperture width of 2.6 mm. Remarks. Except for the protoconch specimen, which is located inside a bryozoan colony, none of the vermetid tubes studied is attached to a substrate. Most probably they have been associated with poriferans, which based on the numerous spicules present in the rock must have been quite abundant in the palaeo-environment.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8E2B9CC7AFFAA4FB24.taxon	description	Figs 27 F – J 2014 Serpulorbis sp. Hansen & Surlyk: 339, fig. 6: 1, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96AFF8E2B9CC7AFFAA4FB24.taxon	materials_examined	Diagnosis. Young shell regularly curled with coarse transverse ribbing on concave flanks and equally coarse spiral ribbing on abapical side. Transition between the two sides sharp. Derivation of name. Refers to the contrasting sculpture of spiral ribs and transverse ribs on coiled shell. Type material. Holotype MGUH 33187 is an external and internal mould of a coiled juvenile shell. Paratype MGUH 33188 is an external mould of a coiled juvenile shell attached to a cephalopod and found at Skeldervig. Paratype MGUH 33189 is from just south of Højerup Church, while paratype MGUH 33190 is from Rødvig, Stevns. All three paratypes belong to the same horizon as the holotype. Additional material. ØSM. 10042 - 199 - c, ØSM. 10042 - 244 - c, and 29 informally catalogued specimens with sample numbers SH. 2. A, SH. 41, SH. 42, SR. 778, SR. 779, SR. 780, SR. 783. B, SR. 787. D, SR. 914, SR. 916. A, SR. 926. B, SR. 934. A, SR. 984, SR. 987, SR. 1017, SR. 1054, SR. 1055, SR. 1056, SR. 1057. A – B, SR. 1058. A – B (2 specimens), SR. 1060. A – B, SR. 1076. A – B, SR. 1110, SR. 1111. A – B, SR. 1119, SR. 1121, SR. 1127 (2 specimens). A last small specimen is found together with the holotype. Type stratum and type locality. Lithified top of the upper Maastrichtian Højerup Member of the Tor Formation from just north of Holtug Quarry. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig, Skeldervig, Højerup Church, Mandehoved and Holtug Quarry, Stevns. Description. Young shell regularly coiled, very low with flattened spire attached to hard substrate (cheilostome bryozoans); whorl strongly widening adapically on peripheral side, increasing attachment surface; lateral flank slightly concave, changing to moderately convex on abapical side. Aperture obliquely oval. At least third whorl carrying pronounced, fairly coarse spiral ribs on abapical side, abruptly changing to equally strong and evenly distributed, forwardly inclined oblique transverse ribs. Shell wall moderately thin, strongly thickening peripherally on apical side. Mature shell irregularly coiled; abapical surface covered by more or less unevenly distributed, fairly coarse spiral ribs and fine growth lines; sides covered by pronounced and fairly regular growth increments or ribs and weak to nearly effaced spiral ribs. Spiral ribs may be substituted by relatively fine and short transverse ribs. Measurements. Coiled young shell, paratype MGUH 33189, 1.1 mm high and 2.8 mm wide with an aperture width of 1.0 mm. Remarks. The uncoiled teleoconch resembles somewhat Vermetus sp. described above from the succeeding Rødvig Formation, but differs in the pronounced division between spiral ribbing in the abapical side and transverse ribbing or growth increments on the lateral sides and in the coarser transverse ribs. The general outline of the present species resembles some specimens of the recent Thylacodes adamsii (Mørch, 1859) except for the strong transverse ribs. To the knowledge of the author this is the oldest described possible representative of this genus, which previously has been reported from the mid to late Danian de Vigny deposits in France (Pacaud et al. 2000).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96BFF8E2B9CC446FD0DF8A1.taxon	description	Figs 27 K – M	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96BFF8E2B9CC446FD0DF8A1.taxon	materials_examined	Material. Specimen MGUH 33191 is a fragmentary 2.4 mm wide external mould. Occurrence. Thylacodes? sp. was collected from the top of the Maastrichtian Tor Formation at Holtug Quarry, Stevns Klint. Description. Shell low turbiniform, sinistral with flattened apex seemingly attached to a slender bryozoan branch. Protoconch heterostrophic, consisting of two whorls on which no microsculpture can be observed. Teleoconch whorls expanding slowly, flattened on adapical side, becoming more convex on lateral and abapical side. Adapical side of whorls with strong transverse ribs curving from the adapical suture and backwards into an initially indistinct, but progressively deeper sinus situated halfway between sutures; transverse ribs continue forward and down to the periphery of the last whorl, where they are replaced rather abruptly by seven moderately coarse spiral ribs. Transverse rib number per half-whorl approximating 15 on third whorl. Remarks. Thylacodes? sp. differs from T. contendis n. sp. described above by the sinistral coiling and a coarser transverse ribbing. This may though just reflect ecophenetic differences within the species; especially as only a single specimen is known of the present taxon.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96DFF882B9CC716FB3CF954.taxon	materials_examined	Type species. Laxispira lumbricalis Gabb, 1877 (type by monotypy). Cretaceous, New Jersey, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96DFF8B2B9CC644FBE5FE69.taxon	description	Fig. 27 O	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96DFF8B2B9CC644FBE5FE69.taxon	materials_examined	Material. MGUH 33193, an external mould in the unformally catalogued sample SH. 3. B – C, and a last mould without number in the old collections at the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Tor Formation at Højerup Church and Holtug Quarry, Stevns. Description. Protoconch not known. Teleoconch loosely coiled with round aperture and apparently without keel. Shell sculpture of well defined, densely spaced granular spiral ribs crossed by densely spaced growth lines. Spiral ribs may become slightly coarser on abapical side of whorl. Measurements. Largest specimen with aperture diameter of 11.3 mm. Remarks. The taxon has a strong resemblance to the Maastrichtian L. libycus Quaas, 1902 from Egypt and the Late Maastrichtian L. pinguis Holzapfel and L. cochleiformis (Müller) from Holland as figured by Wenz (1939), but differs by its strongly and evenly beaded spiral ribs. It also resembles Laxispira monilifera Sohl, 1964 b from the Late Cretaceous of Mississippi, but differs by a more evenly beaded spiral sculpture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96EFF8B2B9CC3EDFB20FD1F.taxon	materials_examined	Type species. Triforis grignonensis Deshayes, 1864 (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96EFF8A2B9CC20FFC7DFDDD.taxon	description	Figs 27 P – R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96EFF8A2B9CC20FFC7DFDDD.taxon	materials_examined	Diagnosis. Very slender shell with sigmoid collabral threads and no spiral sculpture on protoconch whorls and with opisthocline, truncated transverse ribs on early teleoconch whorls. Derivation of name. Refers to the type locality at Skeldervig, Stevns Klint. Type material. Holotype MGUH 33194 is a nearly complete specimen preserved as an external mould. Paratype MGUH 33195 comes from the same stratum as the holotype and was collected at Holtug Quarry, Stevns Klint. Paratype MGUH 33196 is from the Cerithium Limestone Member at Skeldervig, Stevns Klint. Additional material. An external mould from Holtug is deposited in the uncatalogued old collections at the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member of the Rødvig Formation at Skeldervig, Stevns Klint. Occurrence. Lower Danian Cerithium Limestone Member of the Rødvig Formation at Skeldervig and Holtug Quarry. Lower Danian Korsnaeb Member north of Holtug at Stevns Klint. Description. Protoconch planktotrophic with around 2 ½ moderately to weakly convex whorls; protoconch height approximately 0.4 mm and width 0.3 mm. Sculpture on embryonic shell unknown. Last protoconch whorl carrying approximately 14 fairly coarse, slightly sigmoid collabral threads, the main adapical part opisthocyrt with a more sharply prosocyrt curvation at abapical suture. Spiral threads absent. Transition to teleoconch not visible. Spire slender turriform, sinistrally coiled, the first two teleoconch whorls seemingly smooth and moderately to weakly convex. Succeeding whorls flattened with sharp to later more gradual transition to base; whorl height corresponding to less than half the width. Base strongly flattened on early teleoconch whorls, becoming more convex with growth. Aperture relatively small, narrow with straight and smooth columella carrying large columellar and parietal lip. Outer lip slightly thickened. Canal not visible or preserved. Growth lines fine, opisthocline, nearly straight and densely distributed. Teleoconch sculpture consisting of 18 to around 26 opisthocline transverse ribs per whorl and wide and somewhat effaced spiral ribs and furrows. Transverse ribs only distinct on abapical two spiral ribs of early whorls; fading out before completely disappearing on later whorls. Spiral sculpture composed of two narrow and low spiral ribs abapically, divided by shallow furrow and succeeded adapically by moderately deep spiral furrow situated on abapical half of whorl. Adapical part of whorl carrying two wide and indistinct spiral bands divided by weak spiral furrow. Whorl base characterized by eight or more weak and closely lying spiral ribs, becoming fainter towards columella. Measurements. Holotype MGUH 33194 is 9.6 mm high and 2.4 mm wide, consisting of 20 whorls (protoconch included). Remarks. The pronounced opisthocline axial sculpture with well developed, though truncated transverse ribs may indicate that the Danish taxon from the Cerithium Limestone Member most likely does not belong to this genus sensu stricto. The material, though, is too poor for the erection of a new genus and the taxon is at the moment assigned Epetrium sensu lato.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96FFF8A2B9CC4ABFBFCF9A1.taxon	materials_examined	Type species. Murex tubercularis Montagu, 1803 (type by monotypy). Recent, European seas.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96FFF952B9CC7D9FC9CFAB4.taxon	description	Figs 27 T – W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A96FFF952B9CC7D9FC9CFAB4.taxon	materials_examined	Diagnosis. Shell high and slender, more turriculate than cyrtoconoid; teleoconch whorls relatively wide with four spiral ribs of which only the 2 nd and 3 rd are tuberculated; transverse ribs strong and rounded, approximately 12 per whorl. Derivation of name. The name, which translates ‘ long rope’, refers to the slender and rope-like outline of the shell. Type material. The holotype, MGUH 33198 is an external mould with preserved protoconch, while paratype MGUH 33199 is an external mould from the lithified top of the Tor Formation at Stevns Klint just north of Stevns Fyr and paratype MGUH 33200 is a well preserved protoconch mould from the very top of the Cerithium Limestone Member at Rødvig. Paratype MGUH 33201 is a protoconch from the Cerithium Limestone Member, 40 to 50 cm above the K-Pg boundary at Rødvig, Stevns Klint. Additional material. Four fragmentary moulds from the informally catalogued samples SH. 281. A and C, SR. 406, SR. 744 and SR. 919, and a mould without number from north of the lighthouse Stevns Fyr. All are deposited in the collections at the Natural History Museum of Denmark. Type stratum and type locality. Holotype MGUH 33198 is collected from the lithified interval encompassing the K-Pg Boundary at Skeldervig, Stevns Klint. Occurrence. Top Højerup Member of the Maastrichtian Tor Formation to the top of the lower Danian Cerithium Limestone Member of the Rødvig Formation at Stevns Klint. Description. Shell high, extremely slender, the teleoconch only widening very slightly abapically. Protoconch turriform multispiral, consisting of 5 to 5 ½ smooth and convex whorls, evenly widening down to last two whorls at which the size increase lessens. The last whorls may show very weakly developed abapical spiral lirae and around 25 to 30 indistinct opisthocline threads per whorl. Last whorl carrying one or two weakly developed spiral ribs, which continue on to the teleoconch whorls. Transition from protoconch to teleoconch rather abrupt. The teleoconch consists of up to 12 relatively low and wide whorls with the whorl height corresponding to just over 60 % of width. Teleoconch whorls flatly convex, weakly reticulate with four spiral ribs and around 12 strong and rounded transverse ribs centrally on whorl. First spiral rib weak and narrow, bordering deep suture; second rib strongest, followed by a slightly weaker third spiral rib, both well rounded and coarsely tuberculated; fourth rib fairly weak, located just above suture, on the last whorl continuing out onto sharp keel. Transverse ribs generally slightly opisthocline, but may become orthocline to weakly prosocline. Base of shell flat to slightly concave, smooth but for weak and closely spaced spiral ribs and growth lines. Measurements. Holotype MGUH 33198, comprising 12 whorls, is 4.0 mm high and 0.9 mm wide with a 1.05 mm high and 0.4 mm wide protoconch. Remarks. Cerithiopsis restemlongis n. sp. is distinguished from most species belonging to this genus by its very high and slender spire and the presence of only two tuberculated ribs, which are well rounded in cross-section. It resembles the Danish middle Danian Cerithiopsis cf. rosenkrantzi (Ravn, 1933) sensu Nützel (1998), but is distinguished by the distinctly lower whorls and seemingly slightly higher number of transverse ribs per whorl. It differs from the middle Danian Cerithiopsis rosenkrantzi (Ravn, 1933) sensu stricto from Denmark by the presence of only two tuberculated spiral ribs and by the flatter whorls. It also appears very close to a rather poorly preserved species of Cerithiopsis from the Campanian of Spain figured by Kiel (2001), but appears to differ by a less inflated protoconch whorl and a markedly larger protoconch size. It has a strong resemblance to Zaclys? sp. sensu Nützel (1998) from the middle Danian limestone deposits at Faxe, Denmark. It differs by having a much higher protoconch with more convex whorls and by a flatter teleoconch whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A970FF952B9CC4E4FC95F874.taxon	description	Fig. 27 X	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A970FF952B9CC4E4FC95F874.taxon	materials_examined	Material. A single very fragmentary specimen, MGUH 33202. Occurrence. Lithified top of the Maastrichtian Tor Formation just south of Højerup Church at Stevns Klint. Description. Teleoconch whorls flattened, twice as wide as high and separated by rather indistinct sutures. Base flattened. Columella smooth without folds. Sculpture consists of three moderately weak spiral ribs with strongly developed, rounded tubercles numbering around 16 per whorl. Transverse sculpture only present in the form of a weak, opisthocyrt organization of the tubercles. Measurements. MGUH 33202, consisting of two teleoconch whorls, 0.65 mm high and nearly 0.6 mm wide. Remarks. The specimen strongly resembles the Danian Cerithiopsis subcylindracea (Briart & Cornet, 1873) from Belgium with which it may be conspecific. The Danish taxon is kept under open nomenclature due to the extremely poor material and the slight stratigraphic separation.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A971FF942B9CC1D4FEBAFEF7.taxon	materials_examined	Type species. Seila (Notoseila?) angusta Tembrock, 1964 (type by original designation). Early Oligocene, Paris Basin in France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A971FF972B9CC02BFEEFFE14.taxon	description	Figs 27 Y – AE, 28 A	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A971FF972B9CC02BFEEFFE14.taxon	materials_examined	Diagnosis. Protoconch of five whorls with densely spaced oblique axial ribs and spiral ribs on the last half whorl. Teleoconch with flat whorls carrying four spiral ribs, the third generally much stronger than the others while the fourth is covered by succeeding whorl. Derivation of name. In honour of the Carlsberg Foundation, which financed this project. Type material. The holotype MGUH 33203 is an external mould collected by A. Rosenkrantz, 20 Sep. 1927. Paratype MGUH 33204 comes from Rødvig, while paratype MGUH 33205 is from Skeldervig. Both are from the same stratigraphic level as the holotype. Additional material. ØSM. 10042 - 111, 201 internal and external moulds collected in connection with this study (sample number: SH. 107. A – B, SH. 142, SH. 174, SH. 183. B, SH. 206, SH. 231, SH. 251, SH. 254, SH. 274. A – C, SH. 274. C (3 specimens), SH. 281. A, SH. 287. B, SH. 287. C (2 specimens), SH. 288, SH. 289, SH. 290, SH. 291. A, SH. 291. B, SH. 292, SH. 295. A, SH. 310, SH. 318. A – B, SH. 320, SH. 325. A, SH. 328. A – B, SH. 329, SH. 330, SH. 331, SH. 332, SH. 333. A – B, SH. 334. A – B, SH. 349, SH. 350, SH. 352. B, SH. 353, SH. 359, SH. 377, SH. 382 (4 specimens), SH. 385. B (2 specimens), SH. 385. B – C, SH. 385. C (2 specimens), SH. 386. C, SH. 388 (2 specimens), SH. 394, SH. 396, SH. 404, SH. 405, SH. 410. A – B, SH. 410. B, SH. 414. C, SH. 415. A, SH. 419. B, SH. 424, SH. 425, SH. 437. A, SH. 445, SH. 465 (2 specimens), SH. 466, SH. 467, SH. 468. A, SH. 473, SH. 485. C, SR. 143, SR. 144, SR. 146, SR. 147. A – B, SR. 148, SR. 150, SR. 151, SR. 152. A – B (2 specimens), SR. 152. A, SR. 264, SR. 275 (2 specimens), SR. 276. A – B (4 specimens), SR. 277. C (2 specimens), SR. 279, SR. 280, SR. 281, SR. 283, SR. 284, SR. 287 (2 specimens), SR. 288 (2 specimens), SR. 290, SR. 294 (2 specimens), SR. 295 (4 specimens), SR. 296. A – B (3 specimens), SR. 297 (2 specimens), SR. 298. A (3 specimens), SR. 298. B, SR. 301 (2 specimens), SR. 302, SR. 303. A – B (2 specimens), SR. 305. A – B, SR. 307. A, SR. 308. A – B, SR. 309. A, SR. 310. B (2 specimens), SR. 310. B – C (2 specimens), SR. 311. A (2 specimens), SR. 311. A – B, SR. 320, SR. 340, SR. 359, SR. 360. A – B, SR. 361, SR. 369, SR. 372, SR. 374, SR. 381, SR. 394, SR. 398, SR. 399, SR. 400, SR. 401, SR. 402, SR. 403. B, SR. 404. A – B, SR. 406, SR. 432, SR. 439, SR. 449, SR. 451, SR. 464, SR. 473, SR. 480, SR. 490, SR. 494, SR. 495, SR. 496, SR. 501, SR. 507, SR. 508, SR. 509. A – B, SR. 513, SR. 517, SR. 519. A, SR. 521, SR. 522, SR. 523, SR. 524, SR. 529, SR. 548, SR. 552, SR. 562, SR. 567, SR. 569, SR. 577, SR. 578. B – C, SR. 587. A – B, SR. 588. B, SR. 591, SR. 614, SR. 619, SR. 623, SR. 624, SR. 641, SR. 652, SR. 653, SR. 654, SR. 655 (2 specimens), SR. 656 (2 specimens), SR. 685 (2 specimens), SR. 698. A, SO. 185, DN. 19 and DN. 26. B), and an unspecified number found in the collections of the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member of the Rødvig Formation just north of Kirkevig, Stevns Klint. Occurrence. This species occurs from the lower half to the top of the lower Danian Cerithium Limestone at Stevns Klint. It is also known from the ‘ dead layer’ in Dania and Vokslev quarries, Northern Jutland. Description. The shell is high turriculate, more than five times higher than wide. The last whorl equals close to a quarter of the total shell height and the aperture approximately 1 / 8 the total height. Protoconch multispiral, high conical with five moderately convex whorls separated by fairly deep sutures. First 1 ½ whorls smooth, the following covered by up to around 22 oblique collabral threads. Two weak upper and a strong abapical spiral rib appear less than half a whorl from teleoconch. Transition to teleoconch fairly gradual, being suggested by the disappearance of collabral threads. Teleoconch consisting of up to 11 flat whorls with two narrow adapical spiral ribs of equal strength and a normally much stronger abapical rib situated on a weak carina somewhat above a narrow suture. On a few specimens the spiral ribs are of equal strength or the adapical one becomes slightly stronger than the other two. Distance between the crest of all three ribs equal; interspaces between ribs becoming wider than ribs on later whorls. A fourth spiral rib occurs just below abapical suture, rib typically more or less completely covered by succeeding whorl. Base of whorl flat. Aperture rounded rectangular with a short, smooth and straight columella with the short canal turned left. Measurements. The largest complete specimen measures 7.4 mm in height and 1.4 mm in diameter. Protoconch approximating 0.4 mm in width and 0.8 mm in height. Remarks. Thereitis carlsbergi n. sp. differs from the Selandian Thereitis monbergi (Ravn, 1939) from Denmark by the presence of spiral ribs on the last protoconch whorl, the absence of axial riblets on the teleoconch, the wider interspace between the spiral ribs and by the relatively higher spire. It is distinguished from the Selandian Thereitis weinbrechti Schnetler, 2001 from Denmark by the absence of spiral ribs before the last protoconch whorl and by the stronger third spiral rib on the teleoconch. It is furthermore distinguished from the middle Danian Thereitis tricingulata (Ravn, 1933) from Faxe, Denmark, by having only five protoconch whorls and possibly also by a slightly different spiral sculpture, although the latter may be due to the limited material available (two specimens) of T. tricingula.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A972FF972B9CC305FBC9F872.taxon	description	Figs 28 B – D 2014 Thereitis sp. Hansen & Surlyk: 339, fig. 5: 19, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A972FF972B9CC305FBC9F872.taxon	materials_examined	Diagnosis. Protoconch of five whorls with densely spaced oblique collabral threads and two spiral ribs after initial 1 ½ whorl. Teleoconch with flat whorls carrying four spiral ribs, the adapical one generally much stronger than the others while the fourth is covered by succeeding whorl. Siphonal canal strongly oblique. Derivation of name. Refers to its status as the first representative from the Cretaceous. Type material. The holotype MGUH 33206 is a protoconch and initial teleoconch whorl preserved as an external and internal mould. Paratype MGUH 33207 comes from Holtug Quarry, while paratype MGUH 33208 was collected near Stevns Fyr, Stevns Klint. The two paratypes are preserved as external moulds and come from the lithified top of the Højerup Member. Additional material. Six moulds with the informal sample numbers SR. 734. B, SR. 741. B, SR. 990. A – B, SR. 997. B – C, SR. 1090. A – B and SR. 1109 were collected in connection with this study. Type stratum and type locality. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Holtug Quarry, Stevns Klint. Occurrence. Top of the Maastrichtian Tor Formation at Rødvig, Holtug Quarry and north of Stevns Fyr, Stevns Klint. Description. Protoconch just over 0.5 mm long and 0.35 mm wide, multispiral, high conical with five moderately convex whorls separated by fairly deep sutures. First 1 ½ whorls smooth, the following covered by up to around 22 oblique collabral threads. Two sharply defined spiral ribs located close to the adapical and abapical suture, respectively, appearing together with the axial riblets. The adapical spiral rib moves down to the periphery on the last two protoconch whorls. Teleoconch consisting of up to 12 flat whorls with four spiral ribs of slightly varying strength. Spiral rib 1 usually distinctly stronger than abapical ones and situated just below adapical suture. Spiral rib 2 and 3 separated by wide interspace; spiral rib 3 located immediately above abapical suture, in part merging with weak spiral rib 4 situated just below suture line. Whorl base flat. Aperture rounded rectangular with a short, smooth and straight columella terminated by short and strongly transverse canal. Aperture takes up less than 20 % of total shell length. Measurements. The two largest specimens measure 4.3 mm in height (protoconch excluded) and approximately 1 mm in diameter and consist of 12 teleoconch whorls. Aperture height of MGUH 33207 is approximately 0.5 mm, while the lower whorl takes up around 0.9 mm of the teleoconch height. Remarks. This species resembles the early Danian Thereitis carlsbergi n. sp. described above, but differs by the smaller size, by the more typical protoconch sculpture, by a stronger left-turn of the apertural canal and by the spiral ribs, where the adapical one is distinctly stronger than the others and situated immediately below the suture, while a wide interspace separates spiral rib 2 and 3, whereas spiral ribs 3 and 4 are nearly fused together due to the slightly lower location of spiral rib 3. It differs from the Selandian (Late Palaeocene) T. weinbrechti Schnetler, 2001 of Denmark by the fewer protoconch whorls with their slightly sparser axial sculpture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A974FF902B9CC716FCA2FF34.taxon	materials_examined	Type species. Cerithium metula Lovén, 1846 (designation by Sars (1878) when erecting Lovenella Sars, 1878, a name already in use). Recent, Europe. Remarks. It was suggested by Nützel (1998) that all fossil taxa with a planktotroph multispiral protoconch may have to be assigned their own genus or reassigned to other genera as all extant species of Cerithiella are characterized by a lecithotroph larval shell. It is clear that the present definition of the genus is rather wide and it seems plausible that the planktotrophic taxa in general would be closer related to each other than to the lecithotrophic forms. I do not think that there is any basis for a higher taxonomical reassignment of the planktotrophic forms within the cerithiopsids, as was suggested on a tentative basis by Nützel (1998). Actually there are all grades of intermediates between the two groups and even within a single recent species like Cerithiella metula (Lovén, 1846) as re-evalu- ated by Bouchet & Warén (1993) a surprisingly large variation in the protoconch morphology may be observed, depending on the environment or geographical location.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A974FF912B9CC39BFED7FA9A.taxon	description	Fig. 28 E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A974FF912B9CC39BFED7FA9A.taxon	materials_examined	Material. Specimen MGUH 33209 is an external mould of the protoconch and first five teleoconch whorls. Occurrence. Lithified top of the Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Protoconch 0.7 mm high and 0.4 mm wide, multispiral, high conical with 5 ½ convex whorls separated by fairly deep sutures. First three whorls strongly expanding, following whorls of nearly equal width. First whorl smooth, the following covered by two peripheral to abapical spiral threads and up to 30 opisthocyrt and opisthocline collabral threads. Teleoconch very slender, nearly parallel-sided, narrower than protoconch; whorls weakly, but distinctly convex, their height corresponding to their width. Teleoconch sculpture dominated by three to five spiral ribs, the three central ones strongest. Weakly developed and slightly prosocline transverse ribs are present between spiral ribs, giving the whorls a slightly reticulate sculpture. Measurements. Specimen MGUH 33209 is 2.3 mm high and 0.4 mm wide, consisting of 10 whorls. Remarks. The taxon differs from Thereitis cretacea n. sp. described above by the extremely slender spire with its narrow, but clearly convex teleoconch whorls carrying up to five spiral ribs. With only one available specimen so far it is unclear whether these differences are all true diagnostic characters or if some may reflect abnormal interspecific variation.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A975FF902B9CC064FD61F93B.taxon	description	Figs 28 F – I 1902 Cerithium Moltkianum Ravn: 223 – 224, pl. 1, fig. 23. 1903 Cerithium Moltkianum Ravn — Ravn: 380, 394 – 395.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A975FF902B9CC064FD61F93B.taxon	materials_examined	Diagnosis. Protoconch with four whorls, first whorl not inflated, the last with around 15 strongly opisthocline collabral threads per half whorl and a weak spiral thread on upper part. Teleoconch columella lacking fold; siphonal termination evenly rounded; spiral ribs 1 and 2 closer than 2 and 3; base carrying only one primary spiral rib. Material. MGUH 33210, MGUH 33211, ØSM. 10042 - 128, the four informally catalogued specimens with sample numbers SR. 295 (2 specimens), SR. 303. A – B and SR. 637. A – B, and 25 external and in a few cases also internal moulds without number belong to the Cerithium Limestone Member. ØSM. 10042 - 381 - a and an external mould informally catalogued by sample number SH. 385. B derives from the basal Korsnaeb Member of the Stevns Klint Formation. Those without numbers are all from the collections of the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member at Stevns Klint. Occurrence. Cerithiella moltkianum (Ravn, 1902) ranges from the lower Danian Cerithium Limestone Member and up into the Korsnaeb Member. It is moderately common in the southern to northern part of the Stevns Klint exposure. A few additional specimens have been collected by C. Heinberg from the lower Danian ‘ dead layer’ at Vokslev, Northern Jutland. Description. Protoconch turriform multispiral with four convex whorls; first 2 ½ whorls possibly smooth, though this may be due to preservation; first whorl not inflated. Last two protoconch whorls with fairly closely spaced opisthocline and moderately opisthocyrt collabral threads, number of collabral threads approximately 35 per whorl. Last protoconch whorl carrying weak spiral thread located at strongest curvature of collabral threads on upper part of whorl. Transition to teleoconch seemingly rather gradual. Protoconch approximately 1.0 mm high and 0.5 mm wide. Teleoconch whorls flat sided with a more or less pronounced, sharp keel located slightly above narrow and very shallow abapical suture; height of whorls corresponding to around 38 to 40 % of whorl width. Base weakly convex. Aperture rounded rectangular, abapically continuing out into oblique siphonal canal; outer lip with broad and fairly deep sinus just above keel; columella lacking fold. Whorl sculpture strong, reticulate, consisting of three primary spiral ribs and a number of slightly weaker axial ribs carrying tubercles at connecting points. Spiral rib 1 not starting before third teleoconch whorl after which it lies closer to spiral rib 2 than 2 lies to the distinctly stronger spiral rib 3 situated on the generally weak keel. Axial ribs numbering between 9 and 10 per half whorl on third teleoconch whorl, terminating just below keel. Number of axial ribs increasing slowly abapically while no secondary spiral ribs are added before the 18 th teleoconch whorl. The base carries a single primary spiral rib slightly below the keel and is succeeded by a wide shallow furrow and densely spaced, weak spiral ribs covering the rest of the base. Measurements. Largest specimen just over 48 mm high and 9.2 mm in diameter with 23 teleoconch whorls. Remarks. C. moltkianum (Ravn, 1902) differs from the diagnostic features of Cerithiella as defined by Gründel (1980) and Nützel (1998) by the lack of columellar folds. Even so the other characters such as the protoconch sculpture clearly indicate a strong relation with this group, and with the general variation in strength and number of columellar folds I do not hesitate to assign it here.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A975FF922B9CC667FC49FD80.taxon	description	Figs 28 J – O	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A975FF922B9CC667FC49FD80.taxon	materials_examined	Diagnosis. Protoconch with four and a half whorls, with collabral threads but no or only very weakly developed spiral ribs on last whorl. Teleoconch whorls with strong keel; columellar plate not strongly developed; columella with weak fold; base carrying only one primary spiral rib. Transverse sculpture fairly dense. Derivation of name. Name of the Danish island Sealand from which it derives. Type material. The holotype MGUH 33212 is an external mould presenting the last protoconch whorl and 13 teleoconch whorls. Paratype MGUH 33213, consisting of an external and fragments of the internal mould, comes from the same locality and horizon, while paratype MGUH 33214, an external mould of protoconch and teleoconch, was collected from Vokslev Quarry, Northern Jutland. Paratype MGUH 33215 is an external spire mould from the Cerithium Limestone Member at Holtug, Stevns Klint. Additional material. ØSM. 10042 - 178 - a – c, eight specimens collected in connection with this study and found in the informally catalogued samples SH. 105. A – B, SR. 296. B, SR. 476. A – C, SR. 565, SR. 620. A – B, SO. 93, DN. 1 and DN. 52. A – B, and 15 external and in a few cases internal moulds deposited at the Natural History Museum of Denmark. An additional specimen from Dania Quarry is deposited in the private collection of K. I. Schnetler. Type stratum and type locality. Cerithium Limestone Member at Skeldervig, Stevns Klint. Occurrence. Cerithiella selandica n. sp. ranges from the lower part of the lower Danian Cerithium Limestone Member and up into the lower part of the Korsnaeb Member. It is moderately common in the southern to northern part of the Stevns Klint exposure and is also found in the corresponding layers at Vokslev and Dania in Northern Jutland. Description. Protoconch turriform multispiral with approximately 4 ½ convex whorls; initial whorl seemingly smooth, the following carrying around 14 to 18 fine, strongly opisthocline and somewhat opisthocyrt collabral threads per half whorl. Last whorl with two faintly developed spiral threads centrally. Transition to teleoconch very gradual, showing a continuation of spiral ribs and densely spaced collabral threads, the former becoming much stronger and adjoined by weak abapical one located slightly adapical to suture. Protoconch 0.9 mm high and 0.5 mm wide. First teleoconch whorls high with marked keel and concomitant deeply V-shaped sutural groove, suture narrow and shallow. Succeeding whorls flat sided with a pronounced, sharp keel located slightly above narrow abapical suture; height of whorls corresponding to nearly 50 % of whorl width. Base weakly convex. Aperture rounded subrectangular to pentagonal, abapically continuing out into relatively long siphonal canal; siphonal canal strongly oblique; outer lip thin with broad and fairly deep sinus just above keel; parietal lip strongly developed but thin; columellar plait low and broad, located on lower part of slightly concave columella, accentuating weak canal. Whorl sculpture initially weakly reticulate with densely spaced collabral threads and three spiral ribs, two centrally on the whorl and a weaker abapical one located slightly above suture. Collabral threads describing forwardly directed sinus around abapical spiral thread. Later whorls strongly reticulate, reticulation consisting of 12 to 16 transverse ribs per whorl on third or fourth teleoconch whorl, crossed by slightly stronger spiral ribs. Additional spiral ribs are added to three initial ones, which become the second to fourth spiral ribs with the fourth moving down below the suture line and the third and strongest situated on keel. Spiral ribs 2 and 3 closer than 1 and 2. Transverse ribs terminating just below keel. Number of transverse ribs increasing abapically to around 40 ribs on the 20 th teleoconch whorl, while the tubercles at the rib junctions and the transverse and adapical two spiral ribs become weaker. Third spiral rib becoming higher and more sharply delineated on later whorls. Base carrying a single primary spiral rib succeeded by a weak, but wide furrow and densely spaced weak spiral ribs covering the whole lower base. Measurements. Largest specimen, MGUH 33213, is 31 mm high and 7.1 mm in diameter with 18 teleoconch whorls. Remarks. Cerithiella selandica n. sp. closely resembles the Palaeocene Cerithiopsis sp. 1 sensu Kollmann & Peel (1983) from West Greenland, but C. selandica n. sp. has fewer transverse ribs and the fourth primary spiral rib is not exposed before the final whorl. It differs from the Palaeocene Cerithium zigzag Grönwall & Harder, 1907 from Denmark by the less concave spire profile; a slightly sharper transition between base and canal and by a denser transverse ribbing. Furthermore the growth lines appear in general to be straighter with a weaker bend at transition to base. This species may look somewhat similar to Cerithiella moltkianum treated above, but differs in that spiral ribs 2 and 3 are closer to each other than spiral ribs 1 and 2; the number of transverse ribs on the adapical teleoconch whorls is normally lower and the whorls are generally higher and with a more pronounced keel. It is furthermore distinguished from the Late Cretaceous Cerithium tectiforme Binkhorst, 1861 from Holland by the distinctly lower number of transverse ribs per whorl, though they are clearly closely related. The teleoconch of Cerithiella selandica n. sp. closely resembles that of Orthochetus Cossmann, 1889, which recently has been revised by Darragh (2011). Darragh (2011) assigned several Palaeogene taxa to this genus based solely upon their teleoconch morphology and prominent keel, although recognizing that some species among the Cerithiella strongly resembled them but for their multispiral protoconch. Orthochetus is characterized by a paucispiral protoconch. The type species Orthochetus leufroyi (Michelin, 1825) from the Eocene of France and the morphologically most similar species Orthochetus pagoda (Chapman & Crespin, 1934) from the Eocene of Australia are furthermore distinguished by their aperture with its narrow and relatively long anterior canal and prominent columellar plate. The aperture may actually be more important as a diagnostic character from Cerithiella than the prominent keel as a few recent species like Cerithiella amblytera (Watson, 1880) also developes a keel. The Maastrichtian C. tectiforme Binkhorst, 1861 from Holland, the early Palaeocene C. sp. 1 sensu Kollmann & Peel (1983) from Greenland and the Palaeocene Cerithium zigzag Grönwall & Harder, 1907 from Denmark were reassigned to Orthochetus in spite of a lack of known protoconchs. They are all more likely belonging to Cerithiella sensu lato due to their stratigraphical and morphological closeness with the present species, but a definite generic assignment must wait until their protoconchs have been uncovered. Undoubted species of Orthochetus are restricted to the Eocene to perhaps the Oligocene of Europe and Australia (see Darragh 2011).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A977FF922B9CC3F8FBF8FA5E.taxon	description	Figs 28 P – Q 2014 Cerithiella sp. Hansen & Surlyk: 339, fig. 6: 4, tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A977FF922B9CC3F8FBF8FA5E.taxon	materials_examined	Material. MGUH 33216, MGUH 33217 and three fragmentary external moulds of teleoconchs with the informal sample numbers SR. 739, SR. 777 and SR. 912. B – C. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig and Holtug Quarry, Stevns Klint. Description. Teleoconch slender with a slightly stepped outline due to deeply impressed and broad suture. Whorls strongly flattened, rounded subrectangular in outline with a height corresponding to 2 / 5 the width; peripheral keel located just above abapical suture succeeded by weak abapical keel covered by adapical wall of succeeding whorl. Base weakly convex. Columella strong and relatively short with no or only very weakly developed columellar fold. Aperture not preserved. Teleoconch sculpture composed of strong reticulate pattern of up to around 30 sharp and slightly prosocyrt transverse ribs crossed by four equally strong spiral ribs adapically to base. Strongest spiral rib located on keel. Transverse ribs terminating at sutures. Small tubercles present at connecting points between ribs. Base with one primary spiral rib located on abapical keel and succeeded by increasingly weaker spiral ribs axially. Growth lines fine, their course corresponding to that of the transverse ribs. Measurements. Largest specimen, MGUH 33216, consisting of 5 ½ teleoconch whorls is 13 mm high and 6.7 mm wide. Remarks. The taxon strongly resembles C. moltkianum (Ravn, 1902), but differs by the staircase outline and the more pronounced reticulation with four instead of three spiral ribs on each whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A977FF922B9CC71AFC59F957.taxon	materials_examined	Type species. Eumetula dilecta Thiele, 1912 (type by monotypy). Recent, Antarctic.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A977FF9D2B9CC64BFEB4FBF1.taxon	description	Fig. 28 R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A977FF9D2B9CC64BFEB4FBF1.taxon	materials_examined	Material. A single mould, MGUH 33218. Occurrence. The taxon is known exclusively from the lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Protoconch 0.4 mm high and 0.35 mm wide, conical multispiral with approximately 3 ½ convex whorls. First 1 ½ (embryonic) whorls seemingly smooth, subsequent whorls covered by up to around 35 closely spaced and very fine opisthocyrt collabral threads per whorl. A keel or coarse spiral rib appears on first post-embryonic whorl, while a second is added on last protoconch whorl. Teleoconch slender turriculate, with evenly increasing whorl width; whorls weakly convex, constricted at narrow adapical suture bordered by spiral ribs; whorl height corresponding to half or slightly more than half the width. Transition to smooth flat base sharp, marked by subperipheral carination. Base flattened, smooth. Aperture unknown. Teleoconch sculpture dominated by coarse and weakly opisthocline transverse ribs, strongest centrally on whorl, interrupted adapically and at carination abapically, giving an impression of a more convex whorl; rib number increasing gradually with growth from around 14 on first teleoconch whorl. A more or less distinctly developed spiral rib and succeeding furrow occur adjacent to adapical suture. Interspaces between transverse ribs characterized by an additional five secondary and very weak spiral ribs. Measurements. Specimen MGUH 33218 is 3.6 mm high and 0.9 mm wide, consisting of 3 ½ protoconch and eight teleoconch whorls. Remarks. Eumetula sp. may be the oldest taxon assigned this genus so far, moving the generic range back to the Maastrichtian (see Gründel 1980). Eumetula sp. somewhat resembles the Belgian Cerithium regularicostatum Briart & Cornet, 1873 from the Danian, which was reassigned to Harrisianella Olsson, 1929 by Glibert (1973). Both taxa clearly do not belong to this genus as revised by Kowalke (2001: p. 260 – 261) and differ by much more convex whorls; the lack of an adapical subsutural spiral furrow and, at least with regard to the Danish taxon, by the densely sculptured protoconch whorls. The Danish taxon differs from Eumetula regularicostatum by the rather weakly convex teleoconch whorls and a more pronounced spiral rib demarcating the base. It is clearly distinguished from the Selandian Harrisianella subglabra Schnetler, 2001 from Denmark by the fewer and strongly sculptured protoconch whorls and by the more compact teleoconch whorls with their abapical spiral rib. The Selandian species does not belong within Harrisianella for the same reasons as for the species above and due to the high, multispiral protoconch.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A978FF9D2B9CC529FB3CFB23.taxon	materials_examined	Type species. Triforis bitubulatus Baudon, 1856 (type by monotypy). Eocene, Paris Basin in France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A978FF9C2B9CC45EFC6FFE68.taxon	description	Fig. 28 S 1898 Cerithium oblique-costulatum nov. sp. Kaunhowen: 62, pl. 6, fig. 4. 1933 Cerithiopsis obliquecostulata n. sp. — Ravn: 52 – 53, pl. 4, figs 12 a – b, 13 a – b.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A978FF9C2B9CC45EFC6FFE68.taxon	materials_examined	Diagnosis. Protoconch with six to seven convex whorls carrying in excess of 20 fine collabral ribs per whorl. Teleoconch with flattened whorls sculptured by strong and slightly opisthocline transverse ribs and around three very weak, but wide spiral ribs. Material. The K-Pg boundary strata is represented by specimen MGUH 33219 and six somewhat fragmentary moulds of teleoconch from the uncatalogued old collections at the Natural History Museum of Denmark. Occurrence. The material here treated derives from the lithified top of the Cretaceous Tor Formation at Rødvig, Stevns. This species is additionally known from the lower Danian (?) bryozoan limestone at Limburg in Holland and from the middle Danian limestone at Faxe, Denmark (Kaunhowen 1898; Ravn 1933). Description of boundary material. Protoconch not preserved. Teleoconch moderately slender with flat to slightly concave and weakly keeled whorls. Keel located approximately 1 / 3 the whorl height from abapical suture. Suture not deep, marked by narrow sutural ramp. Transition from whorl side to weakly convex base abrupt. Whorl subrectangular in cross-section with smooth columella lacking columellar fold. Aperture not known. Teleoconch whorls carrying between 16 and 20 coarse and sharp-ridged opisthocline and slightly opisthocyrt transverse ribs, becoming less pronounced and more rounded on later whorls. Transverse ribs crossed by three weakly developed spiral ribs; one forming adapical shoulder on whorl, followed by second rib located approximately 1 / 3 of whorl height from adapical suture. The weak keel carries the third and strongest spiral rib. Measurements. MGUH 33219 is 2.3 mm high and 1.3 mm wide with six teleoconch whorls. Remarks. This taxon from the uppermost Maastrichtian chalk at Stevns Klint is somewhat fragmentary, but the teleoconch appears completely in agreement with the middle Danian Cerithiopsis obliquecostulata Ravn, 1933 from Denmark, which is clearly a junior subjective synonym of the only slightly older Cerithium obliquecostulatum Kaunhowen, 1898 from the lower Danian limestone deposits in Holland.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A979FF9C2B9CC090FA83FDDA.taxon	materials_examined	Type species. Cerithium bimoniliferum Sandberger, 1858 (type by original designation). Late Oligocene, Germany.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A979FF9C2B9CC3C1FB98F9AE.taxon	description	Figs 28 T – V	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A979FF9C2B9CC3C1FB98F9AE.taxon	materials_examined	Diagnosis. Fairly broad-spired with strongly flattened whorls; periphery located close to abapical suture; transition to base sharp. Protoconch with a single spiral thread. Derivation of name. In honour of C. Heinberg, who found the holotype and has dedicated much of his scientific effort into the study of the fauna and environments at the Danish Cretaceous-Palaeogene boundary. Type material. The holotype MGUH 33220 is a nearly complete external mould with protoconch. Paratypes MGUH 33221 and MGUH 33222 are from the Cerithium Limestone Member at Skeldervig, Stevns Klint. Type stratum and type locality. The lower Danian Cerithium Limestone Member at Hulerne (The Caves) near Storedal at Stevns Klint. Occurrence. The lower Danian Cerithium Limestone at Stevns Klint. Description. Protoconch planktotrophic turriculate, 0.85 mm high and 0.5 mm wide, consisting of around 5 ½ moderately convex whorls. Embryonic whorl slightly inflated, smooth. Second whorl too worn for a clear view of any potential sculpture. Succeeding whorls carrying 13 – 14 slightly opisthocline and opisthocyrt collabral threads per half whorl. The collabral threads may just cross single spiral thread appearing at periphery near abapical suture on final two whorls. Teleoconch with flattened whorls with sharp transition to flat base. Aperture only partly preserved, appearing to have a straight columella. Teleoconch sculpture reticulate with around 16 opisthocline transverse ribs per whorl on first teleoconch whorl. Transverse ribs crossed by three or four spiral ribs, the strongest located on keel slightly above abapical suture and succeeded by weaker abapical rib. Connecting points marked by tubercles. Base with strong spiral rib close to abaxial margin; otherwise smooth. Measurements. The holotype MGUH 33220 is 3.3 mm high and 1.2 mm wide, consisting of five teleoconch whorls in addition to protoconch. Remarks. The taxon appears to resemble the Bohemian Cerithium ternatum Reuss, 1845, but differs at least by the better developed spiral and transverse ribs in contrast to the tubercle sculpture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A979FF9F2B9CC7EDFB0EFE4C.taxon	description	Fig. 28 W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A979FF9F2B9CC7EDFB0EFE4C.taxon	materials_examined	Material. Specimen MGUH 33223 is an external mould of the teleoconch. Occurrence. Lithified top of the Cretaceous Højerup Member at Stevns Klint. Description. Shell moderately slender with strongly flattened and low whorls separated by narrow and indistinct suture located on periphery on preceding whorl. Whorl height corresponding to less than half the width. Peripheral transition to flat base sharp. Teleoconch sculpture weakly reticulate with five spiral ribs crossed by between 30 and 34 prosocline and slightly prosocyrt transverse ribs with low tubercles at connecting nodes. Adapical two spiral ribs stronger and more widely spaced than succeeding three. An additional spiral rib accentuating transition to base visible on last whorl. Base carrying at least seven spiral ribs decreasing in strength adaxially. Measurements. MGUH 33223 is a teleoconch lacking the apex and measuring 5.9 mm in height and 2.5 mm in width. It consists of just over six whorls. Remarks. It closely resembles Vatopsis heinbergi n. sp. described above from the Cerithium Limestone Member, but differs by the distinct spiral sculpture on the base and by having five spiral ribs instead of four. Further material is needed before the taxonomic status of this taxon may be satisfactorily unravelled.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97AFF9F2B9CC3DCFAA9FD0E.taxon	materials_examined	Type species. Ageria gaultina Abbass, 1973 (type by original designation). Early Cretaceous, Kent in England.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97AFF9E2B9CC20DFD86FEA0.taxon	description	Figs 28 X – Y, 29 A – B	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97AFF9E2B9CC20DFD86FEA0.taxon	materials_examined	Diagnosis. Early teleoconch whorls strongly and fairly densely sculptured by sharp transverse ribs mostly lacking tubercles; last whorl widening with base contour shifting from flat to strongly convex. Derivation of name. Refers to the type locality Skeldervig at Stevns Klint. Type material. The holotype MGUH 33226 is a nearly complete external mould without protoconch. Paratype MGUH 33224 is from the Højerup Member at Rødvig, while paratype MGUH 33225 comes from the Højerup Member north of Kulsti Rende, Stevns Klint. Additional material. A mould in the informally catalogued sample SO. 5. A, and a fragmentary external and internal mould from the uncatalogued old collections at the Natural History Museum of Denmark. Type stratum and type locality. The lithified top of the Maastrichtian Højerup Member at Skeldervig, Stevns Klint. Occurrence. This species is known exclusively from the lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, Skeldervig and north of Kulsti Rende, Stevns Klint. Description. Protoconch unknown. Teleoconch slender turriculate, with evenly increasing whorl width; whorls weakly convex, separated by narrow suture; whorl height corresponding to half or slightly more than half the width. Transition to smooth flat base sharp, marked by weak carina. A second weaker carina may appear on base of adult specimens. Base on early whorls flattened, becoming more convex with growth. Columella smooth. Aperture unknown. Teleoconch sculpture dominated by coarse and weakly opisthocline transverse ribs; rib number increasing gradually with growth from around 14 on initial teleoconch whorls. Rare large and low, weakly developed varices occur, becoming slightly more common on later whorls. A more or less distinctly developed spiral rib and succeeding furrow occur adjacent to adapical suture. Interspaces between transverse ribs characterized by an additional five secondary and very weak spiral ribs. Later whorls smooth but for weak spiral lirae crossed by fine, opisthocyrt and slightly opisthocline growth lines. Base with slightly stronger spiral lirae. Measurements. Holotype MGUH 33226 is 4.5 mm wide and 14.4 mm high, consisting of 14 ½ teleoconch whorls. Remarks. The Danish late Maastrichtian Ageria skeldervigensis n. sp. resembles the Maastrichtian A. weeksi Wade, 1926 from the Ripley Formation of southern United States, but is distinguished by the higher number of tubercle rows on the later whorls and the apparently two and not three smooth protoconch whorls. It differs from the Maastrichtian Ageria gankinensis Kaim et al., 2004 from northern Russia by the more convex base and a finer teleoconch sculpture with weaker tubercles. It is furthermore distinguished from the two British late Albian species A. gaultina Abbass, 1973 and A. costata (Sowerby, 1827) by the distinctly less pronounced spiral ribs and growth line sculpture and by a slightly more convex base. Ageria skeldervigensis n. sp. may resemble Ageria laxa n. sp. described below from the succeeding Cerithium Limestone Member, but differs by the nearly effaced tubercles on each transverse rib; a relatively early disappearance of the teleoconch sculpture and by a generally slightly higher number of transverse ribs per whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97BFF9E2B9CC0D8FBF0F874.taxon	description	Figs 29 C – E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97BFF9E2B9CC0D8FBF0F874.taxon	materials_examined	Diagnosis. An Ageria with approximately two smooth protoconch whorls. Teleoconch strongly and fairly densely sculptured; last whorl widening with base contour shifting from flat to strongly convex. Derivation of name. Refers to the strong widening of the last whorl seen on large specimens. Type material. The holotype MGUH 33227 is a nearly complete external mould with protoconch. Paratype MGUH 33228 is from the Cerithium Limestone Member at Stevns Klint. Additional material. ØSM. 10042 - 133 and ØSM. 10042 - 388 - a comes from the Crab Layer in the Korsnaeb Member. A mould in the informally catalogued sample SR. 296. B – C and an additional three external and internal moulds without numbers are from the Cerithium Limestone Member and belong to the Natural History Museum of Denmark. Type stratum and type locality. The Crab Layer, a hardground related lithified bed approximately two metres up in the Korsnaeb Member of the lower Danian Stevns Klint Formation at Stevns Klint. Occurrence. The lower Danian Cerithium Limestone Member of the Rødvig Formation and up into the Crab Layer, located about two metres up in the Korsnaeb Member of the Stevns Klint Formation at Stevns. Stevns Klint: Rødvig, Korsnaeb, Skeldervig, Højerup and Holtug Quarry. Description. Protoconch 0.3 mm high and 0.3 mm wide, consisting of around two smooth, slightly convex whorls forming a continuation of teleoconch spire. Teleoconch very slender coeloconoid; initial whorls slightly convex, becoming gradually more flattened abapically. Whorls moderately low, height on teleoconch whorl 16 corresponding to approximately 45 % the width. Suture narrow, distinct, but not deep. Keel sharp, located at transition to base at abapical suture line. Base flat with short anterior canal. Last whorl on large specimens may increase dramatically in width, becoming more convex with the keel changing to a more central location on the whorl before disappearing just behind aperture. Aperture rounded subrectangular with smooth, slightly concave columella. Teleoconch strongly sculptured adapically, becoming smoother abapically with only weak spiral ribs on base of inflated final whorl. First whorl with five to six strong transverse ribs on a half whorl, crossed by two weaker spiral ribs, forming two weak tubercles on transverse ribs. A third, narrower adapical spiral rib is added on third whorl. Spiral ribs increase rapidly in number while becoming finer. Transverse ribs may increase slowly or faster in number abapically, while tubercles become more pronounced, arranged in up to four rows. Keel carrying narrow spiral rib, which may be slightly serrated due to densely spaced growth lines. Base with single primary spiral rib slightly adaxially of keel rib and succeeded by closely spaced, weaker secondary spirals. Growth lines closely and evenly spaced, nearly straight adapically of keel, weakly opisthocyrt. Measurements. The holotype MGUH 33227 is 16.2 mm high and 4.8 mm wide with 19 whorls. Largest examined specimen is 15.4 mm wide. Remarks. The Danish early Danian Ageria laxa n. sp. resembles the Maastrichtian A. weeksi Wade, 1926 from the Ripley Formation of southern United States, but is distinguished by the higher number of tubercle rows on the later whorls and the apparently two and not three smooth protoconch whorls. It differs from the Maastrichtian Ageria gankinensis Kaim et al., 2004 from northern Russia by having a more convex base, a stronger developed anterior canal and apparently by having only two protoconch whorls. It is furthermore distinguished from the two British late Albian species A. gaultina Abbass, 1973 and A. costata (Sowerby, 1827) by the distinctly less pronounced spiral ribs and growth line sculpture and by a slightly more convex base. The differences between Ageria laxa n. sp. and Ageria skeldervigensis n. sp. are presented above in the remarks for the latter species.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97DFF982B9CC3C2FE52FA9A.taxon	description	Figs 29 F – G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97DFF982B9CC3C2FE52FA9A.taxon	materials_examined	Material. A moderately well preserved external and internal mould (MGUH 33229), which may be mislaid, and a poorly preserved external mould MGUH 33230. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Rødvig and Holtug Quarry, Stevns Klint. Description. Protoconch not known. Teleoconch whorls flattened, relatively high, the height corresponding to about 60 % of width, separated by shallow suture. Keel on last whorl very sharp, delineated adapically by wide furrow. Base strongly flattened. Aperture subrectangular, slightly flaring with two low spiral folds on outer lip. Teleoconch sculpture changing with growth; initial whorls carrying about 14 coarse, slightly opisthocline transverse ribs fading out towards sutures; ribs terminating abapically towards weak spiral rib delineating beaded keel. Transverse ribs fading out on later whorls, being replaced by three beaded spiral ribs; adapical keel ramp carrying three weak spiral threads. Growth lines opisthocyrt and slightly opisthocline. Measurements. Teleoconch on disappeared specimen at least 11.2 mm high and 3.7 mm wide, consisting of nine whorls. Remarks. Ageria sp. resembles Ageria gankinensis Kaim et al., 2004 from the Late Maastrichtian of Siberia, but differs by the denser sculpture with three instead of two primary beaded spiral ribs adapical to the keel. It seems to differ from Ageria skeldervigensis n. sp. described above from the same deposits by the relatively high whorls and extremely sharp keel, but a very limited material prevents any definite conclusions as the differences could reflect ecophenotypic variation.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97DFF982B9CC486FD2FF91C.taxon	materials_examined	Type species. Scalaria angarensis Ryckholt, 1854 (type by original designation). Paleocene, Belgium. Remarks. The sharp costae used as a diagnostic feature by Wenz (1940: p. 802) are not always present as illustrated by the Middle Danian C. faxensis Ravn, 1933 and the Selandian C. johnstrupi (Mørch, 1874), both from Denmark. The former has broad, rounded transverse ribs, while the latter features both rounded ribs and more costae-like ribs. The species are in all other regards typical members of this genus and the presence or absence of costae should thus only be used as a general diagnostic feature.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97DFF9B2B9CC61CFC61FA20.taxon	description	Figs 29 H – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97DFF9B2B9CC61CFC61FA20.taxon	materials_examined	Diagnosis. Blade-like costae not strongly shouldered, closely spaced with up to around 22 per whorl; costae only slightly flexed at transition to basal carina. Umbilicus open, relatively wide. Derivation of name. Refers to the relatively wide umbilicus characterizing this species. Type material. The holotype MGUH 33232 was collected by A. Rosenkrantz 21 July 1943. Paratype MGUH 33233 is from the upper part of the Tor Formation exposed just north of Holtug Quarry, and was collected by Claus Heinberg in 1995. Paratype MGUH 33231 is from the Cerithium Limestone Member at Holtug Quarry, and was collected by A. Rosenkrantz in 1940. Additional material. A specimen with informal sample number SR. 781, and an uncatalogued specimen, both with part of shell preserved, derives from the lithified top of the Tor Formation; ØSM. 10042 - 90 - a – b, two specimens with informal sample numbers DN. 21 and DN. 49, and four specimens without number comes from the ‘ dead layer’ in Dania Quarry, Northern Jutland, and from the Cerithium Limestone Member at Rødvig, Holtug Quarry and Højerup at Stevns Klint. ØSM. 10042 - 236 - a – b and five additional specimens with no numbers derive from the Korsnaeb Member at Stevns Klint and Dania. Type stratum and type locality. Basal Korsnaeb Member, or more precisely from the Korsnaeb Member infilled Thalassinoides burrows in the Cerithium Limestone Member at Harvig, Stevns Klint. Occurrence. Lithified top of the Maastrichtian Tor Formation at Holtug Quarry, Stevns Klint; the lower Danian Cerithium Limestone Member and contemporaneous ‘ dead layer’ from Stevns Klint and Dania Quarry, respectively, and their Korsnaeb Member outfilled Thalassinoides burrows. Description. Shell spire high turriculate with well rounded whorls only just meeting neighbouring whorls. Sutures deep. Base of whorl with flattened disc or carina surrounding deep and moderately wide umbilicus, umbilicus taking up approximately 15 % of whorl width. Aperture broadly ovoid with slightly developed varix; height corresponding to around 75 % of width. Shell surface covered by very fine and dense spiral lirae and between 15 and 22 blade-like costae per whorl; costae composed of multiple lamellae, flexed subsuturally and, to a lesser degree, at transition to basal carina and to umbilicus; costae distinct on carinal disc. Protoconch not known. Measurements. Holotype specimen, MGUH 33232, more than 35.4 mm high and 19.6 mm wide. Remarks. Coniscala umbilica n. sp. is easily distinguished from most other species of Coniscala by its high lamellar ribs, the open and relatively wide umbilicus and the distinct costae on the carinal disc. It is probably most similar to the Palaeocene Cavoscala crispn Gorbach, 1972 from the Crimea, Russia, from which it differs by the coarser and sparser transverse ribbing. More material of the Crimean taxon may perhaps show them to represent different phenotypes of the same species. Coniscala umbilica n. sp. resembles the slightly younger C. faxensis Ravn, 1933 from the middle Danian of Denmark, and C. johnstrupi (Mørch, 1874) from the Selandian of Denmark, but differs by the high and blade-like costae, slightly weaker spiral lirae, the generally wider umbilicus and possibly by a less pronounced basal disc. It strongly resembles the Late Maastrichtian Scalaria contorta Kaunhowen, 1898 from Holland, which was later assigned to Confusiscala Boury, 1909 by Abdel-Gawad (1986), but according to P. v. Knippenberg from Holland (pers. comm. 2009) it lacks the wide umbilicus of the Danish taxon. Coniscala umbilica n. sp. resembles species of the contemporaneous North American genus Striaticostatum Sohl, 1963, and here especially S. bexarense (Stephenson, 1941), but differs by the wider umbilicus, the less honeycombed and more blade-like costae and by the distinctly less backwardly turned costae on the basal carina (compare with Sohl 1964 a). Striaticostatum should probably be regarded as a subgenus of Coniscala.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97EFF9B2B9CC759FE82F973.taxon	materials_examined	Type species. Scalaria australis Lamarck, 1822 (subsequent designation by Boury 1886). Recent, New South Wales to Western Australia.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97EFF9A2B9CC7ACFACBFDDC.taxon	description	Figs 29 L – N	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97EFF9A2B9CC7ACFACBFDDC.taxon	materials_examined	Material. MGUH 33234 and MGUH 33235, both external and internal moulds. Occurrence. The lower Danian Cerithium Limestone Member at Stevns Klint. Description. Protoconch high conical, approximately 0.4 mm high and 0.3 mm wide, consisting of around 2 ½ moderately convex whorls. Embryonic whorl small, smooth or granulate. Larval whorls 1 ½ to two in number, carrying around 30 opisthocline collabral threads per whorl. Teleoconch moderately slender with highly convex whorls separated by very deep suture. Whorl width nearly twice as large as height. Transition to flattened base marked by basal carina. Base smooth but for weak and wide spiral furrows. No umbilicus present. Aperture rounded ovoid with weakly concave columella. Teleoconch sculpture dominated by around 14 sharp-ridged transverse ribs per whorl; ribs crossed by five or six weaker spiral ribs or threads, the abapical four much stronger than adapical ones. Measurements. Specimen MGUH 33234 is 3.6 mm high and 1.25 mm wide, consisting of protoconch and 5 ¾ teleoconch whorls. Remarks. Opalia sp. differs from the early Palaeocene Cerithiscala sp. of Kollmann & Peel (1983) from Greenland by the higher number of spiral threads. Likewise it is easily distinguished from the middle Danian Opalia tricincta (Ravn, 1933) from Denmark, by the finer sculpture, the more slender outline and the more convex whorls. It differs from the Selandian Opalia poulseni (Ravn, 1939) from Denmark by the finer sculpture and broader outline, and from the Selandian Opalia hauniensis (Ravn, 1939), also from Denmark, by the stronger spiral sculpture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97FFF9A2B9CC3DCFE08FCEF.taxon	materials_examined	Type species. Scalaria richardi Dautzenberg & Boury, 1897 (type by original designation). Recent, southwest European part of the North Atlantic.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97FFF9A2B9CC213FACDF837.taxon	description	Figs 29 O – Q	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A97FFF9A2B9CC213FACDF837.taxon	materials_examined	Diagnosis. Shell small; conical protoconch with around 35 opisthocyrt and strong collabral threads per whorl; teleoconch whorls rather weakly convex with weakly opisthocline transverse ribs and no spiral ribs or threads. Derivation of name. Refers to the small size compared to the typical species of Claviscala. Type material. The external mould MGUH 33236 consisting of the entire protoconch and first several teleoconch whorls is selected as holotype. Paratype MGUH 33237 is an external mould from the lithified top of the Højerup Member at Rødvig, Stevns Klint. Additional material. ØSM. 10042 - 354, six specimens with the informal sample numbers SR. 808, SR. 906, SR. 996. A – B, SR. 997. B – C, SR. 1048 and SR. 1065. B – C, and likely also SR. 982, SR. 1050 and SR. 1051. A – B. Type stratum and type locality. Lithified top of the Højerup Member of the Tor Formation exposed just south of Højerup Church, Stevns Klint. Occurrence. This species is known exclusively from the lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, just south of Højerup Church, and north of Kulsti Rende at Stevns Klint. Description. Protoconch conical multispiral with four convex whorls, holotype 0.65 mm high and 0.4 mm wide. First protoconch whorl seemingly smooth, the following covered by up to around 35 closely spaced opisthocyrt collabral threads per whorl. Transition to teleoconch seemingly sharp, seen as a change in sculpture and a decrease in whorl expansion. Teleoconch spire slender, conical with evenly increasing whorl width; initial whorl moderately convex, succeeding ones flattened, constricted just above narrow and relatively shallow abapical suture; whorl height corresponding to approximately 80 % of width. Transition to smooth flat base abrupt, marked by subperipheral carination. Aperture not known. Whorls carrying coarse, sharp and weakly opisthocline transverse ribs, their number increasing very slowly with growth from around 14 on first teleoconch whorl. Measurements. The holotype MGUH 33236 is 2.9 mm high and at least 0.9 mm wide, consisting of four protoconch and five teleoconch whorls. Remarks. Claviscala minor n. sp. is unusually small for this genus, but otherwise its features correspond quite well with the generic diagnosis. This species is easily distinguished from the Early Cretaceous Claviscala clementina (Michelin, 1833) from France by the higher density of transverse ribs, which are also more transversely directed. It differs from the Late Cretaceous Claviscala sp. of Squires and Saul (2003 a) from California by a slightly higher whorl convexity, a sharper transition to the base of the last whorl and the opisthocline nature of the transverse ribs. It differs from the Cretaceous Claviscala darwishi Abbass, 1963 from Egypt by the much less convex whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A980FF652B9CC26CFAC7FC9E.taxon	materials_examined	Type species. Scalaria magnifica Sowerby, 1844 (subsequent designation by Boury 1909). Recent, Pacific coast of Japan.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A980FF652B9CC1D4FAEDFD2C.taxon	description	Fig. 29 R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A980FF652B9CC1D4FAEDFD2C.taxon	materials_examined	Material. A single external mould, MGUH 33238, lacking protoconch and aperture. Occurrence. The Cerithium Limestone at Vokslev Quarry in northern Jutland. Description. Teleoconch spire acutely conical, whorls weakly convex, relatively high, the height corresponding to 65 % of the width; whorls constricted just above abapical suture. Suture located just abapically of outer sharp margin of basal disc. Transition to smooth flat basal disc sharp, marked by subperipheral carination. Aperture not known. Teleoconch sculpture consisting of sharp crested, inverted sigmoid transverse ribs numbering around 16 on sixth whorl, while increasing to 30 on 11 th whorl; ribs starting slightly abapically of adapical suture, reaching all the way down to the basal disc. A shoulder or adapical tubercle appears on adapical end of ribs on later whorls. No spiral ribs present. Measurements. Preserved teleoconch 7.1 mm high and 1.4 mm wide, consisting of 11 teleoconch whorls. Remarks. The taxon differs from the Maastrichtian C. minor n. sp. described above by the wide spiral band without transverse ribs on the adapical part of the whorls and the more sigmoid form of the transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A980FF642B9CC29DFDE4FE15.taxon	description	Figs 29 S – V	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A980FF642B9CC29DFDE4FE15.taxon	materials_examined	Diagnosis. Shell slender with generally strongly convex whorls and flattened base. Teleoconch carrying five to six well developed spiral ribs on early whorls and around 35 transverse ribs per whorl. Transverse ribs typically only slightly weaker than spiral ribs, but may fade out on later whorls. Derivation of name. In honor of the late Alice Rasmussen from Faxe. Type material. The holotype MGUH 33239 is a nearly complete external and internal mould lacking protoconch. Paratype MGUH 33240 is an external mould of apex from the lithified top of the Højerup Member at Holtug. Additional material. Eight external and internal moulds without numbers are found in the old collections of the Natural History Museum of Denmark. Type stratum and type locality. The lithified top of the upper Maastrichtian Højerup Member at Holtug, Stevns Klint. Occurrence. Lithified top of the Maastrichtian Højerup Member, Tor Formation, at Rødvig, Mandehoved and Holtug Quarry, Stevns Klint. Description. Shell very high and slender turriculate. Protoconch high conical, consisting of about three whorls. Embryonic whorl appearing smooth and slightly inflated. Larval whorls with dense pattern of around 45 to 50 fine opisthocline and forward concave collabral threads per whorl. Teleoconch fairly thin-shelled; whorls strongly convex with impressed suture, width corresponding to between 2 and 2 ½ times the height. Transition to base moderately sharp, marked by slight angle, becoming more gradual on later whorls. Base flattened to weakly convex on early whorls, with moderately sharp transition to short but well developed canal; base becoming more convex on later whorls. Aperture subrectangular with short and concave columella terminating abapically in columellar fold. Columellar lip covered by thin callus. Siphonal canal deep. Early teleoconch whorls covered by reticulate pattern of fine and sharply defined spiral ribs crossed by equally strong transverse ribs. Spiral ribs numbering five to six above abapical suture and one below. Transverse ribs slightly opisthocyrt, numbering around 35 per whorl or slightly below, terminating at abapical spiral rib. Whorl sculpture on larger specimens consisting of seven to eight fine but strong spiral ribs. Base smooth, while canal carries weak and low spiral ribs. Growth lines on whorls weakly opisthocyrt and opisthocline. Measurements. Largest specimen consisting of four teleoconch whorls 9.05 mm high and 4.5 mm wide. Holotype specimen MGUH 33239 is more than 7.6 mm high (protoconch lacking) and 2.65 mm wide with 7 ¾ teleoconch whorls. Remarks. A Danish middle Danian specimen strongly resembling Amaea aliceae n. sp. is illustrated by Damholt & Rasmussen (2005: p. 40) as Amaea elegans (Ravn, 1902). It is unclear whether they could represent the same taxon as the middle Danian specimen lacks the protoconch. Even so it is clear that neither belongs to A. elegans as evidenced by the presence of no more than around six spiral ribs, which are rather strongly developed, and by the smooth and less strongly delineated base. Amaea aliceae n. sp. seem to differ from the Danish middle Danian Acrilla sp. 1 sensu Lauridsen & Schnetler (2014) by the fine collabral threads on the protoconch and much denser sculpture of transverse teleoconch ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A981FF642B9CC304FECAF901.taxon	description	Fig. 29 W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A981FF642B9CC304FECAF901.taxon	materials_examined	Diagnosis. Shell slender with strongly convex whorls. Protoconch consisting of four whorls. Teleoconch with flattened, smooth base. Spiral ribs four to six in number, slightly stronger than transverse ribs. Transverse ribs straight. Derivation of name. Refers to the Stevns Klint exposure at which it was found. Type material. The nearly complete external mould MGUH 33241 is selected as holotype. Additional material. In addition to the type specimen another three fragmentary external moulds with informal sample numbers SH. 386. A – B, SH. 394 and SR. 311. A have been examined. Type stratum and type locality. Cerithium Limestone Member just north of Højerup Church, Stevns Klint. Occurrence. Amaea stevniensis n. sp. has been collected from 25 to 30 centimetres above the base of the Rødvig Formation (base of the P 1 a Foraminiferal Subzone) in the Cerithium Limestone Member at Højerup Church, and from 30 to 40 centimetres above the base of the Rødvig Formation (Pα Foraminiferal Zone) at Rødvig, Stevns Klint. Description. Shell high turriform. Protoconch multispiral, approximately 0.6 mm high and 0.35 mm wide, consisting of four smooth whorls. Apex somewhat blunt with low convex whorls; convexity increasing gradually to the last and strongly convex protoconch whorl. Teleoconch whorls strongly convex, with a height corresponding to 2 / 3 the width. Transition to slightly thickened base on last whorl marked by distinct cord. Aperture rounded ovoid with slight angle at transition from outer lip to columella. Aperture height corresponding to ¼ the shell height. Outer lip with weak varix. Teleoconch sculpture cancellate, consisting of six sharp and moderately strong spiral ribs; the adapical weakest, sometimes completely effaced. Spiral ribs crossed by slightly weaker and less closely spaced, opisthocyrt to nearly straight, orthogonal or slightly prosocline transverse ribs. Number of transverse ribs per whorl 20 to 22 on first three whorls. Interconnecting points with weakly developed tubercles. Weak varices may occur. Base smooth. Measurements. Holotype MGUH 33241 2.3 mm high and 1.0 mm wide, consisting of eight whorls. Remarks. Amaea stevniensis n. sp. differs from the Selandian Acrilla fenestrata Ravn, 1939 by the smaller protoconch; the orthogonal transverse ribs and the more convex whorl outline. It differs from Amaea? sp. C treated below by the lower number of transverse ribs per whorl and the absent to weakly developed and rare varices. It is distinguished from the Danish mid-Danian Acrilla sp. 1 of Lauridsen & Schnetler (2014: p. 66) by the slightly more convex whorls and seemingly also by the transverse ribs not being stronger than the spirals. It is distinguished from Acrilla sp. 3 of Lauridsen & Schnetler (2014: p. 67) from the same place and horizon as the one before, by the higher number of spirals.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A981FF672B9CC678FD5FFD49.taxon	description	Figs 29 X, 30 A – B	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A981FF672B9CC678FD5FFD49.taxon	materials_examined	Material. MGUH 33242, a mould with informal sample number SR. 918. B – C, and an external mould lacking number. All belong to the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig and Holtug Quarry, Stevns Klint. Description. Protoconch turriform multispiral with five to 5 ½ whorls, width of whorls increasing steadily through the first four whorls after which only a slight increase may be observed. Protoconch whorls convex, very finely collabrally ribbed; ribs slightly opisthocyrt, between 34 and 40 per half whorl, fewer and coarser on last whorl. Teleoconch whorls weakly convex, width only increasing very slightly abapically; whorls separated by moderately deep suture. Whorl width corresponding to 1.5 times the height. Transition to base marked by weak angle. Base fairly narrow, slightly flattened, with no umbilicus. Aperture ovoid in shape. Whorl sculpture reticulate with narrow and sharp transverse and spiral threads and well developed tubercles at connecting nodes. Spiral threads five to six in number, while transverse threads numbers 9 to 10 on a half whorl. Measurements. Most complete specimen, MGUH 33242, containing 16 whorls, measures 7.7 mm in height and 1.1 mm in width; the protoconch measuring 1.1 mm in height and 0.6 mm in width. Remarks. Amaea sp. A appears to be characterized from other species of this genus by the extremely narrow spire. It differs additionally from Amaea stevniensis n. sp. described above by the conical outline of the protoconch and the opisthocline transverse ribs on the teleoconch.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A982FF662B9CC454FD95FF18.taxon	description	Fig. 30 E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A982FF662B9CC454FD95FF18.taxon	materials_examined	Material. Specimen ØSM. 10042 - 373 represents a fragmentary external mould of the teleoconch. Occurrence. The taxon belongs to the Maastrichtian Tor Formation and here probably from the basal Højerup Member or the uppermost Sigerslev Member, where it is cut by the early Danian lithified hardground layer in the northern part of Stevns Klint. Description. Protoconch unknown. Teleoconch very slender, turriculate, with strongly convex whorls separated by deep suture; whorls 1.6 times as wide as high. Distinct carina delimiting basal disc just visible above abapical suture. Varices narrow and frequent, tending to appear after every whorl. Teleoconch sculpture reticulate, consisting of more than 30 transverse ribs per whorl, crossed by slightly weak- er spiral threads. Transverse ribs sharp-ridged, opisthocyrt, bending forward adapically; ribs continuing down onto basal disc. Spiral threads close-spaced, numbering around 18 on spire whorls; adapical five to six very weak and closely spaced, succeeding ones stronger, alternating between coarser and finer threads. Connecting points between spirals and transverse ribs marked by weak tubercle. Basal disc covered by fine and closely spaced spiral threads on at least marginal part. Measurements. Specimen ØSM. 10042 - 373 consists of eight teleoconch whorls and measures 20.2 mm in height and more than 6 millimetres in width. Remarks. As pointed out by Squires and Saul (2003 b) the high frequency of varices is somewhat unusual for taxa belonging to Amaea. Allowing for the lack of the protoconch this taxon seems in all other respects to belong within this genus to which it is provisionally assigned. Amaea? sp. C has a strong resemblance to Amaea elegans (Ravn, 1902) from the middle Danian Coral Limestone at Faxe, Denmark. It is readily distinguished by the much finer and close-laying transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A982FF672B9CC3B0FBC1FB25.taxon	description	Figs 30 C – D	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A982FF672B9CC3B0FBC1FB25.taxon	materials_examined	Material. MGUH 33243 and MGUH 33244, both external moulds of the protoconch. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig and Højerup Church, Stevns Klint. Description. Protoconch low turriform, multispiral, with five whorls; width increasing steadily from apex to fifth whorl. Whorls convex and low, the height corresponding to approximately half the width. Sculpture consisting of fine, opisthocyrt and slightly opisthocline collabral threads numbering around 35 on fourth whorl. Collabral threads becoming slightly coarser on last whorl, on which the number drops to around 30. Measurements. Specimen MGUH 33243 just over 1.0 mm high and 0.6 mm wide. Remarks. The specimens resemble Amaea sp. A described above from the same horizon, but differ in the relatively more compact protoconch with its lower and comparatively wider whorls. Although the difference is quite marked further material may show them to represent two extremes of the same species.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A983FF662B9CC000FB54FDD7.taxon	materials_examined	Type species. Scalaria eschrichti Holböll in Møller (1842) (subsequent designation by Bouchet & Warén 1986). Recent, northwest Atlantic. Remarks. Bouchet & Warén (1986: p. 526) regarded Hemiacirsa Boury, 1890, Plesioacirsa Boury, 1909 and Pseudacirsa Kobelt, 1903 as junior subjective synonyms of Acirsa Mørch. This opinion is followed here.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A983FF662B9CC3CBFA35F973.taxon	description	Figs 30 F – G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A983FF662B9CC3CBFA35F973.taxon	materials_examined	Diagnosis. Protoconch conical multispiral with close to four whorls. Spire moderately broad with relatively convex whorls, smooth base and eight to nine coarse and slightly opisthocline transverse ribs per half whorl. Spirals few and very weak. Derivation of name. Refers to the upper Maastrichtian Tor Formation at Stevns Klint from the top of which it was collected. Type material. The nearly complete external mould MGUH 33245 is selected as holotype. Paratype MGUH 33246 is from the same locality and horizon. Type stratum and type locality. Lithified top of the Højerup Member, Tor Formation, at Rødvig, Stevns Klint. Occurrence. Lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Description. Shell relatively broad conical in outline. Protoconch multispiral, conical, consisting of 3 ½ to four smooth and moderately convex whorls. Protoconch height 0.5 mm and width 0.5 mm. Transition to teleoconch sharp, marked by the appearance of coarse transverse ribs. Teleoconch whorls weakly to moderately convex with periphery at or slightly above abapical suture; height corresponding to 2 / 5 the width. Transition to flattened base more or less sharp. Aperture rounded subrectangular with straight columella bearing narrow and smooth callus; aperture height taking up 30 % of shell height on holotype. Teleoconch sculpture dominated by fairly coarse, slightly sharp-ridged transverse ribs numbering eight to nine per half whorl on first teleoconch whorl; ribs slightly opisthocline, adapically terminating in weak spiral rib bordering adapical suture. Abapical termination at sharp transition to base. Spiral ribs on first teleoconch whorl crossed by four to five weak, but moderately coarse spiral ribs, the most abapical located on transition to smooth base. Measurements. Holotype MGUH 33245 1.3 mm high and 0.8 mm wide, consisting of the initial five whorls of the spire. Remarks. Acirsa torensis n. sp. differs from the Selandian Acirsa elatior (von Koenen, 1885) from Denmark by the less convex whorls with fewer and weaker transverse ribs. It is distinguished from the Late Cretaceous Acirsa clathrata Sohl, 1964 a from the U. S. A. by a relatively wider spire with a smooth base and stronger spiral sculpture.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A983FF602B9CC7AFFE0AFAA4.taxon	description	Fig. 30 H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A983FF602B9CC7AFFE0AFAA4.taxon	materials_examined	Material. Specimen MGUH 33247 is an external mould consisting of seven teleoconch whorls. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Mandehoved, Stevns Klint. Description. Protoconch not preserved. Teleoconch whorls very weakly convex, twice as wide as high and separated by deep linear suture located immediately abapically to whorl periphery. Transition to convex base rather gradual. Aperture unknown. Teleoconch sculpture consisting of around 13 sharp ridged, low, axially oriented transverse ribs; ribs tapering out at transition to base. Ribs crossed by fine, orthocline to slightly prosocline growth lines and up to around 15 weak and densely spaced spiral striae. Periphery with coarser spiral furrow, marking transition to coarser, scattered spiral striae on base. Measurements. MGUH 33247 is just over 2.0 mm wide with a preserved height of 6.2 mm. Remarks. Acirsa sp. differs from Acirsa torensis n. sp. described above by the apparently lower number of transverse ribs, which are less opisthocline in nature, the less convex whorls and the seemingly more slender spire. On the other hand these differences could merely reflect ontogenetic changes as Acirsa sp. is represented by a single teleoconch, while the material of A. torensis n. sp. is limited to two juvenile specimens. Acirsa sp. differs from the Early Cretaceous A. miyakoensis (Nagao, 1934) and A. ofunatoensis Kase, 1984 from Japan by the finer transverse ribs and less sharp transition to the base. It appears close to the Late Cretaceous Acirsa culmosa Sohl, 1964 b from the USA, but differs by the slightly less convex whorls with apparently a much less dense sculpture of spiral striae on the base and a complete termination of the transverse ribs immediately abapically of the periphery. It is characterized from the slightly younger Selandian Acirsa elatior (von Koenen, 1885) from Denmark by a more slender spire and less convex whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A985FF602B9CC499FC51F9C9.taxon	materials_examined	Type species. Scala elata Thiele, 1925 (type by monotypy). Recent, off South Africa.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A985FF632B9CC738FEB1FBD4.taxon	description	Figs 30 I – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A985FF632B9CC738FEB1FBD4.taxon	materials_examined	Diagnosis. Shell profile largely unchanged from protoconch to teleoconch; protoconch high-spired with approximately 30 collabral ribs; teleoconch with 15 to 17 relatively low and sharp transverse ribs and no spiral ribs. Derivation of name. In honour of the Carlsberg Foundation for its financial support of this work. Type material. Holotype MGUH 33248 is an external mould with well preserved details, while paratype MGUH 33249 from the same locality and horizon is a nearly complete external mould with slightly poorer preservation of details. Paratype MGUH 33250 is a nearly complete mould from the same strata and locality as the holotype. Additional material. In addition to the type material another five external moulds from the same locality and horizon were examined (Informal sample numbers SR. 747, SR. 982, SR. 1052, SR. 1122 (2 specimens )). Type stratum and type locality. Lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, Stevns Klint. Occurrence. Lithified top of the Maastrichtian Tor Formation at Rødvig, Stevns. Description. Protoconch consisting of four convex whorls, widening rather rapidly, the last measuring 0.3 millimetres in width. Initial whorl apparently smooth. Succeeding whorls carrying up to in excess of 30 opisthocline and slightly opisthocyrt collabral ribs reaching from suture to suture. Transition to teleoconch sharp, marked by decreasing whorl expansion and a shift from fine collabral ribs to coarse transverse ribs. Teleoconch spire high and narrow with low, convex whorls; whorl height corresponding to 2 / 3 the width. Transition to flattened base marked by clear angle; weakly developed subsutural collar may be present. Aperture ovoid with concave columella with callus. Teleoconch sculpture consisting of 15 to 17 strong and moderately sharp varix-like transverse ribs per whorl. Aperture unknown. Measurements. Paratype specimen MGUH 33250 is 3.0 mm high and at least 0.8 mm wide with two preserved protoconch and eight teleoconch whorls. Remarks. Opaliopsis carlsbergi n. sp. resembles the Campanian Opaliopsis faearium (Dockery, 1993), but differs in having a more high-spired protoconch with a width less than half that of the latter. It is clearly distinguished from the Maastrichtian O. aff. faearium sensu Nützel (1998) from the USA by the much coarser collabral ribs on the protoconch, which are also less opisthocyrt. Opaliopsis carlsbergi n. sp. differs furthermore from the Albian O. aff. dupinianum (d’Orbigny, 1842) sensu Kiel (2006) from Madagascar by the lack of spiral ribs and by a more even transition from protoconch to teleoconch with regard to shell outline. It also resembles the Albian Opaliopsis franciscae (Collignon, 1949), but differs in the absence of spiral ribs and the higher number of transverse ribs on the first teleoconch whorl, which only reach ten on O. franciscae (see Kiel 2006: p. 463). The slightly younger middle Danian limestone deposits of Denmark include the species Graphis danica Ravn, 1933 and Graphis sp. 1 sensu Lauridsen & Schnetler (2014: p. 71). They closely resemble Opaliopsis carlsbergi n. sp., but may be differentiated on strength of the protoconch, which is smooth and paucispiral, lacking the fine collabral ribs characterizing Opaliopsis.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A986FF632B9CC4E5FC52F9E6.taxon	materials_examined	Type species. Murex colus Linnaeus, 1758 (type by monotypy). Recent, Indo Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A986FF632B9CC5C5FD19FB09.taxon	materials_examined	Superfamily Buccinoidea Rafinesque, 1815	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A986FF622B9CC71AFDA2FBD4.taxon	description	Figs 30 L – M	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A986FF622B9CC71AFDA2FBD4.taxon	materials_examined	Diagnosis. Teleoconch moderately slender with eight strongly convex and rounded transverse ribs per whorl, the ribs forming weak rounded shoulder. Transverse ribs crossed by sharp spiral riblets with moderately broad interspaces. Derivation of name. In honour of K. I. Schnetler, who has contributed much to our knowledge of the Cainozoic gastropod faunas from Denmark. Type material. Holotype MGUH 33251 is an external and internal mould lacking protoconch. Paratype MGUH 33252 is a teleoconch from the dead layer in Dania Quarry, Northern Jutland. Additional material. Five external moulds from Stevns Klint are located in the collections of the National History Museum of Denmark. A mould catalogued by the informal sample number DN. 9. A – B comes from Dania Quarry, Northern Jutland. Another three specimens from the same locality and belonging to the private collection of K. I. Schnetler have been examined. Type stratum and type locality. Cerithium Limestone Member south of Stevns Fyr, Stevns Klint. Occurrence. Sporadically occurring in the Cerithium Limestone Member along the southern part of Stevns Klint (Skeldervig to slightly south of Stevns Fyr) and contemporaneous dead layer in Dania Quarry, Northern Jutland. Description. Protoconch unknown. Teleoconch moderately high-spired, consisting of weakly convex whorls; whorls low, their height corresponding to nearly half their width. Narrow suture located in distinct angle between succeeding whorls, or slightly below widest part of preceding whorl. Height of last whorl corresponding to around 60 % of shell. Aperture elongated ovoid with moderately gradual transition to narrow siphonal canal. Inner and outer lips and columella smooth, with no folds. Teleoconch sculpture dominated by around eight strong, rounded transverse ribs partly aligned from whorl to whorl. Ribs giving the whorls a more convex outline. Interstices wider than ribs. Spiral riblets strong and sharp; strongest and least crowded four ribs running at whorl periphery. Around four smaller ribs are located between these and adapical suture on some specimens, while others have fewer and stronger ribs. Transverse ribs fading out at transition to base, which is dominated by close lying spiral riblets, becoming gradually finer towards distal end of canal. Measurements. Most complete specimen, MGUH 33252, 13.6 mm high and 6.2 mm wide, consisting of initial six teleoconch whorls, but lacking tip of siphonal canal. Remarks. This taxon differs from the middle Danian Fusinus faxensis (Ravn, 1902) from Faxe Quarry, Denmark, by the slightly stronger and more closely spaced transverse ribs and the less crowded spiral riblets. It resembles the Early Palaeocene Fusinus sp. (sensu Kollmann & Peel 1983) from Greenland, but differs by a less tapering siphonal canal and a slightly more compact spire. It appears distinctly more common in the ‘ dead layer’ of Northern Jutland than in the Cerithium Limestone of Stevns Klint, suggesting a preference for a more deep-water and muddy environment than inhabited by most of the gastropods from the Danish area.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A987FF622B9CC5C4FBE2FB06.taxon	materials_examined	Type species. Fusus intortus Lamarck, 1803 b (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A987FF6D2B9CC47AFE5BFEFC.taxon	description	Figs 30 N – O	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A987FF6D2B9CC47AFE5BFEFC.taxon	materials_examined	Material. A nearly complete external mould, MGUH 33253, and two incomplete moulds from the uncatalogued old collections at the Natural History Museum of Denmark. The latter two are collected by A. Rosenkrantz at Kirkebraade (19.091927) and Skeldervig (August 1944). Occurrence. Very rare in the Cerithium Limestone Member, where it has been found at Skeldervig and Højerup, Stevns Klint. Description. Protoconch conical with between 1 ½ and 2 smooth, convex whorls separated by deep suture; protoconch measuring 0.55 mm in width and 0.35 mm in height. Teleoconch spire slender with convex whorls separated by moderately deep suture; whorl height less than half the width. Aperture narrowly ovoid with concave, seemingly smooth columella and oblique and wide siphonal canal; height of aperture and siphonal canal corresponding to 45 % of shell height. Siphonal canal short, only half as long as aperture. Fissure or weak pseudoumbilicus present. Siphonal canal with rounded distal end. Teleoconch sculpture dominated by around eight moderately strong, convex and more or less orthogonal transverse ribs per whorl. Transverse ribs crossed by six fairly broad and flat spiral cords between sutures; transverse ribs highest at periphery, fading out toward sutures; not organized in axial rows. Spiral cords strongest at whorl periphery. Growth lines slightly sigmoid. Measurements. MGUH 33253 is 9.2 mm high and approximately 3.6 mm wide, consisting of protoconch and six teleoconch whorls. Remarks. It is not possible to verify whether this taxon carries teeth on the outer lip or not. According to Wenz (1943: p. 1264) the genus should be characterized by a slightly oblique protoconch and a sharply cut distal end of the siphonal canal. Neither feature is observed in the present taxon, which otherwise is well in accord with the diagnostic features of this genus.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A988FF6D2B9CC5E1FBC5FAEB.taxon	materials_examined	Type species. Fusus subaffinis d’Orbigny, 1850 b (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A988FF6D2B9CC03DFCC5FBF1.taxon	description	Figs 30 P – Q	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A988FF6D2B9CC03DFCC5FBF1.taxon	materials_examined	Material. A single nearly complete external mould. MGUH 33254 consists of the protoconch and initial six teleoconch whorls collected south of Højerup, Stevns Klint. Occurrence. The Cerithium Limestone Member south of Højerup, Stevns Klint. Description. Protoconch obtusely conical, consisting of two smooth whorls with distinct sutural angle; initial whorl small, not inflated. Teleoconch high-spired, consisting of moderately convex whorls; whorls low, their height corresponding to nearly half their width. Narrow suture located in distinct angle between succeeding whorls, and bordered by weak subsutural collar. Aperture unknown. Teleoconch sculpture dominated by around 10 strong, rounded transverse ribs partly aligned from whorl to whorl. Ribs giving the whorls a more convex outline. Interstices more or less equal in width to ribs. Spiral riblets strong and sharp, numbering around six. Transverse ribs fading out at transition to base, which is dominated by close lying spiral riblets, becoming gradually finer towards canal. Measurements. Specimen, MGUH 33254 is more than 6.5 mm high and 2.6 mm wide, consisting of the protoconch and initial six teleoconch whorls. Exposed part of protoconch is 0.6 mm wide and 0.4 mm high. Remarks. Streptochetus? sp. B differs from Streptochetus sp. A described above from the same beds by the more opisthocline transverse ribs and the weak subsutural collar.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A988FF6C2B9CC41DFD31FF19.taxon	description	Figs 30 R – T	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A988FF6C2B9CC41DFD31FF19.taxon	materials_examined	Material. MGUH 33255, MGUH 33256, a mould in the informally catalogued sample SH. 287. A – B, and three fragmentary external and internal moulds without numbers are found in the collections of the Natural History Museum of Denmark. Of the latter three two have been collected by A. Rosenkrantz at Skeldervig (22 – 28 July 1943) and Rødvig (1924), while the third was collected by S, B. Andersen at Holtug (21 July 1970). Occurrence. A rare species occurring in the Cerithium Limestone Member from Stevns Klint, and in the contemporaneous ‘ dead layer’ at Vokslev, Northern Jutland. Description. Protoconch low conical with 2 ½ apparently smooth and moderately convex whorls. Teleoconch spire moderately slender with strongly convex whorls and a fairly shallow sinuous suture. Siphonal canal at least as high as aperture. Outer lip of aperture smooth. Columella carrying four weak columellar folds arranged two and two. Teleoconch sculpture dominated by strong and moderately narrow transverse ribs with no constriction at adapical suture. Ribs numbering 9 or 10 on early whorls, becoming fewer on later whorls. Transverse ribs crossed by very strong and sharp spiral cords, adapical three weaker than succeeding ones on whorl periphery. Peripheral spiral cord, generally number four from adapical suture, widest and most pronounced, succeeded by two slightly weaker cords abapically on spire whorls. Spiral cords coarse on base. Growth lines nearly orthocline, with weak prosocyrt sinus at adapical suture. Measurements. Protoconch measures 0.65 mm in width and 0.65 mm in height on MGUH 33255. Remarks. The assignment to Latirulus is very tentative due to the presence of four instead of three columellar folds, and as many as 2 ½ protoconch whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A989FF6C2B9CC001FA94FE4A.taxon	materials_examined	Type species. Turbinella bronni Michelotti, 1847 (subsequent designation by Cossmann 1901: p. 22). Miocene, Italy.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A989FF6C2B9CC0B6FA81F93E.taxon	description	Figs 30 U – W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A989FF6C2B9CC0B6FA81F93E.taxon	materials_examined	Diagnosis. Species of Dolicholatirus with a relatively short shell only three times as high as wide, and with a very fine spiral sculpture, sparse transverse ribbing and a smooth outer lip. Derivation of name. In honour of Mr. Lorenz, who found the holotype. Type material. Holotype MGUH 33257 is a nearly complete external and fragmentary internal mould collected by a Cand. [Candidate of Science?] Lorenz in 1881. Paratype MGUH 33258 is an external mould with preserved protoconch from the Cerithium Limestone Member at Skeldervig, Stevns Klint. Additional material. 38 external and internal moulds without numbers are found in the collections of the Natural History Museum of Denmark. Type stratum and type locality. The Cerithium Limestone Member of the Rødvig Formation at Stevns Klint. Occurrence. The Cerithium Limestone Member from the southern to central part of Stevns Klint (Skeldervig and up to Barmhjertigheden). Description. Protoconch obtusely conical, consisting of 2 ½ smooth whorls; exposed height approximating 0.8 mm and width measuring 0.7 mm. Teleoconch high-spired fusiform; whorls twice as wide as high, convex with slight adapical constriction. Sinuous suture relatively deep, emphasized by weak subsutural inflation on succeeding whorl. Last whorl taking up nearly 2 / 3 of shell height. Aperture and siphonal canal high and narrow, corresponding to half the shell. Outer lip smooth on inner side. Inner lip with two strong and sharp columellar folds. Teleoconch sculpture starting with 9 – 10 broad and strong, slightly prosocline transverse ribs, later decreasing in number to two or three per whorl; ribs largely organized into axial rows. Adapical end of transverse ribs nearly effaced, increasing in strength abapically to centrally between sutures, where a weak shoulder is formed. Ribs fading out on base, completely disappearing at transition to canal. Transverse ribs crossed by fine and numerous spiral cords, which may be completely effaced on older specimens, but usually increases in strength on base. Growth lines slightly sigmoid to nearly orthocline. Measurements. The holotype MGUH 33257 is 30.0 mm high and 9.6 mm wide with 8 or 9 teleoconch whorls. The largest shell fragment in the museum collections measures 12.6 mm in width. Remarks. Dolicholatirus lorenzi n. sp. differs from the late Cretaceous D. torquatus Sohl, 1964 a from the U. S. by the decreasing number of transverse ribs on later whorls and the much finer and weaker spiral sculpture. It is distinguished from the Danian D. striatulus (Briart & Cornet, 1871) from Belgium by the stronger columellar folds, lack of pseudoumbilicus and by the adapical effacement of the transverse ribs. It differs from the Danish middle Danian Dolicholatirus sp. 1 sensu Lauridsen & Schnetler (2014: p. 94) by fewer transverse ribs per whorl and finer spiral ribs. The external part of the shell may resemble that of Fusinus schnetleri n. sp. but for a stronger spire, the very slightly sigmoid growth lines and the generally finer and often nearly effaced spiral cords on the whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A989FF6F2B9CC6F2FDD5FF2C.taxon	materials_examined	Type species. Vexillum plicatum Röding, 1798 accepted as Voluta plicarium Linnaeus, 1758 (subsequent designation by Woodring 1928: p. 244). Recent, Indo-Pacific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98AFF6F2B9CC06CFA32F9C4.taxon	description	Figs 30 X, 31 A – C	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98AFF6F2B9CC06CFA32F9C4.taxon	materials_examined	Diagnosis. Protoconch obtusely conical with around two and a half moderately inflated whorls. Teleoconch small, slender with around 12 to 14 distinct transverse ribs per whorl on adapical five whorls, changing to fine but unevenly cancellate sculpture on later whorls. Derivation of name. Compound word for obtuse (stolidus) and young (tener), referring to the obtusely conical protoconch. Type material. Holotype MGUH 33259 is a nearly complete mould with protoconch collected by A. Rosenkrantz in 1943. Paratype MGUH 33260 is an external mould collected by K. I. Ingemann in 1986 from Dania Quarry, Northern Jutland. Additional material. Two moulds with informal sample numbers SH. 278 and SR. 376, and 33 uncatalogued and more or less fragmentary moulds are found in the collections of the Natural History Museum of Denmark. Type stratum and type locality. The lower Danian Cerithium Limestone Member at Skeldervig, Stevns Klint. Occurrence. Relatively common in the Cerithium Limestone Member at Stevns Klint and the corresponding ‘ dead layer’ at Dania, Northern Jutland. Description. Protoconch obtusely conical with around 2 ½ low, inflated whorls, possible smooth. Transition to teleoconch sharp, marked by distinct change in spire angle and the appearance of transverse ribs. Teleoconch slender, fusiform with high, flattened whorls, which are only around 1.5 times as wide as high. Whorls separated by marked suture. Whorls widest centrally or slightly adapically from here and characterized by distinct constriction slightly abapically of adapical suture. Last whorl takes up approximately 2 / 3 the total shell height, while aperture corresponds to around 40 %. Aperture narrow with weakly convex outer margin and sigmoid inner margin; columella carrying four strong folds, their distance and strength decreasing abapically from parietal region to wide siphonal canal. Teleoconch sculpture on first five whorls dominated by around 12 to 14 moderately sharp and strong transverse ribs per whorl, which due to adapical constriction appear strongest just over 1 / 3 the whorl height from adapical suture, forming a slight shoulder on the whorl. Transverse ribs cancellated by relatively weak and densely spaced spiral threads. Transverse ribs becoming finer and closely spaced on later whorls; in some specimens even replaced by distinct growth lines of a like strength with the spiral threads. Measurements. Holotype MGUH 33259 is 14 mm high and 4.8 mm wide, consisting of six teleoconch whorls. Protoconch measuring 0.7 mm in width and 0.5 mm down to teleoconch suture. Remarks. The Danish taxon corresponds well to the subgenus Vexillum Röding, 1798 except in the multispiral protoconch, which differs from the typical form by its obtusely conical shape. According to Stilwell (2003) the oldest members of the subgenus Vexillum appears around the Cretaceous-Palaeogene boundary, and the Danish taxon could therefore perhaps represent an early form or merely a close relative.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98AFF6E2B9CC734FF72FB25.taxon	description	Figs 31 D – G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98AFF6E2B9CC734FF72FB25.taxon	materials_examined	Diagnosis. Protoconch with 3 to 3 ¼ whorls. Teleoconch whorls with around seven strong and generally not sharply ridged costae, which form distinct adapical shoulder. Derivation of name. In honour of Alfred Rosenkrantz, who collected the material and worked with the Palaeocene gastropod faunas from Greenland and Denmark. Type material. Holotype MGUH 33261 is an inner and outer mould collected by A. Rosenkrantz 22 – 28 July 1943. Paratype MGUH 33262 is an external mould from Skeldervig, while paratype MGUH 33263 is an external mould showing the aperture and collected somewhere between Knøsen and Harvig. Both paratypes were collected by A. Rosenkrantz from the Cerithium Limestone Member. Additional material. A single external mould lacking number belongs to the Natural History Museum of Denmark. Type stratum and type locality. The Cerithium Limestone Member of the Rødvig Formation from Skeldervig at Stevns Klint. Occurrence. The Cerithium Limestone Member along the southern half of Stevns Klint. Description. Protoconch multispiral, high conical, consisting of 3 to 3 ¼ smooth whorls of relatively low convexity. Suture fairly deep. Protoconch around 0.8 mm high and 0.65 mm wide. Transition to teleoconch abrupt, marked by appearance of transverse ribs. Teleoconch high fusiform with flattened whorls separated by moderately deep suture marked by slight swelling of subsutural wall of succeeding whorl. Last whorl moderately convex with gradual transition from whorl side to base. Canal short but well developed, turrid-like. Aperture high elongated, corresponding to just over half the shell height; adapically narrowly pointed, while continuing out into short but well developed and wide siphonal canal. Outer lip moderately convex, seemingly lacking teeth or lirae internally. Inner lip smooth except for three centrally placed and distinct columellar folds increasing in strength adapically on slightly curved columella. Teleoconch sculpture consisting of weak striae subsuturally and on transition to canal, and around seven strong costae per whorl. Costae strongest peripherally on whorl, fading gradually out on adapical part of base and very abruptly at mid-whorl, completely disappearing shortly below adapical suture. This abrupt termination results in a marked pseudo-shoulder around mid-whorl and a flat to weakly concave shoulder slope on an otherwise flat whorl. Growth lines weakly sigmoid, nearly orthogonal on spire whorls. Measurements. Most complete specimen, paratype MGUH 33262, 6.8 mm high and 3.2 mm wide, consisting of 3 ¼ protoconch whorl and 4 teleoconch whorls. Remarks. This species differs from the typical members of the genus Vexillum Röding, 1798 by the sparse and exceptionally strong transverse rib sculpture forming a distinct shoulder before fading out close to the adapical suture. The typical costellariid sculpture, at least on early teleoconch whorls, is one of numerous and much finer transverse ribs. The generic assignment is for this reason rather tentative.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98BFF6E2B9CC455FAEFF83D.taxon	description	Fig. 31 H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98BFF6E2B9CC455FAEFF83D.taxon	materials_examined	Material. A single external mould, MGUH 33264. Occurrence. Lithified top of the Maastrichtian Højerup Member at Højerup, Stevns Klint. Description. Protoconch high conical, 1.1 mm high and 1.0 mm wide, consisting of close to four smooth, weakly convex whorls. Transition to teleoconch sharp. Teleoconch fusiform with moderately convex and high whorls separated by shallow suture. Whorl width corresponding to just over 1.5 times the height. Whorls with pronounced shoulder and adapical restriction, giving the spire a stepped outline. Whorl periphery located slightly abapically of mid-whorl. Last whorl corresponding to approximately 75 % of shell height, ending abapically in short and fairly broad canal. Aperture unknown. Teleoconch sculpture dominated by 9 – 10 coarse and rounded costae beginning just below adapical restriction and continuing down onto shell base, where they fade out at transition to canal. Costae crossed by spiral lirae numbering around eight on spire whorls, but covering the whole last whorl evenly. Lirae fading somewhat out on costae, but this may be due to wear. Growth lines very weakly sigmoid. Measurements. MGUH 33264 is 7.3 mm high and 3.5 mm wide, consisting of protoconch and 2 ½ teleoconch whorls. Remarks. The taxon looks very similar to the Danien Vexillum? rosenkrantzi n. sp. described above and more material may perhaps show them to be conspecific. Vexillum? sp. differs by having four protoconch whorls and a slightly higher number of transverse ribs. The adapical whorl collar may also be slightly more pronounced.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98DFF682B9CC252FAD4FC20.taxon	materials_examined	Genus Columbarium Martens, 1881 Type species. Pleurotoma (Columbarium) spinicincta Martens, 1881 (type by monotypy). Recent, Queensland in Australia.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98DFF6B2B9CC558FCC8FF34.taxon	description	Fig. 31 I 1880 Fusus heberti Briart & Cornet: 20, pl. 14, fig. 8 a – d.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98DFF6B2B9CC558FCC8FF34.taxon	description	1973 Columbarium heberti (Briart & Cornet, 1870) — Glibert: 64 – 65, pl. 7, fig. 19. For more synonymy see Darragh (1969: p. 72).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98DFF6B2B9CC558FCC8FF34.taxon	materials_examined	Diagnosis. Protoconch of 1 ½ small whorls, embryonic whorl deviated slightly from axis. Teleoconch keel with scales. Sculpture of strong and low transverse costae and three sharp and strong spiral cords, one on keel, the second halfway between keel and abapical suture, and the third hidden by succeeding whorl on spire. Danish material. Specimen MGUH 33265 is a fragmentary mould of the last two whorls. Type stratum and type locality. The lectotype (Briart & Cornet 1880: fig. 8 d) as suggested by Darragh (1969) comes from the Danian Calcaire de Mons, Hainaut, Belgium. Occurrence. In Denmark it has been found in the Cerithium Limestone Member at Stevns Klint. Outside Denmark it is known from the Maastrichtian at Maastricht in Holland and from the Danian (Montian) in Belgium and Poland (Darragh 1969). Description of Danish material. Whorls angular with flattened shoulder ramp and pronounced shoulder at periphery, succeeded abapically by two weaker carinas. Suture moderately deep, wavy, located immediately adapically of abapical carina. Aperture sub-ovoid with thick callus and outer lip, extending abapically into siphonal canal of unknown length. Columella straight. Teeth or columellar folds absent. Teleoconch sculpture consisting of perhaps 7 or 8 low and broad transverse costae fading out towards abapical carina and crossed by three lamella-like spiral cords on shoulder and carina. Growth lines fine and densely spaced, prosocline on shoulder ramp, becoming orthocline on whorl flanks. Measurements. No clear size can be given for the present fragmentary specimen, but the preserved part of the two whorls are 9.1 mm wide and 8.1 mm high. Remarks. The characteristic shape of the shell makes for a fairly confident assignment to this species even though the specimen is very fragmentary. It seems to only have occurred in the Danish area in the form of strays from the more shallow-water environments further to the south.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98EFF6B2B9CC5C4FAECFB06.taxon	materials_examined	Type species. Mitra fusoides Lea, 1833 (subsequent designation by Cossmann 1889: p. 186). Eocene, Alabama, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98EFF6A2B9CC475FD0AFDDC.taxon	description	Figs 31 K – N 1902 Fasciolaria glabra n. sp. Ravn: 231 – 232, pl. 2, figs 19 – 20. 1903 Fasciolaria glabra Ravn — Ravn: 381, 394 – 395.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98EFF6A2B9CC475FD0AFDDC.taxon	materials_examined	Diagnosis. Whorls flat and smooth with narrow adapical shoulder accentuated by spiral rib; whorl height corresponding to 40 % of width. Outer lip smooth or with weak, beaded lirae. Canal moderately long and narrow. Material. Syntype MGUH 116, MGUH 33267, MGUH 33268, ØSM. 10042 - 80, ØSM. 10042 - 81, 30 specimens with informal sample numbers SH. 178, SH. 202, SH. 325. C, SH. 393, SH. 407. A – B, SH. 510, SS. 179. A – B, SS. 189. A – B, SS. 203. A – B, SR. 304. A – B, SR. 313, SR. 373, SR. 397, SR. 403. A (2 specimens), SR. 465, SR. 481, SR. 569, SR. 621. A – B, SR. 644, SO. 93, SO. 111, DN. 3, DN. 31. A – B, DN. 32. A – B, DN. 50 and DN. 51. A – B, and an additional 78 external and internal moulds in the uncatalogued old collections of the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member at Stevns Klint. Occurrence. Conomitra glabra (Ravn, 1902) ranges from the base of the Cerithium Limestone Member of the Rødvig Formation and up into the base of the succeeding Korsnaeb Member of the Stevns Klint Formation at Stevns. Additionally it is found in the upper part of the ‘ dead layer’ at Vokslev and Dania in Northern Jutland. Description. Shell fusiform, width corresponding to between 40 and 45 % of height. Spire taking up one third of shell height. Protoconch rounded conical, 0.7 mm wide and 0.55 mm high, consisting of 2 ¼ smooth whorls. Transition to teleoconch indistinct. Teleoconch with up to around six weakly convex whorls with narrow but distinct shoulder accentuating adapical suture; whorls low, height corresponding to approximately 40 % of width. Aperture high and narrow, the height taking up 55 % of shell height, while the width corresponds to just over one quarter the aperture height. Outer lip smooth or with around 10 weakly developed lirae or rows of teeth. Columella with four strong and sharp columellar folds, the adapical one located on transition to parietal region. Canal well developed. Teleoconch smooth but for a weakly developed spiral rib on shoulder and fine, slightly prosocyrt growth lines. Measurements. Largest specimen at least 12.4 mm high. Remarks. Conomitra glabra resembles the approximately contemporaneous C. montense Glibert, 1973 from Belgium, but differs in the more pronounced shoulder, the less compact shell and less developed lirae on outer lip. It differs from the Danian Conomitra sp. 2 sensu Kollmann & Peel (1983) from Greenland through the higher whorls and seemingly more slender canal and from the middle Danian Conomitra sp. sensu Ravn (1933) from Denmark in the pronounced suture and less convex whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF6A2B9CC736FABAF934.taxon	materials_examined	Type species. Ancilla buccinoides Lamarck, 1803 a (type by original designation). Eocene, Paris Basin in France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF6A2B9CC3DCFAE3FD0E.taxon	materials_examined	Type species. Mitra groenlandica Beck in Møller, 1842 (subsequent designation by Fischer 1884). Recent, Arctic Sea.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF6A2B9CC272FF71FA7A.taxon	description	Fig. 31 O	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF6A2B9CC272FF71FA7A.taxon	materials_examined	Material. MGUH 33269, consisting of mould and replaced shell, is the only available specimen from the studied interval. Occurrence. The described specimen comes from the Cerithium Limestone Member at Holtug Quarry, Stevns Klint. Undoubted Volutomitra quinqueplicata Ravn is known from the middle Danian Coral Limestone of Faxe in Denmark (Fig. 31 P). Description. Shell very slender, turriculid with high flattened whorls, whorl height corresponding to approximately 70 % of the width. Suture narrow but distinct. Transition to weakly convex base very gradual. Teleoconch surface smooth except for nearly orthogonal, very weakly opisthocyrt growth lines. Measurements. MGUH 33269 consists of the last 2 ½ teleoconch whorls and measures 30.9 mm in height and at least 11.6 mm in width. Remarks. The very fragmentary material lacking apex and aperture makes the assignment somewhat tentative. The large and slender shell with its fairly weakly convex and smooth whorls covered by nearly orthogonal growth lines is on the other hand fairly distinctive for at least this genus among the early Palaeocene gastropods of Denmark.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF752B9CC667FE83FAEC.taxon	description	Figs 31 Q – R 1902 Ancilla Milthersii n. sp. Ravn: 234 – 235, pl. II, fig. 18.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A98FFF752B9CC667FE83FAEC.taxon	materials_examined	Diagnosis. Protoconch conical with just over two convex whorls. Teleoconch shell elongated with weakly ex- pressed sutures and no spire furrows. Columella almost vertical, bearing four to six narrow, oblique plaits. Parietal callus very thin except in front of adapical part of aperture, forming weak tongue adapically of aperture. Aperture taking up slightly over half the shell height; its outer lip lacking a labral denticle. Material. Holotype MGUH 33270 (cast), MGUH 33271, a mould with the informal sample number SH. 192. A – B, and two uncatalogued specimens deposited in the collections of the Natural History Museum, Copenhagen. Type stratum and type locality. The single specimen of Ravn (1902), MGUH 33270, and therefore the holotype, is from the Cerithium Limestone Member at Stevns Klint. The last known location for the specimen was in the collections of DGU [= Danmarks Geologiske Undersøgelse], which is now part of the semi-private institution GEUS in Denmark. In spite of several attempts at contacting GEUS for more information on its whereabouts, no information has come to hand, and the specimen may very well have become lost. Occurrence. Rare in the Cerithium Limestone Member at Stevns Klint. Description. Shell small, narrowly ovate with bluntly pointed apex. Protoconch low conical with just over two convex whorls, about 0.45 mm high and 0.65 mm wide. Teleoconch spire weakly convex with flat whorls covered by thin callus obscuring sutures. Last whorl contributing around 78 % of shell height, moderately convex, evenly contracted anteriorly, with broad uncallused zone between abapical edge of spire callus and posterior fasciole. Aperture elongate-ovate, strongly notched anteriorly, taking up about 54 % of shell height. Outer lip with evenly curved anterior margin lacking labral denticle. Columella thickened, gently concave, with four to six narrow plaits continuing into the aperture. Columellar callus bordered adapically by shallow anterior fasciole furrow, furrow terminating at aperture. Middle fasciole slightly wider, terminating just adapical to columella, and divided from the former by more or less distinct groove. Posterior fasciole widest, often indistinct due to effacement of adapical groove, groove terminating at adapical aperture angle. Spire grooves, including ancillid groove and band absent. Parietal callus very thin but for weak, rather narrow and nearly straight-sided tongue reaching up from aperture to spire callus, widening just before reaching the latter. Measurements. Holotype MGUH 33270 is 3.3 mm wide and 7.8 mm high, composed of protoconch and about four teleoconch whorls. Remarks. Ravn (1902) did not explain the reason for the taxon name, but it was clearly in honour of the geologist and collector of the specimen, Poul Christian Vilhelm Madsen Milthers, who found it in the year 1900. In spite of what was indicated by Vermeij (2001: p. 508) Spirancilla milthersii does not posses a labral denticle, but has an evenly curved anterior outer lip margin.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A990FF752B9CC4BCFBD1F9AE.taxon	materials_examined	Type species. Exilia pergracilis Conrad, 1860 (type by monotypy). Eocene, Alabama in the USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A990FF742B9CC7D7FC01FD64.taxon	description	Figs 31 S – T	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A990FF742B9CC7D7FC01FD64.taxon	materials_examined	Material. A single nearly complete external mould, MGUH 33272. Occurrence. Cerithium Limestone Member at Skeldervig, Stevns Klint. Description. Protoconch high conical with around 3 ¼ to 3 ½ weakly to moderately convex and smooth whorls. Protoconch 0.6 mm wide and 0.65 mm high. Teleoconch slender, high-spired, with moderately convex and thin-shelled whorls having a width corresponding to twice the height. Suture deep. Last whorl with gradual transition to convex base and moderately short canal. Aperture relatively short and narrow, the height taking up 36 % of the shell height, and with a width corresponding to 30 % of the height. Columella long, slightly curved and smooth, lacking columellar folds. Siphonal canal moderately wide and deep, not terminating in a notch. Teleoconch sculpture reticulate with moderately strong spiral and transverse ribs forming tubercles at nodes. First teleoconch whorl only with opisthocyrt and somewhat opisthocline transverse ribs, the number amounting to about 24 per whorl. Following whorl with three and later five spiral ribs in addition to and nearly as strong as transverse ribs. Spiral ribs numbering 15 on last whorl. Transverse ribs die out on adapical part of base. Measurements. MGUH 33272 is 5.5 mm high and 1.6 mm wide, consisting of protoconch and 4 ½ teleoconch whorls. Remarks. The specimen corresponds well with the diagnostic features of Exilia but for the short aperture, which takes up much less than half the shell height. Even so it is clear that it is closely related to species like the slightly younger Exilia crassistria (von Koenen, 1885). Therefore it is assigned to this genus. Exilia sp. differs from the Palaeocene E. crassistria (von Koenen, 1885) from Denmark by the coarser sculpture and slightly shorter aperture. It resembles the Selandian Parvisipho (Andonia) crispatus Ravn, 1939 from Denmark, but differs by the relatively higher protoconch and the higher number of transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A991FF742B9CC394FB93FCD6.taxon	materials_examined	Type species. Latiromitra specialis Locard, 1897 (type by monotypy). Recent, NE Atlantic.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A991FF742B9CC2C5FE27F976.taxon	description	Figs 31 U – W	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A991FF742B9CC2C5FE27F976.taxon	materials_examined	Material. A single external and internal mould, MGUH 33273. Occurrence. Extremely rare in the Cerithium Limestone Member. Skeldervig at Stevns Klint. Description. Protoconch with low dome-like apex; composed of between 1 and 3 smooth whorls. Transition between protoconch and teleoconch not pinpointed, but located at change in spire angle around 1 and 1 ½ whorls from apex or at beginning of weak spiral three whorls from apex. Teleoconch moderately slender, the whorls flattened with narrow shoulder demarcating deep adapical suture. Whorl height just surpassing whorl width. Aperture narrow with adapical channel, smooth outer lip and two moderately developed and widely spaced columellar folds; aperture and short siphonal canal corresponding to just over a third of the shell height. Teleoconch sculpture dominated by 14 – 16 low but sharp, weakly opisthocline and opisthocyrt transverse ribs per whorl, disappearing on last whorl. Spiral sculpture consisting of two adapical spiral cords and furrows and additional spiral cords on siphonal part of last whorl. Indistinct growth lines very weakly sigmoid, adapically following pattern of transverse ribs. Measurements. The nearly complete specimen MGUH 33273 is 12.9 mm high and 4.6 mm wide, consisting of nine whorls in total. Remarks. This specimen with its fusiform shell and predominantly axial sculpture shouldered at intersection between axial ribs and spiral cord (see Bouchet & Kantor 2000), closely resembles species in the genus Latiromitra Locard, 1897, to which it is tentatively assigned. More knowledge of the aperture and protoconch is needed for a confident assignment.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A992FF772B9CC3DCFEA8FCEF.taxon	materials_examined	Type species. Pleurotoma longiforma Aldrich, 1897 (subsequent designation by Gardner 1935). Oligocene, Mississippi in the USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A992FF712B9CC213FE0FFB9E.taxon	description	Figs 32 B – G 1902 Pleurotoma Cerithiorum n. sp. Ravn: 236 – 237, pl. III, fig. 4. 1902 Pleurotoma Steenstrupii n. sp. — Ravn: 237, pl. II, figs 23 – 24.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A992FF712B9CC213FE0FFB9E.taxon	materials_examined	Diagnosis. Protoconch smooth with no terminating part carrying transverse riblets. Teleoconch whorls weakly convex with shallow shoulder sulcus. Siphonal canal moderately short and wide. Sculpture development variable but shells typically carries fine spirals and around 10 to 12 transverse nodes below sinus. Material. Lectotype MGUH 123, MGUH 119, MGUH 33275, MGUH 33276, MGUH 33277, MGUH 33278, ØSM. 10042 - 109 - a – b, ØSM. 10042 - 116, 42 moulds with the informal sample numbers SH. 146, SH. 147, SH. 189, SH. 194. A, SH. 196, SH. 203. A – B, SH. 259, SH. 276, SH. 280, SH. 281. C – D, SH. 287. B, SH. 287. B – C, SH. 288, SH. 301. A – B, SH. 304. A – B, SH. 385. A, SH. 385. B, SH. 451, SS. 190. A – B, SR. 291, SR. 293, SR. 306, SR. 310. B, SR. 353, SR. 366. A – B, SR. 367, SR. 368, SR. 416, SR. 475, SR. 503, SR. 506, SR. 514, SR. 542. B, SR. 564, SR. 620. A – B, SR. 739, SO. 108. A – B, SO. 112. A – B, SO. 113, SO. 128, DN. 27. A – B and DN. 53. A – B, and around 60 specimens without number, which are deposited in the collections at the Natural History Museum of Denmark. Type stratum and type locality. The original work by Ravn (1902) was based on four specimens from the Cerithium Limestone Member at Stevns Klint, but only the one figured (Ravn 1902: pl. III: 4) is known to this day. This specimen, MGUH 123, is here selected as lectotype. The type material of Pleurotoma steenstrupii Ravn, 1902, here regarded as a subjective synonym, derives from the same locality and stratum. Occurrence. Common in the Cerithium Limestone Member at Stevns Klint and contemporaneous ‘ dead layer’ from Northern Jutland. This species may also occur sparingly in the lithified top of the Cretaceous Højerup Member. Description. Protoconch 0.8 to 1.0 mm high and 0.8 to 0.9 mm wide; high conical with 3 – 3 ½ smooth whorls expanding slightly faster than succeeding teleoconch whorls. Teleoconch high fusiform with weakly convex whorls with rounded to moderately sharp shoulder slightly abapical of whorl center; steep shoulder slope weakly concave to nearly straight. Aperture narrow, taking up around 45 % of shell height. Sinus broadly, but moderately deep V-shaped with rounded apex located on lower half of shoulder slope. Teleoconch sculpture of generally 10 – 12 moderately distinct to completely effaced transverse nodes or short ribs below sinus, crossed by normally distinct but fine spiral cords. A more pronounced spiral rib typically borders adapical suture; rib often granulate due to growth lines. Remarks. Orthosurcula s. l. cerithiorum differs slightly from the typical Orthosurcula species by the generally weakly developed shoulder sulcus, the typically distinct spirals and transverse sculpture and by the lack of transverse sculpture on the last protoconch-whorl. Except for the last feature all of these characters are found to vary within the same species as outlined below. This species is highly variable with regard to sculpture and shape of the shoulder slope. Most specimens have very weakly developed shoulder sulcus, but some feature a more concave slope outline better resembling the typical deep sulcus so characteristic of the genus. Transverse ribs, when present, are mostly found on the early teleoconch whorls, fading out on succeeding whorls. The fine, sharp spiral cords are generally strongly developed, but may become nearly effaced, resulting in a rather smooth surface especially on older specimens. It has thus not been possible to find any differentiating characters between this species and Pleurotoma steenstrupii Ravn, 1902, which is here regarded as a subjective synonym of T. (O.) cerithiorum (Ravn, 1902). A few very fragmentary specimens of a turriculid closely resembling the present species have been found in the latest Maastrichtian Højerup Member at Stevns Klint. Until better material has been uncovered they are here regarded as conspecific.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A994FF712B9CC45AFAB4FA91.taxon	materials_examined	Type species. Pleurotoma crenulata Lamarck, 1804 (type by original designation). Eocene, Paris Basin in France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A994FF702B9CC488FD88FCF4.taxon	description	Figs 32 H – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A994FF702B9CC488FD88FCF4.taxon	materials_examined	Diagnosis. Protoconch with two and a half whorls. Teleoconch relatively low, compact, with strongly carinated whorls with distinctly concave shoulder slope. Transverse ribs not overly falcate in form. Growth lines with rather wide and shallow sinus. Derivation of name. In honour of A. Rosenkrantz, who collected the type material and worked on the gastropod faunas from the Palaeogene of Denmark and Greenland. Type material. The holotype, MGUH 33279, is a nearly complete external mould collected by A. Rosenkrantz on 25 July 1943. Paratype MGUH 33280 was collected from the Cerithium Limestone Member at Skeldervig, Stevns Klint. Additional material. Two external moulds have been examined from the uncatalogued old collections of the Natural History Museum of Denmark. Both are collected by A. Rosenkrantz at Skeldervig (18 October 1943) and between Knøsen and Harvig (25 July 1943) at Stevns. Type stratum and type locality. Cerithium Limestone Member from somewhere between Knøsen and the first ladder at Harvig, Stevns Klint. Occurrence. Rather rare in the Cerithium Limestone Member at Skeldervig and south of Knøsen, Stevns Klint. Description. Protoconch 0.65 mm high and 0.7 mm wide, conical, consisting of 2 ½ smooth, weakly convex whorls separated by shallow suture. Transition to teleoconch fairly sharp, marked by the appearance of transverse nodes. Teleoconch shell small, compact and moderately thick with low carinated whorls two and a half times as wide as high. Shoulder carina pronounced, located approximately 1 / 3 of whorl height from abapical suture; shoul- der slope wide, convex, delineated by slight spiral swelling adjacent to adapical suture. Aperture elongated with rather gradual transition to wide, oblique siphonal canal, combined height corresponding to 3 / 5 of shell height. Lips smooth. Columella concave, abaxially deflected to the left. Teleoconch sculpture dominated by around 11 short transverse ribs or nodes per whorl; ribs sometimes developed into short spike where crossed by shoulder carina, fading out on abapical half of smooth shoulder slope and on adapical part of base. Transverse rib may form weak node on subsutural swelling. Spiral threads weak, strongest on base, fading out at periphery. Growth lines describing moderately weak and widely open, V-shaped sinus with apex at shoulder. Measurements. Largest specimen, holotype MGUH 33279 is 6.0 mm high and 2.7 mm wide, consisting of protoconch and three teleoconch whorls. Remarks. The generic assignment is tentative due to the rather open sinus and the resulting weakly developed falcate shape of the transverse ribs.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A995FF702B9CC2FCFEAAFBCF.taxon	materials_examined	Type species. Surcula (Cochlespira) boeggildi Ravn, 1939 (type by original designation). Paleocene, Copenhagen in Denmark.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A995FF732B9CC533FE18FCBC.taxon	description	Figs 32 L – N	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A995FF732B9CC533FE18FCBC.taxon	materials_examined	Diagnosis. Protoconch low conical with 3 ½ convex smooth whorls, followed by a sculptured whorl carrying around 18 weak but sharp transverse ribs. Teleoconch with sharply developed carina, coarse spiral threads or cords and sharp transverse ribs. Derivation of name. Refers to the generally well developed and sharp transverse ribs characterizing this species. Type material. The holotype MGUH 33281 is a nearly complete part and counterpart of an external mould collected by A. Rosenkrantz in August 1924. Paratype MGUH 33282 is an external mould from the same stratum and locality as the holotype. Additional material. 12 external and internal moulds of the teleoconch are found in the uncatalogued old collections of the Natural History Museum of Denmark. Type stratum and type locality. The Cerithium Limestone Member from just south of the Stevns Fyr at Stevns Klint. Occurrence. Rare in the Cerithium Limestone Member, where it has been found at Skeldervig, Harvig, Højerup and just south of Stevns Fyr at Stevns Klint. A specimen from the contemporaneous ‘ dead layer’ in Dania Quarry, Northern Jutland, has been examined in the collections of K. I. Schnetler. Description. Protoconch moderately low, conical, multispiral, consisting of convex whorls separated by deep suture. Initial 3 ½ whorls smooth, succeeded by weakly carinated whorl with approximately 18 weak, but sharp, slightly opisthocline and distinctly opisthocyrt transverse ribs. Spiral cords weak but present. Apex of U-shaped and widely open growth line sinus and corresponding trajectory of transverse ribs located adapically of carina. Transition to teleoconch rather gradual. Embryonic smooth part of protoconch 1.2 mm high and 1.5 mm wide. Teleoconch strongly elongate pagodiform in outline. Whorls relatively low, their height corresponding to 2 / 5 their width; whorls characterized by strongly developed median and flange-like peripheral carina, serration turned outwards to slightly adapically. Carina adapically bordered by an angularly concave shoulder slope with a nearly horizontal outer part and a slightly narrower inner part parallel to the axis. Aperture moderately wide and long, tapering evenly towards poorly differentiated narrow siphonal canal; combined height corresponding to 3 / 5 the shell height. Columella straight and smooth, with very weak central columellar plait. Outer lip apparently smooth. Teleoconch sculpture consisting of sharp opisthocline and opisthocyrt transverse ribs and finer spiral cords. Transverse ribs rather weak, thread-like on shoulder slope to sometimes completely effaced, becoming stronger around whorl periphery, forming toothed carinal margin. Transverse ribs fade out on adapical part of base. Spiral cords or threads mostly present on base and canal, where they form a more or less dense surface sculpture. Generally three spiral threads are seen on abapical part of early spire whorls, increasing on later whorls. The shoulder slope is generally free of spiral sculpture, but may carry a single lamellar spiral rib just inside carina. Growth line sinus wide, U-shaped and shallow; its apex located at transition between inner and outer slope area. Measurements. Holotype MGUH 33281 is approximately 12.7 mm high and 6.0 mm wide, consisting of around 6 ½ whorls, protoconch included. Remarks. Pseudocochlespira transversa n. sp. differs from the Selandian P. boeggildi (Ravn, 1939) from Denmark by a more pronounced carina and the obtuse protoconch with its deeper suture. It strongly resembles the Selandian P. koeneni (Arkhanguelsky, 1904) from Denmark and Russia, but differs from it by the moderately developed spiral sculpture and by the less opisthocline and carinated nature of the transverse ribs on the last protoconch whorl. It differs from the Selandian P. rosenkrantzi Schnetler, 2001 from Denmark by the more obtuse outline of the protoconch and the sharp transverse ribs and stronger spiral cords on the teleoconch whorls. The growth lines are furthermore less oblique.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A996FF732B9CC2B4FB21FBB6.taxon	materials_examined	Type species. Murex intorta Brocchi, 1814 (type by original designation). Neogene, Northern Italy.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A996FF722B9CC5EAFAE0FD2C.taxon	description	Figs 32 O – S	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A996FF722B9CC5EAFAE0FD2C.taxon	materials_examined	Diagnosis. Protoconch obtusely conical with slightly less than three smooth whorls. Teleoconch with subsutural spiral cord and constriction and in general around 15 to 18 short and sharp transverse ribs per whorl. Derivation of name. Name refers to Denmark from where it is known. Type material. The holotype, MGUH 33283, is a nearly complete external mould collected by A. Rosenkrantz on 18 July 1940. Paratype MGUH 33284 was collected from the Cerithium Limestone Member between Knøsen and Harvig, while paratype MGUH 33285 was found in the same bed at Skeldervig. Additional material. 15 moulds with informal sample numbers SH. 184. A, SH. 192. A, SH. 194. A – B, SH. 293. A, SH. 452, SR. 310. A – B (2 specimens), SR. 365. A – B, SR. 274, SR. 403. A – B, SR. 513, SR. 515, SR. 625, SR. 689. A – B, DN. 2, and 21 more or less fragmentary moulds from the uncatalogued old collections of the Natural History Museum of Denmark have been examined from the Cerithium Limestone Member at Stevns Klint. A single external fragmentary mould, ØSM. 10042 - 272 - b, is known from the Maastrichtian Højerup Member at Stevns Klint. Type stratum and type locality. Cerithium Limestone Member from the fourth Cerithium Limestone trough south of the old church at Højerup, Stevns Klint. Occurrence. Moderately common in the Cerithium Limestone Member at Stevns Klint. This species is also present in the contemporaneous ‘ dead layer’ in Dania Quarry, Northern Jutland, as evidenced by material collected by K. I. Schnetler. Furthermore it seems to range down into the Maastrichtian Højerup Member, though better material may show this to represent a separate species. Description. Protoconch obtusely conical, consisting of 2 ¾ smooth and convex whorls meeting at a marked angle. Transition to teleoconch sharp, marked by the appearance of sharp transverse ribs and finer spiral cords or threads, and by a change in spire angle. Teleoconch rather compact biconic fusiform with distinct stepped outline and short canal. Whorls moderately low, twice as wide as high, with strongly developed shoulder and distinct subsutural spiral swelling and constriction at adapical suture. Shoulder ramp concave. Aperture subtriangular with distinct adapical channel and rather gradual transition to wide and short siphonal canal; siphonal canal with rounded distal termination. Callus rather narrow, on parietal region additionally very thin, allowing the underlying spiral ribs to appear as weak folds. Apertural lips and columella otherwise smooth. Teleoconch sculpture dominated by around 15 to 18 short and sharp, opisthocline and slightly opisthocyrt transverse ribs; ribs highest at shoulder, tapering out towards sutures. Transverse ribs partly effaced at adapical whorl constriction, becoming stronger again on subsutural swelling. Spiral cords fine and close together, increasing in number with whorl size. Growth lines sigmoid, describing wide U-shaped sinus on abapical part of shoulder ramp. Measurements. Largest complete specimen, MGUH 33283, 14.9 mm high and 6.2 mm wide, consisting of protoconch and four teleoconch whorls. Remarks. Pseudotoma danica n. sp. differs from the contemporaneous Pseudotoma sp. figured by Kollmann & Peel (1983) from Greenland by a slightly more compact shell with a weaker shoulder and a more rounded distal termination of the siphonal canal. It is distinguished from the Danish Selandian P. steenstrupi (von Koenen, 1885) by typically sharper and more numerous transverse ribs and a more gradual transition to the siphonal canal.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A997FF722B9CC25EFE9CFC8C.taxon	materials_examined	Genus indet.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A997FF722B9CC6D0FACFF81A.taxon	materials_examined	Type species. Cymatium pahiense Marshall & Murdoch, 1921 (type by original designation). Mid Eocene, New Zealand.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A998FF7D2B9CC1D7FA9AFA98.taxon	description	Figs 33 B – E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A998FF7D2B9CC1D7FA9AFA98.taxon	materials_examined	Diagnosis. Elongate ranellid in shape with two strong columellar plaits and around 14 teeth on outer lip. Canal not very long. Derivation of name. The name, which translates ‘ False Sassia’, refers to its strong resemblance to Sassia faxense Ravn, 1933 in the same deposits. Type material. The holotype, MGUH 33287, is a nearly complete mould missing part of the aperture. Paratype MGUH 33288 was collected at Holtug, Stevns Klint, and comes from the same stratigraphic level as the holotype. Paratype MGUH 33289 is a mould of the aperture coming from the lithified top of the Højerup Member at Skeldervig, Stevns Klint. Additional material. An external mould with the informal sample number SR. 1123. A, and a fragmentary mould without number from the collections of the Natural History Museum of Denmark. Type stratum and type locality. The lithified top of the Maastrichtian Højerup Member at Stevns Klint. Occurrence. Lithified top of the Maastrichtian Højerup Member at Stevns Klint. Description. Protoconch obtusely conical, composed of at least two smooth, not very convex whorls. Suture fairly weak. Teleoconch elongated ranellid in outline with moderately convex whorls separated by distinct suture. Aperture elongated, taking up slightly more than half the shell height. Aperture width corresponding to 1 / 3 of its height. Parietal region with thin callus not completely covering underlying spiral ribs. Columella carrying two strong folds, the abapical one strongest. Denticles on the inner lip, if present, not preserved. Varices after old apertures fairly weak and thin, generally placed with a distance of half a whorl. MGUH 33289, representing an internal mould of the aperture, has a wide and weakly convex outer lip carrying 14 evenly spaced small denticles at the inner margin. The small teeth decrease in size at the abapical end. Teleoconch sculpture cancellate, consisting of thin spiral ribs and adapically slightly stronger transverse ribs, fading out on base. Spiral ribs numbering around 14 from adapical suture to beginning of canal, with between two and five on spire whorls. Transverse ribs very weakly sigmoid, at times slightly prosocline adapically, numbering around 20 per whorl on early whorls. Tubercles formed at connecting points between ribs. Growth lines nearly orthocline to slightly prosocline adapically. Measurements. Protoconch poorly preserved, measuring between 0.75 and 1.2 mm in height and between 1.05 and 1.7 mm in width. Holotype specimen MGUH 33287 is 8.0 mm high and 4.8 mm wide, consisting of five whorls of which the last whorl lacks the basal part of the canal. Remarks. Tatara pseudosassia n. sp. closely resembles the middle Danian Tatara danica Schnetler & Petit, 2006 from Faxe, Denmark, in the external sculpture and elongated ranellid shape, but differs by its slightly more compact shell outline, a higher number of teeth on the outer lip and seemingly by a slightly shorter canal.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A998FF7F2B9CC483FE5BFCBE.taxon	description	Figs 33 F – H	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A998FF7F2B9CC483FE5BFCBE.taxon	materials_examined	Material. A single external and internal mould, MGUH 33290. Occurrence. The lower Danian Cerithium Limestone Member at Skeldervig, Stevns Klint. Description. Protoconch poorly preserved, obtusely conical with at least two smooth and rather weakly convex whorls. Height at least 0.6 mm. Teleoconch slender, mitriform with moderately to weakly convex whorls separated by thin, shallow suture mainly marked by whorl angle. Whorl width corresponding to 1.6 times the height. Last whorl moderately convex with a rather gradual transition to canal. Aperture narrowly lenticular with a height corresponding to slightly over half the shell height. Columella fairly long with two strong columellar folds on adapical half, succeeded abapically by at least four blunt teeth. Callus absent or thin, exposing continuation of spiral sculpture on parietal region. Outer lip thickened into varix with 11 evenly spaced teeth on inner side; teeth strongest on adapical part. Teleoconch sculpture reticulate, consisting of sharp-ridged and weakly prosocline transverse ribs crossed by spiral ribs. Tubercles formed at connecting nodes. Third teleoconch whorl with 4 – 5 primary spiral ribs and around 28 to 30 transverse ribs. Varices occur with a frequency of nearly 2 / 3 per whorl. Measurements. MGUH 33290 is 19.8 mm high and around 10.5 mm wide, consisting of about five teleoconch whorls and the lower two protoconch whorls. Remarks. Tatara sp. differs from T. pseudosassia n. sp. described above by the slender outline and a finer transverse teleoconch sculpture. It seems furthermore to differ from the Danish middle Danian T. danica Schnetler & Petit, 2006 by the presence of more teeth on the outer lip, by the slender form and by the gradual transition from base to canal on the last whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99AFF7F2B9CC6D4FAD2F818.taxon	materials_examined	Type species. Cancellaria elevata Lea, 1833 (subsequent designation by Cossmann 1889: p. 231). Eocene, Alabama, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99AFF7F2B9CC2E2FB24FBE9.taxon	materials_examined	Type species. Cancellaria quantula Deshayes, 1864 (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99AFF7F2B9CC510FCEAF8A4.taxon	description	Figs 33 I – L	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99AFF7F2B9CC510FCEAF8A4.taxon	materials_examined	Material. Two fragmentary moulds, MGUH 33291 and MGUH 33292. Occurrence. Cerithium Limestone Member at Skeldervig and Højerup, Stevns Klint. Description. Protoconch poorly preserved, 0.85 mm high and 1.0 mm wide, low conical, consisting of between 2 and 2 ½ smooth whorls with fairly weak sutural angle. Teleoconch slender, ranellid-like with moderately convex whorls separated by fairly deep suture. Whorls moderately low, their height corresponding to 40 % of the width. Whorl periphery located approximately 2 / 3 of height from adapical suture. Varix strongly developed. Aperture with straight columella carrying two strong columellar folds. Teleoconch sculpture on initial two whorls dominated by 13 to 14 moderately sharp, strong, slightly prosocline transverse ribs, narrower than the interspaces. Number of ribs increasing with size. Transverse ribs crossed by fine spiral threads numbering 5 to 6 on spire whorls and with 12 from suture to mid-way on the base on last whorl. Growth lines sigmoid. Measurements. Specimen MGUH 33292 is 4.4 mm high and 3.1 mm wide, consisting of protoconch and 2 ½ teleoconch whorls, the last whorl lacking base and canal. Remarks. The taxon resembles Sveltella multistriata (Ravn, 1939) from the Danish Selandian, although the whorls are less convex and the shell slightly more slender in outline. It also shows a strong affinity to the Danian Sveltella tobiasseni Schnetler & Petit, 2010 from Greenland, but has only up to 2 ½ protoconch whorls and not 2 ¾; the protoconch suture is less deep and the shell more slender.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99BFF7E2B9CC1D4FCD9FC44.taxon	description	Figs 33 M – N	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99BFF7E2B9CC1D4FCD9FC44.taxon	materials_examined	Material. MGUH 33293, MGUH 33294 and possibly an incomplete external mould with the informal sample number SS. 213. A – C. Occurrence. The Cerithium Limestone Member at Højerup and perhaps Flagbanken, Stevns Klint. Description. Protoconch conical with nearly planispiral apex. It consists of just over three convex and smooth whorls, with a sharp transition to teleoconch marked by the appearance of transverse ribs. Width 0.95 mm; height slightly lower. Teleoconch fairly slender with highly convex whorls and a marked shoulder and adapical ramp; whorls twice as wide as high. Last whorl taking up approximately 70 % of the shell height, abapically forming short canal. Columella with two moderately weak folds. Teleoconch sculpture dominated by 10 to 11 sharp, prosocline transverse ribs, fading out adapically and abapically on ramp and base; ribs highest at shoulder. Ribs crossed by four or five fine spiral threads from shoulder and down on spire whorl, and two additional but weaker spiral threads on ramp. Last whorl covered by 12 more or less evenly distributed primary spiral threads and two weaker ones alternating with the original two spiral threads on ramp. Measurements. MGUH 33293 is 2.6 mm high and 1.6 mm wide, consisting of protoconch and two teleoconch whorls. Remarks. This taxon has a strong morphological resemblance to the Selandian Babylonella ravni (Glibert, 1960) from Denmark, but the protoconch has more whorls and may be slightly more conical in outline, while the number of transverse teleoconch ribs per whorl is in the low end.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99BFF7E2B9CC2B4FA35FBB6.taxon	materials_examined	Type species. Cancellaria costulata Lamarck, 1803 b (type by original designation of Uxia Joussaume, 1887). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99BFF792B9CC5E5FB16FDDC.taxon	description	Figs 33 O – R	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99BFF792B9CC5E5FB16FDDC.taxon	materials_examined	Diagnosis. Protoconch conical with 2 ¾ whorls. Anterior canal of teleoconch moderately long and shallow. Sculpture moderately coarse with around 16 transverse ribs per whorl and four spiral ribs. Derivation of name. In honour of Claus Heinberg, who has been of great help with the initial field work and discussions on the palaeo-environment during the deposition of the Cerithium Limestone Member. Type stratum and type locality. The holotype was collected 50 to 60 centimetres above the Cretaceous-Palaeogene boundary (= the P 1 b planktonic foraminifer zone), corresponding to the top of the Cerithium Limestone Member at Flagbanken, Stevns Klint. Type material. The holotype MGUH 33295 is a nearly complete internal and external mould. Paratype MGUH 33296 is an internal and external mould from the Cerithium Limestone Member just north of the Holtug Quarry. Occurrence. Unitas heinbergi n. sp. is known from the Cerithium Limestone Member north of Holtug Quarry and from the P 1 b planktonic foraminifer zone 50 to 60 centimetres above the Cretaceous-Palaeogene boundary, corresponding to the top of the Cerithium Limestone Member at Flagbanken, Stevns Klint. Description. Protoconch conical with a somewhat flattened or planispiral apex and consisting of 2 ¾ flattened to later convex whorls. Initial whorls smooth, while last half whorl has three weak spiral threads. Suture moderately deep. Protoconch measuring about 1.2 mm in height and 1.3 mm in width. Transition to teleoconch marked by the appearance of strong transverse ribs. Teleoconch slender fusiform with adapically convex whorls separated by deep suture. Spire whorls twice as wide as high. Base less convex, ending in wide canal. Aperture elongated oval in outline, abapically reaching out into moderately long and shallow canal. Columella deflected slightly to the left, with two moderately strong adapical folds and a much weaker abapical fold. Callus moderately thick and wide on parietal region. Outer lip with low varix and seven rounded teeth at an even distance from each other; the abapical one much weaker than others. Teleoconch sculpture dominated by sharp and strong, moderately prosocline transverse ribs numbering around 16 per whorl. Ribs fading out at transition to canal. The transverse ribs are crossed by around four weaker spiral ribs on spire whorls, forming sub-reticulate pattern. Base and especially canal more dominated by spiral ribs. Unevenly distributed and thin varices occur throughout. Measurements. Holotype MGUH 33295 approximately 5.2 mm high and 3.0 mm wide, consisting of protoconch and two teleoconch whorls. Remarks. Unitas heinbergi n. sp. differs from the Danian Unitas malaisei (Briart & Cornet, 1877) from Belgium by the higher number of transverse ribs per whorl, the longer anterior canal and the more adapical location of the central columellar fold relative to the columellar termination. It differs from the middle Danian Unitas anderseni Schnetler & Petit, 2006 and U. aliceae Schnetler & Petit, 2006 from Faxe, Denmark, by the distinctly less elongated teleoconch outline with its coarser sculpture. It appears very closely related to the Palaeocene (Selandian) U. scalata Traub, 1989 from Germany, but differs by a less convex whorl outline and a larger protoconch.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99CFF792B9CC3DCFB5EFD0E.taxon	materials_examined	Type species. Buccinum evulsum Solander in Brander (1766) (type by original designation). Eocene, England.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99CFF792B9CC272FDE3F95A.taxon	description	Figs 34 A – E	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99CFF792B9CC272FDE3F95A.taxon	materials_examined	Material. Two fragmentary moulds, MGUH 33297 and MGUH 33298, the latter preserved as an amalgamation of an internal and external mould due to early dissolution of the shell surface. Occurrence. The Cerithium Limestone Member south of Stevns Fyr and north of Holtug Quarry, Stevns Klint. Description. Protoconch somewhat dome-like obtusely conical, with a small beginning and consisting of 1 ½ to 2 ½ convex whorls. Transition to teleoconch gradual, at least on the inner mould. Teleoconch thin-shelled, ovoid-conical with moderately convex whorls separated by impressed suture. Spire whorls less than half as high as wide, characterized by narrow but distinct subsutural ramp. Aperture moderately narrow; outer lip with 6 or 7 teeth and in some strengthened by weak varix. Short columella with 3 strong columellar folds. Teleoconch sculpture dominated by unevenly weak spiral threads or bands, separated by narrower grooves and increasing in number with whorl size. Shoulder ramp demarcated by stronger spiral band, which may be separated from adapical suture by up to four weaker threads on final whorls. Spirals crossed by prosocline and opisthocyrt growth lines curving strongly forward adapically. At times growth stops form weak collabral threads, resulting in a weakly reticulate sculpture. Weak varices present but unevenly distributed. Measurements. The best preserved specimen MGUH 33297 is 7.7 mm wide and close to 12.5 mm high, but lacks the apex. Remarks. This taxon is morphologically similar to the Palaeocene Admetula eivindi Schnetler & Petit, 2010 from Greenland, but is distinguished through the strength of the more obtuse protoconch, a less marked shoulder ramp, stronger columellar folds and by the presence of varices. It differs from the Selandian Kroisbachia conoidea (von Koenen, 1885) and Admetula latesulcata (von Koenen, 1885), both from Denmark, by the much narrower aperture and stronger columellar folds.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99CFF792B9CC646FB64F88B.taxon	materials_examined	Type species. Kroisbachia haunsbergensis Traub, 1989 (type by original designation). Paleocene, Austria.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99DFF782B9CC1D4FB64FC60.taxon	description	Fig. 34 F	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99DFF782B9CC1D4FB64FC60.taxon	materials_examined	Material. One mould, MGUH 33299. Occurrence. The Cerithium Limestone Member at Skeldervig, Stevns Klint. Description. Protoconch obtusely conical, 1.0 mm wide and 0.75 mm high, consisting of around 2 ¾ smooth and moderately convex whorls. Teleoconch subfusiform to ovoid-conical with an H / W ratio of around 1.3. Last whorl corresponding to nearly 80 % of shell height. Whorls moderately to weakly convex with a narrow subsutural ramp and a whorl periphery located immediately adapical to deep abapical suture. Aperture subrectangular, the height corresponding to nearly 2 / 3 of shell height. Columella moderately long, slightly curved, carrying three strong columellar folds. Siphonal canal short. Teleoconch sculpture composed of weak spirals, strongest adapically around shoulder. Spirals crossed by up to around 35 weak prosocline transverse ribs or growth lines per whorl; ribs fading out at transition to base. Measurements. The available specimen MGUH 33299 is more than 3.4 mm high and 2.7 mm wide, consisting of protoconch and two teleoconch whorls. Remarks. This taxon is clearly distinguished from the Selandian Kroisbachia conoidea (von Koenen, 1885) from Denmark by the relatively larger aperture and stronger columellar folds. It has more resemblance to the Selandian K. peeli Schnetler & Petit, 2010 from Greenland, from which it differs by the stronger columellar folds and higher protoconch whorl number. It differs from the Palaeocene K. haunsbergensis Traub, 1989 from Germany by the absence of well developed spiral ribs on the base and by the much denser transverse ribbing.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99DFF782B9CC5C8FD1BF8A9.taxon	description	Fig. 34 G	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99DFF782B9CC5C8FD1BF8A9.taxon	materials_examined	Material. Specimen MGUH 33300 is an external mould. Occurrence. Lithified top of the Maastrichtian Højerup Member at Holtug. Description. Protoconch not preserved. Teleoconch bucciniform with weakly convex whorls separated by distinct, but fairly shallow suture. Whorls 2 ½ times as wide as high. Last whorl relatively large with evenly rounded sides and poorly developed canal. Aperture moderately wide and high, the height corresponding to at least 60 % of shell height. Siphonal canal short, wide, turned slightly to the left abapically. Columella moderately long, slightly curved, with weak columellar fold centrally. Outer lip slightly thickened but otherwise simple, without teeth. Teleoconch sculpture consisting of fine, slightly irregular, flattened spiral ribs. A single stronger spiral rib borders the adapical suture. Growth lines prosocline and weakly sigmoid. Measurements. MGUH 33300 is 3.8 mm wide and 5.7 mm high, but lacks the apex. Remarks. The taxon differs from typical Mataxa species by the shorter siphonal canal and the presence of only one weak columellar fold. It seems thus not to be a Mataxa in the strict sense, but nonetheless appear closely related. It is here regarded as a Mataxa sensu lato. This taxon differs from the Late Cretaceous Mataxa elegans Wade, 1916 of North America by the more convex whorls, the lack of teeth and by the presence of only one weak columellar fold. It differs from the Maastrichtian Mataxa arida Squires & Saul, 2003 a from California by the much less sigmoid growth lines; the much finer sculpture and by the distinctly shorter siphonal canal.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99DFF782B9CC298FB1DFBD1.taxon	materials_examined	Type species. Mataxa elegans Wade, 1916 (type by original designation). Cretaceous, Tennessee, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99FFF7A2B9CC77AFC6AF83B.taxon	materials_examined	Genus Ravniella Rosenkrantz, 1970 Type species. Cinulia danica Ravn, 1902 (type by original designation). Early Danian, Stevns Klint, Denmark. Remarks. Rosenkrantz (1970) erected the taxon as a subgenus of Tornatellaea Conrad, 1860 due to its close affinity with that genus, but it was later raised to full generic status by Kollmann & Peel (1983). Ravniella Rosenkrantz was already defined in a meeting abstract published by Rosenkrantz (1963) and a type species was selected in a later abstract by Rosenkrantz (1966). Nevertheless the name did not become available before the 1970 publication of Rosenkrantz as meeting abstracts are regarded as invalid media for publication of new taxa (Article 9.9 of the International Code of Zoological Nomenclature).	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99FFF7A2B9CC39AFD6EFAEE.taxon	description	Figs 34 H – I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A99FFF7A2B9CC39AFD6EFAEE.taxon	materials_examined	Material. Specimen MGUH 33301 consists of an external and part of an internal mould. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig, Stevns Klint. Description. Protoconch conical, multispiral, nearly 0.7 mm high and 0.7 mm wide, consisting of around 3 ½ weakly convex and smooth whorls. Suture moderately impressed. Teleoconch weakly to moderately convex with gradual transition to base. Aperture and possible canal not preserved. Teleoconch sculpture consisting of fine spiral striae, the most adapical slightly stronger than succeeding ones. Growth lines appearing nearly orthogonal. Measurements. MGUH 33301 is 1.55 mm high and 0.9 mm wide, consisting of 4 ¼ whorls. Remarks. The fragmentary and sparse material makes a more precise taxonomical assignment of this species very dubious at best. The nearly orthogonal growth lines, the multispiral protoconch and the seeming lack of transverse teleoconch sculpture may suggest a possible connection to the family Buccinidae. It superficially resembles the young shell of Orthosurcula s. l. cerithiorum (Ravn, 1902) treated above, but differs by the growth line pattern and by the slightly smaller size of the protoconch.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A0FF452B9CC7AEFC5AF8DD.taxon	materials_examined	Type species. Modulus disculus Philippi, 1846 (type by monotypy). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A0FF452B9CC1D4FD94F9E2.taxon	description	Figs 34 J – N 1902 Cinulia danica n. sp. Ravn: 238 – 239, pl. III, fig. 7. 1970 Tornatellaea (Ravniella) danica (Ravn) — Rosenkrantz: 431 – 433, fig. 10: 1.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A0FF452B9CC1D4FD94F9E2.taxon	materials_examined	Diagnosis. Protoconch heterostrophic, of 1 ½ smooth whorls. Teleoconch low-spired with strongly developed columellar folds and around eight teeth on outer lip. Whorl sculpture of widely spaced, finely pitted spiral grooves, generally numbering three to five on spire whorls. Type stratum and type locality. Cerithium Limestone Member at Stevns Klint, Denmark. Material. Of the original 12 type specimens Ravn (1902) used for his description, only the one figured, MGUH 126, is known today. The others were not given a number and are no longer possible to separate from the rest of the specimens found in the old collections. For this reason MGUH 126 is here selected as lectotype. Additional material consists of MGUH 33302, MGUH 33303, ØSM. 10042 - 88 - a – b, ØSM. 10042 - 89 - a – b, ØSM. 10042 - 106, more than 130 specimens without numbers from the old collections at the Geological Museum, Natural History Museum of Copenhagen, and 43 new specimens with the informal sample numbers SH. 106. B, SH. 142, SH. 148, SH. 161, SH. 186, SH. 197. A, SH. 205, SH. 206, SH. 285. A – B, SH. 285. A, SH. 289 (2 specimens), SH. 290, SH. 303. A (2 specimens), SH. 324, SH. 325. B – C, SH. 325. C, SH. 334. A – B, SH. 334. C, SH. 352. A – B (6 specimens), SH. 384. A, SH. 410. A – B, SH. 414. B – C, SH. 416, SH. 418. A – B, SR. 168, SR. 225, SR. 278, SR. 283, SR. 286, SR. 307. A – B, SR. 400, SR. 516 (2 specimens), SR. 570. A – C, SR. 642. B, SO. 110. A – B, DN. 26. A – B, DN. 30. A – B, DN. 33. A – B (2 specimens) and DN. 52. A – B. Occurrence. Common in the Cerithium Limestone Member at Stevns Klint and in the contemporaneous ‘ dead layer’ in Northern Jutland. Description. Shell subovate, small, generally below 10 mm high, width / height ratio varying between 57 and 70 %, generally decreasing with size. Protoconch heterostrophic with nearly 1.5 loosely coiled, smooth whorls. Transition to teleoconch sharp, marked by appearance of spiral grooves. Protoconch between 0.6 and 0.8 mm wide. Teleoconch with rather weakly convex whorls and distinct suture. Spire sides nearly straight to at times slightly concave due to an increase in whorl expansion on later whorls. Last whorl corresponding to between 75 and 85 % of shell height. Aperture narrowly oval-rhombic, width approximating 2 / 3 the height. Inner lip with two widely spaced, oblique strong columellar folds and a weaker parietal fold slightly above columella. Outer lip moderately sharp, slightly thickened, carrying around eight sharp teeth, although adapical and abapical ones often appears effaced or missing. Whorl sculpture of widely spaced, finely pitted spiral grooves, generally numbering three to five on spire whorls, adapical groove markedly stronger than succeeding ones. Last whorl typically with around 15 spiral grooves. Measurements. Lectotype MGUH 126 is approximately 6.5 mm high and 4.3 mm wide, consisting of 5 ½ whorls. Remarks. This gastropod, which most likely was a vermivore (Jeffery & Tracey 1997), is one of the more dominant gastropod species in the Cerithium Limestone Member, although it generally does not exceed 2 % of the total number of gastropod specimens.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A1FF442B9CC1D4FDF6F930.taxon	description	Figs 34 O – S 1902 Solarium selandicum n. sp. Ravn: 216, pl. I, figs 5 – 6. 1903 Solarium selandicum Ravn — Ravn: 380, 394 – 395. 1940 Solarium selandicum Ravn — Rosenkrantz: 513.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A1FF442B9CC1D4FDF6F930.taxon	materials_examined	Diagnosis. Protoconch heterostrophic, submerged with around 2 ½ smooth, rounded convex whorls. Teleoconch H / W ratio approximately 0.55. Peripheral keel strongly developed, without sculpture. Adapical side of teleoconch whorl distinctly concave with fine but distinct spiral and radial sculpture; spiral threads strongest close to sutures; growth lines strongly posteriorly turned; beading on umbilical keel coarse. Type stratum and type locality. Cerithium Limestone Member at Stevns Klint, Denmark. Material. Of the eight type specimens used for the description of the species (Ravn 1902: p. 216), only the two figured, MGUH 79 and MGUH 80, are registered and known today. The most complete of the two, MGUH 79, figured by Ravn (1902: pl. I: 5) is here selected as lectotype. MGUH 33304, MGUH 33305, 35 moulds with informal sample numbers SO. 184, SH. 149. A, SH. 190. A – B, SH. 195, SH. 261, SH. 286. A – B, SH. 294. A, SH. 339, SH. 380, SH. 415. A, SR. 142, SR. 145, SR. 146, SR. 152. A (2 specimens), SR. 170. A – B, SR. 286, SR. 292, SR. 295 (2 specimens), SR. 359, SR. 366. A, SR. 391, SR. 394, SR. 467, SR. 479, SR. 492. A – B, SR. 519. A, SR. 519. A – B, SR. 571. A – B, SR. 578. A – B, SR. 629. A, SR. 682. A – B, SR. 699 and DN. 54. A, and around 30 specimens without numbers are deposited in the collections of the Natural History Museum of Denmark. The largest measures 9.0 mm in width. Occurrence. Cerithium Limestone Member at Stevns Klint and the contemporaneous ‘ dead layer’ in Dania and Vokslev quarries, Northern Jutland. Description. Shell low cone-shaped, height / width ratio approximating 0.55. Protoconch heterostrophic, submerged with around 2 ½ smooth, rounded convex whorls. Large, width about 1.0 to 1.1 mm. Transition to teleoconch marked by beginning of sculpture. Teleoconch thick-walled with distinctly concave spire whorls transitioning into rounded peripheral keel at marked abapical suture. Last whorl with sharply rounded peripheral keel and convex base. Aperture rounded to sub-pentagonal, slightly wider than tall, acuminate laterally towards peripheral and umbilical keel. Umbilicus moderately wide, around 25 % of shell width, bordered by strong umbilical keel. A secondary pronounced keel occurs 1 / 3 the distance between umbilical and peripheral keel from umbilicus. Base between two abapical keels strongly concave. Teleoconch sculpture on spire consisting of fine spiral threads and ribs and radial ribs or growth increments; suture bordered on both sides by stronger and weakly beaded spiral rib, beads formed by growth increments, numbering close to 40 on initial teleoconch whorl, increasing gradually on later whorls. Weak closely spaced spiral threads between subsutural ribs, often around six on early whorls, sometimes more on later whorls. Growth lines sigmoid, strongly prosocline on concave part, becoming more orthocline at sutural margins. Peripheral keel and base with slightly prosocline growth lines and no other sculpture. Keel abaxially to umbilical keel beaded, beads rounded, around 21 on last whorl of the 7.1 mm wide paralectotype MGUH 80. Umbilical keel bearing coarse, oblique posteriorly directed nodules or ridges, nodules numbering 12 to 13 on last whorl of the 7.1 mm wide paralectotype MGUH 80. Measurements. MGUH 33305 is 5.9 mm wide and 3.5 mm high, consisting of protoconch and three teleoconch whorls. The species may reach widths of at least 9 mm and have up to 4 ½ whorls. Remarks. D. selandicum is easily distinguished from the Eocene type species D. disculus (Philippi, 1846) from France by the more strongly posteriorly directed growth lines, the more concave upper surface of the whorls, the coarser beading on the umbilical keel and by the stronger collabral and spiral sculpture but lacking an outer spiral furrow on the abapical side of the whorl.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A1FF442B9CC668FC5FF861.taxon	materials_examined	Type species. Bifrontia zanclaea Phillipi, 1844 (type by monotypy). Pliocene, Sicily.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC224FBCAFC26.taxon	materials_examined	Type species. Solarium patulum Lamarck, 1804 (type by original designation). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC555FA88FA01.taxon	description	Figs 35 C – D	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC555FA88FA01.taxon	materials_examined	Material. MGUH 33307, a poorly preserved mould of a spire. Occurrence. Lithified top of the Højerup Member of the Maastrichtian Tor Formation at Mandehoved, Stevns Klint. Description. Protoconch not known. Teleoconch very low-spired with slowly expanding concavo-convex whorls separated by distinct ridge. Sculpture consisting of weak spiral striation and dense, strongly prosocline and slightly prosocyrt growth lines. Measurements. Specimen MGUH 33307 has a diameter of just over 8.5 mm, consisting of at least six teleoconch whorls. Remarks. The poor preservation makes the generic assignment somewhat tentative, even though the low spire with its concave and slowly expanding whorls together with the very fine sculpture strongly supports it.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC789FACAF8C2.taxon	materials_examined	Type species. Turbo quadricarinatus Brocchi, 1814 (subsequent designation by Cossmann 1888: p. 68). Pliocene, Sicily.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC1D4FAF7FCFB.taxon	description	Figs 35 A – B	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF472B9CC1D4FAF7FCFB.taxon	materials_examined	Material. MGUH 33306 and three fragmentary specimens with the informal sample numbers SH. 9, SR. 1080 and SR. 1081, all belongs to the Natural History Museum of Denmark. Occurrence. The specimens were collected from the lithified top of the Tor Formation at Rødvig, Holtug Quarry, Harvig and Højerup Church, Stevns Klint. Description. Protoconch not preserved. Shell consisting of 3 ½ very slightly widening subrectangular and weakly to moderately convex whorls with two sharply distinct, unbeaded keels located at whorl periphery and umbilical margin respectively. Height and width of whorl largely equal. Umbilical keel is in contrast to the peripheral keel completely free of the following whorl. Shell surface smooth except for a few very indistinct spiral cords and weak growth lines. Measurements. The most complete specimen, MGUH 33306, is 4.3 mm in diameter with a whorl height of 1.0 mm on the third whorl. Remarks. This species differs from other species of this genus by the clearly convex spire and the nearly smooth and moderately inflated whorls carrying continuous, unbeaded keels. It resembles somewhat the Late Cretaceous Pseudomalaxis? stantoni Sohl, 1960 from Tennessee, but differs in the distinctly more angular cross-section of the whorls; the absence of a third spiral cord on the outer whorl face and in the slightly more convex apex.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF462B9CC63FFEFAFAD0.taxon	description	Figs 35 E – I	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A2FF462B9CC63FFEFAFAD0.taxon	materials_examined	Diagnosis. Protoconch smooth, nearly perpendicular to teleoconch axis. Teleoconch moderately slender with ap- proximately 50 thin transverse ribs and three to four coarser, slightly less densely spaced spiral ribs adapical of keel. Keel situated one quarter the whorl height from abapical suture. Derivation of name. Refers to the Danian age and its Danish origin. Type material. Holotype MGUH 33308 is a nearly complete external mould. Paratype MGUH 33309 is an external mould and comes from the Cerithium Limestone Member at Skeldervig. Paratype MGUH 33310 is a protoconch from 20 to 25 cm above the K-Pg boundary at Højerup, while the protoconch paratype MGUH 33311 was found 42 to 52 cm above the K-Pg boundary at Rødvig. Paratype MGUH 33312 is an external mould of the spire from Vokslev, Northern Jutland. Additional material. ØSM. 10042 - 89 - c – d, 10 specimens with the informal sample numbers SH. 294. A – C, SH. 398, SH. 446, SH. 499. A – B, SR. 476. A, SR. 624, SR. 631, SR. 641, SR. 668. B and SO. 189, and 3 more or less fragmentary moulds without numbers from the uncatalogued old collections at the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member of the Rødvig Formation at Holtug, Stevns Klint. Occurrence. Middle to upper part of the Cerithium Limestone Member of the Rødvig Formation and basal Korsnaeb Member of the Stevns Klint Formation. This species has been found at Rødvig, Skeldervig, Højerup and at Holtug, all localities at Stevns Klint. Additionally it occurs in material collected by C. Heinberg from the contemporaneous ‘ dead layer’ in Vokslev Quarry, Northern Jutland. Description. Protoconch hyperstrophic, diverging about 125 ˚ from teleoconch axis, smooth with sharp transi- tion to teleoconch. Teleoconch slender, conical; whorls with strong keel ¼ the whorl height from rather indistinct abapical suture; whorl sides flattened to slightly concave adapically and abapically to keel. Last whorl taking up 40 % of shell height. Base with weaker keel, abapically flattened. Aperture rounded, sub-rectangular with slightly thickened lip. Teleoconch sculpture on first whorl consisting of densely spaced weak transverse ribs and three to four spiral ribs, the one on the keel being the strongest. Number of spiral and transverse ribs increasing abapically on shell, the spiral ribs reaching a number of five to six moderately coarse ribs. Transverse ribs numbering around 50 on fourth whorl. Base, except for strong spiral rib on abapical keel, nearly smooth or carrying weak spiral ribs and faint transverse ribs. Measurements. Holotype MGUH 33308 is 4.3 mm high and 1.7 mm wide with 5 ½ teleoconch whorls. Remarks. Mathilda dania n. sp. resembles the Selandian Mathilda carinata (Ravn, 1939) from Denmark, but has distinctly less convex whorls with a stronger keel and fewer strong spiral ribs abapical to the keel. It differs from the Danian Mathilda unicarinata (Ravn, 1933) from Denmark by having only two spirals above the carina on the early whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A3FF462B9CC43AFA81FA2E.taxon	materials_examined	Type species. Cerithium ziczac Eudes-Deslongchamps, 1842 (type by original designation). Early Jurassic, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A3FF412B9CC75FFEBEFE30.taxon	description	Figs 35 J – K	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A3FF412B9CC75FFEBEFE30.taxon	materials_examined	Material. MGUH 33313 and an external mould with the informal sample number SR. 938, collected in connection with this study. Occurrence. Upper 15 centimetres of the upper Maastrichtian Højerup Member at Rødvig, Stevns Klint. Description. Protoconch smooth, apparently hyperstrophic, but only last quarter of a whorl preserved. Transition to teleoconch sharp. Teleoconch thick-shelled, very slender with relatively high, convex, keeled whorls; keel located just abapical to whorl center; whorl height corresponding to approximately 80 % of the width. Suture very faint. First teleoconch whorl with three spiral ribs, a weak adapical one and two stronger spiral ribs located on keel and halfway between keel and abapical suture. A fourth, weak spiral rib is added between spiral rib 1 and keel on second teleoconch whorl. Spiral ribs crossed by faint transverse, largely orthocline transverse ribs numbering around 20 per half whorl on the fourth teleoconch whorl. Measurements. MGUH 33313 is 1.8 mm high and 0.5 mm wide, consisting of part of the protoconch and the first 5 ½ teleoconch whorls. Remarks. Clathrobaculus sp. is the youngest taxon so far, which with any degree of confidence may be assigned to this genus (see Kaim 2004: p. 140). There are, though, two taxa from the slightly younger middle Danian deposits of Denmark that may belong here, but at present their protoconchs are unknown (Lauridsen & Schnetler 2014: pp. 109 – 110). The Danish specimen strongly resembles the Early Cretaceous Clathrobaculus sp. 1 sensu Kaim (2004) from Poland, but differs by the higher number of transverse ribs, which appear weaker and by the later appearance of spiral rib 2.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A4FF412B9CC662FD31F875.taxon	materials_examined	Superfamily Diaphanoidea Odhner, 1914	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A4FF412B9CC22EFBA3F976.taxon	description	Figs 35 L – P	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A4FF412B9CC22EFBA3F976.taxon	materials_examined	Material. MGUH 33314, MGUH 33315, MGUH 33316, and an additional specimen sitting together with MGUH 33315. Occurrence. The lower Danian ‘ dead layer’ exposed in the Dania Quarry, Northern Jutland. Description. Protoconch apparently conical in outline, 0.8 mm wide with smooth, convex whorls separated by deep suture. Teleoconch low spired, rotund with flattened, adpressed spire whorls; whorls nearly four times as wide as high. Last whorl evenly convex all the way to anterior canal, taking up more than 80 % of shell height. Aperture narrow, outer lip strengthened by strong varix reaching from anterior canal to adapical suture of transition between last whorl and penultimate whorl. Outer lip carrying five stronger, elongated teeth on abapical part, succeeded adapically by several more, but very weak teeth fading out completely adapically. Inner lip with moderately wide callus and two sinuses separated by a strong and furrowed ridge. Adapical parietal region carrying strong but low and ridged transverse thickening of the shell. Teleoconch sculpture consisting of around 20 moderately weak spiral grooves, which continue onto the varix. Measurements. Most complete specimen MGUH 33314 is 6.0 mm wide and more than 6.9 mm high, but lacking anterior canal and protoconch apex. It consists of the final protoconch whorl and 3 ½ teleoconch whorls. Remarks. This species differs from the Danian Ringicula pinguis Glibert, 1973 from Belgium by the much more rotund outline, adpressed whorls and furrowed lower ridge on the inner lip. It differs from the Campanian R. abundanta Kiel & Bandel, 2001 a and R. multidentata Kiel & Bandel, 2001 a from Spain by the rotund shape and from the latter also by a different tooth pattern on the outer lip. It differs from the Palaeocene R. erratica Roedel, 1937 from Northern Europe by its lower spire and more flattened spire whorls.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A4FF412B9CC320FB10FCF2.taxon	materials_examined	Genus Ringicula Deshayes in Deshayes & Milne-Edwards (1838) Type species. Auricula ringens Lamarck, 1804 (subsequent designation by Gray 1847). Eocene, France.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A5FF402B9CC1D4FADBFF15.taxon	materials_examined	Type species. Bulla cylindracea Pennant, 1777 (subsequent designation by Herrmannsen 1852). Recent, offshore Norway.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A5FF402B9CC4A1FAA1F836.taxon	description	Figs 35 U – V 2014 Cylichna sp. A Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A5FF402B9CC4A1FAA1F836.taxon	materials_examined	Material. One external mould MGUH 33320. Occurrence. Lithified top of the Maastrichtian Højerup Member in Holtug Quarry, Stevns Klint. Description. Shell deeply involute, inflated cylindrical with deep apical spire truncation. Aperture longer than last whorl; inner lip covered by callus. Columella with at least two columellar folds. Teleoconch sculpture dominated by strong growth lines, crossed by sporadic, very weakly developed spiral grooves. Measurements. The specimen MGUH 33320 is 4.3 mm high and 2.5 mm wide. Remarks. This taxon is clearly distinguished from the other cylichniid taxa from the Danish boundary layers by the two or more columellar folds; the open spire truncation and, perhaps, by the marked growth lines.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A5FF402B9CC005FBADFA79.taxon	description	Figs 35 Q – T	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A5FF402B9CC005FBADFA79.taxon	materials_examined	Diagnosis. Shell rather elongate with pointed posterior end, aperture distinctly higher than aperturally strongly convex penultimate whorl; apex hidden on adult specimens, not sharply truncated. Columellar fold very weak. Surface sculpture of fine spiral grooves. Derivation of name. Refers to the type locality Stevns Klint. Type material. Holotype MGUH 33317 is a nearly complete external mould. Paratype MGUH 33318 is an internal mould and comes from the Cerithium Limestone Member at Skeldervig. Paratype MGUH 33319 comes from the same horizon and locality as the holotype. Additional material. Six specimens with the informal sample numbers SH. 179, SH. 192. B, SR. 608 (2 specimens), SR. 680 and SO. 191, together with three uncatalogued external and internal moulds from Stevns and a single mould from Northern Jutland, which is found in the private collection of K. I. Schnetler. The Stevns material belongs to the Natural History Museum of Denmark. Type stratum and type locality. Cerithium Limestone Member of the Rødvig Formation at Rødvig, Stevns Klint. Occurrence. The Cerithium Limestone Member at Stevns Klint and the stratigrapically equivalent ‘ dead layer’ in Dania Quarry, Northern Jutland. Description. Protoconch unknown. Teleoconch moderately thick-shelled, involute, elongated in shape with weakly curved outer lip and aperturally strongly convex penultimate whorl; apex covered by last whorl on adult specimens. Aperture apically pointed, widened and rounded abapically. Width of last whorl, excluding aperture, corresponding to half the spire height. Columella with very weak plait partly hidden by callus. Sculpture, if present, consisting of weak, fine spiral grooves without pitting, grooves numbering round 30 on holotype MGUH 33317. Measurements. Largest specimen MGUH 33319 is 5.2 mm high and at least 2.8 mm wide. Remarks. As noted by Kaim & Beisel (2005: p. 61) the species of Cylichna are extremely similar and only few features may be used for differentiation on fossil specimens. Among the stratigraphically and geographically comparable taxa Cylichna stevniensis n. sp. is distinguished from the Danian Roxania raristriata (Briart & Cornet, 1887) from Belgium as figured by Glibert (1973: pl. XI: 1) by the distinctly higher aperture opening relative to the apex. It resembles somewhat Cylichna sp. 1 of Kollmann & Peel (1983) from the Danian of Greenland, but the columella is generally longer with a slightly better developed columellar plait.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A7FF422B9CC3C3FDE3FB5C.taxon	description	Figs 35 W – Y 2014 Cylichna sp. B Hansen & Surlyk: tab. 3.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A7FF422B9CC3C3FDE3FB5C.taxon	materials_examined	Material. MGUH 33321, MGUH 33322 and two incomplete moulds with informal sample numbers SR. 784. A – B and SR. 990. B – C, all from the collections of the Natural History Museum of Denmark. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig, Harvig and Stevns Fyr, Stevns Klint. Description. Shell involute, bulbous ovoid with small, shallow spire pit not showing more than latest whorl. Aperture moderately narrow, higher than penultimate whorl. No columellar folds are present on juvenile specimen MGUH 33322. Teleoconch sculpture absent. Measurements. Largest shell, MGUH 33321, 3.7 mm high and 2.4 mm wide. Remarks. This taxon is distinguished from the other Danish cylichniid boundary taxa by the bulbous shell, which may not have a columellar fold. The lack of a columellar fold may, though, be due to the undeveloped juvenile condition of the studied specimen.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A7FF422B9CC45CFECEFA6F.taxon	materials_examined	Type species. Bulla cranchii Fleming, 1828 accepted as Roxania utriculus (Brocchi, 1814) (type by monotypy). Recent, Eastern Atlantic Ocean.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A7FF4D2B9CC492FDD5FD19.taxon	description	Figs 35 Z – AE, 36 A – C	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A7FF4D2B9CC492FDD5FD19.taxon	materials_examined	Material. MGUH 33323, MGUH 33324 and two external and internal moulds found in sample SR. 394 collected for this study. Occurrence. Cerithium Limestone Member at Rødvig, between Knøsen and first ladder at Harvig, Højerup and Barmhjertigheden, Stevns Klint. Description. Protoconch unknown. Teleoconch deeply involute, with inflated, rounded whorls and a spire completely hidden or at most visible through a very narrow opening. Width of penultimate whorl corresponding to nearly 75 % of shell width. Aperture higher than last whorl, nearly twice as wide abapically as on moderately narrow apical half. Inner lip with distinct callus and a relatively well defined columella with weak columellar fold. Teleoconch sculpture consisting of more than 50 fairly distinct spiral grooves on largest specimen. Grooves not pitted. Measurements. Largest specimen, MGUH 33323, 3.6 mm wide and about 5.1 mm high. Remarks. This taxon differs from the early Paleocene Roxania raristriata (Briart & Cornet, 1887) from Belgium and the early Paleocene Roxania clausa (von Koenen, 1885) from Denmark by the rather rounded outline, wider aperture and a denser spiral striation.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A8FF4D2B9CC2B7FB99F923.taxon	description	Figs 36 D – F	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A8FF4D2B9CC2B7FB99F923.taxon	materials_examined	Material. MGUH 33325 and an external and internal mould in sample SR. 736. A collected in connection with this study. Occurrence. Lithified top of the Maastrichtian Højerup Member at Rødvig and Knøsen, Stevns Klint. Description. Shell involute, elongated cylindrical with a thick wall. Spire pit closed, only seen as a slight apical impression. Aperture long and narrow with weakly curved outer and inner lip; inner lip covered by well developed callus reaching down onto short columella. Columella smooth; columellar fold lacking or strongly effaced. Surface with spiral grooves on base. Measurements. MGUH 33325 is 6.55 mm high and 3.5 mm wide. Remarks. This taxon differs from typical species of the genus by lacking spiral grooves on the central part of the shell; by the unpitted grooves and by the rather broad callus on the parietal region. This would suggest that it should not be regarded as a true Ellipsoscapha, but as the material is limited and the taxon furthermore shows a strong resemblance to Ellipsoscapha sp. of Kollmann & Peel (1983) from the Palaeocene of Greenland, it is placed in this genus for the time being. Ellipsoscapha? sp. is distinguished from the Palaeocene Ellipsoscapha sp. of Kollmann & Peel (1983) from Greenland by the restricted spiral sculpture and possible by the presence of a shallow spire pit. It is distinguished from the above described cylichniid taxa from Denmark by the narrow cylindrical shell lacking a columellar fold and spire pit, and by the spiral grooves restricted to the base.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
BF18F633A9A8FF4D2B9CC201FB15FC4B.taxon	materials_examined	Type species. Cylichna striatella Shumard, 1861 (type by original designation). Paleocene, Texas, USA.	en	Hansen, Thomas (2019): Gastropods from the Cretaceous-Palaeogene boundary in Denmark. Zootaxa 4654 (1): 1-196, DOI: https://doi.org/10.11646/zootaxa.4654.1.1
