taxonID	type	description	language	source
364E87DE8677FF8D5AA6FC11DB3626C6.taxon	type_taxon	(type species: Abdosetae hainan Fu, Zhang & MacDermott, 2010, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8D5AA6FC11DB3626C6.taxon	diagnosis	Diagnosis. This genus can be distinguished from all other phrurolithid genera by the male abdomen ventrally with a double longitudinal row of long setae behind the epigastric fold (Fig. 1 B); the chelicerae have a single long spine anteriorly; the male palpal femoral protuberance is not well developed, only slightly swollen; the female epigyne lacks glandular appendages.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8D5AA6FC11DB3626C6.taxon	description	Description. Small spiders. Carapace low, smooth, highest near fovea, widest at coxae II and III. Chelicerae with three well separated promarginal and two close retromarginal teeth, with single long spine anteriorly. All femora without dorsal spine; femur I usually with three prolateral spines distally, femur II usually with two prolateral spines; tibia I usually with six proventral spines and seven retroventral spines, tibia II usually with six pairs of ventral spines; metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines. Abdomen oval, with narrow anterior dorsal scutum in dorsal view and double longitudinal row of long setae behind the epigastral fold in ventral view in males; without dorsal scutum and long setae in female. Male palp: femur only with somewhat swollen, with distal concavity; tibia with one large retrolateral tibial apophysis (prolateral margin often modified), one slender dorsal tibial apophysis and one conspicuous ventral tibial hump; tegulum convex, usually with conspicuous apophysis; sperm duct short, thick and distinct; embolus relatively long, situated at tip of tegulum. Female epigyne: epigynal plate with pair of shallow concavities or flat, copulatory openings concealed in anterior margins of concavities or exposed; copulatory ducts short, connected to two transparent bursae; spermathecae small and usually spherical.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	description	Figs 1 – 2	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	materials_examined	Type material. Holotype ♁, CHINA: Hainan Province: Baoting County, Jiuyi Mountain (25 ° 16 ′ 43.12 ″ N, 111 ° 56 ′ 45.6 ″ E, 734 m a. s. l.), 27 Apr. 2021, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	etymology	Etymology. The specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	diagnosis	Diagnosis. This new species can be separated from other congeneric species except A. ornata (Deeleman-Reinhold, 2001) by the smooth prolateral margin of the retrolateral tibial apophysis without a branch, but differing from A. ornata by: 1) the shorter embolus and tegular apophysis (vs long, cf. Fig. 1 D and fig. 678 in Deeleman-Reinhold [2001]); and 2) the presence of a conductor (vs without a conductor, cf, Fig. 1 E and fig. 678 in Deeleman-Reinhold [2001]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	description	Description. Male (Holotype): total length 1.97, carapace 1.00 long, 0.87 wide; abdomen 0.97 long, 0.65 wide. Eye sizes and interdistances: AME 0.06, ALE 0.06, PME 0.05, PLE 0.08; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.18, PME – PME 0.09, PME – PLE 0.04, PLE – PLE 0.27, ALE – PLE 0.06. EAW 0.37, CRW 0.50, EAW / CRW 0.74, CRW / CW 0.57. MOA 0.20 long, anterior width 0.16, posterior width 0.20. CH 0.10, CH / AME 1.67. Labium 0.09 long, 0.16 wide. Sternum 0.60 long, 0.50 wide. Leg measurements: Ⅰ 3.93 (1.02, 0.39, 1.10, 1.00, 0.42), II 3.22 (0.84, 0.34, 0.78, 0.79, 0.47), III 3.08 (0.80, 0.34, 0.60, 0.79, 0.55), IV 4.48 (1.17, 0.38, 1.03, 1.24, 0.66). Spination: femora without dorsal spines, femur I pl 3, femur II pl 2, tibia Ⅰ pv 6 rv 7, tibia II pv 6 rv 5, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with longitudinal black stripes along marginal of carapace. Abdomen gray, anterior with narrow yellow dorsal scutum anteriorly, laterally with black flanks, posteriorly with black transverse stripe (Fig. 1 A, B). Legs yellow, femora, tibia and metatarsi with black annuli. Palp as in Figs 1 C – F, 2. Femoral apophysis smooth, with slightly swollen near tip. Retrolateral tibial apophysis with smooth prolateral margin, without branches, tapering from base to blunt tip; dorsal tibial apophysis narrower than retrolateral tibial apophysis, with curved tip; ventral tibial hump conspicuous, round. Tegulum oval, narrower than cymbium; tegular apophysis small, with sharp tip. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus short, directed distoventrally. Conductor membranous, with thin base and wide tip. Female. Unknown.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8677FF8F5AA6F8A1DACA22BB.taxon	distribution	Distribution. China (Hainan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8675FF8E5AA6F99BD9AD21B7.taxon	type_taxon	(type species: Otacilia bijiashanica Liu, 2020, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8675FF8E5AA6F99BD9AD21B7.taxon	diagnosis	Diagnosis. This genus is similar to Otacilia in body color and chevron stripes on abdomen, but can be distinguished by the short, slight bent embolus in male palp and small copulatory openings, large and appressed bursa in female epigyne.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8675FF8E5AA6F99BD9AD21B7.taxon	description	Description. Small spiders. Carapace oval, yellow, with several distinct markings resembling flowing water droplets beside fovea. AME smaller than ALE, PME as large as or slightly smaller than PLE. Chelicerae with two (one long and another short) spine anteriorly. Femora I – IV with one dorsal spine, femur I with three prolateral spines, femur II with two prolateral spines; tibia I – II with six pairs of ventral spines (usually unpaired); metatarsus I with four pairs of ventral spines, metatarsus II with four proventral spines and three retroventral spines. Abdomen oval, with distinct chevron stripes posteriorly in dorsal view, with small and thin dorsal scutum anteriorly in male, lacking in female. Male palp: femoral apophysis not well developed, only swollen; tibia with one retrolateral apophysis and one dorsal tibial apophysis; bulb pyriform, wider than cymbium; embolus short, slight bent; conductor present or absent. Female epigyne: epigynal plate with median septum; copulatory openings small; copulatory ducts thick and short, connecting tubes thin and long, glandular appendages digitiform; bursa large and close to each other, spermathecae peanut-shaped.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	description	Figs 3 – 5	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	materials_examined	Type material. Holotype ♁, CHINA: Guangxi Zhuang Autonomous Region: Hezhou City, Fuchuan Yao Autonomous County, Chengbei Town, Xinzhai Village (24 ° 45 ′ 57.7 ″ N, 111 ° 13 ′ 53.85 ″ E, 312 m a. s. l.), 7 May 2021, leg. Yannan Mu. Paratype: 1 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	etymology	Etymology. This species name is a combination of “ brevis ” and “ spina ”, referring to the short, spine-like embolus; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	diagnosis	Diagnosis. This new species resembles A. fabiformis (Liu, Xu, Xiao, Yin & Peng, 2019) in having similarlyshaped spermathecae, but can be recognized by: 1) embolus transverse (vs directed anteriorly, cf. Fig. 4 C and fig. 6 C in Liu et al. [2019]), 2) retrolateral tibial apophysis strongly curved in ventral view, and dorsal tibial apophysis curved towards the prolateral side of the cymbium in dorsal view (vs retrolateral tibial apophysis slightly curved, dorsal tibial apophysis straight, cf. Fig. 4 C, D and fig. 6 D, E in Liu et al. [2019]), 3) median septum narrower (vs wide, cf. Fig. 4 E and fig. 7 B in Liu et al. [2019]), and 4) glandular appendages of copulatory duct small, mastoid-shaped (vs without glandular appendages, cf. Fig. 4 F and fig. 7 C in Liu et al. [2019]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	description	Description. Male (Holotype): total length 2.27, carapace 1.14 long, 0.97 wide; abdomen 1.13 long, 0.82 wide. Eye sizes and interdistances: AME 0.06, ALE 0.07, PME 0.06, PLE 0.07; AME – AME 0.02, AME – ALE 0.01, ALE – ALE 0.14, PME – PME 0.07, PME – PLE 0.04, PLE – PLE 0.28, ALE – PLE 0.05. EAW 0.38, CRW 0.54, EAW / CRW 0.70, CRW / CW 0.56. MOA 0.16 long, anterior width 0.13, posterior width 0.21. CH 0.10. CH / AME 1.67. Labium 0.14 long, 0.16 wide. Sternum 0.70 long, 0.61 wide. Leg measurements: Ⅰ 4.13 (1.10, 0.44, 1.12, 1.03, 0.44), II 3.42 (0.96, 0.39, 0.86, 0.85, 0.36), III 2.92 (0.80, 0.34, 0.61, 0.73, 0.44), IV 4.34 (1.17, 0.40, 0.93, 1.26, 0.58). Spination: femur I d 1 pl 3, femur II d 1 pl 2, femur IV d 1, tibia Ⅰ pv 6 rv 7, tibia II pv 6 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with small, dorsal scutum anteriorly and 5 black transverse chevron stripes posteriorly (Fig. 3 A, B). Leg yellow, with black annuli near tip of femora, tibia, metatarsi I – IV. Palp as in Figs 4 A – D, 5 A – C. Femoral apophysis protruding. Retrolateral tibial apophysis base wide, strongly curved in ventral view, tapering from base to tip; dorsal tibial apophysis curved towards prolateral of cymbium in dorsal view. Tegulum oval, wider than cymbium; tegular apophysis absent. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus short, needle-like, transverse. Conductor absent. Female: total length 2.79, carapace 1.15 long, 1.02 wide; abdomen 1.64 long, 1.28 wide. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.06, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.14, PME – PME 0.07, PME – PLE 0.04, PLE – PLE 0.28, ALE – PLE 0.06. EAW 0.38, CRW 0.54, EAW / CRW 0.70, CRW / CW 0.53. MOA 0.19 long, anterior width 0.12, posterior width 0.20. CH 0.09, CH / AME 1.80. Labium 0.13 long, 0.19 wide. Sternum 0.74 long, 0.65 wide. Leg measurements: Ⅰ 4.19 (1.12, 0.40, 1.18, 1.08, 0.41), II 3.41 (0.91, 0.42, 0.87, 0.85, 0.36), III 3.06 (0.80, 0.38, 0.60, 0.83, 0.45), IV 4.36 (1.18, 0.40, 0.96, 1.27, 0.55). Spination: femur I d 1 pl 3, femur II d 1 pl 2, femur IV d 1, tibia Ⅰ pv 6 rv 6, tibia II pv 5 rv 6, metatarsus Ⅰ – II pv 4 rv 3. Coloration. Carapace darker color than male, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with black triangular pattern anteriorly, with 4 chevron stripes at posterior of abdomen (Fig. 3 C, D). Leg yellows, with black annuli near tip of femora, tibiae and metatarsi I – IV. Epigyne as in Figs 4 E – F, 5 D – E. Epigynal plate weakly sclerotized. Median septum thin. Copulatory openings small, slightly separate, located at centre of plate. Copulatory ducts thick, L-shaped. Connecting tubes thinner than copulatory ducts, straight. Bursa large, asymmetrical. Glandular appendages mastoid-shaped, subtriangular. Spermathecae clavate, transverse, close to each other. Fertilization ducts located along anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8674FF815AA6FE31DB3A279F.taxon	distribution	Distribution. China (Guangxi Zhuang Autonomous Region).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	description	Fig. 6	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	materials_examined	Material examined. 2 ♀, CHINA: Hunan Province: Chenzhou City, Rucheng County, Jiulongjiang National Forest Park, Qinglong Gorge Scenic Spot (25.441143 ° N, 113.804605 ° E, 540 a. s. l.), 13 May 2018, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	diagnosis	Diagnosis. See Liu et al. 2022 a.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	description	Description. See Liu et al. 2022 a. Habitus and genitalia as in Fig. 6.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	distribution	Distribution. China (Hunan Province, Jiangxi Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8679FF835AA6F9B4DC4D27E3.taxon	discussion	Remark. This species is recorded from Hunan province for the first time.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8678FF825AA6F95BDD5D27A0.taxon	discussion	Comments: According to the original description of Otacilia pseudofabiformis, it shares similar characteristics with genus Aculithus, such as peanut-shaped spermathecae and bursae located anteriorly and touching. Therefore, we transfer it to this genus as Aculithus pseudofabiformis (Liang, Li, Yin, Li & Xu, 2021) comb. nov..	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FF7CDD2322DF.taxon	type_taxon	(type species: Phrurolithus zhejiangensis Song & Kim, 1991, by original designation and monotypic).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FF7CDD2322DF.taxon	diagnosis	Diagnosis. This genus can be distinguished from the other phrurolithid genera by the oval PME without a layer of black pigment around the eye cup, lacking distinct longitudinal and radial stripes on the carapace and chevron stripes on the abdomen.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FF7CDD2322DF.taxon	description	Description. Small spiders. Carapace oval, smooth, without distinct cervical groove and radial striae. AME rounded, PME oval, light-coloured, without black pigment, anterior eye row straight, posterior eye row procurved. Chelicerae with two (one long and another short) spines anteriorly. Femur I with two prolateral spines; tibia I with five pairs of ventral spines, tibia II with five proventral spines and four retroventral spines; metatarsus I with four proventral spines and three retroventral spines, metatarsus II with three proventral spines and two retroventral spines. Abdomen oval, with distinct chevron stripes posteriorly in dorsal view, with small and thin dorsal scutum anteriorly in male, absent in female. Male palp: femoral apophysis well developed; tibia with one retrolateral tibial apophysis; sperm duct thin and distinct; embolus short. Female epigyne: epigynal plate with copulatory atrium medially, copulatory openings small; copulatory ducts and glandular appendages slender; spermathecae round.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	description	Figs 7 – 8	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	materials_examined	Type material. Holotype ♀, CHINA: Hunan Province: Zhuzhou City, Yanling County, Shennonggu National Forest Park (26.500841 ° N, 114.069715 ° E, 1060 m a. s. l.), 9 May 2018, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	etymology	Etymology. This specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	diagnosis	Diagnosis. This new species resembles A. zhejiangensis (Song & Kim, 1991) in having similar white PME, but can be recognized by: 1) the larger bursa (vs small, cf. Fig. 7 D and fig. 3 D in Liu et al. [2020]), 2) thicker, shorter copulatory ducts and thicker connecting tubes (vs copulatory ducts thin and long, connecting tubes thin, cf. Fig. 7 D and fig. 3 D in Liu et al. [2020]), 3) clavate-shaped, long and transverse spermathecae (vs globular, cf. Fig. 7 D and fig. 3 D in Liu et al. [2020]), and 4) thicker and shorter glandular appendages (vs thin and long, cf. Fig. 7 D and fig. 3 D in Liu et al. [2020]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	description	Description. Female (Holotype): total length 2.70, carapace 1.12 long, 0.90 wide; abdomen 1.58 long, 1.10 wide. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.06, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.19, PME – PME 0.05, PME – PLE 0.04, PLE – PLE 0.23, ALE – PLE 0.03. EAW 0.35, CRW 0.52, EAW / CRW 0.67, CRW / CW 0.58. MOA 0.19 long, anterior width 0.18, posterior width 0.17. CH 0.11, CH / AME 1.57. Labium 0.14 long, 0.17 wide. Sternum 0.65 long, 0.59 wide. Leg measurements: Ⅰ 3.19 (0.91, 0.38, 0.80, 0.70, 0.40), II 2.69 (0.78, 0.35, 0.59, 0.57, 0.40), III 2.43 (0.66, 0.29, 0.49, 0.60, 0.39), IV 3.56 (0.97, 0.36, 0.83, 0.90, 0.50). Spination: femora I – IV without dorsal spines, femur I pl 2, tibia Ⅰ pv 5 rv 5, tibia II pv 5 rv 4, metatarsus Ⅰ pv 4 rv 3, metatarsus II pv 3 rv 2. Leg yellow, femora I – IV with white tip. Carapace yellow, with indistinct radial groove beside fovea. Abdomen gray, with 4 pairs of black patterns and transverse black stripe before anal tubercle (Fig. 7 A, B). Epigyne as in Figs 7 C – D, 8. Epigynal plate weakly sclerotized, without depression. Median septum thin. Copulatory openings small, separated by median septum, located at centre of plate. Copulatory ducts thick, short. Connecting tubes thinner than copulatory ducts, inflated at middle parts. Bursa large, asymmetrical. Glandular appendages short, transverse. Spermathecae clavate, transverse, close to each other. Fertilization ducts located middle of anterior spermathecal margin. Male: unknown.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867FFF855AA6FCD8DACC2703.taxon	distribution	Distribution. China (Hunan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF875AA6FC6EDA962686.taxon	etymology	Etymology. The generic name “ Beatitas ” is a Latin word meaning happiness, referring to the beautiful appearance of spiders of this genus, which elicits positive feelings. The gender is masculine.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF875AA6FC6EDA962686.taxon	diagnosis	Diagnosis. This new genus can be distinguished from all other Phrurolithidae genera by: 1) the round fovea, 2) the male with one long spine on the anterior surface of the chelicerae (Fig. 11 E, F), but the female with two pairs (one long and one short) (Fig. 12 C), 3) all femora with one dorsal spine near base, and anterior femora without prolateral spines, 4) legs yellow except black femora, and patellae, tibiae and metatarsi with black annuli, 5) embolus thick, strong, directed anteriorly, and 6) atrium diamond-shaped and bursa placed anteriorly, with connecting tubes nearly surrounding spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF875AA6FC6EDA962686.taxon	description	Description. Small spiders, total length 2.27 – 2.86 mm. Carapace oval, granular, black, widest at coxae II, highest near fovea. Cervical groove inconspicuous. Fovea round. PER slightly wider than AER. Chelicerae brown, with one long spine anteriorly in the male, two spines in female. Sternum yellow, granular, without pattern. Legs yellow to black; all femora black, with one dorsal spine near base; tibiae I – II usually with five or six pairs of strong spines; metatarsi I – II usually with four proventral spines and three retroventral spines; posterior tibia and metatarsi lack spines. Abdomen oval, dorsum black, with scutum in males. Male palp: femur with slight swelling near tip; tibia about 1 / 2 length of femur, tibia with one large retroventral apophysis and one long dorsal apophysis; tegulum oval, tegular apophysis small; conductor membranous; embolus strong, extending upwards. Female epigyne: epigynal plate weakly sclerotized, with septum; copulatory openings located at sides of diamond-shaped atrium; spermathecae globular, located posteriorly, nearly surrounded by connecting tubes; glandular appendages absent; pair of oval bursa located anteriorly.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF875AA6FC6EDA962686.taxon	type_taxon	Type species. Beatitas octomaculatus sp. nov. (Ā ỄẸĦ). Composition. Beatitas octomaculatus sp. nov..	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	description	Figs 9 – 13	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	materials_examined	Type material. Holotype ♁, CHINA: Guangxi Zhuang Autonomous Region: Hezhou City, Pinggui district, Guposhan scenic spot (24 ° 37 ′ 47.49 ″ N, 111 ° 31 ′ 46.89 ″ E, 830 m a. s. l.), 11 May 2021, leg. Yannan Mu. Paratypes: 2 ♁ 9 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	etymology	Etymology. This species name is a combination of “ octonarius ” and “ macula ”, referring to the eight light markings on the abdominal dorsum; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	diagnosis	Diagnosis. The male of this new species can be recognized by: 1) the strong and upward-extending embolus, 2) the membranous conductor, 3) the retrolateral tibial apophysis with a bifurcated tip, and curved dorsal tibial apophysis extending towards prolateral side of cymbium. The female can be recognized by: 1) the diamond-shaped atrium on the epigynal plate, 2) the widely separated copulatory openings located at the sides of the atrium, and 3) the globular spermathecae that are almost entirely surrounded by the long connecting tubes.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	description	Description. Male (Holotype): total length 2.27, carapace 1.06 long, 0.89 wide; abdomen 1.21 long, 0.84 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.05, PLE 0.06; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.15, PME – PME 0.05, PME – PLE 0.04, PLE – PLE 0.23, ALE – PLE 0.03. EAW 0.32, CRW 0.45, EAW / CRW 0.71, CRW / CW 0.51. MOA 0.15 long, anterior width 0.15, posterior width 0.15. CH 0.13, CH / AME 2.17. Labium 0.12 long, 0.17 wide. Sternum 0.61 long, 0.56 wide. Leg measurements: Ⅰ 3.22 (0.88, 0.37, 0.84, 0.78, 0.35), II 2.87 (0.80, 0.36, 0.65, 0.65, 0.41), III 2.56 (0.71, 0.32, 0.49, 0.62, 0.42), IV 3.80 (0.98, 0.36, 0.88, 0.97, 0.61). Spination: femora I – IV d 1, tibia Ⅰ pv 6 rv 5, tibia II pv 5 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Coloration. Carapace black, fovea round, with several markings resembling flowing water droplets beside fovea. Abdomen black, covered entirely by dorsal scutum, with four pairs of light markings on abdominal dorsum (Fig. 9 A, B). Legs yellow except black femora; patellae, tibiae and metatarsi I – IV with black annuli (Fig. 10). Palp as in Figs 11 A – D, 13 A – C. Femoral apophysis round, protruded near tip of femur. Retroventral tibial apophysis strong, strongly curved, extending towards ventral side, middle parts wider than base and tip bifurcated as U-shaped; dorsal tibial apophysis digitiform, base wide and tip curved towards prolateral side of cymbium. Tegulum as wide as cymbium, diamond-shaped; tegular apophysis triangular, located at retrolateral tip of tegulum. Sperm duct distinct, U-shaped, tapering from retrolateral side of tegulum to embolus. Embolus strong, tip slightly curved, beak-shaped, directed retroventrally. Conductor large, membranous, close to embolus. Female (One paratype): Total length 2.86, carapace 1.20 long, 1.02 wide; abdomen 1.66 long, 1.37 wide. Eye sizes and interdistances: AME 0.05, ALE 0.08, PME 0.06, PLE 0.07; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.17, PME – PME 0.05, PME – PLE 0.04, PLE – PLE 0.24, ALE – PLE 0.04. EAW 0.35, CRW 0.50, EAW / CRW 0.70, CRW / CW 0.49. MOA 0.16 long, anterior width 0.16, posterior width 0.16. CH 0.14, CH / AME 2.80. Labium 0.15 long, 0.19 wide. Sternum 0.71 long, 0.64 wide. Leg measurements: Ⅰ 3.65 (0.97, 0.41, 0.99, 0.88, 0.40), II 3.10 (0.82, 0.38, 0.75, 0.74, 0.41), III 2.85 (0.79, 0.38, 0.54, 0.68, 0.46), IV 4.05 (1.03, 0.37, 0.98, 1.02, 0.65). Spination: femora I – IV d 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Other characters as for male, except dorsal scutum absent (Fig. 9 C, D). Epigyne as in Figs 12 A – B, 13 D – E. Epigynal plate weakly sclerotized, posterior edge nearly straight. Atrium diamond-shaped. Median septum thin, Y-shaped. Copulatory openings large, horn-shaped, located at lateral margin of atrium. Copulatory ducts short and thick, straight and connected with oval bursa. Glandular appendages absent. Connecting tube long, thinner than copulatory ducts, nearly surrounding entire spermathecae. Bursa oval, at anterior of the plate. Spermathecae globular, slightly separated. Fertilization ducts located anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE867DFF9D5AA6F8E2DB3A215B.taxon	distribution	Distribution. China (Guangxi Zhuang Autonomous Region).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9D5AA6FE5DDD1F23D9.taxon	type_taxon	(type species: Phrurolithus coreanus Paik, 1991, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9D5AA6FE5DDD1F23D9.taxon	diagnosis	Diagnosis. This genus can be distinguished from the other phrurolithid genera by femora I – II lacking prolateral spines, the furcated retrolateral tibial apophysis, and the embolus and conductor contiguous to each other in the male palp; the female epigyne has anteriorly situated copulatory openings and the pair of membranous bursae is smaller than in other genera.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9D5AA6FE5DDD1F23D9.taxon	description	Description. Small spiders. Carapace black, smooth, widest at coxae II and III. Chelicerae with three promarginal and two retromarginal teeth, with one or two (one long and another short) spine anteriorly. Femora I – II with one dorsal spine but without prolateral spine; tibia I – II usually with unpaired ventral spines; metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines. Abdomen oval, without distinct chevron stripes in dorsal view, with large dorsal scutum in males, absent in female. Male palp: femur only somewhat swollen; tibia with two large retrolateral apophyses; tegulum with conspicuous apophysis; embolus long, conductor membranous, contiguous to each other. Female epigyne: epigynal plate with pair of copulatory openings situated anteriorly; copulatory ducts long; spermathecae round and situated posteriorly.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9D5AA6FBD2D9EA2633.taxon	discussion	Comments. We reviewed the type specimens of Phrurolithus subnigerus Fu, Chen & Zhang, 2016, currently placed in Corealithus, and found that its morphological and genitalic characters are the same as those of Otacilia nigerus (Yin, 2012), particularly the shape of the copulatory ducts, connecting tubes, bursa, spermathecae, fertilization ducts, and copulatory openings. We therefore suggest that C. subnigerus syn. nov. should be a junior synonym of O. nigera. Furthermore, O. nigera should be transferred to Corealithus as a new combination, Corealithus nigerus (Yin, 2012) comb. nov..	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9C5AA6F9B5DC662193.taxon	type_taxon	(type species: Grandilithus anyuan Liu & Li, 2022, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9C5AA6F9B5DC662193.taxon	diagnosis	Diagnosis. This genus can be distinguished from the other phrurolithid genera by the carapace white or yellow, with several distinct markings resembling flowing water droplets beside the fovea; abdomen yellow to black, with a short and thin dorsal scutum anteriorly, without chevron stripes posteriorly in dorsal view; the male palpal femur has a well-developed apophysis distally and tibia with single retrolateral apophysis; the female epigyne with gobletshaped median septum and nearly globular-shaped copulatory ducts.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8667FF9C5AA6F9B5DC662193.taxon	description	Description. Medium-sized spiders. Carapace flat, white to yellow, with several distinct markings resembling flowing water droplets beside fovea. Chelicerae with two (one long and another short) spine anteriorly. Femora I – II with one or two dorsal spines; femur I with five to seven prolateral spines and femur II with two to five prolateral spines; tibia I – II usually with more than seven pairs of ventral spines (usually unpaired); metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines. Abdomen oval, without distinct chevron stripes in dorsal view, with small and thin dorsal scutum in male, absent in female. Male palp: femoral apophysis well developed; tibia with one retrolateral apophysis; bulb oval, narrower than cymbium; embolus long, and curved and hook-shaped; conductor absent. Female epigyne: epigynal plate with distinct median septum, copulatory openings small, situated medially on plate; copulatory ducts nearly globular-shaped, glandular appendages digitiform or mastoid-shaped; spermathecae round.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9C5AA6FE15DACA2571.taxon	description	Figs 14 – 15	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9C5AA6FE15DACA2571.taxon	materials_examined	Material examined. 1 ♁ 7 ♀, CHINA: Jiangxi Province: Ganzhou City, Shashi Town, Fengshan National Forest Park (25 ° 44 ′ 33.73 ″ N – 25 ° 45 ′ 13.03 ″ N, 115 ° 00 ′ 36.08 ″ E – 115 ° 00 ′ 56.88 ″ E, 780 – 900 m a. s. l.), 21 Jul. 2022, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9C5AA6FE15DACA2571.taxon	diagnosis	Diagnosis. See Liu et al. (2022).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9C5AA6FE15DACA2571.taxon	description	Description. Female: total length 4.27, carapace 1.80 long, 1.53 wide; abdomen 2.47 long, 1.69 wide. Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.08, PLE 0.10; AME – AME 0.06, AME – ALE 0.02, ALE – ALE 0.33, PME – PME 0.13, PME – PLE 0.06, PLE – PLE 0.43, ALE – PLE 0.10. EAW 0.57, CRW 0.76, EAW / CRW 0.75, CRW / CW 0.50. MOA 0.32 long, anterior width 0.28, posterior width 0.30. CH 0.13, CH / AME 1.08. Labium 0.17 long, 0.25 wide. Sternum 1.02 long, 0.86 wide. Leg measurements: Ⅰ 8.68 (2.14, 0.63, 2.56, 1.95, 1.40), II 6.80 (1.81, 0.52, 1.87, 1.42, 1.18), III 5.85 (1.53, 0.54, 1.28, 1.56, 0.94), IV 8.93 (2.38, 0.63, 2.09, 2.53, 1.30). Spination: femora II – IV d 1, femur Ⅰ pl 6, femur II pl 4, tibia Ⅰ pv 10 rv 10, tibia II pv 9 rv 9, metatarsus Ⅰ pv 5 rv 4, metatarsus II pv 4 rv 3. Legs yellow. Carapace brown, with several shapes resembling flowing water droplets beside fovea. Abdomen black, without pattern (Fig. 14 C, D). Epigyne as in Fig. 15 D – E. Epigynal plate sclerotized, posterior edge nearly straight. Median septum narrow. Copulatory openings small, close to each other, located at centre of plate. Copulatory ducts thick, curved and connected with bursa. Glandular appendages mastoid-shaped. Connecting tubes thinner than copulatory ducts, connected with spermathecae. Bursa large, transparent. Spermathecae globular, located at posterior of plate. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9C5AA6FE15DACA2571.taxon	distribution	Distribution. China (Jiangxi Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9F5AA6FA4ADDD427ED.taxon	type_taxon	(type species: Otacilia armatissima Thorell, 1897, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9F5AA6FA4ADDD427ED.taxon	diagnosis	Diagnosis. This genus can be distinguished from Phrurolithus by having chevron stripes posteriorly on abdomen and two strong, thin tibial apophyses in male palp. It differs from Abdosetae by the absence of a longitudinal row of long setae behind the epigastric fold in males. They also can be separated by lacking a tegular sheet (vs with tegular sheet in Lingulatus Mu & Zhang, 2022) and prolateral tegular apophysis (vs with prolateral tegular apophysis in Xilithus Liu & Li, 2022). This genus is similar to Aculithus, but can be distinguished by the long and curved embolus in the male palp and the distinct copulatory openings, and the large and separated bursae, in the female epigyne.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8666FF9F5AA6FA4ADDD427ED.taxon	description	Description. Small to medium spiders. Carapace oval, yellow to brown, with different pattern in dorsal view. Chelicerae with two (one long and another short) spine anteriorly. Femora I – II with one or two dorsal spines, femora III – IV with one dorsal spine; femur I – II with three to seven prolateral spines; tibia I – II with more than six pairs of ventral spines (usually unpaired); metatarsus I with four pairs of ventral spines, metatarsus II with four proventral spines and three retroventral spines. Abdomen oval, with distinct chevron stripes posteriorly in dorsal view, with small and thin dorsal scutum anteriorly in male, absent in female. Male palp: femoral apophysis well developed; tibia with one retrolateral apophysis and one dorsal tibial apophysis (sometimes absent); bulb oval, as wide as cymbium or wider than cymbium; embolus long, needle-like or hook shaped; conductor present or absent. Female epigyne: median septum present or absent; copulatory openings distinct; copulatory ducts and connecting tubes variable, glandular appendages digital or mastoid-shaped; bursa membranous, spermathecae with different shapes.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8664FF935AA6F8E1DACC217F.taxon	description	Figs 16 – 18	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8664FF935AA6F8E1DACC217F.taxon	materials_examined	Material examined. 1 ♁ 5 ♀, CHINA: Hunan Province: Chenzhou City, Yizhang County, Mangshan National Forest Park, Jiangjunzhai Scenic Spot (24.947814 ° N, 112.941690 ° E, 1262 m a. s. l.), 13 Nov. 2020, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8664FF935AA6F8E1DACC217F.taxon	diagnosis	Diagnosis. This species is different from other Otacilia species by the following characters: 1) the absence of a retrolateral tibial apophysis, 2) tibia with a round, large ventral apophysis, 3) tegular apophysis thick, clavate-shaped in ventral view, 4) cymbium with a large, arch-shaped apophysis at the base retrolaterally, 5) copulatory ducts comma-shaped, and 6) connecting tubes absent.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8664FF935AA6F8E1DACC217F.taxon	description	Description. Male: total length 3.41, carapace 1.77 long, 1.55 wide; abdomen 1.64 long, 1.00 wide. Eye sizes and interdistances: AME 0.11, ALE 0.12, PME 0.09, PLE 0.11; AME – AME 0.04, AME – ALE 0.02, ALE – ALE 0.27, PME – PME 0.12, PME – PLE 0.06, PLE – PLE 0.43, ALE – PLE 0.09. EAW 0.51, CRW 0.77, EAW / CRW 0.66, CRW / CW 0.50. MOA 0.28 long, anterior width 0.25, posterior width 0.31. CH 0.17, CH / AME 1.55. Labium 0.18 long, 0.24 wide. Sternum 1.02 long, 0.84 wide. Leg measurements: Ⅰ 7.47 (1.95, 0.67, 2.26, 1.77, 0.82), II 5.97 (1.59, 0.57, 1.58, 1.39, 0.84), III 4.43 (1.23, 0.43, 1.01, 1.06, 0.70), IV 7.21 (1.97, 0.55, 1.72, 2.05, 0.92). Spination: femur I d 2 pl 4, femur II d 1 pl 2, femur III – IV d 1, tibia Ⅰ pv 7 rv 8, tibia II pv 7 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Legs yellow. Carapace black, with several shapes resembling flowing water droplets beside fovea. Abdomen black, without pattern, a large dorsal scutum in dorsal view (Fig. 16 A, B). Palp as in Fig. 17. Femoral apophysis weakly protruded, near middle part of femur. Tibia with large, round apophysis ventrally. Retrolateral tibial apophysis absent; prolateral tibial apophysis pronounced, round; dorsal tibial apophysis with wide base, tip curved, goose neck-shaped. Cymbium with an arc-shaped apophysis at retrolateral base. Tegulum thinner than cymbium, oval; tegular apophysis long, upwards, clavate in ventral view, thick in retrolateral view. Sperm duct distinct, U-shaped. Embolus short, spike-like, extending towards ventral side in retrolateral view. Conductor absent. Female. See Liu et al. 2019.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8664FF935AA6F8E1DACC217F.taxon	distribution	Distribution. China (Hunan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	description	Figs 19 – 21	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	materials_examined	Type material. Holotype ♁, CHINA: Yunnan Province: Hani-Yi Autonomous Prefecture of Honghe, Pingbian Miao Autonomous County, Daweishan National Forest Park (22 ° 54.317 ′ N, 103 ° 41.961 ′ E, 2124 m a. s. l.), 28 Oct. 2016, leg. Yannan Mu. Paratypes: 4 ♁ 5 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	etymology	Etymology. This species name is a combination of “ allocot ” and “ manubrium ”, referring to the long pedicel; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	diagnosis	Diagnosis. This new species resembles O. taoyuan (Fu, Chen & Zhang, 2016) in having a similar femoral apophysis, but can be recognized by: 1) slender embolus (vs short and thick, cf. Fig. 20 C and figs 9 B, 10 B in Fu et al. [2016]), 2) shorter retrolateral tibial apophysis without distinct curvature (vs long and curved tip, cf. Figs 20 B, D and figs 9 E – F, 10 C in Fu et al. [2016]), 3) curved tip of dorsal tibial apophysis (vs not curved, cf. Fig. 20 B and figs 9 F, 10 C in Fu et al. [2016]), 4) digit-shaped glandular appendages (vs absent, cf. Fig. 20 F and figs 9 H, 10 E in Fu et al. [2016]), and (5) closely situated spermathecae (vs separate, cf. Fig. 20 F and figs 9 H, 10 E in Fu et al. [2016]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	description	Description. Male (Holotype): total length 3.81, carapace 1.82 long, 1.55 wide; abdomen 1.99 long, 1.27 wide. Eye sizes and interdistances: AME 0.12, ALE 0.11, PME 0.10, PLE 0.11; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.29, PME – PME 0.12, PME – PLE 0.07, PLE – PLE 0.50, ALE – PLE 0.09. EAW 0.65, CRW 0.84, EAW / CRW 0.77, CRW / CW 0.54. MOA 0.31 long, anterior width 0.27, posterior width 0.35. CH 0.15, CH / AME 1.25. Labium 0.15 long, 0.27 wide. Sternum 1.02 long, 0.94 wide. Leg measurements: Ⅰ 6.59 (1.67, 0.72, 1.83, 1.58, 0.79), II 5.62 (1.55, 0.66, 1.38, 1.25, 0.78), III 4.97 (1.32, 0.61, 1.00, 1.26, 0.78), IV 6.08 (1.95, 0.65, 1.53, 0.98, 0.97). Spination: femur I d 2 pl 4, femur II d 1 pl 2, femora III – IV d 1, tibia Ⅰ pv 6 rv 7, tibia II pv 6 rv 6, metatarsi Ⅰ – II pv 4 rv 3. Coloration. Carapace blackish-brown, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with small narrow dorsal scutum, tip with large black spot, posterior with four black stripes (Fig. 19 A, B). Legs yellow, with black annuli near tips of femora Ⅰ – IV. Palp as in Figs 20 A – D, 21 A – C. Femoral apophysis protrude, near at middle part of femur. Tibia short, about half length of femur. Retrolateral tibial apophysis base wide, tip narrow and slightly curved, leaf-shaped in retrolateral view; dorsal tibial apophysis base wide, tip lamellar-shaped and curved in retrolateral view. Tegulum wider than cymbium, posterior edge nearly straight; tegular apophysis small, arc-shaped, located centrally at tip of tegulum. Sperm duct wide and distinct, tapering from retrolateral of tegulum to embolus. Embolus tapering from base to tip, slightly curved, extending towards retrolateral side of cymbium. Conductor absent. Female (Paratype): total length 4.09, carapace 1.85 long, 1.56 wide; abdomen 2.24 long, 1.57 wide. Eye sizes and interdistances: AME 0.12, ALE 0.12, PME 0.11, PLE 0.11; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.29, PME – PME 0.11, PME – PLE 0.06, PLE – PLE 0.47, ALE – PLE 0.08. EAW 0.60, CRW 0.83, EAW / CRW 0.72, CRW / CW 0.53. MOA 0.31 long, anterior width 0.26, posterior width 0.34. CH 0.13, CH / AME 1.08. Labium 0.14 long, 0.29 wide. Sternum 1.08 long, 1.01 wide. Leg measurements: Ⅰ 6.85 (1.75, 0.75, 1.96, 1.62, 0.77), II 5.89 (1.54, 0.69, 1.48, 1.37, 0.81), III 5.01 (1.23, 0.62, 1.07, 1.29, 0.80), IV 5.88 (1.61, 0.46, 1.30, 1.55, 0.96). Spination: femur I d 2 pl 4, femur II d 1 pl 2, femur III d 1, femur IV without spine, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Others characters as for male, except color lighter and dorsal scutum absent (Fig. 19 C, D). Epigyne as in Figs 20 E – F, 21 D – E. Epigynal plate weakly sclerotized. Median septum narrow. Copulatory openings small and round, widely separated. Copulatory ducts short and thick, connected to bursa. Connecting tubes long and narrower than copulatory ducts, strongly curved as in spectacles frame. Glandular appendages slightly long, digit-shaped. Bursa large, transparent, irregular-shaped. Spermathecae clavate, close to each other. Fertilization ducts located anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8669FF975AA6FBA3DAC12123.taxon	distribution	Distribution. China (Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	description	Figs 22 – 24	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	materials_examined	Type material. Holotype ♁, CHINA: Guangxi Zhuang Autonomous Region: Hezhou City, Pinggui district, Guposhan scenic spot (24 ° 37 ′ 47.49 ″ N, 111 ° 31 ′ 46.89 ″ E, 830 m a. s. l.), 11 May 2021, leg. Yannan Mu. Paratypes: 3 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	etymology	Etymology. The specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	diagnosis	Diagnosis. This new species resembles O. celata (Fu, Chen & Zhang, 2016) in having a strong retrolateral tibial apophysis and dorsal tibial apophysis, but can be recognized by: 1) the upward bending embolus (vs not bending, cf. Fig. 23 C and figs 1 D, 2 B in Fu et al. [2016]), 2) curved, beak-shaped retrolateral tibial apophysis and dorsal tibial apophysis (vs not curved, cf. Fig. 23 B and figs 1 E, 2 C in Fu et al. [2016]), 3) triangular tegular apophysis (vs semicircular, cf. Fig. 23 C and figs 1 D, 2 B in Fu et al. [2016]), 4) crack-shaped copulatory openings (vs round, cf. Fig. 23 E and fig. 2 D in Fu et al. [2016]), and 5) the longer connecting tubes (vs short, cf Fig. 23 F and figs 1 G, 2 E in Fu et al. [2016]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	description	Description. Male (Holotype): total length 2.13, carapace 1.05 long, 0.92 wide; abdomen 1.08 long, 0.73 wide. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.06, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.15, PME – PME 0.07, PME – PLE 0.04, PLE – PLE 0.27, ALE – PLE 0.05. EAW 0.37, CRW 0.52, EAW / CRW 0.71, CRW / CW 0.57. MOA 0.17 long, anterior width 0.14, posterior width 0.20. CH 0.11, CH / AME 2.20. Labium 0.13 long, 0.17 wide. Sternum 0.66 long, 0.62 wide. Leg measurements: Ⅰ 3.65 (1.00, 0.34, 1.00, 0.90, 0.41), II 3.18 (0.87, 0.37, 0.77, 0.75, 0.42), III 2.83 (0.76, 0.34, 0.62, 0.70, 0.41), IV 3.88 (0.99, 0.34, 0.87, 1.14, 0.54). Spination: femur I d 1 pl 3, femur II d 1 pl 1, femur IV d 1, tibia Ⅰ pv 6 rv 7, tibia II pv 6 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with several shapes resembling flowing water droplets beside fovea. Abdomen off-white, with small narrow dorsal scutum; dorsum with pair of black triangular patterns and four chevron stripes posteriorly (Fig. 22 A, B). Palp as in Figs 23 A – D, 24 A – C. Femoral apophysis well developed, round, near tip of femur. Retrolateral tibial apophysis strong, slightly curved, digital tip curved towards tegulum; dorsal tibial apophysis smaller than retrolateral tibial apophysis, with broad base and thin, beak-shaped tip towards cymbium. Tegulum wider than cymbium, pyriform-shaped; tegular apophysis triangular, located at retrolateral tip of tegulum. Sperm duct distinct, U-shaped, tapering from retrolateral of tegulum to embolus. Embolus short, base wide, tip spine-shaped and upwards. Conductor absent. Female (Paratype): total length 2.22, carapace 1.03 long, 0.88 wide; abdomen 1.19 lo 0 ng, 0.86 wide. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.14, PME – PME 0.06, PME – PLE 0.04, PLE – PLE 0.25, ALE – PLE 0.05. EAW 0.35, CRW 0.50, EAW / CRW 0.70, CRW / CW 0.57. MOA 0.15 long, anterior width 0.13, posterior width 0.18. CH 0.08, CH / AME 1.60. Labium 0.11 long, 0.16 wide. Sternum 0.66 long, 0.58 wide. Leg measurements: Ⅰ 3.45 (0.91, 0.37, 0.94, 0.86, 0.37), II 2.95 (0.79, 0.40, 0.68, 0.70, 0.38), III 2.59 (0.67, 0.33, 0.50, 0.67, 0.42), IV 3.72 (0.99, 0.34, 0.82, 1.03, 0.54). Spination: femur I d 1 pl 3, femur II d 1 pl 1, femur IV d 1, tibia Ⅰ pv 6 rv 7, tibia II pv 6 rv 5, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as for male, except dorsal scutum absent, color darker and body slightly larger (Fig. 22 C, D). Epigyne as in Figs 23 E – F, 24 D – E. Epigynal plate weakly sclerotized, posterior edge W-shaped. Median septum absent. Copulatory openings transverse, crack-shaped, located at middle of plate. Copulatory ducts short and thick, curved and connected with large bursa. Connecting tubes long and curved. Spermathecae globular, close to each other. Fertilization ducts located at posterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE866DFFA85AA6FE8ADB222443.taxon	distribution	Distribution. China (Guangxi Zhuang Autonomous Region)	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	description	Figs 25 – 27	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	materials_examined	Type material. Holotype ♁, CHINA: Yunnan Province: Baoshan City, Longyang District, Lujiang Town (24.928124 ° N, 98.797474 ° E, 1927 m a. s. l.), 23 Nov. 2017, leg. Yannan Mu. Paratypes: 2 ♁ 5 ♀, with same data as holotype; 7 ♁ 6 ♀, Taibaoshan Forest Park (25.118748 ° N, 99.149809 ° E, 1787 m a. s. l.), 21 Nov. 2017, leg. Yannan Mu; 2 ♁ 4 ♀, Taibaoshan Forest Park (25.116509 ° N, 99.153918 ° E, 1759 m a. s. l.), 20 Nov. 2017, leg. Yannan Mu; 1 ♁ 1 ♀, Longling County, Xiaoheishan Nature Reserve (24.556765 ° N, 98.821807 ° E, 1840 m a. s. l.), 28 Nov. 2017, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	etymology	Etymology. This specific name is derived from the Latin word “ horizon ”, referring to the transverse embolus; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	diagnosis	Diagnosis. This new species resembles O. yangi Zhang, Fu & Zhu, 2009 in having a similar position and shape of the copulatory openings, but can be recognized by: 1) presence of a triangular apophysis prolaterally at the base of the retrolateral tibial apophysis (vs absent, cf. Fig. 26 B and fig. 4 in Zhang et al. [2009]), 2) the transverse embolus (vs upwards, cf. Fig. 26 C and fig. 3 in Zhang et al. [2009]), 3) the thicker copulatory ducts (vs thin, cf. Fig. 26 E and fig. 7 in Zhang et al. [2009]), and 4) the absence of glandular appendages (vs clavate-shaped, cf. Fig. 26 F and fig. 8 in Zhang et al. [2009]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	description	Description. Male (Holotype): total length 4.10, carapace 2.01 long, 1.78 wide; abdomen 2.09 long, 1.29 wide. Eye sizes and interdistances: AME 0.15, ALE 0.15, PME 0.12, PLE 0.15; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.34, PME – PME 0.14, PME – PLE 0.08, PLE – PLE 0.54, ALE – PLE 0.10. EAW 0.72, CRW 0.93, EAW / CRW 0.77, CRW / CW 0.52. MOA 0.36 long, anterior width 0.32, posterior width 0.39. CH 0.16, CH / AME 1.07. Labium 0.21 long, 0.29 wide. Sternum 1.12 long, 0.98 wide. Leg measurements: Ⅰ 8.16 (2.01, 0.71, 2.45, 1.87, 1.12), II 7.18 (1.98, 0.76, 1.89, 1.53, 1.02), III 5.91 (1.63, 0.67, 1.35, 1.40, 0.89), IV 9.39 (2.64, 0.87, 2.22, 2.48, 1.18). Spination: femur I d 2 pl 4, femur II d 1 pl 2, femur IV d 1, tibia Ⅰ pv 8 rv 8, tibia II pv 7 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with yellow dorsal scutum and black pattern besides dorsal scutum anteriorly and three black transverse stripes posteriorly (Fig. 25 A, B). Legs yellow. Palp as in Figs 26 A – D, 27 A – C. Femoral apophysis well-developed, located near middle of femur. Prolateral tibial apophysis distinct, semicircular. Retrolateral tibial apophysis base wide, digitiform tip, with small triangular apophysis prolaterally at base and large triangular apophysis retrolaterally at base. Tegulum pyriform; tegular apophysis absent. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus long, transverse, extending to retrolateral margin of cymbium. Conductor absent. Female (Paratype): total length 5.58, carapace 2.21 long, 1.94 wide; abdomen 3.37 long, 2.08 wide. Eye sizes and interdistances: AME 0.15, ALE 0.16, PME 0.13, PLE 0.15; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.35, PME – PME 0.14, PME – PLE 0.09, PLE – PLE 0.58, ALE – PLE 0. 11. EAW 0.76, CRW 1.03, EAW / CRW 0.74, CRW / CW 0.53. MOA 0.38 long, anterior width 0.34, posterior width 0.41. CH 0.14, CH / AME 0.93. Labium 0.18 long, 0.34 wide. Sternum 1.29 long, 1.15 wide. Leg measurements: Ⅰ 9.35 (2.50, 0.77, 2.81, 2.08, 1.19), II 7.53 (2.04, 0.74, 2.02, 1.66, 1.07), III 6.27 (1.63, 0.71, 1.38, 1.60, 0.95), IV 9.28 (2.64, 0.82, 2.23, 2.58, 1.01). Spination: femur I d 2 pl 5, femur II d 1 pl 2, femora III – IV d 1, tibia Ⅰ pv 9 rv 9, tibia II pv 8 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as for male, except with three black chevron stripes and two transverse black stripes, and dorsal scutum absent (Fig. 25 C, D). Epigyne as in Figs 26 E – F, 27 D – E. Epigynal plate sclerotized. Median septum V-shaped. Copulatory openings round, located at middle parts of plate, separated by about 3 times diameter of spermathecae. Copulatory ducts extremely thick, curved and connecting with bursa. Connecting tubes thinner than copulatory ducts, straight, close to each other. Bursa oval, transparent. Spermathecae oval, separated by less than their diameter. Fertilization ducts located on anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8652FFAC5AA6FB25DAC1215B.taxon	distribution	Distribution. China (Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	description	Figs 28 – 30	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	materials_examined	Type material. Holotype ♁, CHINA: Yunnan Province: Dali Bai Autonomous Prefecture, Nanjian Yi Autonomous County, Fenghuang Mountain (24 ° 53 ′ 33.35 ″ N, 100 ° 19 ′ 42.67 ″ E, 2294 m a. s. l.), 26 Jul. 2021, leg. Yannan Mu. Paratypes: 6 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	etymology	Etymology. The specific name is derived from the Latin word “ impletus ”, referring to the satisfaction derived from naming this species; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	diagnosis	Diagnosis. This new species resembles O. celata in having round copulatory openings and globular spermathecae, but can be recognized by: 1) the curved, arc-shaped retrolateral tibial apophysis (vs not curved, cf. Fig. 29 C and fig. 2 B in Fu et al. 2016), 2) the longer, upwards-extending embolus curved towards the retrolateral side of the cymbium (vs short, not curved, cf. Fig. 29 C and fig. 2 B in Fu et al. 2016), 3) the smaller, ridge-shaped tegular apophysis (vs large, triangular, cf. Fig. 29 C and fig. 1 D in Fu et al. 2016), 4) thinner, longer connecting tubes (vs thick and short, cf. Fig. 29 F and fig. 1 G in Fu et al. 2016).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	description	Description. Male (Holotype): total length 2.40, carapace 1.13 long, 1.11 wide; abdomen 1.27 long, 0.84 wide. Eye sizes and interdistances: AME 0.06, ALE 0.08, PME 0.05, PLE 0.08; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.16, PME – PME 0.05, PME – PLE 0.05, PLE – PLE 0.24, ALE – PLE 0.05. EAW 0.38, CRW 0.55, EAW / CRW 0.69, CRW / CW 0.50. MOA 0.19 long, anterior width 0.15, posterior width 0.16. CH 0.11, CH / AME 1.83. Labium 0.06 long, 0.18 wide. Sternum 0.57 long, 0.53 wide. Leg measurements: Ⅰ 4.24 (1.18, 0.47, 1.08, 0.95, 0.56), II 3.66 (1.02, 0.42, 0.84, 0.80, 0.58), III 3.25 (0.87, 0.40, 0.64, 0.80, 0.54), IV 4.95 (1.31, 0.45, 1.13, 1.30, 0.76). Spination: femora I – IV d 1, femur I pl 3, femur II pl 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Coloration. Carapace dark brown, with several shapes resembling flowing water droplets beside fovea. Abdomen black, dorsum nearly entirely covered by scutum, with pair of oval yellow markings near middle and two W-shaped stripes posteriorly (Fig. 28 A, B). Legs yellow, with black annuli near tip of femora and tibiae. Palp as in Figs 29 A – D, 30 A – C. Femoral apophysis protruding slightly, round, at middle parts of femur. Retrolateral tibial apophysis strongly curved, arc-shaped in ventral view; dorsal tibial apophysis smaller than retrolateral tibial apophysis, with broad base and thin tip, leaf-shaped in dorsal view. Tegulum slightly wider than cymbium, oval; tegular apophysis small, triangular, located at retrolateral tip of tegulum. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus short, base wide, triangular, tip curved towards retrolateral side of cymbium tip. Conductor narrow, transparent. Female (One paratype): total length 2.68, carapace 1.29 long, 1.12 wide; abdomen 1.39 long, 1.10 wide. Eye sizes and interdistances: AME 0.06, ALE 0.10, PME 0.06, PLE 0.08; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.18, PME – PME 0.07, PME – PLE 0.06, PLE – PLE 0.31, ALE – PLE 0.06. EAW 0.42, CRW 0.60, EAW / CRW 0.70, CRW / CW 0.57. MOA 0.420 long, anterior width 0.17, posterior width 0.19. CH 0.10, CH / AME 1.67. Labium 0.14 long, 0.19 wide. Sternum 0.82 long, 0.69 wide. Leg measurements: Ⅰ 4.64 (1.51, 0.47, 1.15, 0.97, 0.54), II 3.67 (1.01, 0.42, 0.88, 0.83, 0.53), III 3.25 (0.86, 0.38, 0.68, 0.80, 0.53), IV 4.82 (1.29, 0.46, 1.09, 1.21, 0.77). Spination: femora I – IV d 1, femur I pl 3, femur II pl 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Carapace color and pattern same as male, abdomen black, without dorsal scutum, covered by fluff, several white spots and stripes faint in dorsal view (Fig. 28 C, D). Epigyne as in Figs 29 E – F, 30 D – E. Epigynal plate weakly sclerotized, posterior edge straight. Median septum absent. Copulatory openings round, located at middle of plate. Copulatory ducts not visible. Glandular appendages short, digitiform. Connecting tubes V-shaped, shorter than fertilization duct. Bursa large, transparent, balloon-shaped. Spermathecae globular, close to each other. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8656FFA15AA6FE5DDAC12443.taxon	distribution	Distribution. China (Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	description	Figs 31 – 33	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province: Chengdu City, Chongzhou County, Wenjingjiang Town, Anzihe Conservation Area, Maliugou (30 ° 46.790 ′ N, 103 ° 13.530 ′ E, 1692 m a. s. l.), 3 Oct. 2016, leg. Luyu Wang. Paratypes: 17 ♁ 19 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	etymology	Etymology. The species name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	diagnosis	Diagnosis. This new species resembles O. hamata (Wang, Zhang & Zhang, 2012) in having a similar embolus and dorsal tibial apophysis, but can be recognized by: 1) the slightly protruding femoral apophysis (vs strongly protruded, cf. Fig. 32 A and fig. 9 E in Wang et al. [2012]), 2) the wider retrolateral tibial apophysis, with the tip of the prolateral edge folding inwards (vs thin and not folding, cf. Fig. 32 B – D and fig. 9 F in Wang et al. [2012]), 3) the closely situated spermathecae (vs distinctly separated, cf. Fig. 32 F and fig. 9 I in Wang et al. [2012]), and 4) the smaller, round copulatory openings (vs large and arch-shaped, cf. Fig. 32 E and fig. 9 H in Wang et al. [2012]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	description	Description. Male (Holotype): total length 2.52, carapace 1.23 long, 1.08 wide; abdomen 1.29 long, 0.91 wide. Eye sizes and interdistances: AME 0.06, ALE 0.09, PME 0.05, PLE 0.08; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.16, PME – PME 0.06, PME – PLE 0.07, PLE – PLE 0.30, ALE – PLE 0.04. EAW 0.40, CRW 0.65, EAW / CRW 0.62, CRW / CW 0.53. MOA 0.18 long, anterior width 0.15, posterior width 0.17. CH 0.12, CH / AME 2.00. Labium 0.14 long, 0.18 wide. Sternum 0.77 long, 0.69 wide. Leg measurements: Ⅰ 4.16 (1.17, 0.44, 1.12, 0.97, 0.46), II 3.62 (1.04, 0.43, 0.85, 0.87, 0.43), III 3.36 (0.91, 0.38, 0.65, 0.85, 0.57), IV 4.75 (1.22, 0.47, 1.04, 1.26, 0.76). Spination: femora I – IV d 1, femur I pl 3, femur II pl 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Coloration. Carapace yellow, radial striae distinct beside fovea. Abdomen off-white, with a small and narrow dorsal scutum, a pair of black triangular pattern and four chevron stripes posteriorly (Fig. 31 A, B). Legs yellow. Palp as in Figs 32 A – D, 33 A – C. Femoral apophysis protruding, round, located near middle of femur. Tibia short, about half length of femur. Retrolateral tibial apophysis wide, prolateral margin of tip folding inward; dorsal tibial apophysis slender, curved towards prolateral side of cymbium in dorsal view. Tegulum wider than cymbium, posterior edge straight; tegular apophysis small, triangular, located at retrolateral tip of tegulum. Sperm duct wide and distinct, occupying more than half length of tegulum, tapering from retrolateral side of tegulum to embolus. Embolus short, base triangular, tip transverse and extending towards retrolateral side of cymbium. Conductor subtriangular, transparent. Female (Paratype): total length 2.66, carapace 1.25 long, 1.09 wide; abdomen 1.41 long, 0.96 wide. Eye sizes and interdistances: AME 0.06, ALE 0.09, PME 0.05, PLE 0.08; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.16, PME – PME 0.06, PME – PLE 0.05, PLE – PLE 0.29, ALE – PLE 0.05. EAW 0.39, CRW 0.61, EAW / CRW 0.64, CRW / CW 0.56. MOA 0.18 long, anterior width 0.15, posterior width 0.18. CH 0.09, CH / AME 1.50. Labium 0.16 long, 0.20 wide. Sternum 0.83 long, 0.75 wide. Leg measurements: Ⅰ 4.21 (1.15, 0.46, 1.15, 0.94, 0.51), II 3.50 (1.02, 0.47, 0.84, 0.79, 0.38), III 3.35 (0.87, 0.43, 0.64, 0.84, 0.57), IV 4.71 (1.22, 0.46, 1.02, 1.27, 0.74). Spination: femora I – IV d 1, femur I pl 3, femur II pl 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ – II pv 4 rv 3. Carapace color darker than male, radial striae indistinct, abdomen offwhite, without dorsal scutum, anterior half covered by fluff, several chevron stripes at posterior (Fig. 31 C, D). Epigyne as in Figs 32 E – F, 33 D – E. Epigynal plate weakly sclerotized. Median septum narrow. Copulatory openings small and round, located near middle of plate. Copulatory ducts thick and curved, connecting with bursa. Connecting tubes short, narrower than copulatory ducts. Glandular appendages absent. Bursa large, transparent. Spermathecae globular, close to each other. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865BFFA55AA6FB25DADD20C6.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	description	Figs 34 – 36	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	materials_examined	Type material. Holotype ♁, CHINA: Guangxi Zhuang Autonomous Region: Guilin City, Xing'an County, Gaozhai village (25 ° 53 ' 28.28 ″ N, 110 ° 26 ' 04.78 ″ E, 1964 a. s. l.), 12 Aug. 2018, leg. Yannan Mu. Paratypes: 3 ♁ 7 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	etymology	Etymology. This specific name is the Latin word for “ spine ”, referring to the small spine at the base of the prolateral margin of the retrolateral tibial apophysis in retrolateral view.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	diagnosis	Diagnosis. This new species resembles O. triangula Mu, Jin & Zhang, 2022 in having a similarly shaped embolus and spermathecae, but can be recognized by: 1) the longer retrolateral tibial apophysis, with a spine-shaped apophysis at the base of the prolateral margin and a mamillary process at the base of the retrolateral margin (vs short, apophysis absent, cf. Fig. 35 B, D and fig. 11 B, D in Mu et al. [2022]), 2) the dagger-shaped conductor, with a blunt tip (vs triangular, cf. Fig. 35 C and fig. 11 C in Mu et al. [2022]), 3) the smaller, triangular copulatory openings (vs large, oval, cf. Fig. 35 E and fig. 11 E in Mu et al. [2022]), and 4) the curved, bow-shaped connecting tubes (vs auricular, cf. Fig. 35 F and fig. 11 F in Mu et al. [2022]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	description	Description. Male (Holotype): total length 4.06, carapace 1.95 long, 1.60 wide; abdomen 2.11 long, 1.27 wide. Eye sizes and interdistances: AME 0.12, ALE 0.12, PME 0.09, PLE 0.11; AME – AME 0.06, AME – ALE 0.01, ALE – ALE 0.29, PME – PME 0.14, PME – PLE 0.07, PLE – PLE 0.48, ALE – PLE 0.10. EAW 0.64, CRW 0.89, EAW / CRW 0.72, CRW / CW 0.56. MOA 0.32 long, anterior width 0.28, posterior width 0.34. CH 0.18, CH / AME 1.50. Labium 0.17 long, 0.24 wide. Sternum 1.07 long, 0.95 wide. Leg measurements: Ⅰ 7.52 (2.02, 0.68, 2.20, 1.80, 0.79), II 6.30 (1.71, 0.67, 1.61, 1.48, 0.83), III 5.50 (1.50, 0.62, 1.20, 1.43, 0.75), IV 8.28 (2.30, 0.74, 1.92, 2.31, 1.01). Spination: femur I d 2 pl 4, femur II d 1 pl 3, femora III – IV d 1, tibia Ⅰ pv 7 rv 8, tibia II pv 8 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with two black longitudinal stripes. Abdomen gray, with short narrow dorsal scutum, tip black, posteriorly with four black transverse stripes, venter with an H-shaped black pattern (Fig. 34 A, B). Legs yellow. Palp as in Figs 35 A – D, 36 A – C. Femoral apophysis well developed, nearly at middle of femur. Tibia short, about half length of femur. Retrolateral tibial apophysis long, with spine-shaped apophysis at base of prolateral margin and mamillary process at base of retrolateral margin. Bulb large, pyriform. Tegulum wider than cymbium, tegular apophysis long, tip coracoid. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus thick, strongly curved. Conductor dagger-shaped, tip blunt. Female (Paratype): total length 3.75, carapace 1.85 long, 1.66 wide; abdomen 1.90 long, 1.36 wide. Eye sizes and interdistances: AME 0.11, ALE 0.12, PME 0.09, PLE 0.11; AME – AME 0.06, AME – ALE 0.01, ALE – ALE 0.28, PME – PME 0.17, PME – PLE 0.07, PLE – PLE 0.47, ALE – PLE 0.10. EAW 0.63, CRW 0.96, EAW / CRW 0.66, CRW / CW 0.58. MOA 0.31 long, anterior width 0.25, posterior width 0.35. CH 0.14, CH / AME 1.27. Labium 0.18 long, 0.25 wide. Sternum 1.13 long, 0.96 wide. Leg measurements: Ⅰ 7.02 (1.79, 0.68, 2.21, 1.75, 0.59), II 6.07 (1.63, 0.66, 1.62, 1.41, 0.75), III 5.16 (1.38, 0.62, 1.07, 1.33, 0.76), IV 7.64 (2.06, 0.70, 1.80, 2.11, 0.97). Spination: femur I d 2 pl 4, femur II d 1 pl 3, femora III – IV d 1, tibia Ⅰ pv 7 rv 8, tibia II pv 8 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Legs yellow, with black annuli near tip of femora Ⅰ – IV. Other characters as in male, except carapace darker and dorsal scutum absent (Fig. 34 C, D). Epigyne as in Figs 35 E – F, 36 D – E. Epigynal plate sclerotized, slightly depressed in center, posterior margin W-shaped. Median septum wide. Copulatory openings small and triangular, located at anterior of depressed area, separated by median septum. Copulatory ducts short. Connecting tubes long and curved, bow-shaped. Glandular appendages short, mastoid-shaped. Bursa large, transparent, nearly rectangular. Spermathecae oval, separated by about half their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865FFFA65AA6FEA2DB3A249F.taxon	distribution	Distribution. China (Guangxi Zhuang Autonomous Region).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	description	Figs 37 – 39	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	materials_examined	Type material. Holotype ♁, CHINA: Hubei Province: Shennongjia Forestry District, Hongping Town (31 ° 41 ′ 41.62 ″ N, 110 ° 25 ′ 40.44 ″ E, 1753 m a. s. l.), 24 Oct. 2020, leg. Luyu Wang. Paratypes: 4 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	etymology	Etymology. The species name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	diagnosis	Diagnosis. This new species resembles O. triangula Mu, Jin & Zhang, 2022 in having a similarly shaped embolus and spermathecae, but can be recognized by: 1) the absence of a scar on the embolus (vs with scar, cf. Fig. 38 B, C and fig. 11 B, C in Mu et al. [2022]), 2) the longer retrolateral tibial apophysis, with a slightly curved tip (vs short, not curved, cf. Fig. 38 B and fig. 11 B in Mu et al. [2022]), 3) the rectangular conductor (vs triangular, cf. Fig. 38 C and fig. 11 C in Mu et al. [2022]), and 4) the anteriorly wide, posteriorly narrow median septum (vs anteriorly narrow, posteriorly wide, cf. Fig. 38 E and fig. 11 E in Mu et al. [2022]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	description	Description. Male (Holotype): total length 3.13, carapace 1.62 long, 1.38 wide; abdomen 1.51 long, 1.01 wide. Eye sizes and interdistances: AME 0.09, ALE 0.09, PME 0.09, PLE 0.09; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.23, PME – PME 0.11, PME – PLE 0.06, PLE – PLE 0.43, ALE – PLE 0.08. EAW 0.56, CRW 0.73, EAW / CRW 0.77, CRW / CW 0.53. MOA 0.28 long, anterior width 0.22, posterior width 0.30. CH 0.13, CH / AME 1.44. Labium 0.13 long, 0.24 wide. Sternum 0.88 long, 0.84 wide. Leg measurements: Ⅰ 6.39 (1.67, 0.59, 1.88, 1.50, 0.75), II 5.10 (1.36, 0.55, 1.31, 1.17, 0.71), III 4.24 (1.15, 0.48, 0.92, 1.11, 0.58), IV 6.75 (1.84, 0.60, 1.58, 1.84, 0.89). Spination: femur I d 0 pl 4, femur II d 1 pl 1, femur III d 0, femur IV d 1, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, with two black longitudinal stripes. Abdomen gray, with short narrow dorsal scutum, tip black, with five black transverse stripes posteriorly (Fig. 37 A, B). Legs yellow. Palp as in Figs 38 A – D, 39 A – C. Femoral apophysis well developed, near middle of femur. Tibia short, about half length of femur. Prolateral tibial apophysis distinct, round. Retrolateral tibial apophysis long, tip slightly curved, vertical in retrolateral view. Tegulum pyriform, retrolateral margin straight; tegular apophysis long, clavate-shaped. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus, occupying about half length of tegulum. Embolus thick, strongly curved. Conductor nearly rectangular, membranous. Female (Paratype): total length 4.01, carapace 1.64 long, 1.36 wide; abdomen 2.37 long, 1.58 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.09, PLE 0.10; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.23, PME – PME 0.11, PME – PLE 0.06, PLE – PLE 0.40, ALE – PLE 0. 08. EAW 0.54, CRW 0.71, EAW / CRW 0.76, CRW / CW 0.52. MOA 0.27 long, anterior width 0.23, posterior width 0.29. CH 0.11, CH / AME 1.10. Labium 0.15 long, 0.25 wide. Sternum 0.96 long, 0.88 wide. Leg measurements: Ⅰ 6.12 (1.65, 0.63, 1.78, 1.41, 0.65), II 5.01 (1.33, 0.56, 1.30, 1.13, 0.69), III 4.32 (1.18, 0.51, 0.85, 1.14, 0.64), IV 6.47 (1.67, 0.60, 1.51, 1.82, 0.87). Spination: femur I d 0 pl 4, femur II d 1 pl 2, femora III – IV d 1, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Legs yellow. Other characters as in male, except carapace darker and dorsal scutum absent (Fig. 37 C, D). Epigyne as in Figs 38 E – F, 39 D – E. Epigynal plate sclerotized, slightly depressed in center. Median septum wide anteriorly, narrow posteriorly. Copulatory openings oval, separated by more than 1.5 diameter of spermathecae. Copulatory ducts short. Connecting tubes long and curved, bow-shaped. Glandular appendages short, mastoid-shaped. Bursa large, transparent, kidney-shaped. Spermathecae oval, separated by about 1 / 4 their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE865CFFBA5AA6FB19DA362203.taxon	distribution	Distribution. China (Hubei Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	description	Figs 40 – 42	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	materials_examined	Type material. Holotype ♁, CHINA: Hunan Province: Xiangxi Tujia and Miao Autonomous Prefecture, Yongshun County, Bijiashan (28.983211 ° N, 109.813074 ° E, 331 m a. s. l.), 13 Nov. 2018, leg. Zhaoyi Li. Paratypes: 1 ♁ 6 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	etymology	Etymology. This specific name is derived from the Latin word “ striatus ”, referring to the longitudinal black stripe on carapace.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	diagnosis	Diagnosis. This new species resembles O. acutangula Liu, 2020 in having a similarly shaped embolus, but can be recognized by: 1) the smaller and thinner tegular apophysis (vs large and wide, cf. Fig. 41 C and fig. 7 D in Liu et al. [2020]), 2) the wider, dagger-shaped retrolateral tibial apophysis in retrolateral view, which is curved in dorsal view (vs thin and digitiform, not curved, cf. Fig. 41 B, D and fig. 7 E, F in Liu et al. [2020]), 3) the wider median septum (vs narrow, cf. Fig. 41 E and fig. 8 C in Liu et al. [2020]), 4) the slanted, slightly curved connecting tubes (vs straight, nearly transverse, cf. Fig. 41 F and fig. 8 D in Liu et al. [2020]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	description	Description. Male (Holotype): total length 2.85, carapace 1.37 long, 1.19 wide; abdomen 1.48 long, 0.96 wide. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.06, PLE 0.08; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.21, PME – PME 0.09, PME – PLE 0.05, PLE – PLE 0.35, ALE – PLE 0.08. EAW 0.47, CRW 0.63, EAW / CRW 0.75, CRW / CW 0.53. MOA 0.25 long, anterior width 0.20, posterior width 0.26. CH 0.12, CH / AME 1.50. Labium 0.14 long, 0.21 wide. Sternum 0.82 long, 0.70 wide. Leg measurements: Ⅰ 5.55 (1.40 + 0.52 + 1.65 + 1.34 + 0.64), II 4.26 (1.15, 0.47, 1.15, 0.96, 0.56), III 3.76 (1.00, 0.43, 0.79, 1.00, 0.54), IV 5.73 (1.53, 0.45, 1.34, 1.62, 0.79). Spination: femora I – IV d 1, femur I pl 4, femur II pl 5, tibia Ⅰ pv 7 rv 8, tibia II pv 7 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace gray, with one black longitudinal stripe nearly same width as eye area. Abdomen gray, with a narrow and wedge shape dorsal scutum, large black pattern beside dorsal scutum, four black transverse stripes at posterior of abdomen (Fig. 40 A, B). Legs gray-white. Palp as in Figs 41 A – D, 42 A – C. Femoral apophysis well developed, nearly at middle of femur. Tibia about half length of femur. Prolateral tibial apophysis distinct, round. Retrolateral tibial apophysis base thick, extending towards retrolateral side, then folding upwards, dagger-shaped in retrolateral view, curved in dorsal view. Tegulum oval, wider than cymbium; tegular apophysis oxhorn-shaped in retrolateral view, located at tip of tegulum retrolaterally. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus wide, curved, almost L-shaped. Conductor absent. Female (Paratype): total length 3.24, carapace 1.42 long, 1.26 wide; abdomen 1.82 long, 1.36 wide. Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.07, PLE 0.09; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.23, PME – PME 0.08, PME – PLE 0.05, PLE – PLE 0.34, ALE – PLE 0. 08. EAW 0.48, CRW 0.69, EAW / CRW 0.70, CRW / CW 0.55. MOA 0.25 long, anterior width 0.21, posterior width 0.25. CH 0.10, CH / AME 1.11. Labium 0.15 long, 0.22 wide. Sternum 0.89 long, 0.76 wide. Leg measurements: Ⅰ 5.58 (1.39, 0.58, 1.68, 1.34, 0.59), II 4.55 (1.22, 0.52, 1.18, 1.04, 0.59), III 3.84 (1.00, 0.42, 0.86, 0.97, 0.59), IV 5.82 (1.50, 0.53, 1.38, 1.62, 0.79). Spination: femur I d 2 pl 4, femora II – IV d 1, femur II pl 2, tibia Ⅰ pv 8 rv 8, tibia II pv 8 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Legs yellow, with black annuli near tip of femora, patella, tibia I – IV. Other characters as in male, except carapace darker, abdomen lighter and dorsal scutum absent (Fig. 40 C, D). Epigyne as in Figs 41 E – F, 42 D – E. Epigynal plate sclerotized. Atrium separated into two parts, filled with mating plug. Median septum wide. Copulatory openings located at posterior margin of atrium. Copulatory ducts short, thick. Connecting tubes long, slightly curved. Glandular appendages small, mastoid-shaped. Bursa small, oval, transparent. Spermathecae oval, separated by their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8640FFBD5AA6FD65DACC279F.taxon	distribution	Distribution. China (Hunan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	description	Figs 43 – 45	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	materials_examined	Type material. Holotype ♁, CHINA: Yunnan Province: Baoshan City, Longyang District, Lujiang Town (24.928230 ° N, 98.836184 ° E, 1001 m a. s. l.), 27 Nov. 2017, leg. Yannan Mu. Paratypes: 5 ♁ 3 ♀, with same data as holotype. 5 ♁ 10 ♀, same locality, Baihua Ridge Old Street (25.298096 ° N, 98.786316 ° E, 1983 m a. s. l.), 24 Nov. 2017, leg. Yannan Mu; 1 ♁ 1 ♀, same locality, Dayang Zhai (24.928124 ° N, 98.797474 ° E, 1927 m a. s. l.), 23 Nov. 2017, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	etymology	Etymology. This specific is named for its similarity to O. forcipata Yang, Wang & Yang, 2013.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	diagnosis	Diagnosis. This new species resembles O. forcipata in having a similarly shaped embolus and connecting tubes, but can be recognized by: 1) the absence of a conductor (vs with conductor, cf. Fig. 44 B, C and fig. 9 in Yang et al. [2013]), 2) the presence of a mastoid-shaped apophysis on the retroventral side of the tibia (vs without, cf. Fig. 44 C and fig. 10 in Yang et al. [2013]), 3) the blunt tip of the retrolateral tibial apophysis, with a slender branch at the base prolaterally (vs tip bifurcate, without a branch, cf. Fig. 44 B and fig. 10 in Yang et al. [2013]), and 4) the wider, wavy-shaped posterior edge of the epigynal plate (vs narrow, M-shaped, cf. Fig. 44 E and fig. 13 in Yang et al. [2013]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	description	Description. Male (Holotype): total length 3.46, carapace 1.74 long, 1.52 wide; abdomen 1.72 long, 1.14 wide. Eye sizes and interdistances: AME 0.13, ALE 0.11, PME 0.11, PLE 0.10; AME – AME 0.06, AME – ALE 0.01, ALE – ALE 0.31, PME – PME 0.14, PME – PLE 0.08, PLE – PLE 0.52, ALE – PLE 0.09. EAW 0.65, CRW 0.83, EAW / CRW 0.78, CRW / CW 0.55. MOA 0.28 long, anterior width 0.28, posterior width 0.36. CH 0.17, CH / AME 1.31. Labium 0.17 long, 0.26 wide. Sternum 0.97 long, 0.93 wide. Leg measurements: Ⅰ 7.18 (1.82, 0.63, 2.10, 1.73, 0.90), II 6.12 (1.63, 0.60, 1.64, 1.35, 0.90), III 5.32 (1.41, 0.57, 1.14, 1.35, 0.85), IV 7.95 (2.09, 0.62, 1.88, 2.20, 1.16). Spination: femur I d 2 pl 4, femora II – IV d 1, femur II pl 3, tibia Ⅰ pv 8 rv 8, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace black, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with a narrow and wedge shape dorsal scutum, large black pattern beside dorsal scutum, four black transverse stripes at posterior of abdomen (Fig. 43 A, B). Legs yellow, with black annuli near tip of femora I – IV. Palp as in Figs 44 A – D, 45 A – C. Femoral apophysis well developed, nearly at middle of femur. Tibia about half length of femur. Prolateral tibial apophysis distinct, nearly round. Retrolateral tibial apophysis base wide, tapering from base to end, tip blunt, with slender branch at base prolaterally, curved in dorsal view. Retro-ventral tibia with mastoid-shaped apophysis. Tegulum pyriform, about 1 / 3 anterior area membranous; tegular apophysis absent. Sperm duct distinct, “ U ” - shaped, tapering from retrolateral side of tegulum to embolus. Embolus flat and wide, tip blunt, curved, C-shaped. Conductor membranous. Female (Paratype): total length 3.14, carapace 1.56 long, 1.41 wide; abdomen 1.58 long, 1.09 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.08, PLE 0.09; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.29, PME – PME 0.11, PME – PLE 0.07, PLE – PLE 0.43, ALE – PLE 0. 10. EAW 0.57, CRW 0.77, EAW / CRW 0.74, CRW / CW 0.55. MOA 0.27 long, anterior width 0.25, posterior width 0.30. CH 0.12, CH / AME 1.20. Labium 0.14 long, 0.26 wide. Sternum 0.91 long, 0.94 wide. Leg measurements: Ⅰ 6.73 (1.64, 0.64, 2.02, 1.62, 0.81), II 4.64 (1.23, 0.42, 1.18, 1.13, 0.68), III 4.72 (1.27, 0.54, 1.00, 1.23, 0.68), IV 7.22 (1.87, 0.62, 1.73, 2.04, 0.96). Spination: femora I – IV d 1, femur I pl 5, femur II pl 3, tibia Ⅰ pv 9 rv 9, tibia II pv 6 rv 5, metatarsus Ⅰ pv 5 rv 4, metatarsus II pv 4 rv 3. Legs yellow. Other characters as in male, except carapace darker and dorsal scutum absent (Fig. 43 C, D). Epigyne as in Figs 44 E – F, 45 D – E. Epigynal plate sclerotized. Atrium heart-shaped. Median septum distinct, wide anteriorly, narrow posteriorly. Copulatory openings nearly triangular, located at anterior of atrium, separated by more than 3 times spermathecae diameter. Copulatory ducts short. Connecting tubes long, auriform. Bursa balloon-shaped, transparent. Spermathecae small, round, separated by about one diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8645FFB35AA6F9F2DAC12227.taxon	distribution	Distribution. China (Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	description	Figs 46 – 48	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province: Chengdu City, Chongzhou County, Wenjingjiang Town, Anzihe Conservation Area, Shaoyaogou Protection Station (30 ° 47.577 ′ N, 103 ° 12.683 ′ E, 1641 m a. s. l.), 2 Oct. 2016, leg. Yannan Mu. Paratypes: 2 ♁ 2 ♀, with same data as holotype; 2 ♁ 6 ♀, same locality, Maliugou (30 ° 46.790 ′ N, 103 ° 13.530 ′ E, 1692 m a. s. l.), 3 Oct. 2016, leg. Luyu Wang.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	etymology	Etymology. The species name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	diagnosis	Diagnosis. This new species resembles O. longituba Wang, Zhang & Zhang, 2012 in having a similar-shaped embolus, but can be recognized by: 1) the palmate-shaped retrolateral tibial apophysis (vs rectangular, cf. Fig. 47 B and fig. 5 F in Wang et al. [2012]), 2) the closely situated copulatory openings (vs separated by about one spermatheca diameter, cf. Fig. 47 E and fig. 5 H in Wang et al. [2012]), and 3) distinctly separated spermathecae (vs close to each other, cf. Fig. 47 F and fig. 5 G in Wang et al. [2012]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	description	Description. Male (Holotype): total length 3.28, carapace 1.61 long, 1.42 wide; abdomen 1.67 long, 1.04 wide. Eye sizes and interdistances: AME 0.11, ALE 0.10, PME 0.08, PLE 0.10; AME – AME 0.02, AME – ALE 0.01, ALE – ALE 0.25, PME – PME 0.13, PME – PLE 0.07, PLE – PLE 0.45, ALE – PLE 0.08. EAW 0.59, CRW 0.84, EAW / CRW 0.70, CRW / CW 0.59. MOA 0.29 long, anterior width 0.23, posterior width 0.30. CH 0.14, CH / AME 1.27. Labium 0.16 long, 0.24 wide. Sternum 0.93 long, 0.82 wide. Leg measurements: Ⅰ 6.26 (1.63, 0.62, 1.83, 1.39, 0.79), II 5.17 (1.39, 0.56, 1.35, 1.11, 0.76), III 4.70 (1.24, 0.52, 0.98, 1.24, 0.72), IV 7.02 (1.88, 0.59, 1.62, 1.94, 0.99). Spination: femora I – IV d 1, femur I pl 4, femur II pl 3, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, radial striae distinct, beside fovea. Abdomen gray, with narrow dorsal scutum anteriorly, large black pattern beside dorsal scutum, four black transverse stripes at posterior of abdomen (Fig. 46 A, B). Legs yellow. Palp as in Figs 47 A – D, 48 A – C. Femoral apophysis well developed, nearly at middle parts of femur. Tibia about half length of femur. Prolateral tibial apophysis distinct, triangular. Retrolateral tibial apophysis flat, palmate-shaped, tip blunt, curved towards cymbium. Tegulum nearly diamond-shaped; tegular apophysis small, semicircular. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus needle-like, transverse. Conductor absent. Female (Paratype): total length 3.79, carapace 1.68 long, 1.44 wide; abdomen 2.11 long, 1.48 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.08, PLE 0.09; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.24, PME – PME 0.12, PME – PLE 0.07, PLE – PLE 0.42, ALE – PLE 0. 12. EAW 0.56, CRW 0.79, EAW / CRW 0.71, CRW / CW 0.55. MOA 0.28 long, anterior width 0.21, posterior width 0.30. CH 0.15, CH / AME 1.50. Labium 0.16 long, 0.25 wide. Sternum 0.97 long, 0.92 wide. Leg measurements: Ⅰ 6.35 (1.58, 0.66, 1.84, 1.53, 0.74), II 5.32 (1.40, 0.57, 1.35, 1.27, 0.73), III 4.62 (1.22, 0.52, 0.97, 1.21, 0.70), IV 6.78 (1.78, 0.64, 1.53, 1.86, 0.97). Spination: femora I – IV d 1, femur I pl 4, femur II pl 3, tibia Ⅰ pv 7 rv 8, tibia II pv 7 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Legs yellow. Other characters as in male, except carapace darker and dorsal scutum absent (Fig. 46 C, D). Epigyne as in Figs 47 E – F, 48 D – E. Epigynal plate sclerotized, with small depression centrally. Median septum absent. Copulatory openings oval, close to each other, located at centre of atrium. Copulatory ducts long, inverted V-shaped. Connecting tubes short, auriform. Bursa balloon-shaped, membranous. Spermathecae small, round, separated by about half their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8649FFB25AA6FD81DADD27B1.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	description	Figs 49 – 51	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province: Mianyang City, Pingwu County, Baima Tibetan Town (32 ° 42 ′ 47.92 ″ N, 104 ° 22 ′ 35.71 ″ E, 1778 m a. s. l.), 15 Oct. 2020, leg. Luyu Wang. Paratypes: 2 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	etymology	Etymology. This specific is named for its similarity to O. dentigera Mu & Zhang, 2021.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	diagnosis	Diagnosis. This new species resembles O. dentigera in having a similarly shaped embolus, but it can be recognized by: 1) the semicircular prolateral tibial apophysis (vs triangular, cf. Fig. 50 A and fig. 5 A in Mu & Zhang [2021]), 2) the retrolateral tibial apophysis narrowing abruptly near the middle, with a digitiform tip (vs tapering from base to tip, cf. Fig. 50 B and fig. 5 F in Mu & Zhang [2021]), 3) the oval spermathecae (vs clavate, cf. Fig. 50 F and fig. 5 I in Mu & Zhang [2021]), and 4) the absence of glandular appendages (vs clavate, cf. Fig. 50 F and fig. 5 I in Mu & Zhang [2021]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	description	Description. Male (Holotype): total length 2.69, carapace 1.28 long, 1.12 wide; abdomen 1.41 long, 0.94 wide. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.08, PLE 0.09; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.18, PME – PME 0.11, PME – PLE 0.04, PLE – PLE 0.34, ALE – PLE 0.07. EAW 0.45, CRW 0.61, EAW / CRW 0.74, CRW / CW 0.54. MOA 0.22 long, anterior width 0.18, posterior width 0.26. CH 0.12, CH / AME 1.71. Labium 0.14 long, 0.20 wide. Sternum 0.72 long, 0.67 wide. Leg measurements: Ⅰ 4.42 (1.21, 0.45, 1.22, 1.06, 0.48), II 3.81 (1.04, 0.41, 0.93, 0.88, 0.55), III 3.31 (0.91, 0.37, 0.67, 0.86, 0.50), IV 4.71 (1.25, 0.44, 1.07, 1.30, 0.65). Spination: femora I d 1 pl 4, femur II d 1 pl 1, femur IV d 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 4, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow, radial striae indistinct, with black stripes surrounding submargin of carapace. Abdomen gray, with dorsal scutum, with black line beside dorsal scutum anteriorly and four black transverse stripes at posterior of abdomen (Fig. 49 A, B). Legs yellow, with black annuli near tip of femora, patellae and tibiae I – IV. Palp as in Figs 50 A – D, 51 A – C. Femoral apophysis well developed, nearly at tip of femur. Prolateral tibial apophysis distinct, semicircular. Retrolateral tibial apophysis base wide, narrowing abruptly near middle, tip digitiform, with short branch prolaterally at base. Tegulum wider than cymbium, pyriform; tegular apophysis small. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus needle-like, curved, transverse, extending to retrolateral margin of cymbium. Conductor absent. Female (Paratype): total length 2.95, carapace 1.31 long, 1.15 wide; abdomen 1.64 long, 1.16 wide. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.07, PLE 0.09; AME – AME 0.02, AME – ALE 0.01, ALE – ALE 0.19, PME – PME 0.10, PME – PLE 0.04, PLE – PLE 0.34, ALE – PLE 0. 07. EAW 0.46, CRW 0.64, EAW / CRW 0.72, CRW / CW 0.56. MOA 0.22 long, anterior width 0.17, posterior width 0.25. CH 0.10, CH / AME 1.43. Labium 0.14 long, 0.21 wide. Sternum 0.81 long, 0.76 wide. Leg measurements: Ⅰ 4.45 (1.15, 0.47, 1.26, 1.05, 0.52), II 3.69 (1.00, 0.42, 0.92, 0.86, 0.49), III 3.27 (0.90, 0.38, 0.65, 0.81, 0.53), IV 4.71 (1.29, 0.41, 1.09, 1.25, 0.67). Spination: femur I d 0 pl 3, femur II d 1 pl 1, femora III – IV d 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as in male, except dorsal scutum absent (Fig. 49 C, D). Epigyne as in Figs 50 E – F, 51 D – E. Epigynal plate sclerotized. Median septum absent. Copulatory openings located near centre of plate, close to each other. Copulatory ducts short, curved upwards and connecting with bursa. Connecting tubes long, thinner than copulatory ducts, curved, circular. Bursa oval, transparent. Spermathecae oval, close to each other. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE864EFFC85AA6F994DADD2193.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	description	Figs 52 – 54	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	materials_examined	Type material. Holotype ♁, CHINA: Yunnan Province: Baoshan City, Tengchong County, Laifengshan National Forest Park, Wenbi Tower (25.014684 ° N, 98.479428 ° E, 1914 m a. s. l.), 12 Dec. 2017, leg. Yannan Mu. Paratypes: 10 ♁ 18 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	etymology	Etymology. The species name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	diagnosis	Diagnosis. This new species resembles O. hamata in having a similarly slanted embolus, but can be recognized by: 1) the larger, arch-shaped tegular apophysis (vs small and triangular, cf. Fig. 53 C and fig. 9 G in Wang et al. [2012]), 2) the absence of a conductor (vs with conductor, cf. Fig. 53 C and fig. 9 G in Wang et al. [2012]), 3) the blunt-tipped retrolateral and shorter dorsal tibial apophyses (vs sharp-tipped retrolateral tibial apophysis tip and long dorsal tibial apophysis, cf. Fig. 53 D and fig. 9 F in Wang et al. [2012]), 4) the smaller, oval copulatory openings (vs large and arc-shaped, cf. Fig. 53 E and fig. 9 H in Wang et al. [2012]), and 5) the smaller, closely situated spermathecae (vs separated by about one spermatheca diameter, cf. Fig. 53 F and fig. 9 I in Wang et al. [2012]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	description	Description. Male (Holotype): total length 3.50, carapace 1.65 long, 1.44 wide; abdomen 1.85 long, 1.46 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.08, PLE 0.09; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.27, PME – PME 0.13, PME – PLE 0.07, PLE – PLE 0.47, ALE – PLE 0.07. EAW 0.59, CRW 0.73, EAW / CRW 0.81, CRW / CW 0.51. MOA 0.27 long, anterior width 0.24, posterior width 0.32. CH 0.17, CH / AME 1.70. Labium 0.15 long, 0.25 wide. Sternum 0.88 long, 0.88 wide. Leg measurements: Ⅰ 5.90 (1.49, 0.59, 1.72, 1.39, 0.71), II 4.94 (1.32, 0.55, 1.29, 1.11, 0.67 +), III 4.25 (1.13, 0.47, 0.91, 1.11, 0.63), IV 5.84 (1.66, 0.49, 1.32, 1.51, 0.86). Spination: femur I d 1 pl 5, femur II d 1 pl 2, femur IV d 1, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsi Ⅰ – II pv 4 rv 3. Coloration. Carapace brown, with several markings resembling flowing water droplets beside fovea. Abdomen gray, with long narrow dorsal scutum, tip black, posterior with two black transverse stripes (Fig. 52 A, B). Legs yellow, with black annuli near tips of femora Ⅰ – IV and tibiae Ⅰ – III. Palp as in Figs 53 A – D, 54 A – C. Femoral apophysis protruding, well-developed, near middle of femur. Tibia short, about 1 / 3 of femur. Retrolateral tibial apophysis short but strong, tip blunt; dorsal tibial apophysis base strong, tip flake-shaped, wider than base in dorsal view, curved in retrolateral view. Bulb large, tegulum wider than cymbium, pyriform; tegular apophysis large, arch-shaped, located at retrolateral tip of tegulum. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus tapering from base to tip, transverse, slightly curved and extending towards retrolateral side of cymbium. Conductor absent. Female (Paratype): total length 4.09, carapace 1.76 long, 1.56 wide; abdomen 2.33 long, 1.56 wide. Eye sizes and interdistances: AME 0.12, ALE 0.12, PME 0.10, PLE 0.11; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.28, PME – PME 0.12, PME – PLE 0.07, PLE – PLE 0.47, ALE – PLE 0.09. EAW 0.63, CRW 0.79, EAW / CRW 0.80, CRW / CW 0.51. MOA 0.30 long, anterior width 0.27, posterior width 0.33. CH 0.15, CH / AME 1.25. Labium 0.15 long, 0.28 wide. Sternum 1.00 long, 0.95 wide. Leg measurements: Ⅰ 6.44 (1.69, 0.68, 1.85, 1.52, 0.70), II 5.28 (1.42, 0.61, 1.40, 1.21, 0.64), III 4.54 (1.19, 0.55, 0.97, 1.14, 0.69), IV 6.76 (1.78, 0.60, 1.54, 1.89, 0.95). Spination: femora I – IV d 1, femur I pl 4, femur II pl 3, tibia Ⅰ pv 8 rv 8, tibia II pv 7 rv 7, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as in male, except color darker and dorsal scutum absent (Fig. 52 C, D). Epigyne as in Figs 53 E – F, 54 D – E. Epigynal plate weakly sclerotized. Median septum trapezoidal. Copulatory openings small and round, separated by about half diameter of spermatheca. Copulatory ducts short, not visible. Connecting tube long and curved. Glandular appendages absent. Bursa large, sclerotized, irregularly-shaped. Spermathecae round, close to each other. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8632FFCD5AA6FE15DAC126F2.taxon	distribution	Distribution. China (Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	description	Figs 55 – 57	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province: Guangyuan City, Qingchuan County, Qingxi Town, Yongjiawan Village (32 ° 25 ′ 59.52 ″ N, 104 ° 45 ′ 35.20 ″ E, 1362 m a. s. l.), 16 Oct. 2020, leg. Zhisheng Zhang. Paratypes: 1 ♁ 3 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	etymology	Etymology. The species name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	diagnosis	Diagnosis. This new species resembles O. horizon sp. nov. in also having a transverse embolus, but can be recognized by: 1) the smaller retrolateral tibial apophysis (vs large, cf. Fig. 56 D and Fig. 34 D), 2) the large dorsal tibial apophysis (vs without dorsal tibial apophysis, cf. Fig. 56 D and Fig. 34 D), 3) the copulatory openings extending posteriorly (vs extending anteriorly, cf. Fig. 56 E and Fig. 34 E), 4) the thinner copulatory ducts, extending anteriorly (vs extending posteriorly, cf. Fig. 56 E and Fig. 34 E), and 5) the distinctly separated connecting tubes (vs close to each other, cf. Fig. 56 F and Fig. 34 F).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	description	Description. Male (Holotype): total length 2.89, carapace 1.45 long, 1.27 wide; abdomen 1.44 long, 0.93 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.08, PLE 0.09; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.22, PME – PME 0.12, PME – PLE 0.06, PLE – PLE 0.39, ALE – PLE 0.08. EAW 0.52, CRW 0.65, EAW / CRW 0.80, CRW / CW 0.51. MOA 0.26 long, anterior width 0.21, posterior width 0.28. CH 0.13, CH / AME 1.30. Labium 0.17 long, 0.20 wide. Sternum 0.83 long, 0.75 wide. Leg measurements: Ⅰ 5.77 (1.46, 0.52, 1.70, 1.38, 0.71), II 4.70 (1.23, 0.48, 1.22, 1.11, 0.66), III 4.02 (1.07, 0.44, 0.82, 1.08, 0.61), IV 6.09 (1.69, 0.47, 1.44, 1.66, 0.83). Spination: femora I – IV d 1, femur I pl 4, femur II pl 2, tibia Ⅰ pv 7 rv 8, tibia II pv 6 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace yellow-brown, with several shapes resembling flowing water droplets beside fovea. Abdomen gray, with narrow dorsal scutum anteriorly, flanks black, posterior with four black transverse stripes (Fig. 55 A, B). Legs yellow. Palp as in Figs 56 A – D, 57 A – C. Femoral apophysis well developed, located at middle of femur. Tibia short, about half length of femur. Retrolateral tibial apophysis short but strong, tip blunt; dorsal tibial apophysis long, extending to middle parts of retrolateral margin of cymbium. Bulb large, tegulum wider than cymbium, pyriform; tegular apophysis absent. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus tapering from base to tip, transverse, extending towards retrolateral of cymbium. Conductor absent. Female (Paratype): total length 3.36, carapace 1.50 long, 1.30 wide; abdomen 1.86 long, 1.36 wide. Eye sizes and interdistances: AME 0.10, ALE 0.09, PME 0.08, PLE 0.09; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.24, PME – PME 0.09, PME – PLE 0.06, PLE – PLE 0.38, ALE – PLE 0.09. EAW 0.51, CRW 0.71, EAW / CRW 0.72, CRW / CW 0.55. MOA 0.26 long, anterior width 0.23, posterior width 0.27. CH 0.16, CH / AME 1.60. Labium 0.13 long, 0.22 wide. Sternum 0.91 long, 0.84 wide. Leg measurements: Ⅰ 5.84 (1.48, 0.57, 1.75, 1.38, 0.66), II 4.66 (1.22, 0.51, 1.19, 1.13, 0.61), III 3.99 (1.04, 0.49, 0.86, 1.02, 0.58), IV 6.25 (1.67, 0.56, 1.45, 1.76, 0.81). Spination: femur I d 2 pl 4, femora II – IV d 1, femur II pl 2, tibia Ⅰ pv 8 rv 8, tibia II pv 8 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as in male, except body larger, color darker and dorsal scutum absent (Fig. 55 C, D). Epigyne as in Figs 56 E – F, 57 D – E. Epigynal plate sclerotized, with shallow depression centrally. Median septum wide. Copulatory openings small and round, towards posteriorly, separated by median septum. Copulatory ducts short, extending anteriorly and connecting with bursa. Connecting tubes thin and straight. Glandular appendages absent. Bursa oval. Spermathecae round, separated by about half their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8637FFC15AA6F8F6DADD242F.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863BFFC15AA6FBBEDA1C26E9.taxon	discussion	Comments: With the discovery of Otacilia species, our study found that the difference between O. dawushan and O. wugongshanica exists in the abdominal color pattern, the angle of the retrolateral tibial apophysis curvature, and the size of the sclerotized fovea on the epigynal plate. The differences between O. zhonglong and O. dawushan are only based on the abdominal color pattern and curvature of the retrolateral tibial apophysis. The differences between O. xingguo and O. zhonglong are recorded as “ … by the ventral abdomen lacking an H-shaped dark brown mark (Fig. 116 C) (vs present) and the connecting tubes are transverse posteriorly (Fig. 116 D) (vs obliquely) ” by Liu & Li (2022). Our study found that these four species share the same characteristics in the male palp and female epigyne, e. g., the shape of the embolus, tegular apophysis, retrolateral tibial apophysis, and femoral apophysis in the male palp, and the atrium, median septum, copulatory openings, copulatory ducts, connecting tubes, and spermathecae in the female epigyne. As a result, we suggest that O. dawushan, O. xingguo, and O. zhonglong should be considered junior synonyms of O. wugongshanica.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863BFFC05AA6F8F1DAA121CB.taxon	type_taxon	(type species: Phrurolithus pennatus Yaginuma, 1967, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863BFFC05AA6F8F1DAA121CB.taxon	diagnosis	Diagnosis. This genus can be distinguished from all other phrurolithid genera by the femur I with some prolateral spines (at least two), the femur II without prolateral spine; the male palp with retroventral tibial apophysis; the female with a pair of blind ducts protruding anteriorly but without bursa.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863BFFC05AA6F8F1DAA121CB.taxon	description	Description. Small spiders. Carapace oval, highest near fovea, widest at coxae II and III. Chelicerae with three well separated promarginal and two close retromarginal teeth, with two spines anteriorly (one long and another short). Femur I usually with two dorsal spines and more than two prolateral spines distally, femur II usually with two prolateral spines but without dorsal spine; tibia I usually with more than six pairs of ventral spines, tibia II usually with six pairs of ventral spines (usually unpaired); metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines. Abdomen oval, black, with large dorsal scutum in dorsal view in males but without in female. Male palp: femur only with somewhat swollen; tibia with one large retrolateral tibial apophysis and one small retroventral tibial apophysis; tegulum oval, sperm duct long, U-shaped; embolus relatively short, situated at tip of tegulum. Female epigyne: internal genitalia with a pair of blind ducts protruding anteriorly but without membranous bursa.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	description	Figs 58 – 60	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	materials_examined	Type material. Holotype ♁, CHINA: Liaoning Province: Benxi City, Huanren Manchu Autonomous County, Wu’nv Mountain (41 ° 18.597 ′ N, 125 ° 23.952 ′ E, 360 m a. s. l.), 6 Jul. 2016, leg. Weitong Wang.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	etymology	Etymology. This specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	diagnosis	Diagnosis. This new species resembles P. splendidus (Song & Zheng, 1992) in having a similarly shaped retroventral tibial apophysis, but can be recognized by: 1) the thinner, needle-shaped embolus (vs thick, nubbyshaped, cf. Fig. 59 C and fig. 3 H in Kamura [2021]), and 2) the shorter, rod-like tegular apophysis (vs arc-shaped, cf. Fig. 59 C and fig. 3 K in Kamura [2021]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	description	Description. Male (Holotype): total length 2.58, carapace 1.23 long, 1.00 wide; abdomen 1.35 long, 0.74 wide. Eye sizes and interdistances: AME 0.08, ALE 0.08, PME 0.07, PLE 0.07; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.20, PME – PME 0.04, PME – PLE 0.03, PLE – PLE 0.26, ALE – PLE 0.06. EAW 0.37, CRW 0.49, EAW / CRW 0.76, CRW / CW 0.49. MOA 0.22 long, anterior width 0.20, posterior width 0.20. CH 0.12, CH / AME 1.50. Labium 0.13 long, 0.18 wide. Sternum 0.71 long, 0.62 wide. Leg measurements: Ⅰ 4.96 (1.27, 0.44, 1.31, 1.17, 0.77), II 3.84 (1.03, 0.38, 0.89, 0.92, 0.65), III 3.35 (0.90, 0.34, 0.72, 0.86, 0.53), IV 5.57 (1.58, 0.42, 1.32, 1.47, 0.78). Spination: femur I d 2 pl 2, femora II – IV d 1, tibia Ⅰ pv 7 rv 7, tibia II pv 6 rv 5, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 2. Coloration. Carapace yellow, oval, radial striae distinct, with sparse iridescent scales in dorsal view. Abdomen darker in color than carapace, with large dorsal scutum covering nearly entire abdomen and sparse iridescent scales dorsally (Fig. 58). Legs yellow. Palp as in Figs 59 – 60. Femoral apophysis protruding slightly, with several setae. Prolateral tibial apophysis distinct, arch-shaped. Retrolateral tibial apophysis base wide, narrowing abruptly at middle and tip digitiform; retroventral tibial apophysis nearly trapeziform. Tegulum narrower than cymbium, oval; tegular apophysis short rod-like. Sperm duct distinct, stomach-shaped, tapering from retrolateral side of tegulum to embolus. Embolus short, transverse. Conductor absent. Female. Unknown.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863AFFC25AA6FDADDAD621B7.taxon	distribution	Distribution. China (Liaoning Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	description	Figs 61 – 63	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	materials_examined	Type material. Holotype ♁, CHINA: Hunan Province: Yongzhou City, Ningyuan County, Jiuyi Mountain (25 ° 16 ′ 43.12 ″ N, 111 ° 56 ′ 45.6 ″ E, 734 m a. s. l.), 27 Apr. 2021, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	etymology	Etymology. This specific name is derived from the Latin word “ mico ”, referring to the twinkling iridescent scales; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	diagnosis	Diagnosis. This new species resembles P. splendidus (Song & Zheng, 1992) in having a similarly shaped embolus, but can be distinguished by: 1) the M-shaped prolateral margin of the retrolateral tibial apophysis, and the shorter, digitiform tip (vs arch-shaped and tip long, cf. Fig. 62 C and fig. 3 I in Kamura [2021]), 2) the horseshoe-shaped retroventral tibial apophysis (vs trapezoid-shaped, cf. Fig. 62 C and fig. 3 I in Kamura [2021]), and 3) the round tegular apophysis (vs nearly rectangular, cf. Fig. 62 B and fig. 3 K in Kamura [2021]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	description	Description. Male (Holotype): total length 2.76, carapace 1.30 long, 1.13 wide; abdomen 1.46 long, 0.79 wide. Eye sizes and interdistances: AME 0.09, ALE 0.09, PME 0.07, PLE 0.08; AME – AME 0.05, AME – ALE 0.01, ALE – ALE 0.22, PME – PME 0.05, PME – PLE 0.05, PLE – PLE 0.30, ALE – PLE 0.06. EAW 0.41, CRW 0.57, EAW / CRW 0.72, CRW / CW 0.50. MOA 0.24 long, anterior width 0.22, posterior width 0.21. CH 0.12, CH / AME 1.33. Labium 0.13 long, 0.20 wide. Sternum 0.80 long, 0.67 wide. Leg measurements: Ⅰ 5.84 (1.50, 0.49, 1.57, 1.34, 0.94), II 4.67 (1.25, 0.44, 1.17, 1.03, 0.78), III 4.09 (1.12, 0.43, 0.88, 1.02, 0.64), IV 6.36 (1.71, 0.53, 1.52, 1.72, 0.88). Spination: femur I d 2 pl 3, femur II d 2, femora III – IV d 1, tibia Ⅰ pv 7 rv 6, tibia II pv 6 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace black, oval, radial striae distinct, with sparse iridescent scales in dorsal view. Abdomen darker in color than carapace, with lighter color at anterior of dorsal scutum, gray transverse stripe medially, and dense iridescent scales posteriorly in dorsal view (Fig. 61). Legs gray. Palp as in Figs 62 – 63. Femoral apophysis protruding slightly near tip, with several setae. Prolateral tibial apophysis distinct, semicircular. Retrolateral tibial apophysis wide basally, prolateral margin curved, M-shaped, digitiform tip short, curved posteriorly; retroventral tibial apophysis horseshoe-shaped. Tegulum thinner than cymbium, oval; tegular apophysis nearly round. Sperm duct distinct, stomach-shaped, tapering from retrolateral side of tegulum to embolus. Embolus short, transverse. Conductor membranous. Female. Unknown.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8638FFC45AA6FE31DACC2123.taxon	distribution	Distribution. China (Hunan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	description	Figs 64 – 66	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	materials_examined	Material examined. 1 ♀, CHINA: Liaoning Province: Dalian City, Zhuanghe County, Xianren Cave Conservation Area (40 ° 00.655 ′ N, 122 ° 58.262 ′ E, 143 m a. s. l.), 4 Jul. 2016, leg. Weitong Wang; 1 ♀, Jilin Province: Tonghua City, Ji’an County, Dalu Town (40 ° 59.591 ′ N, 125 ° 46.597 ′ E, 239 m a. s. l.), 9 Jul. 2016, leg. Weitong Wang.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	diagnosis	Diagnosis. See Kamura, 2021.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	description	Description. See Kim & Lee, 2014. Habitus and genitalia as in Figs 64 – 65.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	distribution	Distribution. China (Jilin Province, Liaoning Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863EFFC45AA6FA8DDBAB27BF.taxon	discussion	Remark. This species is recorded from Jilin province for the first time.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863CFFC65AA6FCA7DC2A2768.taxon	etymology	Etymology. The generic name “ punctus ” is a Latin word meaning short spine, referring to the short spines in the middle of the ventral surface of metatarsus Ⅰ. The gender is masculine.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863CFFC65AA6FCA7DC2A2768.taxon	diagnosis	Diagnosis. This new genus can be distinguished from all other phrurolithid genera by absence of teeth on promargin and retromargin of chelicerae and presence of short, stout spines in the ventral, middle surface of metatarsus Ⅰ. The new genus is similar to Phrurolithus C. L. Koch, 1839 in having a large retroventral tibial apophysis, but can be distinguished by: 1) the presence of a dorsal tibial apophysis (vs without dorsal tibial apophysis, compare Figs 71 C, 76 E with figs 2 C, 3 C, 4 C in Zamani & Marusik 2020); 2) the small, blunt, triangular-shaped tegular apophysis (vs rod-shaped, compare Figs 71 B, 76 B with figs 3 B, 4 B in Zamani & Marusik 2020); 3) the large copulatory openings with sclerotized anterior margin, located in the middle of the epigynal plate (vs small, located at posterior of plate, compare Figs 72 A, 77 A with fig. 2 B in Kamura 2021); and 4) the presence of iridescent scales on the dorsal body surfaces (vs without iridescent scales, compare Figs 68, 75 with fig. 15 A, B in Wang et al. 2015).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863CFFC65AA6FCA7DC2A2768.taxon	description	Description. Small, total length 2.26 – 3.68 mm. Carapace smooth, blackish-brown, widest at coxae II, highest near fovea, with sparse iridescent scales. Cervical groove inconspicuous. Fovea longitudinal, dark-red. PER slightly wider than AER. Eyes moderate in size. Chelicerae yellow, promargin and retromargin without teeth, with one long spine on anterior surface. Endites longer than wide, labium wider than long. Sternum yellow, smooth, without pattern, longer than wide, shield-shaped; precoxal triangles and intercoxal sclerites absent. Legs yellow to black; all femora with one spine and tibia tips with nearly white annuli; tibiae I – II usually with five to six proventral and retroventral spines; metatarsus I – II usually with three to four proventral spines and two to three retroventral spines; posterior legs lack spines; metatarsi III with distal preening brush. Leg formula: 4123. Abdomen oval, dorsum black, with dorsal scutum in males; venter yellow; colulus absent. Male palp: femur with well-developed femoral apophysis near middle; tibia with wide, large, strong retroventral tibial apophysis and strong dorsal tibial apophysis; embolus acicular-shaped; tegulum oval, tegular apophysis small; conductor present. Female epigyne: epigynal plate sclerotized; copulatory openings large and far apart, anterior margin sclerotized, located at middle of plate; median septum absent; spermathecae small, located posteriorly; glandular appendages absent; pair of round or oval bursa anteriorly.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE863CFFC65AA6FCA7DC2A2768.taxon	type_taxon	Type species. Punctus taibai sp. nov. (ÁHƉflff) Composition. Two species, Punctus taibai sp. nov. and Punctus maoxian sp. nov..	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	description	Figs 67 – 73	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	materials_examined	Type material. Holotype ♁, CHINA: Shaanxi Province: Xi’an City, Zhouzhi County, Houzhenzi Town, Taibai Mountain National Nature Reserve (33 ° 53.800 ′ N, 107 ° 47.989 ′ E, 1908 m a. s. l.), 17 May 2018, leg. Zhisheng Zhang. Paratypes: 4 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	etymology	Etymology. The specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	diagnosis	Diagnosis. This new species resembles Punctus maoxian sp. nov. in having a similarly shaped embolus, but can be recognized by: 1) the wider retroventral tibial apophysis, with a triangular apophysis on the prolateral margin, and the wider, simple tip (vs retroventral tibial apophysis tapering from base to tip, prolateral margin smooth, and tip bifurcated, cf. Fig. 71 B – C and Fig. 76 B – C), 2) the longer dorsal tibial apophysis (vs short, cf. Fig. 71 C, E and Fig. 76 E), 3) the blunt, triangular-shaped tegular apophysis (vs arc-shaped, cf. Fig. 71 B and Fig. 76 B), 4) copulatory openings directed mesally, towards each other (vs directed posteriorly, cf. Fig. 72 A and Fig. 77 A), and 5) the round bursa (vs oval, cf. Fig. 72 B and Fig. 77 B).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	description	Description. Male (Holotype): total length 2.64, carapace 1.23 long, 1.03 wide; abdomen 1.41 long, 0.87 wide. Eye sizes and interdistances: AME 0.06, ALE 0.09, PME 0.05, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.16, PME – PME 0.05, PME – PLE 0.04, PLE – PLE 0.25, ALE – PLE 0.04. EAW 0.35, CRW 0.53, EAW / CRW 0.66, CRW / CW 0.51. MOA 0.17 long, anterior width 0.14, posterior width 0.17. CH 0.12, CH / AME 2.00. Labium 0.12 long, 0.17 wide. Sternum 0.72 long, 0.65 wide. Leg measurements: Ⅰ 3.83 (1.01, 0.41, 0.98, 0.90, 0.53), II 3.09 (0.81, 0.37, 0.68, 0.76, 0.47), III 2.92 (0.78, 0.36, 0.60, 0.72, 0.46), IV 4.41 (1.14, 0.46, 1.00, 1.18, 0.63). Spination: femora I – IV d 1, femora I – II pl 1, tibia Ⅰ pv 5 rv 5, tibia II pv 5 rv 4, metatarsus Ⅰ pv 3 rv 3, metatarsus II pv 4 rv 2. Coloration: Carapace brown, oval, with several markings resembling flowing water droplets beside fovea and sparse iridescent scales in dorsal view. Abdomen darker in color than carapace, with large dorsal scutum covering almost entire abdomen, with sparse iridescent scales in dorsal view (Fig. 67 A, B). Legs yellow to black; all tibia tips with nearly white annuli. Palp as in Figs 71, 73 A – D. Femoral apophysis protruding, with several setae, near tip of femur. Retroventral tibial apophysis wide, prolateral margin with triangular apophysis, tip blunt, slightly curved towards tegulum; dorsal tibial apophysis long, strong, tapering from base to tip. Tegulum thinner than cymbium, oval; tegular apophysis small, blunt triangular-shaped. Sperm duct distinct, covering more than half length of tegulum, tapering from retrolateral side of tegulum to embolus. Embolus short, slanting posteriorly. Conductor wide, membranous. Female (Paratype): total length 3.66, carapace 1.46 long, 1.22 wide; abdomen 2.20 long, 1.59 wide. Eye sizes and interdistances: AME 0.07, ALE 0.09, PME 0.06, PLE 0.08; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.19, PME – PME 0.05, PME – PLE 0.06, PLE – PLE 0.29, ALE – PLE 0. 05. EAW 0.41, CRW 0.65, EAW / CRW 0.63, CRW / CW 0.53. MOA 0.21 long, anterior width 0.17, posterior width 0.18. CH 0.14, CH / AME 2.00. Labium 0.14 long, 0.22 wide. Sternum 0.88 long, 0.80 wide. Leg measurements: Ⅰ 3.83 (1.23, 0.52, 1.18, 1.10, 0.61), II 3.09 (1.07, 0.47, 0.88, 0.90, 0.58), III 2.92 (0.96, 0.43, 0.73, 0.82, 0.55), IV 4.41 (1.38, 0.52, 1.22, 1.42, 0.75). Spination: femora I – IV d 1, femur I pl 1, tibia Ⅰ pv 5 rv 6, tibia II pv 5 rv 4, metatarsus Ⅰ pv 4 rv 3, tibia II pv 4 rv 2. Other characters as in male, except dorsal scutum absent (Fig. 67 C, D). Epigyne as in Figs 72, 73 E – F. Epigynal plate sclerotized. Copulatory openings large, far apart, located laterally at middle of plate. Copulatory ducts and connecting tubes short. Bursa round, at anterior of plate. Spermathecae globular, separated by about half length of their diameter. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8620FFDC5AA6FF73DADD27D0.taxon	distribution	Distribution. China (Shaanxi Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	description	Figs 74 – 78	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province: Aba Tibetan and Qiang Autonomous Prefecture, Mao County, Mountain Ecosystem Positioning Station (31 ° 41.539´N, 103 ° 53.809´E, 1885 m a. s. l.), 26 May 2015, leg. Luyu Wang. Paratypes: 11 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	etymology	Etymology. This specific name is derived from the type locality; noun in apposition.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	diagnosis	Diagnosis. This new species resembles Punctus taibai sp. nov. in having a similarly shaped embolus, but can be recognized by: 1) the thin retroventral tibial apophysis, tapering from the base to the tip, with a smooth prolateral margin and bifurcated tip (vs retroventral tibial apophysis wide, with a triangular apophysis on the prolateral margin, and wide, not bifurcate tip, compare Fig. 76 B – C with Fig. 71 B – C), 2) the short dorsal tibial apophysis (vs long, compare Fig. 76 E with Fig. 71 C, E), 3) the arc-shaped tegular apophysis (vs blunt triangular-shaped, compare Fig. 76 B with Fig. 71 B), 4) copulatory openings concealed in anterior margins of concavities (vs directed mesally, towards each other, compare Fig. 77 A with Fig. 72 A), 5) the oval bursa (vs round, compare Fig. 77 B with Fig. 72 B).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	description	Description. Male (Holotype): total length 2.26, carapace 1.07 long, 0.89 wide; abdomen 1.19 long, 0.76 wide. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.05; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.14, PME – PME 0.04, PME – PLE 0.04, PLE – PLE 0.23, ALE – PLE 0.03. EAW 0.31, CRW 0.46, EAW / CRW 0.67, CRW / CW 0.52. MOA 0.14 long, anterior width 0.13, posterior width 0.15. CH 0.10, CH / AME 2.00. Labium 0.11 long, 0.17 wide. Sternum 0.65 long, 0.59 wide. Leg measurements: Ⅰ absent, II 3.07 (0.81, 0.37, 0.68, 0.73, 0.48), III 2.71 (0.72, 0.34, 0.55, 0.68, 0.42), IV 4.07 (1.05, 0.40, 0.96, 1.07, 0.59). Spination: femora II – IV d 1, tibia II pv 5 rv 5, metatarsus II pv 4 rv 2. Coloration. Carapace brown, oval, with several markings resembling flowing water droplets beside fovea and sparse iridescent scales in dorsal view. Abdomen darker in color than carapace, with large dorsal scutum covering nearly entire abdomen and sparse iridescent scales dorsally (Fig. 74 A, B). Legs yellow to black; all tibia tips with nearly white annuli. Palp as in Figs 76, 88 A – D. Femoral apophysis protruding, with several setae, near tip of femur. Retroventral tibial apophysis wide, tapering from base to tip, tip bifurcated, short branch triangular; dorsal tibial apophysis short, strong, tapering from base to tip. Tegulum narrower than cymbium, oval; tegular apophysis small, arc-shaped. Sperm duct distinct, covering more than half length of tegulum, tapering from retrolateral side of tegulum to embolus. Embolus short, slanting posteriorly. Conductor thin, transparent. Female (Paratype): total length 3.68, carapace 1.36 long, 1.16 wide; abdomen 2.32 long, 1.52 wide. Eye sizes and interdistances: AME 0.07, ALE 0.09, PME 0.06, PLE 0.07; AME – AME 0.03, AME – ALE 0.01, ALE – ALE 0.18, PME – PME 0.05, PME – PLE 0.06, PLE – PLE 0.29, ALE – PLE 0. 04. EAW 0.41, CRW 0.62, EAW / CRW 0.66, CRW / CW 0.53. MOA 0.20 long, anterior width 0.18, posterior width 0.17. CH 0.13, CH / AME 1.86. Labium 0.13 long, 0.21 wide. Sternum 0.84 long, 0.74 wide. Leg measurements: Ⅰ 4.54 (1.19, 0.49, 1.21, 1.05, 0.60), II 3.82 (1.03, 0.46, 0.87, 0.87, 0.59), III 3.37 (0.89, 0.42, 0.68, 0.84, 0.54), IV 5.15 (1.34, 0.50, 1.22, 1.35, 0.74). Spination: femora I – IV d 1, femur I pl 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 4, metatarsus Ⅰ pv 4 rv 3, tibia II pv 4 rv 2. Other characters as in male, except dorsal scutum absent (Fig. 74 C, D). Epigyne as in Figs 77, 78 E – F. Epigynal plate sclerotized. Copulatory openings large, anterior margin sclerotized, towards posteriorly, located at middle of plate. Copulatory ducts and connecting tubes short. Bursa oval, at middle of plate. Spermathecae small, globular, close to each other. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8625FFD25AA6FA79DADD2732.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8628FFD55AA6F8B5D91E2297.taxon	type_taxon	(type species: Acrolithus lingyun Liu & Li, 2022, by original designation).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8628FFD55AA6F8B5D91E2297.taxon	diagnosis	Diagnosis. This genus can be distinguished from all other phrurolithid genera by the male palpal tegulum with a spiculate prolateral tegular apophysis and long, needle-like embolus; it can also be identified by the dark, slanted male palpal bulb with a nearly round distal tegular apophysis. Female epigynal plate with a pair of large shallow concavities.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8628FFD55AA6F8B5D91E2297.taxon	description	Description. Medium-sized spiders. Carapace oval, dark brown, highest near fovea, with several distinct markings resembling flowing water droplets beside fovea. Chelicerae with two spines anteriorly (one long and another short). Femora I – IV with one dorsal spine, femur I with three to four prolateral spines, femur II with one or two prolateral spines; tibia I usually with seven pairs of ventral spines, tibia II usually with six pairs of ventral spines; metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines. Abdomen oval, with large dorsal scutum nearly covering entire abdomen in males. Male palp: femoral apophysis well developed; tibia with one retrolateral tibial apophysis and one small retroventral tibial apophysis (absent in some species); tegulum wider than cymbium, with spiculate prolateral tegular apophysis; embolus long, needle-like. Female epigyne: epigynal plate with median septum and pair of large shallow concavities; copulatory ducts longer than connecting tubes; glandular appendages digitiform or mastoid-shaped; spermathecae small and round.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	description	Figs 79 – 81	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	materials_examined	Type material. Holotype ♁, CHINA: Zhejiang Province: Lishui City, Longquan County, Fengyang Mountain National Nature Reserve (27 ° 52 ′ 49.46 ″ N, 119 ° 10 ′ 10.76 ″ E, 1494 m a. s. l.), 29 Apr. 2016, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	etymology	Etymology. This specific name is derived from the Latin word “ pugio ”, referring to the dagger-shaped prolateral tegular apophysis; adjective.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	diagnosis	Diagnosis. This new species resembles X. shijiao Liu & Li, 2022 in having a similar embolus shape, but can be recognized by: 1) the wider, shorter and dagger-shaped prolateral tegular apophysis (vs thin and long, cf. Fig. 80 C and fig. 9 G in Liu et al. [2022]), 2) claviform retroventral tibial apophysis with compressed tip (vs cylindrical tip, cf. Fig. 80 E and fig. 9 I in Liu et al. [2022]), 3) retrolateral tibial apophysis bifurcate, with the short branch nearly triangular (vs short branch almost rectangular, cf. Fig. 80 D and fig. 9 I in Liu et al. [2022]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	description	Description. Male (Holotype): total length 3.58, carapace 1.82 long, 1.57 wide; abdomen 1.76 long, 1.27 wide. Eye sizes and interdistances: AME 0.11, ALE 0.14, PME 0.10, PLE 0.10; AME – AME 0.04, AME – ALE 0.02, ALE – ALE 0.28, PME – PME 0.09, PME – PLE 0.08, PLE – PLE 0.44, ALE – PLE 0.07. EAW 0.59, CRW 0.83, EAW / CRW 0.71, CRW / CW 0.53. MOA 0.26 long, anterior width 0.26, posterior width 0.29. CH 0.19, CH / AME 1.73. Labium 0.24 long, 0.27 wide. Sternum 1.00 long, 0.94 wide. Leg measurements: Ⅰ 6.36 (1.75, 0.60, 1.69, 1.46, 0.86), II 5.61 (1.47, 0.56, 1.40, 1.30, 0.88), III 4.95 (1.36, 0.54, 1.01, 1.25, 0.79), IV 7.06 (1.86, 0.63, 1.66, 1.94, 0.97). Spination: femora I – IV d 1, femur Ⅰ pl 4, tibia Ⅰ pv 7 rv 7, tibia II pv 6 rv 6, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Coloration. Carapace nearly round, red-brown, with several markings resembling flowing water droplets beside fovea. Abdomen lighter in color than carapace, with large dorsal scutum covering almost entire abdomen, with indistinct light triangular marking anteriorly (Fig. 79). Legs brown. Palp as in Figs 80 – 81. Femur apophysis large, well-developed, located near middle. Retroventral tibial apophysis extending towards retrolateral side, with wide base and compressed tip. Retrolateral tibial apophysis with wide base, tip bifurcated with two branches, long branch lamelliform, curved towards cymbium, short branch nearly triangular. Tegulum wider than cymbium, pyriform; tegular apophysis absent. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus long, needle-like, slanting upwards. Conductor absent. Female: unknown.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862FFFD75AA6FD11DAD7225F.taxon	distribution	Distribution. China (Zhejiang Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	description	Figs 82 – 84	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	materials_examined	Type material. Holotype ♁, CHINA: Sichuan Province, Aba Tibetan and Qiang Autonomous Prefecture, Mao County, Mountain Ecosystem Positioning Station (31 ° 41.539´N, 103 ° 53.809´E, 1885 m a. s. l.), 26 May 2015, leg. Luyu Wang. Paratypes: 2 ♁ 2 ♀, with same data as holotype.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	etymology	Etymology. The specific name is taken from the family name of Luyu Wang, the collector of this new species.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	diagnosis	Diagnosis. This new species resembles X. pseudostella (Fu, Jin & Zhang, 2014) in having a similarly shaped embolus and large epigynal atrium, but can be recognized by: 1) the wider and shorter prolateral tegular apophysis (vs thin and long, cf. Fig. 83 B and fig. 4 B in Fu et al. [2014]), 2) the absence of a retroventral tibial apophysis (vs presence, cf. Fig. 83 B, C and fig. 4 B, C in Fu et al. [2014]), and 3) the smaller and rectangular-shaped bursa (vs large and oval, cf. Fig. 83 G and fig. 4 E in Fu et al. [2014]).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	description	Description. Male (Holotype): total length 3.26, carapace 1.62 long, 1.44 wide; abdomen 1.64 long, 1.23 wide. Eye sizes and interdistances: AME 0.10, ALE 0.09, PME 0.13, PLE 0.10; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.24, PME – PME 0.06, PME – PLE 0.06, PLE – PLE 0.37, ALE – PLE 0.05. EAW 0.52, CRW 0.82, EAW / CRW 0.65, CRW / CW 0.56. MOA 0.27 long, anterior width 0.22, posterior width 0.25. CH 0.15, CH / AME 1.50. Labium 0.22 long, 0.27 wide. Sternum 0.93 long, 0.89 wide. Leg measurements: Ⅰ 5.78 (1.59, 0.62, 1.53, 1.26, 0.78), II 4.83 (1.36, 0.54, 1.14, 1.08, 0.71), III 4.22 (1.16, 0.49, 0.84, 1.06, 0.67), IV 6.30 (1.73, 0.56, 1.40, 1.67, 0.94). Spination: femora I – IV d 1, femur Ⅰ pl 3, femur II pl 1, tibia Ⅰ pv 7 rv 7, tibia II pv 7 rv 6, metatarsus Ⅰ – II pv 4 rv 3. Coloration. Carapace red brown, with several shapes resembling flowing water droplets beside fovea. Abdomen lighter color than carapace, with large dorsal scutum covered nearly entire abdomen (Fig. 82 A, B). Palp as in Figs 83 A – E, 84 A – C. Retrolateral tibial apophysis base wide, narrower tip digit-shaped and curved towards cymbium; retroventral tibial apophysis absent. Tegulum wider than cymbium, pyriform-shaped; tegular apophysis absent. Sperm duct indistinct, tapering from retrolateral of tegulum to embolus. Embolus long, needle-like, transverse but curved upwards at end. Conductor absent. Female (Paratype): total length 4.41, carapace 1.85 long, 1.68 wide; abdomen 2.56 long, 1.63 wide. Eye sizes and interdistances: AME 0.11, ALE 0.14, PME 0.11, PLE 0.11; AME – AME 0.04, AME – ALE 0.01, ALE – ALE 0.28, PME – PME 0.08, PME – PLE 0.06, PLE – PLE 0.39, ALE – PLE 0.08. EAW 0.58, CRW 0.90, EAW / CRW 0.64, CRW / CW 0.54. MOA 0.29 long, anterior width 0.25, posterior width 0.29. CH 0.15, CH / AME 1.36. Labium 0.22 long, 0.31 wide. Sternum 1.14 long, 1.07 wide. Leg measurements: Ⅰ 7.03 (1.91, 0.66, 1.95, 1.60, 0.91), II 5.91 (1.63, 0.62, 1.48, 1.29, 0.89), III 5.01 (1.34, 0.57, 0.99, 1.30, 0.81), IV 7.72 (2.10, 0.65, 1.79, 2.07, 1.11). Spination: femora I – IV d 1, femora Ⅰ – II pl 3, tibia Ⅰ pv 7 rv 8, tibia II pv 7 rv 5, metatarsus Ⅰ pv 4 rv 4, metatarsus II pv 4 rv 3. Other characters as for male, except dorsal scutum absent, color darker and body larger (Fig. 82 C, D). Epigyne as in Figs 83 F – G, 84 D – E. Epigynal plate weakly sclerotized, with pair of shallow concavities. Copulatory openings small, located at anterior of atrium. Copulatory ducts slender, slightly curved. Bursa nearly rectangular. Glandular appendages mastoid-shaped. Connecting tubes short, wider than copulatory ducts. Spermathecae globular, widely separated. Fertilization ducts located at anterior margin of spermathecae.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE862DFFEA5AA6F9E9DADD2103.taxon	distribution	Distribution. China (Sichuan Province).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	description	Figs 85 – 86	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	materials_examined	Material examined. 1 ♁ 2 ♀, CHINA: Hunan Province: Chenzhou City, Rucheng County, Jiulongjiang National Forest Park, Qinglong Gorge Scenic Spot (25.394029 ° N, 113.785765 ° E, 300 m a. s. l.), 12 May 2018, leg. Yannan Mu.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	diagnosis	Diagnosis. See Liu et al. 2022 a.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	description	Description. See Liu et al. 2022 a. Habitus and genitalia in Figs 85 – 86.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	distribution	Distribution. China (Hunan and Jiangxi provinces).	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
364E87DE8610FFED5AA6FE64DC4F21D3.taxon	discussion	Remark. This species is recorded from Hunan Province for the first time.	en	Mu, Yannan, Zhang, Feng (2023): 5338. Zootaxa 5338 (1): 1-104, DOI: 10.11646/zootaxa.5338.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5338.1.1
