taxonID	type	description	language	source
ECB9CD00E20E8E367B8C104751AD1D66.taxon	diagnosis	Diagnosis. (1) Body size small to moderate, SVL 30.2 - 39.3 mm in 11 adult male specimens; (2) head width slightly larger than head length, HDW / HDL ratio 1.04 - 1.09; (3) snout pointed in dorsal view; (4) tympanum distinct, moderate-sized, TD / ED ratio 0.42 - 0.60; (5) strong vomerine ridge bearing vomerine teeth; (6) margin of tongue not notched behind; (7) hindlimbs short, heels not meeting, tibio-tarsal articulation reaching the region between tympanum and eye; (8) presence of subarticular tubercles and absence of lateral fringes on fingers, relative finger lengths II <I ≤ IV <III; (9) toes with rudiment of webbing at their bases and without lateral fringes, subarticular tubercles only present at the base of each toe; (10) numerous granules present on dorsal surface of body, several large tubercles present on surface of flanks; (11) presence of a barely visible reddish horn-like tubercle at the edge of the upper eyelid; (12) supratympanic fold distinct, whitish; (13) yellowish brown dorsally, with an incomplete dark triangular marking between eyes and usually an X-shaped marking on back of trunk; (14) ventral surface black brown, with white spots on posterior surface of abandon; (15) males with a single subgular vocal sac; (16) presence of nuptial pads with darker nuptial spines on dorsal surface of the first and second fingers in adult males during breeding season, respectively.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
ECB9CD00E20E8E367B8C104751AD1D66.taxon	description	Description of holotype. Adult male. Body moderate-sized, SVL 38.0 mm; head width slightly larger than head length, HWD / HDL 1.09; snout pointed in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.40, pupil vertical; nostril oblique ovoid; canthus rostralis well developed, forming the beginning of a fleshy, protruding ridge, that continues over the upper eyelid, and transitions into a supratympanic fold that terminates in the scapular region; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.54; large ovoid choanae at the base of the maxilla; presence of vomerine ridge bearing vomerine teeth; margin of tongue not notched posteriorly; internal vocal slits present near the rear of the lower mandible. Radioulna length and hand length 0.24 of SVL; fingers without webbing and lateral fringes, relative finger length II <I <IV <III; tips of fingers slightly dilated, round; presence of subarticular tubercles on finger III, and one subarticular tubercle at the bases of each finger; outer metacarpal tubercles indistinct, inner metacarpal tubercles distinct and observably enlarged. Hindlimbs short, tibio-tarsal articulation reaching the region between tympanum and eye when hindlimb is stretched along the side of the body; heels not meeting when the flexed hindlimbs are held at right angles to the body axis; tibia length 0.41 of SVL and foot length 0.61 of SVL; relative toe length I <II <V <III <IV; tips of toes round and slightly dilated; presence of rudimentary webbing on toes but absence of lateral fringes and tarsal folds; one subarticular tubercle at the bases of each toe; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsal skin texture rough with dense granules; granules forming discontinuous X-shaped ridge with two discontinuous dorsolateral ridges on both sides at the central trunk; several large tubercles present on dorsal surface of flanks, thighs, shanks and forearms; four small tubercles present on the edge of upper eyelid, one of which is more prominent; distinct narrow supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; ventral skin texture smooth, several granules present on surface of abandon, ventral and posterior surface of thighs; pectoral gland small, closer to axilla; single femoral gland on rear of thigh.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
ECB9CD00E20E8E367B8C104751AD1D66.taxon	description	Measurements of holotype (in mm). SVL 38.0, HDL 12.0, HDW 13.1, SNT 4.5, IND 3.9, IOD 3.6, ED 4.8, TD 2.6, TED 2.1, HND 9.1, RAD 9.2, FTL 23.2, TIB 15.6.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
ECB9CD00E20E8E367B8C104751AD1D66.taxon	etymology	Etymology. The specific epithet " dongguanensis " is in reference to the type locality, Dongguan City of the new species. We propose the common English name " Dongguan Horned Toad " and Chinese name " Dong Guan Jiao Chan (东莞角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
ECB9CD00E20E8E367B8C104751AD1D66.taxon	distribution	Distribution and natural history. Currently, Megophrys dongguanensis sp. nov. is only known from Mt. Yinping, Guangdong Province, China. It inhabits flowing montane streams and the nearby forest floor and leaf litter at elevations between 100 - 300 m. Advertisement calls of males were noticed from mid-December until April of the next year just before the rainy season. Males were found calling on rocks in the flowing streams. Tadpoles could be found in this period.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
5D795EE73B28175073523DC3F228900E.taxon	diagnosis	Diagnosis. (1) Body size small, SVL 29.9 - 34.9 mm in 11 adult males, 39.4 - 41.9 mm in two adult females; (2) head width slightly larger than head length, HDW / HDL ratio 1.00 - 1.20; (3) snout rounded in dorsal view, tip of snout slightly sharpened; (4) tympanum distinct, moderate-sized, TD / ED ratio 0.43 - 0.61; (5) strong vomerine ridge bearing vomerine teeth; (6) margin of tongue not notched behind; (7) shanks short, heels not meeting when the flexed hindlimbs are held at right angles to the body axis; tibia-tarsal articulation reaching forward to the region between tympanum and eye when hindlimb is stretched along the side of the body; (8) TIB / SVL ratio 0.35 - 0.42, FTL / SVL ratio 0.53 - 0.62; (9) absence of lateral fringes on fingers, presence of an indistinct subarticular tubercle on the bases of each finger, relative finger lengths II <I <IV <III; (10) toes with rudimentary webbing at their bases and without lateral fringes, subarticular tubercles only present on the bases of each toes; (11) dorsal surface with dense granules, surface of flanks and dorsal surface of limbs with large tubercles; (12) edge of eye lid with a small reddish horn-like tubercle; (13) supratympanic fold distinct, forming a depressed supraaxillary gland above insertion of arm; (14) dorsum beige to dark brown, with indistinct light brown patches, with an incomplete dark triangular marking between eyes; (15) males with a single subgular vocal sac, and dense dark villiform nuptial spines present on dorsal surface of first and second fingers during breeding season, respectively; (16) gravid females bear creamy yellow oocytes.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
5D795EE73B28175073523DC3F228900E.taxon	description	Description of holotype. Adult male. Habitus small, SVL 31.3 mm; head width slightly larger than head length, HDW / HDL 1.12; snout rounded in dorsal view, tip of snout slightly sharpened, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.38; nostril oblique ovoid; pupil vertical; canthus rostralis well developed, forming the beginning of a fleshy, protruding ridge, that continues over the upper eyelid, and transitions into a supratympanic fold that terminates in the scapular region; loreal region vertical; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.44; large ovoid choanae at the base of the maxilla; strong vomerine ridge bearing vomerine teeth; margin of tongue weakly notched posteriorly; internal vocal slits present near the rear of the lower mandible. RAD / SVL 0.22, HND / SVL 0.22; absence of lateral fringes and webbing on fingers, relative finger lengths II <I <IV <III; tip of finger rounded, slightly swollen; presence of a distinct subarticular tubercle on the base of each finger; outer metacarpal tubercles indistinct, inner metacarpal tubercles distinct and observably enlarged. Hindlimbs short, tibio-tarsal articulation reaching forward the anterior margin of tympanum when hindlimb is stretched along the side of the body; heels not meeting when the flexed hindlimbs are held at right angles to the body axis; TIB / SVL 0.37 and FTL / SVL 0.55; relative toe lengths I <II <V <III <IV; tips of toes round and slightly dilated; presence of rudimentary webbing on toes but absence of lateral fringes and tarsal folds; presence of a subarticular tubercle only at the bases of each toes; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsal skin texture smooth with dense granules, some of which forming a weak X-shaped skin ridge on center of trunk; surface of flanks with large tubercles; presence of a small horn-like tubercle at the edge of eyelid; distinct supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm, forming a swollen supraaxillary gland above insertion of arm; ventral skin texture smooth with granules on the surface of abdomen; pectoral gland large, equal size to tip of fingers, closer to axilla; single large femoral gland on rear of thigh.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
5D795EE73B28175073523DC3F228900E.taxon	description	Measurements of holotype (in mm). SVL 31.3, HDL 9.6, HDW 10.8, SNT 3.4, IND 3.4, IOD 2.4, ED 3.7, TD 1.6, TED 1.0, HND 6.9, RAD 7.0, FTL 17.3, TIB 11.6.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
5D795EE73B28175073523DC3F228900E.taxon	etymology	Etymology. The specific epithet " nankunensis " is in reference to the type locality of the new species: Mt. Nankun. We propose the common English name " Nankunshan Horned Toad " and Chinese name " Nan Kun Shan Jiao Chan (南昆山角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
5D795EE73B28175073523DC3F228900E.taxon	distribution	Distribution and habits. Currently, Megophrys nankunensis sp. nov. is known only from the type locality, Mt. Nankun in Longmen County, Guangdong Province, China. It inhabits forest floor, leaf litter and the nearby undergrowth rocky mountainous streams (2 - 3 m wide) surrounded by moist subtropical evergreen broadleaved forests at elevations between 300 - 600 m. Breeding season of M. nankunensis sp. nov. is from October to the following March, males were found calling under the leaf litter or rocks (Fig. 5 A) on the ground in the flowing streams, besides, a pair were observed exposed on the floor in a flowing stream, about 2.5 m wide, prior to amplexus (Fig. 5 B) at 20: 09 P. M. on 20 October 2015. Tadpoles were not observed in this period.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
B6ECDD6CF7DC666526CB3C0EA8369F53.taxon	diagnosis	Diagnosis. (1) Body slender and small-sized, SVL 30.4 - 33.9 mm in nine adult males, 34.1 - 37.5 mm in two adult females; (2) head width slightly larger than head length, HDW / HDL ratio 1.04 - 1.06; (3) snout rounded in dorsal view; (4) eye large, tympanum distinct, moderate-sized, TD / ED ratio 0.50 - 0.59; (5) weak vomerine ridge bearing vomerine teeth; (6) tongue weakly notched posteriorly; (7) hindlimbs slender, heels overlapping when the flexed hindlimbs are held at right angles to the body axis, tibia-tarsal articulation reaching forward to the middle of eye when hindlimb is stretched along the side of the body; (8) absence of lateral fringes on fingers, presence of an indistinct subarticular tubercle on the bases of each finger, relative finger lengths II <I <IV <III; (9) toes with rudimentary webbing at their bases and without lateral fringes, subarticular tubercles only present at the base of toe I and II; (10) dorsal skin rough, presence of black spines on granules of dorsal skin, and occasionally present on canthus rostralis and margin of tympanum, presence of large tubercles on flanks, dorsal body and limbs; (11) four prominent parallel dorsolateral ridges with granules bearing black spines on back of trunk, the middle two ridges forming a X-shaped ridge occasionally; (12) a reddish horn-like tubercle bearing a black spine at its tip at the edge of eye lid; (13) distinct supratympanic fold bearing black spines; (14) beige to brownish red above, with an hollow dark triangle between eyes and a rectangular dark marking on the center of the back of trunk; (15) males with a single subgular vocal sac, and presence of nuptial pads bearing darker nuptial spines on dorsal surface of the first and second fingers in adult males during breeding season, respectively; (16) gravid females bear creamy yellow oocytes.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
B6ECDD6CF7DC666526CB3C0EA8369F53.taxon	description	Description of holotype. Adult male. Habitus slender and small, SVL 32.0 mm; head width slightly larger than head length, HDW / HWL 1.04; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.39; nostril oblique ovoid; pupil vertical; canthus rostralis well developed, forming the beginning of a fleshy, protruding ridge, that continues over the upper eyelid, and transitions into a supratympanic fold that terminates in the scapular region; loreal region vertical; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.52; large ovoid choanae at the base of the maxilla; weak vomerine ridge bearing vomerine teeth; margin of tongue weakly notched posteriorly; internal vocal slits present near the rear of the lower mandible .. RAD / SVL 0.25; absence of lateral fringes and webbing on fingers, relative finger lengths II <I <IV <III; tip of finger rounded, slightly swollen; presence of an indistinct subarticular tubercle on the base of each finger; outer metacarpal tubercles indistinct, inner metacarpal tubercles distinct and observably enlarged. Hindlimbs long, tibio-tarsal articulation reaching forward to the middle of eye when hindlimb is stretched along the side of the body; heels overlapping when the flexed hindlimbs are held at right angles to the body axis; TIB / SVL 0.46 and FTL / SVL 0.62; relative toe lengths I <II <V <III <IV; tips of toes round and slightly dilated; presence of rudimentary webbing on toes but absence of lateral fringes and tarsal folds; presence of a subarticular tubercle only at the bases of the first and second toes; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsum rough with dense granules bearing spines; canthus rostralis, margin of tympanum, supratympanic fold and upper lip with dense spines; presence of large tubercles bearing spines on dorsal surface of body, surface of flanks and dorsal and posterolateral surface of limbs; prominent parallel dorsolateral ridges with granules bearing spines on back of trunk; presence of a horn-like tubercle bearing a spine at its tip at the edge of eye lid; distinct supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; ventral skin texture smooth, the lower lip bears spines; sides of belly with large tubercles; ventral skin texture of thighs smooth with a few small tubercles, posterior surface and surface around anus with large tubercles bearing spines; surface of tibia-tarsal with a few tubercles bearing spines; presence of spines on lateral sides of fingers and toes; pectoral gland moderate-sized, closer to axilla; single femoral gland on rear of thigh, distinctly smaller than pectoral gland.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
B6ECDD6CF7DC666526CB3C0EA8369F53.taxon	description	Measurements of holotype (in mm). SVL 32.2, HDL 11.5, HDW 11.4, SNT 3.6, IND 3.5, IOD 3.3, ED 4.2, TD 2.3, TED 1.7, HND 8.0, RAD 8.1, FTL 20.5, TIB 14.7.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
B6ECDD6CF7DC666526CB3C0EA8369F53.taxon	etymology	Etymology. The specific epithet " jiulianensis " is in reference to the known localities of the new species: Mt. Jiulian and Nankunshan Natuire Reserve located in the Jiulian Mountains range. We propose the common English name " Jiulianshan Horned Toad " and Chinese name " Jiu Lian Shan Jiao Chan (九连山角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
B6ECDD6CF7DC666526CB3C0EA8369F53.taxon	distribution	Distribution and natural history. Currently, Megophrys jiulianensis sp. nov. is known from Mt. Nankun in Guangdong Province and the type locality, Jiulian Nature Reserve in Jiangxi Province, China. It inhabits forest floor, leaf litter and the nearby undergrowth mountainous streams surrounded by moist subtropical evergreen broadleaved forests at elevations between 500 - 800 m. Breeding season of M. jiulianensis sp. nov. is from March to July, males were usually found staying while calling on leaves (Fig. 7 A), about 0.1 - 0.3 m above the ground. After the rain, numerous individuals can be easily found on the road, and a female individual from Mt. Nankun was observed feeding on an earthworm (Fig. 7 B) on 20: 45 p. m., 21 March 2016. Tadpoles could be found all year round. Megophrys jiulianensis sp. nov. is sympatric with M. nankunensis sp. nov. and M. mangshanensis at Mt. Nankun.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
A42AC86A0FE5355E09E6DD6938D92E11.taxon	diagnosis	Diagnosis. (1) Body small-sized, SVL 30.5 - 37.3 mm in 10 adult males; (2) snout rounded in dorsal view; (3) tympanum distinct, moderate-sized, TD / ED ratio 0.43 - 0.57; (4) vomerine ridge and vomerine teeth present; (5) tongue notched posteriorly; (6) absence of lateral fringes and webbing on fingers, presence of narrow lateral fringes and rudimentary webbing on toes; (7) presence of a subarticular tubercle at the base of each finger and toe; (8) hindlimbs slender, heels overlapping, tibio-tarsal articulation reaching between the posterior corner to the center of eye; (9) TIB / SVL ratio 0.45 - 0.51 and FTL / SVL ratio 0.61 - 0.73; (10) dense conical granules present on surface of temporal region, upper lip, and from loreal region to the tip of snout; (11) granules and tubercles on dorsal surface forming a discontinuous X-shaped ridge and a pair of discontinuous dorsolateral ridges on back of trunk; (12) supratympanic fold distinct, whitish tan; (13) brown dorsally, with a dark triangular marking with light yellow edge between eyes, and an X-shaped or V-shaped marking with light yellow edge on the center of the back of trunk; (14) presence of a single subgular vocal sac in males; (15) nuptial pads and nuptial spines invisible in males during breeding season.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
A42AC86A0FE5355E09E6DD6938D92E11.taxon	description	Description of holotype. Adult male. Body size small, SVL 32.5 mm; head length and head width almost isometric, HDW / HDL 0.99; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.37, pupil vertical; nostril oblique ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.48; large ovoid choanae at the base of the maxilla; presence of vomerine ridge bearing vomerine teeth; margin of tongue notched posteriorly; internal vocal slits present near the rear of the lower mandible. RAD / SVL 0.25, HND / SVL 0.24; fingers without webbing and lateral fringes, relative finger length II <I <IV <III; tips of fingers slightly dilated, round; one subarticular tubercle at the bases of each finger; outer and inner metacarpal tubercles distinct, and the inner one observably enlarged. Hindlimbs slender, tibio-tarsal articulation reaching forward to the center of the eye when hindlimb is stretched along the side of the body; heels overlapping when the flexed hindlimbs are held at right angles to the body axis; TIB / SVL 0.49 and FTL / SVL 0.69; relative toe length I <II <V <III <IV; tips of toes round and slightly dilated; toes with narrow lateral fringes, rudimentary webbing; one subarticular tubercle at the bases of each toes; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsal skin texture rough; head surface rough, with small tapered granules densely covering from temporal region, upper lip, loreal region to tip of snout; granules forming discontinuous X-shaped ridge with two discontinuous dorsolateral ridges on both sides at the central trunk; large tubercles on flanks; a horn-like prominent tubercle on the edge of the upper eyelid; distinct supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; ventral skin texture smooth, with several large granules and tubercles on two sides; ventral skin texture of thighs smooth, with a few small tubercles; pectoral gland larger, closer to axilla; single femoral gland on rear of thigh.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
A42AC86A0FE5355E09E6DD6938D92E11.taxon	description	Measurements of holotype (in mm). SVL 32.5, HDL 11.5, HDW 11.4, SNT 3.7, IND 3.5, IOD 3.3, ED 4.2, TD 2.0, TED 1.7, HND 8.0, RAD 7.8, FTL 22.3, TIB 15.9.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
A42AC86A0FE5355E09E6DD6938D92E11.taxon	etymology	Etymology. The specific epithet " nanglingensis " is in reference to the type locality of the new species, Nanling Nature Reserve of the Nanling Mountains. We propose the common English name " Nanling Horned Toad " and Chinese name " Nan Ling Jiao Chan (南岭角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
A42AC86A0FE5355E09E6DD6938D92E11.taxon	distribution	Distribution and natural history. Currently, Megophrys nanglingensis sp. nov. is known from Nanling Nature Reserve and the neighboring Mangshan Nature Reserve (between elevations of 1000 - 1300 m), together with Mt. Qiyun (between elevations of 690 - 1400 m). It inhabits streams in bamboo forests. Males are frequently heard calling during August and December. Tadpoles could be found in this period. Megophrys nanlingensis sp. nov. is sympatric with M. mangshanensis and M. popei in Nanling Nature Reserve and the neighboring Mangshan Nature Reserve.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
D8D4D64FCB28EBF30EA98E6B528E4B41.taxon	diagnosis	Diagnosis. (1) Body size small, SVL 31.0 - 34.1 mm in four adult males and body size moderate, SVL 38.5 - 42.8 mm in nine adult females; (2) tympanum distinct, slightly convex, moderate-sized, TD / ED ratio 0.47 - 0.52; (3) vomerine teeth absent; (4) margin of tongue not notched posteriorly; (5) hindlimbs short, heels not meeting, tibia-tarsal articulation reaching forward to the region between posterior corner of eye and posterior margin of tympanum; (6) TIB / SVL ratio 0.39 - 0.44, FTL / SVL ratio 0.56 - 0.64; (7) fingers without lateral fringes, presence of a subarticular tubercle at the bases of each finger, relative finger lengths II <I = IV <III; (8) toes with rudimentary webbing at their bases and without lateral fringes, subarticular tubercles only present at the base of each toe; (9) numerous granules present on dorsal surface of body, several large tubercles present on surface of flanks and dorsal surface of limbs; (10) presence of a small horn-like tubercle at the edge of the upper eyelid; (11) supratympanic fold distinct, whitish; (12) yellowish brown or reddish brown dorsally, with an incomplete dark triangular marking between eyes and an X-shaped marking on back of trunk; (13) ventral surface greyish brown, ventral surface of abdomen with creamy white nebulous patches and black spots; (14) males with a single subgular vocal sac; (15) gravid females bear creamy yellow oocytes.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
D8D4D64FCB28EBF30EA98E6B528E4B41.taxon	description	Description of holotype. Adult male. Habitus small, SVL 31.0 mm; head width slightly larger than head length, HDW / HWL 1.03; snout rounded in dorsal view, tip of snout slightly sharpened, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.41; nostril oblique ovoid; pupil vertical; canthus rostralis well developed; loreal region vertical; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.47; large ovoid choanae at the base of the maxilla; weak vomerine ridge present, vomerine teeth absent; margin of tongue not notched posteriorly; internal vocal slits present near the rear of the lower mandible. RAD / SVL 0.24, HND / SVL 0.22; absence of lateral fringes and webbing on fingers, relative finger lengths II <I = IV <III; tip of finger rounded, slightly swollen; presence of a distinct subarticular tubercle on the base of each finger; outer metacarpal tubercles indistinct, inner metacarpal tubercles distinct and observably enlarged. Hindlimbs short, tibio-tarsal articulation reaching forward the posterior corner of eye when hindlimb is stretched along the side of the body; heels not meeting when the flexed hindlimbs are held at right angles to the body axis; TIB / SVL 0.43 and FTL / SVL 0.61; relative toe lengths I <II <V <III <IV; tips of toes round and slightly dilated; presence of rudimentary webbing on toes but absence of lateral fringes and tarsal folds; presence of a subarticular tubercle only at the bases of each toes; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsal skin texture rough with dense granules, some of which forming an X-shaped skin ridge on center of trunk; surface of flanks with large tubercles; presence of a small horn-like tubercle at the edge of eye lid; distinct supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; superior margin of tympanum in connect with supratympanic fold; ventral skin texture smooth with granules on the surface of abdomen; pectoral gland large, closer to axilla; single large femoral gland on rear of thigh.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
D8D4D64FCB28EBF30EA98E6B528E4B41.taxon	description	Measurements of holotype (in mm). SVL 30.8, HDL 11.9, HDW 11.7, SNT 3.5, IND 3.0, IOD 2.8, ED 3.5, TD 1.8, TED 1.7, HND 8.5, RAD 7.2, FTL 21.8, TIB 15.1	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
D8D4D64FCB28EBF30EA98E6B528E4B41.taxon	etymology	Etymology. The specific epithet " wugongensis " is in reference to the type locality of the new species in the Wugong Mountains. We propose the common English name " Wugongshan Horned Toad " and Chinese name " Wu Gong Shan Jiao Chan (武功山角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
D8D4D64FCB28EBF30EA98E6B528E4B41.taxon	distribution	Distribution and habits. Currently, Megophrys wugongensis sp. nov. is known from the type locality, Yangshimu Scenic Area, Pingxiang City, Jiangxi Province at approximate 550 m a. s. l., Wugongshan Scenic Area, Ji'an City, Jiangxi Province at approximate 1050 - 1080 m a. s. l., all located in the Luoxiao Mountains in eastern China. All specimens were collected on leaf litter near a stream in the bamboo forest, males were not heard calling. In consideration of the invisible nuptial pad and nuptial spines in all male specimens and the undeveloped fallopian tubes in all female specimens, the breeding season of M. wugongensis sp. nov. still remains unknown. Tadpoles were not observed. Megophrys wugongensis sp. nov. is sympatric with M. jinggangensis in all localities.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
36E013866E1B8AF51686A7CE0120BB61.taxon	diagnosis	Diagnosis. (1) Body size small, SVL 30.1 - 30.8 mm in two adult males and SVL 36.3 mm in two adult females; (2) head length slightly larger than head width, HDW / HDL ratio 0.98 - 0.99; (3) tympanum distinct, moderate-sized, TD / ED ratio 0.51 - 0.58, upper 1 / 4 part of the tympanum concealed by supratympanic fold; (4) vomerine teeth absent; (5) margin of tongue not notched posteriorly; (6) heels overlapping, tibia-tarsal articulation reach forward to the tympanum in males and to the eye in females; (7) TIB / SVL ratio 0.47 - 0.53, FTL / SVL ratio 0.68 - 0.74; (8) fingers without lateral fringes, presence of a subarticular tubercle at the bases of each finger, relative finger lengths II = IV <I <III; (9) toes with rudimentary webbing at their bases and narrow lateral fringes, subarticular tubercles only present at the base of each toe; (10) numerous granules scattered with tubercles present on dorsal surface of body, limbs and surface of flanks, some of which forming a V-shaped, \ / - shaped or X-shaped skin ridge on central back of trunk; (11) presence of a small horn-like tubercle at the edge of the upper eyelid; (12) supratympanic fold distinct; (13) light brown to dark brown dorsally, with a dark triangular marking between eyes; (14) a pair of dark longitudinal and irregular marking with white edges on its upper side on ventrolateral surface of flanks; (15) surface of throat and chest greyish brown with dark brown patches and creamy white spots, surface of abdomen greyish white with creamy white and orange spots; (16) ventral surface of thighs with dense small whitish tubercles.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
36E013866E1B8AF51686A7CE0120BB61.taxon	description	Description of holotype. Adult male. Habitus small, SVL 30.8 mm; head length slightly larger than head width, HDW / HWL 0.98; snout rounded in dorsal view, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eye large, ED / HDL 0.30; nostril oblique ovoid; pupil vertical; canthus rostralis well developed; loreal region vertical; internasal distance slightly larger than interorbital distance; tympanum distinct, moderate-sized, TD / ED 0.56; large ovoid choanae at the base of the maxilla; weak vomerine ridge present, vomerine teeth absent; margin of tongue not notched posteriorly. RAD / SVL 0.25, HND / SVL 0.30; absence of lateral fringes and webbing on fingers, relative finger lengths II = IV <I <III; tip of finger rounded, slightly swollen; presence of a distinct subarticular tubercle on the base of each finger; outer metacarpal tubercles indistinct, inner metacarpal tubercles distinct and observably enlarged. Hindlimbs long, tibio-tarsal articulation reaching forward to the tympanum when hindlimb is stretched along the side of the body; heels overlapping when the flexed hindlimbs are held at right angles to the body axis; TIB / SVL 0.53 and FTL / SVL 0.74; relative toe lengths I <II <V <III <IV; tips of toes round and slightly dilated; presence of rudimentary webbing and narrow lateral fringes on toes but absence of tarsal folds; presence of a subarticular tubercle only at the bases of each toes; presence of a long ovoid inner metatarsal tubercle and absence of outer metatarsal tubercle. Dorsal skin texture rough with dense granules and scattered with small tubercles, some of which forming a \ / - shaped skin ridge on central back of trunk; presence of a small horn-like tubercle at the edge of upper eye lid; distinct supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; upper 1 / 4 part of the tympanum covered by supratympanic fold; ventral skin texture smooth with granules; pectoral gland large, closer to axilla; single large femoral gland on rear of thigh.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
36E013866E1B8AF51686A7CE0120BB61.taxon	description	Measurements of holotype (in mm). SVL 30.1, HDL 11.6, HDW 11.4, SNT 3.5, IND 3.0, IOD 2.8, ED 3.5, TD 18, TED 1.7, HND 8.5, RAD 7.2, FTL 21.8, TIB 15.1.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
36E013866E1B8AF51686A7CE0120BB61.taxon	etymology	Etymology. The specific epithet " mufumontana " is in reference to the type locality of the new species, Mt. Mufu. We propose the common English name " Mufushan Horned Toad " and Chinese name " Mu Fu Shan Jiao Chan (幕阜山角蟾) ".	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
36E013866E1B8AF51686A7CE0120BB61.taxon	distribution	Distribution and habits. Currently, Megophrys mufumontana sp. nov. is known only from Mt. Mufu, Pingjiang County, Yueyang City, Hunan Province, China at approximate 1300 m a. s. l .. All specimens were collected on leaf litter near a stream (about 5 m wide) surrounded by moist subtropical evergreen broadleaved forests, males were not heard calling. Tadpoles were not observed. Because none of the males have nuptial pads developed and none of the females have fallopian tubes and eggs developed, the breeding season of M. mufumontana sp. nov. remains unknown.	en	Wang, Jian, Lyu, Zhi-Tong, Liu, Zu-Yao, Liao, Cheng-Kai, Zeng, Zhao-Chi, Zhao, Jian, Li, Yu-Long, Wang, Ying-Yong (2019): Description of six new species of the subgenus Panophrys within the genus Megophrys (Anura, Megophryidae) from southeastern China based on molecular and morphological data. ZooKeys 851: 113-164, DOI: http://dx.doi.org/10.3897/zookeys.851.29107, URL: http://dx.doi.org/10.3897/zookeys.851.29107
