taxonID	type	description	language	source
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	description	Figs 1, 2, 3, 4, 5, 6, 7, 8	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	diagnosis	Diagnosis. (1) Skin of dorsum shagreen, occipital fold evident, dorsolateral folds low in banded individuals or weakly defined in uniform or irregular dorsal colour patterns, skin on venter coarsely areolate (Fig. 2); (2) tympanic annulus and tympanic membrane present, tympanum prominent, sexually dimorphic, tympanum in males 7 % SVL, tympanum in females 6 % SVL; (3) snout subacuminate in dorsal view (tip of snout pointed), rounded in lateral view (Fig. 2); (4) upper eyelid with several small rounded to subconical tubercles; upper eyelid in males 56 % IOD (in females 59 %) IOD; cranial crests absent; (5) odontophore processes of the vomer partially concealed, oblique in outline, each processes bearing several small, poorly defined teeth; (6) sphenethmoides with acuminated anterior border dorsally, ventrally present an anteriorly rounded acuminated projected border, posterior border horizontal articulated with frontoparietals; posterior border of frontoparietal present enlarged quadrangular process posteriorly in dorsal view; zygomatic ramus of squamosal short and rounded in dorsal view; procesus cultriform of the parasphenoides reaching posterior level of the vomers, acuminated anterior border (Fig. 8); (7) males with vocal slits and subgular vocal sac; lacking nuptial pads; (8) finger I shorter than II, discs on fingers II-IV rounded (Fig. 3); (9) fingers with slightly visible lateral fringes; (10) ulnar tubercles small, low; (11) heel with small tubercles; outer edge of tarsus with small tubercles, more accentuated in males; (12) inner metatarsal tubercle oval elongated, about twice the size of outer metatarsal tubercle that is rounded; (13) toes lacking lateral fringes, webbing absent; Toe V slightly longer than III, discs slightly expanded (Fig. 3); (14) dorsum brown in several shades, with dark extreme yellow or green tones, females generally darker than males that have usually a banded pattern; banded limbs; venter light brown with dark brown flecks; hidden surfaces of the groin, thighs and armpits pinkish to reddish (in life), chest and throat present light brown tones (Figs 4 - 7); (15) SVL in males, 14.9 - 16.4 mm (mean = 13.8, SD = 3.9, n = 32), in females, 13.5 - 19.7 mm (mean = 17.1, SD = 4.41, n = 18); (16) advertisement call with two notes, duration note of 236 - 293 ms, with intervals of 522 - 620 ms; call is comprised by tonal sounds of constant frequency, with a slight upward modulation at end of call; (17) dominant frequency at 2.93 kHz, with two partial harmonics, first one with a range of 5.86 - 5.94 kHz and the second one with a range of 8.79 - 8.96 kHz; (18) call duration ranging from 1052 - 1136 ms.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	description	Description of the holotype (Figs 2, 3, 4). Adult female (DHMECN 14701) with robust body and short limbs; head slightly longer than wide, not as wide as body, head width 40 % of SVL (15.4 mm); head length 41 % of SVL; snout short, with a fleshly terminal rounded tip, subacuminated in dorsal view and rounded in lateral view; eye-nostril distance 9.9 % of SVL; nostrils narrow higher than long, laterally directed; canthus rostralis angular in dorsal and lateral view; loreal region slightly concave; lips rounded; upper eyelid bearing small low rounded tubercles; snout and interorbital region with scattered small low rounded tubercles; upper eyelid width 49.3 % of IOD; tympanic annulus visible on its 3 ⁄ 4 part, with the dorsal margins obscured by supratympanic fold; tympanic membrane present, distinct; tympanum diameter 43.6 % of eye diameter, one subconic postrictal tubercle surrounded by little low tubercles; rounded tubercles along the supratympanic fold and 2 - 3 rounded tubercles surrounded by small tubercles on the temporal region; Choanae small, rounded, not concealed by palatal shelf of maxilla; dentigerous processes of vomers low, distinct, oblique in outline, widely separated, positioned posteromedial to choanae; each vomer bearing 2 or 3 teeth; tongue twice as long as wide, free posteriorly on two-thirds of its length. Temporal folds are formed by a row of small low tubercles, extending from the posterior margin of the upper eyelid to the suprascapular region. Skin on the dorsum with scattered, low-rounded tubercles, small folds formed by little rounded tubercles on upper areas of flanks; lower flanks finely granular; skin on the upper surfaces of forelimbs and hind limbs with indistinct rows of low rounded tubercles. Skin of the throat, chest, and ventral surfaces and limbs is weakly areolate, venter coarsely areolate; discoidal fold weakly defined; cloacal sheath short; skin in the cloacal region smooth. Row of 2 or 3 rounded ulnar tubercles; palmar tubercles low, outer palmar tubercle bifid, V-shaped, approximately twice as large as elongate ovoid thenar tubercle; subarticular tubercles low, well-defined, rounded in ventral view and flattened in lateral view; supernumerary tubercle at the base of fingers present, indistinct; fingers bearing lateral fringes; Finger I shorter than Finger II; discs on Fingers I and II, rounded and slightly expanded; discs more expanded on Fingers III and IV; ventral pads on fingers well-defined by circumferential grooves on Fingers II, III and IV, not evident on Finger I (Fig. 3). Hind limbs robust, tibia length 47.54 % of SVL; foot length 47.67 % of SVL; upper surfaces of hind limbs with low rounded tubercles; heel bearing one to four low rounded tubercles; the outer surface of tarsus bearing a row of four low rounded tubercles; inner tarsal fold present; inner metatarsal tubercle evident, oval, elongate elliptical, three times size of outer metatarsal tubercle; inner plantar surface smooth; subarticular tubercles poorly defined, rounded in ventral view and flattened in lateral view; toes lacking lateral fringes; webbing between toes absent; discs on toes rounded, slightly expanded; all toes with ventral pads defined by circumferential grooves, less distinct on Toe I; relative lengths of toes: I <II <III <V <IV (Fig. 3); Toe V slightly longer than III, disc of Toe III not reaching medial subarticular tubercle of Toe IV; the distal end of the disc on Toe V reaches the medial subarticular tubercle of Toe IV.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	description	Measurements of the holotype (in mm). Adult female, DHMECN 14701, SVL = 15.4; Tibia Length = 7.3; Foot Length = 7.4; Hand Length = 3.8; Head Length = 6.3; Head Width = 6.2; Eye Diameter = 1.9; Tympanum Diameter = 0.8; Forearm Length = 4.2; Snout Length = 5.3; Tarsus Length = 4.2; Thigh Length = 5.5; Upper Arm Length = 3.4; Interorbital Distance = 2.1; upper Eyelid Width = 1.0; Internarial Distance = 1.5; Eye-Nostril distance = 1.5; Wide Finger III = 0.4; Toe IV Width = 0.4.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	description	Call description (Fig. 10). The call description of Pristimantis donnelsoni sp. nov. is based on recordings of a male paratype DHMECN 4774, on 21 March 2007, 19: 00 h, made by JPRP, on the northern flank of Tungurahua Volcano, no environmental parameters were obtained. The calling male was sitting on herbaceous plants on the forest floor. The call is comprised by tonal sounds of constant frequency, with a slight upward modulation at the end of the song. It presents a dominant frequency at 2.93 kHz, with two partial harmonics, first one with a range of 5.86 - 5.94 kHz and the second one with a range of 8.79 - 8.96 kHz. It has a duration ranging from 1052 - 1136 ms, with intervals ranging from 4531 - 5985 ms, emitting at a rate ranging from 8.43 - 10.60 calls / minute. The calls are composed of two notes. The notes have a duration ranging from 236 - 293 ms, with intervals ranging from 522 - 620 ms.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	distribution	Distribution and natural history observations. Pristimantis donnelsoni sp. nov. is known from seven localities on the southern mountains of the Pastaza drainage, Tungurahua Province, Ecuador (Fig. 11). Three localities are located on the Tungurahua Volcano, whereas the others correspond to nearby protected areas (Chamana Reserve and Cerro Candelaria Reserve, Finca Palmonte and Bosque Protector Hacienda Guamag), all at elevations of 2950 - 3800 m. The collection localities range from high Andean montane forest, with trees of the genera Ceroxylum, Clusia, Weinmannia, and others, between 10 - 20 m high, with abundant epiphytes, bromeliads orchids, and mosses, to paramo habitat which is mainly grassland with shrubs in the higher part (Fig. 12). Most of the individuals were found sitting on herbs or on leaf litter on the forest floor, with dense ferns and small herbaceous plants, always near ground level (<30 cm above ground). Calling males are active during the day and sporadically call at night. Sympatric species include Pristimantis aff. devillei, P. modipeplus in Tungurahua Volcano, P. puruscafeum and. P. marcoreyesi and P. lacrimosus group sp. nov., in Cerro Candelaria and Chamana.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	etymology	Etymology. The specific epithet " Pristimantis donnelsoni " is a noun in the genitive case and a patronym for Don Nelson Palacios (Fig. 12 D). We named this new species after him as a tribute to his friendship and collaboration in the first collections of new species described during the last decade and several additional important localities within the upper basin of the Pastaza River. This is a special recognition to him for eternity.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	description	Figs 1, 5, 8, 12, 13, 14, 15, 16, 17	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	diagnosis	Diagnosis. (1) Skin on dorsum shagreen with small scattered, rounded warts; upper flanks with numerous low warts; dorsolateral and w-shaped occipital folds present (Figs 13 - 15); (2) tympanic membrane and tympanic annulus well differentiated (Fig. 12); tympanic annulus not sexually dimorphic; tympanum in males 7 % SVL, tympanum in females 7 % SVL; (3) snout rounded in dorsal, protruding in lateral view (Fig. 15); (4) upper eyelid usually bearing one conical or subconical tubercle and many low tubercles; upper eyelid about in males 63 % IOD, in females 59 % IOD; cranial crests absent; (5) dentigerous processes of the vomer evident, each process bearing 2 - 3 teeth; (6) sphenethmoides on its dorsal view with short blunt anterior border, is short and blunt in ventral view, posterior border present oblique articulation with frontoparietals; posterior border of frontoparietal with an irregular border not projected in dorsal view; zygomatic ramus of squamosal elongated anteriorly, in dorsal view; procesus cultriform of the parasphenoides reaching posterior level of the vomers, rounded anterior border (Fig. 8); (7) males with vocal slits, nuptial pads absent; (8) finger I shorter than finger II; disc on Finger I not expanded; discs on Finger II-IV slightly expanded (Figs 6, 15); (9) fingers with thin lateral fringes; (10) two or three low ulnar tubercles present; (11) heel with low to conical tubercle; outer edge of tarsus with low conical tubercles; (12) inner metatarsal tubercle oval, about 1.5 - 2 x the length of round outer metatarsal tubercle; (13) toes with thin lateral fringes; webbing absent; Toe V slightly longer than Toe III; discs slightly expanded (Figs 15, 16); (14) in life, dorsum brown with darker and lighter markings; venter light grey to black, with or without orange to red spots of different sizes; groin with orange to red spots that are more conspicuous in females than males (see Colour variation; Figs 14, 17); (15) SVL in adult males, 11.8 - 15.6 mm (mean = 13.7, SD = 2.7, n = 30), SVL in adult females 12.3 - 20.2 mm (mean = 16.3, SD = 5.5, n = 13); (16) call composed of 1 to 7 notes, notes have a mean duration of 36.0 + / - 18.3 ms, the mean interval between notes is 43.8 + / - 36.2 ms, emitted at a mean rate of 15.7 + / - 6.1 notes / second. The notes are composed of 2 to 13 pulses. Pulses have a mean duration of 3.9 + / - 1.1 ms, with a mean interval between pulses of 4.2 + / - 2.8 ms, emitted at a mean rate of 202.2 + / - 119.7 pulses / second; (17) dominant frequency of 3.19 + / - 0.05 kHz, with 2 - 6 partial harmonics in the spectrogram; (18) call duration of 165.1 + / - 130.1 ms.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	description	Description of the holotype (Figs 13, 15, 16). Adult female (ZSFQ 0775) robust body; head slightly longer than wide, not as wide as body, head width 35.27 % of SVL (19.7); head length 38 % of SVL; snout rounded in dorsal view, protruding in lateral view; eye-nostril distance 9 % of SVL; with small papilla at tip (Fig. 1); in lateral view, distinct rostral ridge; loreal region slightly concave; nostrils protruding, laterally directed; interorbital area flat, wider than upper eyelid (upper eyelid width 62 % of OID); cranial ridges absent; upper eyelid with one conical and several non-conical tubercles; tympanic membrane well defined, pigmented like surrounding skin; tympanic ring distinct, round; supratympanic fold present, obscuring anterodorsal and posterodorsal edges of the ring; tympanic diameter 46 % of eye length; several low to conical tubercles situated in the area just posterior to tympanum. Choanae are small, with an oval to square shape, not concealed by the palatal shelf of maxillary; the dentigerous process of the vomer oblique, widely separated, posteromedial to choanae, each bearing two small teeth; tongue slightly longer than wide, granular, with a conspicuous notch along the posterior border. The skin of the head is greyish; the dorsum greyish, with scattered small tubercles, some of which are aligned over a W-shaped occipital marking, forming folds; upper flanks with numerous low warts; venter slightly areolate; discoidal fold absent; the cloacal sheath is absent; fingers with thin lateral bangs; length of fingers I <II <IV <III; palmar tubercle round, thenar tubercle oval (Fig. 16); subarticular tubercles round, not prominent; supernumerary palmar tubercles not evident; disc sheath of Finger I not expanded; those of Fingers II-IV slightly expanded; all disc sheaths with nearly elliptical ventral pads defined by grooves (Fig. 16). Hind limbs relatively robust; tibia length 47 % SVL; foot length slightly smaller than tibia length (foot length 44 % SVL); tarsal tubercles present; small, conical tubercle on heel; toes with narrow lateral fringes (Fig. 16); subarticular tubercles round, not prominent; inner metatarsal tubercle oval, two times the size of outer tubercle; few low supernumerary plantar tubercles present (Fig. 16); disc covers of Toes slightly expanded; toes with defined pads; disc pads nearly elliptical; toe lengths I <II <III <V <IV (Fig. 16); the tip of Toe V reaches the middle level of penultimate subarticular tubercle of Toe IV; the tip of Toe III almost reaches the proximal border of penultimate subarticular tubercle of Toe IV.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	description	Measurements of the holotype (in mm). Adult female, ZSFQ 0775. SVL = 19.7; Tibia Length = 9.3; Foot Length = 8.6; Hand Length = 5.0; Head Length = 7.5; Head Width = 7.0; Eye Diameter = 2.4; Tympanum Diameter = 1.1; forearm length = 4.4; snout length = 5.6; Tarsus length = 5.3; Thigh Length = 8.3; Upper arm Length = 3.1; Interorbital Distance = 2.6; Upper Eyelid Width = 1.6; Internarial Distance = 2.1; Eye-Nostril distance = 1.8; Finger III Width = 0.8; Toe IV Width = 0.3.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	description	Call description (Fig. 18). The description is based on the call of an adult male (MZUTI 852), recorded during the night (21: 40 h) by Italo Tapia on 7 June 2012, at the Cordillera de los Guacamayos, Province of Napo, Ecuador. The male was calling from leaf litter at ground level. The air temperature was 12.2 ° C. The recording consisted of 21 calls, 37 notes and 856 pulses. The call of Pristimantis kayi sp. nov. is composed of the emission of several elements with different characteristics, ranging from single notes to pulsed notes. The call (Fig. 1) has a mean dominant frequency of 3.19 + / - 0.05 kHz, with 2 - 6 partial harmonics in the spectrogram. It has a mean duration of 165.1 + / - 130.1 ms. The mean interval between calls is 4565.6 + / - 5742.3 ms, emitted at a mean rate of 32.6 + / - 23.2 calls / minute. Calls are composed of 1 to 7 notes. Notes have a mean duration of 36.0 + / - 18.3 ms. The mean interval between notes is 43.8 + / - 36.2 ms, emitted at a mean rate of 15.7 + / - 6.1 notes / second. The notes are composed of 2 to 13 pulses. Pulses have a mean duration of 3.9 + / - 1.1 ms, with a mean interval between pulses of 4.2 + / - 2.8 ms, emitted at a mean rate of 202.2 + / - 119.7 pulses / second (Table 4). The call of Pristimantis kayi sp. nov. does not present stereotyped notes. Due to their structural elements (i. e. notes-pulses), notes can be classified into three different types: Single note calls (CNS; Fig. 18 A), Pulsed note calls (CNP; Fig. 18 B) and Complex calls (CNP; Fig. 18 C), which contain single (non-pulsed) and pulsed notes in its structure. Temporal and spectral measurements of the different call types are shown in Table 4.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	distribution	Distribution and natural history observations. Pristimantis kayi sp. nov. has been recorded from the Cordillera de Guacamayos, Volcan Sumaco (Napo Province) and Cerro Mayordomo, Machay Reserve, Naturetrek Vizcaya Reserve, Leito Reserve (Tungurahua Province) at an elevational range of 2190 - 3600 m a. s. l. (Fig. 11). Calling males were heard during the day and night. During the night, frogs of Pristimantis kayi sp. nov. were found on the forest floor or perched on low vegetation (always below 60 cm from ground level), including shrubs, grasses (Neurolepis sp.), ferns and Selaginella. The species has been found in primary and disturbed Andean forests, with trees of the genus Weinmannia, Clusia, Alnus and others, of 10 - 15 metres (Fig. 12).	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
10218311988457C8AE068B321C63C959.taxon	etymology	Etymology. The specific epithet " Pristimantis kayi " is a noun in the genitive case and a patronym for Andreas Kay, a German physicist, biologist and friend who spent much of his life documenting and exploring biodiversity, contributing substantially to the conservation of Ecuadorian forests (Fig. 12). Andreas was part of the team that created Fundacion EcoMinga's Dracula Reserva in north-western Ecuador. Additionally, his work shed light on understanding the diversity and endemism patterns of orchids in the Llanganates-Sangay Ecological Corridor, where he passed away in 2019. Some of his amazing photographs can be seen here: https: // www. flickr. com / photos / andreaskay / albums /.	en	Reyes-Puig, Juan Pablo, Urgiles-Merchan, Miguel, Franco-Mena, Daniela, Guayasamin, Juan M., Batallas, Diego, Reyes-Puig, Carolina (2023): Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology. ZooKeys 1180: 257-293, DOI: http://dx.doi.org/10.3897/zookeys.1180.107333, URL: http://dx.doi.org/10.3897/zookeys.1180.107333
