taxonID	type	description	language	source
D3216F85C44D5CB1BF4B59985626654C.taxon	diagnosis	Diagnosis. Pristimantis gretathunbergae sp. nov., a member of the Pristimantis ridens species group (sensu Reyes-Puig et al. 2020), is characterized by the following combination of characters: (1) dorsal skin surfaces slightly areolate, with dispersed tubercles; venter weakly areolate; discoidal fold present, dorsolateral folds absent; (2) tympanum concealed, indistinguishable or poorly distinguished; annulus and tympanic membrane barely visible in males, not visible in females; tympanic fold from the posterior edge of the eye to the arm insertion; (3) snout short, broadly rounded in dorsal view, moderate in length, rounded and slightly protruding in profile; (4) upper eyelid with a single conical to spine-like, some triangular tubercle, ED wider than IoD; cranial crests absent; (5) dentigerous processes of vomers present, prominent, oblique, each bearing from 5 to 10 teeth; (6) vocal slits and nuptial pads absent; (7) Finger I shorter than Finger II; discs on outer fingers truncate, more than twice width of digit proximal to disc; (8) fingers bearing narrow lateral fringes; (9) three to four low ulnar tubercles, barely visible in preservative; (10) heel bearing a conical tubercles, outer edge of tarsus with three to four low and small conical tubercles, inner edge of tarsus lacking tubercles; (11) inner metatarsal tubercle large and elliptical, 4 - 5 x size of outer, ovoid metatarsal tubercle; supernumerary plantar tubercles low; subarticular tubercles conical; (12) toes bearing narrow lateral fringes; webbing absent; Toe V much longer than Toe III; discs as large as those on outer fingers; (13) dorsal ground coloration usually shades of brown with individual tones of red or yellow with or without scattered orange flecks, and / or larger reddish or distinct brown blotches, or light dorsolateral band; (14) venter uniform dirty white (some specimens exhibit dark spotting) or patternless yellow to orange; (15) groin and inner thighs white, yellow or orange-red, some with flecks matching the dorsal ground color or red; (16) blackish iris, some individuals show very dark red iris and / or red-golden speckling; (17) prominent light upper lip in all females and in some males, while other males exhibit some blotches extending from the nose vertically across the lip, however, the upper border of the light-colored lip patches is still demarcated by the darker nose coloration, except in generally light-colored specimens; (18) SVL up to 36.7 mm in males, up to 46.3 mm in females.	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
D3216F85C44D5CB1BF4B59985626654C.taxon	description	Description of the holotype (Figs 4 A, 5). Adult male (SVL 34.6 mm; head approximately as wide as long (HL / HW = 1.11); snout short, broadly rounded in dorsal view, moderate in length, rounded and slightly protruding in profile, eye to nostril distance 10 % of SVL. Canthus rostralis and loreal region slightly concave, nares situated near tip of snout and slightly dorso-laterally directed, clearly visible in frontal and dorsal view, but not ventrally; interorbital area smooth, the upper eyelid is 1.4 of the IoD; a low and conical upper eyelid tubercle, rest of the head with scattered tubercles, but visible only in live specimens, without crests; tympanic annulus slightly visible, tympanum indistinguishable, tympanic annulus concealed by skin, tympanum of moderate size, ratio TY / EW 0.39, supratympanic fold present, from the outer edge of the eye to posterior the insertion point of the jaw, skin around the tympanum with scattered small tubercles; clearly visible choanae rounded and moderate in size, dentigerous processes of vomer in transverse row between choanae, separated by half of a vomer size, with five teeth on right side and seven on left side; vocal slits absent; tongue slightly longer than wide, 2 / 3 attached to mouth floor, shagreen in texture, with an evident papillae at the anterior 1 / 4 of the tongue; dorsal skin surface shagreen with scattered tubercles, ventral surface weakly areolate, without dorsolateral folds, discoidal fold present, extended from level of arm pit to the groin; cloaca partially smooth, granular in the lower part; hands moderate in size, 30 % of SVL, four or five low and small ulnar tubercles; finger II longer than finger I, expanded disks on fingers II, III, & IV; relative lengths of adpressed fingers I <II <IV <III; finger II subequal in size to finger VI, finger II reaching the disc on finger IV when adpressed; finger III disc 2.4 x wider than distal end of adjacent phalanx; subarticular tubercles rounded, and elevated on lateral view, thenar tubercle long, oval and low; palmar and supernumerary present, slightly visible, no nuptial pads, narrow lateral fringes on fingers; hindlimbs of moderate length, TL 51 % of SVL; relative lengths of adpressed toes I <II <III <V <IV; when adpressed, tip of toe I reach tubercle of toe II; disc of toe IV expanded, 1.9 x wider than distal end of adjacent phalanx; narrow lateral fringes on toes; between one and three non-protuberant subarticular tubercles present (one each on toes I and II, two on toes III and V, and three on toe IV); inner metatarsal tubercle elongated; outer metatarsal tubercles slightly pointed and smaller than inner; tarsal ridge absent, outer tarsal tubercles absent; hands and feet without webbing; finger and toe discs broadly expanded.	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
D3216F85C44D5CB1BF4B59985626654C.taxon	description	Measurements of the holotype. SVL 34.6, HW 12.8, HL 14.2, InD 2.4, IoD 4.1, EW 5.4, ED 4.6, EN 3.6, TY 1.9, TL 17.7, FL 16.2, FAL 8.8, HAL 10.2, BW 8.7, 3 FW 0.8, 3 FD 1.5, 3 TW 0.9, 3 TD 1.4, 4 TW 0.7, 4 TD 1.3.	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
D3216F85C44D5CB1BF4B59985626654C.taxon	etymology	Etymology. The specific name is a noun in the genitive case and is a patronym in honor for Greta Thunberg, a Swedish student, and her global climate activism. Greta initiated a " School Strike for Climate Action " outside the Swedish parliament to demand a radical response to the threat by the ongoing climate change. Then sixteen-year-old Thunberg's example has inspired students worldwide to carry out similar strikes called Fridays For Future that started in August 2018. In December 2018 she addressed world leaders at the COP 24 climate talks in Katowice, Poland, with sharp and unmasked words, and equally impressed a global audience in January 2020 with her unpolitical, direct speech down to the point on " Averting a Climate Apocalypse " at the WEF (World Economic Forum) in Davos, Switzerland. Just recently, she publicly slammed the world leaders at the 26 th UN Climate Change Conference of the Parties (COP 26) in Glasgow, November 2021, for not doing enough to meet the demands of the climate emergency. Greta Thunberg represents the authentic voice that exposes the motivations behind the diplomatic curtain of politicians and business stakeholders. Her voice is essential if we want to revert to and maintain a healthy environment on the planet we all share, and not least, learn to respect its magnificent mega-diversity of life that took millions of years to evolve.	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
D3216F85C44D5CB1BF4B59985626654C.taxon	distribution	Distribution. Pristimantis gretathunbergae sp. nov. is endemic to Panama, but it could occur on near mountains along the border in Colombia. Its currently known distribution covers eastern Panama with records from the Darien Mountains within Embera Comarca and the Maje Mountains within Darien and Panama Provinces, including the type locality at Cerro Chucanti. The distribution continues west into Central Panama, including records from Piedras-Pacora Mountains, Panama Province, and Cerro Bruja, Colon Province, both within Chagres National Park. Farther west across the Panama Canal, P. gretathunbergae sp. nov. is present at Altos del Maria, region of Gaita Hills, Panama Oeste Province, and in the region of El Cope, Omar Torrijos National Park, Cocle Province. Color pattern of specimens from Cerro Brewster, not included in the LDA (DFA) analysis, are consistent with the specimens from Maje Mountains in having a cream dorsum coloration, the margin of the upper lip in females yellow, an iris nearly black with pale dots or speckles, venter dirty white, and general stocky body and head. Due to the unique combination of characters of P. gretathunbergae sp. nov., in particular the blackish non-reticulated iris and light, unpatterned upper lip, that differs from any other related rainfrog in Panama and Colombia, we confidently allocate specimens available only as photo vouchers from Cerro Bruja, Colon Province, and Altos del Maria, Gaita Hills, Panama Oeste Province to the same species. The latter two localities substantially reduce the gap to El Cope, Cocle Province, the origin of the most western specimen of our Group 1. So far, we have not received photographic vouchers for the specimen from El Cope, but the low 16 S-divergence of 2.3 % clearly links it to the undescribed species from the Maje Mountains (see above).	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
D3216F85C44D5CB1BF4B59985626654C.taxon	conservation	Conservation. Habitats occupied by P. gretathunbergae sp. nov. are under latent threat. For example, anthropogenic pressure around Cerro Chucanti and the Maje Mountains most likely will lead to declines of populations through habitat destruction (Batista et al. 2020). Similar scenarios are known and can be expected from the other known sites of P. gretathunbergae sp. nov., as they mostly represent restricted montane areas surrounded by agriculture and pastures, and only a few sites are within protected areas (Chucanti Private Reserve, Chagres National Park, General de Division Omar Torrijos Herera). Greta Thunberg's Rainfrog is, thus far, known only from patches of primary forest and slightly disturbed areas. Unfortunately, in the areas surrounding P. gretathunbergae sp. nov. localities, population declines are related to the chytrid fungus (Batrachochytrium dendrobatidis) and pose an additional serious threat (Rebollar et al. 2014; Voyles et al. 2018). Consequently, P. gretathunbergae sp. nov. should be listed as " Vulnerable (VU) " in the global Red List of the IUCN (2018) according to criteria B 2 ab (iii), because: i) its reduced area of occupancy is less than 2000 km 2, ii) it is known from fewer than ten localities, iii) its range is severely fragmented with continuing decline in extent and / or quality of habitat. The Environmental Vulnerability Score (EVS) of this species is 18, placing it in the upper segment of the high vulnerability categories. This score is based on a contributory score of 6 for distribution limited to Central America in the vicinity of the type locality; 8 for ecological distribution, because it is known only from one forest type, and 4 for reproductive mode, because eggs are laid in moist arboreal situations, and tadpoles undergo direct development (Johnson et al. 2015).	en	Mebert, Konrad, Gonzalez-Pinzon, Macario, Miranda, Madian, Griffith, Edgardo, Vesely, Milan, Schmid, P. Lennart, Batista, Abel (2022): A new rainfrog of the genus Pristimantis (Anura, Brachycephaloidea) from central and eastern Panama. ZooKeys 1081: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.1081.63009, URL: http://dx.doi.org/10.3897/zookeys.1081.63009
