identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038B8782B3202B7762FAFB49FDFA530F.text	038B8782B3202B7762FAFB49FDFA530F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus LeConte 1849	<div><p>Genus Tmesiphorus LeConte, 1849</p> <p>Tmesiphorus LeConte, 1849: 75. Type species: Pselaphus carinatus Say, 1824, subsequent designation by Lucas, 1920: 643.</p> <p>Glyptosoma Motschulsky, 1851. Type species: Glyptosoma paulinae by original monotypy.</p> <p>Sintectes Westwood, 1870. Type species: Sintectes carinatus Westwood (= Tmesiphorus westwoodi Raffray), by original monotypy.</p> <p>Syntectes Jeannel, 1949: 203 (misspelling)</p> <p>Note. A detailed synonymic list was provided by Chandler (2001)</p> <p>Diagnosis. Members of this genus are easily distinguished from the other pselaphine species by the following character states: body large and stout; antennae thick; apical three segments of antennae enlarged to form variously sized club in male; maxillary palpi asymmetrical; second and third segments with penicillate spine on outer margin; fourth segment with rounded lobe on outer margin and conical shaped projection at apex; legs with protibiae arcuate and thickened at middle; apex of each tibia with dense yellow setae; tarsal claws paired; abdomen with visible tergites IV–VIII, with paratergites; tergite V longest.</p> <p>Among the Japanese pselaphine genera, Tmesiphorus is very similar in general appearance to Raphitreus Sharp, 1883, but is readily distinguished from it by the combination of larger body, segment IV of the maxillary palpi with a rounded lobe on the outer margin.</p> </div>	http://treatment.plazi.org/id/038B8782B3202B7762FAFB49FDFA530F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
038B8782B3232B7262FAFE2BFB0055AB.text	038B8782B3232B7262FAFE2BFB0055AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus costatus Weise 1877	<div><p>Tmesiphorus costatus Weise, 1877</p> <p>[Japanese name: Misuji-higekata-arizukamushi]</p> <p>(Figs 1–3)</p> <p>Tmesiphorus costatus Weise, 1877: 99 (original description); Raffray, 1908: 374 (catalogue); Raffray, 1911: 157 (catalogue); Jeannel, 1958: 118 (diagnosis); Nomura, 2001: 158 (catalogue); Löbl and Besuchet, 2004: 327 (catalogue); Nomura, 2006: 101 (taxonomic note); Shibata et al., 2013: 97 (catalogue).</p> <p>Type locality. Japan, Honshû, Yamaguchi-ken, Hagi-shi.</p> <p>Type material. Holotype: 1♂, “ Japan, Hiller // Tmesiphorus costatus W.” (MNHU).</p> <p>Material examined. (5♂, 4♀). Japan: [Honshû]: 1♂, Iwate-ken, Miyako-shi, Genbeidaira, 9 VI 2007, Y. Hi- rano leg. (NSMT); 1♂, Ishikawa-ken, Suzu-shi, Misaki-machi, Suzu-jinja, 5–20 V 2018 K. Nakata leg. (NSMT); 1♂, 1♀, Hiroshima-ken, Kamo-gun, Kurose-chô, 2 V 1988, I. Okamoto leg. (NSMT); 1♀, Hiroshima, Kure-shi, 10 IX 1994, I. Okamoto leg. (NSMT); 1♀, same data, but 2 V 1988 (NSMT); 1♂, Hiroshima-ken, Higashihiroshimashi, Saijô-chô, Shitami, 17 IV 2014, K. Maehara leg. (KUM); 1♂, same data, but 24 IV 2014 (KUM); 1♀, same data, but 18 IV 2017 (KUM).</p> <p>Diagnosis. Tmesiphorus costatus is readily distinguishable from the other Japanese congeners by a combination of strong sulcus on each elytron that extends from the median fovea to the posterior 2/3 of the elytra, and tergites IV–VI strongly carinate along their midlines. This species is smaller than the other Japanese Tmesiphorus species found in Honshû, Japan.</p> <p>Redescription. Male (Fig. 1-A). Body length 2.37–2.47 mm. Body dark brown to reddish-brown, elytra testaceous.</p> <p>Head (Figs. 2-A, B) about as long as wide, HL 0.46–0.50 mm, HW 0.44–0.49 mm, almost ovoid in dorsal view, coarsely punctate with setae; frons narrow, with longitudinal groove including fovea narrowed posteriorly to vertex; vertex with two foveae, with short longitudinal carina between vertexal foveae, shallowly depressed around vertexal foveae; eyes large, prominent; occiput rounded, with bundles of yellow setae; antennae (Figs. 1-C, D) slightly thick, antennomeres I thick, elongate, each 2.5 times longer than II; II as long as wide, each slightly wider than III; III–VI each longer than wide, narrowed at base, thickened apically; VII–VIII each transverse; VII each slightly longer than VIII; VIII each slightly wider than VII; IX–XI enlarged to form club, slightly curved inward in outer margin; IX each longer than wide, 1.3 times longer than I, 1.2 times wider than I, narrowed at base, thickened apically, longitudinally depressed with dense short setae on ventral surface; with a depression on ventral surface from basal 1/4 to apex, broadened apically, 1/3 as wide as segmental width on apical margin; X each longer than wide on inner margin, as long as wide on outer margin, longitudinally depressed with sparse setae on ventral side; XI slightly wider than IX at middle, produced outward, narrowed apically from middle, shallowly depressed with sparse setae on ventral side; maxillary palpi asymmetrical; segments II and III each with penicillate spine on lateral margin; segments IV thick, with rounded lobe on lateral margin, apical portion conical.</p> <p>Pronotum (Fig. 2-C) about as long as wide, PL 0.50–0.55 mm, PW 0.50–0.53 mm; dorsum broadly round, widest anterior to middle, narrowed posteriorly, sparsely covered with setae, with distinct median and lateral foveae at base, with prominent convexity extending from anterior margin to median fovea; elytra (Fig. 2-D) much wider than long, EL 0.63–0.65 mm, EW 0.90–0.95 mm, coarsely punctate with setae; each elytron with basal median and inner foveae pubescent, with inner and lateral longitudinal carinae, strongly sulcate from median fovea to posterior 3/4 of elytra; inner carinae extending from between two basal foveae to posterior 3/4 of elytra; lateral carinae extending from anterior to posterior margin; posterior margin with thick brush-like yellow setae; profemora thickest, with white setae on anterior face; mesofemora as thick as metafemora; protibiae thickest, arcuate and thickened at middle, as long as mesotibiae; metatibiae longest; tibiae all with dense yellow setae at apices; tarsi (Fig. 2-F) elongate; tarsomeres II 2 /3 as long as III; protarsal claws asymmetrical, anterior claws each thicker than posterior one.</p> <p>Abdomen (Fig. 2-E) large, AL 0.78 mm, AW 0.94–0.98 mm, wider than long, with sparse setae; tergite IV as long as VI; tergite V longest; tergite VII small; tergite VIII smallest, semicircular; tergites IV–VI each remarkably convex along midline; tergites IV and V each with lateral longitudinal carinae, those of tergite IV extending from anterior margin to posterior margin, those of tergite V extending from anterior to posterior margin; tergite IV with thick brush-like yellow setae along posterior margin.</p> <p>Male genitalia (Fig. 3) 0.31 mm in length, well-sclerotized, small, oval; parameres symmetric, projecting ventrally; each paramere with three setae; median lobe nearly symmetric, strongly projecting ventrally, slender; basal bulb broad, weakly narrowed apically; apical part slightly asymmetric; secondary gonopore small; endophallus elongate, circular at base.</p> <p>Female (Fig. 1-B). BL 2.22–2.39 mm; HL 0.44–0.47 mm; HW 0.41–0.46 mm; PL 0.47–0.50 mm; PW 0.44–0.50 mm; EL 0.56–0.59 mm; EW 0.80–0.88 mm; AL 0.75–0.83 mm; AW 0.88–0.94 mm. Antennae with antennomeres II–VIII more slender than in male, successively broadening towards apices; antennomeres IX–XI not modified, smaller than in male, antennal club unmodified.</p> <p>Distribution. Japan: Honshû (Iwate-ken, Ishikawa-ken, Hiroshima-ken, Yamaguchi-ken).</p> <p>Biology. The specimens were collected by flight intercept traps (FIT) or by sifting leaf litter.</p></div> 	http://treatment.plazi.org/id/038B8782B3232B7262FAFE2BFB0055AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
038B8782B3242B7E62FAFC68FC7656AF.text	038B8782B3242B7E62FAFC68FC7656AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus crassicornis Sharp 1883	<div><p>Tmesiphorus crassicornis Sharp, 1883</p> <p>[Japanese name: Higekata-arizukamushi]</p> <p>(Figs 4–6)</p> <p>Tmesiphorus crassicornis Sharp, 1883: 299 (original description); Raffray, 1908: 374 (catalogue); Raffray, 1911: 157 (catalogue); Jeannel, 1958: 118 (diagnosis); Nomura, 2001: 158 (catalogue); Löbl and Besuchet, 2004: 327 (catalogue); Terayama and Maruyama, 2007: 6 (catalogue); Shibata et al., 2013: 97 (catalogue); Maruyama et al., 2013: 32 (illustration, diagnosis).</p> <p>Type locality. Japan, Kyûshû, Nagasaki-ken, Nagasaki-shi, Suwa-temple [Suwa-jinja].</p> <p>Type material. Syntype: 1♂, “ Tmesiphorus crassicornis, Type D. S. / Suwa Temple. Japan / 31 7 71 G. Lewis (on glue board) // Sharp Coll. / 1905-313.// Type / H. T. (red-bordered disc) // Syn-type (blue-bordered disc) // Tmesiphorus / crassicornis / Sharp / det. Löbl 1976” (NHM).</p> <p>Material examined. (15♂, 20♀). Japan: [Hokkaidô]: 3♂, Sapporo-shi, Hokkaido University, 20 VI 1997, M. Ôhara &amp; K. Mizota leg. (LDN) (KUM); 1♂, 1♀, Ebetsu-shi, Nopporo, Tomambetsu, 29 V 2002, M. Maruyama leg. (LDN) (KUM); 1♂, Shakotan-gun, Raikishi-chô, 7 IX 1996, M. Ôhara leg. (LDN) (KUM); 1♂, Date-shi, Ôtakiku, 24 VIII 2001, M. Maruyama leg. (KUM); [Honshû]: 1♂, 2♀, Miyagi-ken, Sendai-shi, Aobayama, 22 IX 2001, M. Maruyama leg. (LDM) (KUM); 3♂, Yamanashi-ken, Nirasaki-shi, Hosaka-machi, Kamiimai, 6 VI 2004, M. Maruyama leg. (LDN) (KUM); 2♀, Gifu-ken, Kamioka-chô, Morimo, 27 VII 2003, K. Kinomura leg. (LDN, LDO) (KUM); 1♂, 2♀, Gifu-ken, Kuguno-chô, Bijo-kôgen, 5 VIII 2003, K. Kinomura leg. (LDN) (KUM); 1♂, Gifu-ken, Neo-mura, Imamura, 31 VIII 2003, K. Kinomura leg. (LDS) (KUM); 1♂, 1♀, Gifu-ken, Takayama-shi, Nakao, 4 VIII 2003, K. Kinomura leg. (LDM) (KUM); [Shikoku]: 2♂, 9♀, Kagawa-ken, Kotohira-chô, Atago-yama, 1 VI 2001, M. Maruyama leg. (LDS) (KUM); [Kyûshû]: 3♀, Fukuoka-ken, Tagawa-gun, Soeda-chô, Mt. Hiko-san, 11 VIII 2018, S. Inoue leg. (cIN).</p> <p>Diagnosis. This species is similar to T. princeps and T. kinomurai sp. nov. in general appearance, but is distinguished from them by the combination of broader frons, thicker antennae, strongly produced antennomere XI, sparser setae on body surface, and longitudinal carinae on tergite V extend to posterior third.</p> <p>Redescription. Male (Fig. 4-A). Body length 2.90–3.16 mm. Body almost uniformly reddish-brown.</p> <p>Head (Figs. 5-A, B) about as long as wide, HL 0.54–0.59 mm, HW 0.54–0.60 mm, almost ovoid in dorsal view, densely punctate with short setae; frons broad, with longitudinal groove including fovea narrowed posteriorly to vertex; vertex with two foveae, with short longitudinal carina between vertexal foveae, shallowly depressed around vertexal foveae; eyes large, prominent; occiput broad, with angular projection behind eyes, with tufts of short yellow setae; antennae (Figs. 4-C, D) thick, with antennomeres I thick, elongate, each 2.5 times longer than II; II as long as wide; III each slightly wider than long, thickened apically; IV–VIII each wider than long; II–VIII successively broadened towards apices; IX–XI significantly enlarged to form club, strongly curved inward in outer margin; IX each wider than long, oval, two times wider than VIII, depressed with dense setae on ventral surface, with a depression on ventral surface broadened apically from base, as wide as segmental width on apical margin; X each longer than wide, with basal margin 1.2 times wider than apical margin, narrowed apically, with glabrous area with sparse setae on ventral surface; XI each thickest at middle, significantly narrowed at base, strongly produced outward, narrowed apically from middle, depressed with sparse setae on ventral surface, with a depression on ventral surface broadened apically from base to middle, nearly circular; maxillary palpi asymmetrical; segments II and III each with penicillate spine on lateral margin, segment IV thick, with rounded lobe on lateral margin, apical portion conical.</p> <p>Pronotum (Fig. 5-C) slightly wider than long, PL 0.56–0.62 mm, PW 0.60–0.66 mm; dorsum broadly rounded, widest anterior to middle, narrowed posteriorly, covered with short setiferous setae, with distinct median and lateral foveae at base; elytra (Fig. 5-D) much wider than long, EL 0.76–0.84 mm, EW 1.08–1.21 mm, sparsely covered with short, thin setae and weakly punctate; each elytron with basal median and inner foveae pubescent, weakly sulcate from median fovea to posterior 2/3 of elytra; posterior margin with thick brush-like yellow setae; profemora thickest, with white setae on anterior face; mesofemora as thick as metafemora; protibiae thickest, arcuate and thickened at middle, as long as mesotibiae; metatibiae longer than other tibiae; tibiae all with dense yellow setae at apices; tarsi (Fig. 5-F) elongate; tarsomeres II 2 /3 as long as III; protarsal claws asymmetrical, anterior claws each thicker than posterior one.</p> <p>Abdomen (Fig. 5-E) large, bulbous, wider than long, AL 0.98–1.14 mm, AW 1.18–1.28 mm; tergite IV as long as VI; tergite V longest; tergite VII small; tergite VIII smallest and semicircular; tergites IV and V each with lateral longitudinal carinae; those of tergite IV extending from anterior margin to posterior margin; those of tergite V extending from anterior margin to posterior 1/3; tergite IV with thick brush-like yellow setae along posterior margin.</p> <p>Male genitalia (Fig. 6) 0.50 mm in length, well-sclerotized, oval; parameres symmetric, projecting ventrally; each paramere with four setae at apex; median lobe asymmetric, right part longer than left in ventral view, projecting ventrally; basal bulb broad, weakly narrowed apically; secondary gonopore large; endophallus elongate and bifurcate at base, its right projection longer than left.</p> <p>Female (Fig. 4-B). BL 2.82–3.18 mm; HL 0.54–0.58 mm; HW 0.54–0.62 mm; PL 0.54–0.64 mm; PW 0.60– 0.70 mm; EL 0.76–0.84 mm; EW 1.13–1.24 mm; AL 0.98–1.14 mm; AW 1.22–1.38 mm. Antennae moniliform; antennomeres II–VIII longer than in male, successively wider towards apices; antennomeres IX–XI not modified, smaller than in male, antennal club indistinct.</p> <p>Distribution. Japan: Hokkaidô, Honshû, Shikoku, Kyûshû.</p> <p>Biology. This species has been collected from around nest entrances and trails of Lasius (Dendrolasius) spp. (Maruyama et al. 2013). Sharp (1883) stated that this species was collected with a "black ant." This probably indicates Lasius (Dendrolasius) spp. of which all are dark brown in color.</p> </div>	http://treatment.plazi.org/id/038B8782B3242B7E62FAFC68FC7656AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
038B8782B3292B7D62FAFA94FC2A505F.text	038B8782B3292B7D62FAFA94FC2A505F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus princeps Sharp 1883	<div><p>Tmesiphorus princeps Sharp, 1883</p> <p>[Japanese name: Oo-higekata-arizukamushi]</p> <p>(Figs 7–9)</p> <p>Tmesiphorus princeps Sharp, 1883: 299 (original description); Raffray, 1908: 374 (catalogue); Raffray, 1911: 157 (catalogue); Jeannel, 1958: 119 (diagnosis); Nomura, 2001: 158 (catalogue); Löbl &amp; Besuchet, 2004: 327 (catalogue); Terayama &amp; Maruyama, 2007: 6 (catalogue); Shibata et al., 2013: 97 (catalogue); Maruyama et al., 2013:33 (illustration, diagnosis).</p> <p>Type locality. Japan, Honshû, Niigata, Futai.</p> <p>Type material. Holotype: 1♀, “ ♀ / Tmesiphorus / princeps / Type D.S. / 20.9.81 / Lewis. (on glue board) // Type / H. T. (red-bordered disc) // Japan / G. Lewis / 1910–220” (NHM).</p> <p>Material examined. (11♂, 14♀). Japan: [Hokkaidô]: 3♀, Furano-shi, Fukiage-onsen, 11 VI 2008, Y. Kida leg. (KUM); [Honshû]: 1♂, Kanagawa-ken, Isehara-shi, Hinata-rindô, 8 IV 2001, K. Toyoda leg. (cAR); 1♂, same data, but S. Arai leg. (LU) (cAR); 2♀, Ishikawa-ken, Hakusan-shi, Mt. Kuchisanpoudake, 4 VI 2016, K. Nakata leg. (FH) (KUM); 2♀, same data, but 11 X 2015 (CJ); 1♀, Ishikawa-ken, Hakusan-shi, Mt. Eboshi-yama, 17 IX 2016, K. Nakata leg. (PS) (KUM); 1♀, same data, but 4 VI 2017 (FH); Ishikawa-ken, Kanazawa-shi, Tomurobessho, Mt. To- muro-yama, 22 V 2016, K. Nakata leg. (PF) (KUM); 4♂, Gifu-ken, Gifu-shi, Nagaraobusa, 7 VI 2003, K. Kinomura leg. (LM) (KUM); 2♂, same data, but 10 VII 2005 (LM) (KUM); 1♂, 1♀, same data, but 25 VI 2005 (LM) (KUM); 1♂, Gifu-ken, Takayama-shi, Kamitakara-chô, Nakao, 4 VIII 2003, K. Kinomura leg. (LT) (KUM); 1♀, same data, but 11 VIII 2005 (AJ) (KUM); 2♀, Gifu-ken, Takayama-shi, Shoukawa-chô, Isshiki, 5 V 2013, K. Kinomura leg. (AJ) (KUM); 1♂, same data, but Sandani, 28 VIII 2005, K. Kinomura leg. (AJ) (KUM); 1♀, [Kyûshû]: Nagasakiken, Tsushima-shi, Kamiagata-chô, Sasuna, 24 IX 2015, Y. Hisasue leg. (CJ) (cIN).</p> <p>Diagnosis. Tmesiphorus princeps is similar in general appearance to T. crassicornis and T. kinomurai sp. nov. Tmesiphorus princeps is distinguished from T. crassicornis by the combination of long setae covering body surface, slightly more slender antennae, elongate antennomeres IX, and shorter longitudinal carinae of tergite V. It may be separated from T. kinomurai sp. nov. by the combination of stouter body, thicker antennae, enlarged antennomeres IX–XI that form a club, bulb-like abdomen, and shorter longitudinal carinae on tergite V. Tmesiphorus princeps shows local variation in lengths of tergal carinae. The specimens from Hokkaidô tend to have shorter carinae on tergite V.</p> <p>Redescription. Male (Fig. 7-A). Body length 2.96–3.16 mm. Body reddish-brown.</p> <p>Head (Figs. 8-A, B) slightly longer than wide, HL 0.58–0.60 mm, HW 0.54–0.56 mm, almost ovoid in dorsal view, densely punctate with long setae; frons narrowed, with longitudinal groove including small fovea narrowed posteriorly to vertex; vertex with two small foveae, with short longitudinal carina between foveae, shallowly de- pressed around vertexal foveae; eyes large, prominent; occiput rounded, with dense tufts of yellow setae; antennae (Figs. 7-C, D) thick, with antennomeres I thick, elongate, each 2.5 times longer than II; II each slightly longer than III; III–VIII each small, successively broadened towards apices; VII, VIII each wider than long; IX–XI significantly enlarged to form club, curved inward in outer margin; IX each longer than wide, 1.5 times longer than X, slightly thicker than X, shallowly depressed longitudinally with dense setae on ventral surface; with a depression on ventral surface from base to apex, broadened apically, half as wide as segmental width on apical margin; X each as long as wide, nearly quadrate, slightly depressed, with sparse setae on ventral surface; XI each thickest at middle, narrowed at base, narrowed apically from middle, strongly produced outward, slightly depressed with sparse setae on ventral surface; with a depression broadened apically from the base to middle, nearly circular; maxillary palpi asymmetri- cal; segments II and III each with slightly longer penicillate spine on lateral margin; segments IV small, thick, with rounded lobe on lateral margin, apical portion conical.</p> <p>Pronotum (Fig. 8-C) about as long as wide, PL 0.58–0.64 mm, PW 0.60–0.64 mm; dorsum broadly rounded, widest anterior to middle, narrowed posteriorly, densely punctate, with long setae, with distinct median and lateral foveae at base; elytra (Fig. 8-D) much wider than long, EL 0.76–0.84 mm, EW 1.10–1.18 mm, sparsely covered long setae, weakly punctate; each elytron with basal median and inner foveae pubescent, weakly sulcate from median fovea to posterior 2/3 of elytra; posterior margin with thick brush-like yellow setae; profemora thickest, with white setae on anterior face; mesofemora as thick as metafemora; tibiae slender; protibiae thickest, arcuate, thickened at middle, as long as mesotibiae, metatibiae longest; tibiae all with dense yellow setae at apices; tarsi (Fig. 8-F) elongate; tarsomeres II 2 /3 as long as III, protarsal claws asymmetrical, anterior claws each thicker than posterior one.</p> <p>Abdomen (Fig. 8-E) large, bulbous, wider than long, AL 1.00– 1.12 mm, AW 1.20–1.30 mm, covered with long setae; tergite IV as long as tergite VI; tergite V longest; tergite VII small; tergite VIII smallest, semicircular; tergites IV and V with lateral carinae; those of tergite IV extending from anterior margin to posterior margin; those of tergite V short, 1/3 times as long as tergal IV carinae; tergite IV with thick brush-like yellow setae along posterior margin.</p> <p>Male genitalia (Fig. 9) 0.38 mm in length, well-sclerotized, oval; parameres symmetric, projecting ventrally; each paramere with four setae at apex; median lobe asymmetric, strongly projecting ventrally; basal bulb broad, weakly narrowed apically; apex slightly asymmetrical, narrowing, projecting ventrally; secondary gonopore large; endophallus short, curved laterally.</p> <p>Female (Fig. 7-B). BL 2.92–3.22 mm; HL 0.55–0.60 mm; HW 0.54–0.58 mm; PL 0.59–0.62 mm; PW 0.58–0.64 mm; EL 0.76–0.84 mm; EW 1.06–1.20 mm; AL 0.98–1.20 mm; AW 1.19–1.30 mm. Antennae with antennomeres II–VIII slightly longer than in male, more slender than in male, successively wider towards apices; segments IX–XI not modified, smaller than in male, antennal club indistinct.</p> <p>Distribution. Japan: Hokkaidô, Honshû, Tsushima.</p> <p>Biology. This species has been collected from nests of Formica spp., Camponotus spp., Lasius (Chthonolasius) spp. (Maruyama et al. 2013), Manica yessensis (new host record), and Polyergus samurai (new host record). Sharp (1883) wrote that this species was collected with a "black ant". This is probably either a Formica sp. or Camponotus sp., all of which have a dark brown body color among the host ants recorded.</p> </div>	http://treatment.plazi.org/id/038B8782B3292B7D62FAFA94FC2A505F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
038B8782B32A2B7862FAF8C1FE265153.text	038B8782B32A2B7862FAF8C1FE265153.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus kinomurai Inoue & Maruyama & Nomura 2019	<div><p>Tmesiphorus kinomurai sp. nov.</p> <p>[Japanese name: Hoso-higekata-arizukamushi]</p> <p>(Figs 10–12)</p> <p>Tmesiphorus sp. 1, Maruyama et al., 2013: 33 (illustration, diagnosis). Type material. Holotype (placed in NSMT): ♂, Japan: [Honshû], Gifu-ken, Seki-shi, Senbiki, 20 IV 2003, K. Kinomura leg. (AJ), NSMT-I-C- 200334. Paratypes: 1♂, Gifu-ken, Gifu-shi, Mt. Kinka-zan (Alt. 50m), 18 IV 2013, K. Kinomura leg. (AJ) (NSMT); 2♀, same data as holotype (KUM); 1♀, Gifu-ken, Gifu-shi, Nagaraobusa, 7 VI 2003, K. Kinomura leg. (LM) (KUM); 2♀, Gifu-ken, Gifu-shi, Mt. Kinka-zan, 22 IV 2006, K. Kinomura leg. (AJ) (KUM); 1♀, Gifu-ken, Takayama-shi, Shoukawa-chô, Isshiki, 5 V 2013, K. Kinomura leg. (AJ) (KUM).</p> <p>Diagnosis. Tmesiphorus kinomurai sp. nov. is very similar in general appearance to T. princeps. However, T. kinomurai is readily distinguished from T. princeps by the following states: head longer than wide, weakly shining; eyes small; postocular margin longer than eyes; antennae slender; antennomeres IX–XI each longer than wide; XI lanceolate; pronotum slender; tergite V with longitudinal carinae extending to posterior half.</p> <p>Description. Male (Fig. 10-A). Body length 2.62–2.71 mm. Body dark brown to reddish-brown.</p> <p>Head (Figs. 11-A, B) longer than wide, HL 0.5–0.53 mm, HW 0.43–0.48 mm, slender, almost ovoid in dorsal view, with long setae; frons narrowed, with longitudinal groove including small fovea, densely punctate, narrowed posteriorly to vertex; vertex with two small foveae, with short longitudinal carina between vertexal foveae, weakly shining, slightly depressed around the two vertexal foveae; eyes small, prominent; occiput rounded, with tufts of yellow setae; postocular margin two times as long as eyes; antennae (Figs. 10-C, D) more slender, with antennomeres I thick, elongate, each 2.5 times longer than II, 1.5 times wider than II; II each as long as wide; III each slightly as long as wide, narrowed at the base, broadened apically, IV–VI each as long as wide; VII–VIII each transverse; III–VIII successively broadened towards apices; IX–XI enlarged to form club, slightly curved inward in outer margin; IX each longer than wide, slightly wider than I, shallowly depressed longitudinally with dense setae on ventral surface; with a depression from base to apex, 2/3 as wide as segmental width on apical margin; X each longer than wide, slightly slender than IX, with glabrous area with sparse setae on ventral surface; XI each longer than wide, about as thick as IX, slightly produced outward at middle, narrowed apically from middle to apices, lanceolate; shallowly depressed longitudinally from base to middle with sparse setae on ventral surface; maxillary palpi asymmetrical; segments II and III each with slightly long penicillate spine on outside; segment IV thick, with rounded lobe on lateral margin, apical portion conical.</p> <p>Pronotum (Fig. 11-C) slightly longer than wide, PL 0.56 mm PW 0.53–0.55 mm; dorsum broadly rounded, widest to middle, narrowed posteriorly, weakly shining, sparsely covered with long setae, with distinct median and lateral foveae at posterior base; elytra (Fig. 11-D) much wider than long, EL 0.71–0.73 mm, EW 1.00– 1.11 mm, sparsely covered with long setae; each elytron with basal median and inner foveae pubescent, weakly sulcate from median fovea to posterior 2/3 of elytra, weakly carinate from between two foveae to posterior 1/2 of elytra, slightly carinate from outside of median fovea to posterior margin, with thick brush-like of yellow setae along posterior margin; profemora thickest, with white setae at anterior face; mesofemora, metafemora slender; tibiae slender; protibiae thickest, arcuate and thickened at middle, as long as mesotibiae; metatibiae longest; tibiae all with dense yellow setae at apices; tarsi (Fig. 11-F) all slightly slender, elongate; tarsomeres II 2 /3 as long as III; tarsal claws all asymmetrical; protarsal anterior claws each thicker than posterior one; mesotarsal and metatarsal posterior claws slightly longer than anterior claws.</p> <p>Abdomen (Fig. 11-E) large, wider than long, AL 0.82–0.93 mm, AW 1.08–1.18 mm, sparsely covered with long setae, tergite IV as long as tergite VI, tergite V longest, declined toward apex, tergite VII small, tergite VIII smallest, semicircular; tergites IV and V with lateral carinae, those of tergite IV extending from anterior margin to posterior margin, those of tergite V extending from anterior margin to posterior 1/2; tergite IV with thick brush-like of yellow setae along posterior margin.</p> <p>Male genitalia (Fig. 12) 0.36 mm in length, well-sclerotized, more like oval; parameres symmetric, strongly projecting ventrally; median lobe asymmetric, projecting ventrally, thick, curved downwardly from middle to apex; basal bulb broad; apex slightly asymmetrical; secondary gonopore small; endophallus elongate, circular at base, weakly curved dorsally at middle.</p> <p>Female (Fig. 10-B). BL 2.61–2.83 mm: HL 0.50–0.55 mm; HW 0.45–0.49 mm; PL 0.53–0.60 mm; PW 0.53– 0.57 mm; EL 0.69–0.73 mm; EW 1.01–1.10 mm; AL 0.90–0.93 mm; AW 1.05–1.20 mm. Antennae with antennomeres II–VIII slightly longer than in male, successively broadened towards apices; antennomeres IX–XI not modified, smaller than in male, antennal club indistinct.</p> <p>Etymology. This species is named after the collector of the holotype, Mr. Kyôichi Kinomura, who is the super- visor of the first author.</p> <p>Distribution. Japan: Honshû (Gifu-ken).</p> <p>Biology. This species has been collected from nests of Aphaenogaster japonica, A. famelica, and Lasius meridionalis (Maruyama et al. 2013).</p> </div>	http://treatment.plazi.org/id/038B8782B32A2B7862FAF8C1FE265153	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
038B8782B32E2B6762FAFF7DFAF250C3.text	038B8782B32E2B6762FAFF7DFAF250C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmesiphorus okinawensis Inoue & Maruyama & Nomura 2019	<div><p>Tmesiphorus okinawensis sp. nov.</p> <p>[Japanese name: Okinawa-higekata-arizukamushi]</p> <p>(Figs 13–15)</p> <p>Tmesiphorus sp. 2, Maruyama et al., 2013: 34 (illustration, diagnosis).</p> <p>Type material. Holotype (placed in NSMT): ♂, Japan: [Ryûkyû]: Kume-jima, Mt. Daruma–Mt. Ôtake, 15 IV 1995, N. Takahashi leg., NSMT-I-C- 200335. Paratypes: 1♂, Japan: [Ryûkyû]: Kume-jima, Gushikawa-son, Mt. Ôtake, 4 XII 1997, S. Inada leg. (NSMT); 1♀, Okinawa-jima, Ôkuni-rindô, 23 IV 1986, S. Nomura leg. (NSMT); 2♀, Okinawa-jima, Kunigami-son, Yona, 15 II 2010, T. Shimada leg. (CM) (KUM); 1♀, Okinawa-jima, Nago-shi, Mt. Nago-take, 7 IX 2003, S. Arai leg. (NSMT).</p> <p>Diagnosis. Tmesiphorus okinawensis sp. nov. is readily distinguished from the other Japanese species by the combination of stouter body, significantly longer setae on body surface, weakly shining body, and absence of tergal carinae.</p> <p>Description. Male (Fig. 13-A). Body length 2.63 mm. Body reddish-brown; large and stout; dorsal surface polished and weakly shining.</p> <p>Head (Figs. 14-A, B) wider than long, HL 0.43–0.45 mm, HW 0.54–0.55 mm, transverse, weakly shining, covered with long setae; frons short longitudinally, weakly punctate, with short longitudinal groove including large fovea; vertex with two foveae, with short longitudinal carina between vertexal foveae; eyes large, prominent; occiput rounded gently, with tufts of yellow setae; postocular margin slightly longer than eyes, roundly narrowed towards head base; antennae (Figs. 13-C, D) slightly slender, with antennomeres I thick and elongate, two times as long as II; III–V each longer than wide; VI–VIII each transverse, successively broadening towards apices; IX–XI significantly enlarged to form club, curved inward in outer margin; IX each longer than wide, 1.5 times as long as X, thickened apically, shallowly depressed longitudinally from basal 1/4 to apical margin with dese setae on ventral surface, with a depression on ventral surface broad, 3/4 as wide as segmental width; X each about as long as wide, quadrate, about as wide as base of IX, with small glabrous area on ventral surface with sparse setae; XI each about as wide as IX, slightly longer than IX, produced outward at middle, nearly lanceolate, thickened apically from base to middle, narrowed apically from middle to apex, shallowly depressed with sparse setae on ventral surface, with a depression on ventral surface broadened from base to middle, nearly circular; maxillary palpi asymmetrical, segments II and III with penicillate spine on lateral margin; IV thick, with rounded lobe on lateral margin; apical portion conical.</p> <p>Pronotum (Fig. 14-C) about as long as wide, PL 0.55–0.58 mm, PW 0.55–0.56 mm; dorsum broadly rounded, widest posterior to middle, polished and weakly shining, densely covered with long setae, with distinct median and lateral foveae at posterior base; elytra (Fig. 14-D) much wider than long, EL 0.78 mm, EW 1.10–1.15 mm, trapezoidal, covered with long setae; each elytron with basal median and inner foveae pubescent, slightly sulcate from median fovea to elytral midpoint; posterior margin with brush-like of yellow setae; profemora as thick as mesofemora, metafemora most slender; tibiae all elongate, with dense yellow setae at apices; protibiae with white setae on anterior face; mesotibiae longer than protibiae; metatibiae longest; tarsi (Fig. 14-F) all elongate; tarsomeres II 2 /3 as long as III; protarsal claws asymmetrical; anterior claws each thicker than posterior one.</p> <p>Abdomen (Fig. 14-E) wider than long, AL 0.83–0.88 mm, AW 1.15– 1.13 mm, enlarged, surface polished, weakly shining, with long setae, narrowed posteriorly; tergite IV as long as VI; tergite V longest; tergite IV with brush-like of yellow setae along posterior margin; tergites IV and V lacking carinae.</p> <p>Male genitalia (Fig. 15) 0.35 mm in length, well-sclerotized, small, oval, slightly slender; parameres symmetric, projecting ventrally; each paramere with three setae; median lobe asymmetric, right part longer than left in ventral view, projecting ventrally; projection slightly thick; basal bulb broad, narrowed apically, secondary gonopore small; endophallus sclerotized, short, enlarged at base, curved dorsally in lateral view.</p> <p>Female (Fig. 14-B). BL 2.63–2.81mm; HL 0.43–0.46 mm; HW 0.54–0.59 mm; PL 0.54–0.59 mm; PW 0.55– 0.60 mm; EL 0.73–0.78 mm; EW 1.08–1.18 mm; AL 0.94–1.00 mm; AW 1.13–1.20 mm. Antennae slender than in male, with antennomeres XI, II–VIII each longer than wide, successively broadened towards apices; antennomeres IX–XI not modified, smaller than in male, antennal club indistinct.</p> <p>Etymology. The new species is named after its type locality.</p> <p>Distribution. Japan: [Ryûkyû]: Okinawa-jima, Kume-jima.</p> <p>Biology. This species has been collected from nests of Camponotus monju (Maruyama et al. 2013).</p> </div>	http://treatment.plazi.org/id/038B8782B32E2B6762FAFF7DFAF250C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Shota;Maruyama, Munetoshi;Nomura, Shûhei	Inoue, Shota, Maruyama, Munetoshi, Nomura, Shûhei (2019): Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan. Zootaxa 4646 (1): 67-86, DOI: https://doi.org/10.11646/zootaxa.4646.1.4
