taxonID	type	description	language	source
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	description	(Figs 1, 8 a) Types. HOLOTYPE: m #, body, aedeagus and detached abdomen glued on the same card, “ MEXICO, Veracruz Est. Biol. de Los Tuxtlas 18 ° 35 ’ N 95 ° 05 ’ W ” [white label, printed] // “ 4. V. 1991 H. A. Hespenheide ” [white label, printed, partly handwritten] // “ Griburius febriculosus sp. nov. HOLOTYPUS D. Sassi des. ” [red label, printed] // (TAMU). PARATYPES (8 specimens): MEXICO, CHIAPAS: Hwy 199 10 km S Palenque San Manuel Rd 30. V. 1987 at mercury vapor & blacklight (2 m #. ERPC, DSPC); Comitán de Domínguez (2 f #, MNHUB). MEXICO, VERACRUZ: same data of the HT but 28. IV. 1991 (2 f #, TAMU, DSPC); ca 15 km N Catemaco 1000 m N 18 ° 32 ’ W 95 ° 08 ’ 26. IV. 1991 (1 f #, TAMU). “ Mexico … ” [remainder of the label is unreadable] (1 f #, MNHUB). All paratypes are labelled: // “ Griburius febriculosus sp. nov. PARATYPUS D. Sassi des. ” [red label, printed] //.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	materials_examined	Type locality. Estación de Biología Tropical Los Tuxtlas (Veracruz, Mexico).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	etymology	Etymology. Latin adjective meaning feverish. No specific reference to the taxon, but the evening I was working on it, it turned out that my daughter had just got the COVID- 19 disease.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	distribution	Distribution. Mexico.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	diagnosis	Diagnosis. A species of Griburius characterized by unusually strong and dense punctation on the elytra and pronotum. On the latter the punctation tends to become wrinkled. The species seems to show an evident chromatic dimorphism, the males being dorsally yellow with a pattern of black lateral bands, while the females available for the study have entirely black elytra and pronotum. In terms of dorsal colouration, the species vaguely recalls Pachybrachis forreri Jacoby 1889 (see http: // www. chrysomelidae. it / amer-pachy / p. - forreri. html), which however is well distinguished by the shape of the pronotum, much less transverse, by the elytra slimmer and not converging towards the apex, and by the finer punctation.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	description	Description of male. Habitus in Fig. 1 a-c (PT). BL = 3.7 – 4.0 mm, BW = 2.4 – 2.7 mm, PL = 1.4 – 1.6 mm, PW = 2.2 – 2.3 mm. Interocular distance = 0.0 % (inner eyes margins are in contact along median line). Head yellow with vertex and, at times, part of clypeus black. Labrum yellow. Head surface basically devoid of setosity. Surface of frontoclypeal area with few, scattered, shallow-impressed punctation. Upper lobes of eyes in contact along midline, so neither interspace nor mid-cranial suture visible in between. Ocular lines strictly adhering to ocular rim, marked with a line of fine punctures along lower part of upper lobes. Ocular canthus large, angular. Antennae (Fig. 1 h) with first six antennomeres yellow, reminders brownish. Antennomeres 3 - 5 bright, subcylindrical; antennomeres 6 - 11, dull, flattened and more diffusedly setose. Pronotum black with lateral margins yellow. Two trapezoidal spots in posterior part of disc, close to midline, touching posterior margin and median line raising from anterior margin and reaching middle of disc yellow as well. At times, discal spots fused together giving single trilobate marking. Pronotal shape elliptical, scarcely transverse, regularly convex. Lateral margins narrow, visible from above only along posterior half, angularly curved at middle, with maximum width at about middle. Surface dull, covered with coarse strongly impressed punctation evenly distributed across whole pronotal surface. Punctures deep, lengthened, partly coalescent at middle of disc to form longitudinal ridges. Posterolateral impressions well marked just up to midline, so that pronotal outline looks slightly raised in lateral view. Scutellum yellow, triangular with apex truncated in straight line. Surface sparsely micropunctured. Elytron black with large yellow patch along suture, broadened to reach fifth row of punctures on anterior half, at about middle and on apical clivus. In holotype yellow patch on apical clivus separated from anterior one to form independent spot. Epipleuron yellow along anterior half. Elytral outline with sides almost straight and posteriorly convergent. Lateral margin rather large, simultaneously visible from above from apex to humeral callus. Elytral surface slightly flattened on disc, dull with strong, coarse, deeply impressed punctuation, partly coalescent to give rugulose appearance to disc surface. Punctures arranged in almost regular rows above all along inner part of disc. Intervals raised. Postscutellar area perceptibly raised. Humeral callus prominent, impunctate. Epipleuron flat, slightly rugulose. Pygidium totally yellow; surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax black, with prosternal process and most of metasternum yellow. Inner part of metaepisterna bordered with yellow as well. Abdominal ventrites yellow with central part of ventrites 1 - 4 light brown. Medial projection of ventrite 1 yellow. Hypomera almost bare, shiny, covered with dense, strong punctures; mesoepimera and mesoepisterna shiny, with fine, scattered punctation. Remainder of ventral surface matt, covered with thick, short, regularly distributed setae and fine, shallow punctures. Prosternal process large with sides almost straight between anterior coxae; surface almost flat, closely but shallowly punctured, covered with sparse semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite distinct, narrow. Ventrite posterior margin shallowly but perceptibly notched. Median lobe of aedeagus (Fig. 1 m-o) dorsoventrally flattened, terminated by median, long, pointed, apical process, delimited by mildly concave sides. In lateral view apex perfectly straight. Ventral surface marked by median low, linear carina. Setose depressions very shallowly impressed, with few, scattered, very small punctures and long arcuate setae. In dorsal view, in all examined specimens, membranes of endophallus always protruding from apical foramen. Among them, two apical lobes, extending sideways, always well distinguishable. Female. Habitus in Fig. 1 d-f (PT). BL = 3.6 – 4.5 mm, BW = 2.3 – 2.9 mm, PL = 1.2 – 1.6 mm, PW = 2.1 – 2.6 mm. Interocular distance = 8.3 – 8.9 % of BL. In females the head surface is black with the lower margin of clypeus and labrum yellow. The eyes are smaller and more separated along midline, and the head surface is evenly covered with coarse punctation. Additionally, upper parts are totally black, while yellow pattern on ventral parts is more extended, being the abdominal tergites and ventrites totally yellow. The fifth abdominal ventrite in females has a quite large, subelliptical, and deep pit. The bottom of the pit is glabrous, matt, slightly darkened in respect to the surrounding surface, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 1 i) is not pigmented, slender, sickle shaped with slightly curved, not swollen proximal lobe. The distal lobe is slender, almost straight in the first trait and then abruptly curved at apex. The ampulla is not pigmented, short. The duct insertion on the ampulla is short, neither thickened nor pigmented. The sperm gland insertion is rather long and slightly curved. The duct is uniform in size, slender, short, quite rigid, forming a single turn near the vasculum, then straight, devoid of further coils and turns. The insertion on the bursa copulatrix is simple, neither swollen nor pigmented.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB0FC15FCE2B5AF57FFD7C2.taxon	discussion	Remarks. The single male from Comitán de Domínguez is dorsally totally black apart from two triangular yellow spots on the anterior angles of the pronotum, and the scutellum, which is yellow as well. The single female from the same locality also differs from the others in the yellow lateral sides of pronotum and lower part of frontoclypeal surface, up to ocular lobes, yellow as well. Besides, the proximal lobe of the vasculum is fairly swollen at base. However, because of the small number of the available specimens, it is impossible to give any taxonomic significance to this variability.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	description	(Figs 2, 8 b) Types. HOLOTYPE: m #, body, aedeagus and detached abdomen glued on the same card, // “ MEXICO Quintana Roo ” [white label, printed] // “ Celarain Point Cozumel Island 21 - IV- 1960 J. F. C. Clarke ” [white label, printed, partly handwritten] // “ Griburius gracilis sp. nov. HOLOTYPUS D. Sassi des. ” [red label, printed] // (USNMNH). PARATYPES (26 specimens): MEXICO, QUINTANA ROO: same data of the HT (14 m # 12 f #, USNMNH, DSPC, MSNM). All paratypes are labelled: // “ Griburius gracilis sp. nov. PARATYPUS D. Sassi des. ” [red label, printed] //.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	materials_examined	Type locality. Celarain Point on Cozumel Island (Quintana Roo, Mexico).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	etymology	Etymology. Latin adjective meaning “ frail ”, stressing the relatively small size and the poor contrast of the dorsal colouring.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	distribution	Distribution. Mexico. This taxon is likely to be endemic to Cozumel Island (Quintana Roo), which is well known for hosting several endemic animal species (see for instance Cuarón et al. 2004).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	diagnosis	Diagnosis. The dorsal colouration, with a weak contrast between the whitish-yellow light areas and reddish-brown dark ones, characterizes the species quite well, even if, quite rarely, a darker pattern is shown on the central part of the elytra (Fig. 2 e). The most similar species is G. textus, which however differs in the wider and more angular outline of the aedeagal median lobe, in the larger size and the more robust dorsal punctation.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB4FC14FF4AB78F57B8D562.taxon	description	Description of male. Habitus in Fig. 2 a-c (PT). BL = 2.8 – 3.0 mm, BW = 1, 8 mm, PL = 1.1 – 1.6 mm, PW = 1.6 – 1.7 mm. Interocular distance = 0.0 – 3.6 % of BL. Head yellow with vertex, ocular perimeter, lower clypeal margin and antennal insertions brownish. Labrum yellow. Head surface basically devoid of setosity. Vertex shallowly but closely punctured. Surface of frontoclypeal area with scattered, well-impressed punctation above all on ocular canthi and between antennal insertions. Mid-cranial suture shallow, barely detectable between upper lobes of eyes. Upper lobes of eyes close to each other along midline but not really in contact. More rarely, upper ocular lobes touching each other for short distance. Ocular lines usually distinct above all along lower part of upper lobes, marked by row of punctures, strictly adhering to ocular rim. Ocular canthus large, angular. Antennae (Fig. 2 h) with first two antennomeres yellow, reminders brownish. Antennomeres 3 - 5 bright, subcylindrical; antennomeres 6 - 11, dull, more flattened and more diffusedly setose. Pronotum light brown with lateral and anterior margins yellow. Two trapezoidal spots in posterior part of disc, close to midline, touching posterior margin and median line raising from anterior margin and reaching middle of disc yellow as well. Pronotal shape elliptical, scarcely transverse, regularly convex. Lateral margins narrow, visible from above only along posterior third, regularly curved, with maximum width at about middle. Surface moderately shiny with strongly impressed punctation evenly distributed across whole pronotal surface. Posterolateral impressions weakly impressed but still detectable. Scutellum brownish, triangular with apex truncated in straight line. Surface sparsely micropunctured. Elytron light brownish with large yellow patch at center of disc, broadened at middle and reduced to simple stripe along apical clivus. Apical margin yellow as well. Bottom of punctures brown also in lighter areas. Epipleuron yellow. Elytral outline with sides almost straight and convergent posteriorly. Lateral margins narrow, simultaneously visible from above only along posterior half. Elytral surface slightly flattened on disc, moderately shiny with strong, well-impressed punctation distinct up to posterior clivus, arranged in almost regular rows along inner half of elytron, more confused elsewhere. Intervals flat. Postscutellar area slightly raised. Humeral callus prominent, impunctate. Epipleuron smooth, impunctate, with slightly convex surface. Pygidium totally yellow Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax light brown, only metasternum almost completely yellow. Abdominal ventrites yellow with central part of ventrites 1 - 4 light brown. Medial projection of ventrite 1 yellow. Hypomera, almost bare, shiny, covered with dense, strong punctures; mesoepimera and mesoepisterna shiny, almost devoid of punctation. Remainder of ventral surface of thorax matt, covered with thick, short, regularly distributed setae and fine, shallow punctures. Prosternal process large with sides almost straight between anterior coxae, terminated with a round apex; surface almost flat, shallowly punctured, covered with long, semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Ventrite posterior margin shallowly notched. Median lobe of aedeagus (Fig. 2 m-o) dorsoventrally flattened, terminated by triangular apex. In lateral view apex perfectly straight. Ventral surface almost smooth. Setose depressions very shallowly impressed, with surface covered by small punctures and few short setae Female. Habitus in Fig. 2 d-f (PT). BL = 3.6 – 3.8 mm, BW = 2.4 – 2.6 mm, PL = 1.3 – 1.4 mm, PW = 2.1 – 2.3 mm. Interocular distance = 8.3 – 10.5 % of BL. In females the head surface is brownish with a transverse yellow band between the ocular canthi. Two further yellow spots just below the antennal insertions. The eyes are smaller and more separated along midline, and the head surface is evenly covered with coarse punctation and sparse short setae. Sometimes, the yellow patch on elytron tends to change into dark brown shades (Fig. 2 e). The fifth abdominal ventrite in females has a quite large, rounded, and deep pit. The bottom of the pit is glabrous, matt, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 2 i) is scarcely pigmented, sickle shaped with straight, slightly swollen proximal lobe. The distal lobe is slender, regularly curved, distally tapered, and terminated with a slender apex. The ampulla is not pigmented, almost square. The duct insertion is ahort and slightly thickened. The sperm gland insertion is rather long, slender and curved. The duct is uniform in size, short, slender, quite rigid, straight, devoid of coils and turns. The insertion on the bursa copulatrix is simple, neither swollen nor pigmented.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	description	(Figs 3, 8 c) Types. HOLOTYPE: m #, body, aedeagus and detached abdomen glued on the same card, // “ MEX, Chiapas, Parque Laguna Belgica 27 - V- 1987 D A Rider E. G. & T. J. Riley ” [white label, printed] // “ collected at mercury vapor & blacklight ” [white label, printed] // “ Griburius mokaya sp. nov. HOLOTYPUS D. Sassi des. ” [red label, printed] // (TAMU). PARATYPES (42 specimens): MEXICO, CHIAPAS: same data of the holotype (13 m # 3 f #, ERPC, DSPC, TAMU); hwy 195 4.5 km N Ixtapa 3000 ’ 24. V. 1987 D. A. Rider, E. G. & T. J. Riley at mercury vapor and blacklight (8 m # 8 f #, ERPC); Mpio Villa Flores rd. to Agronomos Mexicanos 1200 m (1 f #, ERPC). MEXICO, VERACRUZ: 8 km N Fortin 20. IV. 1978 (1 f #, TAMU). COSTA RICA, GUANACASTE: Finca Jenny 31 km N of Liberia 300 m VI. 1989 (3 m # 2 f #, BMNH, DSPC); 3 km SE R. Naranjo 11. IV. 1993 (1 f #, BYU). HONDURAS, FRANCISCO MORAZÁN: 5 km E Escuela Agricola Panamericana 2. VI. 1993 (1 f #, FSCA). PANAMA, CHIRIQUI: vic Hornito 42 ’’ 14 - 18. V. 1996 (1 f #, TAMU). All paratypes are labelled: // “ Griburius mokaya sp. nov. PARATYPUS D. Sassi des. ” [red label, printed] //.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	materials_examined	Type locality. Parque educativo Laguna Bélgica (Ocozocoautla, Tuxtla Gutiérrez, Chiapas, Mexico).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	etymology	Etymology. Mokaya is the name given to some ancient Mesoamerica’s cultures located in particular along the Pacific Coast of today’s Chiapas, Mexico. The word Mokaya is a noun in the nominative singular, standing in apposition to the generic name (art. 11.9.1.2. I. C. Z. N., 4 th edition). Additional material examined. GUATEMALA: Ciudad de Guatemala 1. V. 2022 (1, GBIF). HONDURAS: Francisco Morazán San Antonio de Oriente, El Zamorano 22. IV. 2020 & 14. V. 2020 (2, GBFI). Further checked data from literature. Jacoby (1889) reported and shortly described a specimen from Guatemala, attributing it to Griburius equestris (Olivier, 1808), which should be considered as belonging to the new species. The datum was also mentioned in Riley et al. (2001) and correctly believed to belong to a taxon distinct from G. equestris. Additionally, this species is reported in Ordóñez-Reséndiz & López-Pérez (2021) for the Mexican States of Puebla, Oaxaca, Veracruz and Yucatan on the basis of material housed in the collections of the Museo de Historia natural de la Ciudad de México. These specimens have probably to be assigned to the new species but, given the difficulty until now to correctly identify the Mexican Griburius species, not comprehensively revised since the late 19 th century, the presence of the new species in the aforementioned Mexican states requires, in my opinion, further confirmation.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	distribution	Distribution. Costa Rica, Guatemala, Honduras, Mexico, Panama.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	diagnosis	Diagnosis. The species is similar in morphology to G. equestris, from which it can be easily distinguished by the different shape of the aedeagal median lobe. Despite the variability of both species in the colouration, some remarkable differences can be pointed out on ventral surface. In males, median projection of the first abdominal ventrite is always yellow (black in G. equestris). Usually, the black spot on the pygidium is smaller, and the fifth abdominal ventrite is totally yellow (central part of it is the same usually black in G. equestris). In females (and in light coloured males) the distinction is easier, being the ventral surface usually almost totally yellow.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	description	Description of male. Habitus in Fig. 3 a-c (HT), 3 f-g (PT). BL = 2.9 – 3.3 mm, BW = 1.7 – 2.0 mm, PL = 1.0 – 1.2 mm, PW = 1.4 – 1.7 mm. Interocular distance = 0.0 – 2.0 %. Head totally yellow, at times vertex and antennal insertions darkened. Labrum yellow as well. Head surface with sparse, very fine punctures and almost devoid of setosity. Mid-cranial suture shallow, but clearly detectable above and below the upper lobes of eyes. Upper lobes of eyes usually in contact along midline, so neither interspace nor mid-cranial suture visible in between. Ocular lines strictly adhering to ocular rim, sometimes marked with line of fine punctures along lower part of upper lobes. Ocular canthus large, angular. Antennae (Fig. 3 h) with first 5 antennomeres yellow, 3 - 5 bright, subcylindrical; 6 - 11 darker, dull, more flattened and more diffusedly setose. Pronotum yellow with two drop-shaped black spots on central part of disc, at sides of midline. Such spots at times smaller and rounded, or angulate, or even completely missing. Sometimes a hint of longitudinal brownish line near posterior margin, in front of scutellum. Pronotal shape elliptical, scarcely transverse, slightly flattened, clearly narrower than elytra. Lateral margins narrow, visible from above only along posterior half, slightly angulate just behind middle, where maximum width occurs. Surface moderately shiny with sparse, strongly impressed punctation evenly distributed across whole pronotal surface. Posterolateral impressions weakly impressed but still detectable. Scutellum black or yellow, regularly rounded at apex or shortly truncated in almost straight line. Surface sparsely micropunctured. Elytron yellow with three rounded black spots on disc, first one near basal margin between first and fifth rows of punctures, second one on humeral callus, third one just behind midline at about same distance from suture and lateral margin. Lateral margin and posterior part of suture black as well. Postmedian spot at times laterally extended to merge into marginal black pattern. Besides, anterior spots can coalesce to form a large transverse black band along basal margin (Fig. 3 f). Again, size of elytral black markings sometimes more or less reduced. Epipleuron totally yellow or, more often, black along posterior half. Elytral outline lengthened, with sides almost straight and slightly convergent posteriorly. Lateral margins narrow, simultaneously visible up to humeral calli when seen from above. Elytral surface moderately shiny with strong, well-impressed punctation distinct up to posterior clivus, arranged in almost regular rows. Intervals flat. Postscutellar area perceptibly raised. Humeral callus prominent, impunctate. Epipleuron, impunctate, slightly rugulose with flat surface. Pygidium totally yellow or with rounded black spot at apex. Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax black except for prosternal process and outer part of hypomera, being yellow. Abdominal ventrites yellow with central part of ventrites 1 - 4 light brown. Medial projection of ventrite 1 yellow. At times, in light coloured specimens, ventral parts totally yellow. Hypomera, mesoepimera and mesoepisterna shiny, with few, well-impressed punctures, almost devoid of setosity. Remainder of ventral surface matt, covered with short, regularly distributed setae and fine, shallow punctures. Prosternal process large, with sides fairly salient, almost straight between anterior coxae; surface almost flat, shallowly punctured, covered with semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Posterior margin of ventrite straight. Median lobe of aedeagus (Fig. 3 m-o) dorsoventrally flattened, terminated by triangular apex with slightly concave sides. In lateral view apex slightly bent dorsally. Ventral surface weakly convex towards apex. Setose depressions barely delimited, with few arcuate setae. Female. Habitus in Fig. 3 d-e (PT). BL = 3.4 – 3.9 mm, BW = 2.1 – 2.4 mm, PL = 1.3 – 1.4 mm, PW = 1.8 – 2.1 mm. Interocular distance = 7.7 – 8.8 % of BL. In females the eyes are smaller and more separated along midline, and mid-frontal suture clearly visible. In addition, dorsal black pattern is generally reduced, and black spots never tend to coalesce. Lastly, ventral parts are often completely yellow. The fifth abdominal ventrite in females has a quite large, rounded, and deep pit. The bottom of the pit is glabrous, matt, usually yellow, but sometimes frankly black, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 3 i) is quite unusual. It is asymmetrical, with no clear-cut separation between a distal and a proximal lobe, hook-shaped, with proximal lobe gradually tapered but terminated with a slightly swollen end. The ampulla is brief, not pigmented. The duct and sperm gland insertions are short and straight. The duct is uniform in size, slender, quite rigid, straight, with a single turn near the vasculum. At the insertion of the ductus, the bursa copulatrix is shaped in a couple of membranous coils.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB5FC18FF55B56F57CFD242.taxon	discussion	Remarks. In Griburius equestris the interocular distance in males is variable, but at least a little gap is always present between the upper lobes of the eyes (Fig. 13 c). On the other hand, in G. mokaya specimens from Mexico the eyes are clearly in contact for a rather long tract (Fig. 3 c). However, it should be noted that in the three Costa Rican males available for study the interocular distance is more similar to that of G. equestris than to the Mexican samples of G. mokaya. Moreover, the dorsal colouration seems to be rather variable in the new species, according to geographic parameters: in the type locality all the studied specimens show the “ typical ” pattern of black spots, while in the population from Ixtapa ventral parts are totally yellow in females, elytral spots are smaller and the pronotum is uniformly yellow (apart from a single male), being devoid of the two black rounded discal markings (Fig. 3 d-e). At the moment it is not considered opportune to assign the populations from Ixtapa and Costa Rica to distinct taxa, but it is hoped that additional material will be available to further investigate the matter.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	description	(Figs 4, 8 d) Types. HOLOTYPE: m #, body, aedeagus and detached abdomen glued on the same card, // “ MEXICO, Nayarit San Blas July 5 1972 K. Stephan leg. ” [white label, printed] // “ Griburius puncturatus sp. nov. HOLOTYPUS D. Sassi des. ” [red label, printed] // (FSCA). PARATYPES (28 specimens): MEXICO, JALISCO: Chamela vic. UNAM 9 - 19. VII. 1993 (1 m #, 1 f #, TAMU); 6 km N Chamela 15 - 17. VII. 2002 (1 m #, BMNH); Puerto Vallarta airport 15. VII. 1987 (1 f #, FSCA); Vulcan Colima 1918 (1 f #, ZSM). NAYARIT: same data of the holotype (5 m # 4 f #, FSCA, DSPC); San Blas (2 f #, MNHUB, ZSM); Navarrete 28. VII. 1953 D. Rockfeller Mexican Expedition 1953 (2 f #, AMNH); El Cora Tepic (1 m # 3 f #, MNHUB); Compostela 27. VI. 1954 (2 f #, AMNH); 15 mi SW Compostela 19. VII. 1984 (1 f #, FSCA); Mecatan NE of Santa Cruz 1. IX. 1963 2000 - 2500 ft (1 f #, FSCA); Santiago de Ixcuintla (1 f #, MNHUB). OAXACA: 2 MI n Candelaria Loxicha 17. VI. 1974 at light (1 f #, TAMU). All paratypes are labelled: // “ Griburius puncturatus sp. nov. PARATYPUS D. Sassi des. ” [red label, printed] //.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	materials_examined	Type locality. San Blas (Nayarit, Mexico).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	etymology	Etymology. Named after the black spotted dorsal yellow surface. Additional material examined. JALISCO: Marina Vallarta Puerto Vallarta 26. vii. 2021 (1, GBIF). SINALOA: Mazatlán 26. VI. 2015 & 27. VI. 2016 & 23. VI- 19. VII. 2017 & 19. VI- 18. VII. 2018 & 26. VI- 1. VII. 2019 & 12. VI. 2020 & 14. VI- 13. VII. 2021 (17, GBIF); Cerritos Mazatlán 8. VII. 2022 (1, GBIF).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	distribution	Distribution. Mexico.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	diagnosis	Diagnosis. The new species is similar in morphology to Griburius gamma. However, the M-shaped brown marking on pronotal disc is always clearly visible in G. gamma, whereas it is missing in the new species. The pronotal punctation is also different, being more closely spaced, more regularly distributed and finer in G. gamma. Again, the elytral dark pattern is usually more extended in G. gamma and the antennae slightly but perceptibly slenderer and lengthened. Additionally, the shape of the aedeagal median lobe is different, in particular on its apical end.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFB9FC1DFF55B20F5750D082.taxon	description	Description of male. Habitus in Fig. 4 a-c (HT). BL = 3.3 – 3.8 mm, BW = 2.0 – 2.3 mm, PL = 1.3 – 1.5 mm, PW = 1.8 – 2.1 mm. Interocular distance = 0.0 – 2.6 % of BL. Head totally yellow. Labrum yellow as well. Vertex very shallowly, minutely punctured. Head surface with sparse punctures and scattered, short setosity above all close to ocular lines and between antennal insertions. Mid-cranial suture shallow, but clearly detectable between upper lobes of eyes. Upper lobes of eyes close to each other along midline usually not really in contact. Ocular lines usually distinct above all along lower part of upper lobes, marked by row of fine but well-impressed punctures, strictly adhering to ocular rim. Ocular canthus large, angular. Antennae (Fig. 4 h) with first 5 antennomeres yellow, 3 - 5 bright, subcylindrical; 6 - 11 darker, dull, more flattened and more diffusedly setose. Pronotum totally yellow. Pronotal shape elliptical, scarcely transverse, regularly convex. Lateral margins narrow, basically not visible from above, regularly curved, with maximum width at about middle. Surface rather matt covered with strong punctuation mixed with much smaller punctures, irregularly distributed, being denser along sides and posterior margin, sparser at center of disc. More rarely pronotal punctures almost missing. Posterolateral impressions well impressed, narrow, obliquely arranged. Scutellum yellow, triangular with apex truncated in straight line. Surface sparsely micropunctured. Elytron yellow with two small, rounded black spots along basal margin, first one between first and second rows of punctures, second one on humeral callus. Further rounded black spot at center of disc, just behind midline. At first third of elytron some punctures larger and deeper, surrounding interstices slightly darkened and perceptibly impressed to give a hint of transverse brownish patch. Apical margin and suture narrowly darkened as well. Bottom of punctures brownish. Epipleuron yellow. Elytral outline with sides almost straight and slightly convergent posteriorly. Lateral margins narrow, simultaneously visible from above only along posterior half. Elytral surface moderately shiny with strong, well-impressed punctation distinct up to posterior clivus, arranged in almost regular rows. Intervals flat. Postscutellar area perceptibly raised. Humeral callus prominent, impunctate. Epipleuron, impunctate, slightly rugulose with slightly convex surface. Pygidium totally yellow Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax and abdomen yellow; only metaepisterna partly brownish. Hypomera, mesoepimera and mesoepisterna shiny almost bare, covered with scattered, strong punctures. Remainder of ventral surface of thorax matt, covered with thick, short, regularly distributed setae and fine, shallow punctures. Prosternal process large, short, with sides almost straight between anterior coxae; surface almost flat and closely but shallowly punctured, covered with long, semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Ventrite posterior margin very shallowly notched. Median lobe of aedeagus (Fig. 4 m-o) dorsoventrally flattened, almost parallel-sided, terminated by small median denticle. Apex straight in lateral view. Ventral surface marked by low, barely distinguishable, longitudinal carina extended along apical half. Setose depressions very shallowly impressed, with few, minute punctures and arcuate setae. Female. Habitus in Fig. 4 d-f (PT). BL = 4.4 – 5.2 mm, BW = 2.7 – 3.3 mm, PL = 1.6 – 2.0 mm, PW = 2.4 – 3.1 mm. Interocular distance = 9.1 – 9.6 % of BL. In females the eyes are smaller and more separated along midline, and the head surface is evenly covered with strong punctation. Additionally, the dark pattern on the anterior third of elytron is often darker, being transformed into a transverse, additional black spot, extended between the first and the fifth rows of punctures, or even broken into two-three separate rounded spots. The suture and the lateral margin, and at times even part of epipleuron, are black as well. The fifth abdominal ventrite in females has a quite large, rounded, and deep pit. The bottom of the pit is glabrous, matt, slightly darkened in respect to the surrounding surface, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 4 i) is weakly pigmented along the basal half of proximal lobe, which is fairly swollen at base. The distal lobe is long, slender, regularly curved and terminated with an acute apex. The ampulla is slightly pigmented, weakly curved. The duct and sperm gland insertions are short and straight. The duct is uniform in size, slender, quite rigid, devoid of coils and turns. The insertion on the bursa copulatrix is slightly swollen and mildly pigmented.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	description	(Figs 5, 8 e) Types. HOLOTYPE: m #, body, aedeagus and detached abdomen glued on the same card, // “ MEXICO Quintana Roo ” [white label, printed] // “ Isla Mujeres 29. iii. 1960 J F G Clarke ” [white label, printed] // “ Griburius textus sp. nov. HOLOTYPUS D. Sassi des. ” [red label, printed] // (USNMNH). PARATYPES (28 specimens): MEXICO, QUINTANA ROO: same data of the HT (14 m # 9 f #, USNMNH, DSPC, MSNM); 30 km W. Puerto Morelos 23 - VI- 2009 MV & blacklight (1 f #, BYU); Quintana Roo or. Tulum 13 - 15. VI. 1994 (1 f #, MDPC); 10.9 km. S Playa del Carmen 1. VI. 1990 (1 f #, FSCA). YUCATÁN: 16 km W Piste, Hwy 180, 19. V. 1987 (1 f #, ERPC); // “ textus m. Yucatan ” [white label, handwritten] // “ 34199 ” [white label, handwritten] // (1 f #, MLUH). BELIZE, Stann Creek Dist.: Rum Point Inn 2.5 mi N Placiencia 21 - 22. VIII. 2001 (1 f #, FCSA). All paratypes are labelled: // “ Griburius textus sp. nov. PARATYPUS D. Sassi des. ” [red label, printed] //.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	materials_examined	Type locality. Isla Mujeres (Quintana Roo, Mexico).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	etymology	Etymology. This epithet was found handwritten on a label pinned under the single specimen housed in Suffrian’s collection at MLUH. It was undoubtedly appointed by the German entomologist who eventually did not find the time to describe the new taxon. Additional material examined. BELIZE, Orange Walk: Indian Church 5. V. 2022 (1, GBIF).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	distribution	Distribution. Belize, Mexico.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	diagnosis	Diagnosis. The species shows a strong punctuation on pronotum and elytra, which is not a common pattern within Griburius. For this characteristic it looks rather like G. febriculosus, but pronotal punctures are always rounded and well separated and there is no tendency to coalesce, giving a rugose appearance to the surface, as observed in this latter species. Besides, the dorsal dark colouration is chestnut or reddish or dark brown, never frankly black like in G. febriculosus. Additionally, males differ in the aedeagal apex (less pointed) and females in the shape of the spermatheca (Figs 1 i and 5 i). G. textus is also similar to G. gracilis, but it differs in the darker dorsal colouration, in the stronger dorsal punctation and in the larger size.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	description	Description of male. Habitus in Fig. 5 a-c (PT). BL = 3.6 – 3.7 mm, BW = 2.2 – 2.3 mm, PL = 1.4 – 1.5 mm, PW = 2.1 mm. Interocular distance = 2.7 – 2.8 % of BL. Head yellow with vertex, antennal insertion, and subtle line between upper lobes of eyes brownish black. Labrum yellow. Whole head surface almost bare, with few, scattered, shallow punctures above all on frontoclypeal surface and ocular canthi. Mid-cranial suture shallow but clearly detectable between upper lobes of eyes. Upper lobes of eyes close to each other along midline, but not in contact (Fig. 5 c). Ocular lines narrow, marked by row of fine punctures, strictly adhering to ocular rim. Ocular canthus angular, not differentiated from the remainder of frontoclypeal surface. Antennae (Fig. 5 h) with antennomeres 1 - 2 yellow; 3 - 5 partly brownish, bright, subcylindrical; 6 - 11 darker, dull, more flattened and more diffusedly setose. Pronotum light brown with lateral and anterior margins yellow. Two trapezoidal spots on posterior part of disc, close to midline, touching posterior margin and median line raising from anterior margin and reaching middle of disc yellow as well. Pronotal shape elliptical, scarcely transverse, regularly convex. Lateral margins narrow, visible from above only along posterior third, regularly curved, with maximum width just behind middle. Surface moderately shiny with coarse strongly impressed punctation evenly distributed across whole pronotal surface. Posterolateral impressions weakly impressed but easily detectable. Scutellum yellow bordered with brown, posteriorly raised, subtriangular with apex truncated in straight line. Surface sparsely micropunctured. Elytron yellow irregularly patched with brown along lateral half of disc, above all along anterior third and posterior area just before apical clivus. Lateral margin and suture narrowly brown; bottom of punctures brown as well. Epipleuron yellow. Elytral outline rather short with sides almost straight and convergent posteriorly. Lateral margins narrow, simultaneously visible from above only along posterior half. Elytral surface slightly flattened on disc, moderately shiny with strong, well-impressed punctation distinct up to posterior clivus, arranged in almost regular rows along inner half of elytron, more confused elsewhere. Intervals slightly raised. Postscutellar area perceptibly raised. Humeral callus prominent, impunctate. Epipleuron smooth, impunctate, with slightly convex surface. Pygidium totally yellow Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax light brown, only metasternum almost completely yellow. Abdominal ventrites yellow with central part of ventrites 1 - 4 light brown. Medial projection of ventrite 1 yellow. Hypomera, almost bare, shiny, covered with dense, strong punctures; mesoepimera and mesoepisterna shiny, almost devoid of punctation. Remainder of ventral surface of thorax matt, covered with thick, short, regularly distributed setae and fine, shallow punctures. Prosternal process large, short, with sides almost straight between anterior coxae; surface almost flat and closely but shallowly punctured, covered with long, semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Ventrite posterior margin very shallowly notched. Median lobe of aedeagus (Fig. 5 m-o) dorsoventrally flattened, terminated by triangular apex with slightly concave sides. In lateral view apex perfectly straight. Ventral surface marked by very low, linear carina extended from apex to basal foramen. Setose depressions very shallowly impressed, with surface covered by small punctures and long arcuate setae. In dorsal view two flattened membranous lobes almost always protruding from apical foramen. Female. Habitus in Fig. 5 d-g (PT). BL = 4.2 – 4.4 mm, BW = 2.9 – 3.0 mm, PL = 1.6 – 1.7 mm, PW = 2.5 – 2.7 mm. Interocular distance = 9.1 – 9.5 % of BL. In females the head surface is brownish with a transverse wavy yellow line, usually discontinuous, between ocular canthi. Lower margin of clypeus and labrum are yellow as well. The eyes are smaller and more separated along midline, and the head surface is evenly covered with coarse punctation. In addition, the brown pattern on the pronotum and elytra is more extended with yellow colour confined to irregular patches along the intervals of the rows of punctures. The fifth abdominal ventrite in females has a quite large, rounded, and deep pit. The bottom of the pit is glabrous, matt, slightly darkened in respect to the surrounding surface, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 5 i) is scarcely pigmented, sickle shaped with straight, swollen proximal lobe. The distal lobe is slender, regularly curved and terminated with a blunt apex. The ampulla is not pigmented, almost square. The duct insertion is short while the sperm gland insertion is rather long, slender and curved. The duct is uniform in size, slender, quite rigid, straight, devoid of coils and turns. The insertion on the bursa copulatrix is simple, neither swollen nor pigmented.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFBCFC03FF4AB0CF52A1D4C2.taxon	discussion	Remarks. Male specimens were available for the study only from the type locality (Isla Mujeres). Mainland material consists only in females which differ in slightly larger size and darker colouration.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA3FC01FF55B38F5349D3A2.taxon	description	(Figs 6, 9 a)	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA3FC01FF55B38F5349D3A2.taxon	materials_examined	Type locality. “ Nord America ” Additional material examined. U. S. A., FLORIDA, Alachua Co.: Gainesville 14. III. 1982 (2, USNMNH); Gainesville 10. X. 1985 (1, FIMU); Sugarfoot Gainesville 20. III. 2022 (1, GBIF). Hernando Co.: Inverness 3. V. 1980 (1, USNMNH). Highlands Co.: Archbold Biol. Sta. 7. IV. 1971 & 13 - 17. IV. 1981 (3, USNMNH); Archbold Biol. Sta. 5. IV. 1998 (2, BYU); Archbold Biol. Sta. 13 - 17. IV. 1981 & 4. IV. 1984 & 12. IV. 1986 (3, FIMU); Archbold Biol. Sta. Red Hill Area 31. III. 2005 at black light (1, USNMNH); Archbold Biol. Sta. Lk. Placid 7. IV. 1983 (1, FIMU). Leon Co.: Tallahassee 3. IV. 1982 at light (1, ERPC); Tallahassee 11. IV. 1976 (1, ERPC). Levy Co.: hwy 24 at Alachua Co. line 9. IV. 1980 (1, ERPC). Marion Co.: 2.5 mi SSE Micanopy 6 - 13. III. 2001 (1, TAMU); Ocala Nat. For. Hopkins Prairie 29. IV. 1988 (1, ERPC); East Silver Springs Shores Malauka Road nr. Meadow Lake 10. IV. 1992 (2, ERPC); SR 200 Paddok Mall on Quercus virginiana 15. IV. 2008 (1, BYU). Polk Co.: Lk. Marion Crk. Rd. 25. III. 1999 (1, USNMNH). St. Johns Co.: St. Augustine 29 ° 55.4 ’ N 81 ° 17.9 ’ W 10. IV. 2004 (2, BYU). Wakulla Co.: Ochlockonee State Park 23. IV. 1986 (1, FIMU). GEORGIA, Johnson Co.: 1 mi E Kite 7. IV. 1977 (1, ERPC). MISSISSIPPI, Jackson Co.: Jackson County 25. III. 2020 (1, GBIF). TEXAS, Bastrop Co.: 7.3 mi S Rosanky 8. V. 2005 (1, DSPC); Buescher St. Pk. 22. IV. 1973 (1, TAMU). Brazos Co.: College Sta. Riley Estate 30 ° 35 ’ 18 ’’ N 96 ° 15 ’ 12 ’’ W 17 - 19. IV. 2006 & 13. IV. 2007 (5, ERPC); College Station Lick Ck. Pk. 13. IV. 1996 (2, ERPC); College Station 21. IV. 1933 (2, TAMU). Burleston Co.: 7 mi NW jct. Hwy 21 on FM 908 30. V. 2004 (1, ERPC). Tyler Co.: 4 mi E Spurger 24 - 25. III. 1995 (2, TAMU). Victoria Co.: 3 mi S Inez 6. V. 1978 (1, FIMU).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA3FC01FF55B38F5349D3A2.taxon	distribution	Distribution. U. S. A.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA3FC01FF55B38F5349D3A2.taxon	diagnosis	Diagnosis. The species is very similar in size, body outline and colour pattern to G. mokaya, but only in a single case the pronotal black spots are missing, while this condition is usual at least in some populations of G. mokaya. Additionally, the elytral spots are always rounded and do not seem to have the tendency to coalesce. The major differences, however, are shown by the different ventral profile of the aedeagal median lobe (Figs 3 o and 6 o) and the different shape of the spermathecal vasculum (Figs 3 i and 6 i).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA3FC01FF55B38F5349D3A2.taxon	description	Description of male. Habitus in Figs 6 c, 6 g. BL = 3.2 – 3.4 mm, BW = 1.9 – 2.1 mm, PL = 1.1 – 1.2 mm, PW = 1.6 – 1.9 mm. Interocular distance = 0.0 – 2.9 % of BL. Head totally yellow, at times vertex and antennal insertions darkened. Labrum yellow as well. Head surface almost devoid of punctation and setosity. Mid-cranial suture shallow, but clearly detectable above and below the upper lobes of eyes. Upper lobes of eyes very close to each other but usually not really in contact, so that narrow interspace and mid-cranial suture are visible in between. Ocular lines strictly adhering to ocular rim, sometimes marked with hint of very fine punctures aligned along lower part of upper lobes. Ocular canthus large, angular, with surface not differentiated by remainder of frontoclypeus. Antennae (Fig. 6 h) with first 5 antennomeres yellow, 3 - 5 bright, subcylindrical; 6 - 11 darker, dull, more flattened and more diffusedly setose. Pronotum yellow with two rounded black spots on central part of disc, at sides of midline. Rarely such spots slightly obliquely elongated, or smaller, but never missing. Sometimes a hint of longitudinal brownish line near posterior margin, in front of scutellum. Pronotal shape elliptical, scarcely transverse, slightly flattened. Lateral margins narrow, visible from above only along posterior half, slightly angulate just behind middle, where maximum width occurs. Surface moderately shiny with usually strongly impressed punctation almost evenly distributed across whole pronotal surface. Sometimes punctures thinner on central part of disc. Posterolateral impressions weakly impressed but easily detectable. Scutellum black, regularly rounded at apex or shortly truncated in almost straight line. Surface sparsely micropunctured. Elytron yellow with three rounded black spots on disc, first one near basal margin between first and fifth rows of punctures, second one on humeral callus, third one just behind midline at about same distance from suture and lateral margin. Posterior part of lateral margin black as well. Elytral black pattern sometimes more or less reduced to almost totally missing, but at least brownish marking on humeral callus always present. Elytral outline lengthened, with sides almost straight and slightly convergent posteriorly. Lateral margins narrow, simultaneously visible along posterior half when seen from above. Elytral surface moderately shiny with strong, well-impressed punctation distinct up to posterior clivus, arranged in almost regular rows. Intervals flat. Postscutellar area slightly raised. Humeral callus prominent, impunctate. Epipleuron, impunctate, slightly rugulose with flat surface. Pygidium yellow with rounded black spot at apex. Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax black with yellow hypomera and prosternal process. Abdominal ventrites black with large yellow margins. Hypomera, mesoepimera and mesoepisterna shiny, with few, well-impressed punctures, almost devoid of setosity. Remainder of ventral surface matt, covered with short, regularly distributed setae and fine, shallow punctures. Prosternal process large, with sides slightly salient, almost straight between anterior coxae; surface almost flat, shallowly punctured, covered with semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Ventrite posterior margin straight. Median lobe of aedeagus (Fig. 6 m-o) dorsoventrally flattened, terminated by triangular apex with regularly convergent, straight sides. In lateral view apex slightly bent dorsally. Ventral surface weakly convex towards apex. Setose depressions barely delimited, with few short setae. Female. Habitus in Fig. 6 a-b (NT), 6 d-f. BL = 3.4 – 3.9 mm, BW = 2.1 – 2.3 mm, PL = 1.2 – 1.3 mm, PW = 1.8 – 2.0 mm. Interocular distance = 7.7 – 8.8 % of BL. In females the eyes are smaller and more separated along midline, and mid-frontal suture clearly visible. The colour pattern is similar to the males one, but in a single studied specimen the pronotal black spots are totally missing. The fifth abdominal ventrite in females has a quite large, rounded, and deep pit. The bottom of the pit is black, glabrous, matt, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 6 i) is subangular, hook-shaped, with proximal section twisted, slender and almost uniform in section. The ampulla is brief, with the duct insertion short and straight. The sperm gland insertion is rather long and curved. The duct is uniform in size, slender, straight. At the insertion of the ductus, the bursa copulatrix shows a couple of membranous coils.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA0FC07FCE2B22F54B3D7C2.taxon	description	(Figs 7, 9 b)	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA0FC07FCE2B22F54B3D7C2.taxon	materials_examined	Type locality. Acapulco (Guerrero, Mexico). Additional material examined. EL SALVADOR, Cuscatlán: El Rosario 4. VII. 1953 & 17 - 20. VII. 1955 (5, USNMNH). Cabañas: Tejutepeque 23. VII. 2021 (1, GBIF). MEXICO, CHIAPAS: 16 km W Ocozocoautla El Aguacero 10. VI. 2009 (1, BYU). COLIMA: 6 mi w Colima 6. VIII. 1978 (1, TAMU); 9.7 km S Colima 335 m 4. VIII. 1988 (2, TAMU). GUERRERO: Acapulco coll. Flohr (3, MNHUB); 38 km W Iguala 18 à 31 ’ N 99 ° 44 ’ W 16 VII. 1995 (6, USNMNH); 10.3 mi S Iguala 23. VII. 1981 (1, TAMU); 8 km S Mazatlan 1130 m 29. VI. 1992 (1, ERPC); 4 mi W Chilpancingo 15. VII. 1984 (3, TAMU); 12 mi S Chilpancingo 12. VII. 1966 (1, TAMU); 3.3 km Cacahuamilpa 1495 m 2. VII. 1992 (2, ERPC); Grutas de Cacahuamilpa 30 km N Taxco 27. VIII. 1993 (1, MNHUB); 6 km W Teloloapan 20. IX. 1989 (1, TAMU); Tierra Colorada 12 mi N Guerrero 2600 ’ 5. VIII. 1954 (1, CNCI). JALISCO: 4 mi S El Tuito 1200 ’ hwy 20010. VIII. 1982 (2, ERPC); Chamela Vic. ESTC UANM 9 - 19. VII. 1993 (3, TAMU); Chamela Res. Sta. 13. VII. 1986 (1, BYU); 18 mi N Barra de Navidad 23. VIII. 1976 (1, BYU). MORELOS: Xochicalco Ruins 18 ° 48 ’ N 99 ° 16 ’ W 14. VII. 1995 (5, USNMNH); Cuernavaca 15. VIII. 1954 (1, USNMNH). OAXACA: 4.4 mi NE San Pedro Mixtepec 16. VII. 1985 (1, TAMU); 2.7 mi NW El Camarón 21 - 22. VII. 1974 (1, TAMU). NAYARIT: 2 km E Punta de Mita 30. VII- 2. VIII. 1991 (4, TAMU). PUEBLA: Izúcar de Matamoros-Cautla Rd. Route 160 km 129 24. VII. 1997 (1, USNMHN).	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA0FC07FCE2B22F54B3D7C2.taxon	distribution	Distribution. El Salvador, Mexico. New from El Salvador.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA0FC07FCE2B22F54B3D7C2.taxon	diagnosis	Diagnosis. The species looks very similar to G. puncturatus. It can be distinguished from the latter by the different (more transverse and more flattened) pronotal shape. Besides, punctures on pronotum are usually denser and finer and the “ gamma-shaped ” dark marking is always visible while in G. puncturatus the pronotum is uniformly yellow. Finally, the aedeagal median lobe looks rather different, above all on its apical section.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
03C01511FFA0FC07FCE2B22F54B3D7C2.taxon	description	Description of male. Habitus in Fig. 7 a-b (HT), 7 c, 7 e. BL = 3.8 – 3.9 mm, BW = 2.3 – 2.4 mm, PL = 1.3 – 1.4 mm, PW = 2.1 – 2.2 mm. Interocular distance = 0.0 – 2.6 % of BL. Head totally yellow. Labrum yellow as well. Vertex very shallowly, minutely punctured. Head surface with sparse punctures and scattered, short setosity above all between antennal insertions. Mid-cranial suture shallow, but clearly detectable between upper lobes of eyes. Upper lobes of eyes close to each other along midline but usually not really in contact. Ocular lines usually clearly visible above all along lower part of upper lobes, strictly adhering to ocular rim. marked by row of fine, shallowly impressed punctures. Ocular canthus large, angular, with surface not differentiated by remainder of frontoclypeus. Antennae (Fig. 7 h) long, yellow with last two antennomeres slightly darkened. Antennomeres 3 - 5 slender, bright, subcylindrical; 6 - 11 slender as well, dull, slightly more flattened and more diffusedly setose. Pronotum yellow with large, M-shaped brown marking extended across pronotal disc, leaving free only anterior and lateral margins. Pronotal shape elliptical, transverse, slightly flattened on disc. Lateral margins narrow, only basal half visible when seen from above, regularly curved, with maximum width at about middle. Surface rather matt, covered with fine punctures rather regularly distributed. Posterolateral impressions well impressed, narrow, obliquely arranged. Scutellum yellow, triangular with apex truncated in straight line. Surface sparsely micropunctured. Elytron yellow with brown markings of variable appearance: “ typical ” pattern consists of six brown spots, two of them, usually almost coalescent, forming a curved marking around scutellar area; three spots sitting on oblique line from humeral callus to beginning of apical clivus; one spot, further back than previous ones, close to lateral margin. All spots often coalescing in different ways giving transversal, more or less discontinued bands. Sometimes part of dark spots missing. Apical margin and suture narrowly darkened as well. Bottom of punctures brownish. Epipleuron yellow. Elytral outline with sides almost straight and slightly convergent posteriorly. Lateral margins narrow, simultaneously visible from above only along posterior half. Elytral surface moderately shiny with fine but well-impressed punctation, distinct up to posterior clivus, arranged in almost regular rows. At about one third of length usually one shallow but well-visible transverse impression, extended from first to fifth rows of punctures. Intervals flat. Postscutellar area slightly raised. Humeral callus prominent, impunctate. Epipleuron impunctate, slightly rugulose with slightly convex surface. Pygidium totally yellow Surface bright, covered with shallow punctures and short, semi-erect pale setae. Ventral parts of thorax and abdomen brown. Abdominal ventrites brown with large yellow band along margins; median projection of abdominal ventrite 1 sometimes yellow as well. Hypomera, mesoepimera and mesoepisterna shiny, almost devoid of punctures and setosity. Remainder of ventral surface of thorax and abdomen covered with rather sparse, regularly distributed setae and fine, shallow punctures. Prosternal process large, with sides slightly salient, almost straight between anterior coxae, terminated with rounded apex; surface almost flat and closely punctured, covered with long, semi-erect setae. Legs totally yellow. Median depression on fifth abdominal ventrite indistinct, hardly detectable. Ventrite posterior margin straight. Median lobe of aedeagus (Fig. 7 m-o) dorsoventrally flattened, almost parallel-sided, terminated by triangular apex with slightly concave sides, devoid of true terminal denticle. Apex slightly bent dorsally in lateral view. Ventral surface smooth, slightly convex. Setose depressions very shallowly impressed, with few, minute punctures and almost straight setae. Female. Habitus in Fig. 7 d, 7 f-g. BL = 4.0 – 4.7 mm, BW = 2.6 – 2.9 mm, PL = 1.4 – 1.5 mm, PW = 2.3 – 2.6 mm. Interocular distance = 7.5 – 8.5 % of BL. In females the eyes are smaller and more separated along midline and antennae are shorter and less distinctive than in males. Additionally, the elytral pattern is darker in some specimens, being blackish instead of brown. The fifth abdominal ventrite in females has a large, rounded, and deep pit. The bottom of the pit is glabrous, matt, slightly darkened in respect to the surrounding surface, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (Fig. 7 i) is scarcely to fairly pigmented along the base of the proximal lobe, sickle shaped with straight, slightly swollen proximal lobe. The distal lobe is slender, regularly curved and terminated with an acute apex. The ampulla is not pigmented, almost square. The duct insertion is short. The sperm gland insertion is rather long and curved. The duct is uniform in size, quite robust and rigid, straight, with a single turn close to the vasculum, then straight up to the bursa copulatrix. The insertion on the bursa copulatrix is simple, neither swollen nor pigmented. Acknowledgments – I would like to gratefully acknowledge Lee H. Herman (American Museum of Natural History, New York), Max Barclay and Michael Geiser (The Natural History Museum, London), Shawn Clark (Arthropod Collection, Monte L. Bean Life Science Museum, Brigham Young University, Provo, UT), Patrice Bouchard (Canadian National Collection of Insects, Ottawa), James H. Boone (Field Museum of Natural History, Chicago), Edward G. Riley (Texas A & M University, College Station), Kyle E. Schnepp (Florida State Collection of Arthropods, Gainesville), Mauro Daccordi (Verona), Karla Schneider and Joachim Händel (Suffrian’s Collection, Martin Luther University, Halle), Antoine Mantilleri (Muséum national d’Histoire Naturelle, Paris), Johannes Frisch, Bernd Jäger and Joachim Willers (Stiftung Museum für Naturkunde, Leibniz-Institut fur Evolutions und Biodiversitätsforschung, Berlin), Maurizio Rigato e Michele Zilioli (Museo civico di Storia Naturale, Milano), Alexander Konstantinov and Maria Lourdes Chamorro (National Museum of Natural History, Smithsonian Institution, Washington, D. C.), Katja Neven (Zoologische Staatssammlung, München) for the loan of material and, in many cases, for the kind and friendly support during my visits in their institutions. This work was supported by the Synthesys Project (http: // www. synthesys. info /) financed by the European Community Research Infrastructure Action, under GB-TAF- 3383, DE-TAF- 4995 and the Systematics Research Fund 2014 / 15 administered by the Linnean Society of London and the Systematics Association.	en	Sassi, Davide (2023): Five new species of the genus Griburius Haldeman from Central America (Coleoptera: Chrysomelidae, Cryptocephalinae). Fragmenta entomologica (Deerfield Beach, Fla.) 55 (1): 117-138, DOI: 10.13133/2284-4880/1465, URL: http://dx.doi.org/10.1111/aej.12773
