taxonID	type	description	language	source
78771043CB72FFA002E1F8BEFCCA6ECE.taxon	type_taxon	Type species: Lithosia leucanioides Walker, 1862, by present designation.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB72FFA002E1F8BEFCCA6ECE.taxon	diagnosis	Diagnosis. Species of the new genus (Figs 1 – 27) are externally very similar to Philippinithosia (Figs 28 – 35). Both the genera have a characteristic creamy or pale ochreous-creamy body and wing colouration, with a forewing pattern represented by the intense brown or blackish suffusion on veins, which can be reduced or absent in some specimens. The only difference found is the androconial tuft on the male foreleg, which consists of elongate and grey scales whereas the corresponding scales of Philippinithosia are ochreous yellow and shorter. In the male genitalia structure, Leucanithosia (Figs 40 – 57) is largely distinct from Tigrioides (Figs 36, 37, 63, 75) and is most similar to the externally fundamentally different Dubatoloviana (Figs 38, 39, 64, 76) due to its short and swollen uncus and the presence of an apical spur on the dorsal section of the valva. However, the male genital capsule of Leucanithosia is characterised by the valvae proximally connected by a commissure and the heavily sclerotised posterior section of the juxta which is laterally fused with transtillae and ventrally connected to the aforementioned commissure whereas Dubatoloviana lacks the intervalval commissure and its juxta is weakly sclerotised. The phallus of the new genus bears a distal serrulate plate laterally whereas it is smooth in Dubatoloviana. In the vesica, Leucanithosia has a stick-shaped distal plate (basal plate of the vesica ejaculatorius) which is absent in Dubatoloviana. The female genitalia of the new genus (Figs 65 – 71) are characterised by the gelatinous and rugose posterior section of the corpus bursae and the large, sack-like and distally membranous appendix bursae situated laterally whereas in Dubatoloviana, the posterior section of the corpus bursae is sclerotised, and the appendix bursae is small, conical, sclerotised and situated postero-laterally.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB72FFA002E1F8BEFCCA6ECE.taxon	description	Description. Adults. Sexual dimorphism limited: female somewhat larger than male and with slightly broader forewing. Antenna dark grey, ciliate in both sexes. Head creamy or ochreous creamy. Thorax creamy. Pro- and metathorax with black spot medio-dorsally. Patagia ochreous creamy. Male foreleg with androconial tuft consisting of elongate and grey scales. Tegula creamy with black spot medially. Forewing elongate and narrow, with almost parallel margins; costal margin straight proximally and medially and downcurved distally; anal margin convex. Forewing ground colour creamy or ochreous creamy. Pattern represented by dark grey or brown suffusion along veins, can be reduced. Forewing cilia creamy or ochreous creamy. Hindwing unicolorous creamy or ochreous creamy, paler than forewing. Abdomen creamy with intense admixture of ochreous scales laterally and distally. Male genitalia. Uncus strongly swollen, distally tapered with a short claw-shaped tip. Tegumen with band-like arms, somewhat downcurved posteriorly. Tuba analis membranous, as broad as half of tegumen length. Vinculum shorter than tegumen, with U-shaped saccus. Dorsal section of valva moderately sclerotised, bearing robust spur and cluster of setae apically. Sacculus broad (equal to valva width), distal saccular process various in shape, not reaching valva apex. Sacculi fused with each other by commissure proximally. Juxta weakly sclerotised and medially membranous anteriorly, with sclerotised and protruding posterior section fused with intervalval commisure and transtillae. Anellus membranous. Phallus cylindrical, with medial protrusion in certain species, and with distal lateral plate densely covered with denticles. Vesica membranous, with several diverticula bearing single spike-like cornuti or clusters of several spines. Vesica ejaculatorius with stick-like basal plate. Female genitalia. Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses elongate and thin, apophysis anterioris shorter than apophysis posterioris. Postvaginal area gelatinous and rugose or with a postvaginal plate. 7 th abdominal sternite forming antevaginal fold in certain species. Ductus bursae elongate, tubular or posteriorly dilated, and forming antrum, rugose. Posterior section of corpus bursae rugose, with sclerotised crests and short postero-lateral protrusion. Anterior section of corpus bursae egg-shaped or almost globular, membranous, and bearing two round or teardrop-shaped signa. Appendix bursae originates postero-laterally, well-separated from corpus bursae, sack-like, gelatinous, and rugose proximally and membranous distally.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB72FFA002E1F8BEFCCA6ECE.taxon	distribution	Distribution. The genus is widely distributed from Northeast India through Indochina and Southwest China to Sundaland including Palawan Island.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB72FFA002E1F8BEFCCA6ECE.taxon	etymology	Etymology. The genus name is an aggregate of the specific epithet of its type species leucanioides and the genus-group name Lithosia Fabricius, 1798. Gender is feminine.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA102E1FF2DFBB76ED1.taxon	description	(Figs 1 – 7, 40 – 44, 65, 66)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA102E1FF2DFBB76ED1.taxon	materials_examined	Type material examined. Lectotype (Figs 1, 65) (hereby designated): female, round label “ Sar ” [awak] / “ A Walker | type | Lithosia | leucanioides | 31 - 227 ” / “ Type | Walker | Journ. Linn. Soc. | Zool. VI. P. 103 | 1862. ” / “ TYPE LEP.: No. 390 | Lithosia | leucanioides | Walker | HOPE DEPT. OXFORD ” / “ 570 ” / “ Collita ” / “ leucanioides ” (OUMNH). Additional material examined. MALAYSIA: 1 male, 3 females, Borneo, Sabah, Mt. Trus Madi, 5 ° 26 ’ N 116 ° 21 ’ E, 900 – 1000 m, iii – iv. 1996, local collectors leg., gen. prep. Nos.: ZSM Arct. 2021 - 001 (male), ZSM Arct. 2021 - 320 (female) (prepared by Volynkin) (MWM / ZSM); 1 male, Sarawak, Kapit Distr., Rumah Tuan Vill., Sut Riv., 03 – 09. iii. 1994, J. Horák leg. (CKC); 1 male, Pahang Distr., Endau Rompin State Park, camping by the Kincin River, 50 m, 2 ° 37 ’ 10.4 ’’ N 103 ° 20 ’ 10.9 ’’ E, 1 – 13. iii. 2011, Karol Bucsek leg., gen. prep. No.: ZSM Arct. 2021 - 080 (prepared by Volynkin (MWM / ZSM); CAMBODIA: 1 male, 20 km SE Koh Kong, Tatai river, 11 ° 34 ’ N 103 ° 07 ’ E, 3 – 19. v. 2005, 50 – 300 m, E. Jendek & O. Šauša leg., gen. prep. No.: No.: ZSM Arct. 2021 - 081 (prepared by Volynkin) (MWM / ZSM); THAILAND: 3 males, 1 female, Lampang Prov., Chae Son NP, 1496 m, 18 ° 51 ’ 23 ’’ N 99 ° 22 ’ E, 1. vi. 2005, K. Černý leg., gen. prep. Nos.: AV 7325, AV 7326 (males), AV 7327 (female) (CKC); 1 female, Ubon Ratchatani Prov., 220 m, border of Nam Yuem NP, 14 ° 25.37 ’ N 104 ° 52.07 ’ E, 2 – 3. xi. 2010, K. Černý leg. (CKC); 1 female, Nakhon Si Tammarat, Khao Luang / Noppitham, 08 ° 47 ’ N 99 ° 42 ’ E, 27. iv. 2008, Plössl leg. (CBP); LAOS: 1 male, Champasak Prov., 27 km ENE of Pakse, near Tad Fane waterfall, 15 ° 11 ’ 01.1 ’’ N 106 ° 07 ’ 40.5 ’’ E, 950 m, local collector leg. (CAV); 1 male, Khamouane [Khammouane Prov.], Phon Hin Boun, 24 – 25. v. 2013, [4] 50 m, 18 ° 10.435 ’ N 104 ° 29.042 ’ E, Thomas Ihle leg., gen. prep. No.: AV 7333 (CKC); VIETNAM: 1 male, Tam Dao, sec. forest, 60 km NW of Hanoi, 1200 m, 21 ° 34 ’ N 105 ° 20 ’ E, 1 – 15. xi. 1992, Sinyaev & Simonov leg., gen. prep. No.: ZSM Arct. 2021 - 312 (prepared by Volynkin) (MWM / ZSM); 1 male, 1 female, Da Nang Province, W of Da Nang City, Tryong Son Mts, Ba Na Mt., 1450 m, 15 ° 59 ’ 56 ’’ N 107 ° 59 ’ 13 ’’ E, local collectors leg., gen. prep. No.: AV 6830 (CAV). Notes. (1) Walker (1862) described the species based on a female but without the number of specimens being mentioned. In order to stabilise the nomenclature, the syntype specimen deposited in the type collection of the OUMNH is hereby designated as the lectotype. (2) The male genitalia of the species slightly vary in the length of the dorsal part of the valva and the distal saccular process as well.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA102E1FF2DFBB76ED1.taxon	diagnosis	Diagnosis. The forewing length is 13.5 – 15.5 mm in males and 15.0 – 17.5 mm in females. The male genitalia structure of L. leucanioides is most similar to L. diehli and the detailed comparison is provided below under the diagnosis of this species.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA102E1FF2DFBB76ED1.taxon	distribution	Distribution. The species is reliably known from Malaysia (Borneo Island and Malay Peninsula) (Holloway 2001; Bucsek 2012), Singapore, Thailand (Černý & Pinratana 2009, partim.), Cambodia, Vietnam and central and southern Laos. The records from northern Laos (Kishida 2020), southern Myanmar (Hampson 1900; Holloway 2001) and south-western China (Yunnan) (Fang 2000) require clarification and may belong to L. witti. The record from Sumatra (Holloway 2001) belongs to either L. diehli or L. inusitata or L. lampunga. The records from the Philippines without islands being specified (Bucsek 2012; Kishida 2020) belong to either L. leptivalva or one of the three species of Philippinithosia. The records from Java (Bucsek 2012) and Xizang, China (Fang 2000) require further clarification as no material available was found during the current study.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA602E1F95AFABA6ADA.taxon	description	(Figs 8 – 10, 45, 46, 67)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA602E1F95AFABA6ADA.taxon	materials_examined	Type material. Holotype (Figs 8, 45): male, [Indonesia] “ Sumatra | Dolok Merangir | 20.4. – 7.5.1981 | Dr. Diehl leg. ” / “ Slide | ZSM Arct. | 2021 - 337 ♁ | A. Volynkin ” (MWM / ZSM). Paratypes. INDONESIA, SUMATRA: 1 female, Aek Tarum, Gunung Malayu, 8. v. 1983, Dr. Diehl leg., gen. prep. No.: ZSM Arct. 2021 - 338 (prepared by Volynkin) (MWM / ZSM); 1 male, Simalungun, Huta Padang, 7. ii. [19] 89, Plössl leg., gen. prep. No.: AV 7320 (prepared by Volynkin) (CBP); 1 female, the same data as previous but 5. ii. [19] 89, gen. prep. No.: AV 7321 (prepared by Volynkin) (CBP).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA602E1F95AFABA6ADA.taxon	diagnosis	Diagnosis. The forewing length is 16.5 mm in males and 18.0 – 19.5 mm in females. The male genital capsule of L. diehli is reminiscent of L. leucanioides but can be distinguished by the longer dorsal part of the valva, which is longer than the tegumen-vinculum complex whereas it is shorter than or equal to the tegumen-vinculum complex in L. leucanioides. Additionally, the distal saccular process of the new species is somewhat more elongate than in the similar congener. The phallus of L. diehli is shorter and narrower than in L. leucanioides (in proportion to the tegumen-vinculum complex length). The vesicae of the two species are similar but that of L. diehli is considerably shorter and narrower than in L. leucanioides. The female genitalia of L. diehli differ from L. leucanioides in the presence of the postvaginal plate, the shorter ductus bursae having a goblet-like dilated posterior section (it is evenly tubular in the congener), and the markedly broader appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA602E1F95AFABA6ADA.taxon	distribution	Distribution. The species is currently known from northern Sumatra Island.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB70FFA602E1F95AFABA6ADA.taxon	etymology	Etymology. The new species is dedicated to the memory of the late Dr. Eduard W. Diehl, a famous researcher of Sumatran insects and collector of the part of the type series. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA602E1FD6CFE336D0A.taxon	description	(Figs 11, 47)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA602E1FD6CFE336D0A.taxon	materials_examined	Type material. Holotype (Figs 11, 47): male, [Indonesia] “ Sumatra Aceh | Westcoast | Kuala Dou near | Calang, 100 m | 20. – 26. II 1996 | leg. A. Kallies ” / “ Slide | AV 7324 ♁ | A. Volynkin ” (MWM / ZSM, ex CKC).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA602E1FD6CFE336D0A.taxon	diagnosis	Diagnosis. The forewing length is 13.5 mm in the male holotype. The male genital capsule of L. paraliaka is similar to L. diehli but readily differs in the broader sacculus with a markedly shorter and proximally broader distal saccular process, which is more S-like curved distally. The phallus of the new species is somewhat longer than in the congener (in proportion to the tegumen-vinculum complex length). Compared to L. diehli, the vesica of L. paraliaka is distally broader, has shorter and semiglobular medial dorsal diverticula (they are utricular in the congener), and a broader distal diverticulum bearing two larger terminal cornuti (whereas L. diehli has a single terminal cornutus). The female is unknown.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA602E1FD6CFE336D0A.taxon	distribution	Distribution. The new species is currently known only from its type locality in coastal north-western Sumatra.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA602E1FD6CFE336D0A.taxon	etymology	Etymology. The specific epithet is derived from the Greek ‘ παραλιακός’ meaning ‘ coastal’ and refers to the occurrence of the new species in the coastal area of north-western Sumatra. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA702E1FA1CFCAD6942.taxon	description	(Figs 12, 13, 48, 49)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA702E1FA1CFCAD6942.taxon	materials_examined	Type material. Holotype (Figs 12, 48): male, “ Myanmar (Burma) | 25 km E Putao | env. Nan Sa Bon vill. | 800 m, 6 – 9. V. 1998 | leg. Murzin & Sinjaev [recte: Sinyaev] ” / “ Slide | ZSM Arct. | 2021 - 319 ♁ | A. Volynkin ” (MWM / ZSM). Paratype. MYANMAR: 1 male, Chin State, Mount Victoria, 2200 – 2500 m, montane pine forest, mid – late October 2012, Dr. V. Kravchenko & team, gen. prep. No.: AV 6745 (prepared by Volynkin) (GMF-B).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA702E1FA1CFCAD6942.taxon	diagnosis	Diagnosis. The forewing length is 15.0 – 17.0 mm in males. The male genital capsule of L. nansabon is reminiscent of L. diehli but distinguished by the narrower uncus, the shorter vinculum, the somewhat shorter (in proportion to the valva length) and almost straight distal saccular process which is curved in the congener. The phallus of the new species is narrower and somewhat shorter than in L. diehli (in proportion to the valva length). The vesica of L. nansabon differs from L. diehli in the presence of a single and broadly conical medial dorsal diverticulum (L. diehli has two narrower and utricular medial dorsal diverticula), and the distal diverticulum directed ventrally perpendicular the proximal axis of the vesica whereas it is directed distally along the axis of the vesica in the congener. The detailed comparison with another similar species, L. witti is provided below in the diagnosis of the latter species. The female is unknown.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA702E1FA1CFCAD6942.taxon	distribution	Distribution. The new species is currently known from northern Myanmar (Kachin and Chin States).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB77FFA702E1FA1CFCAD6942.taxon	etymology	Etymology. The specific epithet is homonymic of the type locality of the new species, Nan Sa Bon village. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA702E1FED4FD4C6E2A.taxon	description	(Figs 14 – 21, 50 – 54, 68)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA702E1FED4FD4C6E2A.taxon	materials_examined	Type material. Holotype (Figs 14, 50): male, “ SW Thailand | Ranong-prov. Ranong, 380 m | 10 ° 01 ’ 32 ’’ N, 98 ° 40 ’ 13 ’’ E | 3. – 4. XII. 2005 leg. K. Cerný [recte: Černý] ” / “ Slide | AV 7328 ♁ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. THAILAND: 1 male, 1 female, the same data as holotype, gen. prep. No.: AV 7329 (prepared by Volynkin) (CKC); 1 female, Phang Nga Prov., Kuraburi distr., 180 m, Ban Tiem, 7. iii. 2005, Khao Pra Me, Karel Černý leg., gen. prep. No.: AV 7330 (prepared by Volynkin) (CKC); 1 male, Phangna, Kuraburi, Num Sai, 6. iii. 2005, Ao Khoei Beach, 5 m, leg. K. Sukhumalind (CKC); 2 males, 1 female, Changwat Nan, 5 km N of Bo Luang, 1000 m, 12. xi. 1999, Márton Hreblay leg., gen. prep. Nos.: ZSM Arct. 2021 - 313 (male), ZSM Arct. 2021 - 314 (female) (prepared by Volynkin) (MWM / ZSM); LAOS: 2 males, 2 females, Khamouane [Khammouane Prov.], Phon Hin Boun, 24 – 25. v. 2013, [4] 50 m, 18 ° 10.435 ’ N 104 ° 29.042 ’ E, Thomas Ihle leg., gen. prep. Nos.: AV 7332 (male), AV 7334, AV 7335 (females) (prepared by Volynkin) (CKC); CHINA: 6 males, 5 females, W Yunnan Prov., Xishuangbanna Dai auton. pref., Puwen, 30 km SSW Simao, 900 m, 22 ° 30 ’ N 100 ° 02 ’ E, 16. iii. – 10. iv. 2000, Brechlin’s loc. coll. leg., gen. prep. Nos.: ZSM Arct. 2021 - 315, ZSM Arct. 2021 - 316 (males), ZSM Arct. 2021 - 317 (female) (prepared by Volynkin) (MWM / ZSM); INDIA: 1 male, 4 females, W. B. [West Bengal], Siliguri, above Sukna, 300 m, 2. vii. 1990, W. Thomas leg., gen. prep. Nos.: ZSM Arct. 2021 - 003 (male), ZSM Arct. 2021 - 318 (female) (prepared by Volynkin) (MWM / ZSM); 1 male, W. B. [West Bengal], Darjeeling above Sukna, ca. 500 m, 7 – 8. vii. 1989, Dr W. Thomas leg., gen. prep. No.: ZSM Arct. 2021 - 339 (prepared by Volynkin) (MWM / ZSM). Note. The male genital capsule of the species is somewhat variable in the length of the distal saccular process, the width of the dorsal part of the valva, and the number of cornuti in the distal diverticulum (from 1 to 4).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA702E1FED4FD4C6E2A.taxon	diagnosis	Diagnosis. The forewing length is 14.0 – 15.5 mm in males and 17.5 – 19.0 mm in females. The male genital capsule of L. witti is most similar to L. nansabon but can be distinguished by the proximally broader uncus and the markedly shorter distal saccular process. The phalli of the two species are alike. Compared to L. nansabon, the vesica of L. witti is downcurved (it is directed distally in the congener), has an additional distal diverticulum armed with a terminal cornutus, and its medial dorsal diverticulum has a terminal cornutus apically. As the female of L. nansabon is unknown, the female genitalia of the new species were compared with the most similar L. leucanioides instead, from which L. witti differs in the presence of an antevaginal fold of the 7 th abdominal sternite, the somewhat shorter ductus bursae, and the markedly longer and broader gelatinous rugose area of the posterior section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA702E1FED4FD4C6E2A.taxon	distribution	Distribution. The new species is widespread and known from Northeast India, Southwest China (Yunnan Province), Thailand (Nan, Ranong and Phang Nga Provinces), and Laos (Khammouane Province).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA702E1FED4FD4C6E2A.taxon	etymology	Etymology. The species is dedicated to the memory of the late Dr Thomas J. Witt (Munich, Germany), a prominent lepidopterist and founder of the Museum Witt Munich who has built one of the largest collections of Heterocera in the world. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA402E1F97CFA946C07.taxon	description	(Figs 22, 23, 55, 70)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA402E1F97CFA946C07.taxon	materials_examined	Type material. Holotype (Figs 22, 55): male, [the Philippines] “ Philippinen / Nord Palawan | S. Vicente, 20 km NEE Roxas | 10 [°] 21 ’ N / 119 [°] 10 ’ E | Mittelgebirgsurwald [mid-mountain rainforest], 400 m | 12. I. – 17. I. 1988, leg. | Černý & Schintlmeister ” / “ coll. Dr. W. Thomas | Mus. Witt München ” / “ Beschtimmungsliste | Lfd. Nr. | 206 | Leucanioides | tigioides | det. K. Cerny 1989 ” / “ Slide | ZSM Arct. | 2021 - 002 ♁ | A. Volynkin ” (MWM / ZSM). Paratypes. THE PHILIPPINES: 3 males, 2 females, the same data as holotype (CKC); 9 males, 4 females, C. Palawan, 5 km E of Napsan, Primary forest clearing, 09 ° 42.256 ’ N 118 ° 31.034 ’ E, 1 – 2. xii. 2007, J. H. Lourens leg., gen. prep. No.: ZSM Arct. 2021 - 322 (female) (prepared by Volynkin) (MWM / ZSM); 1 male, C. Palawan, E of Napsan, prim. for. edge, Salakot Falls, 250 m, 09 ° 50 ’ N 118 ° 37 ’ E, 12. iii. 2006, J. H. Lourens leg. (MWM / ZSM); 1 male, N Palawan, SE Port Barton, Matalangao, 80 m, 10 ° 18 ’ N 119 ° 11 ’ E, 6. iii. 2006, J. H. Lourens leg. (MWM / ZSM); 3 males, 3 females, South Palawan, Mt. Gantung, 200 m, 9 ° 01 ’ N 117 ° 57 ’ E, cypress forest / secondary vegetation, 19 – 21. i. 1988, Černý & Schintlmeister leg. (CKC); 1 male, E Palawan, 8 km W of San Vicente, costal hill forest, 138 m, 10 ° 31.639 ’ N 119 ° 20.096 ’ E, 19. xii. 2007, J. H. Lourens leg. (CKC); 1 male, C Palawan, E of Napsan, prim. for. edge, Salakot Falls, 250 m, 9 ° 50 ’ N 118 ° 37 ’ E, 12. iii. 2006, J. H. Lourens leg. (CKC).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA402E1F97CFA946C07.taxon	diagnosis	Diagnosis. The forewing length is 14.5 – 16.0 mm in males and 15.5 – 17.5 mm in females. The male genital capsule of L. leptivalva is reminiscent of L. diehli, L. paraliaka, L. nansabon and L. witti due to the strongly elongate and narrow dorsal part of the valva but can easily be distinguished from all the aforementioned species by the structure of the uncus, which is proximally swollen but distally narrow (it is evenly swollen in the similar species), the rectangular and heavily sclerotised posterior section of the juxta with a semiglobular and spinulose apical process, the more weakly sclerotised and proximally narrower valva, and the considerably shorter and narrower sacculus with a small and weakly sclerotised, willow leaf-shaped distal saccular process. Compared to the similar species, the phallus of the new species is shorter (in proportion to the tegumen-vinculum complex length), has a medial ventral protrusion, and is distally dilated. The vesica configuration of L. leptivalva is most similar to L. nansabon, from which the new species differs in the reduced proximal section, the proximal cluster of spines situated laterally, the larger dorsal medial diverticulum, the distally tapered distal diverticulum bearing a smaller terminal cornutus, and the presence of the broad, second distal diverticulum. In the female genitalia, L. leptivalva differs from the similar species in the broader and longer antevaginal fold of the 7 th abdominal sternite forming a deep caliciform subostial depression, the markedly broader ductus bursae, and the considerably larger appendix bursae directed anteriorly (it is directed laterally or latero-anteriorly in the congeners).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA402E1F97CFA946C07.taxon	distribution	Distribution. The new species is endemic to the island of Palawan (the Philippines).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB76FFA402E1F97CFA946C07.taxon	etymology	Etymology. The specific epithet is derived from the Greek ‘ λεπτός’ meaning ‘ thin’ and refers to the narrow and weakly sclerotised valva of the new species. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB75FFA502E1FB10FAE2693A.taxon	description	(Figs 24, 25, 56)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB75FFA502E1FB10FAE2693A.taxon	materials_examined	Type material. Holotype (Figs 24, 56): male, [Indonesia] “ Sumatra Sitahoan | 2 ° 39 ’ N, 99 ° 00 ’ E GPS | 1500 m, 26.4.1997 Malicky [leg.] ” / “ Slide | AV 7331 ♁ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. INDONESIA, SUMATRA: 1 male, Siantar-Prapat road, Holzweg 3, Lf 250 w Hg, ca. 3 – 400 m NN, sec. forest, W. Nässig & [E. W.] Diehl leg., 31. i. / 1. + 9. ii. 1984 (MWM / ZSM); 1 male, Parapat (Toba Lake), 1000 m, 1. viii. 1972, light, Erber leg. / coll. Dr. W. Thomas, Mus. Witt München, gen. prep. No.; ZSM Arct. 2021 - 336 (prepared by Volynkin) (MWM / ZSM); 1 male, Utara, Simalungun / Aek Nauli, 2 ° 44 ’ N 98 ° 58 ’ E, 1200 m, 18. vi. 1986, Plössl & Dr Tarmann leg. (CBP); 2 males, Utara, Simalungun / Tiga Dolok, 2 ° 46 ’ N 98 ° 59 ’ E, 1040 m, 04. iii. 1997, Plössl & Dr Tarmann leg. (CBP); 1 male, Utara, Simalungun / Tiga Dolok, 2 ° 47 ’ 98 ° 59 ’ E, 1020 m, 01. vi. 1986, Plössl & Dr Tarmann leg. (CBP); 1 male, Aceh, Singah Mata, 4 ° 09 ’ N 97 ° 30 ’ E, 1720 m, 25. i. 1989, Dr Tarmann & Plössl leg. (CBP); 1 male, Dolok Merangir, 2. v. – 21. v. 1983, Dr Diehl leg. (CKC); 2 males, Popat [Prapat], 1250 m, vii. 1972, Diehl leg. (CKC); 6 males, 15 km NE Prapat, HW 4, 1150 m, 3 – 6. i. 1994, Karel Černý leg. (CKC). Note. This and the next species have the male genital capsule structure considerably different from the other species in the genus due to the reduced dorsal section of the valva having a massive claw-shaped apical spur, and the larger sacculus. However, the presence of the aforementioned apical spur along with the intervalval commissure fused with the heavily sclerotised juxta, as well as the phallus vesica structure confirm their belonging to Leucanithosia.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB75FFA502E1FB10FAE2693A.taxon	diagnosis	Diagnosis. The forewing length is 16.5 – 17.5 mm in males. The male genital capsule of the new species is clearly different from the other congeners in the short and broad dorsal part of the valva having a robust, claw-shaped apical spur, and the broad sacculus with a broad, hook-like curved distal process. Additionally, the uncus of L. inusitata is markedly broader than in the similar species. The phallus and the vesica structures of L. inusitata are most similar to L. lampunga and the detailed comparison is provided below in the diagnosis of the latter species. The female is unknown.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB75FFA502E1FB10FAE2693A.taxon	distribution	Distribution. The new species is known from northern Sumatra Island (Indonesia).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB75FFA502E1FB10FAE2693A.taxon	etymology	Etymology. The specific epithet is derived from the Latin ‘ inusitatus’ meaning ‘ unusual’ and refers to the peculiar configuration of the male genital capsule. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFA502E1FE0CFB476C9A.taxon	description	(Figs 26, 27, 57, 71)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFA502E1FE0CFB476C9A.taxon	materials_examined	Type material. Holotype (Figs 26, 57): male, “ Indonesia, Sumatra, | Bandar Lampung, | Bukit Rigis Mts., | Gunung Terang, 1022 m, | 5 ° 3 ’ 43 ’’ S, 104 ° 27 ’ 6 ’’ E, 13. – | 15. XI. 2003 leg. Černý ” / “ Slide | AV 7322 ♁ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. INDONESIA, SUMATRA: 5 males, 3 females, the same data as holotype, gen. prep. No.: AV 7323 (female) (CKC); 1 male, South Sumatra, 900 m, Lake Ranau, Tebah Hemi, 4 ° 56 ’ 03 ’’ S 103 ° 57 ’ 15 ’’ E, 18. xi. 2003, Černý leg. (CKC).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFA502E1FE0CFB476C9A.taxon	diagnosis	Diagnosis. The forewing length is 16.0 – 16.5 mm in males and 19.5 – 20.0 mm in females. The male genital capsule of the new species is vaguely reminiscent of L. inusitata but can easily be distinguished by the longer and narrower uncus, the narrower and hook-like curved dorsal part of the valva gradually terminating with a thorn-shaped apical spur (it is claw-shaped in the congener), and the thin, S-like curved and apically club-shaped distal saccular process. Additionally, compared to L. inusitata, the arms of the tegumen of L. lampunga are posteriorly broader, the vinculum is narrower, and the posterior section of the juxta is more convex and serrulate (it is smooth in the congener). The phalli of the two species are similar but that of the new species has narrower distal plate covered with larger denticles. In the vesica structure, L. lampunga is distinguished from L. inusitata by the narrower diverticula bearing smaller cornuti.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFA502E1FE0CFB476C9A.taxon	distribution	Distribution. The new species is currently known from southern Sumatra Island (Indonesia).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFA502E1FE0CFB476C9A.taxon	etymology	Etymology. The specific epithet is derived from the Indonesian province of Lampung, where the holotype and most of the paratypes were collected. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFAA02E1FAACFA9A6BB3.taxon	type_taxon	Type species: Philippinithosia lourensi sp. n., by present designation.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFAA02E1FAACFA9A6BB3.taxon	diagnosis	Diagnosis. Species of the new genus (Figs 28 – 35) are externally very similar to Leucanithosia (Figs 1 – 27) but in males, the androconial tuft on the foreleg consists of ochreous yellow and shorter scales, which are elongate and grey in Leucanithosia. The male genital capsule of Philippinithosia (Figs 58 – 62) is characterised by the distally dilated and swollen dorsal part of the valva and the distal saccular process, which both are densely covered with minute setae, the feature characteristic of the genus. Additionally, compared to Leucanithosia (Figs 40 – 57), Philippinithosia lacks the intervalval commissure, and its juxta is weakly sclerotised whereas in Leucanithosia, the valvae are proximally connected by a commissure, and the posterior section of the juxta is heavily sclerotised and laterally fused with transtillae and ventrally connected to the intervalval commissure. The uncus of the new genus is proximally narrow and distally dilated and swollen whereas it is evenly swollen and basally dilated in Leucanithosia. The phallus of Philippinithosia is broader than in Leucanithosia and lacks the distal serrulate plate. The vesicae of the two genera display no fundamental differences. The female genitalia ground plan of Philippinithosia (Figs 72 – 74) is similar to Leucanithosia (Figs. 65 – 71) but in the current new genus, the appendix bursae is weakly expressed whereas the bursa copulatrix of Leucanithosia is clearly subdivided into a corpus bursae and a sack-like appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFAA02E1FAACFA9A6BB3.taxon	description	Description. Adults. Sexual dimorphism limited: female somewhat larger than male and with slightly broader forewing. Antenna dark grey, ciliate in both sexes. Head creamy or ochreous creamy. Thorax creamy. Pro- and metathorax with black spot medio-dorsally. Patagia ochreous creamy. Male foreleg with androconial tuft consisting of ochreous yellow scales. Tegula creamy with black spot medially. Forewing elongate and narrow, with almost parallel margins; costal margin straight proximally and medially and downcurved distally; anal margin convex. Forewing ground colour creamy or ochreous creamy. Pattern represented by dark grey or brown suffusion along veins, reduced in certain species. Forewing cilia creamy or ochreous creamy. Hindwing unicolorous creamy or ochreous creamy, paler than forewing. Abdomen creamy with intense admixture of ochreous scales laterally and distally. Male genitalia. Uncus proximally narow and distally dilated and swollen, with tiny claw-shaped tip. Tegumen weakly sclerotised, with posteriorly dilated arms. Tuba analis membranous, as broad as half of tegumen length. Vinculum shorter than tegumen, with narrowly U-shaped saccus. Valva lobular, somewhat dilated medially. Valva apex dilated, swollen and densely covered with short setae. Sacculus narrow and elongate, distal saccular process dilated and densely setose apically. Juxta moderately sclerotised, trapezoidal with medial anterior incision. Anellus membranous. Phallus short but broad, cylindrical. Proximal section of vesica broadly tubular. Distal section of vesica narrower than proximal one, tubular, terminating with one or two tubular diverticula bearing one or two short but basally broad cornuti apically. Vesica ejaculatorius originates medially, with stick-shaped basal plate. Female genitalia. Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses elongate and thin, apophysis anterioris shorter or equal in length to apophysis posterioris. Postvaginal area membranous. Ductus bursae elongate, tubular, rugose. Corpus bursae broad sack-like with two small and rounded signa, rugose posteriorly, in certain species with short and semiglobular postero-lateral rugose appendix at junction with ductus bursae. Appendix bursae not separated from corpus bursae, represented as broad lateral protrusion with ductus seminalis originating laterally.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFAA02E1FAACFA9A6BB3.taxon	distribution	Distribution. The genus is endemic to the Philippines (absent from Palawan Island).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB74FFAA02E1FAACFA9A6BB3.taxon	etymology	Etymology. The new genus name is an aggregate of the toponym ‘ The Philippines’ and the genus group name Lithosia, and refers to the Philippine Archipelago to which the new genus is endemic. Gender is feminine.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7BFFAB02E1FB48FEF868A6.taxon	description	(Figs 28 – 31, 58, 59, 72)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7BFFAB02E1FB48FEF868A6.taxon	materials_examined	Type material. Holotype (Figs 28, 58): male, “ the Philippines E. Luzon | Sierra Madre 180 m | 5 km W of Gen Nakar | 14 ° 46.703 ’ N 121 ° 36.799 ’ E | prim forest edge | 12.02.2007 K Knoblich | & JH Lourens leg. ” / “ Museum Witt | München ” / “ Slide | ZSM Arct. | 2021 - 085 ♁ | A. Volynkin ” (MWM / ZSM). Paratypes. THE PHILIPPINES: 5 males, 12 females, E Luzon, N. Ec. Bongabon Bgy Laby, Sierra Madre Mingan Mts, 15 ° 38 ’ N 121 ° 15 ’ E, 950 m, 7 – 9. ix. 2005, J. H. Lourens leg., gen. prep. No.: ZSM Arct. 2021 - 323 (female) (MWM / ZSM); 1 male, 1 female, Mindoro, Mt. Halcon, 1000 m, 20 – 25. vi. 2002, local people leg. (MWM / ZSM); 2 males, 820 m, Negros (western Negros Prov.), Mt. Kanlaon, W route via Mambucal, 15. vii. 1996, primary forest, R. Brechlin leg., gen. prep. Nos.: ZSM Arct. 2021 - 004, ZSM Arct. 2021 - 079 (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7BFFAB02E1FB48FEF868A6.taxon	diagnosis	Diagnosis. The forewing length is 14.5 – 16.0 mm in males and 18.0 – 18.5 mm in females. The male genital capsule of the new species is most similar to P. settelei but differs in the longer and distally broader uncus, the somewhat longer vinculum, the narrower setose apex of the valva, and the broader setose apex of the distal saccular process. The phallus of P. lourensi is broader and somewhat narrower than in P. settelei. Compared to P. settelei, the vesica of the new species is longer and proximally broader, and has longer and broader diverticula bearing larger terminal cornuti. The female genitalia of the two species are similar but in P. lourensi, the ductus bursae is longer, the rugose posterior area of the corpus bursae is shorter, and the signa are somewhat smaller than the corresponding structures of P. settelei.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7BFFAB02E1FB48FEF868A6.taxon	distribution	Distribution. The new species is known from the islands of Luzon, Negros and Mindoro (the Philippines).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7BFFAB02E1FB48FEF868A6.taxon	etymology	Etymology. The new species is named after Dr Johannes H. Lourens, a dedicated researcher of the Lepidoptera fauna of the Philippines and collector of the part of the type series including the holotype. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFAB02E1FEF0FC186BFA.taxon	description	(Figs 32, 33, 60, 73)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFAB02E1FEF0FC186BFA.taxon	materials_examined	Type material. Holotype (Figs 32, 60): male, “ the Philippines SE Mindanao | Davao Oriental Aliwagwag | Primary forest 90 m. | 07 ° 43.667 ’ N 126 ° 17.304 ’ E | 30 Apr. + 1 May 2008 | JH Lourens leg. ” / “ Museum Witt | München ” / “ Slide | ZSM Arct. | 2021 - 084 ♁ | A. Volynkin ” (MWM / ZSM). Paratypes. THE PHILIPPINES: 2 males, 1 female, the same data as holotype, gen. prep. No.: ZSM Arct. 2021 - 321 (female) (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFAB02E1FEF0FC186BFA.taxon	diagnosis	Diagnosis. The forewing length is 14.5 – 15.0 mm in males and 17.0 mm in the female. The genitalia of both sexes of the new species are most similar to P. lourensi and the detailed comparison is provided above in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFAB02E1FEF0FC186BFA.taxon	distribution	Distribution. The new species is known from the island of Mindanao (the Philippines).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFAB02E1FEF0FC186BFA.taxon	etymology	Etymology. The new species is named after Prof. Josef Settele (Helmholtz Centre for Environmental Research – UFZ, Halle, Germany), who was very helpful in organizing the second author’s researches in the Philippines and who is actively involved in science-policy processes for the conservation of Lepidoptera and biodiversity in the Philippines as well as globally. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFA802E1FC4CFB4669D6.taxon	description	http: // zoobank. org / urn: lsid: zoobank. org: act: 065083 E 8 - C 4 EB- 4 D 30 - A 9 F 6 - 8 BC 929 BDFE 1 A (Figs 34, 35, 61, 62, 74)	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFA802E1FC4CFB4669D6.taxon	materials_examined	Type material. Holotype (Figs 34, 61): male, “ the Philippines Mindanao | E. Surigao Lianga Hanayan | 13 km W. of Diatagon 350 m | 29 Nov. – 1 Dec. 2005 | 8 ° 42 ’ N 126 ° 05 ’ E | JH Lourens / A. Schintlmeister ” / “ Museum Witt | München ” / “ Slide | ZSM Arct. | 2021 - 082 ♁ | A. Volynkin ” (MWM / ZSM). Paratypes. THE PHILIPPINES: 1 male, 1 female, the same data as holotype, gen. prep. No.: ZSM Arct. 2021 - 324 (female) (prepared by Volynkin) (MWM / ZSM); 5 males, 1 female, Mindanao, Misames Norte Prov., 22 km NE of Claveria, Brg. Mat I, 1050 m, 20. iii. 2009, 8 ° 39.874 ’ N 124 ° 59.516 ’ E, K. Černý & J. Lourens leg. (CKC); 5 males, Mindanao, Bukidnon Prov., 60 km S Talakag, Dominorog, 1271 m, 23 – 24. iii. 2009, 7 ° 56.144 ’ N 124 ° 38.5 ’ E, K. Černý & J. Lourens leg. (CKC); 1 male, 1 female, Mindanao, Surigao del Sur, 4 km E Homburquer, Gata, 170 m, 17 – 18. iii. 2009, 8 ° 42.944 ’ N 126 ° 9.402 ’ E, K. Černý & J. Lourens leg. (CKC); 1 male, Mindanao, Mt. Apo western side, secondary forest, 28 – 30. vii. 1993, 6 ° 57 ’ N 125 ° 16 ’ E, Schintlmeister & Sinyaev leg. (CKC); 3 males, 6 females, Mindanao, Bukidnon [Prov.], 40 km NW Maramag, Dalongdong, 800 m, Talakag, 7 ° 53 ’ N 123 ° 54 ’ E, forest edge, 31. xii. 1991 – 2. i. 1992, Karel Černý leg. (CKC); 5 males, 5 females, the same locality as previous but 1 – 3. x. 1988, Černý & Schintlmeister leg. (CKC); 6 males, 1 female, Mindanao, Bukidnon [Prov.], 45 km NW Maramag, Mt. Binansilang, 1200 m, 7 ° 55 ’ N 124 ° 40 ’ E, mountain forest, 2. x. 1988, Černý & Schintlmeister leg. (CKC); 1 male, 1 female, SE Leyte, S of Libertad Brgy Imelda, Katipunan, 300 m, 28. xi. 2005, 10 ° 43 ’ N 125 ° 04 ’ [E], J. H. Lourens & A. Schintlmeister leg., gen. prep. No.: ZSM Arct. 2021 - 083 (male) (prepared by Volynkin) (MWM / ZSM); 2 females, C Samar, 4 km SE Bachakay, 287 m, 26. iii. 2009, 11 ° 48.025 ’ N 125 ° 14.61 ’ E, K. Černý & J. Lourens leg. (CKC).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFA802E1FC4CFB4669D6.taxon	diagnosis	Diagnosis. The forewing length is 16.0 mm in males and 18.5 – 19.0 mm in females. Philippinithosia reducta externally differs from its congeners in the unicolorous forewing lacking the dark suffusion on veins. The male genital capsule of the new species is most similar to P. settelei but can easily be distinguished by the elongate, footshaped setose apex of the valva (it is globular in the congener), the apically broader distal saccular process covered with larger spinules, and the longer and distally broader uncus. The phallus of P. reducta is somewhat longer than in P. settelei (in proportion to the tegumen-vinculum complex length). In the vesica structure, the new species is distinct from P. settelei by the longer and narrower proximal section, and the presence of a single distal diverticulum bearing a larger terminal cornutus. The female genitalia of P. reducta differ from P. settelei in the longer apophyses anteriores, and the presence of a semiglobular rugose diverticulum of the corpus bursae at the junction with the ductus bursae.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFA802E1FC4CFB4669D6.taxon	distribution	Distribution. The new species is currently known from the islands of Mindanao, Leyte and Samar (the Philippines).	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
78771043CB7AFFA802E1FC4CFB4669D6.taxon	etymology	Etymology. The specific epithet is derived from the Latin ‘ reducta’ meaning ‘ reduced’ and refers to the reduced dark suffusion on the forewing veins. The name is a noun in the nominative singular in apposition.	en	Volynkin, Anton V., Černý, Karel (2023): On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 5352 (2): 151-176, DOI: 10.11646/zootaxa.5352.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5352.2.1
