identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0389886AE949FFBF13991174FBA22562.text	0389886AE949FFBF13991174FBA22562.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudochthonius biseriatus Mahnert 2001	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pseudochthonius biseriatus Mahnert, 2001</p>
            <p>
                 Material examined.   2 males, Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.38211/lat -8.570583)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.38211&amp;materialsCitation.latitude=-8.570583">Furna do Morcego cave</a>
                 , 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR008)  . 
            </p>
            <p> Diagnosis. According to Mahnert (2001)  Pseudochthonius biseriatus , it can be easily distinguished from the other species of the genus by the carapace reticule, as long as broad, absent eyes, anterior margin with a prominent dentate epistome, 16 strong setae, one preocular seta present on each side (Mahnert 2001; Fig. 13); Chelicerae with five setae, fixed finger with 9–11 acute teeth, movable finger with 8–10 teeth, absent spinneret tooth-like in male, serrula exterior with 20 blades, rallum nine setae, first only shorts dentate (see Mahnert 2001; Fig. 14); Tergites whitish-yellow, tergites III–IV with round medial spots, chaetotaxy: 2: 2: 4: 4-6: 6: 6: 6: 6: 4: 4(2ST): 2; Pedipalps, femur 6.4–6.6×, patella 2.2–2.5×, hand 2.3–2.4×, chela 7.5–8.0×, finger 2.3×, fixed finger with 37–41 pointed teeth arranged like offset saw teeth and therefore giving the impression of heterodonty, trichobothrium ist much near to esb than to est, and one sensillum distal to sb on movable finger (see Mahnert 2001; Figs. 15–16); Leg I: femur 6.1–6.2×, 4.2–4.5×, tibia 4.9–5.3×, tarsus 10.4–11.3×; Leg IV: femur+patella 3.1–3.4×, tibia 5.0–5.2×, metatarsus 3.5–3.8×, tarsus 10.6–11.9× (one basal tactile seta). </p>
            <p> Remarks. Measurements and proportion of pedipalps, distribution of trichobothria, and some other taxonomic characters correspond to the description of  P. biseriatus from Minas Gerais, Brazil (Mahnert 2001; Figs. 13–16). The specimens of the cave from Pernambuco (Brazil) slightly differ from the other specimens in the body length (1.649 –1.656), smaller than those described by Mahnert (2001), similarly in the number of teeth on the fingers of the chela, 35 teeth on the fixed finger and 31 on the movable finger, however, these differences can be attributed to intraspecific variations by the geographical distribution of the species, since all the diagnostic characters correspond to the species  P. biseriatus . </p>
            <p>Measurements (mm). Body length: 1.649 –1.656. Carapace: 0.523–0.526/0.438–0.456. Pedipalps: femur: 0.426–0.459/0.114–0.116, patella 0.237–0.241/0.101–0.103, chela: 0.792–0.796/0.157–0.159, movable finger: 0.513 / 0.030. Chelicera: 0.149 / 0.090, movable finger length: 0.125. Leg I: femur: 0.461–0.463/0.070–0.080, patella: 0.273–0.281/0.060–0.080, tibia: 0.251–0.273/0.050–0.070, tarsus: 0.477–0.479/0.050–0.060. Leg IV: femur+patella: 0.689–0.690/0.213–0.232, tibia: 0.441–0–443/ 0.070 –0.090, metatarsus: 0.25–0.27/ 0.070 –0.090, tarsus: 0.493–0.495/0.050–0.070.</p>
            <p> Distribution: Brazil (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE949FFBF13991174FBA22562	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE949FFBE13991458FB9C2054.text	0389886AE949FFBE13991458FB9C2054.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptoditha elegans (Beier 1931)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Cryptoditha aff. elegans</p>
            <p>
                 Material examined.   1 male, Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.38211/lat -8.570583)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.38211&amp;materialsCitation.latitude=-8.570583">Furna do Morcego cave</a>
                 , 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR009)  . 
            </p>
            <p> Diagnosis. According to Chamberlin &amp; Chamberlin (1945) specimens of  Cryptoditha can be recognized by the marginal teeth of chela prominent, acute, and distinctly spaced; sub-equally developed on both fingers; tending to become contiguous basally on the fingers (Chamberlin &amp; Chamberlin 1945; Fig. 6). Carapace smooth, weakly emarginate, and provided with a dentate, epistomal process; with four large eyes. Vestitural setae sparse but robust; tergites uniseriate with about six marginal setae per segment. Coxal spines typical, with four to five each on coxae I and II, intercoxal tubercle absent. Trichobothrium et subterminal, est and it both distinctly distal of median, but well-spaced from et; trichobothrium ist, esb, and eb, forming an oblique basal series. Trichobothrium esb closer to ist than to eb. Trichobothrium st submedial between sb and t (Chamberlin &amp; Chamberlin 1945; Fig. 6). The species  Cryptoditha elegans (Beier, 1931) was described initially from Minas Gerais (Passa Quatro), subsequently from Santa Catarina (Beier 1931a, 1974), and later reported for the State of Mina Gerais (Mahnert 2001). The specimen of the cave from Pernambuco (Brazil) presents some taxonomic characters corresponding to the description of the  C. elegans made by Beier (1931a) and Chamberlin &amp; Chamberlin (1945), such as proportion of pedipalps, and distribution of trichobothria, similarly, to the proportions of the specimens reported by Mahnert (2001). Therefore, due to the lack of specimens from the type locality of the species (see Mahnert 2001), which would allow a reexamination of the type specimens, likewise, only having one individual, it is decided to assign the Pernambuco specimen as  Cryptoditha aff. elegans . </p>
            <p>Measurements (mm). Body length:1.315.Carapace:0.423.Pedipalps: femur: 0.375 / 0.149, patella: 0.186 / 0.086, chela: 0.626 / 0.127, movable finger: 0.408. Chelicera: 0.273 / 0.139, movable finger length: 0.153. Leg I: femur: 0.463 / 0.080, patella: 0.291 / 0.070, tibia: 0.261 / 0.070, tarsus: 0.452 / 0.050. Leg IV: femur+patella: 0.581 / 0.160, tibia: 0.403 / 0.080, metatarsus: 0.223 / 0.060, tarsus: 0.413 / 0.060.</p>
            <p> Distribution. Brazil (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE949FFBE13991458FB9C2054	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE948FFBE13991792FF6324D4.text	0389886AE948FFBE13991792FF6324D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parachernes (Parachernes) nitidimanus (Ellingsen 1905)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Parachernes (Parachernes) nitidimanus (Ellingsen, 1905)</p>
            <p>
                 Material examined.   1 female, Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.38211/lat -8.570583)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.38211&amp;materialsCitation.latitude=-8.570583">Furna do Morcego cave</a>
                 , 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR010)  . 
            </p>
            <p> Diagnosis. According to Ellingsen (1905) and Beier (1932b) can be easily distinguished from the other species of the genus  Parachernes Chamberlin (1931) by the carapace as long as broad 1.14×, gradually narrowing forwards, the short frontal margin straightened. Metazone distinctly granulated, yet a little glossy before the first furrow. Two distinct transverse furrows, the anterior one about in the middle, the posterior one somewhat nearer to the hinder margin than to the anterior furrow. Tergite divided (except XI), slightly granulated, al setae rather long, slightly clavate. Chaetotaxy tergal: 10: 10: 10: 8(1): 8(2): 6(2): 8(2): 8(2): 8(2): 10(2): 4(2ST):2. Genital region opercula with a central group of 18 short, three setae on each side, and ten marginal setae. Chelicerae, female with galea very slender and short, with some fine teeth in and near the extremity on the underside, male with about 6 terminal branches; paired spermatheca, typical of the genus. Pedipalps moderately strong, all segments smooth and glossy. The setae on the inner side of the trochanter and femur are slightly clavate, the other setae of the palps are slightly dentate in the extremity, on the fingers simple with some longer ones intermixed; trochanter and the proximal part of femur slightly shagreened; femur 2.35×, strongly granulated; patella 2.02×, more slightly, rather shagreened. chela with pedicel 3.18×, chela without pedicel 2.47×, hand round about smooth and very glossy, fingers as general smooth and glossy. </p>
            <p>Measurements (mm). Body length: 2.359. Carapace: 0.771 / 0.672. Pedipalps: trochanter: 0.347 / 0.197, femur: 0.544 / 0.235, patella: 0.589 / 0.266, chela with pedicel: 1.064 / 0.384, chela without pedicel: 0.950 / 0.384, movable finger: 0.538. Chelicera: 0.277 / 0.141, movable finger length:0.202. Leg I: femur: 0.105 / 0.127, patella: 0.317 / 0.137, tibia: 0.272 / 0.091, tarsus: 0.308 / 0.071. Leg IV: femur+patella: 0.569 / 0.158, tibia: 0.318 / 0.103, tarsus: 0.255 / 0.052.</p>
            <p> Distribution. Brazil, Saint Vincent, and the Grenadines (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE948FFBE13991792FF6324D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991073FB9C2565.text	0389886AE94FFFB913991073FB9C2565.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceriochernes foliaceosetosus Beier 1974	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ceriochernes foliaceosetosus Beier, 1974</p>
            <p>
                 Material examined.   1 female, Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37° 16’48.8”W, 777 m a.s.l., 19- I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-012)  . 
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            <p> Diagnosis. According to the description made by Beier (1974), the specimens of  Ceriochernes foliaceosetosus Beier, 1974 can be easily distinguished by the carapace reddish-brown, slightly longer than wide, without eyes and eyespots, strongly granulated, the granules of the metazone covered with an epicuticle, reticulate in the form of a rough honeycomb, both transverse furrow narrow and not very deep, but distinct, the anterior curved laterally, the posterior closer to the rear margin, with six setae on the anterior margin, and eight on the posterior margin. All tergites divided, setae clavate and broadly foliated, rounded at the tip, curved and almost sessile, partially covered by epicuticle, with midrib, pleural membrane very densely granulated. Chaetotaxy tergal: 6: 8: 8: 6: 6: 8: 8: 8: 8: 8: 8: 2. Genital region operculum with a set of 12 setae, and ten setae on posterior margin. Chelicerae with six to seven setae on hand, es very short, three blades on the rallum, Serrula with 17 blades, galea with 5 very short branches distally. Pedipalps strongly reticulated, trochanter 1.0×, femur 2.58×, patella 2.15×, fingers not as long as the hand without pedicel, each with about 35–40 marginal teeth, the accessory teeth small and inconspicuous, present on the medial side only on the distal part of the finger, trichobothrium ist midway between isb and it, est proximal of ist; trichobothrium st closest to t. </p>
            <p>Measurements (mm). Body length: 2.021. Carapace: 0.638 / 0.538. Pedipalps: trochanter: 0.344 / 0.244, femur: 0.522 / 0.204, patella: 0.569 / 0.204, chela with pedicel: 0.864 / 0.346, chela without pedicel: 0.833 / 0.346, movable finger: 0.383. Leg I: trochanter: 0.124 / 0.105, femur: 0.127 / 0.093, patella: 0.255 / 0.093, tibia: 0.275 / 0.072, tarsus: 0.283 / 0.053. Leg IV: trochanter: 0.183 / 0.140, femur+patella: 0.488 / 0.152, tibia: 0.377 / 0.109, tarsus: 0.385 / 0.088.</p>
            <p> Distribution. Brazil (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE94FFFB913991073FB9C2565	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFB913991383FB9C201C.text	0389886AE94FFFB913991383FB9C201C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Petterchernes brasiliensis Heurtault 1986	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Petterchernes brasiliensis Heurtault, 1986</p>
            <p>
                 Material examined.   1 male, Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.38211/lat -8.570583)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.38211&amp;materialsCitation.latitude=-8.570583">Furna do Morcego cave</a>
                 , 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR011)  . 
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            <p> Diagnosis. According to Heurtault (1986) and Mahnert (1994) the males of  Petterchernes brasiliensis Heurtault, 1986 can be easily distinguished by the carapace without eyes and without eyespots, dark brown, metazone finely granulated, setae of the carapace and tergites strongly clavate, with two deep transverse furrows, for setae on anterior margin; tergites divided, granulate. Chaetotaxy tergal: 9: 9: 9: 10: 10: 10: 12: 12: 10: 10: 10: 2. Genital region operculum with 32 setae and ten setae on posterior margin. Chelicerae, galea with multiple branches, and three blades in the rallum. Pedipalps coarsely granulate, setae stout and enlarged, trochanter 1.67×, with high dorsal hump, femur 2.22×, abruptly enlarged, patella 2.15×, chela with pedicel 2.28×, chela without pedicel 2.09×, trichobothrium isb, ib, and est at same level, hand with well pronounced internal hump bearing some longer and stouter setae. Legs, tarsus IV without tactile setae, claws smooth and simple, subterminal seta smooth, curve. </p>
            <p>Measurements (mm). Body length: 1.251. Carapace: 0.554 / 0.473. Pedipalps: trochanter: 0.269 / 0.161, femur: 0.477 / 0.214, patella: 0.415 / 0.193, chela with pedicel: 0.766 / 0.335, chela without pedicel: 0.7020 / 0.335, movable finger: 0.418. Leg I: trochanter: 0.137 / 0.085, femur: 0.131 / 0.115, patella: 0.106 / 0.076, tibia: 0.127 / 0.078, tarsus: 0.202 / 0.047. Leg IV: trochanter: 0.153 / 0.121, femur+patella: 0.380 / 0.121, tibia: 0.164 / 0.086, tarsus: 0.276 / 0.058.</p>
            <p> Distribution. Brazil (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE94FFFB913991383FB9C201C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE94FFFBA139915E8FCB424B0.text	0389886AE94FFFBA139915E8FCB424B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geogarypus gollumi Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Geogarypus gollumi n. sp.</p>
            <p>Figs. 1 – 2, 5 – 14</p>
            <p>urn:lsid:zoobank.org:act:3ee4f2bc-5242-443d-9f2c-69baa621e93e</p>
            <p>
                 Material examined.   
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Female</a>
                 holotype: Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Catimbau National Park</a>
                 , E. Barbier leg. (LECA; Ps-013).   Paratypes: Brazil: 1 male, Pernambuco,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Catimbau National Park</a>
                 , E. Barbier leg. (LECA; Ps-014);  1 female, Pernambuco, Meu Rei cave, 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-015) . 
            </p>
            <p> Diagnosis.  Geogarypus gollumi sp. nov. differs from the other species of the genus  Geogarypus Chamberlin, 1930 by the trichobothrium ib basal to est, and is not distal to esb. Fixed finger chelal with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened; female, femur pedipalpal 3.5× (male, 3.5×); patella 2.3× (male, 2.2×); leg I and IV diplotarsate. </p>
            <p>Adults. Both females and males. Carapace and pedipalps dark brown, granulated. Carapace with a pale-yellow median zone, a dark brown longitudinal stripe, which extends from the posterior furrow to the posterior margin. Tergites with several dark brown zones on lateral margins, and the center of each. All legs light yellow. All setae short pin-shaped (Figs. 1 – 2).</p>
            <p>Chelicera (female and male), with five setae on hand, all setae acuminate, ls shorter, fixed finger with one apical tooth, followed by one small tooth, two larger teeth, and two small teeth (Fig. 6). Galea of female slender with nine small branches (Fig. 7), male galea is less slender than the female and simple. Rallum with a long and simple blade (Fig. 8); serrula exterior with 15 blades.</p>
            <p>Pedipalps slender, densely granulated, partially irregular, all setae short pin-shaped (Figs. 9 – 10). Femur 3.5× (male 3.4×), only slightly curved, with a distinct pedicel; patella 2.3× (male 2.2×). Chela with rounded hand, fingers only slightly curved, chela with pedicel 3.6× (male 3.5×), chela without pedicel 3.4× (male 3.3×) longer than broad, fixed finger chelal with seven trichobothria, isb absent, trichobothrium ib basal to est, and is not distal to esb, and movable finger with four trichobothria (Fig. 11). Fixed finger with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened (Fig. 11). Nodus ramosus at the level of st in the movable finger, and distal of ist in the fixed finger; well-developed venom apparatus on both fingers of the chela.</p>
            <p>Carapace lateral margin strongly granulated, with two pairs of horned eyes, with 21 short pin-shaped setae (male, with 23 short pin-shaped setae), four setae on anterior margin, and 12 setae on margin posterior (Fig. 5). Cucullus furrow long, extending to about the level of the first pairs of eyes (Fig. 5); median transverse furrow well defined, and posterior very conspicuous (Fig. 5).</p>
            <p>Region coxal, pedipalpal coxa sparsely granulated, approx. 36 setae, coxa I with 18 setae, Coxa II 22, coxa III 32, coxa IV approx. 38.</p>
            <p>Opisthosoma, tergites granulated. All setae pin-shaped, being longer than the setae of the carapace; and pleural membrane strongly striated; Tergal chaetotaxy of the female holotype, tergites I–XII: 16: 16: 14: 14: 12: 16: 14: 12: 12: 12: 10: 2, female paratype, tergites I–XII: 16: 16: 16: 14: 14: 14: 14: 14: 12: 12: 10: 2, male paratype tergites I–XII: 18: 18: 18: 16: 16: 16: 14: 14: 12: 12: 10: 2. Sternites, scarcely granulated, Sternal chaetotaxy of holotype female, sternites II–XII: 8: (2)6(2): (2)16(2): 16: 16: 16: 16: 16: 14: 12: 2, female paratype, sternites II–XII: 8: (2)6(2): (2)16(2): 16: 16: 16: 14: 14: 14: 12: 2, male paratype, sternites II–XII: 6: (2)9(2): (2)16(2): 16: 16: 16: 16: 16: 14: 12: 2.</p>
            <p>The genital region of the female holotype with four short setae on each side of the anterior operculum (Fig. 12), the posterior operculum with six setae (Fig. 12). Region genital of the male paratype with six setae on the anterior operculum, with lateral sacs thick and strongly pleated, and nine setae on the posterior operculum.</p>
            <p>Legs: all legs light-yellow (Figs. 1 – 2), femur 3.9× (male 3.6×), patella 2.3× (male 2.4×), tibia 2.7× (male 2.5×), Metatarsus 2.8× (male 2.5×), tarsus 3.0× (male 3.1×) (Fig. 13). Leg IV: femur+patella 5.2× (male 5.0×), tibia 3.8× (male 4.2×), metatarsus 2.8× (male 2.7×), tarsus 3.8× (male 3.7×), all setae pin-shaped, tarsus without a seta tactile, subterminal setae acuminate, undivided arolium longer than simple claws.</p>
            <p>Measurements (mm): Female holotype (Paratype in parentheses): body length 1.912 (2.125); carapace 0.581 / 0.473 (0.593 / 0.476); cucullus length 0.168(0.171).Chelicerae 0.221 / 0.126 (0.254 / 0.139);movable finger0.165 (0.178). Pedipalps trochanter: 0.221 / 0.154 (0.224 / 0.158); femur: 0.537 / 0.154; patella 0.354 / 0.159 (0.358 / 0.162); chela with pedicel 0.914 / 0.251 (0.937 / 0.254); chela without pedicel 0.866 / 0.251 (0.893 / 0.254), movable finger 0.457 (0.481). Leg I: trochanter 0.08/0.09 (0.09/0.13), femur 0.235 / 0.061 (0.238 / 0.064), patella 0.141 / 0.063 (0.144 / 0.069), tibia 0.192 / 0.072 (0.195 / 0.085); metatarsus 0.141 / 0.031 (0.144 / 0.034); tarsus 0.094 / 0.041 (0.097 / 0.045). Leg IV: trochanter 0.199 / 0.121 (0.197 / 0.124); femur+patella: 0.416 / 0.126 (0.419 / 0.129); tibia: 0.304 / 0.081 (0.307 / 0.084); metatarsus 0.171 / 0.061 (0.174/0.07); tarsus 0.151 / 0.041 (0.154 / 0.041).</p>
            <p>
                 Male paratypes: Body length 1.609.  
                <a title="Search Plazi for locations around (long 0.46/lat 0.451)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=0.46&amp;materialsCitation.latitude=0.451">Carapace</a>
                 0.451 / 0.460; cucullus length 0.153. Chelicerae 0.219 / 0.123; movable finger 0.153. Pedipalps: trochanter 0.220 / 0.149; femur 0.533 / 0.152; patella 0.350 / 0.153; chela with pedicel 0.893 / 0.247; chela without pedicel 0.863 / 0.247; movable finger 0.446. Leg I: trochanter 0.08/0.090; femur 0.225 / 0.062; patella 0.345 / 0.147; tibia 0.182 / 0.071; metatarsus 0.140 / 0.030; tarsus 0.086 / 0.051. Leg IV: trochanter 0.193 / 0.120; femur+patella 0.415 / 0.123; tibia 0.302 / 0.071; metatarsus 0.169 / 0.060; tarsus 0.149 / 0.040. 
            </p>
            <p> Etymology. The specific epithet (  gollumi ) derives its name from the mythical (fictional) character Gollum, from Middle Earth, in the fantasy literary book “The Lord of the Rings” written by J.R.R. Tolkien, who lived in the cave. </p>
            <p>Distribution. Only known from the type locality (Brazil: Pernambuco, Meu Rei cave).</p>
            <p> Remarks. Currently, only 11 species of the genus  Geogarypus Chamberlin, 1930 are known from South America (Harvey 2013). Individuals of  Geogarypus gollumi sp. nov. differ from the other species of the genus by the absence of trichobothrium isb, likewise, by the trichobothrium ib basal to est, and is not distal to esb. Fixed finger chelal with 26 teeth and four larger teeth (male, with 20 teeth and four larger teeth), movable finger with 11 cusp-shaped teeth (male, with 10 cusp-shaped teeth), and the remaining teeth, basal and flattened. However, when compared to other species with the trichobothria isb absent.  Geogarypus gollumi sp. nov. differs from  Geogarypus bucculentus Beier, 1955 by the fixed finger with four pit-like structures with raised rims between and below eb, esb and est. Lateral margins of tergites swollen and raised. Tarsus of most legs diplotarsate, but some legs possess fused or partially fused tarsus (see Harvey, 1987; Figs. 7 – 8), and by possessing large lateral flanges on the carapace. Also, it differs from  Geogarypus connatus Harvey, 1986 by the pedipalps, carapace and to a lesser extent tergites with large pustules; teeth of chelal fingers closely spaced, slightly retrorse. 5–7 internal accessory teeth present. Serrula exterior of chelicera with 11–12 (male), 11–13 (female) blades; galea of male simple, occasionally with 1 or 2 minute distal spinules, and female with 6–7 distal branches, and tarsus of all legs monotarsate; occasionally a slight suture is evident, but the segment is never articulated. </p>
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	https://treatment.plazi.org/id/0389886AE94FFFBA139915E8FCB424B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB513991383FA1F200B.text	0389886AE943FFB513991383FA1F200B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geogarypus Chamberlin 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to  Geogarypus Chamberlin, 1930 species reported from Brazil </p>
            <p>1 Chela of the pedipalp with eight trichobothria; trichobothrium isb present........................................ (2)</p>
            <p> - Chela of the pedipalp with seven trichobothria; trichobothrium isb absent; femur pedipalpal 3.5× (male, 3.5×); patella 2.3× (male, 2.2×); leg I and IV diplotarsate.......................................................  G. gollumi sp. nov.</p>
            <p>2 Femur of pedipalp more than 3.0× as long as wide, chela of pedipalp 1.6–3.2×; movable finger more than 40–48 teeth, fixed finger with 27–32 teeth............................................................................... (3)</p>
            <p> - Femur of pedipalp 3.0× longer than wide; pedipalp chela 1.8× (see Mello-Leit„o, 1937; fig.1); movable finger with 38 teeth, fixed finger with 32 teeth...........................................................  G. formosus Mello-Leit „o </p>
            <p>3 Femur of pedipalp 3.1–3.5× as long as wide; patella 2.1–2.4× longer than wide; fingers 1.1–1.2× longer than hand with pedicel............................................................................................. (4)</p>
            <p> - Femur of pedipalp more than 3.5× as long as wide; patella thinner, 2.4× longer than wide (see Balzan, 1890; fig. 22); finger shorter (0.54 mm) than the hand with pedicel (0.57 mm) and longer than the chela (0.49 mm)......................................................................................................  G. cuyabanus (Balzan, 1887)</p>
            <p> 4 Femur of the pedipalp 3.0–3.3×, patella 2.4–2.5× as long as wide (see Mahnert, 1979; fig. 87); fingers (0.47–0.48 (0.49–0.55)) longer than hand (0.40–0.42 (0.45–0.49)) with pedicel; fixed finger with 41–48 teeth (distal 1–2 small, half teeth arranged in two rows), movable finger with 27–31 teeth (distal half only pointed) (see Mahnert, 1979; fig. 88b)......................................................................................................  G. amazonicus Mahnert</p>
            <p> - Femur of pedipalp 3.2×, patella 2.3× as long as wide (see Beier, 1932a; fig. 258); chela finger (0.52 mm) as long as hand (0.49) with pedicel; fixed finger with about 27 teeth, movable finger with 22 teeth (more or less clearly pointed on the base).......................................................................................  G. paraguayanus Beier</p>
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	https://treatment.plazi.org/id/0389886AE943FFB513991383FA1F200B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE943FFB61399109DFD2D2110.text	0389886AE943FFB61399109DFD2D2110.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progarypus smaugi Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Progarypus smaugi sp. nov.</p>
            <p>Figs. 3 – 4, 15 – 24</p>
            <p>urn:lsid:zoobank.org:act:003a5f97-862a-4dee-94d1-98722cbad9ff</p>
            <p>
                 Material examined.   
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Male</a>
                 holotype: Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Catimbau National Park</a>
                 , E. Barbier leg. (LECA; Ps-016).   Paratypes: Brazil: 3 males, Pernambuco,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Catimbau National Park</a>
                 , E. Barbier leg. (LECA; Ps-017);  4 females, Pernambuco, Meu Rei cave, 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-018) . 
            </p>
            <p>Diagnosis. Distribution of the trichobothria on the fixed finger chelal, being trichobothrium ist distal de est. Trichobothrium it proximal to est; ib near the level of isb. Nodus ramosus on movable finger chelal proximal to trichobothria t; the robust pedipalps, male femur 0.604 / 0.229 mm, 2.6× (female, 0.681 / 0.251 mm, 2.7×), patella 0.485 / 0.209 mm, 2.3× (female, 0.552 / 0.250 mm, 2.5×). Fixed finger with 39 pointed teeth, movable finger with 30 flattened teeth.</p>
            <p>Adults. Carapace (Figs. 3–4), and pedipalps reddish-brown, carapace in both male and female with pale posterior area, more evident in males than in females, all legs light yellow, all tergites divided (except XI), anterior tergites light-brown colors, chela pedipalpal darker reddish-brown (Figs. 3–4).</p>
            <p>Chelicera with five setae on hand, fixed finger with one apical tooth, and four subapical teeth, movable finger with tooth-like subapical lobe (Fig. 16), rallum with three blade, first and second long toothed (Fig. 17), gale long, with three apical branches (Fig. 18), serrula exterior with 18 blades (female, with 20 blades).</p>
            <p>Pedipalps robust (Figs. 19–20), in both male and female strongly granulated, trochanter 1.3× (female, 1.2×) as long as broad, femur 2.6× (female 2.7×) as long as broad, chela with pedicel 3.2× (female 3.4×), chela without pedicel 3.0× (female 3.2×), inner margin of hand in female rounded (Fig. 20). Movable finger chelal 0.518 mm (female 0.641 mm), fixed finger with 39 pointed teeth (female with 37 pointed teeth), movable finger with 30 flattened teeth (female with 32 flattened teeth) (Fig. 21). Trichobothrium ist distal de est; trichobothrium it proximal to est; ib near the level of isb; trichobothrium st proximal to the middle of the finger, sb closer to b than to st, nodus ramosus on fixed finger distal of est, and in the movable finger chelal proximal to trichobothrium t (Fig. 21), well-developed venom apparatus in both on the fixed finger and on the movable finger chelal (Fig. 21).</p>
            <p>Carapace triangular (Fig. 15), squamose sculpturing, posteriorly reticulate, cucullus shorts, with an indistinct longitudinal furrow, with medial furrow and indistinct posterior furrow (Fig. 15), four large eyes, all setae very short and acuminate, with approx. 38 setae (female approx. 36 setae), four setae on the anterior margin, one seta preocular on each side, with six to twelve setae on the posterior margin (Fig. 15).</p>
            <p>Coxal region: female and male with an apical lobe of coxa pedipalpal with two setae marginal, and three discal setae, approx. 26 setae (female approx. 28), coxa I ca. 22 setae (female ca. 26 setae), Coxa II ca. 24 setae (female ca. 28), Coxa III ca. 30 setae (female ca. 32 setae), Coxa IV ca. 34 setae (female ca. 36 setae).</p>
            <p>Opisthosoma: all tergites are divided (except XI); reticulate sculpture, all setae acuminate, pleural membrane strongly striated. Tergal chaetotaxy of holotype male, tergites I–XI: 6: 8: 8: 8: 6: 6: 4: 4: 6: 6: 4 [4ST]: 2, females paratypes, tergites I–XI: 6: 6–8: 6–8: 8: 6: 6: 6–4: 6–4: 6: 6: 4 [4ST]: 2, males paratypes, tergites I–XI: 6: 6: 8: 8: 6: 6–8: 6–8: 6–8: 6: 6–8: 4 [4ST]: 2, tergite XI with four setae tactile. Sternal chaetotaxy of the male holotype, sternites II–XII: 8(4): (1)6(1): (1)6(1): 8: 8: 8: 8: 8: 10: 10: 2, females paratypes, sternites II–XII: 6: (1)6(1): (1)6(1): 6–8: 6–8: 8: 8–10: 8–10: 10–12: 10: 2, males paratypes, sternites II–XII: 8(4): (1)6(1): (1)6(1): 6: 6–8: 8–10: 8–10: 8–10: 10–12: 10: 2, sternites XI with two setae tactile, pleural membrane III–X with one seta.</p>
            <p>The genital region of the male holotype with eighth setae, and four discal setae on the anterior operculum (Fig. 22), the posterior operculum with six marginal setae (Fig. 22), internal genital structures, indistinctly visible (Fig. 22). Genital region of the female paratype with six central setae on the anterior operculum, and six marginal setae on the posterior operculum.</p>
            <p>Legs: all legs light-yellow (Figs. 23–24), Leg I (Fig. 23): trochanter 1.5× (female 1.7×), femur 2.8× (female 2.7×), patella 2.4× (female 2.6×), tibia 2.4× (female 2.6×), metatarsus 3.2× (female 3.3×), tarsus 3.3× (female 4.8×), Leg IV (Fig. 24): trochanter 1.1× (female 1.2×), femur+patella 2.6× (female 2.7×), tibia 4.0× (female 4.1×), metatarsus 3.1× (female 3.2×), tarsus 3.7× (female 3.8×), subterminal setae smooth, arolium undivided and longer than claws (Fig. 24).</p>
            <p>Measurements (mm):Male holotype (Paratype in parentheses):body length 1.796(1.813); carapace 0.609 / 0.486 (0.612 / 0.489); Chelicerae; movable finger. pedipalps trochanter: 0.255 / 0.191 (0.257 / 0.193); femur: 0.604 / 0.229 (0.607 / 0.230); patella 0.485 / 0.209 (0.487 / 0.210); chela with pedicel 0.929 / 0.289 (0.931 / 0.290); chela without pedicel 0.873 / 0.289 (0.853 / 0.290), movable finger 0.578 (0.581). Leg I: trochanter 0.136 / 0.129 (0.137 / 0.131), femur 0.220 / 0.077 (0.225 / 0.079), patella 0.149 / 0.061 (0.151 / 0.063), tibia 0.159 / 0.065 (0.161 / 0.067); metatarsus 0.131 / 0.041 (0.133 / 0.043); tarsus 0.134 / 0.043 (0.136 / 0.045). Leg IV: trochanter 0.231 / 0.131 (0.235 / 0.133); femur+patella: 0.843 / 0.180 (0.845 / 0.183); tibia: 0.361 / 0.091 (0.365 / 0.093); metatarsus 0.219 / 0.071 (0.221 / 0.070); tarsus 0.189 / 0.051 (0.187 / 0.053).</p>
            <p>Female paratypes: Body length 2.241 –2.250. Carapace 0.691–0.693/0.552–0.553; Chelicerae; movable finger. Pedipalps:trochanter 0.226–0.229/0.210–0.213; femur 0.681–0.683/0.251–0.253; patella 0.550–0.553/0.220–0.223; chela with pedicel1.091–1.093/0.321–0.323; chela without pedicel 1.031–1.033/0.321–0.323; movable finger 0.641 – 0.643. Leg I: trochanter 0.050–0.060/0.136–0.137; femur 0.263–0.265/0.080–0.090; patella 0.167–0.169/0.102– 0.103; tibia 0.215–0.217/0.080–0.090; metatarsus 0.203–0.206/0.060–0.070; tarsus 0.195–0.197/0.041–0.043. Leg IV: trochanter 0.230–0.231/0.130–0.133; femur+patella 0.803–0.805/0.251–0.253; tibia 0.532–0.533/0.110–0.112; metatarsus 0.316–0.319/0.060–0.070; tarsus 0.249–0.251/0.040–0.050.</p>
            <p> Etymology. The specific epithet (  smaugi ) derives its name from the mythical (fictional) dragon, the last of the great dragons remaining in Middle Earth, in the fantasy literary book “The Hobbit” written by J.R.R. Tolkien. </p>
            <p>Distribution. Only known from the type locality (Brazil: Pernambuco, Meu Rei cave).</p>
            <p> Remarks. According to the descriptions by Beier (1931a, 1932b, 1959, 1964) and the taxonomic key presented by Mahnert (2001),  Progarypus smaugi sp. nov. differs from other species of  Progarypus Beier, 1931 as follows: it differs from the type species of the genus  Progarypus ramicola (Balzan, 1887) , from Paraguay, for the femur pedipalpal without pedicel, clearly separated, somewhat concave, femur pedipalps 0.720 / 0.160, patella 0.690 (0.540)/0.201, rallum of chelicera with three denticulate blades. It differs from  Progarypus longipes Beier, 1964 by each tergite with six to eight very short marginal setae; pedipalps relatively very thinned, femur pedipalpal on the apical third with an extended tactile seta, scaly sculpture in the medial area and granular in the rest. Fixed finger chelal with 45 densely spaced teeth, finger movable chelal with 35 pointed teeth in apical third, but completely flat at the base, and  Progarypus marginatus Beier, 1964 from Chile by the eb -esb -isb trichobothria at the base of the fingers, ist is closer to it than to ib; male femur pedipalpal 0.950 / 0.201, 4.7×, patella 0.801 / 0.240, 3.3×. Differ from  Progarypus novus Beier, 1931 from Brazil by the carapace 1.5× as long as wide, gradually narrowing, with truncated anterior margin, and approximately net-like sculpturing. Pedipalps slender, femur 4.0×, as long as wide, sessile, patella 3.5×, chelal fingers slightly shorter than hand without pedicel 0.54 mm. It differs from  Progarypus peruanus Beier, 1959 from Peru by the carapace as long as wide at the base, conically narrowed at anterior margin, densely granulated. Pedipalps almost as long as the body, femur without pedicel 4.3×, patella 3×, fingers clearly longer than hand chela with pedicel and as long as femur, tarsus IV shorter than tarsus I. It differs from  Progarypus oxydactylus (Balzan, 1887) from Paraguay by the carapace clearly longer than wide, finely granulated, and reticulate at the base, pedipalps rather long and thin. Femur 4.1× times as long as wide, weakly pedicelled, finely granulated, and striated, patella 3.2× as long as wide, notably shorter than the femur. Similarly, it differs from the species  Progarypus gracilis Mahnert, 2001 ,  Progarypus liliae Mahnert, 2001 , and  Progarypus setifer Mahnert, 2001 from Brazil by the length of the femur pedipalps 0.7-1.0- 1.21 mm, by the distribution of the trichobothria est, ist, it, et, the length of the fingers of the pedipalpal hand. The new species resembles  Progarypus nigrimanus Mahnert, 2001 , from Brazil however, it differs by the femur pedipalpal length 0.7–1.0 mm; at least one of trichobothrium est, ist, it, et proximal to st, finger relatively shorter, at most 1.3×, pedipalps bicolored, chelal hand darker than femur and patella, femur pedipalpal 4.7–5.3× (length 0.86 mm), patella 3.8–3.9× (length 0.78–0.79mm) longer than broad, carapace anteriorly with tessellated sculpturing. </p>
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	https://treatment.plazi.org/id/0389886AE943FFB61399109DFD2D2110	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB61399117FFB9C25E8.text	0389886AE940FFB61399117FFB9C25E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progarypus setifer Mahnert 2001	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Progarypus setifer Mahnert, 2001</p>
            <p>
                 Material examined.   5 males: Pernambuco, Brazil:  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-019)  .   4 females, Pernambuco,  
                <a title="Search Plazi for locations around (long -37.280224/lat -8.48725)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.280224&amp;materialsCitation.latitude=-8.48725">Meu Rei cave</a>
                 , 08°29’14.1”S, 37°16’48.8”W, 777 m a.s.l., 19-I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-020)  . 
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            <p> Diagnosis. According to the description made by Mahnert (2001), specimens of  Progarypus setifer Mahnert, 2001 can be identified by the combination of characters: the carapace and pedipalps reddish-brown, 1.6× as long as broad, densely granulate, between the numerous larger granula is a clearly granulated microsculpture four setae on anterior margin, two preocular setae each side, five to six setae on posterior margin, setae small, and indistinct (see Mahnert 2001; Fig. 36). Tergal chaetotaxy, tergites I–XII: 4: 5-6: 5-6: 5-6: 5-6: 5-6: 6-8: 6-8: 6-8[2ST]: 8[4ST]: 10[4ST]:2. Region genital of the male with ten marginal, and six discal setae on the operculum anterior, camera genital with 2/2 setae. Region genital of the female with four setae on anterior genital operculum. Chelicera, hand with 5 setae, movable finger with slightly bilobed subapical lobe, galea with 3 apical branches, serrula exterior 18 blades, flagellum 3 setae. Pedipalps: trochanter with a distinct dorsal hump, coarsely granular, femur and patella coarsely granulate, femur of the male 6.2× (female 4.9×), with a short dorsal tactile seta near the base, patella 6.3× (female 4.5×), chela with pedicel 5.4× (female 3.7×), fixed finger chelal with approx. 40 (female 39) pointed teeth, movable finger with 39 (32 female) pointed teeth. Trichobothrium eb, esb, ib, isb at finger base, ist, est, it, et in distal half and distal to st; trichobothrium ist nearly halfway between it and est; trichobothrium st halfway between sb and t (see Mahnert 2001; Figs. 38–40). Leg 1: femur 4.7× (female 3.9×), patella 2.6× (female 2.3×), tibia 6.6× (female 5.2×), metatarsus 5.7× (female 5.0×), tarsus 4.4× (female 5.0×). Leg IV: femur+patella 2.8× (female 2.7×), tibia 5.5× (female 4.8×), metatarsus 5.4× (female 4.4×), with a short tactile seta near i1s base, tarsus 6.3× (female 5.7×), subterminal seta smooth, arolia undivided and longer than claws. </p>
            <p>Measurements (mm). Body length: 2.613 –2.651, carapace 0.914–0.937/0.553–0.571 (0.931–0.950/0.611– 0.651). Pedipalp: femur 1.21–1.23/ 0.170 –0.190 (1.113–1.115/0.230–0.250), patella 1.230–1.250/0.200–0.210 (1.050–1.070/0.230–0.250), hand with pedicel 0.751–0.771/0.342–0.345 (0.801–0.803/0.461–0.471), finger length 1.101 –1.116 (0.981 –0.983), chelal length 1.830 –1.850 (1.720 –1.750). Legs: Leg I: femur 0.550–0.570/0.120–0.140 (0.470 –0.490 / 0.l22–0.123), patella 0.291–0.301/0.110–0.120 (0.253–0.256/0.112–0.113), tibia 0.487–0.489/0.071– 0.073 (0.401 –0.403 /0.08), metatarsus 0.351 –0.353 /0.06–0.07 (0.281 –0.283 /0.06–0.070), tarsus 0.221–0.223/0.050– 0.070 (0.201–0.203/0.040–0.050). Leg IV: femur+patella 0.881–0.883/0.310–0.313 (0.831–0.833/0.301–0.303), tibia 0.680–0.681/0.120–0.123 (0.626–0.629/0.131–0.133), metatarsus 0.411–0.413/0.080 (0.351–0.353/0.080), tarsus 0.311–0.313/0.050 (0.291–0.311/0.050).</p>
            <p> Distribution. Brazil (Harvey 2013; World  Pseudoscorpiones Catalog 2022 ). </p>
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	https://treatment.plazi.org/id/0389886AE940FFB61399117FFB9C25E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
0389886AE940FFB113991427FA1F2141.text	0389886AE940FFB113991427FA1F2141.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progarypus Beier 1931	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to  Progarypus Beier, 1931 species recorded from Brazil </p>
            <p>Adapted and extended from the key made by Mahnert (2001).</p>
            <p> 1. Large species, pedipalpal femur length 1.1 mm (female) to 1.21 mm (male); trichobothria ist, est, it, et in distal finger half, distal to st; finger 1.5× longer than chelal hand with pedicel...........................................  P. setifer Mahnert</p>
            <p>- Smaller species. pedipalpal femur length 0.6–1.0 mm; at least one of trichobothria est, ist, it, et proximal to st, finger relatively shorter, almost 1.3×longer than chelal hand with pedicel...................................................... 2</p>
            <p> 2. Finger shorter than hand with pedicel, trichobothrium ist and est apparently proximal to st; pedipalpal femur 4.6× (length 0.7 3mm), patella 3.5× (length 0.70) longer than broad.................................................  P. novus Beier</p>
            <p>- Finger 1.2 to 1.3× longer than chelal hand; only trichobothria est proximal to st .................................... 3</p>
            <p>3. Pedipalps bicolored................................................................................... 4</p>
            <p>- Pedipalps unicolored, the hand not darker than femur and patella................................................ 5</p>
            <p> 4. Chelal hand darker than femur and patella; pedipalpal femur 4.7–5.3× (length 0.8 6mm), patella 3.8–3.9× (length 0.7 8–0.7 9mm) longer than broad, carapace anteriorly with tessellated sculpturing...................... ..  P. nigrimanus Mahnert</p>
            <p> - Chelal hand lighter than femur and patella; pedipalpal femur 2.6× (female 2.7×) (length 0.604 –0.607 mm), patella 2.19–2.51× (length 0.485 –0.487 mm) longer than broad, carapace anteriorly with strongly granulate sculpture.......  P. smaugi sp. nov.</p>
            <p> 5. Pedipalps and carapace granulate; pedipalpal femur 5.5– 6× longer than broad (length 1.01 mm), patella 4.9–5.4× (length 0.8 5– 1.02 mm): carapace anteriorly granulate, with clear microsculpture between granula, tarsus IV 5.0–5.2 × longer than broad....................................................................................  P. gracilis Mahnert</p>
            <p> - Pedipalps and carapace smooth, pedipalpal femur 5.8× longer than broad (length 1. 12 mm). patella 4.7× (length 1.06mm); tarsus IV 7.7× longer than broad.............................................................  P. liliae Mahnert</p>
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	https://treatment.plazi.org/id/0389886AE940FFB113991427FA1F2141	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bedoya-Roqueme, Edwin;Tizo-Pedroso, Everton;Barbier, Eder;Lira, André Felipe De Araújo	Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder, Lira, André Felipe De Araújo (2023): Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil. Zootaxa 5293 (2): 317-332, DOI: 10.11646/zootaxa.5293.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.6
