taxonID	type	description	language	source
56346EECF6D6501FB765FF4FEED0330F.taxon	description	Figs 12 and 14	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
56346EECF6D6501FB765FF4FEED0330F.taxon	materials_examined	Additional material. ANGOLA • 2 ♂, subadult; same collecting locality as holotype; Dec. 2020; MNCN 50544 and FKH 0499 • 1 ♀; same collecting locality as previous; FKH 0500.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
56346EECF6D6501FB765FF4FEED0330F.taxon	diagnosis	Diagnosis. A medium sized Hemidactylus, with SVL of 43.2 mm (mean) and maximum width of 8.8 mm (Fig. 14). 9 - 11 supralabials and 8 - 10 infralabials. Dorsal pholidosis with 15 - 22 rows of moderate strongly keeled tubercle scales and ventral pholidosis with 32 - 38 smooth and rounded scale rows around midbody. Hemidactylus carivoensis sp. nov. present a moderate, triangular, moderate rounded posteriorly mental scale, two large postmentals followed by two large post-postmentals. Tail with four strongly keeled dorsal tubercles rows dorsally and subcaudal scales small, about one fourth of the tail width, interspersed by large series of horizontal whorls of keeled scales. Males with 15 - 22 continuous precloacal-femoral pores. Five divided scansors beneath first digit of both manus and pes, seven or eight beneath fourth digit of manus, nine beneath the fourth digit of pes. Dorsum darker coloration on the dorsal part of the head that continues along the medial part of the dorsum until the sacrum.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
56346EECF6D6501FB765FF4FEED0330F.taxon	description	Holotype description (Fig. 14). Measurements and meristic characters of the holotype are presented in Table S 7. Adult female with a snout-vent-length (SVL) of 45.16 mm and a tail length (TL) about the same size than SVL 44.79 mm. Body slender, nape distinct. Head slightly narrower than the body and largely elongated (HW / HL 0.65). Canthus rostralis not prominent. Eye diameter (3.04 mm), with vertical pupil and crenulated margin. Supraciliar scales small and rounded. Ear height (0.83 mm). Ear to eye distance slightly larger than orbit diameter (3.46 mm). Snout rounded and pointed. Frontal scales granular and larger than occipital scales. Occipital scales granular interspersed with large number of smooth and conical tubercle scales from eyes to nape. Rostral wider than deep (2.31 vs. 0.93 mm, respectively). Rostral semi-divided posterodorsally, in contact with 1 st supralabial, nostril, two postnasals and one internasal scales. Eleven supralabials and 9 infralabials. First supralabial in contact with the nostril. Nostril circular rounded by rostral, supranasal and two postnasals. Postnasals larger than supranasal. One or two rows of scales between supralabials and the orbit. Mental large, triangular and rounded posteriorly, with two large rectangular postmental scales in broad contact posteriorly to the mental. 4 post-postmental scales, composed by post-postmental slightly smaller than postmental scales in contact with postmentals and 1 st and 2 nd infralabials, and 2 post-postmental half size than lateral post-postmental in contact with postmental scales. Gular scales slightly smaller than ventral scales and granular. Between the gular scales and infralabials a row of enlarged scales is present, decreasing in size until the 5 th infralabial where they become the same size as the gular scales. Body relatively robust and slightly elongated (TRL / SVL 0.39). Ventral scales widely larger than dorsal scales, with 36 scales across the belly. The dorsal pholidosis present heterogenous conical, granular scales interspersed by 17 strongly keeled dorsal tubercle rows of at midbody. Dorsal tubercle rows are separated by 3 granular scales. Tubercle scales reach the posterior part of the eye region where tubercle scales lose the keeling progressively. Tail with four strongly keeled dorsal tubercles rows dorsally and subcaudal scales small, about one fourth of the tail width, interspersed by 22 horizontal whorls of keeled scales. Precloacal scales enlarged and one well-developed postcloacal spur on each side. Fore- and hindlimbs relatively short, stout; forearm short (FL / SVL 0.16); tibia short (CL / SVL 0.18). Short digits and clawed. All digits of manus and pes indistinctly webbed. Scansors beneath each toe composed by 1 st and two terminal scansor undivided. 4 th toes with 3 undivided terminal scansors. Scansors beneath each finger equally divided, with the exception of 1 st and last terminal scansors undivided. Number of scansors: 6 - 6 - 7 - 7 - 7 (right manus), 7 - 8 - 9 - 10 - 7 (right pes). Relative length of digits: V <IV = III> II> I (right manus); V <IV <III> II> I (right pes).	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
56346EECF6D6501FB765FF4FEED0330F.taxon	etymology	Etymology. The species epithet " Hemidactylus carivoensis " refers to the Farm Carivo, an old estate situated along the banks of the mid-lower Coporolo River on the coastal plain of Benguela Province, and where most of the type series was collected. The species proved to be common in the area, and by recognizing the farm, we also acknowledge the ongoing support from the owners to researchers, similar to the Chapmans nearly a century ago.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
56346EECF6D6501FB765FF4FEED0330F.taxon	distribution	Distribution and conservation (Fig. 12 C). The full distribution range of the species remains unknown, even though so far it has only been confirmed to occur across the coastal plain of western Benguela Province. Due to the variable morphology of the taxon known as H. benguellensis and which formerly included material now assigned to H. cinganji sp. nov., it is possible that some historical material was wrongly identified and should instead belong to this newly described species. Spiny savannas are well distributed in western Angola, suggesting that the new species might be common across a wider range than what is currently known, but available data is still poor. However, due to limited material confirmed to belong to this species, we can't calculate the EOO and we regard the conservation status of this species as Data Deficient. Therefore, it is important for future studies to establish the real distribution of H. benguellensis, H. cinganji sp. nov. and H. carivoensis sp. nov. to better determine the conservation status of the species.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
50ECBDF971C65217BB40601239A95CF2.taxon	description	Figs 12, 13	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
50ECBDF971C65217BB40601239A95CF2.taxon	diagnosis	Diagnosis. A large sized Hemidactylus, with maximum SVL of 55 mm (mean) and maximum width of 10.5 mm (Fig. 13). Nine supralabials and 9 - 11 infralabials. Dorsal pholidosis with 13 - 18 rows of moderate strongly keeled tubercle scales and ventral pholidosis with 33 - 41 smooth and rounded scale rows on midbody. Hemidactylus cinganji sp. nov. present a moderate, triangular mental scale, two large postmentals followed by two medium post-postmentals. Tail with six strongly keeled dorsal tubercles rows dorsally and subcaudal scales medium sized, interspersed by 11 horizontal whorls of keeled scales on the original portion of the tail. Males with 26 - 28 continuous precloacal-femoral pores and 1 - 4 postcloacal tail spurs. Six divided scansors beneath first digit of both manus and pes, seven or eight beneath fourth digit of manus, eight or nine beneath the fourth digit of pes. Dorsum presents dark orange-beige coloration across the body with darker coloration on the dorsal part of the head that continues along the medial part of the dorsum until the sacrum.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
50ECBDF971C65217BB40601239A95CF2.taxon	description	Holotype description (Fig. 13). Measurements and meristic characters of the holotype are presented in Table S 6. Adult male with a snout-vent-length (SVL) of 45.05 mm and a tail length (TL) slightly smaller than SVL 42.07 mm. Body slender, nape distinct. Head slightly narrower than the body and moderate head length (HW / HL 0.72). Canthus rostralis not prominent. Eye diameter (3.05 mm), with vertical pupil and crenulated margin. Supraciliar scales small and rounded. Ear height (1.47 mm). Ear to eye distance slightly larger than orbit diameter (4.17 mm). Snout rounded and pointed. Frontal scales granular and larger than occipital scales. Occipital scales granular interspersed with large number of smooth and keeled tubercle scales from eyes to nape. Rostral undivided, in contact with 1 st supralabial, nostril, supranasal and one internasal scales. 9 supralabial and 8 - 9 infralabials. First supralabial and rostral in direct contact with the nostril. Nostril circular rounded by rostral, 1 st supralabial, supranasal, and two postnasals. One row of scales between supralabials and the orbit. Mental large, triangular and rounded posteriorly, with two larges rectangular postmental scales in broad contact posteriorly to the mental. 2 post-postmental scales, in contact with post-postmental slightly smaller than postmental scales and 1 st and 2 nd infralabials. Gular scales slightly smaller than ventral scales and granular. Body relatively robust and slightly elongated (TRL / SVL 0.40). Ventral scales widely larger than dorsal scales, with 32 scales across the belly. The dorsal pholidosis present heterogenous conical, granular scales interspersed by 15 strongly keeled dorsal tubercle rows of at midbody. Dorsal tubercle rows are separated by 3 granular scales. Tubercle scales reach the posterior part of the eye region where tubercle scales remain keeled. Tail with six strongly keeled dorsal tubercles rows dorsally and subcaudal scales medium sized, interspersed by 11 horizontal whorls of keeled scales on the original portion of the tail. Regenerated portion of the tail presents homogeneous scales all around the surface without tubercle scales. 27 precloacal-femoral pores enlarged and 2 well-developed postcloacal spurs on each side. Fore- and hindlimbs relatively short, stout; forearm short (FL / SVL 0.15); tibia short (CL / SVL 0.16). Short digits strongly clawed. All digits of manus and pes indistinctly webbed. Scansors beneath each toe composed by 1 st and terminal scansor undivided with the exception of 4 th and 5 th toes with 3 and 2 undivided terminal scansor, respectively. Scansors beneath each finger equally divided, with exception of 1 st and last terminal scansor undivided. Number of scansors: 5 - 5 - 6 - 7 - 6 (right manus), 6 - 8 - 8 - 9 - 7 (right pes). Relative length of digits: V <IV = III> II> I (right manus); V <IV <III> II> I (right pes).	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
50ECBDF971C65217BB40601239A95CF2.taxon	etymology	Etymology. The name " Hemidactylus cinganji " is a widespread traditional word used in Angolan local languages that represents an ancestral spiritual entity that reincarnates assuming different physical forms in different places and occasions. This name is suitable as the new species corresponds to a taxon that was first described under a different name, then became lost and now resurfaces after its original name had been hijacked by a surrogate sister-species. The species epithet is used as a neuter singular noun in opposition to the generic name.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
50ECBDF971C65217BB40601239A95CF2.taxon	distribution	Distribution and conservation (Fig. 12 B). The full distribution range of the species remains unknown, even though so far it has only been confirmed to occur in the interior of Benguela Province, from Cahata to Passe and Ebanga, above 800 m a. s. l. Due to poor sampling and few material confirmed to belong to this species, we cannot calculate the EOO and thus we regard the conservation status of the species as Data Deficient. Additional data will need to be collected to address its conservation status.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	description	Fig. 15, 16	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	materials_examined	Additional material examined. ANGOLA • 1 ♀; same locality as type material; 11 Aug. 2018; Pedro and Afonso Vaz Pinto; ZMB 90446 • 1 ♂, juv.; same collecting details as previous material; ZMB 90445 • ♀; same locality as type material; 15 Oct. 2020; Pedro Vaz Pinto; MNCN 50536.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	diagnosis	Diagnosis. A robust medium sized Hemidactylus, with SVL of 39.4 mm (mean) and maximum width of 7.4 mm (Fig. 15). 8 - 9 supralabials and 7 - 8 infralabials. Dorsal pholidosis with 15 - 17 rows of moderate keeled tubercle scales and ventral pholidosis with 29 - 32 smooth and rounded scale rows around midbody. Hemidactylus faustus sp. nov. present a moderate, triangular mental scale, two large postmentals followed by two large post-postmentals. Tail with thickness at the base tail with conical tubercle rows laterally. Regenerated tail with regular larger scales. Males with 17 - 19 continuous precloacal-femoral pores. Five or six divided scansors beneath the first digit of both manus and pes, seven beneath the fourth digit of manus, eight or nine beneath the fourth digit of pes. Dorsum coloration with two darker dorsolateral bands from the occiput to the tail, which includes 5 W-shaped darker crossbands in contact with both lateral bands; each dark crossbar is separated by lighter blotches.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	description	Holotype description (Fig. 15). Measurements and meristic characters of the holotype are presented in Table S 8. Adult female with a snout-vent-length (SVL) of 39.39 mm and regenerated tail length (TL) of 22.84 mm. Robust body, nape slightly distinct. Head slightly wider than the body and shorten (HW / HL 0.67). Canthus rostralis not prominent, but slightly marked. Eye diameter (3.05 mm), with vertical pupil and crenulated margin. Supraciliar scales small and pointed. Ear height (0.98 mm). Ear to eye distance slightly larger than orbit diameter (3.17 mm). Snout rounded. Frontal scales granular and of similar size as occipital scales. Occipital scales granular interspersed with large number of smooth and conical tubercle scales from eyes to nape. Rostral wider than deep (1.66 vs. 0.91 mm, respectively). Rostral semi divided posterodorsally, in contact with 1 st supralabial, nostril, two postnasal and one internasal scales. 9 supralabial and 7 infralabials. First supralabial in contact with the nostril. Nostril circular rounded by rostral, supranasal, prenasal and 2 postnasals. Prenasal, postnasal and supranasal same size. One row of scales between supralabials and the orbit. Mental large, triangular, with two larges rectangular postmental scales in short contact posteriorly to the mental. 7 post-postmental scales, composed by 2 post-postmental half size of postmental scales in contact with postmentals and 1 st and 2 nd infralabials, and 5 small post-postmentals in contact with postmental scales. Gular scales half size than ventral scales and granular. Between the gular scales and infralabials a row of enlarged scales is present, decreasing in size towards the 5 th infralabial where they become the same size as the gular scales. Body robust and slightly short (TRL / SVL 0.41). Ventral scales widely larger than dorsal scales, with 31 scales across the belly. The dorsal pholidosis presents heterogenous conical, granular scales interspersed by 16 conical dorsal tubercle rows at midbody. Dorsal tubercle rows are separated by 3 granular scales. Tubercle scales reach the posterior part of the eye region where they lose the keeling progressively. Tubercle in the base of tail is well developed. Tail with lateral conical tubercle rows. Regenerated tail with regular larger scales (after precloacal) enlarged and 2 well-developed postcloacal spurs on each side. Fore- and hindlimbs relatively short, stout; forearm medium sized (FL / SVL 0.23); tibia short (CL / SVL 0.18). Digits short and clawed. All digits of manus and pes indistinctly webbed. Scansors beneath toes and fingers are equally divided and composed by 1 st scansor undivided and variable number of undivided terminal scansors. Number of scansors: 6 - 8 - 7 - 7 - 7 (right manus), 7 - 10 - 11 - 10 - 9 (right pes). Relative length of digits: V <IV <III> II> I (right manus); V <IV <III> II> I (right pes).	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	etymology	Etymology. The name " Hemidactylus faustus " applies to a Latin word that designate ' good luck', evoking the serendipitous discovery of this species. The species epithet is used as a masculine adjective singular. The first specimen was found by Beatriz Vaz Pinto, daughter of PVP, under a small stone which was removed while preparing a campsite. This unexpected find led to further collecting of this new and previously unrecorded form, albeit from a locality that had been regularly surveyed since the mid- 19 th century.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
2B41AE2BFC545F8F83A46D9669CBC588.taxon	distribution	Distribution and conservation (Fig. 16 C). This species is likely a micro-endemic form, strictly associated with the conglomerate inselbergs of Pungo Andongo also known as Pedras Negras (Black Rocks), just north of the mid-Cuanza River in western Malanje Province, Angola (Fig. 16 C). At the moment the only know population occurs on this site, which covers approximately 4,000 ha of huge rocky conglomerate boulders. A similar and nearby inselberg system - Pedras Jingas, albeit smaller, shares identical geological features as Pungo Andongo, being situated some 20 kms to the northeast. It is quite possible that the species is present at Pedras Jingas and also in a few smaller isolated inselbergs nearby, but these areas have not been surveyed yet. All considered, it is likely that H. faustus sp. nov. is contained within about 6,000 ha of suitable habitat in the region. However, due to the limited material confirmed to belong to this species, we cannot calculate the EOO and we regard the conservation status of this species as Data Deficient. This species needs further studies about the real extent of its range and current population trends to better address its conservation status. Due to its small distribution range and highly specialized niche this species may provisionally warrant a threat status.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
291010619C495F4484D73B52A6F19CF7.taxon	description	Figs 8, 9	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
291010619C495F4484D73B52A6F19CF7.taxon	diagnosis	Diagnosis. A medium sized Hemidactylus, with SVL of 45.53 mm (mean) with moderate long snout, 10 supralabials and 8 - 10 infralabials (Fig. 8). Dorsal pholidosis with 11 - 12 rows of moderate dorsal keeled tubercle scales and ventral pholidosis with 28 - 30 smooth scale rows on midbody. The species present a large, triangular mental scale, two large postmentals followed by two enlarged post-postmentals. Base of the tail with four large keeled dorsal tubercle rows and subcaudal scales small, about one fourth of the tail width. Males with 8 continuous precloacal pores. Five divided scansors beneath first digit of both manus and pes, seven beneath fourth digit of manus, seven or eight beneath the fourth digit of pes. Dorsum presents dark coloration with two light brown crossbands from the posterior part of the eye to the sacrum, where the two bands meet each other to form a V-shaped marking.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
291010619C495F4484D73B52A6F19CF7.taxon	description	Holotype description. Measurements and meristic characters of the holotype are presented in Table S 5. Adult female with a snout-vent-length (SVL) of 49.09 mm, a regenerated tail length (TL) of 32.93 mm. Body slender, nape distinct. Head slightly narrower than the body and largely elongated (HW / HL 0.62). Canthus rostralis not prominent, but well-marked. Eye diameter (2.96 mm), with vertical pupil and crenulated margin. Supraciliar scales small and slightly pointed. Ear height (0.93 mm). Ear to eye distance slightly larger than orbit diameter (3.6 mm). Snout rounded. Frontal scales granular and larger than occipital scales. Occipital scales granular with lateral conical a large tubercle scale. Rostral wider than deep (2.21 vs. 1.04 mm, respectively). Rostral semidivided anterodorsally, in contact with 1 st supralabial, nostril, prenasal and one internasal scales. 11 supralabials and 11 infralabials. First supralabial in contact with the nostril. Nostril circular rounded by rostral, supranasal, two postnasal and first supralabial. Postnasals larger than supranasal. Nostril in direct contact with the rostral and 1 st supralabial. One row of scales between supralabials and the orbit. Mental large and markedly triangular, with two large rectangular postmental scales in broad contact posteriorly to the mental. 5 post-postmental scales, composed by post-postmental slightly smaller than postmental scales in contact with postmentals, 1 st and 2 nd infralabials, and 3 small post-postmental in contact with postmental scales. Gular scales and granular smaller than ventral scales. Between the gular scales and infralabials, a row of enlarged scales is present, decreasing in size until the 5 th infralabial where they become the same size as the gular scales. Body relatively slender and elongated (TRL / SVL 0.45). Ventral scales about double size than dorsal scales, with 28 scales across the belly. The dorsal pholidosis present heterogenous conical, granular scales interspersed by 12 keeled dorsal tubercle rows at midbody. Dorsal tubercle rows are separated by 4 - 5 granular scales. Tubercle scales reach the posterior part of head and the nape, where tubercle scales lose the keeling progressively. Base of the tail with four large keeled dorsal tubercle rows dorsally and subcaudal scales small, about one fourth of the tail width. Regenerated tail has no presence of tubercle scales, having largely homogeneous scales along and across the tail. Precloacal scales enlarged and one well-developed postcloacal spurs on each side. Fore- and hindlimbs relatively short, stout; forearm short (FL / SVL 0.17); tibia short (CL / SVL 0.18). Short digits and clawed. All digits of manus and pes indistinctly webbed. Scansors beneath each toe equally divided, with the exception of 1 st and terminal scansor undivided. 4 th and 5 th toes with 2 and 3 undivided terminal scansors, respectively. Scansors beneath each finger equally divided, with the exception of 1 st and two terminal scansor undivided. 1 st and 5 th fingers with 3 undivided terminal scansors. Number of scansors: 5 - 7 - 7 - 8 - 7 (right manus), 5 - 7 - 8 - 8 - 9 (right pes). Relative length of digits: V <IV = III = II> I (right manus); V <IV = III> II> I (right pes).	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
291010619C495F4484D73B52A6F19CF7.taxon	etymology	Etymology. The name " Hemidactylus pfindaensis " derives from the local word " pfinda " which in Kikongo - the main language used in Uige Province and northwestern Angola - refers to a " gallery forest " or a " continuous block of thick forest ", the main habitat type associated with the species.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
291010619C495F4484D73B52A6F19CF7.taxon	distribution	Distribution and conservation (Fig. 9 C). A typical forest gecko, this species has been found in two sites of northern Angola, both in Uige Province, and on a third site in the enclave of Cabinda. Its known presence north and south of the Congo River, suggests a much larger distribution range, that will likely extend to Democratic Republic of the Congo and Republic of the Congo, and possibly also into Gabon. In Angola, it will also likely be present in the Mayombe Forest and Northeastern Forest-Savanna Mosaic. However, due to limited number of records we cannot calculate the EOO and thus we regard the conservation status of the species as Data Deficient, and further studies are suggested to better assess its full distribution and conservation status. Although central African forests are currently threatened by deforestation and human encroachment, the occurrence of this species across a large geographical and altitudinal range, suggests that it is likely common, yet further studies are necessary to evaluate its conservation status.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
BFF4817A5F445915BE9D4DB42D6A31BF.taxon	description	Figs 6, 7	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
BFF4817A5F445915BE9D4DB42D6A31BF.taxon	materials_examined	Material examined. ANGOLA • 2 ♀; Namibe Prov., Santa Marta, Lucira; - 13.87861 °, 12.42444 °; 48 m a. s. l.; 24 May 2019; Pedro Vaz Pinto; FKH 0226 - 7 and MNCN 50538 • 3 ♂; same collecting data as previous material; FKH 0228, MNCN 50539, FKH 0232 • 4 ♀, 1 ♂ same collecting data as previous material; 6 Jul. 2019; Pedro Vaz Pinto and Javier Lobon-Rovira; FKH 0263 - 6, MNCN 50533 • 1 ♀ juv.; same collecting information as previous material; FKH 0268 • 2 ♀; Bentiaba; - 14.17596 °, 12.44689 °; 150 m a. s. l.; 15 Feb. 2020; Pedro Vaz Pinto and Javier Lobon-Rovira; MNCN 50541 and ZMB 90448 • 2 ♂; same collecting information as previous material; FKH 0417 and ZMB 90449 • 3 ♂; Lucira; September 1956; Charles Koch; TM 24447, 24450 - 1 • 1 ♂ and 1 ♀; Benguela Prov., Hanha; - 12.24245 °, 13.71399 °; 39 m a. s. l.; 16 Nov. 2019; Pedro Vaz Pinto; ZMB 90450 - 1.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
BFF4817A5F445915BE9D4DB42D6A31BF.taxon	distribution	Distribution and conservation (Fig. 6 C). This species is considered the sister species of H. bayonii, so far only known to occur along the arid coast from Bentiaba in northern Namibe Province, northwards to Hanha in coastal Benguela Province (Fig. 6). Despite being still poorly known, the species is probably widely distributed, and its habitat does not appear at present to be threatened; however, due to limited number of records we cannot calculate the extent of occurrence (EOO) and thus we regard the conservation status of the species as Data Deficient. Further studies are suggested to better assess its full distribution and conservation status.	en	Lobon-Rovira, Javier, Conradie, Werner, Iglesias, David Buckley, Ernst, Raffael, Verissimo, Luis, Baptista, Ninda, Pinto, Pedro Vaz (2021): Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501, DOI: http://dx.doi.org/10.3897/vz.71.e64781, URL: http://dx.doi.org/10.3897/vz.71.e64781
