identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E23D6D4B1BFFB1395CFD95EFD41CAC.text	03E23D6D4B1BFFB1395CFD95EFD41CAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplitidae Stoyanow 1949	<div><p>Family Acanthohoplitidae Stoyanow, 1949</p> <p>Remarks</p> <p>The supraorder classification follows the account of Hoffmann et al. (2022), grouping all Devonian to Cretaceous ammonoids in the monophylum Ammonoida. The understanding of the superfamily Acanthohoplitoidea follows Sharikadze (2015), and Frau et al. (2020).</p> </div>	https://treatment.plazi.org/id/03E23D6D4B1BFFB1395CFD95EFD41CAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B1BFFB13965FD6AEF561DFE.text	03E23D6D4B1BFFB13965FD6AEF561DFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protacanthoplites Tovbina 1970	<div><p>Genus Protacanthoplites Tovbina, 1970</p> <p>Type species</p> <p>Parahoplites abichi Anthula, 1900; by original designation.</p> </div>	https://treatment.plazi.org/id/03E23D6D4B1BFFB13965FD6AEF561DFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B1BFFB7396FFC35ED4219AA.text	03E23D6D4B1BFFB7396FFC35ED4219AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protacanthoplites abichi (Anthula 1900)	<div><p>Protacanthoplites abichi (Anthula, 1900)</p> <p>Figs 2A–K, 3A, F</p> <p>Conch shape and ontogeny</p> <p>Lectotype IPUW 1900-3 a</p> <p>Understanding of the lectotype was so far based on the hand-drawing of Anthula (1900: pl. IX(VIII) fig. 2a–c) (see Fig. 2A–B). It is here properly illustrated for the first time (Fig. 2C–G). The specimen corresponds to a small-sized, 3D-preserved internal mould with a calcitic phragmocone and calcareous body chamber with trace of aragonitic shell. The body chamber occupies half of the last whorl, but the peristome is not preserved. Measurements are given in Table 1. Shell shape of the lectotype is characterised by a discoidal (Ww/D ~ 0.41), weakly depressed (Ww/Wh ~ 1.08), very evolute (U/Wh ~ 1.18) subophiocone coiling (U/D ~ 0.45).</p> <p>The embryonic (Ammonitella) stage is poorly preserved, and remains poorly characterised. The first visible whorls of the shell develop a reniform whorl section, crateriform umbilicus, spaced and flat-topped ribs angulate at shoulders followed by smooth interspaces. This corresponds to the Royerianum stage that is typical of the early ontogeny of the basal Acanthohoplitidae (Frau et al. 2020).</p> <p>Then, the whorl section progressively becomes depressed, sub-rounded with convex flanks. The umbilical wall is rounded. The ornamentation consists of an alternation of straight to convex, strong primary ribs and variable number, two to four, of sharp atuberculate secondaries, irregular in thickness and irregularly arranged. Primary ribs are gradually thickened on the flank. They develop strong, elongated tubercles, sometimes clavate, located high on the flank, in tight contact with the succeeding whorl. The tuberculate ribs get a distinct hexagonal cross section. The tubercles divide into two, rarely three, branches over the venter. They strengthen as fold-like ribs, notably the posterior branch, and generally join on the tubercles of the other side. A third discrete rib sometimes appears between two ventral ribs. The secondary ribs are simple, rarely bifurcate, and can be coalescent on the primaries forming a polygyrate pattern. Primary ribs can be followed by an approximated and enlarged secondary rib so that they mimic constrictions. By these features, the ontogeny closely resembles the Gargasense stage sensu Frau et al. (2020) of the Acanthohoplitidae Colombiceras crassicostatum but differences are seen in the quadratic whorl section with rounded venter, and distinctive strengthened ribbing and tubercles. This stage is here referred to as the Abichi stage.</p> <p>As growth proceeds, the number of intercalatories decreases, being mostly one to two, and coalescent ribs become sporadic. Primary ribs are dominantly bifurcate, then simple. They thicken along the flank, especially at the umbilical margin, while the tubercles tend to disappear. Ribs distinctly bend forward on the venter. All ribs are flat-topped or cuneiform over the venter. This stage mimics the Crassicostatum stage sensu Frau et al. (2020) of Colombiceras crassicostatum, but differences are seen in the whorl section that is broadly rounded, with subhexagonal rib section with a convex venter lacking ventral attenuation. This sequence is here referred to as the Crassicostatum-like stage.</p> <p>The ontogenetic sequence of Protacanthoplites abichi is illustrated on Fig. 3A.</p> <p>Specimen IPUW 1900-3b</p> <p>The second syntype of Anthula (1900) is here properly illustrated for the first time (Fig. 2H–K). It is a small, 3D-preserved internal mould with a calcitic phragmocone and calcareous body chamber with remains of aragonitic shell. It shows a slight distortion of the shell at the beginning of the body chamber. The latter occupies half of the last whorl. The peristome is not preserved. Measurements are given in Table 2. Shell shape is characterised by a discoidal (Ww/D ~ 0.39), weakly depressed (Ww/Wh ~ 1.02), very evolute (U/Wh ~ 0,91) subvirgacone coiling (U/D ~ 0.35).</p> <p>The first visible whorl of the shell develops the Royerianum stage, changing into the Abichi stage of the lectotype Protacanthoplites abichi. To the difference, the ribbing is more gracile, with discrete spiny tubercles. The venter is narrower. As growth proceeds, the Crassicostatum-like stage initiates on the body chamber, but it is also much more gracile and regular than the lectotype of Protacanthoplites abichi, and develops flexuous primary ribs. By these features, specimen IPUW 1900-3b poorly matches the lectotype.</p> <p>Suture line</p> <p>Anthula (1900: pl. IX (VIII) fig. 2c) provided a partial suture line drawing for Protacanthoplites abichi showing a quinquelobate pattern. The lectotype seems to be immature as it does not show any sign of suture approximation at the end of the phragmocone. The suture morphogenesis of that species has subsequently been examined by Mikhailova (1957: fig. 5; 1958: fig. 1) and Tovbina (1970: fig. 4). According to these authors, the umbilical lobe is tripartite and rather symmetrical. A wide umbilical saddle is divided into two parts, the outer of which in turn becomes bifid. The ventral and dorsal lobes are bifid and complicated by lateral teeth. The ventral lobe has a low median saddle.</p> <p>Age</p> <p>The litho- and biostratigraphy of the Akusha locality was documented by Mordvilko (1962) and Bogdanova &amp; Mikhailova (2016), but none of these authors have reported the type stratum of Protacanthoplites abichi. The species is, however, known to occur in the ‘ Acanthoplites (sic) aschiltaensis – Acanthoplites uhligi Zone’ sensu Mordviko (1962) of Dagestan (Bogdanova &amp; Mikhailova 2016). Correlation of this zone with the SMAZ is still not obtained with certainty, but it may cover part of the Epicheloniceras martini and Parahoplites melchioris standard zones (Luber et al. 2017; Frau, 2020a, 2020b).</p> </div>	https://treatment.plazi.org/id/03E23D6D4B1BFFB7396FFC35ED4219AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B1CFFB6397DFD99E8931CEA.text	03E23D6D4B1CFFB6397DFD99E8931CEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplites Tovbina 1970	<div><p>Genus Acanthohoplites Tovbina, 1970</p> <p>Type species</p> <p>Parahoplites aschiltaensis Anthula, 1900; by subsequent designation of Roman (1938).</p> </div>	https://treatment.plazi.org/id/03E23D6D4B1CFFB6397DFD99E8931CEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B1CFFB9394EFD28EEE01F0B.text	03E23D6D4B1CFFB9394EFD28EEE01F0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplites aschiltaensis (Anthula 1900)	<div><p>Acanthohoplites aschiltaensis (Anthula, 1900)</p> <p>Figs 2L–R, 3B–C, F–J, 4–6</p> <p>Conch shape and ontogeny</p> <p>Lectotype PMU.24105</p> <p>Understanding of the lectotype was so far based on the hand-drawing of Anthula (1900: pl. X(IX) fig. 2a–b) (see Fig. 2L–M). It is here properly illustrated (Fig. 2N–R). The lectotype shows the same state of preservation as that of Protacanthoplites abichi described above. Peristome is not preserved. The body chamber occupies half of the outer whorl. Measurements are given in Table 3. Its shell shape is characterised by a moderate size, discoidal (Ww/D ~ 0.35), weakly compressed (Ww/Wh ~ 0.82), subevolute (U/Wh ~ 0.76) and subvirgacone coiling (U/D ~ 0.33).</p> <p>The ontogenetic sequence develops four stages following the worn embryonic whorls (Fig. 3B) that closely match those of Protacanthoplites abichi described above. The first visible whorls bear the latest part of a typical Royerianum stage that is well visible on the left face. It changes to an Abichi stage almost identical to that of Protacanthoplites abichi, except its more regular pattern of secondary ribs at the beginning. The Abichi stage occupies one whorl and a half. As growth proceeds, the primary ribs lose their tubercles, while secondaries become irregular, and start decreasing. Ribbing changes progressively on the late phragmocone into an alternation of simple, primary ribs, and generally one secondary, irregular, both in length and strength on the flank. The secondaries can be indistinctly branched on the primary ribs in the lower third of the flank or at mid-flank. All ribs are flat-topped or cuneiform over the venter. This is almost similar to the Crassicostatum-like stage of Protacanthoplites abichi, but differences are seen in the progressive change of the whorl section becoming weakly compressed, subrectangular, higher than wide.</p> <p>The Crassicostatum-like stage changes again in the late phragmocone into a regular alternation of strong, slightly flexuous, primary ribs, with a slight retrocurvature at the umbilical margin, and a single secondary starting high on the flank. The maximum whorl width is reached in the lower third of the flank. Twenty-five ventral ribs are observed on the last half whorl. They are distinctly flat-topped over the venter. These features conform to the adult ontogeny of Colombiceras crassicostatum referred to as the Tobleri stage (Frau et al. 2020). The Tobleri stage starts around D ~ 40 mm (estimation) and occupies the rest of the outer whorl.</p> <p>Paralectotype PMU.24106</p> <p>This paralectotype was not figured by Anthula (1900). It corresponds to a fragment of a large-sized phragmocone displaying the features of the Tobleri stage, including a rather regular alternation of strong, moderately flexuous primary ribs, with a slight retrocurvature at the umbilical margin, and a single secondary of variable height (Fig. 4).</p> <p>Paralectotype PMU.24107</p> <p>Anthula (1900: pl. XI(X) fig. 1) provided a hand-drawing of this paralectotype. It is here properly illustrated for the first time (Figs 5A–B, 6A–C). It corresponds to a 3D-preserved internal mould but of larger size compared to the lectotype. Measurements are given in Table 3. Estimated diameter is about 297 mm. Its phragmocone is broken, but the general shell shape is virgacone (U/D ~ 0.39) with extremely discoidal (Ww/D ~ 0.25), very evolute (U/Wh ~ 1.19) and weakly compressed coiling (Ww/ Wh ~ 0.77). Its last preserved sutures are approximated suggesting its maturity.</p> <p>The first preserved whorls bear a late Abichi stage, consisting in strong primaries, simple or bifurcate, bearing tubercle-like thickenings at the point of furcation, and a variable number of strong secondaries. It gives way to a short Crassicostatum-like stage (estimation: D ~ 35 mm), made of strong primaries, simple or bifurcate, with decreasing thickenings on the flank, and generally one strong secondary. All ribs are flat-topped or cuneiform over the venter. Then, a Tobleri stage develops at D ~ 50 mm (estimation) bearing spaced, sharper primary ribs, being slightly flexuous with a slight retrocurvature at the umbilical margin. Secondary ribs are unique, high on the flank, so that they are almost covered by the succeeding whorl. Ribbing changes in the last two whorls into more spaced and enlarged, more flexuous, primaries with a maximum on the body chamber. Primary ribs start with a distinctive retrocurvature at the umbilical margin except approaching the aperture, and are simple, rarely bifurcate, over the flank. The secondaries are unique, and generally start in the middle, or upper flank. This last ontogenetic stage is here referred to as the Aschiltaensis stage (Fig. 3C). Number of ventral ribs cannot be estimated but it seems denser than during the Tobleri stage.</p> <p>Paralectotype PMU.24108</p> <p>Anthula (1900: pl. X(IX) fig. 1) only gave an idealised hand-drawing of the cross section of this paralectotype. A proper illustration is given here (Fig. 6D–H). It corresponds to a half of a moderately large phragmocone. Measurements are given in Table 3. The broken part allows to observe the Abichi stage extending up to D ~ 18 mm (estimation), and then, a part of the Crassicostatum-like stage. Beyond, the ontogeny is rather similar to the lectotype although the Tobleri stage is more robust and shows great affinity with specimen PMU.24107. Indeed, the shell shape differs from the lectotype by a more discoidal (Ww/D ~ 0.33) and weakly compressed (Ww/Wh ~ 0.94) coiling, while the umbilicus is more open (U/Wh ~ 1.19). Specimen PMU.24108 further displays a lower number of ventral ribs (estimated at ~ 21) on the last half whorl.</p> <p>Suture line</p> <p>A partial suture line of Acanthohoplites aschiltaensis was drawn by Anthula (1900: pl. X(IX) fig. 3b) based on the lost syntype. It is of quinquelobate type and shows a deep inner lateral lobe with distinctive central and inner branches. This drawing was commented on by Stoyanow (1949, 1958), and his re-examination of the lectotype PMU.24105 highlighted a distinctive symmetrically trifid first lateral lobe, with some asymmetry in its inner branch, and bifid saddles. The lectotype seems to be mature as it shows suture approximation at the end of the phragmocone. The suture morphogenesis of that species has then been examined by Drushchits (1956), and Bogdanova &amp; Mikhailova (2016) based on individuals from Dagestan. The suture line consists of a bifid ventral lobe with a low bifid median saddle; rather symmetrical umbilical lobe with distinctive lateral branches; symmetrical, short and trifid first umbilical lobe; deep inner lateral lobe with distinctive central and inner branches; bifid dorsal lobe with variable digits; broad and high, asymmetrically bifid external and umbilical saddles. The first umbilical saddle (U1/I) is indistinctly asymmetrically trifid. There is no difference between the suture lines of Protacanthoplites abichi and Acanthohoplites aschiltaensis.</p> <p>Age</p> <p>There is no comprehensive description of the Ashil’ta locality that yielded the type material of Acanthohoplites aschiltaensis. Nevertheless, the Acanthohoplites -bearing beds of Akusha consist of a ~ 25 m-thick interval assigned to the ‘ Acanthoplites aschiltaensis – Acanthoplites uhligi Zone’ (Mordvilko 1962). This is the case in the reference Dargi River section (Bogdanova &amp; Mikhailova 2016: figs 10–11). Most of the taxa from the ‘ Acanthoplites aschiltaensis – Acanthoplites uhligi Zone’ of the Dargi River section derive from a siltstone horizon including fossiliferous phosphatic nodules. Acanthohoplites aschiltaensis co-occurs here with Parahoplites sjogreni Anthula, 1900, Phylloceras (Hypophylloceras) velledae Michelin, 1834, Aconeceras sp., and the problematic Cheloniceras sp. (Bogdanova &amp; Mikhailova 2016). This horizon is of regional significance since it also crops out in the Rubas-chai section, in the vicinity of Khuchni (Bogdanova &amp; Mikhailova 2016: fig. 12). A younger Acanthohoplites -bearing horizon is documented in the upper part of the ‘ Acanthoplites aschiltaensis – Acanthoplites uhligi Zone’ in the Dargi River and Rubas-chai sections. Representatives assigned to Acanthohoplites ex gr. aschiltaensis would be still present in the corresponding horizon but the fauna lacks illustration for further confirmation.</p> <p>Subspecies of Acanthohoplites aschiltaensis</p> <p>Klein &amp; Bogdanova (2013) listed three subspecies of Acanthohoplites aschiltaensis in the Fossilium Catalogus. Their validity is discussed below.</p> </div>	https://treatment.plazi.org/id/03E23D6D4B1CFFB9394EFD28EEE01F0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B12FFB83AF1FD12EF3B1AFA.text	03E23D6D4B12FFB83AF1FD12EF3B1AFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplites aschiltaensis var. aplanatus Sinzow 1908	<div><p>Acanthohoplites aschiltaensis var. aplanatus Sinzow, 1908</p> <p>Acanthohoplites aschiltaensis var. aplanatus is based on two, well-preserved, moderate-sized acanthohoplitids from the Mangyshlak Peninsula (Sinzow 1908: pl. V figs 4–7). The subspecies was subsequently considered as a valid species by Kazansky (1914), as it shows a distinctive morphology (moderately evolute coiling, compressed, quadratic then sub-rectangular whorl section, flattened venter in the juvenile and progressively rounded in the adult with convergent outer flanks on the body chamber, narrowly to broadly rounded umbilical wall) and ornamentation (complex rib and tubercle pattern of the inner whorls, early loss of the feeble lateral tubercles). These features better compare to the character and sculpture of Hypacanthoplites as revised by Kennedy et al. (2000). The species is here transferred to that genus pending further investigation. The individuals figured by Luppov &amp; Drushchits (1958: pl. XLVII fig. 4a–в), Eristavi (1961: pl. II fig. 11) and doubtfully Kazansky (1914: pl. III fig. 48a–c) match Hypacanthoplites aplanatus. The specimen figured by Sinzow (1913: pl. 6 fig. 4) better compares to Acanthohoplites lautum laxa Glazunova (1953); the latter taxon having been tentatively transferred to Egoianiceras Avram, 1974 by Frau et al. (2020).</p> </div>	https://treatment.plazi.org/id/03E23D6D4B12FFB83AF1FD12EF3B1AFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B12FFB83AFFFE9BEFCA1C24.text	03E23D6D4B12FFB83AFFFE9BEFCA1C24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplites aschiltaensis var. rotundatus Sinzow 1908	<div><p>Acanthohoplites aschiltaensis var. rotundatus Sinzow, 1908</p> <p>The subspecies is based on a single, incomplete, moderate-sized phragmocone of an acanthohoplitid from the Mangyshlak Peninsula (Sinzow 1908: pl. 5 figs 2–3). The specimen closely resembles the paralectotype PMU.24108 of Acanthohoplites aschiltaensis (compare with Fig. 6D–H). The rounded whorl section of Acanthohoplites aschiltaensis var. rotundatus cannot be retained as a distinctive conch parameter since its whorl width index (Ww/Wh ~ 1) falls in the variability of the type material. Consequently, it seems unnecessary to separate the subspecies rotundatus from Acanthohoplites aschiltaensis. Other citations of the subspecies rotundatus in the literature differ from true Acanthohoplites aschiltaensis (see synonymy list in Appendix).</p> </div>	https://treatment.plazi.org/id/03E23D6D4B12FFB83AFFFE9BEFCA1C24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
03E23D6D4B12FFB83AE2FB39EEB31BA4.text	03E23D6D4B12FFB83AE2FB39EEB31BA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthohoplites aschiltaensis var. subangulata Luppov 1961	<div><p>Acanthohoplites aschiltaensis var. subangulata Luppov, 1961</p> <p>The subspecies is based on two, well-preserved, acanthohoplitids from the southwest termination of the Gissar Range, Uzbekistan (Luppov 1961: pl. I figs 2a–в, 3a–в). The species is broadly similar to Acanthohoplites aschiltaensis but differs by feebly but distinctive bituberculate primary ribs in the juvenile, a compressed quadratic whorl section with a flattened venter during the growth, and fine ribs in the adult. As such, it better conforms to Hypacanthoplites. The individual figured by Kvantaliani (1971: pl. VIII figs 2a–e, 3a–e) compares to with Hypacanthoplites subangulata while that of Khalilov (1988: pl. XI fig. 4) is of doubtful identification.</p> </div>	https://treatment.plazi.org/id/03E23D6D4B12FFB83AE2FB39EEB31BA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Frau, Camille	Frau, Camille (2023): The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900). European Journal of Taxonomy 888 (1): 137-158, DOI: 10.5852/ejt.2023.888.2237, URL: http://dx.doi.org/10.5852/ejt.2023.888.2237
