taxonID	type	description	language	source
E22D1205F108FFB1FF0799609317FD18.taxon	biology_ecology	Habitat and phenology: — Primary and secondary evergreen broad-leaved forests at elevation of 1400 – 2000 m a. s. l. Creeping epiphyte in canopy of tall trees. Flowering in October – December.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F108FFB1FF0799609317FD18.taxon	distribution	Distribution: — India, Bhutan, Nepal, China, Myanmar, Thailand, Vietnam (Cao Bang, Ha Giang, and Lao Cai provinces).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F108FFB1FF0799609317FD18.taxon	diagnosis	Notes: — Lateral sepals form slender tube at apical half, lower margins fuse along almost all their length, upper margin free at basal half.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F108FFB1FF0799609317FD18.taxon	materials_examined	Studied specimens: — VIETNAM, Cao Bang Province, 27 October 1960, Takhtajan 622 (LE 01055497 http: // en. herbariumle. ru /? t = occ & id = 7253, LE 01055496 http: // en. herbariumle. ru /? t = occ & id = 7252, LE 01055495 http: // en. herbariumle. ru /? t = occ & id = 7251, LE 01055494 http: // en. herbariumle. ru /? t = occ & id = 7250). VIETNAM, Ha Giang Province, Meo Vac 13 Dec 2005, L. Averyanov, P. K. Loc, T. V. Thao, P. V. The, N. S. Khang HAL 8575 (LE 0105549 http: // en. herbariumle. ru /? t = occ & id = 7247). VIETNAM, Ha Giang Province, Yen Minh, November 2004, S. K. Wu, L. K. Phan, X. Gong, J. Y. Xiang, V. T. Nguyen & K. S. Nguyen WP 706 (LE 01055492 http: // en. herbariumle. ru /? t = occ & id = 7248). VIETNAM, Ha Giang Province, Quang Ba 31 October 2019, Truong Ba Vuong, Bui Van Huong BV 441 (VNM 00069926!). VIETNAM, Lao Cai Province, Van Ban 22 March 2002, L. Averyanov, P. K. Loc, D. T. Doan HAL 2629 (LE 01055493 http: // en. herbariumle. ru /? t = occ & id = 7249).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBAFF07999F91F1F446.taxon	biology_ecology	Habitat and phenology: — Dry broad-leaved primary and secondary evergreen and semideciduous forest at elevations 500 – 1000 m a. s. l. Creeping epiphyte. Flowering in March.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBAFF07999F91F1F446.taxon	distribution	Distribution: — India, Myanmar, China, Thailand, Lao, Vietnam (doubtful).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBAFF07999F91F1F446.taxon	discussion	Notes: — According to Seidenfaden (1973) Bulbophyllum forrestii looks similar to B. helenae but actually the median sepal of true B. forrestii is slightly erose to almost entire (vs. irregular erose to denticulate), lip adaxially papillose (vs. smooth in B. helenae).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBAFF07999F91F1F446.taxon	materials_examined	Studied specimens: — CHINA, Yunnan, in the Shweli Valley, June 1912, Forrest G. 8130 (K 000867022 photo!). CHINA, Yunnan, 1917, Forrest G. 17970 (K 000867021 photo!). CHINA, Yunnan, 1923, Forrest, G. 17970 (P 00348028 photo!). LAOS, Xiangkhouang Province, Peak District, Oran Village, around point 19 ° 38 ′ 53 ″ N 103 ° 23 ′ 12 ″ E, primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elevation about 1750 – 1850 m a. s. l., epiphyte on tall mossy tree along ridge edge, not rare, 2 April 2015, N. T. Hiep et al., LA-VN 931 b / TM 1225 (LE 01055537 http: // en. herbariumle. ru /? t = occ & id = 18092, LE 01073555 http: // en. herbariumle. ru /? t = occ & id = 18088 photo).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079ED39475F0C9.taxon	biology_ecology	Habitat and phenology: — Creeping epiphyte on large trees in dry forests at elevation of about 1000 m a. s. l.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079ED39475F0C9.taxon	distribution	Distribution: — Vietnam (Son La Province). India.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079ED39475F0C9.taxon	discussion	Notes: — The species is very close to B. helenae, but can be easily distinguished by the clavate hairs on pedicel, ovary, and abaxial surface of sepals, as well as in having nodular swellings on the thick rhizome. Our discovery represent new national record of this species in the flora of Vietnam.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079ED39475F0C9.taxon	materials_examined	Studied specimens: — INDIA, Nilghiri Hills, s. coll., s. n. (K 000829967 photo!, type of B. nodosum). INDIA, Nilghiri Hills, s. coll., s. n. (K 000829965 photo! http: // specimens. kew. org / herbarium / K 000829965, K 000829966 photo! http: // specimens. kew. org / herbarium / K 000829966, K 000829968 photos! http: // specimens. kew. org / herbarium / K 000829968). VIETNAM, Dien Bien Province, Tua Chua District, Sin Chai Municipality, around point 22 ° 03 ’ 38 ” N 103 ° 19 ’ 56 ” E, primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation of 1350 – 1500 m a. s. l., epiphyte, occasional, 14 December 2010, L. Averyanov et al., CPC 982 a / 3 (LE 01068417 http: // en. herbariumle. ru /? t = occ & id = 15085, LE 01055568 http: // en. herbariumle. ru /? t = occ & id = 7316, LE 01087022 http: // en. herbariumle. ru /? t = occ & id = 18117 photos). VIETNAM, herbarium specimen prepared from plants cultivated in private garden in Ho Chi Minh City, originated from Son La Province, Thuan Chau District, 28 August 2021, Truong Ba Vuong BV 1260 (VNM 00069916, LE 01122925 http: // en. herbariumle. ru /? t = occ & id = 107046 photos), BV 1261 (VNM 00069917), 15 September 2020, Truong Ba Vuong, Ngo Quang Dang, BV 1298 (VNM 00069922).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079CF79298F785.taxon	materials_examined	Type: — VIETNAM, Lai Chau Province, Sin Ho District, Ma Quai Municipality, Can Ti 2 village, around point 22 º 24 ′ 18 ″ N 103 º 22 ′ 45 ″ E, remnants of broad-leaved evergreen closed primary forest on steep mountain slopes composed with sandstone at elevation about 1100 m a. s. l., creeping epiphyte on tall mossy trees, flowers odorless, median sepal and petals white, lateral sepals purple-brown, lip white with finest purple-brown marks, common, 17 Nov. 2006, Nguyen Tien Hiep, L. Averyanov, Pham Van The, HAL 9849 (LE 01055989 http: // en. herbariumle. ru /? t = occ & id = 7777, LE 01087186 http: // en. herbariumle. ru /? t = occ & id = 18711 photos, LE 01087185 http: // en. herbariumle. ru /? t = occ & id = 18710 drawing).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079CF79298F785.taxon	biology_ecology	Habitat and phenology: — Primary and secondary evergreen broad-leaved forests on sandstone at an elevation of about 1100 m a. s. l. Creeping epiphyte in canopy of tall trees. Flowering in November.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079CF79298F785.taxon	distribution	Distribution: — Vietnam (Lai Chau Province). Endemic.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F102FFBBFF079CF79298F785.taxon	discussion	Notes: — Bulbophyllum paraemarginatum differs from B. emarginatum by the color of flowers, the irregular dentate petals (vs. slightly erose), upper margin of lateral sepals free only at base (vs. free at basal half), and entire upper margin of stelidia (vs. stelidia margin erose).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFB9FF079A2A950CF42A.taxon	materials_examined	Type: — VIETNAM, Quang Binh Province, Minh Hoa District, Thuong Hoa Municipality, environs of Mo O O O Village, around point 17 ° 39 ′ 31.7 ″ N 105 ° 54 ′ 48.2 ″ E, primary broad-leaved closed forest on very steep rocky slope of remnant mountain composed of highly eroded solid crystalline limestone at elevation of 400 – 500 m a. s. l., creeping epiphytic on tall tree, not rare, 23 July 2011, N. T. Hiep, L. Averyanov, N. S. Khang, N. Q. Vinh, CPC 3657 (LE 01057821 http: // en. herbariumle. ru /? t = occ & id = 8321, LE 01087269 http: // en. herbariumle. ru /? t = occ & id = 18990 photos, LE 01087271 http: // en. herbariumle. ru /? t = occ & id = 18992 drawing).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFB9FF079A2A950CF42A.taxon	biology_ecology	Habitat and phenology: — Evergreen broad-leaved primary limestone forests at elevation of around 400 – 1000 m a. s. l. Trunk and branch epiphyte. Flowering in June – July.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFB9FF079A2A950CF42A.taxon	distribution	Distribution: — VIETNAM (Quang Nam, Quang Binh, Gia Lai, Kon Tum, and Dak Lak provinces). Thailand.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFB9FF079A2A950CF42A.taxon	materials_examined	Studied specimens: — THAILAND, wild collected plant originated from Thailand, cultivated in private garden of B. Q. Viet in southern Vietnam, AL s. n., 19 October 2017, L. Averyanov (LE 01087720 http: // en. herbariumle. ru /? t = occ & id = 42947 photos). VIETNAM, plant cultivated in Lam Dong province, Di Linh District, originated from Quang Nam Province, 16 June 2020, Truong Ba Vuong, Loc Bao Sang, BV 699 (VNM). VIETNAM, Gia Lai Province, Kon Chu Rang Nature Reserve, 1 June 2016, M. Nuraliev, Nuraliev 1618 (LE 01055514 http: // en. herbariumle. ru /? t = occ & id = 7269). VIETNAM, Quang Binh Province, Minh Hoa District, 29 July 2011, Nguyen Tien Hiep et al., CPC 3919 (LE 01057819 http: // en. herbariumle. ru /? t = occ & id = 8320). VIETNAM, Quang Binh Province, Minh Hoa District, 23 July 2011, Nguyen Tien Hiep et al., CPC 3657 (LE 01057820, http: // en. herbariumle. ru /? t = occ & id = 15154).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	materials_examined	Type: — VIETNAM, N Vietnam without indication of locality, ex Hort. Botanical Garden of the University of Vienna, Anton Sider 250 / 92, holotype: WU 0063106 https: // www. jacq. org / detail. php? ID = 267211 photo!; isotype AMES 00084913 not seen.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	description	Description: — Epiphytic creeping herb. Pseudobulbs 1 - leaved, pale green, narrowly ovoid, oblique, 2 – 2.5 cm tall, tapering to the apex, growing close together, connected by semi-woody rhizome, 2.3 – 2.5 mm in diameter, young pseudobulbs covered by brown fibrous sheaths. Leaves green, slightly twisted, petiolate; petiole 2.3 – 3.7 cm long; leaf blade narrowly oblong, 10 – 13 cm long, 1.8 – 2.2 cm wide, obtuse, slightly unequally 2 - lobed. Inflorescence subumbellate, arising from base of pseudobulbs, erect, slender, green, spotted with purple; peduncle ca. 14.5 cm long with 1 to 2 overlapping tubular bracts at base and 2 distant tubular bracts above; rachis 1.5 – 2 mm long, bears 4 – 5 flowers opening simultaneously; floral bracts 6 – 8 mm long; pedicel and ovary ca. 1.5 cm long, slightly curved. Flower pale yellow with many red dots, more dense on lateral sepals and less in median sepal, petals yellow, margin dark red, lip adaxially dark red mottled with white, adaxially with many red blotches. Median sepal narrowly ovate to ovate, concave, ca. 9 mm long, 4 mm wide, obtuse, margin shortly fimbriate; lateral sepals ca. 2.5 cm long, 4 mm wide (at base) apex cuspidate, upper and lower margin sparsely papillose, outer surface sparely proximally verrucose; twisted and free at base, upper and lower margin turning inward and connate from near base to apex, distally forming narrowing tube. Petals ovate, ca. 2.5 mm long, 2 mm wide, margin fimbriate, apex obtuse. Lip mobile, with white thin ligament connected with apex of column foot, simple, recurved, ligulate, ca. 5 mm long, ca. 1.7 mm wide, apex obtuse to rounded; margin densely papillose; at base adaxially grooved. Column ca. 2 mm tall, at front with triangular wings, column foot ca. 3 mm long, upcurved; stelidia slender, subulate, ca. 6 mm long, slightly upcurved, upper margin with truncate tooth, lower margin sometime finely erose; anther cap rectangular in outline, ca. 1.2 mm long, with umbo at apex; pollinia 4, obovate, ca. 0.8 mm long. Fruit not seen.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	biology_ecology	Habitat and phenology: — Evergreen broad-leaved forests at elevation about 1000 m a. s. l., rare. Branch epiphyte. Flowers at the end of May.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	distribution	Distribution: — Vietnam (Lai Chau Province). Endemic.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	discussion	Notes: — The specimens were found and collected in Sin Ho District, Lai Chau Province, and later were cultivated in private garden in Da Lat Town. Fimbriate median sepal, lateral sepals turning inwards with adnate upper and lower margins, shortly fimbriate free part of upper margin, fimbriate petals, and densely papillose lip are key characters of this species.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F103FFBAFF079D129538F31E.taxon	materials_examined	Studied specimen: — VIETNAM, herbarium specimens prepared on 30 May 2021 by Truong Ba Vuong, Dang Van Son, Nghiem Xuan Son, BV 1255 (VNM 00069914), BV 1256 (VNM 00069915) from plants cultivated in Lam Dong Province, Da Lat Town, originated from Lai Chau Province, Sin Ho District, forest around Sin Ho Town.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF079D3E9491F687.taxon	materials_examined	Type: — MYANMAR, Mawlamyine (Moulmein), Parish 346 (W-Herb. Reichenbach).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF079D3E9491F687.taxon	materials_examined	Type: — MALAY PENINSULA, “ sent alive to KEW by Mr. E. Seimund in 1914 and flowered shortly after ” August 1919. Seimund 337 - 14 (K 000891078 http: // specimens. kew. org / herbarium / K 000891078 photo!).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF079D3E9491F687.taxon	discussion	Notes: — In Vietnam, this species is presented by two varieties. The key for their identification is presented below.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF0798089576F099.taxon	biology_ecology	Habitat and phenology: — Primary broad-leaved forests. Trunk and branch epiphyte. Flowering in January – February.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF0798089576F099.taxon	distribution	Distribution: — Myanmar, China, Thailand, Laos, Vietnam (Hoa Binh Province), Borneo, Malaysia, Sumatra, Java.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB9FF0798089576F099.taxon	materials_examined	Studied specimens: — MALAYSIA, Rasak waud, March 1910, s. coll., 15203 as B. rupicolum (K 000891079 http: // specimens. kew. org / herbarium / K 000891079 photo!). VIETNAM, Hoa Binh Province, 29 March 2001, Nguyen Tien Hiep et al., HAL 299 (LE 01057944 http: // en. herbariumle. ru /? t = occ & id = 8447 & rid = image _ 0016710).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	materials_examined	Type: — VIETNAM, holotype specimen prepared on 29 April 2021 by Truong Ba Vuong, Phan Trong Quyet, BV 1160 (VNM 00069901, LE 01122221 http: // en. herbariumle. ru /? t = occ & id = 82694 photos) from cultivated plant obtained from a private garden in Da Lat Town, Lam Dong Province, originated from Ha Giang Province, Quan Ba District, Quyet Tien Commune, Lung Muoi Moutain.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	etymology	Etymology. The epithet refers to the denticulate lower margin of column wings.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	description	Description: — Epiphytic creeping herb. Pseudobulbs 1 - leaved, green, dark green near apex and chestnut at base, clustering, ovoid to narrowly ovoid, oblique or straight, 1.2 – 3.3 cm tall. Leaves petiolate; petiole 0.6 – 2 cm long, usually twisted at base; leaf blade adaxially green, abaxially green or occasionally purple, oblong to narrowly oblong, 5.5 – 14.5 cm long, 1.5 – 2.1 cm wide, obtuse, slightly retuse. Inflorescence umbellate to subumbellate raceme; peduncle glabrous, purple or green, spotted with purple, slender, arising from base of pseudobulbs, 8 – 15 cm long, with 1 to 3 distant tubular bracts; rachis 1 – 5 mm long, with 2 – 13 flowers opening simultaneously; floral bracts glabrous, 4 – 5.5 mm long; pedicel and ovary glabrous, 1 – 1.4 cm long, ovary with 6 prominent ridges. Flower brownish-yellow to brown, tepals adaxially with red streaks, abaxially with small red dots, lip red above, yellowish below. Median sepal ovate, concave, 7.5 – 9 mm long, 4 – 5 mm wide, with 5 veins, acute, margin sparsely ciliate; lateral sepals glabrous, narrowly ovate, slightly falcate, 16 – 18 mm long, 4 – 5 mm wide, concave, twisted at base, upper and lower margin free at base, turning inward and connate from near base to apex, forming a fusiform tube, tapering to acute apex. Petals narrowly ovate, oblique, with 3 veins, acute, margin fimbriate on apical half. Lip mobile, with white thin ligament connected with column foot apex, simple, slightly, recurved, ligulate, glabrous, ca. 4 mm long, 1.3 mm wide, apex obtuse, grooved at base, abaxially with 2 low keels. Column ca. 2 mm tall, at front with small triangular wing; column foot upcurved, ca. 2.5 mm long; stelidia subulate, acute, slightly falcate, upper margin with an erect acute tooth, lower margin denticulate; anther cap glabrous, ca. 1 mm long and wide, above with a yellow umbo; pollinia 4, ovoid. Fruit not seen.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	biology_ecology	Habitat and phenology: — Epiphyte on large trees at elevation about 1200 m a. s. l. Not common. Flowers in April – May.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	distribution	Distribution: — Vietnam (Bac Kan, Ha Giang, Hoa Binh, Lam Dong, and Ninh Binh provinces).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	discussion	Notes: — The new variety differs from B. taeniophyllum var. taeniophyllum mainly in denticulate column wings. Its morphological comparison with related taxa are presented in table. In addition, there are 3 different forms of this variety which differ in the shape of lateral sepals and the arrangements of flowers on rachis (Fig. 4. A-F).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	materials_examined	Studied specimens (paratypes): — VIETNAM, Bac Kan Province, Cho Don District, Ban Thi Municipality, Phia Khao Village, around point 22 º 17 ’ 31 ’’ N, 105 º 31 ’ 51 ’’ E, Cau Muc Mountain at elevation of around 800 – 900 m a. s. l., primary broad-leaved evergreen closed forest on very steep slopes and cliffs near tops of rocky ridge composed with solid marble-like stratified limestone, epiphyte, leaves below with light purple-violet tint, not common, 24 May 2004, L. Averyanov et al., HAL 4859 a (LE 01057942 http: // en. herbariumle. ru /? t = occ & id = 8445, LE 01087453 http: // en. herbariumle. ru /? t = occ & id = 38856 photos). VIETNAM, Bac Kan Province, Cho Don District, June 2017, C. X. Canh s. n. (LE 01087697 http: // en. herbariumle. ru /? t = occ & id = 42919 photos). VIETNAM, herbarium specimen is prepared on 30 April 2021 by Truong Ba Vuong, Phan Trong Quyet, BV 1156 (VNM 00069904, LE 01122220 http: // en. herbariumle. ru /? t = occ & id = 82693 photos) from plants cultivated in private garden in Da Lat Town, originated from Ha Giang Province, Quan Ba District, Quyet Tien Commune, forest around Tan Tien Village. VIETNAM, Hoa Binh Province, Lac Son District, Ngoc Son Municipality, Khu Village, Bua Coi Mt., around point 20 ° 27 ′ 10 ″ N 105 ° 18 ′ 52 ″ E, in heavily logged closed primary evergreen broad-leaved forest on top ridges of limestone mountains at elevation of around 700 – 790 m a. s. l., 29 March 2011, N. Q. Hieu et al., CPC 1734 a (LE 01057936 http: // en. herbariumle. ru /? t = occ & id = 8441, LE 01087455 http: // en. herbariumle. ru /? t = occ & id = 39090 photos). VIETNAM, herbarium specimen is prepared on 30 April 2021 by Truong Ba Vuong, Phan Trong Quyet, Ngo Quang Dang, BV 1155 (VNM 00069903, LE 01122219 http: // en. herbariumle. ru /? t = occ & id = 82692 photos) from plant cultivated in private garden in Da Lat Town, originated from Lai Chau Province, Sin Ho District. VIETNAM, Lam Dong Province, Da Lat Town, Hang Cop Stream, 12 May 2021, Truong Ba Vuong, Phan Trong Quyet, BV 1171 (VNM 00069902). VIETNAM, Ninh Binh Province, Cuc Phuong National Park, a. 1985, L. Averyanov, LX-VN s. n. (LE 01057945 http: // en. herbariumle. ru /? t = occ & id = 15158, LE 01087456 http: // en. herbariumle. ru /? t = occ & id = 39132 photos). VIETNAM, Ninh Binh Province, Cuc Phuong National Park, 2002, Averyanov s. n. (LE 01087381 http: // en. herbariumle. ru /? t = occ & id = 38701 photos).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	materials_examined	Type: — Lai Chau Province, Sin Ho District, forest around Sin Ho Town, 24 July 2021, Truong Ba Vuong, Truong Quang Tam, BV 1306 (holotype, VNM 00069927).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	description	Descriptions: — Epiphytic creeping herb. Pseudobulbs 1 - leaved, greenish, narrowly ovoid to fusiform, slightly oblique, 3.2 – 4.5 cm, tapering to the apex, covered by remnant fibrous sheaths, pseudobulbs well-spaced, connected by semiwoody rhizome, 2.5 – 7 cm long, 4 mm in diameter, roots from base of pseudobulbs or from nodes of rhizome. Leaves green, petiolate; petiole 2.5 – 3 cm long; leaf blade narrowly oblong, 10 – 11 cm long, 2.5 – 3 cm wide, apex obtuse or retuse. Inflorescence subumbellate, arising from base of pseudobulbs, slender, green with purplish marks; peduncle 20 – 21 cm long; rachis 1 – 1.5 mm long, with 5 – 6 flowers opening simultaneously; floral bract greenish to white with pinkish dots, triangular, 6 – 7 mm long, acute; pedicel and ovary 9 – 10 mm long, pedicel slightly upcurved, green, ovary green with pink dots. Median sepal white or pinkish with pink stripes, lateral sepals yellowish or white, pale green at apical part, with pink stripes, petals whitish to pink, with pink stripes, lip white or dark pink. Median sepal broadly ovate, concave, 6 – 6.6 mm long, 4.5 – 5 mm wide, margin finely erose or dentate, truncate and mucronate. Lateral sepals 27 – 29 mm long, 5 – 6 mm wide, cuspidate, twisted at base, upper margin connate throughout all their length, free at base and along lower margin. Petals narrowly ovate, oblique, falcate 4 – 5 mm long, 2.5 – 3 mm wide, ciliate along the margin at basal half, at apex hairy, apex acute with short seta. Lip glabrous, entire, narrowly ovate, simple, curved, 3.5 – 4 mm long, 2 – 2.5 mm wide, apex rounded, at base adnate to column foot apex by white appendage; adaxially shallowly channeled at base; abaxially with 2 low keels. Column ca. 2 mm long, with deltoid or rounded wings, stigma obovate, ca. 1.5 mm long; column foot ca. 3.5 mm long; stelidia ca. 0.5 mm long, subulate, acute, pointing forward, upper margin with small triangular tooth; anther cap helmet-shaped, ca. 1 mm long, frontal margin ciliate; pollinia 4, obovate. Fruit not seen.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	biology_ecology	Habitat and phenology: — Broad-leaved forest. Trunk or branch epiphyte, commonly along streams. Flowering in July.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	distribution	Distribution: — Vietnam (Lai Chau Province, Sin Ho District). Endemic.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	discussion	Notes: — The morphological comparison with related species is present on the table.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	materials_examined	Type: — INDIA, Khasia, Nungclow, Gibson 31 (K-LINDL K 001368947 http: // specimens. kew. org / herbarium / K 001368947 photo!).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	biology_ecology	Habitat and phenology: — Primary humid evergreen broad-leaved forests at elevations 600 – 2000 m a. s. l. Trunk epiphyte. Flowering in August – December.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	distribution	Distribution: — India, Myanmar, China, Thailand, Laos, Vietnam (doubtful).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	discussion	Notes: — The early records of B. helenae by Averyanov et al. 2015 is actually B. nodosum, so occurrences of this species in Vietnam is doubtful.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F100FFB8FF0799A09558F036.taxon	materials_examined	Studied specimens: — INDIA, Mount Khasia, Hooker & Thomsom, s. n. as C. cornutum (M 0226978 http: // plants. jstor. org / stable / 10.5555 / al. ap. specimen. m 0226978 photo!). INDIA, Sikkim; Sikkim Himalaya, June 1898, R. Pantling, 232 (L. 1484515 https: // data. biodiversitydata. nl / naturalis / specimen / L. 1484515 photo!). INDIA, Sikkim Himalaya, 1892, R. Pantling 232 (MNHN-P-P 00348046 http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00348046 photo!). INDIA, Khasia J. D. Hooker & T. Thomson 6 (MNHN-P-P 00348049 http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00348049 photo!). MYANMAR, Hills around Ku-lu, May 1925, G. Forrest 26609 (E 00123412, https: // data. rbge. org. uk / herb / E 00123412 photo!). THAILAND, Kampaeng Phet, Klong Lan, 1240 m, 29 October 2002, M. van de Bult 601 (L. 1484514, https: // data. biodiversitydata. nl / naturalis / specimen / L. 1484514 photo!). THAILAND, Chiang Rai, Mae Sai, Doi Dtung (Tung), 28 September 2006, J. F. Maxwell 06 - 683 (L. 4175145 https: // data. biodiversitydata. nl / naturalis / specimen / L. 4175145 photo!).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F101FFBFFF0799029437F463.taxon	materials_examined	Type: — Lai Chau Province, Sin Ho District, forest around Sin Ho Town, 24 July 2021, Truong Ba Vuong, Truong Quang Tam, BV 1306 (holotype, VNM 00069927).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F101FFBFFF0799029437F463.taxon	description	Descriptions: — Epiphytic creeping herb. Pseudobulbs 1 - leaved, greenish, narrowly ovoid to fusiform, slightly oblique, 3.2 – 4.5 cm, tapering to the apex, covered by remnant fibrous sheaths, pseudobulbs well-spaced, connected by semiwoody rhizome, 2.5 – 7 cm long, 4 mm in diameter, roots from base of pseudobulbs or from nodes of rhizome. Leaves green, petiolate; petiole 2.5 – 3 cm long; leaf blade narrowly oblong, 10 – 11 cm long, 2.5 – 3 cm wide, apex obtuse or retuse. Inflorescence subumbellate, arising from base of pseudobulbs, slender, green with purplish marks; peduncle 20 – 21 cm long; rachis 1 – 1.5 mm long, with 5 – 6 flowers opening simultaneously; floral bract greenish to white with pinkish dots, triangular, 6 – 7 mm long, acute; pedicel and ovary 9 – 10 mm long, pedicel slightly upcurved, green, ovary green with pink dots. Median sepal white or pinkish with pink stripes, lateral sepals yellowish or white, pale green at apical part, with pink stripes, petals whitish to pink, with pink stripes, lip white or dark pink. Median sepal broadly ovate, concave, 6 – 6.6 mm long, 4.5 – 5 mm wide, margin finely erose or dentate, truncate and mucronate. Lateral sepals 27 – 29 mm long, 5 – 6 mm wide, cuspidate, twisted at base, upper margin connate throughout all their length, free at base and along lower margin. Petals narrowly ovate, oblique, falcate 4 – 5 mm long, 2.5 – 3 mm wide, ciliate along the margin at basal half, at apex hairy, apex acute with short seta. Lip glabrous, entire, narrowly ovate, simple, curved, 3.5 – 4 mm long, 2 – 2.5 mm wide, apex rounded, at base adnate to column foot apex by white appendage; adaxially shallowly channeled at base; abaxially with 2 low keels. Column ca. 2 mm long, with deltoid or rounded wings, stigma obovate, ca. 1.5 mm long; column foot ca. 3.5 mm long; stelidia ca. 0.5 mm long, subulate, acute, pointing forward, upper margin with small triangular tooth; anther cap helmet-shaped, ca. 1 mm long, frontal margin ciliate; pollinia 4, obovate. Fruit not seen.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F101FFBFFF0799029437F463.taxon	biology_ecology	Habitat and phenology: — Broad-leaved forest. Trunk or branch epiphyte, commonly along streams. Flowering in July.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F101FFBFFF0799029437F463.taxon	distribution	Distribution: — Vietnam (Lai Chau Province, Sin Ho District). Endemic.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F101FFBFFF0799029437F463.taxon	discussion	Notes: — The morphological comparison with related species is present on the table.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F106FFBFFF079DF69207F16E.taxon	materials_examined	Type: — INDIA, Khasia, Nungclow, Gibson 31 (K-LINDL K 001368947 http: // specimens. kew. org / herbarium / K 001368947 photo!).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F106FFBFFF079DF69207F16E.taxon	biology_ecology	Habitat and phenology: — Primary humid evergreen broad-leaved forests at elevations 600 – 2000 m a. s. l. Trunk epiphyte. Flowering in August – December.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F106FFBFFF079DF69207F16E.taxon	distribution	Distribution: — India, Myanmar, China, Thailand, Laos, Vietnam (doubtful).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F106FFBFFF079DF69207F16E.taxon	discussion	Notes: — The early records of B. helenae by Averyanov et al. 2015 is actually B. nodosum, so occurrences of this species in Vietnam is doubtful.	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
E22D1205F106FFBFFF079DF69207F16E.taxon	materials_examined	Studied specimens: — INDIA, Mount Khasia, Hooker & Thomsom, s. n. as C. cornutum (M 0226978 http: // plants. jstor. org / stable / 10.5555 / al. ap. specimen. m 0226978 photo!). INDIA, Sikkim; Sikkim Himalaya, June 1898, R. Pantling, 232 (L. 1484515 https: // data. biodiversitydata. nl / naturalis / specimen / L. 1484515 photo!). INDIA, Sikkim Himalaya, 1892, R. Pantling 232 (MNHN-P-P 00348046 http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00348046 photo!). INDIA, Khasia J. D. Hooker & T. Thomson 6 (MNHN-P-P 00348049 http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00348049 photo!). MYANMAR, Hills around Ku-lu, May 1925, G. Forrest 26609 (E 00123412, https: // data. rbge. org. uk / herb / E 00123412 photo!). THAILAND, Kampaeng Phet, Klong Lan, 1240 m, 29 October 2002, M. van de Bult 601 (L. 1484514, https: // data. biodiversitydata. nl / naturalis / specimen / L. 1484514 photo!). THAILAND, Chiang Rai, Mae Sai, Doi Dtung (Tung), 28 September 2006, J. F. Maxwell 06 - 683 (L. 4175145 https: // data. biodiversitydata. nl / naturalis / specimen / L. 4175145 photo!).	en	Dang, Minh Quan, Averyanov, Leonid V., Dang, Van Son, Maisak, Tatiana, Bui, Van Huong, Tu, Bao Ngan, Truong, Ba Vuong (2022): Bulbophyllum section Rhytionanthos (Orchidaceae) in Vietnam with description of new taxa and new national record. Phytotaxa 530 (3): 257-270, DOI: 10.11646/phytotaxa.530.3.1
