taxonID	type	description	language	source
265387929213D174FE848C09F3398E22.taxon	type_taxon	TYPE SPECIES. — Homeokotorella anterioporus n. gen., n. sp.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D174FE848C09F3398E22.taxon	description	E. — The generic name refers to the homeomorphous hooks of the tentacular armature of the type species.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D174FE848C09F3398E22.taxon	diagnosis	DIAGNOSIS. — Scolex elongated, craspedote. Four elongated bothria, without thickened rims, lateral and posterior bothrial margins not fused with peduncle. Bothrial pits absent; hamulate spinitriches along bothrial margins present. Pars bothrialis overlapping pars bulbosa, pars vaginalis shorter than pars bothrialis. Pars bulbosa with short bulbs. Retractor muscles originating at base of bulbs; prebulbar organs and gland-cells inside bulbs absent. Tentacle sheaths straight. Four elongated, slender tentacles, without basal swelling. Basal and metabasal tentacular armature homeoacanthous, homeomorphous; hooks solid. Characteristic basal armature absent, first rows of basal armature increasing in size. Strobila acraspedote, proglottids wider than long, euapolytic; cirrus unarmed, coiled; cirrus sac lateral and elongate, runs along anterior end of proglottid; genital atrium submarginal, located at anterior end of proglottid; seminal vesicles absent; testes numerous, distributed around central ovary, not reaching anterior to cirrus sac, in single layer; ovary positioned centrally; vitelline follicles circumcortical. Parasitic in dasyatid batoids.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D174FE848C09F3398E22.taxon	discussion	REMARKS Within the Tentaculariidae, unlike in Tentacularia Bosc, 1797 with the bothria entirely fused with the scolex peduncle and Nybelinia Poche, 1926, Heteronybelinia Palm, 1999, Mixonybelinia Palm, 1999 and Reimeriella Palm, Morales-Ávila, Galván-Magaña & Haseli, 2020 with a robust, muscular scolex and triangular bothria with fused margins, the new genus possesses elongate bothria with free lateral and posterior margins. Homeokotorella n. gen. therefore closely resembles the two monotypic genera Kotorella Euzet & Radujkovic, 1989 and Kotorelliella Palm & Beveridge, 2002 since the members of these three genera possess an elongate scolex, four bothria each with free lateral and posterior margins, a short pars bulbosa, and a homeoacanthous metabasal armature. Nonetheless, the homeomorphous, rather than heteromorphous hooks of the metabasal and basal tentacular armature easily distinguish it from Kotorella and Kotorelliella. The erection of this genus is consistent with the taxonomy within Tentaculariidae, where each genus possesses its own armature pattern.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	description	(Fig. 1 A-F) urn: lsid: zoobank. org: act: 39 F 66909 - 0486 - 458 E-BB 5 A-E 8 AC 05510 A 9 A	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	materials_examined	TYPE MATERIAL. — Holotype. Malaysia • 1 specimen; Malaysian Borneo, Sabah, off Semporna (fish market); 04 ° 28 ’ 44.09 ” N, 118 ° 37 ’ 00.57 ” E; from stomach of Taeniura lymma 1 (sensu Naylor et al. 2012) (Myliobatiformes, Dasyatidae) (BO- 86); LRP 10983. Paratypes. Malaysia • 1 specimen; same data as for holotype; LRP 4379 • 6 specimens in 5 slides; Malaysian Borneo, Sabah, off Pulau Mabul; 04 ° 14 ’ 44.02 ” N, 118 ° 37 ’ 53.32 ” E; same host as for holotype; LRP 4380, 10984; ZMB E. 7744 - 46.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	etymology	ETYMOLOGY. — The specific name refers to the anterior position of the cirrus sac in this species.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	description	DESCRIPTION [Based on whole mounts of 4 specimens.] Worms 7800 - 7875 (7837, N = 2) long, with 37 - 40 (N = 2) proglottids; scolex slender, craspedote (Fig. 1 A, B), 724 - 781 (739, N = 4) long. Scolex width 290 - 419 (340, N = 4) at level of pars bothrialis, 282 - 378 (320, N = 4) at level of pars vaginalis, 314 - 338 (324, N = 4) at level of pars bulbosa. Bothria 4 in number, elongate, with free lateral and posterior margins (Fig. 1 B). Hamulate spinitriches present along bothrial borders. Pars bothrialis 451 - 523 (497, N = 4) long, slightly overlapping pars bulbosa; pars vaginalis shorter than pars bothrialis, 402 - 499 (441, N = 4) long; tentacle sheaths straight, 20 - 30 (26, N = 2, n = 4) in diameter; muscular ring around basal part of tentacle sheath present. Pars bulbosa 160 - 192 (179, N = 4) long; prebulbar organs and gland cells inside bulbs absent; bulbs 152 - 188 (172, N = 3, n = 11) long, 64 - 80 (72, N = 3, n = 11) wide, bulb width: length ratio 1.0: 2.1 - 2.7 (2.4, N = 3, n = 11); retractor muscles originate at posterior extremity of bulbs; pars post-bulbosa absent. Velum long, 104 - 160 (130, N = 4). Scolex ratio (pars bulbosa: pars bothrialis: pars vaginalis) 1.0: 2.4 - 3.3: 2.1 - 2.8 (1.0: 2.8: 2.5, N = 4). TYMOLOGY Tentacles elongate and slender, 480 - 500 (490, N = 1, n = 2) long; basal swelling absent; tentacle width 20 (N = 2, n = 2) at level of basal region, 24 (N = 2, n = 2) at level of metabasal region, and 20 (N = 1, n = 1) at level of distal region of tentacle. Hooks solid, tentacles with 27 - 31 (29, N = 1, n = 2) rows of hooks. Tentacular armature homeoacanthous, homeomorphous (Fig. 1 C, D). First four to five rows of basal hooks increasing in size (Fig. 1 D), hooks falcate, 4.5 - 10 (7.0, N = 3, n = 3) long, base 3 - 4 (4, N = 2, n = 3) long, a single slender uncinate hook with anterior extension of base seen. Metabasal armature with falcate hooks (Fig. 1 C), 12 - 14 (13, N = 3, n = 7) long, base 4 - 5 (5, N = 2, n = 3) long; distal region of tentacle with falcate hooks, 12 (N = 1, n = 5) long, base 4 - 5 (5, N = 1, n = 5) long. Number of hooks per half spiral row in basal and metabasal region of tentacle 5 - 6 (N = 1, n = 2), 6 (N = 1, n = 1) respectively. Proglottids acraspedote (Fig. 1 E, F), euapolytic; immature and mature proglottids wider than long. Mature proglottids 1 - 5 (3, N = 2) in number, 375 - 445 (410, N = 2, n = 6) long, with maximum width of 525 - 650 (572, N = 2, n = 6); gravid proglottids not observed. Genital pore submarginal in mature proglottids, close to anterior end of proglottids. Cirrus sac elongate, running along anterior end of each proglottid, 44 - 52 (47, N = 1, n = 4) long, 160 - 192 (179, N = 1, n = 4) wide; cirrus sac length: width ratio 1.0: 3.5 - 4.4 (3.8, N = 1, n = 4). Cirrus unarmed, coiled. Seminal vesicles absent. Vas deferens tightly coiled, running anterior medially and posteriorly towards ovarian isthmus. Vagina runs sinuously from ovary towards cirrus sac, then turning laterally along cirrus sac, opening to genital atrium posterior to cirrus sac (Fig. 1 F). Testes round to oval, in a single layer, not reaching anterior to cirrus sac, 40 - 52 (44, N = 2, n = 6) long in diameter, 48 (n = 1) in number (Fig. 1 E). Ovary bilobed, located in centre of proglottid, ovarian lobes 160 - 200 (178, N = 1, n = 4) long, 72 - 92 (83, N = 1, n = 4) wide. Mehlis’ gland posterior to ovarian isthmus, 40 - 52 (46, N = 1, n = 2) in diameter. Vitelline follicles circumcortical (Fig. 1 F), round to oval shaped, 20 - 40 (30, N = 1, n = 5) in diameter.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	etymology	TYMOLOGY	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929213D176FC128E48F2CE897C.taxon	discussion	REMARKS The specimens from the host described as Taeniura lymma 1 are very similar to one another and show only little variability in scolex size, hook sizes and proglottid features. The closely related species K. pronosoma and Ko. jonesi differ from H. anterioporus n. gen., n. sp. in their scolex proportions and bulb ratios (see Palm & Beveridge 2002; Palm 2004). The tentacular armature of these two species, K. pronosoma and Ko. jonesi, is homeoacanthous, heteromorphous in the metabasal and / or basal armature. While K. pronosoma always shows stout, tightly packed hooks with typical, broad diamond-shaped bases on the bothrial tentacle surface, Ko. jonesi has uncinate hooks with an anterior extension of the base on the bothrial and slender spiniform hooks on the antibothrial surface. Homeokotorella anterioporus n. gen., n. sp. displays slender falcate hooks without an anterior extension of the base, which are of similar shape on all tentacle surfaces.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	description	(Figs 2 A-D; 3 A-G) urn: lsid: zoobank. org: act: 989 AEFB 5 - E 48 C- 4 A 3 E-AF 49 - 372588089233	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	materials_examined	TYPE MATERIAL. — Holotype. Malaysia • 1 specimen; from stomach of Lamiopsis tephrodes (Fowler) (Carcharhiniformes, Carcharhinidae) (BO- 74); Malaysian Borneo, Sarawak, off Mukah, 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 11007. Paratypes. Malaysia • 4 specimens; same data as for holotype; LRP 11008, ZMB E. 7719 - 21. Indonesia • 2 specimens (plerocercoids); from stomach wall of Trichiurus lepturus (Scombriformes, Trichiuridae); South Java, off Pelabuhan Ratu; 06 ° 59 ’ 13.9 ” S, 106 ° 32 ’ 37.4 ” E; MNHN-HEL 1926, 1927.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	description	VOUCHER MATERIAL. — Scolex (plerocercoid) of N. pseudafricana n. sp. (voucher specimen) from the stomach wall of Gempylus serpens Cuvier, Pelabuhan Ratu, Indonesia (Jakob & Palm 2006), prepared for SEM. ADDITIONAL SPECIMENS STUDIED. — Several specimens of N. africana (ZMB E. 7722 - 25, 5 slides) from Mullus surmuletus Linnaeus off Kalâat el-Andalous, Tunisia.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	etymology	ETYMOLOGY. — The specific name is related to the similarity of the new species to N. africana.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	description	DESCRIPTION [Based on whole mounts of 3 specimens; plerocercoid from Gempylus serpens observed with SEM.] Worms 28 875 - 48 500 (37 283, N = 3) long, with 160 - 246 (N = 2) proglottids; scolex compact (Figs 2 A; 3 A), craspedote (Fig. 2 A), 555 - 724 (628, N = 3) long. Scolex width 427 - 451 (440, N = 3) at level of pars bothrialis, 427 - 451 (440, N = 3) at level of pars vaginalis, 288 - 338 (319, N = 3) at level of pars bulbosa. Bothria 4 in number, sessile and elongate. Lineate spinitriches along bothrial borders present. Scolex and bothrial surface covered with capilliform (3 - 5) and papilliform (1) filitriches. Pars bothrialis 282 - 386 (346, N = 3) long, overlapping pars bulbosa; pars vaginalis shorter than pars bothrialis, 185 - 370 (252, N = 3) long; tentacle sheaths straight anteriorly, each with a single coil immediately anterior to bulb, 36 - 52 (47, N = 3, n = 10) in diameter; muscular ring around basal part of tentacle sheath not seen. Pars bulbosa 217 - 266 (241, N = 3) long; prebulbar organs and gland cells inside bulbs absent; bulbs 240 - 252 (246, N = 3, n = 6) long, 96 - 104 (99, N = 3, n = 6) wide, bulb width: length ratio 1.0: 2.3 - 2.5 (2.5, N = 3, n = 6); retractor muscles originating at posterior extremity of bulbs; pars post-bulbosa absent. Velum elongate, straight or scalloped, 80 - 113 (99, N = 3) long. Scolex ratio (pars vaginalis: pars bothrialis: pars bulbosa) 1.0: 1.0 - 2.0: 0.6 - 1.4 (1.0: 1.5: 1.1, N = 3). Tentacles elongate, 720 - 1000 (900, N = 1, n = 3) long with c. 40 rows of hooks; basal swelling absent; tentacle width 32 - 36 (N = 1, n = 2) at level of basal region, 40 - 42 (N = 1, n = 2) at level of metabasal region. Hooks solid. Tentacular armature homeoacanthous, homeomorphous (Fig. 2 B, C). Metabasal armature differing distinctly from basal armature. 2 - 3 rows of basal armature with uncinate hooks with distinct anterior extension of base, 10 - 12 (11, N = 2, n = 6) long, base 8 - 10 (9, N = 2, n = 5) long; metabasal hooks larger, slender falcate with small base and regularly curved tip, 20 - 25 long (22, N = 3, n = 14); base 6 - 8 (7, N = 3, n = 15) long; apical hooks of not fully everted tentacles 22 (N = 1) long, base 7 - 8 (7, N = 1, n = 2) long. Number of hooks per half spiral row in basal and metabasal region of tentacle 5 - 6 (5, N = 2, n = 2) and 6 - 7 (6, N = 2, n = 3), respectively. Proglottids acraspedote (Fig. 2 D), apolytic; immature and mature proglottids wider than long. Mature proglottids 2 - 3 (N = 2) in number, 400 - 1125 (690, N = 2, n = 5) long, 675 - 2500 (999, N = 2, n = 4) wide. Gravid proglottids 1000 - 1375 (N = 1, n = 2) long, 2625 wide (N = 1, n = 1). In mature proglottids, genital pore in anterior third of proglottid; genital pores alternate irregularly. Cirrus sac submarginal, elongate, 56 - 72 (64, N = 1, n = 2) long, 168 - 220 (194, N = 1, n = 2) wide, not reaching anterior end of proglottid; cirrus sac length: width ratio 1.0: 2.3 - 3.9 (3.1, N = 1, n = 2). Cirrus unarmed, coiled. Seminal vesicles absent. Vas deferens tightly coiled, running anteriorly medially and then posteriorly towards ovarian isthmus. Testes oval in shape, around 40 in diameter, arranged in single layer; testes number 70 - 75 (73, N = 2, n = 2) per proglottid, 9 testes anterior to cirrus sac (n = 1). Ovary bilobed, located in centre of proglottid, 68 - 189 (106, N = 2, n = 4) long, 140 - 280 (189, N = 2, n = 4) wide. Vitelline follicles circumcortical, round to oval in shape, 24 - 44 (33, N = 2, n = 5) long; 12 - 24 (18, N = 2, n = 5) wide.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929211D178FC6B89CEF5A38F43.taxon	discussion	REMARKS The specimens presented here as a new species are those identified by Schaeffner & Beveridge (2014) as N. africana from Lamiopsis tephrodes (BO- 74). Nybelinia pseudafricana n. sp. most closely resembles N. africana. Both species can be characterized by the presence of several rows of uncinate hooks at the base of the tentacular armature and the presence of falcate hooks along the tentacle. Nonetheless, the new species is distinguished from N. africana by the length of the metabasal hooks (20 - 25 vs 12.5 - 16) and the tentacle length (720 - 1000 vs 200) (see Palm 2004). In this respect, several Tunisian specimens of N. africana isolated from Mullus surmuletus caught off Kalâat el-Andalous, were also used for comparison with their characteristics as follows: scolex 632 - 828 (751, N = 12) long; bulbs 186 - 270 (213, N = 9, n = 9) long, 72 - 96 (86, N = 9, n = 9) wide, bulb width: length ratio 1.0: 1.9 - 3.0 (2.5, N = 9, n = 9); velum 240 - 345 (277, N = 8 n = 8) long; number of rows of hooks 17 - 19 (18, N = 5); metabasal hooks 12 - 17 (15, N = 5, n = 5) long, base 3 - 6 (5, N = 5, n = 5) long; tentacles 258 - 321 (294, N = 4, n = 4) long. Nybelinia pseudafricana n. sp., considering the Tunisian specimens, is also different from N. africana in the number of rows of hooks (40 vs 17 - 19). It is worth mentioning that this feature was not mentioned in the original description of N. africana (see Palm 2004). Regardless of variation in scolex length as can be seen in species of Nybelinia (see Palm 2004), it is of note that the specimens earlier identified as N. africana in Indonesia, including the adult specimens isolated from Alopias superciliosus Lowe as well as Carcharhinus sp. and the plerocercoids isolated from Alepisaurus ferox Lowe, Brama dussumieri Cuvier, Conger cinereus Rüppell, Coryphaena hippurus L., Gempylus serpens Cuvier, Thyrsitoides marleyi Fowler, and Trichiurus lepturus (see Palm 2000, 2004; Jakob & Palm 2006), are re-identified as N. pseudafricana n. sp., and that this species is very common along the southern Java coast, Indonesia. Likewise, the specimens reported as N. africana off Mozambique by Palm et al. (1997) are re-identified as N. pseudafricana n. sp. based on the scolex measurements and the length of their metabasal hooks (20 - 32). Thus, whereas N. pseudafricana n. sp. occurs in the Indo-Pacific region, the distribution of N. africana still remains mainly around Western Africa and in the Mediterranean Sea. Examination of the scolex with SEM identified several rows of tiny pores (Fig. 3 B, D), each parallel to one hook row, and serially placed along the tentacle. This is a novel feature, which had not earlier been described from trypanorhynchs. However, pores have been earlier reported already from the hooks of Molicola horridus (Goodsir, 1841) Dollfus, 1935 and Gymnorhynchus isuri Robinson, 1959 (see Knoff et al. 2004, 2007).	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921FD17AFC0E8FA8F7828CC0.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 5 specimens; from Sphyrna cf. lewini (Griffith & Smith) (BO- 69) (Carcharhiniformes: Sphyrnidae); Malaysian Borneo, off Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 4367 - 4371 • 1 specimen; same data; ZMB E. 7726 • 1 specimen; from Sphyrna lewini 1 (sensu Naylor et al. 2012) (BO- 60); off Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 11005. Japan • holotype and 1 paratype; from S. zygaena off Nagasaki; off Nagasaki; SY 7201. South Australia • 3 specimens; from S. zygaena off Goolwa; SAM; AHC 24958. Malaysia • 1 specimen; from Urogymnus polylepis (Bleeker) (BO- 108) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Kampung Abai, Kinabatangan River, Sabah; 05 ° 41 ’ 10.81 ” N, 118 ° 23 ’ 08.35 ” E; LRP 11006. New Caledonia • 7 plerocercoids; from Saurida undosquamis (Richardson) (Aulopiformes: Synodontidae); off New Caledonia; ZMB E. 7729 - 32. REMARKS Nybelinia sphyrnae was described by Yamaguti (1952) from Sphyrna zygaena off Nagasaki, Japan. Beveridge & Campbell (1996) reported three specimens of this species from S. zygaena from Goolwa, South Australia, providing additional drawings of the scolex and mature proglottid and the first drawing of the tentacle with its hooks. Palm (2004) also reported this species from S. zygaena from Goolwa, South Australia, in his parasite-host checklist. Thereafter, Schaeffner & Beveridge (2014), during a large-scale study focusing on the parasite diversity of elasmobranchs from Malaysian and Indonesian Borneo, reported specimens of N. sphyrnae from S. lewini off Malaysian Borneo, the hosts having been registered in the Global Cestode Database as Sphyrna lewini 1 (BO- 60) and S. cf. lewini (BO- 69). In the present study, in addition to examining the specimens of N. sphyrnae earlier identified by Schaeffner & Beveridge (2014) from S. lewini 1 and S. cf. lewini (BO- 60, 69), we also measured the holotype as well as one paratype from Japan, some specimens from New Caledonia, and those South Australian specimens mentioned in the parasite-host checklist of Palm (2004) to clarify the range of measurements for N. sphyrnae (see Table 1). Furthermore, on the basis of recent examinations and the measurements at hand a proposal for a synonymy of the two species N. sphyrnae and N. jayapaulazariahi is made, both of which have a homeoacanthous, homeomorphous tentacular armature with uncinate hooks, which increase in size towards the metabasal part of the tentacle. In both species, the pars bothrialis is larger than the pars bulbosa and the retractor muscles originate at the base of the bulbs. In Table 1, the measurements of N. jayapaulazariahi from different studies are compared with those of the types and vouchers of N. sphyrnae from different localities. The ranges of the taxonomically important characters of N. jayapaulazariahi overlap with those of N. sphyrnae. Regarding the variation seen for the bulb width to length ratio in the Malaysian Bornean and New Caledonian specimens of N. sphyrnae in comparison to other conspecifics, it is worth mentioning that in all the specimens of N. sphyrnae the values measured for the pars bulbosa as well as the length of bulbs are uniform, for which such differences have rarely been described in trypanorhynchs (see Schaeffner & Beveridge 2013 a), but a variation is seen in the width of the bulbs, which depends on the state of contraction during fixation. This situation can also be considered for the length of the tentacles, however, by re-measuring this character from figure 7 drawn by Reimer (1980), this value is 335 at maximum size and is very close to the lower bound of the range measured (350 - 400) by Yamaguti (1952). None of the taxonomic works carried out on N. jayapaulazariahi mentioned the number of the hook rows along the tentacle. However, about 30 rows can be counted at least from figure 7 of Reimer (1980), which is close to the number of the hook rows counted for the Malaysian Bornean and South Australian specimens of N. sphyrnae. Although Yamaguti (1952) described 12 hook rows along the tentacle for the types, from figure 83 drawn by him, at least 24 rows can be counted. By examining the type specimens, we could count 32 rows of hooks. According to these findings we consider N. jayapaulazariahi, the descriptions of which were based on the larval forms, a new synonym for N. sphyrnae.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921DD17CFC4C8BCCF41C8F62.taxon	description	(Fig. 4 A, B)	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921DD17CFC4C8BCCF41C8F62.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 5 specimens; from Rhinoptera jayakari Boulenger (BO- 83) (Myliobatiformes: Rhinopteridae); Malaysian Borneo, off Sabah, Semporna fish market; 04 ° 28 ’ 44.09 ” N, 118 ° 37 ’ 00.57 ” E; LRP 4375 - 4378, 11002 • 1 specimen; same data; ZMB E. 7713 • 1 specimen; from Maculabatis pastinacoides (Bleeker) (BO- 76) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Sabah, Kampung Tetabuan; 06 ° 01 ’ 10.32 ” N, 117 ° 42 ’ 14.76 ” E; ZMB E. 7712.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921DD17CFC4C8BCCF41C8F62.taxon	description	MEASUREMENTS Long worms, 7850 - 11 776 (10 154, N = 4) long. Scolex 2411 - 3271 (2867, N = 5) long; scolex width 841 - 1121 (979, N = 4, n = 11) at level of pars bothrialis, 518 - 1147 (816, N = 5, n = 1) at level of pars vaginalis, 744 - 1128 (947, N = 5, n = 10) at level of pars bulbosa. Bothria 1287 - 1415 (1358, N = 3, n = 5) long. Pars bothrialis 1342 - 1433 (1391, N = 5) long; pars vaginalis 854 - 1464 (1174, N = 5) long; tentacle sheaths 75 - 121 (92, N = 5, n = 13) in diameter. Pars bulbosa 884 - 1013 (947, N = 5) long; bulbs 793 - 982 (900, N = 5, n = 10) long, 183 - 232 (202, N = 4, n = 8) wide, bulb width: length ratio 1.0: 3.7 - 5.4 (4.5, N = 4, n = 8); pars post-bulbosa present, 42 - 67 (54, N = 5) long. Velum very lobate in its posterior margin (Fig. 4 A), 43 - 537 (400, N = 5, n = 11) long. Scolex ratio (pars bulbosa: pars bothrialis: pars vaginalis) 1.0: 1.4 - 1.6: 1.0 - 1.5 (1.0: 1.5: 1.2, N = 5). Tentacles 1726 - 2025 (1871, N = 3, n = 3) long with 37 - 43 (39, N = 2, n = 4) rows of hooks; tentacle width 51 - 97 (76, N = 5, n = 7) at level of basal region and 69 - 97 (80, N = 5, n = 8) at level of metabasal region. Metabasal armature homeoacanthous, homeomorphous (Fig. 4 B); characteristic basal armature absent. Basal hooks 8 - 21 (15, N = 5, n = 10) long, base 5 - 12 (9, N = 5, n = 10) long. Metabasal hooks 30 - 43 (36, N = 5, n = 19) long, base 10 - 18 (14, N = 5, n = 19) long. Number of hooks per half spiral row in basal and metabasal region of tentacle 7 (N = 3, n = 5). Anterior-most immature proglottid 12 - 18 (15, N = 4, n = 4) long, 640 - 1006 (809, N = 4, n = 4) wide; posterior-most proglottid 165 - 317 (239, N = 4, n = 4) long, 598 - 769 (683, N = 4, n = 4) wide.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921DD17CFC4C8BCCF41C8F62.taxon	discussion	REMARKS Historically, Shipley & Hornell (1906) described this species as Tetrarhynchus equidentatus Shipley & Hornell, 1906 from Brevitrygon walga (Müller & Henle) (therein referred as Himantura walga) off Sri Lanka. After Pintner (1927) redescribed the type, it was transferred to Nybelinia by Dollfus (1930). Wenchuan et al. (1995) reported N. aequidentata from Hemitrygon akajei (Müller & Henle) (therein referred as Dasyatis akajei) from the Taiwan Strait (see also Wenchuan 2007). Palm (1999) redescribed a plerocercoid of this species from Lepturacanthus savala (Cuvier), off Sagar Island, Bay of Bengal. Palm & Beveridge (2002) presented measurements of a postlarva as N. aequidentata isolated from Dendrochirus zebra (Cuvier) off New Caledonia (see also Beveridge et al. 2014). Nonetheless, Palm (2004) stated that the specimens identified and described as N. aequidentata by Palm & Beveridge (2002) were synonymous with N. syngenes (Pintner, 1928) Dollfus, 1930, based on the hook sizes and the length of the bulbs. In the present study, five specimens were collected from Rhinoptera jayakari (BO- 83) off Malaysian Borneo. Rhinoptera jayakari is a new host record for N. aequidentata and a new locality record is also established for this parasite off Malaysian Borneo. Although the examined material was strobilate, the internal organs were invisible and we missed the opportunity to describe the proglottid anatomy of this species.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921DD17CFC4C8BCCF41C8F62.taxon	description	UNIDENTIFIED MATERIAL The following material was examined but was unable to be assigned to a species of Nybelinia: 2 specimens (ZMB E. 7714, 7715) from Chiloscyllium punctatum Müller & Henle (BO- 282) (Orectolobiformes: Hemiscylliidae) off Mukah, Sarawak, Malaysian Borneo (02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E); 1 specimen (LRP 11003) from Carcharhinus sealei (Pietschmann) (BO- 55) (Carcharhiniformes: Carcharhinidae) off Mukah, Sarawak, Malaysian Borneo (02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E); 1 specimen (LRP 11004) from Carcharhinus sorrah (Müller & Henle) (BO- 70) off Mukah, Sarawak, Malaysian Borneo (02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E).	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921BD17CFF3E8FC8F5B58E81.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 7 specimens; from Carcharhinus cf. leucas (Müller & Henle) (BO- 75, 97, 101) (Carcharhiniformes: Carcharhinidae); Malaysian Borneo, off Sabah,; 06 ° 01 ’ 10.32 ” N, 117 ° 42 ’ 14.76 ” E; LRP 10976 - 10982 • 2 specimens; same data; ZMB E. 7717, 7718 • 4 specimens; from Carcharhinus cf. limbatus (Müller & Henle) (BO- 54, 58); Malaysian Borneo, off Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 4372 - 4374, 10975 • 1 specimen; same data; ZMB E. 7716.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921BD17CFF3E8FC8F5B58E81.taxon	description	MEASUREMENTS Long worms, 25 140 - 40 654 (30 112, N = 5) long with 188 - 206 (197, N = 2) proglottids. Scolex 787 - 1324 (1097, N = 5) long; scolex width 305 - 720 (585, N = 11, n = 11) at level of pars bothrialis, 348 - 549 (478, N = 11, n = 1) at level of pars vaginalis, 305 - 518 (423, N = 11, n = 11) at level of pars bulbosa. Bothria 396 - 567 (469, N = 11, n = 13) long. Pars bothrialis 384 - 622 (494, N = 11) long; pars vaginalis 342 - 518 (424, N = 11) long; tentacle sheaths 33 - 48 (43, N = 11, n = 11) in diameter. Pars bulbosa 348 - 451 (398, N = 11) long; bulbs 308 - 417 (350, N = 11, n = 27) long, 85 - 121 (103, N = 11, n = 27) wide, bulb width: length ratio 1.0: 2.9 - 4.3 (3.4, N = 11, n = 27); pars post-bulbosa present, 24 - 159 (72, N = 11) long. Velum 165 - 305 (218, N = 11, n = 11) long. Scolex ratio (pars bulbosa: pars bothrialis: pars vaginalis) 1.0: 0.9 - 1.5: 0.8 - 1.3 (1.0: 1.2: 1.1, N = 11). Tentacles 528 - 690 (595, N = 11, n = 14) long with 31 - 36 (33, N = 5, n = 5) rows of hooks; tentacle width 27 - 39 (33, N = 11, n = 11) at level of basal region and 27 - 36 (32, N = 11, n = 11) at level of metabasal region. Metabasal armature homeoacanthous, heteromorphous; characteristic basal armature absent. Basal uncinate hooks on bothrial tentacle surface 8 - 10 (9, N = 5, n = 5) long, base 10 - 12 (11, N = 5, n = 5) long; basal uncinate hooks on antibothrial tentacle surface 5 - 6 (6, N = 5, n = 5) long, base 7 - 8 (7, N = 5, n = 5) long; metabasal uncinate hooks on bothrial tentacle surface 14 - 15 (15, N = 5, n = 5) long, base 13 - 14 (14, N = 5, n = 5) long; metabasal uncinate hooks on antibothrial tentacle surface 10 - 12 (11, N = 5, n = 5) long, base 9 - 10 (9, N = 5, n = 5) long. Number of hooks per half spiral row in basal and metabasal region of tentacle 5 - 6 (N = 3, n = 5). Mature proglottids 66 - 78 (72, N = 2) in number, 152 - 518 (387, N = 8, n = 8) long, with maximum width of 281 - 793 (462, N = 8, n = 8). Cirrus sac 45 - 66 (52, N = 9, n = 9) long, 172 - 302 (236, N = 9, n = 9) wide; cirrus sac length: width ratio 1.0: 3.3 - 6.0 (4.6, N = 9, n = 9). Testes 21 - 67 (39, N = 11, n = 14) long, 16 - 43 (27, N = 11, n = 14) wide; testis number around 80 per proglottids (N = 1), 3 - 9 (6, N = 10, n = 10) testes anterior to cirrus sac. Ovary 120 - 135 (127, N = 2, n = 2) long, 52 - 75 (64, N = 2, n = 2) wide. Vitelline follicles 12 - 52 (25, N = 11, n = 11) long, 9 - 18 (13, N = 11, n = 11) wide.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921BD17CFF3E8FC8F5B58E81.taxon	discussion	REMARKS Schaeffner & Beveridge (2014) reported H. estigmena from C. limbatus (BO- 54, 58) off Malaysian Borneo. Heteronybelinia estigmena is a well described and a cosmopolitan tentaculariid species (see Palm 2004). The scolex measurements of the Malaysian Bornean specimens fall within the range of the measurements given by Palm (1999) and Palm & Beveridge (2002). This case is also seen for the measurements of the strobila.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921BD17CFC038D29F3B78CC0.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 2 specimens; from Carcharhinus falciformis (Müller & Henle, 1839); BO- 132, 133; (Carcharhiniformes: Carcharhinidae); Malaysian Borneo, off Sabah; 04 ° 14 ’ 44.02 ” N, 118 ° 37 ’ 53.32 ” E; LRP 11009, 11010.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
26538792921BD17CFC038D29F3B78CC0.taxon	discussion	REMARKS Tentacularia coryphaenae is well described and a cosmopolitan tentaculariid species (see Palm 2004). Schaeffner & Beveridge (2014) reported T. coryphaenae from Carcharhinus falciformis (BO- 133) from Malaysian Borneo. Another shark specimen of C. falciformis (BO- 132) was also infected with T. coryphaenae.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929218D17FFF598A0DF4F68D21.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 4 specimens; Pastinachus solocirostris Last, Manjaji & Yearsley (BO- 164, 256) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Mukah; Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; ZMB E. 7741 • 1 specimen; same data; LRP 10991 • 3 specimens; same data; Sematan, Sarawak; 01 ° 48 ’ 15.45 ” N, 109 ° 46 ’ 47.17 ” E; LRP 10988 - 10990 • 1 specimen, from Pateobatis uarnacoides (Bleeker) (BO- 118, 261) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Sabah; 06 ° 01 ’ 10.32 ” N, 117 ° 42 ’ 14.76 ” E; LRP 10987 • 1 specimen; same data; Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; ZMB E. 7742 • 1 specimen; same data; 10992 1 specimen; from Maculabatis gerrardi (Gray) (BO- 49) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 10986 • 2 specimens; from Himantura tutul Borsa, Durand, Shen, Alyza, Solihin & Berrebi (BO- 47) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Mukah, Sarawak; 02 ° 53 ’ 52.16 ” N, 112 ° 05 ’ 44.12 ” E; LRP 10985 • 3 specimens; same data; ZMB E. 7743.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929218D17FFF598A0DF4F68D21.taxon	discussion	REMARKS	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929218D17EFF7A8D89F799891C.taxon	materials_examined	MATERIAL EXAMINED. — Malaysia • 7 specimens; from Neotrygon orientalis Last, White & Serét (BO- 366, 368) (Myliobatiformes: Dasyatidae); Malaysian Borneo; LRP 10997 - 11001 • 1 specimen; same data; ZMB E. 7740 • 3 specimens; from Neotrygon orientalis (BO- 27); Malaysian Borneo; off Sematan, Sarawak; 01 ° 48 ’ 15.45 ” N, 109 ° 46 ’ 47.17 ” E; LRP 10993 - 1099 • 1 specimen; same data; ZMB E. 7738 • 1 specimen, LRP 10996 from Pateobatis cf. jenkinsii (Annandale) (BO- 339) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Sarawak; 02 ° 30 ’ 07.34 ” N, 110 ° 40 ’ 16.82 ” E • 1 specimen; same data; ZMB E. 7739 • 1 specimen, from Maculabatis macrura (BO- 93, 337) (Myliobatiformes: Dasyatidae); Malaysian Borneo, off Sarawak; 02 ° 30 ’ 07.34 ” N, 110 ° 40 ’ 16.82 ” E • 1 specimen; same data; Sabah, off Kudat; 06 ° 48 ’ 60.00 ” N, 116 ° 53 ’ 54.00 ” E; ZMB E. 7736, 7737. MEASUREMENT Long worms, 17 625 - 21 000 (19 312, N =) long with 46 - 69 (60, N = 3) proglottids. Scolex 1320 - 1666 (1446, N =) long; scolex width 330 - 427 (373, N = 3, n = 3) at level of pars bothrialis, 241 - 330 (274, N = 3, n =) at level of pars vaginalis, 322 - 402 (330, N = 3, n = 3) at level of pars bulbosa. Pars bothrialis 515 - 668 (604, N = 3) long; pars vaginalis 724 - 1046 (869, N = 3) long. Pars bulbosa 266 - 322 (290, N = 3) long; bulbs 240 - 304 (265, N = 3, n = 12) long, 84 - 108 (96, N = 3, n = 12) wide, bulb width: length ratio 1.0: 2.3 - 3.6 (2.8, N = 3, n = 12). Velum 250 - 314 (287, N = 3, n = 3) long. Scolex ratio (pars bulbosa: pars bothrialis: pars vaginalis) 1.0: 1.8 - 2.4: 2.7 - 3.2 (1.0: 2.1: 2.9, N = 3). Metabasal armature homeoacanthous, heteromorphous. Metabasal armature with uncinate hooks on bothrial surface, 14 - 16 (15, N =, n = 10) long, base 8 - 10 (9, N = 2, n = 10) long; antibothrial hooks of metabasal armature falcate, 14 - 15 (14, N = 2, n = 5) long, base 4 - 5 (4, N = 1, n = 5) long. Basal hooks heteroacanthous atypical. Bothrial hooks of basal armature 8 - 10 (9, N = 1, n = 5) long, base 6 - 8 (7, N = 1, n = 5) long; antibothrial hooks of basal armature 4 - 8 (6, N = 1, n = 5) long, base 1 - 2 (1, N = 1, n = 5) long. Mature proglottids wider than long, 175 - 275 (224, N = 1, n = 5) long, 650 - 700 (680, N = 1, n = 5) wide; fully gravid proglottids longer than wide, 750 - 1150 (920, N = 1, n = 5) long, 400 - 625 (510, N = 1, n = 5) wide, size increasing towards end of strobila. Testes 52 - 60 (57, N = 1, n = 10) in diameter, 35 - 50 (42, N = 1, n = 10) in number. Vitelline follicles 20 - 24 (22, N = 1, n = 5) in diameter.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
265387929218D17EFF7A8D89F799891C.taxon	discussion	REMARKS Palm & Beveridge (2002) erected the monotypic genus Kotorelliella to accommodate Ko. jonesi by describing a single plerocercoid from Taeniura lymma (Forsskål) off Heron Island, Queensland, Australia. Although the specimens of Malaysian Borneo were adult, because of the poor condition of the strobilae, the internal anatomy of the proglottids could not be easily seen and hence there was no opportunity to describe the proglottids. Schaeffner & Beveridge (2014) reported Ko. jonesi from Pateobatis jenkinsii (therein referred as Himantura jenkinsii) (BO- 335) and Pateobatis cf. jenkinsii (therein referred as Himantura jenkinsii) (BO- 339) from Malaysian Borneo. The host specimens coded as BO- 366 and BO- 368 have been identified morphologically as Neotrygon kuhlii but not registered in the Global Cestode Database. BO- 27 is a code for Ne. orientalis in the Global Cestode Database. The only species of Neotrygon known from Borneo is Ne. orientalis. Neotrygon kuhlii is now known only from the region of the Solomon Island (Last et al. 2016 b). The record cited from Schaeffner & Beveridge (2014) predates the revision of the host species by Last et al. (2016 b), but all of the records from Borneo are now referable to Ne. orientalis, not to Ne. kuhlii. Therefore, a new host record, Ne. orientalis, is established for Ko. jonesi. Compared to the holotype of Ko. jonesi, which is a plerocercoid, the adult specimens of the present study have a smaller scolex size (1320 - 1666 vs 1910). Other scolex features are also smaller. Such a situation was also reported for other trypanorhynchs, e. g. Progrillotia dasyatidis Beveridge, Neifar & Euzet, 2004 (see Beveridge et al. 2004; Marques et al. 2005). This may relate to the size and the physiological condition of the intermediate host, from which it is transmitted by predation into its definitive hosts.	en	Palm, Harry Wilhelm, Haseli, Mohammad (2023): Tentaculariids (Cestoda, Trypanorhyncha) of elasmobranchs from Malaysian Borneo. Zoosystema 45 (18): 513-529, DOI: 10.5252/zoosystema2023v45a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a18.pdf
