taxonID	type	description	language	source
44270927FFA857446C8EFA2CFC899D39.taxon	diagnosis	Diagnosis (sensu Kury et al. 2022). Inner tip of Cx IV of male without lateral gap but also not touching stigmatic area, because Cx IV is elongate (except for the Caelopygus - Deltaspidium - Progonyleptoidellus clade and Acanthogonyleptes Mello-Leit ã o, 1922; contrasting with tip touching stigmatic area in most K 92, or Cx IV projected laterally, creating a gap in some Gonyleptinae). Fe IV of female robust but elongate, substraight, armed with retroventral, proventral, and dorsal rows of spines (except in Caelopygus elegans (Perty, 1833) and Moreiranula picta (Soares & Soares, 1982); contrasting with Fe IV short in Cearininae and Hernandariinae and elongate, substraight, and weakly armed in Sodreaninae). Ta IV with nine or more articles (except Parampheres bimaculatus (Mello-Leit ã o, 1932), Acanthogonyleptes fulvigranulatus (Mello-Leit ã o, 1922), Moreiranula saprophila (Mello-Leit ã o, 1922); contrasting with 7 – 8 tarsal articles in most K 92).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAA57416C8EFF71FA749E55.taxon	description	urn: lsid: zoobank. org: act: 33 E 35006 - 4961 - 4734 - B 156 - 9 AFEB 5666051 “ Undescribed genus CAE- 01 ”, Kury et al. 2022: 80.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAA57416C8EFF71FA749E55.taxon	etymology	Etymology. Cajango is the name of the protagonist of the novel Corpo Vivo (1962) by the Brazilian writer Adonias Filho (1915 – 1990), which is set in the forests of Camacan (the type locality of the type species). Gender masculine.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAA57416C8EFF71FA749E55.taxon	type_taxon	Type species. Cajango camacanensis sp. nov., hereby designated.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAA57416C8EFF71FA749E55.taxon	diagnosis	Diagnosis. Ocularium armed with a pair of robust, slightly divergent spines in both sexes (Figs. 4 a, b, 7 c, g, 11 a, b). Armature of area III sexually dimorphic: low tubercles in males (Fig. 4 a), spines in females (Fig. 4 d). Tr IV of males with a pair of robust, divergent, retroapical spines, unique within clade K 92 (Figs. 7 a, e). Male genitalia: Apical prongs of penis VP without ears (Figs. 6 a, c). Stylus and ventral process of glans inserted almost in parallel to each other (Fig. 6 b). MS A and B robust, subequal (Fig. 6 b). VP subrectangular, elongate, and with a deep parabolic cleft on the apical border that defines two tapering prongs (Figs. 6 a, c). Lateral distal borders of VP not forming a flange (Fig. 9 b). Basal lobe of VP well developed towards dorsal surface, protruding laterally only in C. camacanensis sp. nov., with robust macrosetae A 1 – A 3 and B 1, all subequal (Figs. 6 a, b, c). Supernumerary A 4 only in some individuals of C. pestifer sp. nov. MS C 1 – C 3 slender and twisted, all clustered on distal part of lateral border of VP (Fig. 6 a). No flange, MS E 1 – E 2 very short, almost collinear with C 1 – C 3, although slightly more ventral. D 1 minute, aligned with MS C. Ventral surface of VP covered with simple microsetae (type 1 of Kury 2016). Podium of truncus deeply concave on the sides which are formed by a soft wrinkled region. Flabellum diamond-shaped, with borders finely serrated (coarsely serrated in C. pestifer sp. nov.). Stylus inserted on glans in acute angle with its ventral process. Stylus straight without well-marked head. Glans sac cubic.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAA57416C8EFF71FA749E55.taxon	distribution	Distribution (Fig. 2). All species of Cajango gen. nov. occur in a small area in southern Bahia state, between de Contas and Jequitinhonha Rivers.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	description	(Figs. 3 – 6) urn: lsid: zoobank. org: act: 3421 AEC 1 - DF 0 E- 43 C 7 - 85 DE- 14 BDAAB 44501	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	etymology	Etymology. Species name refers to the toponym Camacan, the type locality and setting of the novel Corpo Vivo, matching the generic name.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	materials_examined	Type series. BRAZIL, Bahia State: ♂ holotype (MNRJ 19390), 24 ♂, 21 ♀, 1 juv. paratypes (MNRJ 19150; one couple subsampled to MZSP), Camacan, RPPN da Serra Bonita, 11 - 13. vi. 2009, A. Chagas Jr, A. B. Kury, D. R. Pedroso, A. P. L. Giupponi & V. Dill leg.; 1 ♀ paratype (MNRJ 19392), same locality, 14. iii. 2008, V. Dill leg.; 1 ♂. 1 ♀. 1 juv. paratypes (MNRJ 19391), same locality, 2 - 3. x. 2011, A. Pérez-González & B. Huber leg.; 3 ♂ paratypes (MNRJ 9054), same locality, 29. iii. 2012, V. Dill leg., 1 ♂ paratype (MZSP 73452), same locality, 24 - 26. vi. 2009, R. Pinto-da-Rocha, C. Bragagnolo & M. B. DaSilva leg.; 1 ♀ paratype (MZSP 73453), same data; 1 ♂ paratype (MZSP 72779), same locality, 1. xi. 2014, A. A. Nogueira leg.; 5 ♂ paratypes (MZSP 73454), Camacan, RPPN da Serra Bonita, Trilha das Bromélias (15 ° 23 ’ 23.1 ” S 39 ° 34 ’ 03.6 ” W), 3. ix. 2014, J. P. P. Pena-Barbosa leg.; 4 ♂, 2 ♀ paratypes (MZSP 73455), Camacan, RPPN da Serra Bonita, Trilha do Riacho (15 ° 23 ’ 17.1 ” S 39 ° 33 ’ 59.2 ” W), 1. xi. 2014, A. A. Nogueira leg.; 5 ♂, 4 ♀ paratypes (MZSP 73456), same locality, 2. xi. 2014, S. Outeda-Jorge leg.; 8 ♂, 4 ♀ paratypes (MZSP 73457), same locality, 2. xi. 2014, A. A. Nogueira leg.; 1 ♀ paratype (MNRJ 8688), Una, REBIO de Una, (15 ° 10.3597 ’ S, 39 ° 3.2953 ’ W), 179 m, 12 - 13. vi. 2015, A. P. Pinto leg; 1 ♂ paratype (MZSP 42090), Arataca, Serra das Lontras, Palmeiras ranch, 1 - 7. iii. 2009, A. Camacho leg. Other material examined. BRAZIL, Bahia State: 1 ♂ brittle, badly damaged (MNRJ 18928), Ilhéus, Ilha do Barbosa, without further data.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	distribution	Distribution. Brazil, Bahia State: Camacan, Ilhéus and Una. This species is sympatric with C. pestifer sp. nov. in Ilhéus and with Cajango vindicator sp. nov. in Camacan.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	diagnosis	Diagnosis. Body background color in both sexes brown; scutal areas with several granules (Figs. 3 a, c). Anal operculum with large, rounded apophysis in male, much smaller apophysis in females (Figs. 3 a, c). Cx IV with retrolateral, apical, spiniform apophysis in both sexes, larger in males (Figs. 3 a, c). Fe IV of both sexes without dorso-basal apophysis. Basal lobes of VP projected to the sides as triangles (Fig. 6 c). Flabellum quadrangular, finely serrated (Fig. 6 a).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFAD574D6C8EFB92FE429815.taxon	description	Description of male holotype. Measurements: CL = 3.4, CW = 4.2, AL = 4.3, AW = 7.8. Pp = 9.1. Legs, I: Fe = 6.4, Ti = 4.5, Mt = 7.2; II: Fe = 14.9, Ti = 12.0, Mt = 15.3; III: Fe = 11.0, Ti = 6.4, Mt = 11.8; IV: Fe = 14.5, Ti = 11.2, Mt = 20.1. Dorsum. Large and slender animal, with granulated tegument and elongate legs in predominantly brown hues. DS gamma pyriform, with posterior border slightly concave. Carapace pentagonal, with ocularium at its anterior portion, but set far away from anterior margin of scutum. Ocularium moderately high, armed with two high parallel spines. Frontal hump also moderately elevated, with two small acuminate tubercles. Anterior border of scutum with one tubercle each side. Areas of dorsal scutum all covered with moderate-sized tubercles, each with a larger paramedian pair, growing larger posteriorly, so that the pair in area III is the largest. Mesotergum divided into three areas; area I divided in broadly connected halves. Area V and free tergites each with a transverse row of 6 – 14 tubercles. Anal operculum with a very large, median, lanceolate apophysis. Legs. Cx IV much larger than the other coxae taken together, slanted. Stigmatic area with elongate axis, with short branches, sternite II square. Cx IV oblique, visible beyond scutum in dorsal view, with strong, straight and uniramous prolateral apical apophysis and strong retrolateral apical apophysis. Tr IV with two retroapical apophyses forming a V. Fe IV with PV and RV rows of small tubercles increasing in size apically, RL surface with two stouter spines proximally (RL 1 and RL 2, Fig. 5), two more in the middle (RL 5 and RL 6), and one more distally (RL 9); other rows formed by minute granules. Tarsal counts 6 (3) / 15 (4) / 8 / 10. Color (in 75 % ethanol). Background color of dorsal scutum Strong Yellowish Brown (# 74), with lighter patches of Moderate Yellow (# 87) along the grooves and arborescent in lateral margins of scutum. Mesotergal granules Brilliant Greenish Yellow (# 98). Chelicerae and pedipalps Brilliant Greenish Yellow (# 98) and densely reticulated in Moderate Olive (# 107). Legs I – III up to femora predominantly Light Olive (# 106), with lighter granulation. Cx IV background Dark Grayish Red (# 20), with apex and rest of podomeres Very Dark Purplish Red (# 260). Apical rim of Tr IV Dark Reddish Orange (# 38). Ventral Cx and stigmatic area predominantly Strong Greenish Yellow (# 99), with dense honeycombed Brownish Gray (# 64). Male genitalia (Figs. 6 a – c). Basal lobes of VP robust, leaning upon the truncus, projected to the sides as triangles. MS A 1 to A 3 and B 1 all robust, subequal, and set at the rim of basal lobe, forming an arch. D 1 set close to E 2. Flabellum quadrangular, finely serrated. Female (Figs. 3 c, d, 4 d). Similar to male in color, proportions, and armature. It differs from male by: (1) armature of Cx IV: weaker prolateral apical apophysis and no retrolateral apical apophysis; (2) Fe IV with only rows of smaller spines; (3) smaller apophysis on anal operculum; and (4) scutal area III with a pair of strong spines instead of low tubercles.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA1574C6C8EFD55FF0D9815.taxon	description	(Figs. 7 – 9) urn: lsid: zoobank. org: act: B 1 AA 95 C 2 - 0684 - 4283 - 84 A 6 - D 8 A 7 F 683 E 7 F 7	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA1574C6C8EFD55FF0D9815.taxon	etymology	Etymology. From the Latin adjective pestĭfer, ĕra, ĕrum (destructive, baleful, noxious). Named thus due to it being collected as part of the campaigns of SEPFA (Service of Studies and Research on the Yellow Fever), and the name also refers to the rage of the character Cajango in the aforementioned novel.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA1574C6C8EFD55FF0D9815.taxon	materials_examined	Type series. BRAZIL, Bahia State: ♂ holotype, 1 ♂, 2 ♀ paratypes (MNRJ 5430), Ilhéus, viii. 1944, Campaign of Serviço de Estudos e Pesquisas sobre a Febre Amarela leg.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA1574C6C8EFD55FF0D9815.taxon	diagnosis	Diagnosis. Body background color yellow in both sexes, with carapace and area III more strongly shaded in brown; scutal areas only with a pair of paramedian granules each and a few scattered extra granules (Figs. 7 a, b). Anal operculum in both sexes with a small median tubercle (Figs. 7 e, f). Cx IV of male without retrolateral apical apophysis (Fig. 7 e). Fe IV with dorso-basal, anvil-shaped apophysis (Fig. 7 a). Basal lobes of VP only very gently projected to the sides as small triangles (Fig. 9 e). Flabellum quadrangular, but this outline is distorted because of the deep serration (Fig. 9 c).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA1574C6C8EFD55FF0D9815.taxon	description	Description of male holotype. Measurements: CL = 3.8, CW = 4.7, AL = 4.7, AW = 10.4. Pp = 9.7. Legs, I: Fe = 6.9, Ti = 5.2, Mt = 8.2; II: Fe = 14.7, Ti = 11.2, Mt = 16.0; III: Fe = 11.3, Ti = 6.4, Mt = 11.9; IV: Fe = 15.2, Ti = 11.0, Mt = 18.7. Dorsum. Large and slender animal, with smooth tegument and elongate legs in predominantly yellow hues. DS gamma pyriform, with posterior border slightly concave. Carapace cordiform, with ocularium at its anterior portion, but set far away from anterior margin of scutum. Ocularium moderately high, armed with two slightly divergent spines. Frontal hump also moderately elevated, with two small low spines. Anterior border of scutum with one tubercle each side. Dorsal scutum glossy, except for one row of scattered tubercles on each lateral area and one transverse row on each area I to III, each row with an outstanding paramedian pair, much larger on area III. Mesotergum divided into three areas; area I divided into broadly connected halves. Area III with a paramedian pair of blunt tubercles and a few scattered granules. Area V and free tergites each with a transverse row of 4 – 12 granules. Anal operculum without any apophysis. Legs. Cx IV much larger than the others taken together, slanted. Stigmatic area with elongate axis, with short branches, sternite II narrow and rounded. Cx IV oblique, visible beyond scutum in dorsal view, with strong, bent prolateral apical apophysis, provided with a small secondary branch. Without a retrolateral apical apophysis. Tr IV with two greatly developed retroapical apophyses forming a V. Fe IV with several stout and curved DO apophyses all along the podomere; strong pectination at RL 1, RL 2, isolated stout apophyses, from RL 3 to RL 10; PV and RV rows of small tubercles increasing in size apically. Tarsal counts 6 (3) / 14 (4) / 8 /?. Color (in 75 % ethanol). Dorsal scutum, chelicerae, and pedipalpi background Brilliant Greenish Yellow (# 98). Scutum with Grayish Brown (# 61) patches covering most of carapace (including ocularium), anterior border of scutum and region around main armature of area III. Chelicerae and pedipalpi with sparse Grayish Brown (# 61) reticulation. Legs I – III predominantly Moderate Olive (# 107). Leg IV background Dark Grayish Olive (# 111) in most regions getting darker to Olive Black (# 114). Ventral Cx and stigmatic area predominantly Light Olive (# 106) with dense mottling in Dark Olive (# 108). Male genitalia (Figs. 9 a – e). Basal lobes of VP leaning upon the truncus, but mediocre, little projected to the sides. MS A 1 to A 3 and B 1 all robust, strongly lanceolate, subequal in length and arranged into two groups: A 1 and B 1 more distal than A 2 and A 3. D 1 set far from E 2, midway between clusters of MS C and A. Solea of glans clearly projected apically. Flabellum fan-shaped, with few deep serrations. Female (Figs. 7 b, d, f). Very similar to male in habitus and colors, except for: (1) scutal area III with a pair of strong spines instead of low spines; (2) Cx IV much smaller, mostly hidden under scutum in dorsal view and without strong dorso-apical apophysis; (3) armature of Tr IV much weaker; (4) dorsal scutum outline gamma classical; (5) Fe IV not curved to the midline and without larger individual spines, although still armed with rows of substantial spines.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA2574E6C8EFF71FADB9D18.taxon	description	(Figs. 10 – 13) urn: lsid: zoobank. org: act: 8304 FD 3 B-BE 9 F- 42 D 0 - ABE 5 - 729928 E 8 BE 79	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA2574E6C8EFF71FADB9D18.taxon	etymology	Etymology. Species name is the Latin noun vindĭcātor, ōris (avenger) used in apposition, and it refers to the revenge of the character Cajango in the novel Corpo Vivo, in allusion to the generic name.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA2574E6C8EFF71FADB9D18.taxon	materials_examined	Type series. BRAZIL, Bahia State: ♂ holotype, 2 ♀ paratypes (MNRJ 9053), Camacan, Highway BR 101, 28. iii. 2012, V. Dill leg; 1 ♂ paratype (MZSP 73451), Uruçuca, Parque Estadual Serra do Conduru (14 ° 29 ’ 02.8 ” S 39 ° 06 ’ 10.4 ” W), 27. viii. 2014, S. Outeda-Jorge leg.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA2574E6C8EFF71FADB9D18.taxon	diagnosis	Diagnosis. Body background color yellow in both sexes, with carapace and area III more strongly shaded in brown; scutal areas only with a pair of paramedian granules each, and a few scattered extra granules (Figs. 10 a, c). Anal operculum with a large lanceolate apophysis, stouter in male (Figs 10 b, d). Sternite II narrow and rounded. Cx IV of male without retrolateral apical apophysis (Fig. 10 b). Fe IV without dorso-basal apophysis (Fig. 12). Basal lobes of VP not projected to the sides (Fig. 13 c). Flabellum quadrangular, finely serrated (Fig. 13 a).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA2574E6C8EFF71FADB9D18.taxon	description	Description of male holotype. Measurements: CL = 4.2, CW = 5.0, AL = 5.6, AW = 10.7. Pp = 9.7. Legs, I: Fe = 7.0, Ti = 4.3, Mt = 7.4; II: Fe = 15.8, Ti = 11.8, Mt = 15.8; III: Fe = 11.7, Ti = 6.4, Mt = 10.4; IV: Fe = 16.6, Ti = 10.5, Mt = 18.7. Dorsum. Large and slender animal, with smooth body and elongate legs in predominant yellow hues. DS gamma pyriform, with posterior border slightly concave. Carapace cordiform, with ocularium at its anterior portion, but set far away from anterior margin of scutum. Ocularium only moderately high, armed with two slightly divergent spines. Frontal hump also moderately elevated, with two small acuminate tubercles. Anterior border of scutum with one tubercle each side. Dorsal scutum glossy, except for one row of scattered tubercles on each lateral area and one transverse row on each area I and II, each row with a highlighted paramedian pair. Mesotergum divided into three areas; area I divided into broadly connected halves. Area III with a paramedian pair of blunt tubercles and a few scattered granules. Area V and free tergites each with a transverse row of 4 – 12 granules. Anal operculum with a very large, median, lanceolate apophysis. Legs. Cx IV much larger than the others taken together, slanted. Stigmatic area with elongate axis, with short branches, sternite II narrow and rounded. Cx IV oblique, visible beyond scutum in dorsal view, with strong and straight prolateral apical apophysis, provided with a small secondary branch. Retrolateral apical apophysis lacking. Tr IV with two retroapical apophyses forming a V. Fe IV with PV and RV rows of small tubercles increasing in size apically, RL surface with two stouter spines in the middle (RL 4 and RL 6) and two more distally (RL 7 and RL 9), other rows formed by minute granules. Tarsal counts 6 (3) / 14 (4) / 8 / 10. Color (in 75 % ethanol). Dorsal scutum, chelicerae and pedipalps and Tr I – III background Light Yellow (# 86). Scutum with Brownish Gray (# 64) patches covering most of carapace (including ocularium), anterior border of scutum and region around main armature of area III. Chelicerae and pedipalpi with sparse Brownish Gray (# 64) reticulation. Tr I – III heavily mottled in Deep Brown (# 56). Legs I – III predominantly Brownish Gray (# 64). Leg IV background Deep Brown (56) in some regions getting darker to Brownish Black (# 65). Anal operculum Brownish Black (# 65). Ventral Cx and stigmatic area predominantly Moderate Greenish Yellow (# 102) to Moderate Yellow (# 87) with honeycombed reticulation in Moderate Olive Brown (# 95). Reticulation much denser on sternites. Male genitalia (Figs. 13 a, b, c). Basal lobes of VP leaning upon the truncus, but mediocre, not projected to the sides. MS A 1 to A 3 and B 1 all robust, subequal and arranged into two groups: A 1 and B 1 more distal than A 2 and A 3. Solea of glans somewhat projected apically. Flabellum attenuate quadrangular, finely serrated. Female (Figs. 10 c, d, 11 d). Very similar to male in habitus and colors, except for: (1) paramedian armature of area III represented by high capitate spines instead of squat blunt tubercles; (2) tubercles on row at lateral areas higher; (3) Cx IV much smaller, mostly hidden under scutum in dorsal view and without strong dorso-apical apophysis; (4) armature of Tr IV much weaker; (5) dorsal scutum outline gamma classical; (6) Fe IV not curved to the median line and without larger individual spines, although still armed with rows of substantial spines.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA7574B6C8EFBD0FA749C4D.taxon	description	Key to the genera of Sodreaninae (sensu Kury et al. 2022)	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	description	urn: lsid: zoobank. org: act: C 662 C 684 - A 0 C 1 - 459 C-A 8 B 9 - 905 D 9 E 91 CA 23 “ Undescribed genera SOD- 01, SOD- 02 ” — Kury et al. 2022: 79.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	etymology	Etymology. From the Greek Τῑτάν (one of the titans, divine beings descending from the primordial deities) + leptes (truncation of generic name Gonyleptes), referring to the very large size of its species. Gender masculine.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	type_taxon	Type species. Titanoleptes hyperion sp. nov.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	diagnosis	Diagnosis. Coxa IV of male without retrolateral apical apophysis (Fig. 16 c), except in T. cupidensis (Fig. 20 a). Trochanter IV of male armed with a strong, retrolateral, apical apophysis (Figs. 21 a, c). Patella and tibia IV covered with yellow acuminate tubercles (Figs. 20 d, 24 b). Femur IV of male either substraight (Fig. 19 a) or strongly curved to the midline on basal 1 / 4 in dorsal view (Figs. 17 a, c). Tibia IV armed with rows of acuminate tubercles (Figs. 20 d, 24 b). VP of penis with ear-flaps small, triangular, ectal vertex connected but not fused with flange. Flabellum quadrangular, as a stylized shell, proximal sides gently arched, non-serrate, distal half extremely reduced, with distal sides coalesced into one, arched and deeply serrate (Fig. 22 e).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	distribution	Distribution (Figs 14 – 15). Southern Bahia State and Espírito Santo State. All four species of Titanoleptes occur in the WWF ecoregion NT 0103 (Bahia coastal forests) and two subsets of two species each are distributed on opposite margins of the Doce River.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFA757546C8EF96AFA749B71.taxon	discussion	Internal and external relationships of the four species of Titanoleptes. At a first glance, one could easily identify two forms in Titanoleptes gen. nov.: (1) T. eros sp. nov. and T. cupidensis are gracile and have a median apophysis on free tergite III in both sexes, substraight femur IV, and naked glans; (2) T. hyperion sp. nov. and T. calcar are robust and lack an apophysis on free tergite III of males, have a heavily curved femur IV, and have a unique scutum covering the glans. The parapatrid pattern of these two pairs — T. eros + T. cupidensis and T. hyperion + T. calcar are found on opposite margins of the Doce River (Figs. 13 – 14) — is highly suggestive of a sister relationship between each of the members of these pairs, regardless of whether the pairs themselves form a more inclusive clade or not. This hypothesis, though, is not corroborated by any of the analyses. The cladistic analysis of Kury et al. (2022) explored different parameters to assess the arrangement of the four Titanoleptes in the phylogeny of Sodreaninae. The favored analysis using the setk script (with implied weights and K value ca. 12) resolved Titanoleptes as a sister clade of Friburgoia / Urodiabunus, with an internal branching sequence of T. eros, T. cupidensis, and then T. hyperion + T. calcar. IE settings of K = 1 and 5, plus equal weights, yielded the same result. However, Bayesian inference did not recover Titanoleptes at all, finding only the clade T. hyperion + T. calcar and then recovering T. eros and T. cupidensis in a trichotomy with the rest of the Sodreaninae. We have decided not to arrange the four species of Titanoleptes into two genera, despite the weak support for their monophyly.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB857516C8EFEB1FB639BD9.taxon	description	(Figs 16 – 18) urn: lsid: zoobank. org: act: 8 D 80 C 035 - 705 B- 46 AF-B 253 - 3 C 5633 E 186 A 5	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB857516C8EFEB1FB639BD9.taxon	materials_examined	Type series. Metagonyleptes calcar: BRAZIL, Bahia State: ♂ holotype (SMF RI 888), “ Bahia ”, without further locality data; examined by photograph. Moojenia bicalcarata: BRAZIL, Espirito Santo State: 2 ♂, 2 ♀ syntypes (MNRJ 58209 [and not “ 55094, ” as incorrectly reported in the original description]), Colatina, M. Rosa leg., 1936 – 1937, examined.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB857516C8EFEB1FB639BD9.taxon	diagnosis	Diagnosis. Body color mahogany with black reticle (Fig. 16 a). Ocularium armed with two small spherical tubercles of lighter color contrasting with background (Figs. 16 b, e, g). Scutal area III armed with two contrasting dark brown and low but strong cones, placed closely together (Figs. 16 e, f). Free tergite III unarmed in males (Fig. 16 b). RD 6 of male Fe IV straight (Figs. 16 a, b); FT II-III of female with yellow cones (Figs. 16 g, h). Glans sac projected apically and forming an incomplete circular wall around the stylus (Fig. 18 b, e).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB857516C8EFEB1FB639BD9.taxon	materials_examined	Material examined. BRAZIL, Bahia State: 1 ♂ (MNRJ 18851), Boa Nova, 19. xi. 2006, C. Canedo leg.; 1 ♂, 1 ♀ (MNRJ 19153), Camacan, RPPN da Serra Bonita, 11 - 13. vi. 2009, A. Chagas Jr, A. B. Kury, D. R. Pedroso, A. P. L. Giupponi & V. Dill leg.; 1 ♂ (MNRJ 2133), same loc., 800 m, 14. iii. 2008, V Dill leg.; 1 ♂ (MNRJ 8786), same loc., 2 - 3. x. 2011, A. Pérez-González & B. Huber leg.; 1 ♂, 1 ♀ (MNRJ 6998), Itabuna, x. 1995, P. S. Fonseca, & L. P. Gonzaga leg.; 4 ♂, 3 ♀ (MNRJ 17725), Porto Seguro, Arraial d’Ajuda, Fazenda do Sr. Valter, (16 ° 32.304 ’ S / 039 ° 08.592 ’ W), 24 - 27. ii. 2005, A. B. Kury et al. leg.; 1 ♂ (MNRJ 19185), Porto Seguro, Estação Vera Cruz, ii. 2003, E. Coutinho leg.; 1 ♂, 1 ♀ (MNRJ 19152), Porto Seguro, Parque Nacional do Pau Brasil, 14 - 17. vi. 2009, A. Chagas Jr, A. B. Kury, D. R. Pedroso, A. P. L. Giupponi & V. Dill leg.; Espirito Santo State: 3 ♂, 4 ♀ (MNRJ 18934), Conceição da Barra, FLONA do Rio Preto, Córrego das Bestas, (- 18.37159 ° - 39.84930 °), 45 m, 19. x. 2005, A. P. L. Giupponi, A. B. Kury, T. Bernabé, M. Milleri, R. Rodrigues & T. Souza leg.; 1 ♂, 1 ♀ (MNRJ 16298), Conceição da Barra, FLONA do Rio Preto, Trilha da Jaqueira, (18.35828 ° S, 39.84559 ° W), 31 m, 18. x. 2005, A. P. L. Giupponi, A. B. Kury, T. Bernabé, M. Milleri, R. Rodrigues & T. Souza leg.; 2 ♂ (MNRJ 17582), Linhares, Reserva Florestal da Companhia Vale do Rio Doce, 28. viii. 1981, C. A. G. Cruz, E. Izecksohn & O. L. Peixoto leg.; 8 ♂, 7 ♀ (MNRJ 4550), Linhares, Reserva Florestal da Companhia Vale do Rio Doce, 06. xii. 1978, A. Peracchi, M. Peracchi & B. O. Fraga leg.; 1 ♂ (MNRJ-HS 0794), same loc., 13 - 15. x. 1979, E. Izecksohn & Barbosa leg.; 1 ♀ (MNRJ 5416), Pedro Canário, Rio Itaúna do Norte, Fazenda do Caboclo do Sr. Alberto Castro (18 ° 14 ’ 55 ” S, 40 ° 02 ’ 48 ” W), viii. 1961, H. N. Cunha leg.; 1 ♂, 2 ♀, 1 juv. (MNRJ 6910), Sooretama, REBIO de Sooretama, i. 1979; 2 ♂, 1 ♀ (MNRJ 8724), same loc., 27 - 28. ix. 2011, A. Pérez-González & B. Huber leg.; 2 ♂, 1 ♀ (MNRJ 17818), same loc., Córrego do Rodrigues, 18. iv. 2006, T. Bernabé, A. P. L. Giupponi, A. B. Kury, C. S. Costa, E. G. Vasconcelos & E. Wienskoski leg.; 1 ♂ (MNRJ 17854), same loc., Paraisópolis, (- 18.9733 °, - 40.2389 °), 101 m, 19. iv. 2006, T. Bernabé, A. P. L. Giupponi, A. B. Kury, C. S. Costa, E. G. Vasconcelos & E. Wienskoski leg.; 11 ♂, 18 ♀ (MNRJ 17850), same loc., Quirinão, 20. iv. 2006, T. Bernabé, A. P. L. Giupponi, A. B. Kury, C. S. Costa, E. G. Vasconcelos & E. Wienskoski leg.; 9 ♂, 12 ♀ (MNRJ 17842), same loc., Trilha da Sede, (- 19.0552 °, - 40.1468 °), 18. iv. 2006, T. Bernabé, A. P. L. Giupponi, A. B. Kury, C. S. Costa, E. G. Vasconcelos & E. Wienskoski leg.; 3 ♂, 3 ♀ (MNRJ 18929), Sooretama, xii. 1973, A. Peracchi leg.; 4 ♂, 1 ♀ (MNRJ-HS 235), same loc., 28. iii. 1959, R. G. Kloss leg.; 3 ♂, 3 ♀ (MNRJ 18929), same loc., xii. 1973, A. Peracchi leg.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB857516C8EFEB1FB639BD9.taxon	distribution	Literature records. Espirito Santo State, “ Colatina ” (currently São Gabriel da Palha), São José River (B. Soares 1944 h). Mello-Leit ã o (1942 b) also mentions Goitacazes, but all syntypes are labeled “ Colatina ”. Distribution (Fig. 14) Brazil, Bahia State, Boa Nova; Camacan; Itabuna; Porto Seguro. Espírito Santo State, Colatina; Conceição da Barra; Linhares; Pedro Canário; São Gabriel da Palha; Sooretama.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD57516C8EFE18FC399ECE.taxon	description	(Figs 19 – 22) urn: lsid: zoobank. org: act: D 0 BE 58 AC- 7 FFD- 4 BEB- 86 B 6 - F 7 B 86 CA 94637	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD57516C8EFE18FC399ECE.taxon	materials_examined	Type series. BRAZIL, Espirito Santo State: ♀ holotype (MZSP OS 2913, examined by photograph), “ Linhares ” [currently Sooretama]: Chapada de Cupido. Material examined. BRAZIL, Espirito Santo State: 1 ♀ (MNRJ 9097), Linhares, Reserva Florestal da Companhia Vale do Rio Doce, 28. viii. 1981, C. A. G. Cruz, E. Izecksohn & O. L. Peixoto, leg.; 2 ♂, 1 ♀ (MNRJ 17869), Sooretama, REBIO de Sooretama, (- 19.0012 ° - 40.1392 °), 93 m, 21. iv. 2006, T. Bernabé, A. P. L. Giupponi, A. B. Kury, C. S. Costa, E. G. Vasconcelos & E. Wienskoski leg.; 1 ♂ (MACN AK 076), Sooretama, REBIO de Sooretama, x. 1962.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD57516C8EFE18FC399ECE.taxon	diagnosis	Diagnosis. Ocularium in both sexes with two high erect spines, concolorous with background (Figs. 19, a, b, h). Scutal area III in both sexes armed with small, paramedian, rounded tubercles (Figs. 19 b, d). Free tergite III armed with a median spiniform apophysis in both sexes (Figs. 19 b, h). Coxa IV of male with strong, retrolateral, apical apophysis, clearly longer than the prolateral one (Fig 20 a). Trochanter IV of male without proximal prolateral apophysis (Fig 20 a). Glans free, without a circular wall around stylus (Fig. 22 g).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD57516C8EFE18FC399ECE.taxon	distribution	Distribution. Brazil, Espírito Santo State (ES), Linhares; Sooretama.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	description	(Figs 23 – 26) urn: lsid: zoobank. org: act: EB 41 BA 5 F- 6 C 2 C- 4 E 04 - 97 CD-DBD 02 ABADBAA	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	etymology	Etymology. Noun in apposition. In Greek mythology, Eros was the god of love, whose Roman counterpart was Cupid. Thus, this name mirrors another species of the genus, C. cupidensis.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	materials_examined	Type series. BRAZIL, Espirito Santo State: ♂ holotype (MNRJ 18440) ,, Santa Teresa, REBIO Augusto Ruschi, Cachoeira II, (19 ° 54.396 ’ S, 40 ° 33.722 ’ W), 11. viii. 2006, D. Mota leg.; 2 ♂, 2 ♀, 7 juv. paratypes (MNRJ 7077), Santa Teresa, Estação Biológica de Santa Lúcia, xi. 2006, J. Wesley leg.; 2 ♂, 1 ♀ paratypes (MNRJ 22), Santa Teresa, 22. xii. 1980, E. Izecksohn & O. L. Peixoto leg.; 3 ♂, 1 ♀ paratypes (MNRJ 18927), Santa Teresa, 3 - 5. iv. 1981, C. A. G. Cruz, O. L. Peixoto, S. Silva, A. Reis & E. Izecksohn leg.; 1 ♂ paratype (MNRJ 18941), Santa Teresa, 27 - 28. ix. 1984, O. L. Peixoto, C. A. G. Cruz & E. Izecksohn leg.; 1 ♂ paratype (MNRJ 18951), Santa Teresa, REBIO de Santa Lúcia, 16 - 20. i. 2006, AracnoLab leg.; 2 ♂ paratypes (MNRJ-HS 872), Santa Teresa, 24. viii. 1974, A. L. Perachi leg; 1 ♀ paratypes (MZSP 73450), Santa Teresa, REBIO Augusto Ruschi, 24 - 30. v. 2008, F. Carbayo leg.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	diagnosis	Diagnosis. Ocularium in both sexes with two high erect spines, concolorous with background (Fig. 23 c). Scutal area III armed with small paramedian rounded tubercles (Figs. 23 d, e). Free tergite III armed with a median apophysis in both sexes (Figs. 23 a, f). Coxa IV of male without retrolateral apophysis (Fig. 23 b). Trochanter IV of male with proximal prolateral conical apophysis forming short pincer with coxal apophysis (Fig. 24 a). Glans of penis free, without a circular wall around stylus (Fig. 26 d).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	distribution	Distribution. Brazil, Espírito Santo State, Santa Teresa.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFBD575F6C8EFAEDFDD09DBC.taxon	description	Description of male holotype. Measurements: CL = 2.9, CW = 3.6, AL = 3.9, AW = 6.3. Pp = 8.4. Legs, I: Fe = 4.8, Ti = 3.2, Mt = 5.1; II: Fe = 10.5, Ti = 8.0, Mt = 9.8; III: Fe = 8.7, Ti = 4.6, Mt = 7.3; IV: Fe = 12.1, Ti = 7.5, Mt = 12.1. Dorsum. Dorsal scutum outline gamma pyriform in dorsal view, quite flattened, so that Cx IV is much thicker than scutum in lateral view. Carapace mid-area cordiform, with narrow and low ocularium. Ocularium armed with a pair of paramedian, strong, divergent spines. Frontal hump low, armed with a pair of small granules. Two ozopores, first slit-like, second rounded. Mesotergum divided into 4 scutal areas, although areas III and IV are extensively fused to each other, except for the lateral thirds, where faint grooves are visible. Area I long, divided into broadly joined halves. Area II projected into area I. Area III strongly arched frontwards and moderately encased into area II. Carapace smooth and unarmed, area I with a transverse row of five granules on each side, area II with a similar row of 10 granules, without any especially developed in the middle. Areas III-IV with two rows of a few granules plus a paramedian pair of larger rounded capitate tubercles, these forming a trapezoid with posterior tubercles much smaller than anterior ones, and more widely spaced. Area V (posterior border of scutum) with a transverse row of granules. Lateral margins of scutum with a row of tubercles stretching from mid-carapace to area V, and which increase in size toward the posterior. Each free tergite with a row of larger granules, free tergite III with an immense spiniform apophysis. Posterior border of dorsal scutum with arched concavity. Venter. Cx I free, other connected by tubercular bridges. Cx IV much larger than the others combined, without any ventro-apical tubercle. Stigmatic area posterior border deeply concave. Free tergites each with a transverse row of granules. Stigmatic area with very long axis and large, elliptical, slanted stigmata. Pedipalps. Moderately elongate, as long as body. Tr with thin neck, 1 + 3 setiferous tubercles dorsal and ventral. Fe with a row of 3 extremely weak ventral setiferous tubercles and much stronger meso-distal spine. Pa strongly curved, unarmed. Ti and Ta flattened ventrally. Spination of Ti: mesal IiIi, ectal IiIi (last two geminated). Spination of Ta: mesal IiIi, ectal IiIi. Legs. All legs slender, only leg IV with significant armature. Dorsal apophyses of Cx I and II: Cx I anterior single, triangular erect, posterior larger, conical, directed backwards. Cx II anterior single, triangular erect, posterior flat triangular with small accessory tubercle. Tr IV with prodorsal median conical apophysis, which connects with apophysis of Cx IV and retrolateral distal conical apophysis of the same size. Otherwise only with a few small and scattered ventral tubercles. Legs I to III entirely unarmed. Fe I – II straight, III slightly arched. Fe IV: elongate, curved to the median line at 1 / 2 length; neither curved to ventral nor to dorsal. Proventral: row of numerous, small, acuminate tubercles all along the femur length. Retroventral: row of numerous, small, acuminate tubercles + unarmed mid-region + 6 larger tubercles distally. Retrodorsal: row of tubercles, being the basalmost and distalmost ones larger, with RD 9 the largest. Dorsal: row of small tubercles, of which Do 6 to Do 8 more outstanding. Prodorsal: row of numerous small acuminate tubercles all along the femur length. Pa IV with many rows of tubercles, Ti IV as well, but tubercles are coarser. Mt IV smooth, thin, and slender. Tarsal counts: 6 (3) - 6 (3) / 10 (3) - 10 (3) / 7 - 7 / 8 - 8. Color (in 75 % ethanol). Body and appendages background Strong Greenish Yellow (# 99) with some darker mottling, especially on carapace. Leg IV considerably darker starting on apical portion of Cx. Venter as dorsum, with unobtrusive honeycomb reticulation. Male genitalia (Figs. 26 a – d). VP trapezoid, and distal border with a moderately deep parabolic cleft. Field of type 1 microsetae occupying most of VP except distal corners and not extending dorsally into podium but rather to latero-basal lobe. Basal lobe as a large, rounded polygon projecting dorsally from VP. Macrosetae (MS) A and B moderately long, forming an arch on latero-basal lobe, B only slightly smaller than A 1 – A 3. MS C 1 – C 3 elongate, straight basally, but distally helical. MS D 1 minute, midway between C and A, E 1 – E 2 short, situated on flange. Flabellum diamond-shaped, with ventral half outline gently serrated. Glans sac short, multi-folded, forming apically a solea projected as a short and blunt dorsal process. Stylus stalk arising from ventralmost part of glans sac; stylus inserted on stalk in candelabrum-mode. Stylus distally provided with distal denticles, which more apically form a large spiny protuberance. Female (MNRJ 7077). The female is essentially similar to male in build, appendages, and armature of dorsal scutum. Main differences are in the apophysis of free tergite III and in the armature of leg IV, which are much reduced. Tarsal counts: 6 (3) / 9 (3) / 7 / 8.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	description	(Figs 27 – 29) urn: lsid: zoobank. org: act: 508 E 2223 - 6 F 21 - 4317 - 95 B 7 - B 4 F 5 A 74 FE 962	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	etymology	Etymology. From the Greek mythological name Ὑπερίων, one of the twelve Titans, children of Gaia and Uranus. Noun in apposition. This name is a nod to the genus name.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	materials_examined	Type series. BRAZIL, Espirito Santo State: ♂ holotype (MNRJ 9076) ,, Domingos Martins, Pedra Azul, Fazenda Firmino Bellon & Lurdinha Bernabé Bellon, (- 20.408169 °, - 41.022323 °), 1400 m, 27. vii. 2015, T. Bernabé, A. P. L. Giupponi, P. Omena & T. Souza leg ..; 5 ♂, 6 ♀, 2 juv. paratypes (MNRJ 9104), Domingos Martins, Braço do Sul, Faz. Ademir Bernabé (- 20.488619 °, - 41.055213 °) 1150 m, 28. vii. 2015, T. Bernabé, A. P. L. Giupponi, A. B. Kury, P. Omena & T. Souza leg.; 1 ♂ paratype (MNRJ 18148), Santa Teresa, REBIO Augusto Ruschi, Roda d’Água, (19 ° 54.575 ’ S, 40 ° 32.531 ’ W), 26. viii. 2006, E. Soeiro leg.; 1 ♀ paratype (MNRJ 18464), Santa Teresa, REBIO Augusto Ruschi, Tracomal, 7. ix. 2006, T. Bernabé leg.; 1 ♂, 1 ♀ paratypes (MNRJ 18621), Santa Teresa, REBIO Augusto Ruschi, Preguiça, (19 ° 54.640 ’ S, 40 ° 32.500 ’ W), 25. viii. 2006, T. Bernabé leg.; 1 ♂, 1 ♀ paratypes (MNRJ 18643), same data; 1 ♂ paratype (MNRJ 18669), ES, Santa Teresa, REBIO Augusto Ruschi, 26. viii. 2006, T. Bernabé, D. Mota, T. Souza & E. Soeiro leg.; 1 ♂ paratype (MNRJ 17958), Santa Teresa, REBIO Augusto Ruschi, Roda d’Água, (19 ° 54.575 ’ S, 40 ° 32.531 ’ W), 26. viii. 2006, D. Mota leg.; 1 ♂ paratype (MNRJ 17975), same loc. and date, T. Souza leg,; 1 ♀ paratype (MZSP 73441), Santa Teresa, REBIO Augusto Ruschi, Trilha da Preguiça (S 19 ° 54 ’ 50,8 ” W 40 ° 33 ’ 8 ”), 30. iii. 2015, A. R. Benedetti leg,; 1 ♀ paratype (MZSP 73446), same locality, 30. iii. 2015, A. A. Nogueira leg.; 1 ♂ paratype (MZSP 73447), same data; 1 ♂ paratype (MZSP 73442), Santa Teresa, REBIO Augusto Ruschi, Trilha da Cachoeira (S 19 ° 54 ’ 50.8 ” W 40 ° 33 ’ 8 ”), 30. iii. 2015, A. A. Nogueira leg; 1 ♀ paratype (MZSP 73445), same locality, 30. iii. 2015, M. P. Silva leg; 1 ♀ paratype (MZSP 73444), same locality, 02. iv. 2015, M. P. Silva leg.; 1 ♀ paratype (MZSP 73443), Santa Teresa, REBIO Augusto Ruschi, Trilha da Roda d’Água (S 19 ° 53 ’ 40,1 ” S W 40 ° 32 ’ 35,6 ”), 03. iv. 2015, A. A. Nogueira leg.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	diagnosis	Diagnosis. Body color in both sexes dark brown, almost black (Figs. 1 e, f). Ocularium with two small, spherical, pale yellow tubercles that contrast with background (Fig. 27 b). Scutal area III armed with small, contrasting, pale yellow, paramedian, rounded tubercles (Fig. 27 b). Free tergites I – III unarmed in both sexes (Figs. 1 e, f). Femur IV of male strongly curved to the midline at proximal fourth, armed at least with two strong retrolateral spines; RD 6 of male Fe IV strongly curved (Fig. 27 a). Glans with a circular wall around stylus (Figs. 29 a, c, d, e).	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	distribution	Distribution. Brazil, Espírito Santo State, Domingos Martins; Santa Teresa.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
44270927FFB257656C8EFAEAFC5F9CC9.taxon	description	Description of male holotype. Measurements: CL = 3.3, CW = 4.8, AL = 5.9, AW = 9.4. Pp = 9.0. Legs, I: Fe = 5.7, Ti = 3.8, Mt = 5.8; II: Fe = 9.7, Ti = 8.0, Mt = 10.1; III: Fe = 7.8, Ti = 5.2, Mt = 8.1; IV: Fe = 12.0, Ti = 7.9, Mt = 12.4. Dorsum. Robust animal, with strong legs. Outline of DS gamma pyriform in dorsal view, quite flattened, so that Cx IV is much thicker than scutum in lateral view. Mesotergum divided into four areas, none entirely separated, of which areas III and IV are widely fused to each other. Posterior border of DS with trapezoid concavity. Carapace mid-area pentagonal, with narrow and low ocularium. Ocularium armed with a pair of paramedian small spherical tubercles. Frontal hump low, armed with a pair of small granules. Mesotergum divided into 4 scutal areas, although all of them show fusion on the sides due to obliteration of grooves. Areas III and IV moreover are extensively fused to each other. Area I long, divided into halves, broadly joined. Area II projected a little into area I. Area III strongly arched frontwards and only moderately encased into area II. Carapace smooth and unarmed, area I with a transverse row of three granules on each side; area II with a similar row of four granules, without any especially developed pair in the middle. Areas III-IV with a row of a few granules plus a paramedian pair of larger, rounded, capitate tubercles, forming a transversal line on area IV. Area V with a transverse row of a few granules. Lateral margins of scutum with a row of minute tubercles at areas I to IV. Each free tergite with a row of minute granules, free tergites I – III unarmed. Posterior border of dorsal scutum with elongate trapezoid concavity. Two ozopores, first slit-like, second rounded. Venter. Cx I free, the other connected to the adjacent ones by tubercular bridges. Cx IV much larger than the other coxae combined. Stigmatic area posterior border deeply concave. Free tergites each with transverse row of granules. Stigmatic area with very long axis and large, elliptical slanted stigmata. Pedipalps. Moderately elongate, shorter than body. Tr with thin neck, 1 + 3 setiferous tubercles dorsal and ventral. Fe with a row of three extremely weak ventral setiferous tubercles and much stronger meso-distal spine. Pa curved, unarmed. Ti and Ta flattened ventrally. Spination of Ti: mesal IiIi, ectal IiIi (last two geminated). Spination of Ta: mesal IiIi, ectal IiIi. Legs. All legs slender, only leg IV with considerable armature. Fe I – II straight, III – IV arched. Femur III with multiple rows of small acuminate tubercles. Tr IV with prodorsal, median, conical apophysis which connects with apophysis of Cx IV and immense, retrolateral, distal, conical apophysis. Fe IV: elongate, curved to the middle at 1 / 3 length; uniformly curved to dorsal. Proventral: row of numerous, small, acuminate tubercles all along the femur length. Retroventral: row of numerous, small, acuminate tubercles, plus four larger tubercles distally. Retrodorsal: only four spines, with RD 5 and RD 10 the largest. Dorsal: row of small tubercles, with the five distalmost much larger. Prodorsal: row of numerous small acuminate tubercles all along the femur length. Pa IV with many rows of tubercles, Ti IV as well, but tubercles are coarser. Mt IV smooth, thin and slender. Tarsal counts: 6 (3) - 6 (3) / 10 (3) - 10 (3) / 7 - 7 / 10 - 10. Color (in 75 % ethanol): Body and appendages background Grayish Brown (61), lighter on scutum and tergites. Leg IV considerably darker starting at apical portion of Cx. Venter Light Brown (57), with unobtrusive, lighter, honeycombed reticulation. In vivo it is considerably darker, with lighter granules more contrasting. Arthrodial membrane between Cx / Tr IV Very Light Blue (180); between free tergites Dark Grayish Blue (187). Male genitalia (Figs. 29 a – e). VP roughly T-shaped, with basal lobes narrow, and distal border with a wide parabolic cleft. Large latero-distal flange of VP projected, forming a pair of triangular ears. Field of type 1 microsetae occupying most of VP with mid-ventral part extending into podium. Basal lobe as a large square projecting dorsally from VP. Macrosetae (MS) A and B well-developed, forming an arch on latero-basal lobe, B the same size as A 1 – A 3. MS C 1 – C 3 elongate, straight basally, but helical distally. MS D 1 very small, inserted at 1 / 3 of the distance between C and A / B, E 1 – E 2 small, on flange. Flabellum subtriangular, with ventral half outline deeply serrated. Glans sac organized as an elaborate calyx, surrounding most of the stylus. Female (MNRJ 9104). The female is essentially similar to male in color, build, appendages, and armature of dorsal scutum. Main differences are in the weak development and reduced armature of leg IV and the absence of yellow tubercles on the patellae and tibiae. Tarsal counts: 6 (3) / 9 (3) / 7 / 8.	en	Kury, Adriano B., Bernabé, Tiago (2023): Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae). Zootaxa 5351 (4): 401-434, DOI: 10.11646/zootaxa.5351.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5351.4.1
