taxonID	type	description	language	source
03A487B62F23FFC7FF6CFA79FA6EFBCC.taxon	type_taxon	Type: — Eperua falcata Aublet (1775: 369).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F23FFC7FF6CFA79FA6EFBCC.taxon	description	Small to very large trees (2.0 – 50.0 m tall); trunk with bark lenticellate, fissured, striate, scaled, or smooth; branchlets glabrous, rarely puberulent, puberulous to strigulose in E. oleifera. Stipules 2, intrapetiolar, entirely joined, joined at the base, or free, pellucid-punctate, foliaceous or non-foliaceous, when foliaceous ovate-falcate, obovate-falcate, elliptic-falcate, falcate to semiorbicular, base attenuate, subcordate to cordate, when non-foliaceous ovate to lanceolate, base truncate, persistent to caducous. Leaves alternate and distichous, paripinnate, 1 - to 7 - jugate, usually glabrous, petioles, rachis and petiolules terete to canaliculate, glabrous, rarely puberulous in E. oleifera; blades coriaceous to chartaceous, glabrous, pellucid-punctate to epunctate, discolorous or not, equilateral, or inequilateral and falcate, elliptic, ovate to lanceolate, apex usually attenuate to acuminate, but acute, obtuse to rounded in two species, base symmetrical to asymmetrical, or oblique, usually lower pair base more rounded and upper pair base more cuneate, margin flat to revolute, secondary venation with one or two intramarginal veins, usually vein closer to the margin not continuous, main vein straight or curved in falcate leaflets, prominent to depressed on the adaxial side, 10 – 40 secondary veins, spacing uniform, angle uniform, intersecondary veins conspicuous, tertiary veins regular polygonal reticulate, conspicuous to inconspicuous, areoles smaller and concave or larger and plane. Inflorescences a raceme or panicle, terminal, axillary, sometimes ramiflorous or cauliflorous, short and erect or long and pendulous, glabrous or with indumentum, lateral racemes alternate, distichous or spirally arranged, peduncle sessile up to 4.0 cm long in short and erect inflorescences, or up to 3.0 m long in long and pendulous inflorescences; one bract at the base of the pedicel; one pair of bracteoles attached to different portions of the pedicels, imbricate, free, not enclosing the developed flower buds, apex gland present or absent. Flowers alternate and spirally arranged, pedicellate, hypanthium cup-shaped or tubular, inner wall with nectar-producing glands; sepals four, free, unequal, the outer ones larger, apex gland present or absent, inner surface glabrous, outer surface glabrous or with indumentum, inner sepals scarious marginally; petals five, adaxial petal superinvolute forming a tube or involute but not forming a tube, flabellate, oblate to broadly obovate, apex rounded to truncate, base attenuate to truncate, white, cream, greenish, lilac, purple, pink to red, glabrous or with indumentum in E. venosa, other four petals reduced to petalodia, up to 7.5 mm long; stamens 10, all fertile or up to 5 staminodial, inserted when the adaxial petal is tubular, exserted in the non-tubular ones, filaments heteromorphic, joined basally in a tube, or in a diadelphous sheath with dorsal one free, usually glabrous on the free part, anthers all equal or dimorphic; ovary with stipe inserted centrally in the hypanthium, glabrous or with indumentum, style usually longer than the largest filaments, glabrous, adaxial groove present, stigma obtuse to capitate, entire or 2 - lobed. Fruit legume, with stipe, equilateral or inequilateral and falcate, wood, flattened, glabrous or with indumentum, dehiscent, valves twisted after opening, seeds 1 - few, flattened.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F23FFC7FF6CFA79FA6EFBCC.taxon	etymology	Etymology: — Eperua is derived from the fruit shape of its type species, E. falcata. The term “ eperu ” is a Galibi name for the fruit of E. falcata (Aublet 1775), which translates to “ sabre ” in English. This refers to a curved-blade sword (Collins 2023), resembling the curved shape of the fruit.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	materials_examined	Type: — VENEZUELA. Amazonas: Municipio Autónomo Maroa, cuenca del río San Miguel, alto caño Mee, bana al sur de la serranía de Cariche, 106 m, 2 ˚ 44 ’ 14 ” N 66 ˚ 20 ’ 7 ” W, 18 September 2011, Romero G. A. et al. 4079 (holotype VEN not seen, isotypes GH not located, MO not seen, TFAV not seen). Treelet 2.0 – 5.0 m tall. Trunk not seen. Stipules free, non-foliaceous, caducous. Leaves 2 - jugate; petioles 1.7 – 4.0 cm long; rachis 2.0 – 4.5 cm long; petiolules 6.4 – 9.0 mm long; blades 8.2 – 14.0 × 3.7 – 6.6 cm, coriaceous, glabrous, pellucid-punctate, discolorous, slightly inequilateral, lower pair oblong, elliptic to narrowly elliptic, upper pair elliptic to obovate, apex apiculate, obtuse, rounded to emarginated, mucronate, base asymmetrical (upper pair) to symmetrical, lower pair base obtuse, upper pair base obtuse to cuneate, margin flat, secondary venation with one intramarginal vein, main vein slightly curved, depressed to prominent on the adaxial surface, tertiary veins conspicuous, areoles larger, concave. Inflorescences cauliflorous or ramiflorous, racemose or cauliflorous flowers, erect, strigulose, 1.5 – 11.0 cm long; bracts caducous, not seen; bracteoles 7.0 – 8.0 × 7.0 – 8.0 mm, ovate, apex gland absent, puberulent externally, caducous, attached to the lower portion of the pedicels; pedicel ca. 10 mm long, diameter not seen, not twisted, pubescent; buds not seen. Flowers: hypanthium 5.0 mm long, diameter not seen, cup-shaped, inequilateral, pubescent; sepal 2.3 – 2.7 × 0.8 – 0.9 cm, oblong to ovate, unequal, the outer ones larger, cucullate, apex gland present on the outer ones, purple, pubescent, inner sepals scarious marginally; adaxial petal 5.0 × 7.0 cm, flabellate, tubular, apex rounded, crisped, base attenuate, purple, abaxially glabrous, adaxially with two white, lanose structures near the margins; petalodia 6.0 – 7.0 × 2.6 – 3.5 mm; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, filaments 2.3 – 4.0 cm long, sheath inequilateral, glabrous, anthers 6.5 – 7.5 × 2.0 – 3.0 mm, rectangular; ovary 7.0 – 8.0 × 2.5 mm, oblong-obovate, sericeous, ovule not seen, stipe 2.0 – 3.0 mm long, glabrous, style ca. 3.5 cm long, stigma bilobate. Legumes not seen (description based on Romero & Aymard 2019 and selected materials). Phenology: — Flowering in September and November; fruiting in February.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	distribution	Distribution: — Venezuela (Amazonas). Habitat: — White sand shrubby communities in low Venezuelan bana, common in the ecotone bana - forest, on white sand soil, at 106 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	description	Conservation status: — Categorized as Data Deficient (DD) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022) due to the lack of access to its collections. Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	etymology	Etymology: — The specific epithet relates to its habitat in the banas (Romero & Aymard 2019). Vernacular names: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	description	Uses: — Unknown. Taxonomic notes: — E. banaensis and E. obtusata are distinct from other species of the genus due to their leaflets with rounded, obtuse, acute to emarginate apices. All other species in the genus have attenuate, acuminate to caudate apices. Morphologically, they share 2 - jugate leaves, short and erect inflorescence, tubular corolla, inserted stamens, and the same pattern of indumentum on reproductive organs. Among the species with tubular corolla, E. bananesis and E. obstusata are the only ones with the combination glabrous sheath and indumented ovary (Table 2). We recognize E banaensis as a treelet from low Amazonian campinarana or bana, while E. obtusata as a tall tree found in the tall campinarana. Additionally, E. banaensis differs from E. obtusata by the elliptic to obovate leaflets (vs. oblong to elliptic in E. obtusata), larger [7.0 – 8.0 × 7.0 – 8.0 mm] bracteoles (vs. smaller [ca. 2.0 × 1.0 – 2.0 mm] in E. obtusata), and longer [5.0 mm long] hypanthium (vs. shorter [2.0 – 3.0 mm long] in E. obtusata), petal with two white, lanose structures near the margins (vs. absent in E. obtusata), ovary twice the length [7.0 – 8.0 mm long] of the stipe [2.0 – 3.0 mm long] (vs. relatively the same length [5.0 mm long] as the stipe [4 mm long] in E. obtusata), glabrous stipe (vs. sericeous in E. obtusata), and bilobate stigma (vs. truncate in E. obtusata). Romero & Aymard (2019) referred the collection Colella M. et al. 1864 as E. obtusata, and did not mention Guanchez-Meza F. J. 2163. Here we recognized these collections also as E. banaensis for being treelets from the bana and having elliptic to obovate leaflets. In addition to the type collection, we recognized one more sample (Coomes D. 207) as E. obtusata. This collection differs from E. banaensis by being tall trees from the tall campinarana with elliptic to obtuse leaflets.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F24FFCBFF6CFB37FABCFECB.taxon	materials_examined	Selected specimens: — VENEZUELA. Amazonas, Casiaquire, rio Casiaquire, dos vueltas abajo de Capihuara, 06 February 1991, Collela M. et al. 1864 (K, NY). San Fernando de Atabapo, Zona 10, cuesta de arenisca de grano fino al pié del cerro Duida, 360 m elev., 3 ° 23 ’ 0 ” N, 65 ° 40 ’ 0 ” W, 10 November 1982, Guanchez-Meza F. J. 2163 (MO).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	description	(Figures 10, 17, 18)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	materials_examined	Type (designated by Cowan 1975): — BRAZIL. Pará: In sylvis inundatis ad insulam Marajo, May 1820, Martius C. F. P. s. n. (lectotype K [000555103] image!, isolectypes LE [00002232] image!, M [0215266] image!, M [0215267] image!, M [0215268] image!, M [0215270] image!). Tree 3.0 – 33.0 m tall. Trunk 18.0 – 31.8 cm in diameter, bark brown, striate. Stipules 1.1 – 14.2 (– 18.3) × 0.5 – 10.0 (– 13.0) mm, free, foliaceous or non-foliaceous, caducous to persistent. Leaves 2 (– 3) - jugate; petioles 0.6 – 3.1 (– 6.8) cm long; rachis (1.3 –) 2.0 – 6.4 (– 9.3) cm long; petiolules 4.4 – 12.3 mm long; blades (3.1 –) 6.4 – 17.6 (– 22.4) × 2.8 – 9.2 cm, coriaceous, glabrous, sparsely pellucid-punctate, sometimes pellucid-punctate or epunctate, slightly discolorous, equilateral, lower pair elliptic to ovate, rarely lanceolate, smaller, upper pair elliptic, apex attenuate to acuminate, sometimes acute, mucronate, base symmetrical, rarely slightly asymmetrical, with one pair of glands, lower pair base obtuse, rarely rounded, subcordate or cordate, upper pair base obtuse, rarely rounded or cuneate, margin flat, secondary venation with two intramarginal veins, vein closer to the margin not continuous, sometimes continuous, main vein straight, prominent near the base, sometimes depressed, on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal, rarely axillary or ramiflorous, panicle or raceme, erect, puberulous, trichomes white, 1.2 – 3.6 cm long, lateral racemes 0.9 – 2.6 cm long, alternate and spirally arranged or distichous; bracts 3.0 – 3.7 × 4.5 mm, oblate, apex gland absent, puberulous, trichomes white, caducous, rarely persistent; bracteoles 3.3 – 5.0 × 2.1 – 6.3 mm, oblate, sometimes broadly ovate, apex gland absent, puberulent externally, glabrous within, sometimes puberulent within, trichomes white, persistent, rarely caducous, attached to the lower middle portion of the pedicels; pedicel 3.3 – 9.5 mm long, 1.0 – 2.2 mm in diameter, not twisted, puberulent, trichomes white; buds 1.0 – 2.3 cm long, 0.6 – 1.7 cm in diameter, puberulous, trichomes white. Flowers: hypanthium 1.6 – 3.6 mm long, 2.5 – 7.6 mm in diameter, cup-shaped, inequilateral, puberulent to puberulous, trichomes white; sepal 1.3 – 3.0 × 0.5 – 1.3 cm, oblong to obovate, unequal, the outer ones more robust, the dorsal one larger, cucullate, apex gland absent, green, puberulent, inner sepals scarious marginally, trichomes white; adaxial petal 3.8 – 8.0 × 3.0 – 7.3 cm, flabellate, tubular, apex rounded, crisped, base attenuate, lilac with white nectar guide, pink, rose, purple, purple-red to red, glabrous; petalodia 0.8 – 7.3 × 0.4 – 1.4 mm; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 3.0 – 5.2 cm long, shorter filaments 1.7 – 3.1 cm long, sheath inequilateral, shorter side 5.5 – 17.3 mm long, longer side 8.9 – 24.0 mm long, sheath puberulous, free filament glabrous, trichomes white, anthers 3.2 – 5.0 × 0.7 – 2.3 mm, rectangular; ovary 5.3 – 11.8 × 1.8 – 5.1 mm, falcate, rarely obovate or oblong-obovate, glabrous, 1? ovule, stipe 2.8 – 9.5 (– 13.4) mm long, glabrous, style 2.4 – 3.8 cm long, stigma capitate, sometimes obtuse. Legumes 7.6 – 15.4 × 4.6 – 8.6 cm, stipe 1.0 – 2.5 mm long, oblong, apex obtuse, sometimes truncate, rarely rounded, apiculate, margin thickened, brown, rarely redbrown or dark-brown, with transversal veins, glabrous, foveate. Seeds 1 – 2 per fruit, 4.0 – 7.3 × 3.2 – 4.7 cm, ovate to inequilateral ovate, dark-brown. Phenology: — Flowering from March to August; fruiting throughout the year.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	distribution	Distribution: — Brazil (Pará and Amapá), and one collection from French Guiana without locality. Habitat: — Floodplain forests, mainly in igapó forest, but also in várzea forest. It is also found in the Amapá (Brazil) savannas in river-associated vegetation.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	description	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 129,115.212 km 2 and the estimated area of occupancy is 148.000 km 2. Occurrence in protected areas: — Floresta Nacional de Caxiuan „ (Brazil, Pará) and Floresta Nacional do Amapá (Brazil, Amapá) (Silva et al. 2015).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	etymology	Etymology: — The specific epithet relates to its 2 - jugate leaves. Vernacular names: — apá (Rabelo B. V. et al. 3307, Rosa N. A. 1288), muirapiranga (Oliveira E. 4929, Cruz E. D. 1123), muirapironga (Oliveira E. 4929).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	description	Uses: — it is described as potentially ornamental (Souza L. A. G. de 13 _ 97). Taxonomic notes: — Eperua bijuga differs from all other species by its 2 - jugate leaves, non-falcate leaflets with flat margin and one pair of glands at the adaxial base, tubular corolla, puberulous inflorescences, and persistent bracteoles. Among the species with tubular corolla, E. bijuga shares some characteristics with E. glabriflora, E. grandiflora subsp. grandiflora, E. grandiflora subsp. ciliata, E. jenmanii, E. oleifera, and E. purpurea, such as a non-glabrous sheath and glabrous ovary. However, E. bijuga is distinguished by its falcate ovary (vs. oblong or oblong-obovate in the others). Eperua bijuga is most morphologically similar to E. glabriflora. Both species share 2 - jugate leaves, non-falcate leaflets with two intramarginal veins, and a corolla of similar size and shape. However, E. bijuga differs by its puberulous to puberulent inflorescence, buds, sepals, and hypanthium (vs. glabrous in E. glabriflora), and smaller and persistent bracteoles (vs. larger and caducous in E. glabriflora).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F28FFCFFF6CF9C9FE50FE7F.taxon	materials_examined	Selected specimens: — BRAZIL. Amapá, Ferreira Gomes, Mangabeira, 29 May 2019, Rocha A. E. S. & Costa-Neto S. V. da 2165 (MG). Macapá, braço do rio Macacoari, July 1982, Rabelo B. V. & Cardoso J. O. 1261 (HAMAB, MG, NY, US). Porto Grande, Cupixi, Estrada Perimetral Norte, 15 December 1985, Rabelo B. V. et al. 3307 (HAMAB, INPA, NY). Tartarugarzinho, Rio Apurena, 22 July 1951, Fróes R. de L. & Black G. A. 27607 (IAN, NY, R). Pará, Afuá, rio urucu, 14 November – 04 December 1992, Maciel U. N. et al. 2147 (MG). Ananindeua, vila Marituba, Igarapé Marituba, 03 May 1991, Almeida S. S. et al. 458 (MG). Belém, Tapan „, beira da estrada, 02 May 1975, Oliveira E. 6291 (MG, RB); IPEAN. Reserva do Aurá. L. 166 – 70 - sn, 03 June 1968, Pires J. M. & Silva N. T. 11785 (HUEFS, IAN, RB, MFS). Breves, Araurá. Rio Maratana, afluente do rio aramé (Ilha Breves), margem inundada, 01 December 1922, Ducke W. A. s. n. RB 16927 (P, RB, U). Melgaço, Floresta Nacional de Caxiuan „, Ima 6, rio Curuzinho, 1 ° 72 ’ 25 ” S 51 ° 43 ’ 11 ” W, 20 April 2004, Freitas M. A. de et al. 1093 (INPA, MG, NY, RB). Moju, Experimental Field Station of Embrapa Amazônia Oriental, PA- 150, km 34, 2 ° 10 ’ 50 ” S 48 ° 0 ’ 0 ” W, 15 May 2008, Torke B. M. 432 (IAN, RB). Mosqueiro, Rio Pará, 20 June 1943, Ducke W. A. 1260 (IAN, F, K, MG, MO, NY, R, US); unknown locality, 13 October 1923, Ducke W. A. s. n. RB 16928 (RB, U). Muaná, Trajeto Monte Alegre / Santo André, 24 April 1982, Dantas M. & Silva N. T. da 1182 (IAN). Portel, FLONA de Caxiuan „, margem do igarapé Caquajó, 1 ° 57 ’ 00.0 ” S 51 ° 35 ’ 00.0 ” W, 08 July 2007, Félix-da-Silva M. M. et al. 241 (IAN, MG); regi „ o do Anapú, Rio Tapacú, 10 May 1956, Fróes R. de L. 32776 (IAN); regi „ o do Anapú, Rio Maparauá, 05 August 1956, Fróes R. de L. 32938 (IAN, MG). Salvaterra, Ilha do Marajó, Rio Paracauarí, margem do campo de pastagem do Sítio Caçador, 06 September 1969, Oliveira E. 4929 (IAN); Marajó, margem direita do Rio Jubim, próximo da Rod. Salvaterra-Joanes, a 16 Km de Joanes, 24 April 1980, Rosa N. A. 3628 (INPA, MG, NY). Santa Isabel do Pará, Caraparú, 25 April 1977, Silva M. G. da 3391 (MG). Soure, insula Marajó, 13 April 1927, Ducke W. A. s. n. RB 20312 (K, RB, U, US). Tailândia, Fazenda Borba Gato, near Rio Acará, 30 km W of Tomé-Açú-Paragominas road, approx. 30 km S of Tomé-Açú, 2 ° 40 ’ 00.0 ” S 48 ° 35 ’ 00.0 ” W, 08 November 1980, Daly D. C. et al. 889 (INPA, IAN, K, MG, NY, US). Tomé Açu, margem do rio Acará, 02 January 1978, Nascimento O. C. do 398 (MG, NY, US). Rio de Janeiro (cultivated), Rio de Janeiro, Arboreto do Jardim Botânico do Rio de Janeiro, 09 August 1991, Fontella H. & Caruso J. 2770 (RB). FRENCH GUIANA (cultivated?). unknown locality, undated, s. c. s. n. P 03465139 (P).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	description	(Figure 8; Fortes et al. 2023 Figures 3 – 5)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	type_taxon	Type: — BRAZIL. Maranh „ o, Caxias, Povoado Morro Agudo, Segundo Distrito, rio Itapecuru, área rural, 4 ° 32 ’ 18 ” S 43 ° 03 ’ 46 ” W, 24 March 2022, Gonçalves A. S. 02 (holotype RB [01458708]!, isotypes INPA!, UEC!, US!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	discussion	Tree or treelet (1.6 –) 3.5 – 11.5 m tall. Trunk 10.0 – 15.7 cm in diameter, bark grayish to brownish, smooth to striate, lenticellate. Stipules 15.1 – 40.9 × 11.8 – 22.7 mm, entirely joined, foliaceous, semiorbicular, persistent. Leaves (4 –) 5 – 7 - jugate; petioles 2.0 – 3.8 cm long; rachis 15.0 – 24.0 cm long; petiolules 5.3 – 8.4 mm long; blades 7.5 – 14.1 × 3.5 – 5.2 cm, coriaceous, glabrous, inconspicuous pellucid-punctate to epunctate, not discolorous, equilateral, lower pair ovate, middle pair ovate to elliptic, upper pair elliptic, apex attenuate to acuminate, mucronate, base asymmetrical, lower pair base rounded, middle pair base rounded to obtuse, upper pair base rounded to obtuse, margin revolute, secondary venation with two intramarginal veins, vein closer to the margin not continuous, main vein straight, depressed on the adaxial surface, tertiary veins slightly conspicuous. Inflorescences terminal, raceme, erect, tomentose, whitish, 10.0 – 20.0 cm long; bracts 8.4 × 4.8 mm, ovate, cucullate, apex gland absent, tomentose, whitish, caducous; bracteoles 9.8 – 12.7 × 6.5 – 9.8 mm, ovate, cucullate, apex gland absent, tomentose externally and within, whitish, persistent, attached to the lower portion of the pedicels; pedicel 18.0 – 24.4 mm long, 3.0 – 4.8 mm in diameter, not twisted, tomentose, whitish; buds 1.3 – 1.5 cm long, 1.0 cm in diameter, tomentose, whitish. Flowers: hypanthium 5.5 – 7.8 mm long, 7.7 – 10.6 mm in diameter, cup-shaped, equilateral, tomentose, whitish; sepal 2.3 – 2.9 × 1.3 – 1.7 cm, elliptic, unequal, the outer ones larger, cucullate, apex gland absent, greenish-white, tomentose, inner sepals scarious marginally, whitish; adaxial petal 2.3 – 3.3 × 5.5 – 6.5 cm, oblate, non-tubular, apex rounded, base truncate, white, glabrous; petalodia 2.7 – 8.8 × 1.1 – 3.0 mm; stamens exserted, joined basally in a tube, longer filaments 6.0 cm long, shorter filaments 5.1 cm long, tube equilateral, 2.8 – 3.4 mm long, glabrous, anthers 10.8 – 12.4 × 2.6 – 2.9 mm, rectangular; ovary 9.6 – 13.0 × 3.3 – 3.7 mm, oblanceolate, tomentose, greenish, stipe 5.8 – 10.5 mm long, tomentose, style 2.7 – 4.9 cm long, glabrous, stigma capitate. Legumes 22.0 × 7.0 cm, stipe 2.6 cm long, elliptic-falcate, apex obtuse, apiculate, dorsal margin alate, yellowish-green, veins absent, tomentose, yellowish. Seeds 4 per fruit, 3.2 × 2.0 cm (immature), obovate, whitishbrown. Phenology: — Flowering in March and September; fruiting in September.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	distribution	Distribution: — Described as having a distribution restricted to the Eastern region of Maranh „ o (Brazil) in the Itapecuru and Munin Basins in the Cerrado Biome (Fortes et al. 2023). Additional collections (Lui Y. et al. 2 and 178) of E. cerradoensis were found near the type location, in the municipality of Altos (Piauí, Brazil) within the Parnaiba Basin. Although the locality is considered part of the Caatinga Biome, it is situated at the border of the Cerrado Biome (IBGE 2019). Habitat: — The only species not occurring in the Amazonian region. E. cerradoensis was collected in the phytofisiognomies of cerradão (a type of savanna woodland), secondary forest, ombrophilous lowland forest with a predominance of palm trees, and in the open ombrophilous forest in the Cerrado Biome; and in semideciduous seasonal forest within the Caatinga Biome. It grows in stony clayey soils and is found at elevations ranging from 57 to 152 meters.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	description	Conservation status: — Categorized as Vulnerable (VU) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 7,323.777 km 2 and the estimated area of occupancy is 24.000 km 2. Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	etymology	Etymology: — The specific epithet alludes to its occurrence in the Cerrado Biome (Fortes et al. 2023). Vernacular names: — Embira de sapo (Almeida A. B. 134), imbira de sapo (Almeida A. B. 152), pitu (Oliveira D. 66), pracateira (Lui Y. et al. 2, Marinho M. A. O. 606, 629, Santos R. S. 296).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2CFFCDFF6CFDABFE94FECB.taxon	description	Uses: — In the type locality, Povoado Morro Agudo, it is mainly used to make charcoal and fences (Fortes et al. 2023). It also can be used in house construction, but this is not a common usage because the local population reported that the plant is very favorable to wood-dwelling termites (Fortes et al. 2023). Taxonomic notes: — Eperua duckeana, E. froesii, E. cerradoensis, E. reddeniae, and E. schomburgkiana are the only species within the short and erect inflorescence group that have a non-tubular corolla, exserted stamens, and falcate fruit with a 3: 1 proportion (vs. tubular corolla, included stamens, non-falcate fruit with a 1: 1 to 2: 1 length / width proportion in other short and erect inflorescence species). They also share multijugate leaves with straight leaflets and inflorescence with dense indumentum. Furthermore, E. cerradoensis shares white petals with E. duckeana and E. schomburgkiana; glabrous stamens joined at the base forming a tube with E. froesii; and tomentose ovary with E. duckena. The similarities and differences between species of this group are summarized in Table 3. Eperua cerradoensis is a distinct species by the completely glabrous stamens joined basally in a tube, tomentose ovary and fruits, and other vegetative characteristics. It is found primarily in the Cerrado Biome, differing from other species with distribution restricted to the Amazonian region (see Table 3, and Figure 8). Specimens examined: — BRAZIL. Maranhão, BRAZIL. Maranh „ o, Caxias, Povoado Morro Agudo, Segundo Distrito, área rural, 4 ° 32 ’ 18.2 ” S 43 ° 03 ’ 45.5 ” W, 18 September 2020, Gonçalves A. S. 01 (IAN, HABIT). Chapadinha, Cajazeiras, conglomerado MA- 255, subunidade 1, subparcela 5, indivíduo 16, F 6, 60 m elev., 3 ° 46 ’ 48 ” S 43 ° 33 ’ 36 ” W, 30 May 2018, Marinho M. A. O. 606 (UB); Cajazeiras, conglomerado MA- 255, subunidade 3, subparcela 10, indivíduo 11, F 8, 60 m elev., 3 ° 46 ’ 48 ” S 43 ° 33 ’ 36 ” W, 30 May 2018, Marinho M. A. O. 629 (UB). Nina Rodrigues, Mangueira, conglomerado MA- 202, subunidade 1, subparcela 1, indivíduo 4, F 6, 57 m elev., 3 ° 25 ’ 12 ” S 43 ° 44 ’ 24.1 ” W, 05 September 2018, Almeida A. B. 134 (UB); Mangueira, conglomerado MA- 202, subunidade 3, subparcela 10, indivíduo 13, F 8, 57 m elev., 3 ° 25 ’ 12 ” S 43 ° 44 ’ 24 ” W, 05 September 2018, Almeida A. B. 152 (UB). Timbiras, comunidade Morada Nova, conglomerado MA- 305, subunidade 2, subparcela 1, indivíduo 10, F 6, 94 m elev., 4 ° 8 ’ 24 ” S 43 ° 44 ’ 24 ” W, 02 May 2018, Oliveira D. 66 (UB); Bacabalzinho, conglomerado MA- 304, subunidade 3, subparcela 1, indivíduo 10, F 8, 91 m elev., 4 ° 08 ’ 24 ” S 43 ° 55 ’ 12 ” W, 28 March 2017, Santos R. S. 296 (UB). Piauí, Altos, Paleobotanical Site of St. Benedict, located in the Homoni village and is 19 km southwest of the headquarters of the municipality of Altos-PI, access made by BR 343 to the county seat, 152 m elev., 5 ° 7 ’ 10.92 ” S 42 ° 31 ’ 4 ” W, 15 March 2018, Lui Y. et al. 2 (HUEFS); Lui Y. et al. 178 (HUEFS).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	description	(Figures 8, 19, 20)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	type_taxon	Type: — BRAZIL. Amazonas: Rio Urubú abaixo da cachoeira Iracema, 7 July 1941, Ducke W. A. 732 (holotype US [00001144]!, isotypes F [0057800] image!, IAN [010263]!, MG [017957]!, MO [954041] image!, NY [1171178] image!, R sheet I [000054531]!, & sheet II [000054531 a]!, SI [002003] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	description	Tree 3.5 – 35.0 m tall. Trunk 5.0 – 60.0 cm in diameter, bark brownish, reddish-brown to grayish-brown, smooth to striate, lenticellate. Stipules 5.3 – 38.7 × (1.3 –) 3.4 – 26.9 mm, free, foliaceous, caducous to persistent. Leaves 3 – 5 - jugate; petioles 0.6 – 2.9 cm long; rachis (3.7 –) 6.8 – 19.5 (– 24.0) cm long; petiolules 3.6 – 10.4 mm long; blades 4.7 – 14.7 × 2.0 – 10.3 cm, coriaceous, glabrous, pellucid-punctate, rarely inconspicuous pellucid-punctate, discolorous, sometimes slightly discolorous, equilateral, lower pair elliptic, sometimes lanceolate, rarely ovate or broadly elliptic, middle pair elliptic, sometimes lanceolate, rarely narrowly elliptic, upper pair elliptic, rarely lanceolate, obovate, or narrowly elliptic, apex attenuate to acuminate, mucronate, base asymmetrical, rarely oblique, lower pair base cuneate, obtuse, rounded to subcordate, middle pair base obtuse to rounded, sometimes cuneate to subcordate, rarely cordate, upper pair base obtuse to cuneate, margin flat, secondary venation with two intramarginal veins, sometimes one, vein closer to the margin not continuous, main vein straight, very prominent on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal, rarely axillary or on the branchlets, panicle, sometimes raceme, erect, tomentose, brownish-yellow, 6.0 – 20.0 cm long, lateral racemes 2.3 – 18.0 cm long, alternate and distichous; bracts caducous, not seen; bracteoles 1.9 – 5.0 × 2.3 – 4.2 mm, ovate, rarely semiorbicular, cucullate, apex gland absent, tomentose externally, glabrous within, brownish-yellow, falling early, attached to the lower middle portion of the pedicels; pedicel 15.0 – 23.3 mm long, 1.0 – 2.8 mm in diameter, not twisted, tomentose to pubescent, brownish-yellow or coppery; buds 0.8 – 1.8 cm long, 0.4 – 0.9 cm in diameter, tomentose, brownish-yellow. Flowers: hypanthium 3.3 – 10.6 mm long, 2.9 – 7.4 mm in diameter, cup-shaped, sometimes tubular, equilateral, tomentose, brownish-yellow; sepal 0.8 – 1.9 × 0.3 – 0.9 cm, elliptic, sometimes oblong or ovate, unequal, the outer ones larger, cucullate, apex gland absent, greenish, green, greenish with rose spots, white to cream, tomentose externally, brownish-yellow, inner sepals scarious marginally; adaxial petal 1.2 – 2.4 × 1.9 – 3.3 cm, oblate, non-tubular, apex truncate or rounded, base truncate, white, whitish, greenish, cream to white-lilac, glabrous; petalodia 0.4 – 1.6 × 0.4 – 1.0 mm; stamens, exserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 1.7 – 4.9 cm long, shorter filaments 1.2 – 3.4 cm long, sheath slightly inequilateral to equilateral, shorter side 3.0 – 8.1 mm long, longer side 6.1 – 8.7 mm long, sheath entirely tomentose, free filaments glabrous, brownish-yellow, shorter stamens anthers 2.7 × 0.7 mm, rectangular, longer stamens anthers 2.2 – 4.1 × 0.7 – 1.3 mm, rectangular; ovary 5.9 – 9.9 × 1.8 – 4.5 mm, oblaceolate, tomentose, brownish-yellow, stipe 2.9 – 9.1 mm long, tomentose, style 2.7 – 4.3 cm long, stigma obtuse to capitate. Legumes 7.5 – 22.0 × (1.3 –) 4.0 – 8.4 cm, stipe 1.1 – 2.5 cm long, with remnants of the androecium, oblanceolate-falcate, apex obtuse, sometimes acute, apiculate, margin entire, dull brown, veins absent, sometimes with inconspicuous transversal veins, tomentulose, rarely tomentose or tomentulose to glabrescent with long scattered trichomes, brownish-yellow. Seeds 2 – 4 per fruit, 3.1 – 4.5 × 1.7 – 3.0 cm, ovate to oblong, reddish-brown. Phenology: — Flowering in March and from May to December with a peak from August to September; fruiting from January to March and from May to December with a peak in August.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	distribution	Distribution: — Brazil in the states of Amazonas and Pará. Habitat: — Mainly terra-firme forest, sometimes in igapó and várzea forests, and campinaranas, on sandy and clayey soil, from 7 to 250 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	description	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 88,736.968 km 2 and the estimated area of occupancy is 204.000 km 2. Occurrence in protected areas: — Reserva Florestal Adolpho Ducke (Brazil, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	etymology	Etymology: — The specific epithet is a tribute to Adolpho Ducke, who revised and described new species and varieties for the genus Eperua (Ducke 1932, 1940, Cowan 1975). Vernacular names: — envira iodo (Marra D. M. & Neves A. F. 78), muirapiranga branca (Coelho D. s. n. INPA 20747, Rodrigues W. A. & Chagas J. 1725, Rodrigues W. A. & Osmarino 6769), muirapiranga da folha miúda (Rodrigues W. A. & Chagas J. 1725, Rodrigues W. A. & Lima J. 2233, Rodrigues W. A. & Osmarino 6769), murajiboia (Mello F. C. de & Ribamar s. n. INPA 58309), pau de óleo (Silva S. F. da 205 a).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	description	Uses: — Used in constructions due to its heavy, soft, and resinous wood (Rodrigues W. A. & Chagas J. 1725). Taxonomic notes: — Eperua duckeana, E. froesii, E. cerradoensis, E. reddeniae, and E. schomburgkiana are the only species with short and erect inflorescence that exhibit a non-tubular corolla, exserted stamens, falcate fruit with a length-to-width ratio of 3: 1 (vs. tubular corolla, included stamens, non-falcate fruit with a length-to-width ratio of 1: 1 to 2: 1 in other short and erect inflorescence species). They also share multijugate leaves with straight leaflets and inflorescence covered in dense indumentum. Furthermore, E. duckeana stands out as a distinct species due to its stamens fused at the base in a sheath of 9 + 1, tomentose sheath and ovary, and tomentulose fruits. These features and other vegetative characteristics contribute to its differentiation from other species. Additionally, E. duckeana has a specific distribution in the Central Amazonia region. Please refer to Table 3 for a summary of the similarities and differences among these species.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F2EFFF0FF6CFE37FA95FC3B.taxon	materials_examined	Selected specimens: — BRAZIL. Amazonas, Itapiranga, rio Pitinga, margem direita, 24 August 1979, Cid-Ferreira C. A. et al. 703 (INPA, NY, US). Manaus, Reserva Florestal Adolpho Ducke ,, L 4 após km 3350, 129 m elev., 2 ° 56 ’ 59 ” S 59 ° 56 ’ 10 ” W, 21 January 2022, Fortes E. A. et al. 190 b (INPA, RB, UEC, US); Reserva Florestal Adolpho Ducke, Igarapé Ipiranga, pique do INPA, sentido oeste-leste, 2 ° 53 ’ 0 ” S 59 ° 58 ’ 0 ” W, 05 July 1993, Ribeiro J. E. L. S. et al. 1037 (HUEFS, INPA, K, MO, NY, R, RB, SP, U); ZF 3, Distrito Agropecuário da SUFRAMA, rodovia BR 174, Km 64, depois 34 Km Leste na ZF 3, fazenda Esteio, reserva nº 1302, árvore nº 775, 2 ° 26 ’ 0 ” S 59 ° 48 ’ 0 ” W, 24 September 1981, Nascimento J. R. M. et al. 1302.775 (INPA, NY); IFAM, Campus Manaus-Zona Leste, 05 November 2011, Kinupp V. F. et al. 4542 (EAFM). Presidente Figueiredo, no entorno do Lago da Rebio Uatum „, Balbina, 13 August 2008, Silva M. C. R. da et al. 40 (INPA). S „ o Sebasti „ o do Uatum „, RDS Uatum „, Igapó de topografia alta, situado nas margens do rio Abacate, 7 m elev., 2 ° 11 ’ 37 ’’ S 58 ° 43 ’ 10 ’’ W, 15 May 2016, Lobo G. S. 14 (INPA). Pará, Oriximiná, cachoeira da Porteira, entrada Perimetral Norte, 1 ° 3 ’ 17.4 ” S 57 ° 2 ’ 28 ” W, 09 June 2000, Faria S. M. de et al. 2053 (HSTM, RB); Porto Trombetas, rio Mapuera, cachoeira da Égua, Ilha, 11 December 1907, Ducke W. A. 9088 (RB). Rio de Janeiro (cultivated), Seropédica, Embrapa-Agrobiologia, 14 September 2001, Faria S. M. de & C. C. F. da 2186 (RB).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	description	(Figures 11, 21, 22)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	materials_examined	Type (designated by Cowan 1975): — FRENCH GUIANA. Cayenne, s. d. [July 1762 – July 1764], Aublet J. B. C. F. s. n. (lectotype BMinflorescence [000952284] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	diagnosis	Tree 8.0 – 40.0 m tall. Trunk 20.0 – 70.0 cm in diameter, bark grayish to reddish, smooth, lenticelate. Stipules 1.4 – 4.3 × 1.3 – 3.1 mm, joined, non-foliaceous, caducous to persistent. Leaves (2 –) 3 – 4 - jugate; petioles 1.2 – 9.7 cm long; rachis 1.2 – 11.9 cm long; petiolules 2.5 – 6.5 mm long; blades 2.6 – 17.5 × 2.4 – 6.8 cm, chartaceous, sometimes coriaceous, glabrous, minutely pellucid-punctate, not all areoles have one dot, not discolorous to discolorous, inequilateral, all pairs elliptic-falcate, apex acuminate, sometimes caudate, mucronulate, base asymmetrical, lower pair base obtuse to rounded, middle pair base obtuse, rounded or cuneate, upper pair base obtuse to cuneate, rarely rounded, margin flat, rarely slightly revolute, secondary venation with one intramarginal vein very close to the margin, main vein curved, depressed, rarely prominent near the base on the adaxial surface, tertiary veins conspicuous, areoles larger, plane. Inflorescences terminal, rarely axillary, panicle, main axis and lateral racemes pendulous, main axis strigulose, sometimes glabrous, lateral racemes tomentose, brownish, 66.0 – 300.0 cm long, lateral racemes (1.1 –) 2.5 – 8.8 (– 11.0) cm long, alternate and spirally arranged; bracts 2.9 – 3.2 × 1.3 – 1.8 mm, ovate, apex gland absent, tomentose, brownish, caducous; bracteoles 2.3 – 6.1 × 2.3 – 4.0 mm, broadly ovate, sometimes ovate, apex gland absent, tomentose, brownish, caducous, attached to the higher portion of the pedicels; pedicel 8.3 – 19.3 mm long, 0.7 – 1.5 mm in diameter, twisted, tomentose, brownish; buds 0.8 – 1.8 cm long, 0.4 – 0.9 cm in diameter, tomentose, brownish. Flowers: hypanthium 3.6 – 5.3 mm long, 3.3 – 6.3 mm in diameter, cup-shaped, equilateral, tomentose, brownish; sepal 1.0 – 1.8 × 0.5 – 1.2 cm, elliptic to oblong, unequal, the outer ones larger, cucullate, apex gland absent, pink, red, redidish-green to greenish, tomentose, inner sepals scarious marginally, brownish; adaxial petal 1.0 – 2.0 × 1.2 – 2.5 cm, oblate, non-tubular, apex rounded, base truncate or subcordate, gibbous, pink, pale-pink, purple, reddish, greenish to red, glabrous; petalodia 0.5 – 1.9 × 0.7 – 1.2 mm; stamens exserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 3.7 – 4.6 cm long, shorter filaments 1.5 – 2.4 cm long, sheath equilateral, 2.1 – 7.3 mm long, sheath and free filaments base villous, middle to upper free filaments glabrous, brownish, anthers 5.7 – 6.0 × 1.6 – 2.0 mm, rectangular; ovary 4.7 – 8.8 × 2.2 – 4.4 mm, oblong to obovate, sericeous, brownish, stipe 3.6 – 6.2 mm long, sericeous, style 2.2 – 4.5 cm long, stigma capitate. Legumes 19.0 – 38.0 × 5.1 – 10.8 cm, stipe 1.8 – 2.6 cm long, elliptic-falcate, sometimes lanceolate-falcate, apex acute, margin entire to slightly thickened, dark brown, veins absent or with some slightly conspicuous veins, strigulose, rarely glabrous, trichomes brownish. Seeds 1 – 4 per fruit, 3.5 – 4.6 × 1.9 – 2.4 cm, oblong, ovate to irregular, dark-brown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	description	Phenology: — Flowering and fruiting throughout the year, with a flowering peak from September to December and a fruiting peak from October to December.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	distribution	Distribution: — Occurring throughout the Guyana Shield: Brazil (Amapá), French Guiana (Cayenne, Saint-Laurent-du-Maroni), Guyana (Barima-Waini, Cuyuni-Mazaruni, Demerara-Mahaica, East Berbice-Corentyne, Pomeroon-Supenaam, Potaro-Siparuni, Upper Demerara-Berbice), Surinam (Brokopondo, Coronie, Kabalebo, Nickerie, Para, Paramaribo, Patamaca, Sipaliwini), and Venezuela (Bolívar). Cultivated in Trinidad and Tobago, Indonesia, and Panama. Habitat: — Non-flooded, inundated and periodically inundated forests, and savannas. It can occur from sea level up to elevations of 500 m, and sometimes even up to 800 meters. It grows mainly on white, brown, and mixed sand soils, although it can also be found on bauxite, laterite, and clayey soils. Along with E. grandiflora, E. falcata comprise the Eperua dominant forest known as wallaba forest in Guyana and Surinam.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	description	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for this species is 501,331.224 km 2, and the estimated area of occupancy is 664.000 km 2. Occurrence in protected areas: — Parque Nacional Montanhas do Tumucumaque (Brazil, Amapá), Réserve Naturelle des Nouragues (French Guiana, Régina), Iwokrama Rainforest Reserve (Guyana, Potaro-Siparuni), Kaieteur National Park (Guyana, Potaro-Siparuni), Mabura Hill Forest Reserve (Guyana, Upper Demerara-Berbice), Brownsberg Nature Reserve (Surinam, Brokopondo), and Central Surinam Nature Reserve (Surinam, Kabalebo).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	etymology	Etymology: — The specific epithet is derived from its elliptic-falcate fruits (Bentham 1870). Vernacular names: — apa (Irwin H. S. et al. 48277, Irwin H. S. et al. 48408, Irwin H. S. & Pires J. M. 48613, Maguire B. et al. 48408), apa-roxo (Pires J. M. & Westra L. Y. T. 48876), bioudou or bi-oudou (BAFOG 51, 7062, 7064, 7254, 7298, Mori S. A. et al. 23384, Petrov I. s. n. P 03465196, Vieillescazes A. 616), biouolou-wata (BAFOG 143), birihoedoe (Stahel G. 22), bi-udu or biudu (Hoffman B. 6522, Puig H. 10263, 10285), caraota (Blanco C. A. 603, Meijeraan J. W. 42), falcaha (Angel G. 19), itoeli-walaba (Stahel G. 66), kergwalaba (Jimenez-Saa J. H. 14330), mahomillo-negro (Steyermark J. A. 86692), ouapa (Oldeman 1212), rosa-de-montaña (Foldats E. 2645), soft-wallaba (Berlo M. van collections, Redden K. M. et al. 1050, 1058, 1059), tapaka (Grenand P. 610, Puig H. 10263, 10285), walaba (Berlo M. van collections, Bhikhi C. R. et al. 79, Borsboom N. W. J. 12025, 12044, B. W. 1171, Florsch ¸ tz J. & Florsch ¸ tz P. A. 217, Jimenez-Saa J. H. 14275, 43360 (1627), Lindeman J. C. & Roon A. C. 1980 / 860, Lobato M. 9, Maguire B. 24778, Ogtrop F. van 1, Reuder & Roberts 12329, Sabajo P. H. & Roberts L. 11192, Vreden C. C. J. 13673), walaba-kharemeroe (Stahel G. 22), walaba-koeleroe (Stahel G. 23), walaba-of-bijlhout (Versteeg G. M. 393), wallaba (Andel T. R. van et al. collections, BAFOG 6033, Evans R. et al. 2154, Evans R. & Lewis G. 1871, Kelloff C. L. et al. 1300, Kelloff C. L. & Roesel C. C. 1155, Junker L. 507, Little E. L. Jr. 16825, McDowell T. & Gopaul D. 2451, Persaud A. C. 243, Pipoly J. J. & Boyan R. 8701, Redden K. M. et al. 4572, Stoffers A. L. et al. 62), wapa (Angel G. 19, 158, 176, BAFOG 6033, Dutrève B. 437, Garnier F. A. 138, Hallé F. 4143, Hoff M. 5895, Kodjoed J. - F. 140, Larpin D. 187, Loubry D. 1811, Mori S. A. et al. 23384, Paget D. 49, Petrov I. 124, Prévost M. F. 4221, 4232, Puig H. 10285, Sauvain M. 743, Thiel J. collections, Vieillescazes A. 616), wapa blone (Thiel J. collections), wapa-de-crique (BAFOG 143), wapa-graj (Bena P. 1274), wapa-gras (BAFOG 1274), wapa-huileux (BAFOG 1274), wapa-rouge (Thiel J. collections), warapa (Andel T. R. van et al. 665), watapa (Andel T. R. van et al. 1007), water-wallaba (Redden K. M. et al. 3265), white-sand-wallaba (Hoffman B. 4599), zwarte-walaba (Stahel G. 22, 22 a).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	description	Uses: — Eperua falcata is highly valued for its wood, which is extensively used in Guyana and Surinam for general construction, bridge construction, light and telephone poles, shutter windows, and roof shingles. The following herbarium specimens provided information of the utilization of its wood: Andel T. R. van et al. 665, 1007, 3073, Berlo M. collections, Bhikhi C. R. et al. 79, Redden K. M. et al. 1050, 1058, 1059. Taxonomic notes: — Eperua falcata is the only species with long and pendulous inflorescences with pendulous lateral racemes (Table 4). Morphologically, it shares the following similarities with E. leucantha and E. venosa: joined stipules, leaflets with secondary venation with one intramarginal vein very close to the margin, long and pendulous inflorescences, non-tubular corolla, exserted stamens joined in a diadelphous sheath, and sericeous ovary. Eperua falcata also shares multijugate leaves with E. leucantha and non-white flowers with E. venosa. However, notable differences set E. falcata apart from these two species. Eperua falcata has smaller adaxial petals (1.0 – 2.0 × 1.2 – 2.5 cm) compared to E. leucantha (1.9 – 3.3 × 2.9 – 6.3 cm) and E. venosa (2.5 – 3.4 × 4.7 – 6.0 cm). Additionally, E. falcata has minutely pellucid-punctate leaflets with not all areoles having a dot, whereas E. leucantha and E. venosa have pellucid-punctate leaflets with each areola having at least one dot. Further distinctions include the lower and middle pairs of leaflets in E. falcata tending to have less asymmetrical and more rounded to obtuse bases (vs. more asymmetrical, obtuse to cuneate, rarely rounded in E. leucantha), longer [(1.1 –) 2.5 – 8.8 (– 11.0) cm long] and pendulous lateral racemes (vs. shorter [0.5 – 2.5 (– 3.1) cm long] and patent to erect in E. leucantha), brownish buds (vs. grayish white in E. leucantha), pink to purple petals (vs. white in E. leucantha), villous sheath (vs. sericeous in E. leucantha). From E. venosa, E. falcata also differs by multijugate leaves (vs. always 1 – 2 (– 3) - jugate in E. venosa), brownish buds (vs. grayish-white in E. venosa), glabrous adaxial petal (vs. sericeous at the base in E. venosa), and villous staminal sheath (vs. sericeous in E. venosa). Eperua falcata and E. leucantha are allopatric species (Figure 11). Eperua falcata occurs in the Guyana Shield, with its westernmost occurrence in the Venezuelan state of Bolívar. It is a widespread and extensively collected species, although no flowering collections have been reported beyond the Bolívar state. In contrast, in Venezuela, E. leucantha occurs exclusively in the southeastern part of the Amazonas state. Eperua falcata is parapatric with E. venosa, which is restricted to the region of Gran Sabana in Venezuela, while E. falcata borders this region (Figure 11).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F13FFF5FF6CFC65FE44F8AB.taxon	materials_examined	Selected specimens: — BRAZIL. Amapá, Laranjal do Jari, Parque Nacional do Tumucumaque, 0 ° 16 ’ 35 ” N 53 ° 6 ’ 24 ” W, 14 January 2005, Pereira L. A. & Cardoso J. O. 825 (RB). Oiapoque, nas margens da rodovia BR 156 entre Oiapoque e Calçoene, a 21 km de Oiapoque, 3 ° 40 ’ 33.2 ” N 51 ° 46 ’ 14.1 ” W, 04 December 2021, Fortes E. A. & Silva G. S. da 165 (IAN, RB). CAMEROON (cultivated). Victoria Botanical Garden, 12 February 1927, Fairchild D. G. s. n. (US 1716064, BR 0000017324229). FRENCH GUIANA. Cayenne, Camopi, Trois sauts (haut Oyapock), saut Mansa, 31 December 1974, Grenand P. 610 (CAY, US). Cayenne, Quesnel-Chantier F. R. G., 11 December 1979, J. Thiel 619 (CAY); Exploitation forestière F. R. G., 02 February 1981, Thiel J. 792 (CAY). Kourou, montagne des Singes, 23 November 1981, Cremers G. A. 7659 (CAY, P, US). Montsinéry-Tonnegrande, crique Serpent, rive gauche à 1 m au dessus de la crique, February 1954, BAFOG 143 (U). Régina, station des Nouragues (cam inselberg), trail CE-CO, 40 – 20 m elev., 4 ° 5 ’ 30 ” N 52 ° 41 ’ 0 ” W, 23 September 2007, Geiselman C. K. 94 (NY). Roura, placer Boulanger, bassin de la Comté, 12 November 1979, Thiel J. 582 (CAY). Saint-Élie, Saut Takari-Tanté, bassin du Sinnamary, 34 m elev., 4 ° 37 ’ 0 ” N 52 ° 56 ’ 0 ” W, 17 November 1989, Hoff M. 5895 (CAY); Saut Dalles, bassin du Sinnamary, 0 m elev., 4 ° 33 ’ 0 ’’ N 52 ° 54 ’ 0 ’’ W, 01 November 1991, Loubry D. 1287 (CAY). Saint-Georges, Saut Maripa, bassin de l’Oyapock, August 1973, Garnier F. A. 138 (CAY). Saint-Laurent-du-Maroni, Mana, Chantier Laussat — S. F. M., bassin de La Mana, 28 November 1979, Thiel J. 593 (CAY). Maripasoula, rivière Grand Inini, bassin de l’Inini, 3 ° 40 ’ 0 ” N 53 ° 50 ’ 0 ” W, 07 July 1990, Sabatier D. & Prévost M. F. 3070 (CAY). Saint-Laurent-du-Maroni, bassin du Bas-Maroni, 14 April 1961, Aubréville A. 1961 / 236 (P, U, US). Sa ̧ l, Monts La Fumée, 200 – 400 m elev., 3 ° 37 ’ 0 ” N 53 ° 12 ’ 0 ” W, 12 October 1982, Boom B. M. & Mori S. A. 1985 (CAY). GUYANA. Barima-Waini, Kariakau, Kariako Village, Barama River, 145 m elev., 7 ° 22 ’ 0 ” N 59 ° 42 ’ 0 ” W, 22 December 1995, Andel T. R. van et al. 665 (U, WAG); Morebo, Barima River, N. W. district, December 1908, Anderson C. W. 122 (RB). Cuyuni-Mazaruni, Bartica, Essequibo River, Moraballi Creek, near Bartica, 23 September 1929, Sandwith N. Y. 329 (RB, U, US). Imbaimadai vicinity, Mazaruni River, 0.55 miles South West of Partang River base camp, 485 m elev., 5 ° 41 ’ 17 ” N 60 ° 17 ’ 10.2 ” W, 23 November 2002, Redden K. M. et al. 1238 (U, US); Pakaraima Mts., Mazaruni River, NW of Chi-Chi Falls, along stream between two tepuis, approx. 1.5 km above base camp 2, 762 m elev., 5 ° 35 ’ 48.6 ” N 60 ° 12 ’ 49 ” W, 12 February 2004, Redden K. M. et al. 1643 (U, US); Mazaruni River, unnamed falls (possible Carin Crow Falls), 0.67 miles W of base camp, 61 m elev., 5 ° 57 ’ 4.3 ” N 59 ° 19 ’ 5.2 ” W, 09 October 2004, Redden K. M. et al. 3265 (US); Kako River, 536 m elev., 5 ° 31 ’ 29 ” N 60 ° 50 ’ 19 ” W, 12 May 2009, Redden K. M. et al. 6552 (NY, US). Demerara-Mahaica, Soesdyke-Linden Hwy, 0 – 3 km N Kuru-Kuru Cr, 16 – 19 km S Georgetown-Timehri Hwy, 10 – 20 m elev., 6 ° 22 ’ 0 ” N 58 ° 15 ’ 0 ” W, 10 November 1991, Gillespie L. J. 4250 (INPA, MO, US). East Berbice-Corentyne, White Hill on Corentyne River, 10 – 50 m elev., 5 ° 6 ’ 0 ” N 57 ° 19 ’ 0 ” W, 24 April 1990, McDowell T. & Gopaul D. 2451 (U, US). Pomeroon-Supenaam, along road between Wyaka and Mainstay, 7 ° 15 ’ 0 ” N 58 ° 31 ’ 59.9 ” W, 23 April 1989, Gillespie L. J. & Persaud H. 1169 (US). Potaro-Siparuni, Iwokrama Rainforest Reserve, N side of Burro-Burro R., 60 – 90 m elev., 4 ° 33 ’ 0 ” N 58 ° 50 ’ 0 ” W, 27 September 1995, Clarke H. D. 255 (CAY, U, US); Pakaraima Mts, Mt. Wokomung, Suruwabaru Creek, 1 – 2 km from juncture with Yuarba River, 675 – 750 m elev., 5 ° 2 ’ 0 ” N 59 ° 54 ’ 0 ” W, 07 November 1993, Henkel T. W. et al. 4142 (CAY, U, US); Kaieteur National Park, above falls, 474 m elev., 5 ° 10 ’ 28.0 ” N 59 ° 28 ’ 54.1 ” W, 29 August 2006, Redden K. M. et al. 3860 (CAY, US). Upper Demerara-Berbice, Berbice, Dubulay ranch, 10 m elev., 5 ° 35 ’ 0 ” N 57 ° 53 ’ 0 ” W, 19 October 1997, Jansen-Jacobs M. J. 5623 (U, US). Mabura Hill Forest Reserve, 13 km S of Mabura Hill, 60 m elev., 5 ° 10 ’ 0 ’’ N 58 ° 42 ’ 0 ’’ W, 13 April 1996, Clarke H. D. 1633 (CAY, U, US). Upper Takatu-Upper Essequibo, Cut Line AB from Lethem-Kurupukari road on 270 °, 75 m elev., 4 ° 34 ’ 58 ” N 58 ° 44 ’ 45 ” W, 21 April 1992, Pennington R. T. et al. 372 (CAY, U, US). INDONESIA (cultivated). Java, Borgor City, cultivated in the Botanic Gardens Buitenzorg-Java, 30 May 1941, Wit I. i. H. C. de 70 a (L). PANAMA (cultivated). Panamá, Ciudad de Panamá, Parque Natural Summit, 80 m elev., 15 February 2008, Ibáñez A. et al. 5358 (MA). SURINAM. Brokopondo, Brownsberg, Boschreserve (Forest Reserve), sectie O, 10 September 1917, B. W. 3210 (IAN, U). Coronie, Vosberg, January 1947, Burger D. 8 (U, WAG). Kabalebo, Central Surinam Nature Reserve, lower Rechter Cooppename River, 50 – 75 m elev., 4 ° 21 ’ 41.2 ” N 56 ° 31 ’ 8.2 ” W, 23 February 2004, Clarke H. D. & Rhodes J. 11085 (U). Nickerie, unknown city, Corantijne, 11 December 1949, Landsbosbeheer 902 (U). Para, Carolina, Blakawatra, 5 ° 22 ’ 59 ” N 54 ° 55 ’ 0 ” W, 28 November 1921, B. W. 5524 (U). Oost, Powakka, Property of Makosi family, 5 ° 26 ’ 38.4 ” N 55 ° 4 ’ 37 ” W, 21 May 2012, Berlo M. van et al. 63 (L). Zuid, Zanderij, September 1942, Stahel G. 22 (IAN, RB, U, WAG); banks of Sabakoe Creek, ca. 100 m W from Zanderrij to Kraka road, 4.9 km from intersection with Zanderij Hwy, 25 m elev., 5 ° 25 ’ 0 ” N 55 ° 11 ’ 30 ” W, 10 July 1994, Evans R. & Lewis G. 1871 (CAY, IAN, INPA, MO, P, RB, U, US). Jodensavanne-Mapane kreek area (Surinam R.), Bij kamp aan Mapane-kreek, 19 September 1953, Lindeman J. C. 4649 (F, U). Paramaribo, Forest surrounding of Carolina, 50 km, South-East of Paramaribo, 11 August 1874, Outer R. W. den 963 (L, U). Patamaca, B. S. H. ekspl. Patamaca, 02 November 1967, Borsboom N. W. J. 12025 (WAG). Sipaliwini, Boven Saramaca, Tafelberg (Table Mountain), 15 September 1944, Maguire B. 24778 (F, U, US). Boven Suriname, Berlijn, January 1844, Focke H. C. 945 (U). Coeroeni, vinicity of camp on W bank of Zuid River, across river from central Surinam Nature Preserve, ca. 10 km straight-line distance SSE of Kayserberg airstrip, 240 m elev., 3 ° 0 ’ 3 ” N 56 ° 27 ’ 3 ” W, 14 June 2003, Rosário C. S. & Rosário D. O. 2011 (MG, MO). Kabalebo, Tafelberg (Table Mountain), summit, trail from Augustus Creek to Lisa Creek, ca. 0.5 km from Lisa Creek, ca. 200 m from western rim of tepuí, 575 m elev., 3 ° 55 ’ 0 ” N 56 ° 12 ’ 30 ” W, 25 June 2001, Evans R. 3209 (MO, US). Nickerie, Fallawatra, 03 November 1971, Jimenez-Saa J. H. 14275 (U). Tapanahony, vicinity of Ulemari River, 13 km upstream from its confluence with Litani River, 150 m elev., 3 ° 13 ’ 17 ” N 54 ° 15 ’ 31 ” W, 04 April 1998, Evans R. & Peckham H. 2814 (L, U). TRINIDAD AND TABAGO (cultivated). Botanical Gardens, 19 August 1927, Broadway W. E. s. n. US 1343487 (US). VENEZUELA. Bolívar, El Dorado, ± 65 km S. of El Dorado, 200 m elev., 25 August 1966, Meijeraan J. W. 42 (WAG). 90 km from El Dorado towards Sta. Elena, 28 December 1956, Foldats E. 2645 (US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	description	Figures 8, 23 – 25)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	type_taxon	Type: — BRAZIL. Pará: Rios Pacajá and Muirapiranga, Km 3,5 – 6,5, line SW of Ilha de Breu, 23 September 1965, Prance G. T. et al. 1458 (holotype IAN [117052]!, isotypes K [000807973] image!, MO [1780623] image!, NY [01171288]!, U [0083421] image!, US [00243011]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	diagnosis	Diagnosis: — Eperua froesii has short and erect inflorescences, non-tubular corollas, and exserted stamens. It differs from species with the same type of inflorescence, corolla, and stamen by the combination (see Table 3): stipules entirely joined, epunctate to inconspicuous pellucid punctate leaflets, flat to slightly revolute margins, depressed main vein on the adaxial surface, inconspicuous tertiary venation, brownish-yellow to reddish-brown inflorescence indumentum, white to light-red petals, stamens joined basally in a tube, glabrous stamens, ovary, and fruit. Tree 20.0 – 25.0 m tall. Trunk and bark not seen. Stipules entirely joined, caducous, 5.6 – 44.5 × 10.1 – 44.0 mm, foliaceous, or 4.0 – 5.2 × 3.0 – 4.7 mm, non-foliaceous, persistent to caducous. Leaves 4 – 5 - jugate; petioles 1.0 – 2.5 cm long; rachis 11.0 – 23.0 cm long; petiolules 4.2 – 7.6 mm long; blades 5.3 – 13.4 × 2.5 – 5.8 cm, coriaceous, glabrous, epunctate to inconspicuous pellucid-punctate, not discolorous to slightly discolorous, equilateral, lower pair elliptic to ovate, middle pairs elliptic, narrowly elliptic to ovate, upper pair elliptic, apex attenuate to acuminate, base symmetrical to asymmetrical, lower pair base obtuse to rounded, middle pair base obtuse to rounded, upper pair base obtuse to cuneate, margin flat to slightly revolute, secondary venation with one or two intramarginal veins (sometimes inconspicuous in developed leaves), vein closer to the margin continuous or not continuous, main vein straight, depressed on the adaxial surface, tertiary veins inconspicuous, areoles smaller concave. Inflorescences terminal or axillary, panicle, erect, puberulous, pubescent to tomentose (strigulose in infructescence), brownish-yellow to reddish-brown, 3.2 – 6.1 (– 7.2 in infructescence) cm long, lateral racemes 1.0 – 2.7 (– 6.0 in infructescence) cm long, alternate and distichous; bracts 3.6 – 4.1 × 2.6 – 4.4 mm, ovate, concave, apex gland absent, strigose to tomentose, brownish-yellow to reddish-brown; bracteoles 4.5 – 9.0 × 2.9 – 6.1 mm, ovate to oblate, cucullate, apex gland absent, pubescent to tomentose, brownish-yellow to reddish-brown, persistent to caducous, attached to the lower middle portion of the pedicels; pedicel 4.0 – 6.6 mm long, 1.6 – 3.4 mm in diameter, not twisted, tomentose, brownish-yellow to reddish-brown; buds 0.7 – 1.9 cm long, 0.5 – 1.0 cm in diameter, tomentose, brownish-yellow to reddish-brown. Flowers: hypanthium 4.3 – 4.9 mm long, 4.8 – 6.4 mm in diameter, cup-shaped, equilateral, tomentose, brownish-yellow to reddish-brown; sepal 1.0 – 1.5 × 0.5 – 0.8 cm, elliptic to oblong, unequal, the dorsal one larger, cucullate, apex gland absent, green, tomentose, tomentulose to puberulous externally, brownish-yellow to reddish-brown, inner sepals scarious marginally; adaxial petal 0.9 – 1.4 × 1.2 – 1.8 cm, oblate, non-tubular, apex rounded, base truncate, white to light-red, glabrous; petalodia 0.6 – 0.8 × 0.4 – 0.5 mm; stamens exserted, joined basally in a tube, filaments 1.9 – 2.5 cm long, tube equilateral, 1.0 – 1.6 mm long, glabrous, anthers 4.6 – 5.4 × 1.0 – 1.3 mm, rectangular; ovary 5.0 – 8.6 × 2.6 – 3.1 mm, oblong to obovate, glabrous, stipe 5.9 – 9.5 mm long, glabrous, style 1.2 (pre-anthesis) – 2.0, stigma capitate, papillose. Legumes 13.2 – 23.5 × 3.8 – 9.8 cm, stipe 2.0 – 2.5 cm long, oblanceolate, apex obtuse, apiculate, margin slightly thickened, dark-brown, veins absent, glabrous. Seeds not seen. Phenology: — Flowering in September and October; fruiting in October and November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	distribution	Distribution: — Occurring in Pará (Brazil) along the Acaraí River, a tributary of the Xingu River, and Pacajá, Anapu, and Muirapiranga (connected with Alto Anapu River) rivers, which are tributaries of the Tocantins River. Habitat: — Floodplain forests, likely occurring in both igapó and várzea forests. The floodplain forests of the Xingu and Tocantins rivers are considered Igapó due to their clear-water rivers (Junk 2011), while the floodplains near Portel in the Anapu river region are classified as várzea forests (Junk 2011) due to Amazon River influence.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	description	Conservation status: — Categorized as Near Threatened (NT) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 21,015.730 km 2 and the estimated area of occupancy is 16.000 km 2. Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	etymology	Etymology: — The specific epithet is a tribute to Ricardo de Lemos Fróes, a Brazilian botanist (1891 – 1960) who collected the first two specimens of E. froesii in 1955 – 56, along with many other species of Eperua. Vernacular names: — Ipê do Bento (Prance G. T. et al. 1458).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F16FFFEFF6CF897FAFCFC2F.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua froesii, E. cerradoensis, E. duckeana, E. reddeniae, and E. schomburgkiana are part of the group with short and erect inflorescences, non-tubular corolla, exserted stamens, falcate fruit with a length / width ration of 3: 1 (vs. tubular corolla, included stamens, non-falcate fruit with a length / width ratio of 1: 1 to 2: 1 in other species with short and erect inflorescence). They also share multijugate leaves with straight leaflets and inflorescence with dense indumentum. E. froesii shares characteristics such as stamens joined in a glabrous tube with E. cerradoensis, and glabrous ovary with E. schomburgkiana. Please refer to Table 3 for a summary of the similarities and differences among these species. Eperua froesii is this group`s only species with entirely glabrous stamens, ovary, and fruits. It was previously identified as E. schomburgkiana mainly by the glabrous ovary and fruit. However, besides the allopatric distribution (E. froesii is restricted to the Eastern Amazonia, and E. schomburgkiana occurs in the Guyana Shield, see Figure 8), E. froesii is a different species by the entirely joined stipules (vs. free in E. schomburgkiana), stamens joined in a tube (vs. in a diadelphous sheath 9 + 1 in E. schomburgkiana), and glabrous stamens (vs. irregularly tomentose sheath in E. schomburgkiana). The separation of the species is also supported by phylogenetic studies using nuclear and ribosomal data (ter Steege et al. 2023 in press, Fortes et al. in prep). We chose the collection Prance G. T. et al. 1458 as the holotype over Fróes R. de L. 32452 because the former collection has more preserved flowers. Both collections could potentially be the type of E. froesii because they have glabrous stamens joined basally in a tube and glabrous ovary, which are the key characteristics to differentiate E. froesii from E. schomburgkiana and other species of the group. Eperua froesii shows some contrasting morphological characteristics within the species with short and erect inflorescence and non-tubular corollas. While most species have clearly white petals (E. cerradoensis, E. duckeana, and E. schomburgkiana) and E. reddeniae has non-white petals (pink to purple), E. froesii shows a range of petal colors from white (Prance G. T. et al. 1458) to light-red (Fróes R. de L. 32418). Another variation is the stipule type, which is entirely joined in E. froesii, ranging from foliaceous (Figure 23 d – f, 25 b, c) to non-foliaceous (Figure 25 d). Foliaceous stipules are found in flowering branches (Fróes R. de L. 32418, Prance G. T. et al. 1458), and the non-foliaceous stipules are found in flowering, fruiting and sterile branches (Fróes R. de L. 32418, 32452, Prance G. T. et al. 1458, 1652). Further collections are needed to fully understand the petal color and the stipule patterns in E. froesii. Specimens examined (Paratypes): — BRAZIL. Pará, Portel, regi „ o do Anapú, rio Pracajaí, à beira do rio, 10 October 1956, Fróes R. de L. 33051 (IAN [097476]!, K [000807978] image!). Rio Acaraí, afl. do Xingú, regi „ o onde foi feito um levantamento estatístico florestal pelo IAN, SPVEA e FAO, 29 November 1955, Fróes R. de L. 32418 (INPA [0005049]!, NY [1171155] image!, R [collection number 106671]!); Fróes R. de L. 32452 (IAN [090190]!); rios Pacajá and Muirapiranga, west bank of rio Pacajá., 15 October 1965, Prance G. T. et al. 1652 (F [V 0111469 F]!, IAN [117165]!, K [000807974, 000807976] image!, NY [01171289]!, U [0181543] image!, US [00243010]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	description	(Figures 12, 26, 28)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	type_taxon	Type: — GUYANA. Near Chodikar Landing, Amongst Acarai foothills, 250 m elev., 17 October 1952, Guppy N. G. L. 430 (holotype NY sheet I [00004378] image!, & sheet II [00004379] image!; isotypes K sheet I [000555111] image!, & sheet II [000555109] image!, RB sheet I [00539601]!, sheet II [00545384]!, US [00001145]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	materials_examined	Type: — BRAZIL: Pará: Rio Trombetas, Cachoeira Porteira, estrada que dá acesso para Perimetral Norte, km 7, mata de várzea de igarapé, 8 June 1978, Silva N. T. & Santos M. R. 4758 (holotype MG [061723]!, isotypes MO [1714824] image!, NY [00004377] image!, RB [00141558]!, US [00090950]!). Tree 7.0 – 37.0 (– 51.0) m tall. Trunk 20.0 – 60.0 cm in diameter, bark variegated to grey-brown, striate, lenticellate. Stipules free, 10.3 – 25.0 (– 30.0) × 6.9 – 14.0 (– 20.0) mm, foliaceous, or 0.8 – 1.4 × 0.4 – 0.8 mm, free, non-foliaceous, caducous. Leaves 3 – 5 - jugate; petioles 0.8 – 4.2 cm long; rachis 6.4 – 19.0 cm long; petiolules (1.3 –) 3.7 – 9.1 mm long; blades 6.0 – 15.1 × 2.6 – 7.8 cm, chartaceous to coriaceous, glabrous, epunctate to inconspicuous pellucid-punctate, discolorous, equilateral to inequilateral, lower pair ovate, sometimes lanceolate or elliptic, middle pair ovate to elliptic, sometimes lanceolate, upper pair elliptic, apex acuminate to caudate, base symmetrical to asymmetrical, sometimes slightly asymmetrical, lower pair base subcordate, obtuse, subcordate to rounded, middle pair base obtuse to rounded, upper pair base obtuse to cuneate, rare rounded, margin flat, secondary venation with one or two intramarginal veins, vein closer to the margin continuous or not, main vein straight to slightly curved, prominent to depressed on the adaxial surface, tertiary veins conspicuous to inconspicuous, areoles smaller to larger, concave. Inflorescences terminal, panicle, main axis pendulous, lateral racemes patent to erect, glabrous, 93.0 – 300.0 cm long, lateral racemes 1.7 – 5.2 cm long, alternate and spirally arranged; bracts 0.9 – 1.9 × 2.2 – 2.3 mm, oblate, cucullate, apex gland absent, glabrous, ciliate, caducous; bracteoles 2.3 – 2.9 × 1.4 – 2.5 mm, ovate, cucullate, apex gland absent, ciliate, caducous, attached to the lower portion of the pedicels; pedicel 16.7 – 27.0 mm long, 1.0 – 2.4 mm in diameter, not twisted, glabrous; buds 0.7 – 1.1 cm long, 0.4 – 0.6 cm in diameter, glabrous. Flowers: hypanthium 4.5 – 7.3 mm long, 3.5 – 7.8 mm in diameter, cup-shaped to tubular, equilateral, glabrous; sepal 1.1 – 2.2 × 0.5 – 1.2 cm, elliptic to oblong, unequal, the outer ones larger, cucullate, apex gland absent, purplish-green, glabrous; adaxial petal 1.5 – 3.0 × 2.2 – 5.0 cm, flabellate, non-tubular, apex rounded, base truncate, red, rose, pink to cream at base, cream to greenish at the base and pink marginally, glabrous; petalodia 0.5 – 1.6 × 0.4 – 0.9 mm; stamens exserted, joined basally in a tube, longer filaments 5.0 – 7.8 cm long, shorter filaments 2.3 – 4.0 cm long, tube equilateral to slightly inequilateral, 2.1 – 3.8 mm long, anthers 5.0 – 9.0 × 0.7 – 2.0 mm, rectangular; ovary 5.2 – 12.5 × 2.2 – 3.5 mm, oblanceolate, glabrous, 5 ovules, stipe 5.5 – 10.9 mm long, glabrous, style 4.2 – 6.0 cm long, stigma capitate to obtuse. Legumes 17.0 – 30.5 × 4.8 – 10.0 cm, stipe 2.1 – 4.6 cm long, falcate-elliptic to falcate-oblanceolate, apex obtuse, apiculate, margin thickened, sometimes dorsal margin alate, with transversal veins or veins absent, glabrous, dark-brown to reddish-brown. Seeds 4 per fruit, 4.6 – 5.8 × 2.3 – 2.6 cm, oblate, oblong, dark-brown. Phenology: — Flowering in February, May to July, September, and October; fruiting in January, May, and from September to November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	distribution	Distribution: — Brazil (Pará and Roraima), Guyana (Upper Takatu-Upper Essequibo, East Berbice-Corentyne, and Upper Demerara-Berbice). One sterile collection made further in Surinam (Oldenburger F. H. F. et al. 1203) was sampled in the full ribosomal phylogeny for the genus (ter Steege et al. 2023 in press) and was nested within Eperua glabra specimens. It resembles E. glabra by the vegetative characteristics, but fertile collections are needed in Surinam to morphologically confirm the presence of the species in this country. Habitat: — Upland (terra-firme) and inundated (gallery, and igapó) forests on clayey, brown and white mixed sandy, and brown sandy soils, from 200 to 250 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	description	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The occurrence extent of estimated for the species is 316,238.548 km 2 and the estimated area of occupancy is 40.000 km 2. Occurrence in protected areas: — Parque Nacional Montanhas do Tumucumaque (Brazil), Mabura Hill Forest Reserve (Guyana, Upper Demerara-Berbice).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	etymology	Etymology: — The specific epithet relates to its entirely glabrous inflorescences and flowers. Vernacular names: — Brazil: espadeiro (Rodrigues I. A. et al. 1000). Guyana: wallaba (Redden K. M. et al. 1054) [although it is more often used to refer to E. falcata], wataba (Redden K. M. et al. 1054), wataja (Guppy N. G. L. 430).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua glabra and E. praesagata were described by Cowan (1957, 1985, respectively). These are the only entirely glabrous species within Eperua. They have long and pendulous inflorescence, non-tubular corolla, and exserted stamens, and share multijugate leaves with equilateral leaflets with E. rubiginosa. In Cowan`s (1985) diagnosis of E. praesagata, its differentiation from E. glabra was based mainly on organ sizes (including filaments, anthers, stipules, and petioles sizes), besides the number of leaflets, and petal colour (see Table 5), but the key characteristics in the genus that allow to differentiate species are related to organ indumentum and bracteole position. Moreover, the organ size variations and leaflets number are not sufficient for distinguishing between the two species, particularly due to the limited number of collections available. Apart from the holotypes of both species, there are three additional flowering collections with entirely glabrous inflorescences and flowers. However, it is not possible to confirm if it is E. praesagata or E. glabra because these new collections have overlapping characteristics between both species (see Table 5). There are seven fruiting collections (including the type collection of E. glabra, and one paratype of E. praesagata) with glabrous fruits in glabrous, long, and pendulous inflorescences. Still we did not recognize a pattern that allows us to separate them into two species. The type locality of E. glabra is the Acarai Mountains on the Brazil-Guyana border, while the type locality of E. praesagata is in the Trombetas River in Oriximiná, Pará, Brazil (see Figure 12). Indeed, Cowan (1985, pg. 293) reported that the type locality of E. glabra is at a “ relatively short distance north of the type locality ” of E. praesagata. Subsequent collections of entirely glabrous specimens were made in various locations, including at the type locality of E. glabra, in the Acari Mountains and Chokidar river (Henkel T. W. et al. 4655, Redden K. M. et al. 3187, 3192, Smith A. C. 2882); further North from Brazil / Guyana border in the Mabura Hill Forest Reserve region (Polak A. M. 218, Redden K. M. et al. 1054); also, in Marapi River near the type locality of E. praesagata (Pena B. S. 499,); and one collection further West in Roraima state (Brazil), but also near the border between Guyana and Brazil (Rodrigues I. A. et al. 1000). These two species are indeed sympatric and occupy the same habitat. Collections were made in upland (terra-firme) and inundated (várzea, gallery) forests in the Acarai Mountains and the Chokidar river. The holotype of E. praesagata (Silva N. T. & Santos M. R. 4758) was collected in inundated forests, but the paratype (Cid-Ferreira C. A. et al. 1419) is from upland forests. In conclusion, there is no reason to keep E. praesagata as a different species from E. glabra: their morphology overlaps, as well as their distributions and habitat; and they appeared paraphyletic in the phylogeny when considered different taxa (ter Steege et al. 2023 in press, Fortes et al. in prep.,). For all these reasons, they are synonymized here. There is one collection, Silva F. A. et al. 869 (MG [245424]!), from Januari River in the Estaç „ o Ecológica Gr „ o- Pará, Roraima, Brazil, which bears a resemblance to E. glabra by the long and pendulous inflorescence with glabrous main axis, and glabrous stamens and ovary. However, it differs from E. glabra by its puberulent lateral racemes and bracts, puberulous bracteoles, pedicel, buds, and hypanthium, and puberulent to glabrescent sepals (Figure 27). Phylogenetically, this collection is nested within E. glabra (Fortes et al. in prep.). Further collections and studies are needed to determine this collection’s taxonomic rank.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F1DFFE3FF6CFC1AFB0DFBBF.taxon	materials_examined	Selected specimens: — BRAZIL. Pará, Oriximiná, rio Trombetas, Tapagem, margem direita a 15 Km acima da mineraç „ o Santa Patrícia, 09 July 1980, Cid-Ferreira C. A. et al. 1419 (EAFM, INPA, MG, MO, NY, RB, US); Distrito de Marapi, rio Marapi, ponto 32, 07 September 1974, Pena B. S. 499 (IAN). Roraima, S „ o Jo „ o da Baliza, BR 210, Km 135, a 6 Km do rio Jatapú, lado esquerdo, 0 ° 44 ’ 14 ” N 59 ° 15 ’ 15 ” W, 16 January 1983, Rodrigues I. A. et al. 1000 (IAN). GUYANA. Upper Demerara-Berbice, Waraputa compartment, c. 25 km S of Mabura, 5 ° 15 ’ 0 ” N 58 ° 45 ’ 0 ” W, 09 January 1991, Polak A. M. 218 (U); Mabura Hill Forest Reserve, 5 ° 9 ’ 33.8 ” N 58 ° 41 ’ 51.8 ” W, 20 October 2002, Redden K. M. et al. 1054 (U, US). Upper Takatu-Upper Essequibo, Acarai Mts, Watuwau Creek 8 – 10 km upstream of juncture with Chodikar River, 1 ° 22 ’ 0 ” N 58 ° 42 ’ 0 ” W, 22 February 1994, Henkel T. W. et al. 4655 (CAY, U, US); Chodikar River, 239 m elev., 1 ° 23 ’ 39.3 ” N 58 ° 47 ’ 7.5 ” W, 04 May 2004, Redden K. M. et al. 3187 (NY, US); Chodikar River, 243 m elev., 1 ° 24 ’ 53.3 ” N 58 ° 47 ’ 43.2 ” W, 04 May 2004, Redden K. M. et al. 3192 (US); basin of Chodikar Creek (Essequibo tributary), 1 ° 18 ’ 0 ” N 58 ° 49 ’ 59 ” W, 8 – 22 January 1938, Smith A. C. 2882 (F, P, U, US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F00FFE0FF6CFBEBFAACFDE2.taxon	materials_examined	Type (designated by Cowan 1975): — BRAZIL. Amazonas: Estrada do Aleixo, 3 April 1936, Ducke W. A. 158 (lectotype US [00001143]!, isolectotypes A not seen, F [0057801] image!, K [000555105] image!, MO [954335] image!, NY [00004376] image!, R sheet I [000054528]!, & sheet II [R 000054528 a]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F00FFE0FF6CFBEBFAACFDE2.taxon	description	Tree 3.0 – 37.5 m tall. Trunk 10.0 – 54.3 cm in diameter, bark dark-yellow-brown, yellowish, brownish, lenticellate to slightly scaled. Stipules 1.3 – 3.7 × 0.7 – 2.7 mm, free, non-foliaceous, caducous to persistent. Leaves 2 (3) - jugate; petioles 0.9 – 6.4 cm long; rachis 1.7 – 15.5 cm long; petiolules 4.0 – 13.8 mm long; blades 6.2 – 23.0 × 2.7 – 17.0 cm, coriaceous, glabrous, epunctate, rarely pellucid-punctate, not discolorous, rarely discolorous, equilateral, lower pair elliptic, sometimes ovate, rarely broadly ovate, smaller, middle pair elliptic, upper pair elliptic, rarely broadly elliptic, obovate or ovate, apex acuminate, mucronate, base symmetrical, lower pair base obtuse to rounded, rarely subcordate to cordate, middle pair base obtuse to subcordate, upper pair base obtuse, sometimes rounded, rarely subcordate or cuneate, margin revolute, sometimes flat to slightly revolute at the base, secondary venation with two intramarginal veins, vein closer to the margin continuous, main vein straight, depressed on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal or axillary, sometimes on ramiflorous, panicle, rarely raceme, erect, glabrous, 3.0 – 14.9 cm long, lateral racemes 1.0 – 6.4 (– 10.6) cm long, alternate and distichous, sometimes spirally arranged; bracts 4.7 – 5.3 × 5.6 – 7.2 mm, broadly ovate, concave, apex gland present, glabrous, caducous; bracteoles 5.3 – 10.3 × 3.1 – 6.5 mm, ovate, rarely broadly ovate, cucullate, apex gland present, glabrous, caducous, attached to the lower middle portion of the pedicels; pedicel (2.0 –) 3.0 – 9.3 mm long, 0.9 – 3.4 mm in diameter, not twisted, glabrous; buds 0.7 – 1.7 cm long, 0.4 – 0.9 cm in diameter, glabrous. Flowers: hypanthium (1.2 –) 2.0 – 4.2 mm long, (1.2 –) 2.1 – 7.0 mm in diameter, cup-shaped, inequilateral, glabrous; sepal 0.8 – 2.0 × 0.3 – 1.1 cm, elliptic to oblong, unequal, the outer sepals more robust, the dorsal one larger, cucullate, apex gland present on the outer ones, purple, sometimes pale-purple to pink-reddish, or greenish, glabrous, pellucid-punctate, inner sepals scarious marginally; adaxial petal 2.4 – 5.6 × 2.1 – 6.7 cm, flabellate, tubular, apex rounded, cripsed, base attenuate, purple, lilac to red or purple, red, greenish to white, glabrous; petalodia 0.4 – 3.5 × 0.2 – 0.8 mm, white to pink with inner white; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 2.0 – 4.2 cm long, shorter filaments 1.4 – 2.8 cm long, sheath inequilateral, shorter side 2.7 – 10.0 mm long, longer side 4.4 – 13.6 mm long, sheath tomentose, pubescent to puberulous, free filaments glabrous, brownish-yellow, shorter stamens anthers 1.1 – 2.8 × 0.5 – 1.8 mm, rectangular, longer stamens anthers 1.7 – 3.6 × 0.6 – 2.2 mm, rectangular; ovary 3.3 – 7.5 × 1.7 – 4.4 mm, obovate, glabrous, fovoate or tomentose, brownish-yellow, 2 – 4 ovules, stipe 3.1 – 7.7 mm long, glabrous or tomentose, style 1.9 – 3.5 cm long, stigma obtuse to capitate. Legumes 3.5 – 15.4 × 5.2 – 13.8 cm, stipe 0.9 – 2.6 cm long, obovate to orbicular, sometimes oblongelliptic, apex obtuse to truncate, sometimes rounded, dorsal margin thickened, brown to dark-brown, veins absent, rarely with some inconspicuous transversal veins, glabrous, foveate to rugose. Seeds 1 per fruit, 4.3 – 10.8 × 3.8 – 7.7 cm, oblate, orbicular to obovate, brown to dark-brown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F00FFE0FF6CFBEBFAACFDE2.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 52,541.927 km 2 and the estimated area of occupancy is 228.000 km 2.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F00FFE0FF6CFBEBFAACFDE2.taxon	discussion	Eperua glabriflora has two varieties: E. glabriflora var. glabriflora and E. glabriflora var. gynopubescens.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	description	(Figures 10, 29, 30) Sepal purple, sometimes pale-purple to pink-reddish; Adaxial petal purple, lilac to red; ovary glabrous, fovoate, stipe glabrous. Phenology: — Flowering throughout the year, except in October, with a peak in July; fruiting from February to March and from May to November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	distribution	Distribution: — Brazil (Amazonas, one collection in Pará). Habitat: — Upland forest, both in terra-firme forest and campinarana, on the sand and clayish soils, from sea level to 125 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	description	Occurrence in protected areas: — Reserva Florestal Adolpho Ducke, Reserva Biológica de Uatum „, Reserva Florestal ZF- 3 (Brazil, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	etymology	Etymology: — Eperua glabriflora was first described as a variety of E. bijuga. The glabrous pedicel, hypanthium, and sepals of E. glabriflora differentiate it from the type variety, then the epithet glabriflora probably derives from that characteristic.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	vernacular_names	Vernacular names: — espadeira (Ducke W. A. s. n. RB 20313), morapiranga (Boom B. M. et al. 8699), muirapiranga (Cid-Ferreira C. A. et al. 271, Cid-Ferreira C. A. et al. 4, Coêlho L. F. s. n. INPA 6516, Ducke W. A. 14, 14 a, 158 (IIª col), s. n. RB 23733, Elias J. & Aloisio 462, Ferreira E. 57 - 29, Haroldo s. n. INPA 57650, Jorge s. n. INPA 57656, INPA 57656, Krukoff B. A. 8006, Magalhães F. M. M. 223, Mello F. C. de s. n. INPA 55353, Monteiro O. P. & Mello F. s. n. INPA 50065, Oliveira A. A. de et al. 1032, Oliveira A. R. de s. n. INPA 58701, Pessoal do C. P. F. INPA 6065, INPA 6280, Prance G. T. et al. 23446, Reis L. Q. s. n. INPA 57645, INPA 57694, Ribeiro J. E. L. S. 881, Rodrigues W. A. & Osmarino 6932, Rodrigues W. A. & Osmarino 7005, Rodrigues W. A. & Osmarino 7903, Rodrigues W. A. 562, s. n. INPA 1043, Silva S. S. da et al. 3402.2791.2), muirapiranga da folha grande (Oliveira A. R. de collections, Rodrigues W. A. & Osmarino 5982, Rodrigues W. A. & Osmarino 7921).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	description	Uses: — general construction (Rodrigues W. A. s. n. INPA 1043). Taxonomic notes: — Eperua glabriflora, along with E. grandiflora subsp. ciliata and E. purpurea are the only species with tubular corolla and glabrous inflorescences. Eperua glabriflora differs from them and all other species by the combination of 2 - jugate leaves, non-falcate leaflets with revolute margin, tubular corolla, glabrous inflorescence, and caducous bracteoles attached to the lower middle portion of the pedicels (Table 2). Eperua glabriflora resembles E. bijuga by the corolla size and shape, 2 - jugate leaves, and the non-falcate leaflets with two intramarginal veins. However, it differs by the revolute leaflet margins (vs. flat in E. bijuga); glabrous inflorescence, buds, hypanthium, and sepals (vs. puberulous to puberulent in E. bijuga); larger and caducous bracteoles (vs. smaller and persistent in E. bijuga); and shorter obovate ovary (vs. longer and falcate in E. bijuga).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F03FFE6FF6CFDC1FB25F80A.taxon	materials_examined	Specimens examined: — BRAZIL. Amazonas, Itacoatiara, Km 75 – 70 da rod. Manaus-Itacoatiara, 17 October 1963, Oliveira E. 2709 (IAN). Itapiranga, rio Uatum „, margem esquerda, subindo o rio, atrás do igarapé do Catitu, 18 July 1979, Cid-Ferreira C. A. et al. 484 (INPA, MG, NY, RB 00140230, US). Manaus, Distrito Agropecuário da SUFRAMA rodovia BR- 174, km 64 depois de 23 km leste na ZF 3, fazenda Esteio, reserva no. 1202, arv. No. 3150, 2 ° 24 ’ 0 ” S 59 ° 52 ’ 0 ” W, 03 July 1980, Costa J. C. da et al. S. n INPA 1202.3150 (NY); Reserva ZF 3, BR 174, Km 63, área do Projeto TEAM, parcela do Cabo Frio sub-parcela 16, indivíduo n ° 444, 24 August 2004, Oliveira et al. A. C. A. 369 (INPA); Reserva Florestal Adolpho Ducke, quadra 21, tree # 2896, 07 May 1965, Rodrigues W. A. & Osmarino 6932 (INPA, US). Novo Air „ o, km 23, KM 19, 18 November 2012, Kinupp V. F. et al. 4552 (EAFM). Presidente Figueiredo, estrada da UHE de Balbina, entrando 8 Km no vicinal ao lado do campo de pouso, 1 ° 30 ’ 0 ” S 60 ° 0 ’ 0 ” W, 22 September 1986, Cid-Ferreira C. A. et al. 8239 (INPA, MG, MO, NY, UFACPZ, US); ramal ZF 5 (SUFRAMA), BR- 174, Km 83, 2 ° 25 ’ 23 ” S 60 ° 17 ’ 36 ” W, 11 June 2011, Marra D. M. et al. 584 (EAFM); Rebio Uatum „, ramal da Morena, corredeira da Anta próximo ao Projeto de Desenvolvimento Sustentável (PDS), 1 ° 0 ’ 0 ” S 59 ° 0 ’ 0 ” W, 06 July 2007, Sakagawa S. & Mesquita J. R. 366 (INPA, RB 00716854). Pará, Lago do Faro, cabeceira de Inajá, lugar húmido e ligeiramente [...]?, 27 January 1927, Ducke W. A. s. n. RB 20313 (RB, U). Rio de Janeiro (cultivated), Rio de Janeiro, Jardim Botânico do rio de Janeiro, Arboreto canteiro 26 ª, 04 February 2020, Fortes E. A. & Camelo M. De C. 150 (RB).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	description	(Figures 10, 31)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	materials_examined	Type: — BRAZIL. Amazonas, Itapiranga, rio Pitinga, margem direita, 24 August 1979, Cid-Ferreira C. A. et al. 659 (holotype INPA [0034257]!, isotypes MG [076037]!, MO [1780613] image!, NY [1171206] image!, R [000049194]!, RB [00266154], US [00242870] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	diagnosis	Diagnosis: — Eperua glabriflora var. gynopubescens is similar to var. glabriflora, except for the indumentum of the ovary. The new variety can be differentiated from the type one by the flowers with glabrous or tomentose ovary in the same inflorescence (vs. always glabrous in var. glabriflora). Sepal greenish; Adaxial petal purple, red, greenish to white; ovary and stipe tomentose or glabrous in the same inflorescence.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	description	Phenology: — Flowering in April, August, and September; fruiting in August.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	distribution	Distribution: — Brazil (Amazonas). Habitat: — Campinarana, gallery and terra-firme forests.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	description	Occurrence in protected areas: — Reserva Florestal Adolpho Ducke (Brazil, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	etymology	Etymology: — The specific epithet relates to its tomentose ovary. Vernacular names: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	description	Uses: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE7FF6CFF33FBAFFC07.taxon	description	Specimens examined: — BRAZIL. Amazonas, Manaus, Reserva Florestal Adolpho Ducke, Manaus-Itacoatiara, km 26, ramal ao Acará, 2 ° 53 ’ 0 ” S 59 ° 58 ’ 0 ” W, 25 April 1988, Ramos J. F. 1855 (HUEFS [0197835] image!, INPA [0179539]!, MG!, MO [1721976]!, NY [1171186] image!, SP [024676] image!, RB [00140240]!, U [0061219] image!); rio Tarumá, 07 August 1949, Fróes R. De L. 24926 (IAN [050817, 50817 ª]!). Pedra Branca, rio Urubú, 17 September 1949, Fróes R. De L. 25266 (IAC [collection number 25266], IAN [144635]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE8FF6CFC6BFCC7FCBA.taxon	description	(Figures 9, 32, 33, 34)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE8FF6CFC6BFCC7FCBA.taxon	materials_examined	Type (designated by Cowan 1975): — FRENCH GUIANA. Unknown locality, s. d. [July 1762 – July 1764], Aublet J. B. C. F. s. n. (lectotype BM [000952282] image!, isolectotypes P [00835933] image!, P [00835934] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE8FF6CFC6BFCC7FCBA.taxon	materials_examined	Type (designated by Fortes & Mansano 2022): — FRENCH GUIANA Gourdonville, Kourou R., 8 September 1914, Benoist R. 1595 (lectotype P [00835930] image!, isolectotypes, P [00835931] image!, P [00835932] image!). Tree 3.0 – 40.0 m tall. Trunk 20.0 – 100.0 cm in diameter, bark gray, greyish-black, reddish, red, lenticellate, smooth to caled. Stipules 1.9 – 11.3 (– 27.1) × 1.3 – 10.0 (– 21.2) mm, free, foliaceous, sometimes non-foliaceous, caducous to persistent. Leaves 2 – 3 (– 4) - jugate; petioles 0.6 – 2.6 cm long; rachis 3.0 – 11.7 cm long; petiolules 1.4 – 9.3 mm long; blades 3.8 – 15.8 × 1.6 – 6.9 (– 9.6) cm, coriaceous, glabrous, epunctate, inconspicuous pellucid-punctate to minutely pellucid-punctate, discolorous to not discolorous, equilateral, lower pair ovate, elliptic to lanceolate, middle pair narrowly elliptic to elliptic, upper pair elliptic to narrowly elliptic, apex attenuate, acuminate to caudate, base symmetrical to oblique, par of glands absent or with one pair of glands, lower pair base obtuse, rounded to subcordate, middle pair base obtuse to rounded, upper pair base obtuse, margin flat to slightly revolute, rarely very revolute or very revolute, rarely flat, secondary venation with one or two intramarginal veins, vein closer to the margin continuous or not, main vein straight, prominent to depressed on the adaxial surface, tertiary veins conspicuous to inconspicuous, areoles smaller, concave. Inflorescences terminal or axillary, panicle or raceme, erect, pubescent to tomentose, sometimes puberulous, or pubescent, puberulous to glabrescent, or glabrous, brownish-yellow, 2.3 – 9.8 cm long, lateral racemes 1.2 – 6.0 cm long, alternate and distichous; bracts 2.1 – 4.7 × 1.6 – 4.0 mm, ovate to broadly ovate, cucullate, apex gland present or absent, tomentose, pubescent to puberulous externally, glabrous within, or glabrous, ciliate, brownish-yellow, caducous to persistent; bracteoles 2.3 – 3.9 × 2.0 – 3.7 mm, or 1.9 – 3.0 × 1.5 – 3.4 mm, or 4.7 – 5.6 × 3.8 – 5.9 mm, ovate to broadly ovate, cucullate, apex gland present or absent, pubescent to tomentose, rarely puberulous externally, glabrous within, or pubescent to puberulous on both sides to glabrous within, brownish-yellow, persistent, attached to the higher portion of the pedicels or at the base of the hypanthium, at slightly different levels; pedicel 3.0 – 10.3 mm long, 0.5 – 1.6 mm in diameter, not twisted, puberulous, pubescent to tomentose, or glabrous, brownish-yellow; buds 0.4 – 1.3 cm long, 0.3 – 0.6 cm in diameter, pubescent to puberulous, sometimes glabrescent, or glabrous, brownish-yellow. Flowers: hypanthium 1.5 – 2.8 mm long, 1.9 – 4.1 mm in diameter, cup-shaped, inequilateral to equilateral, tomentose, pubescent to puberulous, or glabrous, brownish-yellow; sepal 0.5 – 1.4 × 0.2 – 0.8 cm, elliptic, ovate, lanceolate to oblanceolate, unequal, the dorsal one larger, cucullate, apex gland present or absent, green, purple-green, purplish, brown-purple, brown to burgundy, pubescent, puberulous to glabrescent, or glabrous, trichomes brownish-yellow; adaxial petal 1.5 – 4.3 × 1.4 – 5.1 cm, flabellate, tubular, apex rounded, base attenuate, pale-pink, pink, purple, pale-purple, lilac, whitish inner, to white with purple guide nectar, glabrous; petalodia 0.2 – 1.2 × 0.2 – 0.5 mm; stamens, inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 0.9 – 2.7 cm long, shorter filaments 0.5 – 1.3 cm long, sheath inequilateral, shorter side 2.5 – 5.0 mm long, longer side 4.0 – 11.0 mm long, sheath irregularly tomentose, pubescent to puberulous, sometimes irregularly villous to entirely villous, or pubescent to puberulous, or irregularly villous, free filament glabrous, free filament glabrous, brownish-yellow, anthers 1.1 – 2.6 × 0.3 – 1.1 mm, rectangular; ovary 1.7 – 5.4 × 1.3 – 3.7 mm, obovate, glabrous or sericeous, stipe 1.6 – 5.1 mm long, glabrous, style 1.2 – 2.1 cm long, stigma obtuse, papillate. Legumes 6.0 – 12.4 × 4.3 – 9.2 cm, stipe 0.8 – 1.4 cm long, oblong, elliptic to obovate, apex truncate, obtuse to rounded, margin thickened, dark-brown, transversal veins present or absent, glabrous or puberulous, very rugose. Seeds 1 – 2 per fruit, 3.4 – 4.5 × 4.9 – 7.4 cm, oblate, dark-brown. Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 378,125.077 km 2 and the estimated area of occupancy is 208.000 km 2.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F04FFE8FF6CFC6BFCC7FCBA.taxon	description	Eperua grandiflora has three subspecies presented here: E. grandiflora subsp. grandiflora, E. grandiflora subsp. guyanensis, and E. grandiflora subsp. ciliata subsp. nov.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	description	Leaves, margin flat to slightly revolute, rarely very revolute. Inflorescences pubescent to tomentose, sometimes puberulous; bracts tomentose, pubescent to puberulous externally, glabrous within; bracteoles 2.3 – 3.9 × 2.0 – 3.7 mm, pubescent to tomentose, rarely puberulous externally, glabrous within; pedicel puberulous, pubescent to tomentose; buds pubescent to puberulous, sometimes glabrescent. Flowers: hypanthium tomentose, pubescent to puberulous; sepal pubescent, puberulous to glabrescent; stamens sheath irregularly tomentose, pubescent to puberulous, sometimes irregularly villous to entirely villous; ovary glabrous. Legumes glabrous. Phenology: — Flowering in January, April, May, and from July to November; fruiting from January to April, in July, and from September to December.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	distribution	Distribution: — Brazil (Amapá), French Guiana (Cayenne, Saint-Laurent-du-Maroni), Guyana (Upper Demerara-Berbice), and Surinam (Sipaliwini). Habitat: — Mainly in upland forests, sometimes in floodplains, from sea level to 990 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	description	Occurrence in protected areas: — Parque Nacional Montanhas do Tumucumaque (Brazil, Amapá), Réserve Naturelle Nationale de la Trinité (French Guiana, Saint-Laurent-du-Maroni), Parque Nacional do Monte Roraima (Brazil-Roraima, Guyana- Cuyuni-Mazaruni), Mabura Hill Forest Reserve (Guyana, Upper Demerara-Berbice).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	etymology	Etymology: — The specific epithet relates to the species bigger flower compared with Parivoa tomentosa Aubl. (synonym of Crudia tomentosa (Aubl.) J. F. Macbr.). But, after the combination of Pariova into Eperua the epithet became meaningless, as E. jenmanii is the species that has the biggest flowers in the genus. Vernacular names: — apá (Dambrós L. A. 307, Rabelo B. V. et al. 3308), bii adu (Djuka) (Lescure J. P. 746), wap seinı (Grenand P. 3082), wapa (Thiel J. collections), wapa tapaka (Way „ pi) (Lescure J. P. 746).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua grandiflora differs from other species with short and erect inflorescence and tubular corolla by the combination of the following characteristics: free stipules, intramarginal vein not very close to the margin, inflorescence with dense indumentum (subsp. grandiflora and subsp. guyanensis) or glabrous (subsp. ciliata), persistent bracteoles inserted high on the pedicels or at the base of the hypanthium, and non-glabrous sheath (Table 2). According to Cowan (1975), the type subspecies differs from the subsp. guyanensis mainly by the flat leaflet margins (vs. revolute in subsp. guyanensis), glabrous ovary (vs. sericeous in subsp. guyanensis), and its distribution range along the Brazil-French Guiana border, coastal French Guiana and south-central Suriname (vs. central to south-central Guyana). Most specimens of each subspecies follow the pattern above, but we found two specimens with intermediary morphotypes: Pipoly J. J. & Boyan R. 8799 from Guyana, with revolute leaflet margins and glabrous ovary, and Redden K. M. et al. 1053 from Guyana, with flat leaflet margins and puberulous fruits. Considering only the indumentum of the ovary, the first collection can be identified as subsp. grandiflora, and the second one as subsp. guyanensis (puberulous fruits indicate a non-glabrous ovary). In the phylogenetic analysis, the subspecies appeared as a separated lineage, with the specimen Redden K. M. et al. 1053 nesting within subsp. guyanensis (Fortes et al in prep). Although this is a phylogenetic signal, there is an overlap in the distribution range and the vegetative characteristics, which justify the maintenance of subsp. guyanensis as an infraspecific category of E. grandiflora. Most specimens of E. grandiflora subsp. grandiflora has irregularly tomentose, pubescent to puberulous sheaths, but we found one specimen with irregularly villous sheath (W. J. Hahn 3744) and one with entirely villous sheath (Goldenberg R. et al. 2632, Figure 34). These specimens were recorded in the same distribution range as the type subspecies. These described characteristics do not support them as a different species or an infraspecific category of E. grandiflora. Additionally, in the phylogenetic analysis, these two collections appeared as sister groups within E. grandiflora subsp. grandiflora (Fortes et al. in prep).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0BFFE9FF6CFCE7FD74FB6B.taxon	materials_examined	Selected specimens: — BRAZIL. Amapá, Lourenço, Calçoene, rio Araguari, 2 ° 13 ’ 0 ” N 51 ° 58 ’ 0 ” W, 02 September 1961, Pires J. M. et al. 50673 (F, IAN, K, MG, NY, R, S, SP, U, US). Pedra Branca do Amapari, Parque Nacional Montanhas do Tumucumaque, rio Amapari, acima da base Jupará, 1 ° 11 ’ 16 ” N 52 ° 22 ’ 8 ” W, 15 September 2018, Goldenberg R. et al. 2632 (RB, UPCB). Porto Grande, Cupixi, a cerca de 40 km a sudoeste da localidade de Serra do Navio, proximidade do rio Cupixi, 13 m elev., 0 ° 37 ’ 0 ” N 52 ° 10 ’ 0 ” W, 19 November 1983, Dambrós L. A. 307 (HRB, INPA, RB). Serra do Navio, Parque Nacional das Montanhas do Tumucumaque, margens do rio Amapari acima da base do ICMBio, 115 m elev, 1 ° 10 ’ 26 ” N 52 ° 21 ’ 34 ” W, Forzza R. C. et al. 9951 (RB, HAMAB, UPCB, NY). FRENCH GUIANA. Cayenne, Cayenne, Quesnel-Chantier F. R. G., 10 December 1979, Thiel J. 603 (CAY); exploitation forestière F. R. G., km 8, 10 March 1981, Thiel J. 815 (CAY). Kourou, rivière Kourou, bassin du Kourou, 15 October 1981, Thiel J. 890 (CAY); montagne des Singes - 1, km 8, région littorale, 21 October 1981, Thiel J. 900 (CAY). Macouria, Matiti, Point-Quadrat MAT, 5 ° 0 ’ 0 ” N 52 ° 38 ’ 0 ” W, 20 November 2001, Prévost M. F. 4278 (CAY, MO). Montsinéry-Tonnegrande, Montagne des Chevaux, Carrière SCC, 28 m elev., 4 ° 44 ’ 34 ” N 52 ° 26 ’ 12 ” W, 11 January 2009, Tostain O. et al. 2457 (CAY). Régina, station des Nouragues (camp inselberg), trail CE near camp, 20 – 40 m elev., 4 ° 5 ’ 30 ” N 52 ° 41 ’ 0 ” W, 08 August 2008, Geiselman C. K. 165 (NY). Roura, route de l’Est, km 0.5, bassin de l’Orapu, 23 April 1981, Thiel J. 826 (CAY). Saint-Élie, piste de St. Elie, CD- 21, 100 m elev., 5 ° 0 ’ 0 ” N 53 ° 10 ’ 0 ” W, 15 September 1987, Hahn W. J. 3744 (CAY, U, US, WAG). Sinnamary, crique Paracou, 03 November 1996, Hallé F. 4425 (MPU, U); piste de Saint-Élie, km 11.5, 14 October 1980, Thiel J. 736 (CAY). Gourdonville, unknown locality, 08 September 1914, Benoist R. 1595 (P). Saint-Laurent-du-Maroni, Mana, St-Elie Réserve Naturelle des Montagnes de la Trinité-Plateau- Tabulaire, bassin de la Mana, 400 – 630 m elev., 4 ° 35 ’ 0 ” N 53 ° 21 ’ 0 ” W, July 1999, Poncy O. 2652 (CAY). GUYANA. Upper Demerara-Berbice, Mabura hill and vinicity, 50 – 60 m elev., 5 ° 25 ’ 0 ” N 58 ° 40 ’ 0 ” W, 5 ° 25 ’ 0 ” N 58 ° 40 ’ 0 ” W, Pipoly J. J. & Boyan R. 8799 (U, US). SURINAM. Sipaliwini, Nickerie, Fallawatra, 18 January 1972, Jimenez-saa J. H. 1698 (L. B. B. 14431) (U).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	materials_examined	Type: — GUYANA. Bartica-Potaro. 83 mi Bartica-Potaro road, 23 June 1933, Tutin T. G. 245 (holotype US 00001146!, isotypes, BM 000952283 image!, K 000555101 image!, U 0003225 image!). Leaves margin very revolute, rarely flat. Inflorescences pubescent, puberulous to glabrescent; bracts tomentose, pubescent to puberulous externally, glabrous within; bracteoles 1.9 – 3.0 × 1.5 – 3.4 mm, pubescent to puberulous on both sides to glabrous within; pedicel puberulous, pubescent to tomentose; buds pubescent to puberulous, sometimes glabrescent. Flowers: hypanthium tomentose, pubescent to puberulous; sepal pubescent, puberulous to glabrescent; stamens sheath entirely pubescent to puberulous; ovary sericeous. Legumes puberulous.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	description	Phenology: — Flowering from February to April, in June, July, and from September to November; fruiting in January, April, May, August, and October.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	distribution	Distribution: — Guyana (Cuyuni-Mazaruni, Essequibo Islands – West Demerara, Pomeroon-Supenaam, Potaro-Siparuni, Upper Demerara-Berbice) and Surinam (Nickerie). Habitat: — White sand ecosystems in wallaba forest, from sea level to 200 m elev, on white and brown sand, sometimes on white sand-brown loamy sand-bauxite mosaic.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	description	Occurrence in protected areas: — Mabura Hill Forest Reserve (Guyana, Upper Demerara-Berbice), Pibiri Forest Reserve (Guyana, Upper Demerara-Berbice).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	etymology	Etymology: — The specific epithet relates to its distribution in Guyana. Vernacular names: — ituri-wallaba (Ek R. C. & Zagt R. 855, Forest Department 2311, Redden K. M. et al. 1053, Sandwith N. Y. 314, Tutin T. G. 245), wapa-courbaril (Coup C. CL 203).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	description	Uses: — Guyana: firewood (Tutin T. G. 245), light poles (Redden K. M. et al. 1053), fences, posts, electricity poles, and roof shingles (H. ter Steege pers. obs.). Taxonomic notes: — See taxonomic notes of the type subspecies.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0AFFEEFF6CFB57FCDFFE5B.taxon	materials_examined	Selected specimens: — GUYANA. Cuyuni-Mazaruni, Bartica, Essequibo River, Moraballi Creek, near Bartica, 20 September 1929, Sandwith N. Y. 314 (RB, U). Essequibo-Demerara Ocidental, 4 km from Santa Mision, 0 – 200 m elev., 6 ° 30 ’ 0 ” N 58 ° 23 ’ W, 25 November 1992, G ̂ rts-van Rijn A. R. A. et al. 403 (CAY, U, US). Pomeroon-Supenaam, Waraputa compartment, c. 25 km S of Mabura, 0 – 100 m elev., 5 ° 15 ’ 0 ” N 58 ° 45 ’ 0 ” W, 10 October 1991, Polak A. M. et al. 364 (U). GUYANA. Potaro-Siparuni, Kuribrong, 112 m elev., 5 ° 21 ’ 0 ” S 59 ° 15 ’ 0 ” W, April 2010, Zartman C. E. 7928 (INPA, RB). Upper Demerara-Berbice, Mabura Hill area, 150 – 200 m elev., 5 ° 20 ’ 0 ” N 58 ° 40 ’ 0 ” W, 05 June 1986, Pipoly J. J. 7560 (CAY, F, INPA, MO, P, U, US, WAG); Mabura Hill Forest Reserve, main trail, 69 m elev., 5 ° 9 ’ 33.8 ” N 58 ° 41 ’ 51.8 ” W, 20 October 2002, Redden K. M. et al. 1053 (U, US). SURINAM. Nickerie, Arkonikreek, Nickerie rivier, 28 August 1976, Teunissen P. A. 16010 (U).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	description	(Figures 9, 35, 36, 37)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	materials_examined	Type: — GUYANA. Potaro-Siparuni, Potaro River below Waratuk Falls, 107 m elev., 5 ° 16 ’ 57 ” N 59 ° 21 ’ 39 ” W, 05 September 2006, Redden K. M. et al. 4029 (holotype US [00963900]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	diagnosis	Diagnosis: — Eperua grandiflora subsp. ciliata shares with other subspecies of E. grandiflora the free stipules, 2 – 3 - jugate leaves, and persistent bracteoles attached to the higher portion of the pedicels or at the base of the hypanthium. It also shares irregularly villous sheath, and glabrous ovary and fruits with subsp. grandiflora. Eperua grandiflora subsp. ciliata differs from other subspecies by the glabrous inflorescences, pedicel, hypanthium, and sepals (vs. with indumentum in subsp. grandiflora and subsp. guyanensis), bracts and bracteoles glabrous with ciliate margin (vs. with indumentum in subsp. grandiflora and subsp. guyanensis), and larger [4.7 – 5.6 × 3.8 – 5.9 mm] bracteoles (vs. smaller [1.3 – 3.9 × 1.5 – 3.7 mm] in subsp. grandiflora and subsp. guyanensis). Leaves, margin flat. Inflorescences glabrous; bracts glabrous, ciliate; bracteoles 4.7 – 5.6 × 3.8 – 5.9 mm, glabrous, ciliate; pedicel glabrous; buds, glabrous. Flowers: hypanthium, sepal glabrous; stamens, sheath irregularly villous; ovary glabrous. Legumes glabrous.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	description	Phenology: — Flowering in September.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	distribution	Distribution: — Potaro River in the Esequibo River basin (Guyana, Potaro-Siparuni). It is sympatric with E. jenmanii, which also occurs in the same basin. Habitat: — It occurs in floodplain (riparian) forests, and wallaba forests, on white sand soils from 107 to 498 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	description	Occurrence in protected areas: — Kaieteur National Park (Guyana).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	etymology	Etymology: — The specific epithet relates to its ciliate bracts and bracteoles. Vernacular names: — unknown	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFFEEFF6CFD87FE8DF963.taxon	description	Uses: — unknown. Taxonomic notes: — Eperua grandiflora subsp. ciliata is known by only three specimens which shows some sympatry with E. grandiflora subsp. guyanensis. In the upcoming phylogeny of the genus E. grandiflora subsp. ciliata was sampled with one accession that appears as a sister group to the other two subspecies of E. grandiflora (Fortes et al. in prep.). This lineage grouping the three subspecies of E. grandiflora received maximum phylogenetic support. Specimens examined (paratype): — GUYANA. Cuyuni-Mazaruni, Pakaraima Mts., Mazaruni River, trail approx. 0.8 miles North from mining camp, 570 m ele., 6 ° 5 ’ 4 ” N 60 ° 39 ’ 2 ” W, 21 February 2004, Redden K. M. et al. 2059 (NY [03777031] image!, US [01952126]!). Potaro-Siparuni, Tukeit Landing below Kaieteur Falls on Potaro River, 498 m elev., 5 ° 12 ’ 16 ” N 59 ° 27 ’ 11 ” W, 02 September 2006, Redden K. M. et al. 3962 (CAY [109805] image!, US [00963896]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	description	(Figures 13, 38, 39)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	materials_examined	Type (designated by Cowan 1975): — GUYANA. Essequibo River, August 1884, Jenman G. S. 2154 (lectotype K [000555100] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	materials_examined	Type (designated by Fortes & Mansano 2022): Unknown locality, s. d., Martin J. s. n. (lectotype BM [000065079] image!, isolectotype F [0111407] image!)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	materials_examined	Type (designated by Fortes & Mansano 2022): SURINAM. Saramaca, May 1924, Febis 6468 (lectotype U [0003226] image!, isolectotypes U [0003227] image!, U [0003231] image!) Tree 5.0 – 30.0 m tall. Trunk 8.5 – 79.6 cm in diameter, bark gray to pale-brown. Stipules 3.7 – 25.4 (– 31.2) × 2.3 – 31 mm, joined at the base, sometimes free or entirely joined, foliaceous or non-foliaceous, persistent. Leaves 3 – 5 - jugate; petioles 0.8 – 3.0 cm long; rachis 6.0 – 20.0 (– 25.8) cm long; petiolules 6.0 – 10.4 mm long; blades 5.5 – 16.2 × 2.2 – 6.6 cm, coriaceous, glabrous, epunctate to pellucid-punctate, discolorous, equilateral, lower pair elliptic, ovate to lanceolate, middle pair elliptic to narrowly elliptic, upper pair elliptic to narrowly, apex acuminate to attenuate, mucronate, rarely caudate, base symmetrical, with one pair of inconspicuous abaxially gland, lower pair base obtuse to rounded, rarely subcordate, middle pair base obtuse to rounded, upper pair base obtuse to cuneate, margin flat to slightly revolute, secondary venation with two intramarginal veins, vein closer to the margin continuous, main vein straight, depressed, prominent near the base to entirely prominent on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave, to larger, plane. Inflorescences terminal, axillary or cauliflorous, erect, raceme, rarely panicle, puberulent or glabrous, trichomes white, 1.3 – 7.2 cm long; bracts (1.7 –) 3.0 – 5.3 (– 8.5) × (1.1 –) 3.5 – 5.8 mm, ovate, broadly ovate to oblate, cucullate, apex gland absent, puberulent externally, glabrous within or glabrous in both sides, ciliate, trichomes white, persistent; bracteoles (2.3 –) 3.6 – 9.0 × (1.0 –) 2.0 – 6.9 mm, ovate to broadly ovate, apex gland absent, puberulent externally, glabrous within or glabrous in both sides, ciliate, trichomes white, persistent, attached to the lower portion of the pedicels at slightly different levels; pedicel 4.5 – 16.9 mm long, 1.0 – 2.4 mm in diameter, not twisted, puberulent, sometimes puberulent to glabrescent or glabrous, trichomes white; buds 0.8 – 2.2 cm long, 0.4 – 1.1 cm in diameter, puberulous or glabrous, trichomes white. Flowers: hypanthium 1.8 – 4.8 mm long, 2.9 – 6.8 mm in diameter, cup-shaped, inequilateral, puberulent, sometimes puberulent to glabrescent, or glabrous, trichomes white; sepal 1.4 – 3.1 × 0.4 – 1.3 cm, oblong, elliptic to obovate, unequal, the dorsal one larger, cucullate, apex gland present on the outer ones, pink, magenta, purple to reddish-purple, puberulent to glabrescent or glabrous, scarious marginally, trichomes white; adaxial petal 3.2 – 11.7 × 4.0 – 9.6 cm, obovate to broadly obovate, tubular, apex rounded, crisped, base attenuate, gibbous, lavender, purple to pink, whitish marginally, glabrous; petalodia 0.3 – 5.9 × 0.3 – 1.3 mm, lavender to white; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 3.0 – 7.6 cm long, shorter filaments 2.2 – 5.6 cm long, sheath inequilateral, shorter side 9.1 – 15.0 (– 23.0) or 4.5 – 13.0 mm long, longer side 10.3 – 26.0 or 6.1 – 18.0 mm long, sheath irregularly villous, pubescent to puberulous, free filaments glabrous, trichomes white, shorter stamens anthers 1.4 – 3.0 × 0.8 – 1.6 mm, rectangular, longer stamens anthers 2.5 – 4.1 × 1.0 – 2.6 mm, rectangular; ovary 4.0 – 11.0 × 2.4 – 5.1 mm, obovate, glabrous, stipe 3.4 – 10.0 mm long, glabrous, style 2.5 – 4.9 cm long, stigma obtuse to capitate, papillose. Legumes 6.0 – 17.0 × 5.6 – 11.2 cm, stipe 1.0 – 2.7 cm, oblong to obovate, apex truncate, rounded to obtuse, dorsal side apiculate, dorsal margin thickened, dark redsish-brown to reddish-brown, with transversal veins, sometimes inconspicuous, glabrous, rugose. Seeds 1 – 2 per fruit, 4.6 – 6.0 × 2.5 – 3.9 cm, elliptic, ovate to oblong, reddish-brown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	distribution	Distribution: — E. jenmanii occurs in Guyana (Cuyuni-Mazaruni, Upper Demerara-Berbice), mainly in the Mazaruni River and lower Esequibo River in Esequibo River basin region, and Venezuela (Bolívar), mainly in the Caura, Paragua, and Caroni rivers in the Orinoco River basin region. There are some sterile collections in French Guiana (Sabatier D. 1879, Sauvain M. 479) and Surinam (Febis 6468, Hoffman B. 6146) that resembles E. jenmanii, but flowering and fruiting collections are needed to confirm the distribution of the species in those areas .. Habitat: — Floodplain (riverine) forests, sometimes reaches upland forests and wallaba forests, from 5 - 225 (– 1220) m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 264,200.096 km 2 and the estimated area of occupancy is 176.000 km 2.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	vernacular_names	Vernacular names: — arekuna (Cardona F. 2158), baboen walaba (Febis 6468), caraoto (Beruordi 2130, Stergios B. 11719), itoeli walaba (Stahel G. 334), tumay-yek (Steyermark J. A. 60376), opá (Cardona F. 2158), opá-yek (Steyermark J. A. 60376, 60754), water wallaba (Redden K. M. et al. 2074).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F0DFF90FF6CF95FFAE0F888.taxon	materials_examined	Specimens examined: — GUYANA. Cuyuni-Mazaruni, Essequibo River, W bank, Wolga 1 km S, to lower 0.5 km of Kamwatta Creek, 5 m elev., 6 ° 27 ’ 0 ” N 58 ° 36 ’ 0 ” W, 01 April 1993, Henkel T. W. et al. 1863 (US). Upper Demerara-Berbice, Berbice River, 230 km upriver from mouth, N of Kwakwani, Aroiama mining Co. land concession, 70 m elev., 5 ° 39 ’ 36 ” N 58 ° 0 ’ 0 ” W, May 1995, SI / MAB 60818 (US). VENEZUELA. Bolívar, Cedeño, Majawana village along the Río Majagua, tributary of the Río Erebato, 05 March 1992, Boom B. M. 10297 (NY); Ciudad Guayana, orillas de Caroni and upstream from the woods of Ikabaru, October 1947, Cardona F. 27015 (US). Wooded slopes of Quebrada O-paru-má, between Santa Teresita de Kavanayén and Río Pacairao (tributary of rio Mouak), 1065 – 1220 m elev., 20 – 21 November 1944, Steyermark J. A. 60376 (F). Eperua jenmanii has two subspecies: E. jenmanii subsp. jenmanii and E. jenmanii subsp. sandwithii.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	description	Inflorescences puberulent; bracts and bracteoles puberulent externally, glabrous within, trichomes white; pedicel puberulent, sometimes puberulent to glabrescent, trichomes white; buds puberulous, trichomes white. Flowers: hypanthium puberulent, sometimes puberulent to glabrescent; stamens sheath shorter side 9.1 – 15.0 (– 23.0) mm long, longer side 10.3 – 26.0 mm long. Phenology: — Flowering in May and from July to December; fruiting in February, March, September, and November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	distribution	Distribution: — Esequibo River basin in Guyana (mainly Mazaruni River and lower Esequibo River region) and there are two collections in Venezuela bordering Guyana in Cuyuni River and El Dorado road. It is cultivated in Trinidad and Tobago. Habitat: — Mainly floodplains (riverine) forests, sometimes in dry evergreen forest (wallaba forest), on alluvial, laterite, red sand, mixed brown and white sand soils, at sea level to 550 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	description	Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	etymology	Etymology: — The specific epithet is a tribute to G. S. Jenamn, who was the superintendent of the botanical gardens and government botanist at Georgetown in Guyana during 1879 – 1902 (Stafleu & Cowan 1979), and collected the type specimen. Vernacular names: — fine leaf walaba (Hoffman B. & Roberts L. 2502), ituri wallaba (Tutin T. G. 459), mahomillo (Steyermark J. A. 86665), water wallaba (Redden K. M. et al. 3213, Henkel T. W. & Chin M. 478).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	description	Uses: — Guyana: used for firewood and roof shingles (Tutin T. G. 459). Taxonomic notes: — The two E. jenmanii subspecies, namely jenmanii and sandwithii, can be differentiated from other species with 3 - to 5 - jugate leaves and non-falcate leaflets by their secondary venation with two intramarginal veins and vein closer to the margin continuous. The two subspecies have the largest petals in the genus. Among the species with tubular corolla, E. jenmanii distinguishes by its persistent bracts (vs. caducous in E. bijuga, E. glabriflora, E. banaensis, E. obtusata, and E. oleifera), bracteoles attached to the lower portion of the pedicels (vs. at the base of the hypanthium or to the higher portion of the pedicels in E. grandiflora and E. oleifera, or the middle one in E. purpurea), villous to glabrescent sheath (vs. glabrous in E banaensis and E. obtusata, puberulous in E. bijuga, puberulous to pubescent in E. glabriflora, and in E. grandiflora subsp. guyanensis, tomentose in E. manausensis), and glabrous ovary (vs. sericeous in E. banaensis, E. grandiflora subsp. grandiflora, and E. obtusata).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F73FF91FF6CF8F6FF27FA4B.taxon	materials_examined	Specimens examined: — GUYANA. Cuyuni-Mazaruni, Kamakusa, Mazaruni River, 0.59 miles SE of base camp, 105 m elev., 5 ° 55 ’ 0 ” N 59 ° 52 ’ 59 ” W, 30 September 2004, Redden K. M. et al. 3213 (U, US). Pakaraima Mts., SW-side of Kamarang, 450 m elev., 5 ° 52 ’ 0 ” N 60 ° 38 ’ 0 ” W, 09 November 1979, Maas P. J. M. & Westra L. Y. T. 3981 (U, US). Demerara-Mahaica, Ramsaroop Farm-resort 2 km E of Timehri Airport, vicinity of main house, 5 – 15 m elev., 6 ° 30 ’ 0 ” N 58 ° 15 ’ 0 ” W, 25 January 1992, Hoffman B. & Capellaro C. 844 (INPA, US). Essequibo Islands-West Demerara, Groete River 2 – 4 km either side of mouth of Black Creek, 8 – 16 m elev., 6 ° 37 ’ 0 ” N 58 ° 36 ’ 0 ” W, 23 August 1993, Henkel T. W. & Williams R. 2715 (U, US). Pomeroon-Supenaam, Takutu concession of Willems Timber Comp, 0 – 100 m elev., 7 ° 0 ’ 0 ” N 58 ° 49 ’ 59 ” W, 12 November 1991, Polak A. M. et al. 520 (U). GUYANA. Potaro-Siparuni, Garraway Stream, 102.5 miles on Bartica-Potaro road, 38 m elev., 5 ° 22 ’ 25.2 ” N 59 ° 7 ’ 20.6 ” W, 12 March 2004, Redden K. M. et al. 2202 (US). Upper Demerara-Berbice, Rockstone, 15 July – 1 August 1921, Gleason H. A. 837 (US). Unknown city, Essequibo River from Monkey Jump to Persaud timber concession, 5 m elev., 6 ° 19 ’ 60 ” N 58 ° 33 ’ 0 ” W, 22 May 1993, Henkel T. W. & Williams R. 2128 (US). TRINIDAD AND TOBAGO (cultivated). Trindade, s. c. 1350 (US). VENEZUELA. Bolívar, Río Venamo, between río Cuyuni (Anacoco) and Cano Apanao, 18 July 1983, Stergios B. et al. 6152 (US); along road S of El dorado between km 42 and 65, 229 m elev., 26 July 1960, Steyermark J. A. 86665 (S, US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	materials_examined	Type: — VENEZUELA. Bolivar: Raudaul Guaiquinima, 200 m elev., 16 – 19 January 1952, Maguire B. 33136 (holotype NY [00004381] image!; isotypes F [0057802] image!, GH not seen, P [00252890] image!, RB [00539604]!, US [00001147]!). Inflorescences glabrous; bracts and bracteoles glabrous, ciliate, pedicel, buds, and hypanthium glabrous; stamens sheath shorter side 4.5 – 13.0 mm long, longer side 6.1 – 18.0 mm long. Phenology: — Flowering in January, March, April, May, July, August, September, and November; fruiting in March, May, September, October, and December.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	distribution	Distribution: — Unlike the type subspecies it occurs mainly in the Orinoco River basin in Venezuela (Caura, Caroni, Paragua rivers). But it was also collected in Brazil’s upper Uraricoera River and the Kako River in Guyana. Habitat: — Floodplain forests (e. g. in Mora dominated forest), on white sand and laterite soils, at 107 – 1000 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	description	Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	etymology	Etymology: — The epithet sandwithii is a tribute to N. Y. Sandwith, a botanist who collected and revised the genus Eperua in Guyana (Cowan 1975, Sandwith 1931). Vernacular names: — concha-de-suela (Blanco C. A. 598, Liesner R. L. & González A. C. 5860), opai-yek (Maguire B. et al. 46734), tehuwadu (Maiongong name, Milliken W. 2106), tumai-yek (Steyermark J. A. 75537).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua jenmanii subsp. sandwithii differs from the type subspecies by its glabrous bracts, bracteoles, inflorescence, and hypanthium (vs. puberulent in subsp. jenmanii), and the stamens sheath tends to be smaller (vs. larger in subsp. jenmanii). Cowan (1975) described this subspecies as having epunctate leaflets, but the isotype at RB herbarium clearly shows pellucid-punctate glands in the leaflets. Furthermore, observations revealed that both subspecies have blades ranging from epunctate to pellucid-punctate.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F72FF96FF6CF9B7FEF5FC0B.taxon	materials_examined	Specimens examined: — BRAZIL. Roraima, Alto Alegre, environs of the Maiongon village close to the FUNAI PIN Uaikas on the (alto) rio Uraricoera, 300 m elev., 3 ° 33 ’ 0 ” N 60 ° 28 ’ 0 ” W [inaccurate coordinate not in upper Uraricoera River, not added to map], September 1994, Milliken W. 2106 (INPA, K, MIRR, NY); vicinity of Uaicá airstrip, rio Uraricoera, river margin, 2 ° 53 ’ 0 ” N 63 ° 36 ’ 0 ” W – 3 ° 33 ’ 0 ” N 63 ° 11 ’ 0 ” W, 07 March 1971, Prance G. T. et al. 10871 (F, INPA, K, MG, NY, R, U, US). VENEZUELA. Bolívar, Aripao, rio Caura, 5 – 20 km sur (rio arriba) del campamento las Pavas (salto Para), 220 – 240 m elev., 6 ° 15 ’ 0 ” N 64 ° 25 ’ 0 ” W, May 1982, Morillo G. & Liesner R. L. 8886 (US); Caño Pablo, tributary of Río Caura, ca 6 – 9 km east of Río Caura ca 10 km east south of Las Pavas (Salto Para), 240 m elev., 6 ° 14 ’ 0 ” N 64 ° 23 ’ 0 ” W, 10 May 1982, Liesner R. L. & Morillo G. 13972 (US). Barceloneta, Sierra Ichun: María Espuma (Salto Ichun) along the rio Ichun, tributary of rio Paragua, 500 – 625 m elev., 4 ° 46 ’ 0 ” N 63 ° 18 ’ 0 ” W, 29 December 1961, Steyermark J. A. 90416 (US). Gran Sabana, Chimatá Massif, vinicity of base camp, near Río Tirica, lower southwestern slopes of Chimantá-tepuí (Torono-tepuí), 1000 m elev., 24 May 1953, Steyermark J. A. 75537 (F).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	description	(Figures 11, 40, 41)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	materials_examined	Type (designated by Cowan 1975): — BRAZIL. Amazonas: Rio Negro, moist forest at Uanauaca, January 1852, Spruce R. 2021 (lectotype K [000555107] image!, isolectotypes P [00835936] image!, P [00835937] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	description	Tree 6.0 – 32.1 m tall. Trunk 6.0 – 50.0 cm in diameter, bark grayish to brown, smooth to striate, lenticellate. Stipules 2.5 – 4.0 × 1.2 – 3.5 mm, joined, non-foliaceous, caducous. Leaves 2 – 3 (– 4) - jugate; petioles 1.0 – 5.9 cm long; rachis (2.0 –) 4.0 – 15.5 cm long; petiolules 3.6 – 11.1 mm long; blades 6.5 – 19.0 × 3.1 – 9.0 cm, chartaceous, sometimes coriaceous, glabrous, pellucid-punctate, each areole has at least one conspicuous dot, not discolorous, slightly discolorous to discolorous, inequilateral, lower pair elliptic-falcate, middle pair elliptic-falcate, upper pair elliptic-falcate, apex acuminate, mucronate, base asymmetrical, lower pair base obtuse to rounded, rarely subcordate, middle pair base obtuse to cuneate, rarely rounded, upper pair base cuneate, sometimes obtuse, margin flat, sometimes slightly revolute, secondary venation with one intramarginal vein very close to the margin, main vein curved, prominent near the base, sometimes depressed on the adaxial surface, tertiary veins conspicuous, areoles larger, plane. Inflorescences terminal or axillary, panicle, main axis pendulous, lateral patent to erect, racemes main axis strigulose, puberulous to pubescent, lateral racemes strigulose to tomentulose, grayish-white, 46.5 – 300.0 cm long, lateral racemes 0.5 – 2.5 (– 3.1) cm long, alternate and spirally arranged; bracts 1.4 × 1.4 mm, ovate, apex gland absent, tomentose, grayish-white, caducous; bracteoles 1.6 – 5.1 × 0.8 – 1.2 mm, linear to ovate, apex gland absent, tomentulose, grayish-white, caducous, attached to the middle portion of the pedicels; pedicel (6.0 –) 8.7 – 17.0 (– 22.1) mm long, 1.1 – 1.8 mm in diameter, twisted, strigulose to tomentulose, rarely puberulous, grayish-white; buds 0.9 – 2.3 cm long, 0.6 – 1.9 cm in diameter, strigulose to tomentulose, rarely puberulous to glabrescent, grayish-white. Flowers: hypanthium 2.7 – 6.1 mm long, 3.4 – 6.4 mm in in diameter, cup-shaped, equilateral, strigulose to tomentulose, grayish-white; sepal 1.4 – 2.5 × 0.4 – 1.3 cm, oblong to elliptic, unequal, the outer ones larger, cucullate, apex gland absent, green to light-green, puberulous to strigulose, sometimes tomentulose, inner sepals scarious marginally, grayish-white; adaxial petal 1.9 – 3.3 × 2.9 – 6.3 cm, oblate, non-tubular, apex rounded, base truncate, white, rarely greenish-white, glabrous; petalodia (0.4 –) 0.8 – 2.3 × (0.2 –) 1.0 – 1.8 mm; stamens exserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 1.1 – 3.7 cm long, shorter filaments 3.4 – 5.0 cm long, sheath equilateral, 3.0 – 6.4 mm long, sheath sericeous, sometimes tomentose, free filaments glabrous, brownish-yellow, anthers 3.8 – 6.8 × 0.9 – 3.0 mm, rectangular; ovary 5.2 – 9.2 × 2.0 – 3.5 mm, narrowly obovate, sericeous, brownish-yellow, 7 ovules, stipe 2.3 – 7.0 mm long, sericeous, style 3.0 – 4.6 cm long, stigma capitate, papilate. Legumes 15.0 – 28.5 × 5.0 – 10.0 cm, stipe 0.3 – 1.0 cm long, elliptic-falcate, sometimes narrowly elliptic-falcate, apex acute, margin entire, brownish-yellow to reddish-brown, veins absent, sometimes with inconspicuous transversal veins, strigulose, sometimes puberulous, trichomes white. Seeds 2 – 7 per fruit, 2.9 – 6.3 × 1.5 – 2.6 cm, elliptic to oblong, dark-brown. Phenology: — Flowering from January to May, in July, and from September to December, with a peak in October and November; fruiting from January to April, in June, July, and from September to December.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	distribution	Distribution: — Likewise E. purpurea, E. leucantha occurs in the northwest of Amazonas state in Brazil, south of Amazonas state in Venezuela, and southwest of Colombia Amazonas, Guainía, Vaupés, and Vichada departments. Habitat: — Very frequent in the Amazonian forests known as sabatena, varillal, or campinaranas on white and brown sand mixed soils, but also reported in terra-firme (Aymard et al. 2009) and floodplains forests, from 50 to 370 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 268,709.161 km 2 and the estimated area of occupancy is 252.000 km 2. Occurrence in protected areas: — Resguardo Indígena Atabapo (Colombia, Guainía), Reserva Nacional Natural Puinawai (Colombia, Guainía), Gran Resguardo Indígena del Vaupés (Colombia, Vaupés), Reserva Natural Agua Linda (Colombia, Vichada), Reserva Biológica do Morro dos Seis Lagos (Brazil, Amazonas), Parque Nacional do Pico da Neblina (Brazil, Amazonas), and Parque Nacional Serrania La Neblina (Venezuela, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	etymology	Etymology: — The specific epithet relates to the white color of its petals. Vernacular names: — acan „ (Cavalcante P. B. 669, Rodrigues W. A. 905), dzawaka (Stropp J. & Assunção P. 200, 289, 342, 344, 345, 640), espadeira (Nascimento O. C. do et al. 29), iauácano (Rodrigues W. A. 1075), iuacan „ (Rodrigues W. A. 905), jauacaná (Spruce R. 2021), jauácano (Ducke W. A. s. n. RB 23289), kanoneyoa-ke (Guanano) (Acero E. & Rodríguez R. 948), kieraiku (Landinez A. 7), o-k ̂ - ges (Kubeo) (Schultes R. E. & Cabrera I. 17872), palocolorado, palo-rojo (Landinez A. 7), shaquiparo (Delgado H. 30), t ̂ - ee (Puinave) (Schultes R. E. & Cabrera I. 17872), tuic (Imamoto M. 15), yaauaka (Stergios B. 18584), yabakaike (Kubeo) (Acero E. & Rodríguez R. 948), yacuana (Marcano-Berti L. & Salcedo P. 40 - 979), yaguácana (Clark H. L. 6983, Stergios B. 18409), yaguácana-blanco (Clark H. L. 7019, 7020, 7023), yaguácana rojo (Clark H. L. 7021), yaguana (Williams L. 14390), yahuana (Baniba) (Williams L. 14390, 14942), yauácano (Ducke W. A. 164, s. n. RB 23732), yaua-kuan „ (Trivellato C. & Gonçalves G. G. 31), yévaro blanco (Cárdenas-López D. 21951), yévaro sabanero (Cárdenas-López D. 21936).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	discussion	Uses: — Venezuela: bridge construction (Williams L. 13939, 15801), poles (Williams L. 13939), the wood is very durable even when buried into the ground, and it is very heavy and not useful in watercrafts (Williams L. 15801). Colombia: timber (Cárdenas-López D. 21951). According to Souza L. A. G. de et al. 28, it has ornamental potential especially because of its flowers. Taxonomic notes: — Eperua leucantha is morphologically similar to E. falcata and E. venosa, with which it shares joined stipules, secondary venation with one intramarginal vein very close to the margin, long and pendulous inflorescences, non-tubular corolla, exserted stamens joined in a diadelphous sheath, and sericeous ovary (Table 4). Additionally, E. leucantha shares multijugate leaves with E. falcata, and erect lateral racemes and sericeous sheath with E. venosa. Differences between E. leucantha and E. falcata are listed in the taxonomic notes of the last species. Eperua leucantha differs from E. venosa by its shorter [2.7 – 6.1 mm long] hypanthium (vs. longer [6.8 – 11.3 mm long] in E. venosa), and white glabrous petal (vs. deep rose-colored petals basally sericeous in E. venosa).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F75FF95FF6CFC77FE96F83B.taxon	materials_examined	Selected specimens: — BRAZIL. Amazonas, S „ o Gabriel da Cachoeira, porto de Camanaús, 18 October 1978, Madison M. T. et al. 397 (INPA); S „ o Felipe, rio Negro, 11 October 1952, Fróes R. de L. 28826 (IAN); alto rio Negro, caatinga da Ilha das Flores, 17 February 1959, Rodrigues W. A. 905 (INPA, U, US). COLOMBIA. Amazonas, El Encanto, corregimiento departamental de la Chorrera, comunidad de Santa Rosa, río Igará-Parana, 151 m elev., 1 ° 45 ’ 40 ” N 72 ° 37 ’ 16 ” W, 15 March 2020, Cárdenas-López D. & Marín N. 52950 (COAH). Guainía, Cacahual, comunidad indígena Cacahual, río Atabapo sector cñ. Garza, 11 May 2007, Cárdenas-López D. 20677 (COAH). Inírida, Puerto Inírida, via aeropuerto, vitina km 4, 05 March 2010, Castro F. 8510 (COAH, UDBC). La Guadalupe, comunidad indígena El Porvenir, alto río Negro, zona limítrofe entre Colombia, Brasil y Venezuela, 14 October 2009, Cárdenas-López D. et al. 24050 (COAH). Maimachi, Serrania de Naquen, Caño Ima, 350 m elev., 2 ° 12 ’ 0 ” N 68 ° 12 ’ 0 ” W, 26 July 1992, Cortés-Ballen R. et al. 149 (COAH, COL, UDBC). Pana Pana, río Guaviare, Barranco Murciélago, 100 – 600 m elev., 25 November 1976, Delgado H. 30 (UDBC). Vaupés, Mitú, rio Vaupés, Miriti, 01 November 1993, Martínez X. et al. 3476 - 2 (COL). Yavaraté, Río Yutica, 0 ° 49 ’ 57 ” N 69 ° 37 ’ 9 ” W, 12 February 1979, Acero E. & Rodríguez R. 948 (COL, UDBC). Vichada, Reserva Natural Agua Linda, afloramiento rocoso adyacente a la Reserva Morrocoy y Bosque de galería del Caño Morrocoy, 07 April 2008, Castro F. 5010 (UDBC). VENEZUELA. Amazonas, Casiaquire, ríos Pacimoni - Yatua, Río Yatua near Cano Tauavaca, 100 – 140 m elev., 04 December 1953, Maguire B. et al. 36508 (F). Maroa, rio Guainía, 127 m elev., 17 February 1942, Williams L. 14390 (F, IAN, US). San Carlos de Río Negro, 20 km s of conf. of rio Negro and brazo Casiquiare, 4.3 km NNE of San Carlos on Solano road, 119 m elev., 1 ° 56 ’ 0 ” N 67 ° 3 ’ 0 ” W, 21 February 1979, Clark H. L. 7019 (US); Neblina Base camp, rio Mawarinuma, 140 m elev., 0 ° 50 ’ 0 ” N 66 ° 10 ’ 0 ” W, 27 November 1984, Boom B. M. & Weitzman A. L. 5186 (INPA, U, US). Yavita, along road from Maroa to Yavita, ca. 7 km from Maroa, 2 ° 54 ’ 42 ” N 67 ° 26 ’ 55 ” W, 25 February 1998, Acevedo-Rodríguez P. et al. 10408 (US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	description	(Figure 9, Fortes et al. 2023 Figures 6, 7)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	materials_examined	Type: — BRAZIL. Amazonas, Manaus, Reserva Florestal Adolpho Ducke, trilha do Jardim Botânico MUSA, planta marcada JBN 634, 08 June 2017, Cabral F. N. et al. 1561 (holotype INPA [0284318]!, isotype RB [01459755]!). Tree 20.0 m tall. Trunk 38.0 cm in diameter, bark brownish, lenticellate. Stipules 1.8 – 1.9 × 0.8 – 1.4 mm, free, nonfoliaceous, ovate to lanceolate, caducous. Leaves 2 – 3 - jugate, glabrous; petioles 0.6 – 2.9 cm long; rachis 3.5 – 14.6 cm long; petiolules 6.2 – 9.1 mm long; blades 5.3 – 12.8 × 3.5 – 6.5 cm, coriaceous, glabrous, inconspicuous pellucid-punctate to pellucid-punctate, discolorous, equilateral, lower pair ovate, middle pair elliptic, upper pair elliptic, apex attenuate to acuminate, base symmetrical, lower pair base subcordate to cordate, middle pair base subcordate to rounded, upper pair base subcordate, rounded to obtuse, margin revolute to flat at the apex, secondary venation with two intramarginal veins, vein closer to the margin continuous, main vein straight, prominent only near the base on the adaxial surface, tertiary veins conspicuous. Inflorescences axillary, panicle, erect, tomentose, brownish-yellow, 4.6 – 8.9 cm long, lateral racemes 2.1 – 3.4 cm long, alternate and distichous; bracts not seen, caducous; bracteoles 4.9 – 5.0 × 3.0 – 3.2 mm, ovate, apex gland present, tomentose externally, pubescent to tomentose within, brownish-yellow, caducous, attached to the lower middle portion of the pedicel; pedicel 17.5 – 18.9 mm long, 1.0 mm in diameter, not twisted, tomentose, brownish-yellow; buds 0.7 cm long, 0.4 cm in diameter, tomentose, brownish-yellow. Flowers: hypanthium 3.8 mm long, 2.8 mm in diameter, cup-shaped, equilateral, tomentose, brownish-yellow; sepal 1.6 × 0.5 cm, oblong to elliptic, unequal, the dorsal one larger, cucullate, apex gland absent, pink, tomentose, scarious marginally, brownish-yellow; adaxial petal 3.8 × 4.7 cm, flabellate, tubular, apex rounded, base attenuate, pink to purple, glabrous; petalodia not seen; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 3.1 cm long, shorter filaments 2.0 cm long, sheath inequilateral, shorter side 6.5 mm long, longer side 8.0 mm long, sheath tomentose, brownish-yellow, free filaments glabrous, anther 2.2 × 0.8 mm, rectangular; ovary 7.1 × 3.7 mm, obovate, densely tomentose to villous, brownish-yellow, stipe tomentose, style 4.0? cm long, glabrous, stigma capitate. Legumes 11.3 – 15.9 × 7.7 – 8.3 cm, stipe 0.9 – 1.4 mm long, obovate to irregularly elliptic, apex obtuse, apiculate, dorsal margin slightly thickened, brownish-red, with inconspicuous transversal veins, puberulous to puberulent, brownish. Seeds 1 – 2 per fruit. Phenology: — Flowering in June; fruiting in January and June.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	distribution	Distribution: — Brazil (Amazonas) Habitat: — Upland (terra-firme) forest at 114 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	description	Conservation status: — Categorized as Data Deficient (DD) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022) due to its recently circumscription and being known by only one specimen. Occurrence in protected areas: — Reserva Florestal Adolpho Ducke (Brazil, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	etymology	Etymology: — The specific epithet is related to its first collection in the Museu da Amazônia, a botanical garden located in the city of Manaus (Brazil, Amazonas) and associated with the Adolpho Ducke Forest Reserve (Fortes et al. 2023).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	vernacular_names	Vernacular names: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF9AFF6CFF33FEC6F897.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua manausensis differs from other species with short and erect inflorescence and tubular corolla by the following combination of characteristics: free stipules, two intramarginal veins, vein closer to the margin continuous, tomentose inflorescence, caducous bracteoles attached to the lower middle portion of the pedicels, tomentose sheath, tomentose to villous ovary. Additionally, it differs from other species with tubular corolla by its 2 – 3 - jugate leaves, and cordate to subcordate leaflet base. Specimens examined (Paratypes): — BRAZIL. Amazonas, Manaus, Reserva Florestal Adolpho Ducke, MUSA, planta marcada JBN 634 ao lado direito da Casa-Oficina, à 20 m da trilha Branca, 114 m elev., 3 ° 0 ’ 22.9 ” S 59 ° 56 ’ 22.7 ” W, 27 January 2022, Fortes E. A. & Viana G. P. 194 b (HUEFS, IAN, INPA, MG, NY, RB [01458706]!, UEC, US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	description	(Figure 13, 16 b, c, e, 42)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	materials_examined	Type: — VENEZUELA. Amazonas: Capihuara, Rio Casiquiare, 3 October 1943, Williams L. 16183 (lolotype US [00001148]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	description	Tree 10.0 – 20.0 m tall. Trunk, bark smooth, lenticellate. Stipules 7.3 × 2.3 mm, free, non-foliaceous, caducous. Leaves 2 - jugate; petioles 2.6 – 4.5 cm long; rachis 3.4 – 5.3 cm long; petiolules 4.5 – 7.5 mm long; blades 8.0 – 13.0 × 4.0 – 5.9 cm, coriaceous, glabrous, pellucid-punctate, discolorous, equilateral, lower pair elliptic to oblong, upper pair elliptic, apex rounded to obtuse, base asymmetrical, lower pair base obtuse to cuneate, upper pair base cuneate, margin flat to slightly revolute, secondary venation with one intramarginal vein, main vein straight, prominent to depressed on the adaxial surface, tertiary veins conspicuous, areoles larger, plane. Inflorescences axillary or ramiflorous, raceme, erect, strigulose, brownish- yellow, 7.5 – 9.5 cm long; bracts not seen, caducous; bracteoles ca. 2.0 × 1.0 – 2.0 mm, ovate, strigulose to tomentose externally, brownish-yellow, caducous, attached to the lower portion of the pedicel; pedicel 8.0 – 10.0 mm long, 0.8 – 1.1 mm in diameter, not twisted, strigulose, brownish-yellow; buds 0.5 cm long, 0.2 cm in diameter, strigulose, brownish-yellow. Flowers: hypanthium 2.0 – 3.0 mm long, 3.3 – 4.6 mm in diameter, cup-shaped, equilateral, strigulose, brownish-yellow; sepal 1.6 – 2.0 × 0.4 cm, oblong to elliptic, unequal, the outer ones larger, apex gland present, strigulose, inner ones scarious marginally, brownish-yellow; adaxial petal 3.5 – 5.0 × 6.5 cm, flabellate, tubular, apex rounded, base attenuate, glabrous; petalodia 4.0 – 5.0 × 1.5 – 1.8 mm; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, filaments 11.5 – 26.0 cm long, sheath inequilateral, 6.5 – 10.0 mm long, glabrous, anther 2.6 – 3.5 × 1.2 – 1.5 mm, rectangular; ovary 5.0 × 2.0 mm, obovate, sericeous, brownish-yellow, stipe 4.0 mm long, sericeous, style 3.4 cm long, stigma truncate. Legumes not seen. Phenology: — Flowering in October.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	distribution	Distribution — Casiquiare River region in Venezuela (Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	biology_ecology	Habitat: — Tall campinarana and terra-firme forest, on sandy soil, from 106 to 306 m elev. Coomes & Gribb (1996) cited E. obtusata as one of the dominant species in the tall campinarana of La Esmeralda (Venezuela).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	conservation	Conservation status: — Categorized as Data Deficient (DD) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022) due to the lack of access to its collections. Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	etymology	Etymology: — Eperua obtusata and E. banaensis are the only species with leaflets with an obtuse apex, and the specific epithet obtusata relates to that characteristic.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	vernacular_names	Vernacular names: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F79FF98FF6CF8F3FDA6FC53.taxon	discussion	Uses: — Unknown. Taxonomic notes: — Eperua banaensis and E. obtusata are the only species in the genus with rounded, obtuse, acute to emarginate leaflet apex in the genus, whereas all other species have an attenuate, acuminate to caudate apex. They share 2 - jugate leaves, short and erect inflorescence, tubular corolla, inserted stamens, and the same pattern of reproductive organ indumentum. Among the species with a tubular corolla, E. banaensis and E. obstusata are distinguished by the combination of glabrous sheaths and indumented ovaries (Table 2). Differences between them are listed in the taxonomic notes of E. banaensis. Specimens examined: — VENEZUELA. Amazonas, Casiaquire, Capihuara, 03 October 1943, Williams L. 16183 (US). San Fernando de Atabapo, La Esmeralda, 5 Km North of village, near savanna Morichal, 106 m elev., 3 ° 10 ’ 0 ’’ S 65 ° 33 ’ 0 ’’ W, undated, Coomes D. 207 (K).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7BFF99FF6CFB8FFB50FD16.taxon	materials_examined	Type (designated by Cowan 1975): — BRAZIL. Amazonas: Borba, Rio Madeira, 16 January 1930, Ducke W. A. RB 23290 (lectotype R [000028782]!, isolectotypes F-frag. [0057804] image!, G not seen, INPA collection number 234506 image!, K [000555093] image!, K [000858688] image!, NY [00004382] image!, NY [00004383] image!, NY [01185435] image!, P [00252888] image!, RB sheet I [00585686]!, & sheet II [00539606]!, S collection number S-R- 8709 image!, U [0003233] image!, US [00001149]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7BFF99FF6CFB8FFB50FD16.taxon	description	Tree 18.0 – 30.0 m tall, or treelet, or shrub 1.5 – 9.0 m tall. Trunk 3.0 – 60.0 cm in diameter, bark grayish, brown with grey spots, striate. Stipules 1.5 – 4.3 × 1.3 – 4.0 mm, joined, non-foliaceous, persistent. Leaves (2 –) 3 – 4 (– 5) - jugate; petioles 0.8 – 4.2 cm long; rachis 1.6 – 16.6 cm long; petiolules 2.2 – 8.1 mm long; blades 4.6 – 13.7 × 2.6 – 7.7 (– 10.6) cm, coriaceous to chartaceous, glabrous, very nitid adaxially, conspicuous pellucid-punctate, discolorous, sometimes not discolorous, equilateral, lower pair elliptic, sometimes broadly elliptic to ovate, middle pair elliptic, sometimes broadly elliptic to ovate, upper pair elliptic, sometimes broadly elliptic to ovate, apex acuminate, sometimes retuse, base symmetrical, sometimes asymmetrical, lower pair base rounded, subcordate to cordate, or obtuse to rounded, rarely cordate, middle pair base rounded, subcordate to cordate, or obtuse, sometimes rounded, upper pair base obtuse, rounded, subcordate to cordate, or obtuse to cuneate, sometimes rounded, margin flat, secondary venation with one intramarginal vein very close to the margin, main vein straight, depressed on the adaxial surface, tertiary veins very conspicuous, prominent on the adaxial surface, rarely not prominent, or conspicuous to inconspicuous, not prominent on the adaxial surface, rarely prominent, areoles conspicuous, larger, plane. Inflorescences terminal or ramiflorous, sometimes axillary, panicle, sometimes raceme, erect, strigose to tomentose, rarely pubescent, brownish-yellow, 3.0 – 12.3 cm long, lateral racemes 0.9 – 6.2 cm long, alternate and distichous; bracts 1.0 – 2.8 × 0.5 – 1.8 mm, ovate, rarely broadly ovate, apex gland absent, tomentose to tomentulose externally, glabrous within, brownish-yellow, caducous to persistent; bracteoles 1.0 – 2.8 × 0.8 – 1.9 mm, ovate, apex gland absent, tomentose externally, glabrous within, brownish-yellow, persistent, attached to the higher portion of the pedicels or at the base of the hypanthium; pedicel 3.7 – 14.1 mm long, 0.4 – 1.4 mm in diameter, not twisted, tomentulose to tomentose, sometimes tomentulose, rarely pubescent, brownish-yellow; buds 0.5 – 1.0 cm long, 0.3 – 0.7 cm in diameter, tomentose, brownish-yellow. Flowers: hypanthium 0.9 – 3.4 mm long, 1.2 – 4.0 mm in in diameter, cup-shaped, equilateral, tomentulose to tomentose, sometimes strigose, brownish-yellow; sepal 0.4 – 1.0 × 0.2 – 0.7 cm, elliptic to ovate, rarely oblong, unequal, the dorsal one larger, cucullate, apex gland absent, pale-green, yelowish-green to reddish-pink, strigulose, tomentulose to tomentose externally, scarious marginally, brownish-yellow; adaxial petal 1.7 – 3.8 × 2.0 – 5.7 cm, flabellate, tubular, apex rounded, crisped, base attenuate, gibbous, lilac, bluish-lavender, purple, pink-purple to pink, glabrous; petalodia 0.1 – 1.3 × 0.1 – 0.6 mm; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 0.7 – 1.8 cm long, shorter filaments 1.5 – 3.7 cm long, sheath inequilateral to slightly inequilateral, shorter side 2.2 – 6.7 mm long, longer side 1.7 – 8.5 mm long, sheath villous, sometimes pubescent basally, free filaments glabrous, brownish-yellow, shorter stamens anthers 0.3 – 1.2 × 0.3 – 1.0 mm, square to ovate, apiculate, longer stamens anthers 0.9 – 2.0 × 0.2 – 1.2 mm, rectangular, apiculate, rarely ovate with acute apex; ovary 1.4 – 7.6 × 0.8 – 3.5 mm, obovate, glabrous, 3 – 8 ovules, stipe 1.4 – 4.2 mm long, glabrous, style 1.3 – 2.5 cm long, stigma capitate to obtuse, villous to papillose. Legumes 6.5 – 11.8 × 3.3 – 6.0 cm, stipe 0.5 – 1.8 cm long, oblanceolate to obovate, apex obtuse to truncate, apiculate, margin entire, cinnamon, brown to purplish-brown, veins absent, glabrous, pitted. Seeds 1 – 2 per fruit, 2.9 × 1.4 cm, elliptic, dark-brown, nitid, pitted reticulate.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7BFF99FF6CFB8FFB50FD16.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 155,436.182 km 2 and the estimated area of occupancy is 112.000 km 2. Eperua oleifera has to varieties: E. oleifera var. oleifera and E. oleifera var. campestris.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	description	(Figures 14, 44, 45) Tree 18.0 – 30.0 m tall. Blades: lower pair base obtuse to rounded, rarely cordate, middle pair base obtuse, sometimes obtuse to cuneate, sometimes rounded, tertiary veins conspicuous to inconspicuous, not prominent in the adaxial surface, rarely prominent. Phenology: — Flowering in January, from May to August, and in November; fruiting in July and August.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	distribution	Distribution: — Southeast of Amazonas state in the Madeira River basin (Brazil). Habitat: — Mainly in upland (terra-firme) forests on sandy soils, but also in flooded forests at ca. 100 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	description	Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	etymology	Etymology: — The specific epithet relates to the oil resin that is extracted from its trunk (Ducke 1932). Vernacular names: — jacaré-copahiba (Ducke W. A. 213, 235, s. n. RB 23290, s. n. RB 20217), copaíba-jacaré (Coêlho L. F. s. n. INPA 39594), copaíba preta (Coêlho L. F. s. n. INPA 42008).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua oleifera can be differentiated from other species of Eperua by the 3 - jugate leaves with straight and adaxially nitid and conspicuous pellucid-punctate leaflets. Among the species with tubular corollas, E. oleifera differs by the combination tomentose to strigose inflorescences, strigulose, tomentulose to tomentose sepals, bracteoles attached to the higher portion of the pedicels or at the base of the hypanthium, villous sheath, and glabrous ovary (see Table 2).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF99FF6CFD73FAC4F993.taxon	materials_examined	Selected specimens: — BRAZIL. Amazonas, Borba, Axinim, rio madeira, 07 May 1936 – 05 July 1936, Ducke W. A. 213 (F, K, MO, NY, R, S, US). Manicoré, distrito de Santo Antônio do Matupi, área de manejo do Sr. Amadeu S. Vidal, faz. M „ o de Bronze, rodovia transamazônica 234, 106.7 m elev., 7 ° 54 ’ 31 ” S 61 ° 36 ’ 28 ” W, 25 August 2014, Amoêdo A. B. et al. 49 (EAFM). Maués, along rio Apoquitaua, just above mouth of rio Pacoval, 3 ° 47 ’ 0 ” S 57 ° 56 ’ 0 ” W, 27 July 1983, Zarucchi J. L. et al. 3209 (F, INPA, HFSL, MG, MO, NY, RB, UFACPZ, US). Rio de Janeiro (cultivated), Rio de Janeiro, Jardim Botânico do Rio de Janeiro, 01 July 1989, Gonçalves V. F. s. n. RB 422083 (RB).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	description	(Figures 14, 43, 45)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	materials_examined	Type: — BRAZIL: Amazonas: Borba, Rio Madeira, campo arenoso, 17 January 1930, Ducke W. A. s. n. RB 23291 (holotype RB sheet I [00539605]!, sheet II [00585835]!, & sheet III [00585836]!; isotypes NY [00004384] image!, P [00835938] image!, R [000028783]!, U [0003232] image!, US [00001150]!). Treelet, or shrub 1.5 – 9.0 m tall. Blades: lower pair base rounded, subcordate to cordate, middle pair base rounded, subcordate to cordate, upper pair base obtuse, rounded, subcordate to cordate, tertiary veins very conspicuous, prominent in the adaxial surface, rarely not prominent.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	description	Phenology: — Flowering mainly in April, from June to September, and in November, with a flowering peak in August; fruiting from May to August, in October, and November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	distribution	Distribution: — Southeast of Amazonas state in the Madeira River basin (Brazil). Habitat: — White sand campina and campinarana, in upland and fooldplains areas, at ca. 100 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	description	Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	etymology	Etymology: — The specific epithet relates to its habitat in the campinas. Vernacular names: — copaiba jacaré (Pessoal do Proj. Radam 13234).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua oleifera var. campestris differs from the type variety by being a treelet or shrub (vs. tall tree in var. oleifera) with the lower and middle pair of leaflets having a rounded, cordate to subcordate base (vs. rounded to obtuse, rarely cordate or subcordate, in var. oleifera), and very conspicuous tertiary veins (vs. conspicuous to inconspicuous in var. oleifera). Eperua oleifera var. campestris presents strigose to tomentose inflorescence (vs. tomentose, indumentum tending to be denser, in var. oleifera), and it is the only reproductive difference, justifying its permanency as a variety.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7AFF9DFF6CF9CFFED0FD5F.taxon	materials_examined	Selected specimens: — BRAZIL. Amazonas, Apuí, rodovia Transamazônica, igarapé Coca-cola, 7 ° 26 ’ 5 ” S 61 ° 2 ’ 23 ” W, 18 August 2009, Assunção P. A. C. L. & Carvalho F. A. de 1118 (INPA). Borba, rio Madeira, 09 November 1935, Ducke W. A. 13 (K). Manaus, rio Negro basin September 1948, Schultes R. E. & López F. 10337 (IAN, NY, US); 5 Km S of Borba, 4 ° 23 ’ 0 ” S 59 ° 35 ’ 0 ” W, 26 June 1983, Zarucchi J. L. et al. 2842 (HFSL, INPA, K, MG, MO, NY, RB, UFACPZ, US); Axinim, basin of rio Abacaxis, ca. 4 km from mouth of igarapé Surubim, 4 ° 31 ’ 0 ” S 58 ° 34 ’ 0 ” W, 04 July 1983, Zarucchi J. L. et al. 2941 (INPA, K, MG, MO, NY, RB, UFACPZ, US). Manicoré, BR 230, rod. Transamazônica, a 243 Km de Humaitá, 7 ° 40 ’ 0.1 ” S 61 ° 10 ’ 0 ” W, 24 April 1985, Cid-Ferreira C. A. 5791 (F, HRCB, INPA, K, MG, MO, RB, US). Maués, campina Socoró, 4 ° 17 ’ 2 ” S 58 ° 34 ’ 40 ” W, 07 August 2015, Demarchi L. O. et al. 529 (INPA). Novo Aripuan „, BR 230, rod. Transamazônica entre os Kms 350 e 400 ao leste de Humaitá, 30 April 1985, Cid-Ferreira C. A. 5913 (F, HRCB, INPA, K, MG, MO, NY, RB, US); Nova Prainha, 23 July 1976, Mota C. D. A. da & Monteiro O. P. s. n. INPA 60692 (INPA). Nova Olinda do Nortes, rio Abacaxis, Igarapé do Socoró, campina Socoró, 4 ° 17 ’ 2 ” S 58 ° 34 ’ 40 ” W, 07 August 2015, Holanda A. S. S. et al. 799 (INPA). Rio de Janeiro (cultivated), Rio de Janeiro, Arboreto do Jardim Botânico do Rio de Janeiro, nº 317, canteiro A, séc. XXV, 25 April 1985, Carvalho I. R. & Ricardo C. C. s. n. RB 243379 (INPA, RB).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	description	(Figures 14, 46, 47)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	materials_examined	Type (designated by Cowan 1975): — BRAZIL. Amazonas: Panuré, Rio Negro, Uaupés, September 1852, Spruce R. 2577 (lectotype K [000555096] image!, isolectotype P-flower [00835940] image!). Tree (2.0 –) 15.0 – 30.0 (– 50.0) m tall. Trunk 30.0 – 100.0 cm in diameter, bark grayish, smooth, lenticelate, to fissured. Stipules 1.9 – 4.2 × 1.5 – 4.1 mm, joined, non-foliaceous, persistent. Leaves 2 – 3 - jugate; petioles 0.7 – 3.8 cm long; rachis 0.7 – 10.7 cm long; petiolules 2.8 – 8.4 mm long; blades 4.0 – 14.3 × 2.2 – 6.7 (– 10.5) cm, coriaceous, upper surface glabrous, nitid, lower surface densely papillate-ceriferous except on the marginal vein, pellucid-punctate, strongly discolorous, equilateral, lower pair elliptic, sometimes ovate, rarely broadly elliptic, middle pair elliptic, sometimes ovate, rarely broadly elliptic, upper pair elliptic, rarely broadly elliptic or ovate, apex acuminate, retuse, base symmetrical, lower pair base obtuse to rounded, middle pair base obtuse to rounded, sometimes subcordate, rarely cordate, upper pair base obtuse, sometimes rounded, margin flat to slightly revolute, secondary venation with one intramarginal vein very close to the margin, main vein straight, depressed on the adaxial surface, tertiary veins inconspicuous to conspicuous only adaxially, areoles inconspicuous. Inflorescences terminal, rarely axillary, panicle or raceme, erect, glabrous, 2.2 – 9.0 cm long, lateral racemes 1.2 – 9.6 cm long, congested racemes, alternate and distichous; bracts 0.9 – 3.2 × 0.4 – 3.2 mm, ovate, apex gland absent, glabrous, pellucid punctate externally, sericeous, pubescent to glabrescent within, brownish-yellow, persistent, rarely caducous; bracteoles 1.4 – 3.3 (– 8.9) × 0.4 – 3.2 mm, ovate, apex gland absent, glabrous externally, pubescent to sericeous within, ciliate, brownish-yellow, persistent, attached to the middle portion of the pedicels at slightly different levels; pedicel 7.4 – 20.0 mm long, 0.5 – 1.2 mm in diameter, not twisted, glabrous; buds 0.7 – 1.4 cm long, 0.3 – 1.0 cm in diameter, glabrous. Flowers: hypanthium 1.6 – 4.1 mm long, 1.6 – 4.8 mm in diameter, cup-shaped, equilateral, glabrous; sepal 0.7 – 1.5 × 0.3 – 0.8 cm, elliptic to ovate, rarely obovate, unequal, the outer ones larger, cucullate, apex gland absent, purple to red, glabrous, pellucid-punctate; adaxial petal 1.9 – 4.4 × 2.9 – 5.6 cm, flabellate, tubular, apex rounded, base attenuate, gibbous, pink purple to purple, glabrous; petalodia 0.2 – 1.5 × 0.1 – 0.7 mm; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 1.5 – 3.2 cm long, shorter filaments 0.8 – 1.6 cm long, sheath inequilateral, shorter side 3.2 – 7.7 mm long, longer side 4.2 – 8.9 mm long, sheath and one-third of the free filaments villous, brownish-yellow, shorter stamens anthers 0.4 – 1.0 × 0.4 – 1.0 mm, square, apiculate, longer stamens anthers 1.2 – 1.8 × 0.4 – 1.0 mm, rectangular; ovary 2.4 – 5.0 × 0.9 – 2.5 mm, obovate, glabrous, 3 ovules, stipe 1.6 – 3.6 mm long, glabrous, style 1.2 – 3.0 cm long, stigma capitate. Legumes 11.0 – 17.0 × 3.9 – 7.9 cm, stipe 0.5 – 1.0 cm long, obovate, apex obtuse to truncate, apiculate, dorsal margin alate, fuscous, veins absent or with inconspicuous veins, glabrous. Seeds 2 per fruit, 3.8 – 4.4 × 1.9 – 3.0 cm, elliptic to obovate, dark-brown to reddish-brown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	description	Phenology: — Flowering from January to June and from August to November, with a pick in October and November; fruiting from January to May and from September to November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	distribution	Distribution: — Northwest of Amazonas state in Brazil, south of Amazonas state in Venezuela, and southwest of Colombia Amazonas, Guainía, and Vaupés departments.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	biology_ecology	Habitat: — Frequent in the Amazonian forests known as campinarana or varillal on white and brown sand mixed soils, but also reported in terra-firme (Aymard et al. 2009), and gallery forests, from 79 to 320 (– 750) m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 195,184.753 km 2 and the estimated area of occupancy is 216.000 km 2. Occurrence in protected areas: — Resguardo Indígena Río Atabapo (Colombia, Guainía), Parque Nacional Natural Río Puré (Colombia, Amazonas), Parque Nacional do Pico da Neblina (Brazil, Amazonas), and Parque Nacional Serranía La Neblina (Venezuela, Amazonas).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	etymology	Etymology: — The specific epithet relates to its petal color. Vernacular names: — backaike (Roa T. A. 945), boo-ro-go (Desano), boo-toó-ke (Gwanano) (Schultes R. E. & Cabrera I. 19324), gebaro (Roosmalen M. G. M. van et al. 1305), gibarú (Lima H. C. de et al. 3221), guapa (Córdoba M. P. et al. 93, Etter A. 3, Stergios B. 18387), iacana (Nascimento O. C. do et al. 101), iebaro (Fróes R. de L. 21206, Maguire B. et al. 60112, Pires J. M. 468), jébaru (Spruce R. 2577), poé-koó (puinave), vá-pa (Kuripako) (Schultes R. E. et al. 17955), waapa (Stropp J. & Assunção P. 234), wapa-panari (Cárdenas-López D. 21898), yé-ba (Schultes R. E. et al. 17955), yebaro (Cárdenas-López D. et al. 15460, Liesner R. L. 4049, Schultes R. E. et al. 18268), yébaro (Ducke W. A. s. n. RB 23288), yebaroh (Liesner R. L. 3742), yeberro (Marcano-Berti L. & Salcedo P. 65 - 979), yeiaro (Stergios B. et al. 9814), yevaro (Maguire B. et al. 41776), yévaro (Berry P. E. et al. 5737, Clark H. L. 7011, Huber O. 3376, Stergios B. & Matínez O. 9491, Stergios B. et al. 9967, Williams L. 13990), yévaro-morado (Cárdenas-López D. et al. 21916, 21937, 21946), yéparo (Baniba) (Huber O. 3376).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	description	Uses: — Colombia: the bark is used for Yuruparí horns (Schultes R. E. et al. 17955), ashes from the bark is used to make pans and pots (Roa A. T. 376); the sticky resin (colorless) in wood is used to catch birds (Schultes R. E. et al. 17955); ornamental use (Acero E. & Rodríguez R. 945), timber (Cárdenas-López D. 21898). Venezuela: very durable wood used for general construction and bridges (Williams L. 13990). Taxonomic notes: — Eperua purpurea is the only species within Eperua with strongly discolorous and abaxially densely papillate-ceriferous leaflets, which, combined with the 3 - jugate leaves straight leaflets, make it one of the most easily recognizable species. Among the species with tubular corolla, E. purpurea differs by the glabrous inflorescence, hypanthium and sepals, persistent bracteoles attached to the middle portion of the pedicels, and villous sheath (see Table 2).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F7EFF80FF6CFC8BFED7F8AB.taxon	materials_examined	Selected specimens: — BRAZIL. Amazonas, Santa Isabel do Rio Negro, Tapuruquara, basin of rio Negro, 27 October 1971, Prance G. T. et al. 15609 (INPA, K, MG, NY, P, R, U, US). S „ o Gabriel da Cachoeira, rio Içana, próximo a Serra do Grilo, 05 November 1987, Farney C. de S. et al. 1899 (HAMAB, INPA, K, NY, US); rio Marié, Tapuruquara Mirim, 16 October 1978, Madison M. T. et al. 319 (INPA, RB); Uaupés, R. Negro, estrada Perimetral Norte, 13 March 1975, Pires J. M. & Marinho L. R. 15783 (IAN). Rio de Janeiro (cultivated), Jardim Botânico do Rio de Janeiro, 1246 - XXVI-E, undated, s. c. s. n. RB 89490 (RB). COLOMBIA. Amazonas, Leticia, PNN Amacayacu, 12 August 1995, Etter A. 3 (COAH). Guainía, Cacahual, comunidad indígena Cacahual, zona de rebalse del río Atabapo, 18 August 2004, Cárdenas-López D. et al. 15350 (COAH). Inírida, río Atabapo, Caño Chaquita, localidad La Esperanza, 23 May 1979, Rodríguez R. & Acero E. 207 (COL, UDBC). La Guadalupe, corremiento de San Felipe, comunidad El Porvenir, alto rio Negro, em cercanias de la Piedra del Cocui (zona limiftrofe entre Colombia, Brasil y Venezuela), 200 m elev., 1 ° 19 ’ 12 ” N 66 ° 53 ’ 4 ” W, 15 October 2009, Cárdenas-López D. et al. 24076 (COAH, NY). Maimachi, Serrania de Naquen, alrededores del campamento La Planada, 320 m elev., 2 ° 12 ’ 0 ” N 68 ° 11 ’ 59 ” W, 14 August 1992, Cortés-Ballen R. et al. 336 (COAH, COL, UDBC). Pana Pana, bosque ubicado a 15 min, caminando desde la rivera del río Isana, a 15 min en lancha desde la comunidad de Punta Tigre, 09 May 2014, Aymard G. 14571 (COAH). Puerto Colombia, corregimiento de Cano Colorado, medio rio Guainia, Cano Mahimachi, Serrania de Naquen, 300 m elev., 2 ° 9 ’ 13.4 ” N 68 ° 14 ’ 7 ” W, 20 October 2009, Cárdenas-López D. et al. 24415 (COAH, US). San Filipe, rio Negro, below confluence of rio Guainia and rio Casiquiare, 192 m elev., 24 October 1952, Schultes R. E. et al. 17955 (COL, U, US). Vaupés, Mitú, confluencia de los ríos Vaupés y Papurí, Yaravaté, bosque entre el río Vaupés y el caño Ardilla, 120 m elev., 0 ° 37 ’ 0 ” N 69 ° 12 ’ 0 ” W, 30 November 2004, Bernal R. et al. 3683 (COL). VENEZUELA. Amazonas, Casiaquire, pueblo de Maroa, 90 m elev., 2 ° 43 ’ 0 ” N 67 ° 32 ’ 0 ” W, 24 February 1979, Huber O. 3376 (US). Maroa, rio Guainia, along trail to Guzman Blanco 1 – 5 km, Southeast of Moroa, 120 – 140 m elev., 09 October 1957, Maguire B. et al. 41776 (IAN, RB, U, US). San Carlos de Río Negro, Neblina Base camp, rio Negro, Neblina Base camp, rio Mawarinuma, 140 m elev., 0 ° 50 ’ 0 ” N 66 ° 10 ’ 0 ” W, 28 November 1984, Boom B. M. & Weitzman A. L. 5185 (F, INPA, MO, U, US); 18 - rio Yatuá, 84 m elev., 1 ° 16 ’ 33.3 ” N 66 ° 6 ’ 51.1 ” W, 05 February 2005, Redden K. M. et al. 3560 (US). Yavita, along road from Maroa to Yabita, ca. 700 m from Yabita, 2 ° 54 ’ 42 ” N 67 ° 26 ’ 55 ” W, 19 February 1998, Acevedo-Rodríguez P. et al. 10235 (US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	description	(Figures 8, 48, 49, 50)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	materials_examined	Type: — GUYANA. Cuyuni-Mazaruni, Mazaruni River, small islands along river, 90 m elev., 6 ° 0 ’ 12.6 ” N 59 ° 13 ’ 28.9 ” W, 11 October 2004, Redden K. M. et al. 3274 (holotype US [00808373]!, isotypes CAY [109816, 109817] image!, INPA [0220628]!)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	diagnosis	Diagnosis: — Eperua reddeniae has short and erect inflorescences, non-tubular corollas, and exserted stamens. It differs from species with the same type of inflorescence, corolla, and stamen by the combination (see Table 3): stipules joined at the base, inconspicuous pellucid punctate to epunctate leaflets, slightly revolute margins, coppery inflorescence indumentum, pink to purple petals, stamens joined basally in a diadelphous sheath, dorsal one free, sheath irregularly tomentose in the middle-high portion and glabrous at the base, tomentose ovary, tomentulose fruits with entire margins.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	description	Tree 8 – 20.0 m tall. Trunk 20.0 cm in diameter, bark not seen. Stipules 10.9 – 19.2 × 6.3 – 12.2 mm, joined at the base, foliaceous, persistent. Leaves 4 – 5 - jugate; petioles 1.0 – 1.5 cm long; rachis 10.2 – 15.0 cm long; petiolules 6.2 – 8.2 mm long; blades 5.3 – 11.8 × 2.8 – 5.0 cm, coriaceous, glabrous, pellucid-punctate to epunctate, discolorous, equilateral, lower pair elliptic to ovate, middle pair elliptic, upper pair elliptic, apex attenuate to acuminate, mucronate, base symmetrical to asymmetrical, lower pair base subcordate to obtuse, middle pair base obtuse, upper pair base obtuse, margin flat to slightly revolute, secondary venation with two intramarginal veins, sometimes one, vein closer to the margin not continuous, main vein straight, prominent on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal or axillary, panicle, erect, tomentose, coppery, 13.1 – 16.0 cm long, lateral racemes 2.1 – 11.8 cm long, alternate and distichous; bracts caducous, not seen; bracteoles 2.1 – 3.8 × 2.3 – 2.4 mm, ovate, cucullate, apex gland absent, tomentose on both sides, coppery, caducous, attached to the lower middle portion of the pedicels; pedicel 18.4 – 26.2 mm long, 1.3 – 2.1 mm in diameter, not twisted, tomentose, coppery; buds 1.1 – 1.5 cm long, 0.6 – 0.7 cm in diameter, tomentose, coppery. Flowers: hypanthium 4.3 – 6.3 mm long, 4.9 – 6.1 mm in diameter, cup-shaped, equilateral, tomentose, coppery; sepal 1.9 – 2.7 × 0.6 – 1.2 cm, elliptic to oblong, unequal, the outer ones larger, cucullate, apex gland absent, green, tomentose externally, coppery, inner sepals scarious marginally; adaxial petal 2.0 – 2.1 × 3.2 – 4.0 cm, oblate, non-tubular, apex rounded, base truncate, pink to purple, glabrous; petalodia 0.5 – 0.8 × 0.8 – 1.0 mm; stamens, exserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 4.2 – 5.0 cm long, shorter filaments 3.0 cm long, sheath equilateral, 5.2 – 8.0 mm long, sheath irregularly tomentose in the middle-high portion and glabrous at the base, free filaments glabrous, coppery, anthers 5.2 – 6.6 × 1.0 – 1.5 mm, rectangular; ovary 9.2 – 12.0 × 4.0 – 4.2 mm, oblaceolate, tomentose, cream (fresh) to coppery (dried), stipe 8.8 – 15.0 mm long, tomentose, style 3.5 – 5.0 cm long, stigma obtuse to capitate. Legumes 21.0 × 7.6 cm, stipe not seen, oblanceolate-falcate, apex acute, apiculate, margin entire, coppery, veins absent, tomentulose, coppery. Seeds, 4.3 × 2.6 cm, ovate to oblong. Phenology: — Flowering in September and October.; fruiting in October	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	distribution	Distribution: — Guyana (Cuyuni-Mazaruni). Distribution and habitat: — Floodplain (gallery) forests on mixed brown and white sand soils, at 63 – 90 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	conservation	Conservation status: — Categorized as Data Deficient (DD) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022) as it is known by only two specimens. Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	etymology	Etymology: — The specific epithet is a tribute to Karen Redden, a botanist who studied Detarieae groups and the Guyana Shield Flora, besides collecting a lot of species of Eperua (over eighty specimens and 11 species), including the type specimen of this new species. Vernacular names: — Described as having the common name “ tricel ” in the Redden K. M. et al. 3260 collection label. However, a similar name, “ trysil ”, is the vernacular name of the locally quite common species, Pentaclethra macroloba Kuntze, and likely the name “ tricel ” was mistakenly coined by the collectors (ter Steege pers. obs.)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	description	Uses: — Unknown. Taxonomic notes: — Eperua reddeniae, E. froesii, E. cerradoensis, E. duckeana and E. schomburgkiana are the only species with short and erect inflorescences that have non-tubular corolla, exserted stamens, falcate fruit with a 3: 1 length / width proportion (vs. tubular corolla, included stamens, non-falcate fruit with a 1: 1 to 2: 1 length / width proportion in other short and erect inflorescence species). They also share multijugate leaves with straight leaflets and inflorescence with dense indumentum. Additionally, E. reddeniae shares caducous bracteoles and stamens joined in a diadelphous sheath 9 + 1 with E. duckeana and E. schomburgkiana; and irregularly tomentose sheath with E. schomburgkiana. Similarities and differences between species of this group are summarized in Table 3. The flower of E. reddeniae is most similar to E. duckeana by the non-glabrous sheath, ovary, and fruits, and the diadelphous stamen sheath. Besides the allopatric distribution (E. reddeniae is restricted to Guyana, and E. duckeana to the Central Amazonia, Figure 8), E. reddeniae differs from E. duckeana mainly by the coppery inflorescence indumentum (vs. brownish-yellow in E. duckeana), pink to purple petals (vs. white in E. duckeana), and irregularly tomentose sheath (vs. entirely tomentose in E. duckeana).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F63FF81FF6CF897FDCBF78B.taxon	materials_examined	Specimens examined (Paratypes): — GUYANA. Cuyuni-Mazaruni, Mazaruni River, unnamed falls (possible Carin Crow Falls), 0.26 miles NW of base camp, 63 m elev., 5 ° 57 ’ 25 ” N 59 ° 18 ’ 41.8 ” W, 09 September 2004, Redden K. M. et al. 3260 (US [00889812]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF85FF6CFF33FED4FA4A.taxon	description	(Figures 12, 51, 52)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF85FF6CFF33FED4FA4A.taxon	materials_examined	Type (designated by Cowan 1975): — SURINAM. Without locality, s. d., Hostmann F. W. 229 (lectotype U [0003237] image!; isolectotypes K [000555108] image!, LE [00002234] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF85FF6CFF33FED4FA4A.taxon	description	Tree 6.0 – 30.0 (– 40.0) m tall. Trunk 12.0 – 50.0 (– 91.0) cm in diameter, bark grayish, lenticellate. Stipules 1.2 – 3.5 (– 4.1) × 0.7 – 2.2, free, or 1.3 – 7.7 × 3.0 – 5.0 mm, joined at the base, non-foliaceous, caducous. Leaves (2 –) 3 (– 4) or (3 – 5) 4 – 5 - jugate; petioles (0.4 –) 0.8 – 3.8 (– 8.0) cm long; rachis (2.3 –) 4.0 – 15.2 (– 18.5) or 7.3 – 25.0 cm long; petiolules (2.0 –) 4.9 – 12.5 mm long; blades 5.1 – 16.0 (– 19.0) × 2.3 – 7.8 cm, coriaceous, glabrous, epunctate to inconspicuous pellucid-punctate, rarely pellucid-punctate, not discolorous to discolorous, equilateral, lower pair elliptic, sometimes lanceolate or ovate, middle pair elliptic, rarely lanceolate or ovate, upper pair elliptic, rarely lanceolate or obovate, apex attenuate to acuminate, mucronate, sometimes caudate, base symmetrical, lower pair base obtuse to rounded, rarely subcordate, middle pair base obtuse, rarely rounded, subcordate or cordate, upper pair base obtuse to cuneate, margin flat to slightly revolute, rarely revolute, secondary venation with one intramarginal vein, sometimes two, the vein closer to the margin not continuous, or, two intramarginal veins, sometimes one, the vein closer to the margin continuous or not, main vein straight, depressed on the adaxial surface, tertiary veins inconspicuous to conspicuous, areoles smaller, concave. Inflorescences terminal, sometimes axillary, panicle, main axis pendulous, lateral racemes erect, rarely patent, tomentulose, coppery, 46.0 – 280.0 (– 500.0) or 40.0 – 150.0 cm long, lateral racemes 1.8 – 11.5 (– 16.5) cm long, alternate and spirally arranged; bracts not seen; bracteoles 1.9 – 3.8 × 1.2 – 2.3 mm, ovate, cucullate, apex gland absent, tomentose, coppery, caducous, attached to the lower portion of the pedicels; pedicel 9.7 – 33.0 mm long, 0.6 – 2.0 or 2.0 – 3.3 mm in diameter, not twisted, tomentose, coppery; buds 0.6 – 2.1 cm long, 0.3 – 1.2 cm in diameter, tomentose, coppery. Flowers: hypanthium 3.1 – 9.6 mm long, 2.8 – 5.9 or 4.9 – 7.0 mm in diameter, cup-shaped, equilateral, tomentose, coppery; sepal 1.0 – 1.8 (2.1) × 0.3 – 1.1 or 1.7 – 2.3 × 0.6 – 1.0 cm, elliptic to oblong, unequal, the outer ones larger, dorsal one cucullate, apex gland present on the outer ones, coppery, tomentose, inner sepals scarious marginally, coppery; adaxial petal 1.0 – 2.6 × 1.6 – 5.0 cm, oblate, non-tubular, apex rounded, crisped, base truncate, reddish, pink, magenta, purple-red to red, glabrous, punctate, papillose; petalodia 0.3 – 2.5 × 0.2 – 1.2 mm; stamens exserted, joined basally in a tube, longer filaments 3.0 – 6.8 cm long, shorter filaments 2.2 – 5.5 cm long, tube equilateral, 0.4 – 4.8 mm long, glabrous, sometimes tomentulose in the sinuses, coppery, anthers 4.1 – 9.3 × 0.4 – 1.6 mm, rectangular; ovary 4.2 – 13.0 × 1.6 – 4.3 mm, oblanceolate, tomentose, coppery, 5 ovules, stipe 3.9 – 15.5 mm long, tomentose, style 2.0 – 7.4 cm long, stigma capitate. Legumes 13.4 – 30.3 × 3.3 – 10.0 cm, stipe 1.0 – 4.9 cm long, with flower remnants, falcate-oblanceolate, apex acuminate, acute to rounded, dorsal margin thickened, ventral margin entire, coppery, veins absent, tomentose, coppery. Seeds 1 – 3 per fruit, 2.0 – 5.0 × 0.7 – 3.0 cm, oblate, oblong, ovate to orbicular, dark-brown, centrally reddish-brown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF85FF6CFF33FED4FA4A.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 632,928.628 km 2 and the estimated area of occupancy is 436.000 km 2. Eperua rubiginosa has to varieties presented here: E. rubiginosa var. rubiginosa and E. rubiginosa var. grandiflora.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	description	Stipules 1.2 – 3.5 (– 4.1) × 0.7 – 2.2, free. Leaves (2 –) 3 (– 4); rachis (2.3 –) 4.0 – 15.2 (– 18.5) cm long; secondary venation with one intramarginal vein, sometimes two, the vein closer to the margin not continuous. Inflorescences 46.0 – 280.0 (– 500.0) cm long; pedicel 0.6 – 2.0 mm in diameter. Flowers: hypanthium 2.8 – 5.9 mm in diameter; sepal 1.0 – 1.8 (2.1) × 0.3 – 1.1 cm.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	biology_ecology	Phenology: — Flowering throughout the year with a flowering pick from August to December; fruiting throughout the year, mainly from October to November.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	distribution	Distribution: — Brazil (Amapá, Pará), French Guiana (Cayenne, Saint-Laurent-du-Maroni), Guyana (Cuyuni-Mazaruni, Potaro-Siparuni, Upper Demerara-Berbice, Upper Takatu-Upper Essequibo), and Surinam (Brokopondo, Kabalebo, Marowijne, Para, Sipaliwini).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	biology_ecology	Habitat: — Mainly in floodplains forests (igapó, swamp forest), sometimes in terra-firme forest, rarely in wallaba forest, on white and brown sand, stony clayey, sandy stony, and laterite soils, from sea level to 350 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	conservation	Occurrence in protected areas: — Parque Indígena Tumucumaque, Parque Nacional Montanhas do Tumucumaque (Brazil), Réserve Naturelle de Nouragues (French Guiana, Cayenne).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	etymology	Etymology: — The specific epithet relates to its inflorescence, flowers, and fruit color indumentum.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	vernacular_names	Vernacular names: — apa (Irwin H. S. & Westra L. Y. T. 47471, Pires J. M. & Westra L. Y. T. 48849), bioudou (BAFOG 6239, BAFOG 7831, BAFOG 7937, IFAT 7831, IFAT 7937), eturi wallaba (Redden K. M. et al. 6012), franga (boni), ouapa (créole) (Schnell R. A. A. 11456), iperana-apá (Rosa N. A. 1183), ituri wallaba (Fanshawe D. B. 4840), oever wallaba (Maguire B. et al. 54014), ouapa (nom créole) (Oldeman R. A. A. 1552), tapaka (Grenand P. 1521), uapá (Candido J. s. n. IAN 44221, IAN 44221 a), walaba (BAFOG 219, Landsbosbeheer 311, Lanjouw J. 1209, 738, Lanjouw J. & Lindeman J. C. 324, Mennega A. M. W. 102), wallaba (Evans R. et al. 1929, Junker L. 249), wapa (Angel G. 126, Fleury M. 1129, Hoff M. 5964, Oldeman R. A. A. 1520, Thiel J. 891, 892), wapa riviére (Angel G. 126, Bordenave B. 228, Paget D. 51, Thiel J. 891, 892, Prévost M. F. 4038, Benoist R. s. n. P 03465703).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	discussion	Uses: — Brazil: wood used for house construction (Fortes E. A. collections), shingles, fencing, and poles (Pires J. M. & Westra L. Y. T. 48849). Taxonomic notes: — Eperua rubiginosa shares with E. glabra the multijugate leaves with straight leaflets, long and pendulous inflorescences, erect lateral racemes, caducous bracteoles attached to the lower portion of the pedicels, non-tubular corolla, and exserted stamens joined in a tube. However, E. rubiginosa differs by the tomentulose inflorescence (vs. glabrous in E. glabra) and tomentose ovary and fruit (vs. glabrous in E. glabra) (see Table 4).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F66FF88FF6CF9B7FEA0F8DF.taxon	materials_examined	Selected specimens: — BRAZIL. Amapá, Macapá, Fazendinha, Parque Zoobotânico, IEPA, rodovia JK, 10 ki, empreendimento AHE, Santo Antônio do rio Jari, 17 August 2010, Carim M. J. V. & Guimarães J. R. S. 533 (HAMAB). Oiapoque, nas margens da rodovia BR 156 entre Oiapoque e Calçoene, a 23 km de Oiapoque, 3 ° 42 ’ 11.8 ” N 51 ° 46 ’ 24.1 ” W, 04 December 2021, Fortes E. A. & Silva G. S. da 167 (IAN, RB); nas margens do rio Oiapoque próximo à ponte BR-GF, lado brasileiro, 3 ° 51 ’ 22.1 ” N 51 ° 49 ’ 29.7 ” W, 05 December 2021, Fortes E. A. & Silva G. S. da 168 (IAN, RB). Pedra Branca do Amapari, Parque Nacional Montanhas do Tumucumaque, acima da Base Jupará, 105 m elev., 1 ° 11 ’ 16 ” S 52 ° 22 ’ 8 ” W, 08 November 2018, Goldenberg R. et al. 2522 (MG, RB); rio Amapari, between Munguba and Serra do Navio, 0 ° 42 ’ 0 ” N 51 ° 45 ’ 0 ” W, 25 September 1961, Pires J. M. et al. 51182 (IAN, INPA, K, MG, NY, U, US). Serra do Navio, rio Araguari, between camps 12 and 13, 04 October 1961, Pires J. M. et al. 51501 (IAN, MG, NY, US). Pará, Óbidos, rio Paru do Oeste (Tirió), barranco à beira do rio, 26 June 1960, Cavalcante P. B. 870 (MG, NY). Oriximiná, distrito de Porto Trombetas, km 6, cachoeira da Porteira, 1 ° 5 ’ 23 ” S 57 ° 2 ’ 84 ” W, 17 September 2003, Faria S. M. de et al. 2450 (HSTM, MBM, RB). FRENCH GUIANA. Cayenne, Cayenne, unknown locality, October 1910, 1948, Santini 1948 (L). Kourou, riviére Kourou, bassin du Kourou, 10 October 1980, Thiel J. 729 (CAY). Régina, station des Nouragues (cam inselberg), trail XV detour near RG on Grand Plateau, 40 – 20 m elev., 4 ° 5 ’ 30 ” N 52 ° 41 ’ 0 ” W, 10 October 2008, Geiselman C. K. 169 (NY). Roura, rivière la Counamama, 4 ° 31 ’ 0 ” N 52 ° 16 ’ 0 ” W, 06 October 2000, Prévost M. F. 4038 (CAY). Saint-Élie, Saut takari-tante, bassin du sinnamary, m elev., 4 ° 37 ’ 0 ” N 52 ° 56 ’ 0 ” W, 19 November 1989, Hoff M. 5964 (CAY). Saint-Georges, crique Gabaret, 3 ° 55 ’ 0 ” N 51 ° 47 ’ 59 ” W, 07 December 2005, Delnatte C. 325 (CAY, COL, MPU). Sinnamary, pointe Combi, région littorale, riviére Sinnamary, 24 November 1979, Thiel J. 588 (CAY). Saint-Laurent-du-Maroni, Grandi-Santi, 25 August 1961, Schnell R. A. A. 11456 (CAY, P, U, US). Mana, unknown locality, 06 December 1979, Thiel J. 601 (CAY). Maripasoula, camp no. 1 ouman fou langa soula — bassin du haut marouini Saint-Laurent-du-Maron, 2 ° 52 ’ 59.9 ” N 54 ° 0 ’ 0 ” W, 24 August 1987, Granville J-J. de 9666 (CAY, US). GUYANA. Cuyuni-Mazaruni, Iwokrama Rainforest Reserve, between Pakutau Falls and Ireng Creek on Siparuni R., 75 m elev., 4 ° 45 ’ 0 ’’ N 59 ° 01 ’ 0 ’’ W, 09 November 1995, Clarke H. D. 442 (US). Potaro-Siparuni, Kaieteur National Park, plot Study # 1, NE from end of airstrip at headwaters of Korume Creek, 300 – 350 m elev., 5 ° 10 ’ 0 ” N 59 ° 28 ’ 59.9 ” W, 11 – 19 March 1997, Kelloff C. L. & McKee G. S. 1246 (US) Upper Demerara-Berbice, Fairview Landing E bank of Essequibo River, near end of Mabura road, 91 m elev., 4 ° 40 ’ 0 ” N 58 ° 40 ’ 0 ” W, 19 July 1990, McDowell T. 3271 (INPA, US). Upper Takatu-Upper Essequibo, Acarai Mts., 4 km S of Sipu River, area near camp at base of Acarai Mts., 245 m elev., 1 ° 23 ’ 0 ” N 58 ° 56 ’ 0 ” W, 08 September 1998, Clarke H. D. et al. 7639 (U, US); Essequibo region, Kuyuwuni river watershed, Airshalton Landing, upstream 2 – 3 km N & S banks, 250 m elev., 2 ° 03 ’ 00 ” N 59 ° 10 ’ 0 ” W, 22 November 1993, Henkel T. W. et al. 3100 (U, US); Essequibo River, 222 m elev., 2 ° 15 ’ 21.1 ” N 58 ° 19 ’ 26.5 ” W, 27 April 2004, Redden K. M. et al. 3130 (US);. SURINAM. Brokopondo, van Blommesteijn Lake, Island south, 48 m elev., 4 ° 33.8 ’ 0 ” N 55 ° 7.2 ’ 0 ” W, 11 April 2011, Bhikhi C. R. et al. 873 (U). Kabalebo, Central SURINAM Nature Reserve, unnamed tributary of Coppename River 200 m South of Driefoetoe Sula, 50 – 75 m elev., 4 ° 45 ’ 0 ” N 59 ° 1 ’ 0 ” W, 10 March 2004, Clarke H. D. & Rhodes J. 11375 (US). Marowijne, Albina, ad ripas fluv. Marowijne. S of Albina, 19 September 1948, Lanjouw J. & Lindeman J. C. 324 (IAN, U). Para, unknown locality, undated, Voltz F. s. n. U 1302936 (U). Sipaliwini, Coeroeni, N side of Kuruni Island, E of trail from airstrip to N branch of Kuruni River around island, ca. 30 airline km E of confluence on Corantijn River, 180 m elev., 3 ° 22 ’ 12 ” N 57 ° 20 ’ 36 ” W, 10 November 1994, Evans R. et al. 1929 (CAY, IAN, INPA, P, RB, U, US). Goddo, unknown locality, 19 August 1908, Tresling J. H. A. T. 334 (RB, U). Raleigh vallen- Voltzberg Nature Reserve on the Coppename River, 145 kms SW of Paramaribo, vicinity of Lolopasi Hut, left bank of the Coppename River, 21 November 1976, Mori S. A. & Bolten A. 8673 (U).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	materials_examined	Type: — SURINAM. Fluv. Gonini, August 1903, Versteeg G. M. 48 (holotype U [0003236] image!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	description	Stipules 1.3 – 7.7 × 3.0 – 5.0 mm, joined at the base. Leaves (3 –) 4 – 5 - jugate; rachis 7.3 – 25.0 cm long; secondary venation with two intramarginal veins, sometimes one, the vein closer to the margin continuous or not. Inflorescences 40.0 – 150.0 cm long; pedicel 2.0 – 3.3 mm in diameter. Flowers: hypanthium 4.9 – 7.0 mm in diameter; sepal 1.7 – 2.3 × 0.6 – 1.0 cm.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	biology_ecology	Phenology: — Flowering in March, May, June, August, and October; fruiting in January, February, March, and October.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	distribution	Distribution: — Previously known in Surinam (Marowijn and Sipaliwini), here we recognize two collections occurring in French Guiana (Redden K. M. et al. 6005, Schnell R. A. A. 11456).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	biology_ecology	Habitat: — Floodplains (riparian vegetation and riverbanks) forests on loamy sandy soil, below 100 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	conservation	Occurrence in protected areas: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	etymology	Etymology: — The specific epithet relates to its flower that is larger than the type variety.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	vernacular_names	Vernacular names: — panga (Andel T. R. van et al. 4979).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	discussion	Uses: — Surinam: used with Pikin fóo kaka in herbal baths for protection from cutlass slashes (Andel T. R. van et al. 4979). Taxonomic notes: — The most noticeable difference between the two varieties is in the pedicels’ diameter. Eperua rubiginosa var. grandiflora has a thicker pedicel (> 2 mm in diameter), and the type variety has a thinner pedicel (> 2 mm in diameter, mainly bellow 1.5 mm). The flower of E. rubiginosa var. grandiflora looks more robust and it has generally larger sepals. Vegetatively, although there is no precise difference, var. grandiflora has bigger joined stipules (vs. free in var. rubiginosa), generally 4 or 5, rare 3, leaflets pairs (vs. generally 3, sometimes 4, in var. rubiginosa), larger rachis (vs. shorter in var. rubiginosa), and in most of the cases, secondary venation with two intramarginal veins (vs. one in var. rubiginosa).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6BFF89FF6CF80BFB46FB1A.taxon	materials_examined	Selected specimens: — FRENCH GUIANA. Saint-Laurent-du-Maroni, Maroni River near St. Larent., 59 m elev., 5 ° 28 ’ 45 ” N 54 ° 3 ’ 43 ” W, 05 May 2007, Redden K. M. et al. 6005 (CAY, US). GUYANA. Cuyuni-Mazaruni, Mazaruni River, unnamed falls (possible Carin Crow Falls), 0.25 miles NW of base camp, 59 m elev., 5 ° 57 ’ 23.2 ” N 59 ° 18 ’ 43 ” W, 09 October 2004, Redden K. M. et al. 3261 (INPA, COL, US); Mazaruni River, small islands around falls, 49 m elev., 5 ° 55 ’ 57.9 ” N 59 ° 6 ’ 35.3 ” W, 12 October 2004, Redden K. M. et al. 3286 (US); Mazaruni River, 1.15 miles NE of base camp, 16 m elev., 6 ° 06 ’ 50.1 ” N 58 ° 53 ’ 53.9 ” W, 14 October 2004, Redden K. M. et al. 3304. SURINAM. Marowijne, Albina, Bigiston, Gonkíiki / Poitikondée, 0? ° 52 ’ 26 ” N 54 ° 7 ’ 48 ” W 09 March 2006, Andel T. R. van et al. 4979 (U). Sipaliwini, Jacob Kondre, Saramacca river, 16 June 1944, Maguire B. 23804 (F, U, US). Gonini, 25 February 1918, Gonggrijp J. W. 3700 (U); fluv. Gonini, 11 August 1903, Versteeg G. M. 48 (CAY, U).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	description	(Figures 8, 53)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	materials_examined	Type (designated by Cowan 1975 / Fortes & Mansano 2022): — GUYANA. Essequibo & Rupununy, 1838, Schomburgk Rob. Ser. 1 517 (lectotype K [000555112] image!, isolectotypes BM [000065064] image!, G [00365223] image!, G [00365208] image!, K [000555113] image!, K [000555114] image!, P [00252893] image!, W not seen).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	description	Tree (3.0 –) 4.0 – 35.0 m tall. Trunk and bark not seen. Stipules (6.2 –) 10.3 – 30.0 (– 44.0) mm, free, foliaceous, persistent to caducous. Leaves (3 –) 4 – 7 - jugate; petioles 1.0 – 3.6 cm long; rachis (7.3 –) 10.0 – 23.5 (– 30.0) cm long; petiolules 5.2 – 11.9 mm long; blades 5.9 – 18.5 × (1.7 –) 2.7 – 7.2 cm, coriaceous, glabrous, pellucid-punctate, sometimes inconspicuous pellucid-punctate, discolorous to not discolorous, equilateral, lower pair elliptic, narrowly elliptic to ovate, rarely lanceolate, middle pair elliptic, narrowly elliptic to ovate, rarely lanceolate, upper pair elliptic to narrowly elliptic, apex attenuate, acuminate to caudate, base symmetrical to asymmetrical, lower pair base obtuse to rounded, sometimes subcordate, middle pair base obtuse to rounded, upper pair base obtuse to cuneate, margin flat, secondary venation with one intramarginal vein, sometimes two intramarginal veins, vein closer to the margin not continuous, main vein straight, prominent only near the base to entirely prominent on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal, rarely axillary, panicle, rarely raceme, erect, tomentose, coppery, 4.0 – 11.5 cm long, lateral racemes 1.9 – 7.2 cm long, alternate and distichous; bracts 4.1 – 5.9 × 3.4 – 5.6 mm, ovate, broadly ovate to orbicular, apex gland absent?, tomentose externally, coppery, caducous; bracteoles 4.5 – 6.3 × 3.0 – 5.0 mm, ovate, cucullate, apex gland absent, tomentose externally, coppery, caducous, attached to the middle portion of the pedicels; pedicel 10.8 – 26.2 mm long, 1.4 – 2.7 mm in diameter, not twisted, tomentose, copery; buds 1.0 – 2.2 cm long, 0.5 – 1.0 cm in diameter, tomentose, coppery. Flowers: hypanthium 3.8 – 6.4 mm long, 4.7 – 7.3 mm in diameter, cup-shaped, equilateral, tomentose, coppery; sepal 1.6 – 2.6 × 0.4 – 1.7 cm, elliptic to oblong, unequal, the dorsal one larger, apex gland absent, tomentose externally, coppery, inner sepals scarious marginally; adaxial petal 2.0 – 3.2 × 2.9 – 5.0 cm, oblate, non-tubular, apex rounded, base truncate, white, glabrous; petalodia 1.1 – 1.8 × 1.1 – 1.6 mm; stamens exserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 3.0 – 4.5 cm long, shorter filaments 2.9 – 3.5 cm long, sheath equilateral, 7.0 – 12.0 mm long, sheath irregularly tomentose on both sides to glabrous externally and entirely tomentose in the inner surface, free filaments glabrous, coppery, anthers 4.0 – 6.1 × 1.0 – 2.2 mm, rectangular; ovary 6.0 – 15.0 × 2.4 – 5.0 mm, oblanceolate, glabrous, stipe 6.3 – 13.7 mm long, glabrous, style 2.2 – 3.7 cm long, stigma capitate, papillose. Legumes 12.0 – 23.0 × 4.0 – 7.5 cm, stipe 1.3 – 2.0 cm long, oblanceolate-falcate, apex obtuse, apiculate, margin entire, brown to reddish brown, veins absent, glabrous, foveate. Seeds 1 – 3 per fruit, 3.5 – 5.0 × 1.8 – 3.6 cm, elliptic, oblong to irregularly oblong, brown to reddish-brown. Phenology: — Flowering in March and from June to December, with a peak in September; fruiting throughout the year, except in June and August.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	distribution	Distribution: — Throughout Guyana (Cuyuni-Mazaruni, East Berbice-Corentyne, Essequibo Islands-West Demerara, Macouria, Potaro-Siparuni, Upper Demerara-Berbice, Upper Takatu-Upper Essequibo) and in Surinam (Nickerie, Sipaliwini).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	biology_ecology	Habitat: — Mainly in floodplain forests (e. g. Mora forests and swamps), sometimes in wallaba forests, on alluvial soil, predominately white sand, but also brown sand mixed, loamy and laterite soils, from sea level to 130 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	conservation	Conservation status: — Categorized as Least Concern (LC) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 61,910.701 km 2 and the estimated area of occupancy is 156.000 km 2. Occurrence in protected areas: — Iwokrama Rainforest Reserve (Guyana, Potaro-Siparuni) and Conservation International concession on the Essequibo River (Guyana, Upper Takatu-Upper Essequibo).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	etymology	Etymology: — The specific epithet is a tribute to Schomburgk, who first collected the species in Guyana (Bentham 1870).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	vernacular_names	Vernacular names: — bimiti wallaba (Mori S. A. et al. 8162), clump wallaba (Redden K. M. et al. 6011), cutlass (Redden K. M. et al. 3258, 3272), ituri wallaba (Mori S. A. et al. 8162), machetta (Mori S. A. et al. 8162), maseta (Redden K. M. et al. 3258), museta (Redden K. M. et al. 3272), wallaba (Anderson C. W. 182), water wallaba (Mori S. A. et al. 8162, Pennington R. T. 373, Persuad A. C. 14, 166, 183, Schulz J. P. 7708).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	discussion	Uses: — unknown. Taxonomic notes: — Eperua schomburgkiana, E. cerradoensis, E. duckeana, E. froesii, and E. reddeniae are the only short and erect inflorescence species that have non-tubular corolla, exserted stamens, falcate fruit with a 3: 1 proportion (vs. tubular corolla, included stamens, non-falcate fruit with a 1: 1 to 2: 1 length / width proportion in other short and erect inflorescence species). They also share multijugate leaves with straight leaflets and inflorescence with dense indumentum. Similarities and differences between species of this group are summarized in Table 3. Eperua schomburgkiana is a well-differentiated species by the stamens joined basally in a sheath 9 + 1, irregularly tomentose sheath, glabrous ovary and fruits, besides other vegetative characteristics (see Table 3).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6AFF8FFF6CFB48FC87F843.taxon	materials_examined	Selected specimens: — GUYANA. Cuyuni-Mazaruni, Bartica, Moorie island in the Essequibo River near Bartica, 5 – 25 m elev., 18 August 1976, Mori S. A. et al. 8162 (U, US); Essequibo River, Moraballi Creek, near Bartica, 28 August 1929, Sandwith N. Y. 142 (RB, P, U, US). W bank of Essequibo River 0 – 2 km south of Wolga settlement, 0 – 15 m elev., 6 ° 27 ’ 0 ” N 58 ° 37 ’ 59 ” W, 24 December 1992, Henkel T. W. & Chin M. 669 (U, US); Marshall Falls, Mazaruni River, 37 m elev., 6 ° 17 ’ 5.6 ” N 58 ° 43 ’ 15.5 ” W, 06 March 2004, Redden K. M. et al. 2115 (US). East Berbice - Corentyne, Canje River, Cow Savanna E of Digitima Creek, 1 – 25 m elev., 5 ° 36 ’ 0 ” N 57 ° 35 ’ 0 ” W, 12 April 1987, Pipoly J. J. et al. 11448 (CAY, MO, U, US). Essequibo Islands-West Demerara, Esequibo, along bank of river, 7.5 km. from Lanaballi, 1 – 10 m elev., 6 ° 60 ’ 0 ” N 57 ° 75 ’ 0 ” W, 10 September 1992, Tiwari S. 851 (NY). Potaro-Siparuni, Iwokrama Forest Reserve, Burro-Burro River, 3.42 miles from Whitewater base camp, 65 m elev., 4 ° 40 ’ 58 ” N 58 ° 50 ’ 47.3 ” W, 02 November 2002, Redden K. M. et al. 1142 (US); Micobe Road and adjacent forest next to Potaro River, 22 m elev., 5 ° 20 ’ 29 ” N 58 ° 58 ’ 51 ” W, 14 September 2006, Redden K. M. et al. 4231 (NY, US). Upper Demerara-Berbice, Lower 3 km of Macouria River, N and S banks, 15 m elev., 6 ° 25 ’ 0 ” N 58 ° 34 ’ 0 ” W, 07 December 1992, Henkel T. W. & Chin M. 388 (U, US); Essequibo River, E bank, upstream 2 – 4 km from Bartica, island shore and mainland, 0 – 5 m elev., 6 ° 25 ’ 0 ” N 58 ° 35 ’ 0 ” W, 31 March 1993, Henkel T. W. et al. 1798 (US); CI concession, Himara Creek, 130 m elev., 3 ° 22 ’ 0 ” N 58 ° 11 ’ 0 ” W, 14 September 2008, Redden K. M. et al. 6011 (US). Upper Takatu-Upper Essequibo, Conservation International concession on the Essequibo River, 108 m elev., 3 ° 35 ’ 18 ” N 58 ° 17 ’ 58 ” W, 21 January 2007, Redden K. M. et al. 5138 (F, NY, U, US). Essequibo & Rupununy, 1838, Schomburgk Rob. Ser. I 517 (BM, G, K, P, W). SURINAM. Nickerie, Corantijn, ten N. mondingen New River, 01 September 1935, Rombouts H. E. 170 (U). Sipaliwini, Wonatobo falls, Corantijn River, 26 June 1956, Schulz J. P. 7708 (U, US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	description	(Figures 11, 54, 55)	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	materials_examined	Type: — VENEZUELA. Bolívar: Angell Falls, undated, Kunhardt H. R. Jr. 20 (holotype NY [00004386]!).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	description	Tree 10.0 – 30.0 m tall. Trunk 60.0 cm in diameter, bark not seen. Stipules 3.0 – 4.3 × 2.0 – 3.7 mm, joined, non-foliaceous, caducous, caducous. Leaves 1 – 2 (– 3) - jugate; petioles 1.3 – 3.4 cm long; rachis 2.4 – 5.1 cm long; petiolules 6.5 – 9.4 mm long; blades 6.6 – 21.0 × 3.3 – 9.3 cm, coriaceous, glabrous, pellucid-punctate, each areole has at least one conspicuous dot, not discolorous, equilateral to inequilateral, lower pair elliptic, slightly falcate, apex acuminate, base symmetrical to asymmetrical, lower pair base obtuse, upper pair base obtuse to cuneate, margin flat, secondary venation with one intramarginal vein very close to the margin, main vein curved to straight, prominent to depressed on the adaxial surface, tertiary veins very conspicuous, areoles larger, plane. Inflorescences terminal or axillary, panicle, main axis pendulous, lateral racemes erect, main axis puberulent to strigulose, lateral racemes strigose, pubescent to tomentulose, grayish-white, 145.0 – 162.0 cm long, lateral racemes 2.0 – 5.2 cm long, alternate and spirally arranged; bracts not seen; bracteoles 4.1 – 5.8 × 3.8 – 5.1 mm, ovate to oblate, cucullate, apex gland absent, strigose externally, grayish-white, caducous, attached to the middle to high portion; pedicel 7.0 – 15.0 mm long, 2.5 – 7.6 mm in diameter, not twisted?, pubescent, strigose to tomentose, grayish-white; buds 2.0 cm long, 1.0 cm in diameter, pubescent to tomentose, grayish-white. Flowers: hypanthium 6.8 – 11.3 mm long, 4.2 – 6.5 mm in diameter, tubular to cup-shaped, equilateral, pubescent, strigose to tomentose, grayish-white; sepal 1.6 – 2.2 × 0.6 – 2.2 cm, elliptic to oblong, unequal, the outer ones larger, cucullate, apex gland absent, green with purple spots, pubescent, strigose to tomentose, scarious marginally, grayish-white; adaxial petal 2.5 – 3.4 × 4.7 – 6.0 cm, oblate, non-tubular, apex rounded, base attenuate, gibbous, deep-rose, sericeous at the base, glabrous in other parts, pellucid-punctate; petalodia 0.4 – 0.8 × 0.5 – 0.8 mm; stamens exserted, joined basally in a diadelphous sheath, dorsal one free sheath equilateral, 6.7 – 10.0 mm long, sheath sericeous to tomentose, free filaments glabrous, reddish-white, anthers 4.0 – 9.0 × 1.5 – 1.9 mm, rectangular; ovary 7.3 – 10.0 × 3.2 – 4.9 mm, oblanceolate, densely sericeous, reddish-white, stipe 3.5 – 7.0 mm long, tomentose to sericeous, style length not seen, stigma capitate to bilobate. Legumes oblanceolate-falcate, puberulous (young fruit). Seeds not seen.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	biology_ecology	Phenology: — Flowering in August and September; fruiting in February.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	distribution	Distribution: — Endemic to Venezuela (Bolívar), known only in the region of Gran Sabana.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	biology_ecology	Habitat: — Forests, being frequent in forest borders (Maguire B. et al. 53597), it is also found bordering savanna (Steyermark J. A. 115541), from 87 to 1160 m elev.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	conservation	Conservation status: — Categorized as Vulnerable (VU) according to the IUCN criteria (IUCN 2012, IUCN Standards and Petitions Committee 2022). The extent of estimated occurrence for the species is 6,325.304 km 2 and the estimated area of occupancy is 16.000 km 2. Occurrence in protected areas: — Parque Nacional Canaima (Venezuela, Bolívar).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	etymology	Etymology: — The specific epithet relates to its very conspicuous tertiary veins on the adaxial blade surface.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	vernacular_names	Vernacular names: — Unknown.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	discussion	Uses: — Unknown. Taxonomic notes: — Eperua venosa is the only species in the genus with basally sericeous petals. It is morphologically similar to E. leucantha and E. falcata, with which it shares joined stipules, secondary venation with one intramarginal vein very close to the margin, long and pendulous inflorescences, non-tubular corolla, exserted stamens joined in a diadelphous sheath, and sericeous ovary (Table 4). Besides that, E. venosa shares non-white petals with E. falcata, patent to erect lateral racemes and sericeous sheath with Eperua leucantha. However apart from the petal indumentum, Eperua venosa differs from E. falcata and E. leucantha by other characteristics that are listed in their taxonomic notes.	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
03A487B62F6FFF8CFF6CFF33FB15F8F3.taxon	materials_examined	Selected specimens: — VENEZUELA. Bolívar, Gran Sabana, Angel Falls, undated, Kunhardt H. R. 20 (NY), 7 kms al norte de la Misión de Santa Teresita de Kavanayén, 1160 m elev., 20 February 1978, Steyermark J. A. 115541 (F). Rio Uiri-yuk, El Foco, alto Río Cuyuni, 30 August 1962, Maguire B. et al. 53514 (NY, US). Triana Savanna, cerro Pitón, cordillera Epicara, rio Chicanán, 05 September 1962, Maguire B. et al. 53597 (RB, U, US).	en	Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V., Mansano, Vidal F. (2023): A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae). Phytotaxa 617 (1): 1-127, DOI: 10.11646/phytotaxa.617.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.617.1.1
