taxonID	type	description	language	source
0389886AE949FFBF13991174FBA22562.taxon	materials_examined	Material examined. 2 males, Pernambuco, Brazil: Furna do Morcego cave, 8 ° 34 ’ 14.1 ” S; 37 ° 22 ’ 55.6 ” W, 556 m a. s. l., 09 - V- 2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR 008).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBF13991174FBA22562.taxon	diagnosis	Diagnosis. According to Mahnert (2001) Pseudochthonius biseriatus, it can be easily distinguished from the other species of the genus by the carapace reticule, as long as broad, absent eyes, anterior margin with a prominent dentate epistome, 16 strong setae, one preocular seta present on each side (Mahnert 2001; Fig. 13); Chelicerae with five setae, fixed finger with 9 – 11 acute teeth, movable finger with 8 – 10 teeth, absent spinneret tooth-like in male, serrula exterior with 20 blades, rallum nine setae, first only shorts dentate (see Mahnert 2001; Fig. 14); Tergites whitish-yellow, tergites III – IV with round medial spots, chaetotaxy: 2: 2: 4: 4 - 6: 6: 6: 6: 6: 4: 4 (2 ST): 2; Pedipalps, femur 6.4 – 6.6 ×, patella 2.2 – 2.5 ×, hand 2.3 – 2.4 ×, chela 7.5 – 8.0 ×, finger 2.3 ×, fixed finger with 37 – 41 pointed teeth arranged like offset saw teeth and therefore giving the impression of heterodonty, trichobothrium ist much near to esb than to est, and one sensillum distal to sb on movable finger (see Mahnert 2001; Figs. 15 – 16); Leg I: femur 6.1 – 6.2 ×, 4.2 – 4.5 ×, tibia 4.9 – 5.3 ×, tarsus 10.4 – 11.3 ×; Leg IV: femur + patella 3.1 – 3.4 ×, tibia 5.0 – 5.2 ×, metatarsus 3.5 – 3.8 ×, tarsus 10.6 – 11.9 × (one basal tactile seta).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBF13991174FBA22562.taxon	discussion	Remarks. Measurements and proportion of pedipalps, distribution of trichobothria, and some other taxonomic characters correspond to the description of P. biseriatus from Minas Gerais, Brazil (Mahnert 2001; Figs. 13 – 16). The specimens of the cave from Pernambuco (Brazil) slightly differ from the other specimens in the body length (1.649 – 1.656), smaller than those described by Mahnert (2001), similarly in the number of teeth on the fingers of the chela, 35 teeth on the fixed finger and 31 on the movable finger, however, these differences can be attributed to intraspecific variations by the geographical distribution of the species, since all the diagnostic characters correspond to the species P. biseriatus.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBF13991174FBA22562.taxon	description	Measurements (mm). Body length: 1.649 – 1.656. Carapace: 0.523 – 0.526 / 0.438 – 0.456. Pedipalps: femur: 0.426 – 0.459 / 0.114 – 0.116, patella 0.237 – 0.241 / 0.101 – 0.103, chela: 0.792 – 0.796 / 0.157 – 0.159, movable finger: 0.513 / 0.030. Chelicera: 0.149 / 0.090, movable finger length: 0.125. Leg I: femur: 0.461 – 0.463 / 0.070 – 0.080, patella: 0.273 – 0.281 / 0.060 – 0.080, tibia: 0.251 – 0.273 / 0.050 – 0.070, tarsus: 0.477 – 0.479 / 0.050 – 0.060. Leg IV: femur + patella: 0.689 – 0.690 / 0.213 – 0.232, tibia: 0.441 – 0 – 443 / 0.070 – 0.090, metatarsus: 0.25 – 0.27 / 0.070 – 0.090, tarsus: 0.493 – 0.495 / 0.050 – 0.070.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBF13991174FBA22562.taxon	distribution	Distribution: Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBE13991458FB9C2054.taxon	materials_examined	Material examined. 1 male, Pernambuco, Brazil: Furna do Morcego cave, 8 ° 34 ’ 14.1 ” S; 37 ° 22 ’ 55.6 ” W, 556 m a. s. l., 09 - V- 2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR 009).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBE13991458FB9C2054.taxon	diagnosis	Diagnosis. According to Chamberlin & Chamberlin (1945) specimens of Cryptoditha can be recognized by the marginal teeth of chela prominent, acute, and distinctly spaced; sub-equally developed on both fingers; tending to become contiguous basally on the fingers (Chamberlin & Chamberlin 1945; Fig. 6). Carapace smooth, weakly emarginate, and provided with a dentate, epistomal process; with four large eyes. Vestitural setae sparse but robust; tergites uniseriate with about six marginal setae per segment. Coxal spines typical, with four to five each on coxae I and II, intercoxal tubercle absent. Trichobothrium et subterminal, est and it both distinctly distal of median, but well-spaced from et; trichobothrium ist, esb, and eb, forming an oblique basal series. Trichobothrium esb closer to ist than to eb. Trichobothrium st submedial between sb and t (Chamberlin & Chamberlin 1945; Fig. 6). The species Cryptoditha elegans (Beier, 1931) was described initially from Minas Gerais (Passa Quatro), subsequently from Santa Catarina (Beier 1931 a, 1974), and later reported for the State of Mina Gerais (Mahnert 2001). The specimen of the cave from Pernambuco (Brazil) presents some taxonomic characters corresponding to the description of the C. elegans made by Beier (1931 a) and Chamberlin & Chamberlin (1945), such as proportion of pedipalps, and distribution of trichobothria, similarly, to the proportions of the specimens reported by Mahnert (2001). Therefore, due to the lack of specimens from the type locality of the species (see Mahnert 2001), which would allow a reexamination of the type specimens, likewise, only having one individual, it is decided to assign the Pernambuco specimen as Cryptoditha aff. elegans.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBE13991458FB9C2054.taxon	description	Measurements (mm). Body length: 1.315. Carapace: 0.423. Pedipalps: femur: 0.375 / 0.149, patella: 0.186 / 0.086, chela: 0.626 / 0.127, movable finger: 0.408. Chelicera: 0.273 / 0.139, movable finger length: 0.153. Leg I: femur: 0.463 / 0.080, patella: 0.291 / 0.070, tibia: 0.261 / 0.070, tarsus: 0.452 / 0.050. Leg IV: femur + patella: 0.581 / 0.160, tibia: 0.403 / 0.080, metatarsus: 0.223 / 0.060, tarsus: 0.413 / 0.060.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBE13991458FB9C2054.taxon	distribution	Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE948FFBE13991792FF6324D4.taxon	materials_examined	Material examined. 1 female, Pernambuco, Brazil: Furna do Morcego cave, 8 ° 34 ’ 14.1 ” S; 37 ° 22 ’ 55.6 ” W, 556 m a. s. l., 09 - V- 2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR 010).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE948FFBE13991792FF6324D4.taxon	diagnosis	Diagnosis. According to Ellingsen (1905) and Beier (1932 b) can be easily distinguished from the other species of the genus Parachernes Chamberlin (1931) by the carapace as long as broad 1.14 ×, gradually narrowing forwards, the short frontal margin straightened. Metazone distinctly granulated, yet a little glossy before the first furrow. Two distinct transverse furrows, the anterior one about in the middle, the posterior one somewhat nearer to the hinder margin than to the anterior furrow. Tergite divided (except XI), slightly granulated, al setae rather long, slightly clavate. Chaetotaxy tergal: 10: 10: 10: 8 (1): 8 (2): 6 (2): 8 (2): 8 (2): 8 (2): 10 (2): 4 (2 ST): 2. Genital region opercula with a central group of 18 short, three setae on each side, and ten marginal setae. Chelicerae, female with galea very slender and short, with some fine teeth in and near the extremity on the underside, male with about 6 terminal branches; paired spermatheca, typical of the genus. Pedipalps moderately strong, all segments smooth and glossy. The setae on the inner side of the trochanter and femur are slightly clavate, the other setae of the palps are slightly dentate in the extremity, on the fingers simple with some longer ones intermixed; trochanter and the proximal part of femur slightly shagreened; femur 2.35 ×, strongly granulated; patella 2.02 ×, more slightly, rather shagreened. chela with pedicel 3.18 ×, chela without pedicel 2.47 ×, hand round about smooth and very glossy, fingers as general smooth and glossy.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE948FFBE13991792FF6324D4.taxon	description	Measurements (mm). Body length: 2.359. Carapace: 0.771 / 0.672. Pedipalps: trochanter: 0.347 / 0.197, femur: 0.544 / 0.235, patella: 0.589 / 0.266, chela with pedicel: 1.064 / 0.384, chela without pedicel: 0.950 / 0.384, movable finger: 0.538. Chelicera: 0.277 / 0.141, movable finger length: 0.202. Leg I: femur: 0.105 / 0.127, patella: 0.317 / 0.137, tibia: 0.272 / 0.091, tarsus: 0.308 / 0.071. Leg IV: femur + patella: 0.569 / 0.158, tibia: 0.318 / 0.103, tarsus: 0.255 / 0.052.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE948FFBE13991792FF6324D4.taxon	distribution	Distribution. Brazil, Saint Vincent, and the Grenadines (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991073FB9C2565.taxon	materials_examined	Material examined. 1 female, Pernambuco, Brazil: Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 012).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991073FB9C2565.taxon	diagnosis	Diagnosis. According to the description made by Beier (1974), the specimens of Ceriochernes foliaceosetosus Beier, 1974 can be easily distinguished by the carapace reddish-brown, slightly longer than wide, without eyes and eyespots, strongly granulated, the granules of the metazone covered with an epicuticle, reticulate in the form of a rough honeycomb, both transverse furrow narrow and not very deep, but distinct, the anterior curved laterally, the posterior closer to the rear margin, with six setae on the anterior margin, and eight on the posterior margin. All tergites divided, setae clavate and broadly foliated, rounded at the tip, curved and almost sessile, partially covered by epicuticle, with midrib, pleural membrane very densely granulated. Chaetotaxy tergal: 6: 8: 8: 6: 6: 8: 8: 8: 8: 8: 8: 2. Genital region operculum with a set of 12 setae, and ten setae on posterior margin. Chelicerae with six to seven setae on hand, es very short, three blades on the rallum, Serrula with 17 blades, galea with 5 very short branches distally. Pedipalps strongly reticulated, trochanter 1.0 ×, femur 2.58 ×, patella 2.15 ×, fingers not as long as the hand without pedicel, each with about 35 – 40 marginal teeth, the accessory teeth small and inconspicuous, present on the medial side only on the distal part of the finger, trichobothrium ist midway between isb and it, est proximal of ist; trichobothrium st closest to t.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991073FB9C2565.taxon	description	Measurements (mm). Body length: 2.021. Carapace: 0.638 / 0.538. Pedipalps: trochanter: 0.344 / 0.244, femur: 0.522 / 0.204, patella: 0.569 / 0.204, chela with pedicel: 0.864 / 0.346, chela without pedicel: 0.833 / 0.346, movable finger: 0.383. Leg I: trochanter: 0.124 / 0.105, femur: 0.127 / 0.093, patella: 0.255 / 0.093, tibia: 0.275 / 0.072, tarsus: 0.283 / 0.053. Leg IV: trochanter: 0.183 / 0.140, femur + patella: 0.488 / 0.152, tibia: 0.377 / 0.109, tarsus: 0.385 / 0.088.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991073FB9C2565.taxon	distribution	Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991383FB9C201C.taxon	materials_examined	Material examined. 1 male, Pernambuco, Brazil: Furna do Morcego cave, 8 ° 34 ’ 14.1 ” S; 37 ° 22 ’ 55.6 ” W, 556 m a. s. l., 09 - V- 2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR 011).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991383FB9C201C.taxon	diagnosis	Diagnosis. According to Heurtault (1986) and Mahnert (1994) the males of Petterchernes brasiliensis Heurtault, 1986 can be easily distinguished by the carapace without eyes and without eyespots, dark brown, metazone finely granulated, setae of the carapace and tergites strongly clavate, with two deep transverse furrows, for setae on anterior margin; tergites divided, granulate. Chaetotaxy tergal: 9: 9: 9: 10: 10: 10: 12: 12: 10: 10: 10: 2. Genital region operculum with 32 setae and ten setae on posterior margin. Chelicerae, galea with multiple branches, and three blades in the rallum. Pedipalps coarsely granulate, setae stout and enlarged, trochanter 1.67 ×, with high dorsal hump, femur 2.22 ×, abruptly enlarged, patella 2.15 ×, chela with pedicel 2.28 ×, chela without pedicel 2.09 ×, trichobothrium isb, ib, and est at same level, hand with well pronounced internal hump bearing some longer and stouter setae. Legs, tarsus IV without tactile setae, claws smooth and simple, subterminal seta smooth, curve.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991383FB9C201C.taxon	description	Measurements (mm). Body length: 1.251. Carapace: 0.554 / 0.473. Pedipalps: trochanter: 0.269 / 0.161, femur: 0.477 / 0.214, patella: 0.415 / 0.193, chela with pedicel: 0.766 / 0.335, chela without pedicel: 0.7020 / 0.335, movable finger: 0.418. Leg I: trochanter: 0.137 / 0.085, femur: 0.131 / 0.115, patella: 0.106 / 0.076, tibia: 0.127 / 0.078, tarsus: 0.202 / 0.047. Leg IV: trochanter: 0.153 / 0.121, femur + patella: 0.380 / 0.121, tibia: 0.164 / 0.086, tarsus: 0.276 / 0.058.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991383FB9C201C.taxon	distribution	Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	description	Figs. 1 – 2, 5 – 14 urn: lsid: zoobank. org: act: 3 ee 4 f 2 bc- 5242 - 443 d- 9 f 2 c- 69 baa 621 e 93 e	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	materials_examined	Material examined. Female holotype: Pernambuco, Brazil: Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 013). Paratypes: Brazil: 1 male, Pernambuco, Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 014); 1 female, Pernambuco, Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 015).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	diagnosis	Diagnosis. Geogarypus gollumi sp. nov. differs from the other species of the genus Geogarypus Chamberlin, 1930 by the trichobothrium ib basal to est, and is not distal to esb. Fixed finger chelal with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened; female, femur pedipalpal 3.5 × (male, 3.5 ×); patella 2.3 × (male, 2.2 ×); leg I and IV diplotarsate. Adults. Both females and males. Carapace and pedipalps dark brown, granulated. Carapace with a pale-yellow median zone, a dark brown longitudinal stripe, which extends from the posterior furrow to the posterior margin. Tergites with several dark brown zones on lateral margins, and the center of each. All legs light yellow. All setae short pin-shaped (Figs. 1 – 2). Chelicera (female and male), with five setae on hand, all setae acuminate, ls shorter, fixed finger with one apical tooth, followed by one small tooth, two larger teeth, and two small teeth (Fig. 6). Galea of female slender with nine small branches (Fig. 7), male galea is less slender than the female and simple. Rallum with a long and simple blade (Fig. 8); serrula exterior with 15 blades. Pedipalps slender, densely granulated, partially irregular, all setae short pin-shaped (Figs. 9 – 10). Femur 3.5 × (male 3.4 ×), only slightly curved, with a distinct pedicel; patella 2.3 × (male 2.2 ×). Chela with rounded hand, fingers only slightly curved, chela with pedicel 3.6 × (male 3.5 ×), chela without pedicel 3.4 × (male 3.3 ×) longer than broad, fixed finger chelal with seven trichobothria, isb absent, trichobothrium ib basal to est, and is not distal to esb, and movable finger with four trichobothria (Fig. 11). Fixed finger with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened (Fig. 11). Nodus ramosus at the level of st in the movable finger, and distal of ist in the fixed finger; well-developed venom apparatus on both fingers of the chela. Carapace lateral margin strongly granulated, with two pairs of horned eyes, with 21 short pin-shaped setae (male, with 23 short pin-shaped setae), four setae on anterior margin, and 12 setae on margin posterior (Fig. 5). Cucullus furrow long, extending to about the level of the first pairs of eyes (Fig. 5); median transverse furrow well defined, and posterior very conspicuous (Fig. 5). Region coxal, pedipalpal coxa sparsely granulated, approx. 36 setae, coxa I with 18 setae, Coxa II 22, coxa III 32, coxa IV approx. 38. Opisthosoma, tergites granulated. All setae pin-shaped, being longer than the setae of the carapace; and pleural membrane strongly striated; Tergal chaetotaxy of the female holotype, tergites I – XII: 16: 16: 14: 14: 12: 16: 14: 12: 12: 12: 10: 2, female paratype, tergites I – XII: 16: 16: 16: 14: 14: 14: 14: 14: 12: 12: 10: 2, male paratype tergites I – XII: 18: 18: 18: 16: 16: 16: 14: 14: 12: 12: 10: 2. Sternites, scarcely granulated, Sternal chaetotaxy of holotype female, sternites II – XII: 8: (2) 6 (2): (2) 16 (2): 16: 16: 16: 16: 16: 14: 12: 2, female paratype, sternites II – XII: 8: (2) 6 (2): (2) 16 (2): 16: 16: 16: 14: 14: 14: 12: 2, male paratype, sternites II – XII: 6: (2) 9 (2): (2) 16 (2): 16: 16: 16: 16: 16: 14: 12: 2. The genital region of the female holotype with four short setae on each side of the anterior operculum (Fig. 12), the posterior operculum with six setae (Fig. 12). Region genital of the male paratype with six setae on the anterior operculum, with lateral sacs thick and strongly pleated, and nine setae on the posterior operculum. Legs: all legs light-yellow (Figs. 1 – 2), femur 3.9 × (male 3.6 ×), patella 2.3 × (male 2.4 ×), tibia 2.7 × (male 2.5 ×), Metatarsus 2.8 × (male 2.5 ×), tarsus 3.0 × (male 3.1 ×) (Fig. 13). Leg IV: femur + patella 5.2 × (male 5.0 ×), tibia 3.8 × (male 4.2 ×), metatarsus 2.8 × (male 2.7 ×), tarsus 3.8 × (male 3.7 ×), all setae pin-shaped, tarsus without a seta tactile, subterminal setae acuminate, undivided arolium longer than simple claws.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	description	Measurements (mm): Female holotype (Paratype in parentheses): body length 1.912 (2.125); carapace 0.581 / 0.473 (0.593 / 0.476); cucullus length 0.168 (0.171). Chelicerae 0.221 / 0.126 (0.254 / 0.139); movable finger 0.165 (0.178). Pedipalps trochanter: 0.221 / 0.154 (0.224 / 0.158); femur: 0.537 / 0.154; patella 0.354 / 0.159 (0.358 / 0.162); chela with pedicel 0.914 / 0.251 (0.937 / 0.254); chela without pedicel 0.866 / 0.251 (0.893 / 0.254), movable finger 0.457 (0.481). Leg I: trochanter 0.08 / 0.09 (0.09 / 0.13), femur 0.235 / 0.061 (0.238 / 0.064), patella 0.141 / 0.063 (0.144 / 0.069), tibia 0.192 / 0.072 (0.195 / 0.085); metatarsus 0.141 / 0.031 (0.144 / 0.034); tarsus 0.094 / 0.041 (0.097 / 0.045). Leg IV: trochanter 0.199 / 0.121 (0.197 / 0.124); femur + patella: 0.416 / 0.126 (0.419 / 0.129); tibia: 0.304 / 0.081 (0.307 / 0.084); metatarsus 0.171 / 0.061 (0.174 / 0.07); tarsus 0.151 / 0.041 (0.154 / 0.041). Male paratypes: Body length 1.609. Carapace 0.451 / 0.460; cucullus length 0.153. Chelicerae 0.219 / 0.123; movable finger 0.153. Pedipalps: trochanter 0.220 / 0.149; femur 0.533 / 0.152; patella 0.350 / 0.153; chela with pedicel 0.893 / 0.247; chela without pedicel 0.863 / 0.247; movable finger 0.446. Leg I: trochanter 0.08 / 0.090; femur 0.225 / 0.062; patella 0.345 / 0.147; tibia 0.182 / 0.071; metatarsus 0.140 / 0.030; tarsus 0.086 / 0.051. Leg IV: trochanter 0.193 / 0.120; femur + patella 0.415 / 0.123; tibia 0.302 / 0.071; metatarsus 0.169 / 0.060; tarsus 0.149 / 0.040.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	etymology	Etymology. The specific epithet (gollumi) derives its name from the mythical (fictional) character Gollum, from Middle Earth, in the fantasy literary book “ The Lord of the Rings ” written by J. R. R. Tolkien, who lived in the cave.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	distribution	Distribution. Only known from the type locality (Brazil: Pernambuco, Meu Rei cave).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.taxon	discussion	Remarks. Currently, only 11 species of the genus Geogarypus Chamberlin, 1930 are known from South America (Harvey 2013). Individuals of Geogarypus gollumi sp. nov. differ from the other species of the genus by the absence of trichobothrium isb, likewise, by the trichobothrium ib basal to est, and is not distal to esb. Fixed finger chelal with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened. However, when compared to other species with the trichobothria isb absent. Geogarypus gollumi sp. nov. differs from Geogarypus bucculentus Beier, 1955 by the fixed finger with four pit-like structures with raised rims between and below eb, esb and est. Lateral margins of tergites swollen and raised. Tarsus of most legs diplotarsate, but some legs possess fused or partially fused tarsus (see Harvey, 1987; Figs. 7 – 8), and by possessing large lateral flanges on the carapace. Also, it differs from Geogarypus connatus Harvey, 1986 by the pedipalps, carapace and to a lesser extent tergites with large pustules; teeth of chelal fingers closely spaced, slightly retrorse. 5 – 7 internal accessory teeth present. Serrula exterior of chelicera with 11 – 12 (male), 11 – 13 (female) blades; galea of male simple, occasionally with 1 or 2 minute distal spinules, and female with 6 – 7 distal branches, and tarsus of all legs monotarsate; occasionally a slight suture is evident, but the segment is never articulated.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	description	Figs. 3 – 4, 15 – 24 urn: lsid: zoobank. org: act: 003 a 5 f 97 - 862 a- 4 dee- 94 d 1 - 98722 cbad 9 ff	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	materials_examined	Material examined. Male holotype: Pernambuco, Brazil: Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 016). Paratypes: Brazil: 3 males, Pernambuco, Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 017); 4 females, Pernambuco, Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 018).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	diagnosis	Diagnosis. Distribution of the trichobothria on the fixed finger chelal, being trichobothrium ist distal de est. Trichobothrium it proximal to est; ib near the level of isb. Nodus ramosus on movable finger chelal proximal to trichobothria t; the robust pedipalps, male femur 0.604 / 0.229 mm, 2.6 × (female, 0.681 / 0.251 mm, 2.7 ×), patella 0.485 / 0.209 mm, 2.3 × (female, 0.552 / 0.250 mm, 2.5 ×). Fixed finger with 39 pointed teeth, movable finger with 30 flattened teeth. Adults. Carapace (Figs. 3 – 4), and pedipalps reddish-brown, carapace in both male and female with pale posterior area, more evident in males than in females, all legs light yellow, all tergites divided (except XI), anterior tergites light-brown colors, chela pedipalpal darker reddish-brown (Figs. 3 – 4). Chelicera with five setae on hand, fixed finger with one apical tooth, and four subapical teeth, movable finger with tooth-like subapical lobe (Fig. 16), rallum with three blade, first and second long toothed (Fig. 17), gale long, with three apical branches (Fig. 18), serrula exterior with 18 blades (female, with 20 blades). Pedipalps robust (Figs. 19 – 20), in both male and female strongly granulated, trochanter 1.3 × (female, 1.2 ×) as long as broad, femur 2.6 × (female 2.7 ×) as long as broad, chela with pedicel 3.2 × (female 3.4 ×), chela without pedicel 3.0 × (female 3.2 ×), inner margin of hand in female rounded (Fig. 20). Movable finger chelal 0.518 mm (female 0.641 mm), fixed finger with 39 pointed teeth (female with 37 pointed teeth), movable finger with 30 flattened teeth (female with 32 flattened teeth) (Fig. 21). Trichobothrium ist distal de est; trichobothrium it proximal to est; ib near the level of isb; trichobothrium st proximal to the middle of the finger, sb closer to b than to st, nodus ramosus on fixed finger distal of est, and in the movable finger chelal proximal to trichobothrium t (Fig. 21), well-developed venom apparatus in both on the fixed finger and on the movable finger chelal (Fig. 21). Carapace triangular (Fig. 15), squamose sculpturing, posteriorly reticulate, cucullus shorts, with an indistinct longitudinal furrow, with medial furrow and indistinct posterior furrow (Fig. 15), four large eyes, all setae very short and acuminate, with approx. 38 setae (female approx. 36 setae), four setae on the anterior margin, one seta preocular on each side, with six to twelve setae on the posterior margin (Fig. 15). Coxal region: female and male with an apical lobe of coxa pedipalpal with two setae marginal, and three discal setae, approx. 26 setae (female approx. 28), coxa I ca. 22 setae (female ca. 26 setae), Coxa II ca. 24 setae (female ca. 28), Coxa III ca. 30 setae (female ca. 32 setae), Coxa IV ca. 34 setae (female ca. 36 setae). Opisthosoma: all tergites are divided (except XI); reticulate sculpture, all setae acuminate, pleural membrane strongly striated. Tergal chaetotaxy of holotype male, tergites I – XI: 6: 8: 8: 8: 6: 6: 4: 4: 6: 6: 4 [4 ST]: 2, females paratypes, tergites I – XI: 6: 6 – 8: 6 – 8: 8: 6: 6: 6 – 4: 6 – 4: 6: 6: 4 [4 ST]: 2, males paratypes, tergites I – XI: 6: 6: 8: 8: 6: 6 – 8: 6 – 8: 6 – 8: 6: 6 – 8: 4 [4 ST]: 2, tergite XI with four setae tactile. Sternal chaetotaxy of the male holotype, sternites II – XII: 8 (4): (1) 6 (1): (1) 6 (1): 8: 8: 8: 8: 8: 10: 10: 2, females paratypes, sternites II – XII: 6: (1) 6 (1): (1) 6 (1): 6 – 8: 6 – 8: 8: 8 – 10: 8 – 10: 10 – 12: 10: 2, males paratypes, sternites II – XII: 8 (4): (1) 6 (1): (1) 6 (1): 6: 6 – 8: 8 – 10: 8 – 10: 8 – 10: 10 – 12: 10: 2, sternites XI with two setae tactile, pleural membrane III – X with one seta. The genital region of the male holotype with eighth setae, and four discal setae on the anterior operculum (Fig. 22), the posterior operculum with six marginal setae (Fig. 22), internal genital structures, indistinctly visible (Fig. 22). Genital region of the female paratype with six central setae on the anterior operculum, and six marginal setae on the posterior operculum. Legs: all legs light-yellow (Figs. 23 – 24), Leg I (Fig. 23): trochanter 1.5 × (female 1.7 ×), femur 2.8 × (female 2.7 ×), patella 2.4 × (female 2.6 ×), tibia 2.4 × (female 2.6 ×), metatarsus 3.2 × (female 3.3 ×), tarsus 3.3 × (female 4.8 ×), Leg IV (Fig. 24): trochanter 1.1 × (female 1.2 ×), femur + patella 2.6 × (female 2.7 ×), tibia 4.0 × (female 4.1 ×), metatarsus 3.1 × (female 3.2 ×), tarsus 3.7 × (female 3.8 ×), subterminal setae smooth, arolium undivided and longer than claws (Fig. 24).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	description	Measurements (mm): Male holotype (Paratype in parentheses): body length 1.796 (1.813); carapace 0.609 / 0.486 (0.612 / 0.489); Chelicerae; movable finger. pedipalps trochanter: 0.255 / 0.191 (0.257 / 0.193); femur: 0.604 / 0.229 (0.607 / 0.230); patella 0.485 / 0.209 (0.487 / 0.210); chela with pedicel 0.929 / 0.289 (0.931 / 0.290); chela without pedicel 0.873 / 0.289 (0.853 / 0.290), movable finger 0.578 (0.581). Leg I: trochanter 0.136 / 0.129 (0.137 / 0.131), femur 0.220 / 0.077 (0.225 / 0.079), patella 0.149 / 0.061 (0.151 / 0.063), tibia 0.159 / 0.065 (0.161 / 0.067); metatarsus 0.131 / 0.041 (0.133 / 0.043); tarsus 0.134 / 0.043 (0.136 / 0.045). Leg IV: trochanter 0.231 / 0.131 (0.235 / 0.133); femur + patella: 0.843 / 0.180 (0.845 / 0.183); tibia: 0.361 / 0.091 (0.365 / 0.093); metatarsus 0.219 / 0.071 (0.221 / 0.070); tarsus 0.189 / 0.051 (0.187 / 0.053). Female paratypes: Body length 2.241 – 2.250. Carapace 0.691 – 0.693 / 0.552 – 0.553; Chelicerae; movable finger. Pedipalps: trochanter 0.226 – 0.229 / 0.210 – 0.213; femur 0.681 – 0.683 / 0.251 – 0.253; patella 0.550 – 0.553 / 0.220 – 0.223; chela with pedicel 1.091 – 1.093 / 0.321 – 0.323; chela without pedicel 1.031 – 1.033 / 0.321 – 0.323; movable finger 0.641 – 0.643. Leg I: trochanter 0.050 – 0.060 / 0.136 – 0.137; femur 0.263 – 0.265 / 0.080 – 0.090; patella 0.167 – 0.169 / 0.102 – 0.103; tibia 0.215 – 0.217 / 0.080 – 0.090; metatarsus 0.203 – 0.206 / 0.060 – 0.070; tarsus 0.195 – 0.197 / 0.041 – 0.043. Leg IV: trochanter 0.230 – 0.231 / 0.130 – 0.133; femur + patella 0.803 – 0.805 / 0.251 – 0.253; tibia 0.532 – 0.533 / 0.110 – 0.112; metatarsus 0.316 – 0.319 / 0.060 – 0.070; tarsus 0.249 – 0.251 / 0.040 – 0.050.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	etymology	Etymology. The specific epithet (smaugi) derives its name from the mythical (fictional) dragon, the last of the great dragons remaining in Middle Earth, in the fantasy literary book “ The Hobbit ” written by J. R. R. Tolkien.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	distribution	Distribution. Only known from the type locality (Brazil: Pernambuco, Meu Rei cave).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.taxon	discussion	Remarks. According to the descriptions by Beier (1931 a, 1932 b, 1959, 1964) and the taxonomic key presented by Mahnert (2001), Progarypus smaugi sp. nov. differs from other species of Progarypus Beier, 1931 as follows: it differs from the type species of the genus Progarypus ramicola (Balzan, 1887), from Paraguay, for the femur pedipalpal without pedicel, clearly separated, somewhat concave, femur pedipalps 0.720 / 0.160, patella 0.690 (0.540) / 0.201, rallum of chelicera with three denticulate blades. It differs from Progarypus longipes Beier, 1964 by each tergite with six to eight very short marginal setae; pedipalps relatively very thinned, femur pedipalpal on the apical third with an extended tactile seta, scaly sculpture in the medial area and granular in the rest. Fixed finger chelal with 45 densely spaced teeth, finger movable chelal with 35 pointed teeth in apical third, but completely flat at the base, and Progarypus marginatus Beier, 1964 from Chile by the eb - esb - isb trichobothria at the base of the fingers, ist is closer to it than to ib; male femur pedipalpal 0.950 / 0.201, 4.7 ×, patella 0.801 / 0.240, 3.3 ×. Differ from Progarypus novus Beier, 1931 from Brazil by the carapace 1.5 × as long as wide, gradually narrowing, with truncated anterior margin, and approximately net-like sculpturing. Pedipalps slender, femur 4.0 ×, as long as wide, sessile, patella 3.5 ×, chelal fingers slightly shorter than hand without pedicel 0.54 mm. It differs from Progarypus peruanus Beier, 1959 from Peru by the carapace as long as wide at the base, conically narrowed at anterior margin, densely granulated. Pedipalps almost as long as the body, femur without pedicel 4.3 ×, patella 3 ×, fingers clearly longer than hand chela with pedicel and as long as femur, tarsus IV shorter than tarsus I. It differs from Progarypus oxydactylus (Balzan, 1887) from Paraguay by the carapace clearly longer than wide, finely granulated, and reticulate at the base, pedipalps rather long and thin. Femur 4.1 × times as long as wide, weakly pedicelled, finely granulated, and striated, patella 3.2 × as long as wide, notably shorter than the femur. Similarly, it differs from the species Progarypus gracilis Mahnert, 2001, Progarypus liliae Mahnert, 2001, and Progarypus setifer Mahnert, 2001 from Brazil by the length of the femur pedipalps 0.7 - 1.0 - 1.21 mm, by the distribution of the trichobothria est, ist, it, et, the length of the fingers of the pedipalpal hand. The new species resembles Progarypus nigrimanus Mahnert, 2001, from Brazil however, it differs by the femur pedipalpal length 0.7 – 1.0 mm; at least one of trichobothrium est, ist, it, et proximal to st, finger relatively shorter, at most 1.3 ×, pedipalps bicolored, chelal hand darker than femur and patella, femur pedipalpal 4.7 – 5.3 × (length 0.86 mm), patella 3.8 – 3.9 × (length 0.78 – 0.79 mm) longer than broad, carapace anteriorly with tessellated sculpturing.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB61399117FFB9C25E8.taxon	materials_examined	Material examined. 5 males: Pernambuco, Brazil: Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 019). 4 females, Pernambuco, Meu Rei cave, 08 ° 29 ’ 14.1 ” S, 37 ° 16 ’ 48.8 ” W, 777 m a. s. l., 19 - I- 2016, Catimbau National Park, E. Barbier leg. (LECA; Ps- 020).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB61399117FFB9C25E8.taxon	diagnosis	Diagnosis. According to the description made by Mahnert (2001), specimens of Progarypus setifer Mahnert, 2001 can be identified by the combination of characters: the carapace and pedipalps reddish-brown, 1.6 × as long as broad, densely granulate, between the numerous larger granula is a clearly granulated microsculpture four setae on anterior margin, two preocular setae each side, five to six setae on posterior margin, setae small, and indistinct (see Mahnert 2001; Fig. 36). Tergal chaetotaxy, tergites I – XII: 4: 5 - 6: 5 - 6: 5 - 6: 5 - 6: 5 - 6: 6 - 8: 6 - 8: 6 - 8 [2 ST]: 8 [4 ST]: 10 [4 ST]: 2. Region genital of the male with ten marginal, and six discal setae on the operculum anterior, camera genital with 2 / 2 setae. Region genital of the female with four setae on anterior genital operculum. Chelicera, hand with 5 setae, movable finger with slightly bilobed subapical lobe, galea with 3 apical branches, serrula exterior 18 blades, flagellum 3 setae. Pedipalps: trochanter with a distinct dorsal hump, coarsely granular, femur and patella coarsely granulate, femur of the male 6.2 × (female 4.9 ×), with a short dorsal tactile seta near the base, patella 6.3 × (female 4.5 ×), chela with pedicel 5.4 × (female 3.7 ×), fixed finger chelal with approx. 40 (female 39) pointed teeth, movable finger with 39 (32 female) pointed teeth. Trichobothrium eb, esb, ib, isb at finger base, ist, est, it, et in distal half and distal to st; trichobothrium ist nearly halfway between it and est; trichobothrium st halfway between sb and t (see Mahnert 2001; Figs. 38 – 40). Leg 1: femur 4.7 × (female 3.9 ×), patella 2.6 × (female 2.3 ×), tibia 6.6 × (female 5.2 ×), metatarsus 5.7 × (female 5.0 ×), tarsus 4.4 × (female 5.0 ×). Leg IV: femur + patella 2.8 × (female 2.7 ×), tibia 5.5 × (female 4.8 ×), metatarsus 5.4 × (female 4.4 ×), with a short tactile seta near i 1 s base, tarsus 6.3 × (female 5.7 ×), subterminal seta smooth, arolia undivided and longer than claws.	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB61399117FFB9C25E8.taxon	description	Measurements (mm). Body length: 2.613 – 2.651, carapace 0.914 – 0.937 / 0.553 – 0.571 (0.931 – 0.950 / 0.611 – 0.651). Pedipalp: femur 1.21 – 1.23 / 0.170 – 0.190 (1.113 – 1.115 / 0.230 – 0.250), patella 1.230 – 1.250 / 0.200 – 0.210 (1.050 – 1.070 / 0.230 – 0.250), hand with pedicel 0.751 – 0.771 / 0.342 – 0.345 (0.801 – 0.803 / 0.461 – 0.471), finger length 1.101 – 1.116 (0.981 – 0.983), chelal length 1.830 – 1.850 (1.720 – 1.750). Legs: Leg I: femur 0.550 – 0.570 / 0.120 – 0.140 (0.470 – 0.490 / 0. l 22 – 0.123), patella 0.291 – 0.301 / 0.110 – 0.120 (0.253 – 0.256 / 0.112 – 0.113), tibia 0.487 – 0.489 / 0.071 – 0.073 (0.401 – 0.403 / 0.08), metatarsus 0.351 – 0.353 / 0.06 – 0.07 (0.281 – 0.283 / 0.06 – 0.070), tarsus 0.221 – 0.223 / 0.050 – 0.070 (0.201 – 0.203 / 0.040 – 0.050). Leg IV: femur + patella 0.881 – 0.883 / 0.310 – 0.313 (0.831 – 0.833 / 0.301 – 0.303), tibia 0.680 – 0.681 / 0.120 – 0.123 (0.626 – 0.629 / 0.131 – 0.133), metatarsus 0.411 – 0.413 / 0.080 (0.351 – 0.353 / 0.080), tarsus 0.311 – 0.313 / 0.050 (0.291 – 0.311 / 0.050).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB61399117FFB9C25E8.taxon	distribution	Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022).	en	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
