identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039987ADFFC6E02AD398FC51FD9FFEBC.text	039987ADFFC6E02AD398FC51FD9FFEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Achagua Rindge 1983	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Achagua Rindge, 1983</p>
            <p> Type Species:  Achagua obsoleta Rindge, 1983 ; original designation. </p>
            <p> Re-diagnosis. Both Rindge (1983) and Pitkin (2002) provided diagnostic comments for  Achagua , but with the discovery of three undescribed species, it is necessary to revisit these diagnoses. The mostly bipectinate antennae of male  Achagua , with the apical third to quarter being simple filiform, is a shared characteristic among related nacophorine genera, e.g., Cargolia and Gabriola, and thus, does not disambiguate closely related taxa. In males, the two processes of the uncus in conjunction with the subtriangular anellar processes appear to be the most reliable diagnostic features (Figs. 5a, 6a, 7a, 8a). Pitkin (2002) listed the strongly spatulate (or capitate) dorsal process of the uncus as an apomorphy for the genus, but two of the new species described herein possess a non-spatulate dorsal process (Figs. 7a, 8a). In females, the large, elongate, apically pointed, denticulate signum may be diagnostic, but only a single preparation of  A. obsoleta (Fig. 9) and two preparations of  A. cooperae (Fig. 10) were available for study. Superficially, the pearly-white ground color, large size, brown to black wing margins, and lack of welldefined antemedial and postmedial lines seem to also define  Achagua . Further, the multi-locus phylogenetic results of Murillo-Ramos et al. (2019) and Brehm et al. (2019) also support the recognition of  Achagua as distinct from related genera. </p>
            <p> Distribution (Fig. 11).  Achagua inhabit the tropical and subtropical moist broadleaf forests of Central and South America from the Sierra de los Tuxtlas of Veracruz, Mexico (  A. cooperae ), east to the Guiana Highlands (  A. velata ), and southward through the eastern Andes of Colombia, Ecuador, and Peru to at least the Bolivian Yungas (  A. magna ). </p>
            <p>Biology. Thus far, no life history information is known for the genus. Adults are infrequently collected at light.</p>
            <p> Remarks. In  Achagua , the uncus is characterized by two distinct processes. However, there is some disagreement regarding the nomenclature of these processes. Rindge (1983) referred to the much larger dorsal process as the pseudouncus, but it is unclear how he arrived at this conclusion. Alternatively, I adopt the terminology used by Pitkin (2002) and refer to these processes based on their orientation, specifically as the dorsal or ventral processes of the uncus. </p>
            <p> Campos (2001) described the genus “  Achagua ” for a group of freshwater crabs. As the name was already preoccupied by  Achagua Rindge (the subject of this work), “  Achagua ” Campos was rendered a junior homonym. Campos and Magalh„es (2004) synonymized “  Achagua ” Campos with  Eudaniela Pretzmann , such that this junior homonym has not been in contemporary use. </p>
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	https://treatment.plazi.org/id/039987ADFFC6E02AD398FC51FD9FFEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matson, Tanner A.	Matson, Tanner A. (2023): The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae). Zootaxa 5352 (4): 565-576, DOI: 10.11646/zootaxa.5352.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5352.4.7
039987ADFFC5E02AD398FEC9FDE3FC20.text	039987ADFFC5E02AD398FEC9FDE3FC20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Achagua obsoleta Rindge 1983	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Achagua obsoleta Rindge, 1983</p>
            <p>(Figs. 1, 5, 9, 11)</p>
            <p> Distribution (Fig. 11).  Achagua obsoleta is known from the Magdalena Valley montane forests of Colombia. </p>
            <p>Biology. Life history information is wanting. Adults are known to fly in August.</p>
            <p> Material Examined.   COLOMBIA • ♁; holotype; Cundinamarca, 3 km. north of Alban,  Finca San Pablo ; elev. 1800 m; 1–12 Aug. 1967; P. &amp; B. Wygodzinsky leg.; Genitalia: FHR 19129; AMNH _ IZC 00353008, AMNH _ IZC 00353009; AMNH  •  3♁; paratypes; same data as holotype; Genitalia: TAM-2023-270; AMNH •   ♀; Ob. Rio Negro,  Ost Colombia ; elev. 800 m; Fassl. leg.;  Geometridae genitalia slide No. 20288 ♀; BMNH (E)  #   275238  Digitally scanned; Joicey Bequest. Brit. Mus. 1934-120; NHMUK010293435; NHMUK  . </p>
            <p> Remarks. Only male genitalia were described and illustrated in the original description of  A. obsoleta , however, genitalia of both sexes were described and illustrated in Pitkin (2002). Genitalia of both sexes are also illustrated in this work (Figs. 5, 9). </p>
            <p> While not readily seen in Figure 5,  A. obsoleta , like  A. magna , also has a distinct postmedial digitate protuberance along the costal margin of the valve. </p>
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	https://treatment.plazi.org/id/039987ADFFC5E02AD398FEC9FDE3FC20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matson, Tanner A.	Matson, Tanner A. (2023): The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae). Zootaxa 5352 (4): 565-576, DOI: 10.11646/zootaxa.5352.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5352.4.7
039987ADFFC5E02ED398FC75FE75FB5B.text	039987ADFFC5E02ED398FC75FE75FB5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Achagua magna Matson 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Achagua magna Matson ,  n. sp.</p>
            <p>(Figs. 2, 6, 11, 12)</p>
            <p>LSID: EBF2D227-50CC-420C-83F7-2939D2F9C3AD</p>
            <p> Diagnosis.  Achagua magna is most similar to  A. obsoleta . While  A. magna inhabits the eastern slopes of the central Andes in Ecuador, Peru, and Bolivia,  A. obsoleta is thus far only known from the northern Andes in Colombia.  Achagua magna (forewing, 21–22 mm) is noticeably larger than  A. obsoleta (forewing, 19–20 mm), and though subtle, the pearly-white ground color of the forewing upperside appears to be marked more strongly with intermittent black scaling over the veins; especially in the antemedial and postmedial areas.  Achagua magna also appears to bear a more strongly marked, brown to black, terminal area of the hindwing. In males, the lateral margin of the large, subtriangular, anellar processes appears to be less deeply concave in  A. magna than in  A. obsoleta . </p>
            <p> Achagua magna can be distinguished from both  A. cooperae and  A. velata by the absence of a conspicuous antemedial black spot along the forewing inner margin. Additionally, the male genitalia of  A. magna (Fig. 6) bear a spatulate dorsal process of the uncus, a costa with a postmedial digitate protuberance, and a denticulate patch of cornuti on the vesica. In  A. cooperae (Fig. 7) and  A. velata (Fig. 8), the dorsal process of the uncus is not spatulate or obviously swollen, the costa lacks a postmedial digitate protuberance, and the vesica lacks cornuti. There are other, more subtle differences that can be observed in Figures 6–8, which further serve to differentiate  A. magna from  A. cooperae and  A. velata . </p>
            <p>Description. MALE. Forewing length, 21–22 mm (n = 8).</p>
            <p>Head. Antenna mostly bipectinate, but with gradually diminishing rami that are absent in distal quarter of antenna; scales above fuscous, rami ochreous. Vertex mostly white with central gray spot; frons thinly scaled, white. Labial palpus decumbent, 1.5x diameter of eye, and with mixture of white and gray scales on outer surface and white on inner surface. Chaetosemata in transverse row; cephalic collar mostly white with few gray scales.</p>
            <p>Thorax. Patagium, tegula, and mesothorax admixture of gray and white scales. Legs mostly white and mottled with gray; epiphysis well-developed; hind tibia with large hair pencil; tibial spur formula 0–2–4.</p>
            <p>Forewing. Pearly-white; widely scattered with inconspicuous light gray scales and with intermittent black scaling over parts of veins. Basal, costal, and terminal area of outer margin broadly maculated with brown scales; basal area, apical area, and tornus, more uniformly brown. Underside more uniform pearly-white and without brown basal scaling. Apical area with more diffuse brown scaling than upperside.</p>
            <p>Hindwing. Pearly-white except for brown and black terminal band. Underside patterned similar to upperside, but darkened area around outer margin more diffuse and tornal area much whiter.</p>
            <p>Abdomen. First segment white with pair of black elongate dorsal spots, second segment black with narrowly triangular white median patch, remainder of abdomen with mixture of white, gray, black, and brown scales; white below. Third sternite of male abdomen with comb of setae.</p>
            <p>Genitalia (Fig. 6). Uncus abruptly widened at base; dorsal process thin through middle and apically spatulate; ventral process thumb-like. Base of gnathos subquadrangular, with upcurved, heavily sclerotized, pointed apical projection; apex of projection lightly dentate. Valve large and elongate with heavily sclerotized costa bearing postmedial digitate protuberance. Anellar processes large and subtriangular with elongate, slender posterior projection; lateral margin only slightly concave. Juxta with elongate, medial cylindrical process. Vesica with large denticulate basal patch and weakly sclerotized medial patch.</p>
            <p>FEMALE. Unknown.</p>
            <p>Type Material.</p>
            <p>Holotype</p>
            <p> PERU • ♁; Cusco,  Quincemil ; elev. 2400 ft; Aug. 1962; L.E. Pena leg.; AMNH _ IZC 00353021; AMNH. </p>
            <p>Paratypes (8 ♁)</p>
            <p>PERU • 5♁; same collection data as holotype; Genitalia: FHR no. 12173, TAM-2023-255, TAM-2023-268; AMNH _ IZC 00353022– AMNH _IZC 00353025, USNMENT01771249; AMNH, USNM • ♁; Cuzco, Cosnipata; (-12.9018°, -71.4117°); elev. 724 m; 28 Aug. 2016; Gunnar Brehm leg.; BOLD Process ID: PEMOA404-16; JPM • ECUADOR • ♁; Zamora Chinchipe; (-3.97388°, -79.084°); elev. 1925 m; 29 Oct. 2002; Nadine Hilt &amp; Claudia Ramenda leg.; BOLD Process ID: NGEOE359-12; JPM • BOLIVIA • ♁; N.P. Carrasco; elev. 900m; (17°06’44”S 65°33’55”W); 05 Oct. 2010; Aare Lindt leg.; Genetic Voucher: TTBO002; TUZ.</p>
            <p> Distribution (Fig. 11).  Achagua magna inhabits the eastern slopes of the central Andes from the Eastern Cordillera real montane forests of Ecuador south through the Peruvian and Bolivian Yungas. </p>
            <p>Biology. Life history information is wanting. Adult records are from August and October.</p>
            <p>Etymology. The specific epithet magna is from the Latin “magnus,” meaning “large,” as this species is the largest known member of the genus.</p>
            <p> Molecular characterization. This species is represented in BOLD by the BIN: BOLD:ABW8871 (n =3, Peru, Ecuador, Bolivia). Specimens from Bolivia and Peru differ from the only sequenced Ecuadorian individual by about 1.2%. The distance to the nearest neighbor,  Achagua cooperae is about 5.9%, however, the presumable sister species of  A. magna ,  A. obsoleta , has not been sequenced. </p>
            <p> Remarks. Multi-locus molecular data for this species were used in the phylogenetic studies of Murillo-Ramos et al. (2019) and Brehm et al. (2019). However, the sequenced individual of  Achagua was misidentified as  A. obsoleta in these studies. This is unsurprising considering the similarity between  A. magna and  A. obsoleta , and the fact that  A. obsoleta was the only described  Achagua at that time. </p>
            <p> Achagua magna is the name given to the undescribed species mentioned in Rindge (1983). While Rindge initially mentioned his series of  A. magna as being from Ecuador in the distribution section of his generic treatment, he later referred to it as being from Peru in the diagnosis and remarks sections of his species’ treatment of  A. obsoleta . After examining specimens that Rindge likely studied at the AMNH, where he curated  Lepidoptera , there were no specimens from Ecuador, only from Peru. However, during this study, material from Ecuador was later confirmed from other museum collections. </p>
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	https://treatment.plazi.org/id/039987ADFFC5E02ED398FC75FE75FB5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matson, Tanner A.	Matson, Tanner A. (2023): The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae). Zootaxa 5352 (4): 565-576, DOI: 10.11646/zootaxa.5352.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5352.4.7
039987ADFFC1E020D398FA98FB8CFD25.text	039987ADFFC1E020D398FA98FB8CFD25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Achagua cooperae Matson 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Achagua cooperae Matson ,  n. sp.</p>
            <p>(Figs. 3, 7, 10–12)</p>
            <p>LSID: 2053905A-C10D-464D-A68F-694BF3E7E038</p>
            <p> Diagnosis.  Achagua cooperae is currently the only member of the genus known from Central America. This species bears a conspicuous antemedial black spot along the forewing inner margin and a weakly gray, discal, reniform spot that are absent in both  A. magna and  A. obsoleta . The male genitalia of  A. cooperae lack a spatulate dorsal process of the uncus, a postmedial digitate protuberance on the costa, and cornuti on the vesica, all of which are found in  A. magna and  A. obsoleta . </p>
            <p> Achagua cooperae is thought to be closely related to  A. velata . One noticeable distinction is the thickness of the black terminal band on the hindwing, which tends to be thinner in  A. cooperae compared to  A. velata . While male  A. cooperae share many genitalic similarities with  A. velata , the dorsal process of the uncus appears to be knob-like in  A. cooperae , whereas in  A. velata , it is more uniform in shape (Figs. 7a, 8a).  Achagua cooperae and  A. velata may also be separated by their COI barcode (see Molecular characterization). </p>
            <p>Description. MALE. Forewing length, 17–18 mm (n = 8).</p>
            <p>Head. Antenna mostly bipectinate, but with gradually diminishing rami that are absent in distal third of antenna; scales above white and light gray, rami dark gray to black. Vertex white; frons mostly light gray. Labial palpus short, decumbent, 1.5x diameter of eye, light gray and white. Chaetosemata in transverse row; cephalic collar mostly white with few black scales.</p>
            <p>Thorax. Patagium, tegula, and mesothorax admixture of gray and white scales. Legs mostly white and mottled with gray; epiphysis well-developed; hind tibia with large hair pencil; tibial spur formula 0–2–4.</p>
            <p>Forewing. Pearly-white; widely scattered with inconspicuous light gray scales. Basal and costal areas lightly maculated with gray to brown scales. Subtle, light gray, transverse antemedial line, and weakly gray, discal, reniform spot. Inner margin with antemedial black spot. Terminal area broadly maculated with brown scales and bearing undulating, subterminal pale stripe within. Underside patterned as in upperside but darkened areas more diffuse.</p>
            <p>Hindwing. Pearly-white except for blackened terminal area. Underside patterned as in upperside but darkened area more diffuse.</p>
            <p>Abdomen. Admixture of gray and white. Third sternite of male abdomen with comb of setae.</p>
            <p>Genitalia (Fig. 7). Uncus abruptly widened at base; dorsal process larger, gradually enlarging to pronounced swelling at distal third and with knob-like apex; ventral process large and thumb-like. Base of gnathos subquadrangular with upcurved, heavily sclerotized, and pointed apical projection; projection lightly papillated. Valve elongate, quadrate, and large with heavily sclerotized costa; apex strongly falcate. Anellar processes large and triangular, directed inward, and with apical recurved hooks. Juxta with medial, elongate cylindrical process with acuminate apex. Vesica with small medial sclerotized fold; cornuti absent.</p>
            <p>FEMALE. Forewing length, 19 mm (n = 2). Outwardly undifferentiated from male.</p>
            <p>Genitalia (Fig. 10). Papillae anales rounded; posterior apophysis 2.5x longer than anterior apophysis. Lamella antevaginalis cordiform; ostium opening into short, lightly sclerotized ductus bursae. Corpus bursae posteriorly narrow, opening into ovoid anterior portion. Signum large and invaginated, appearing somewhat cone-like; surface lightly denticulate.</p>
            <p>Type Material.</p>
            <p>Holotype</p>
            <p> COSTA RICA • ♁; Alajuela, ACG [Area De Conservación Guanacaste],  Rincon Rain Forest , casa de  Oscar Albergue ; (10.866°, -85.326°); elev. 725 m; 14 Feb. 2010; R. Franco &amp; H. Cambronero leg. (light trap);  Sample IDs : 10-SRNP-105189;  Bold Process ID: BLPDQ563 -10;  GenBank : HQ934011; USNMENT01771270; USNM. </p>
            <p>Paratypes (6♁, ♀)</p>
            <p>COSTA RICA • 2♁, ♀; same collection data as holotype; Sample IDs: 10-SRNP-105551, 10-SRNP-105757, 10-SRNP-105758; Bold Process IDs: BLPDQ925-10, BLPDR132-10, BLPDR133-10; GenBank: HQ934144, HQ934347, HQ934348; Genitalia: TAM-2023-289; USNMENT01771269, USNMENT01771271; USN- MENT01771272; USNM • ♁; Alajuela, ACG [Area De Conservación Guanacaste], Rincon Rain Forest, túnel de Oscar Albergue; (10.868°, -85.327°); elev. 708 m; 14 Feb. 2010; S. Rios &amp; F. Quesada leg. (light trap); Sample ID: 10-SRNP-105667; Bold Process ID: BLPDR042-10; GenBank: HQ934257; USNMENT01771273; USNM • 3♁; Puntarenas, 35 km NE of San Vito at Las Alturas Field Station; elev. 4800 ft; [27–29] Apr. 1992; C. Snyder leg. (at light); Genitalia: F.H.R. No. 21326 and TAM-2023-251; AMNH _IZC 00353026 to AMNH _IZC 00353028; AMNH.</p>
            <p>Other Material Examined.</p>
            <p>MEXICO • ♁; Veracruz, Los Tuxtlas, Sierra Sta. Martha, Arroyo Claro, Neck Point; 19 Mar. 1977; R. Sánchez S. leg.; Genitalia: TAM-2023-315; CNIN • ♀; Veracruz, Est. Biol. de Los Tuxtlas; Alt. 170 m; 01 Apr. 1985; P. Sinaca leg.; Genitalia: TAM-2023-316; 10194; CNIN.</p>
            <p> Distribution (Fig. 11).  Achagua cooperae is known to inhabit the Isthmian-Pacific and Atlantic moist forests of Costa Rica, but its distribution in other parts of Central America remains uncertain. Notably, two individuals were collected from the Sierra de los Tuxtlas, a remote volcanic range situated along the southeastern coast of the Veracruz Gulf in Mexico. These are tentatively regarded as conspecific (see Remarks). </p>
            <p> Biology. The life history of  A. cooperae remains unknown. Adult are known to fly from February through April. </p>
            <p> Etymology.  Achagua cooperae , is named in honor of Loretta Faye Cooper, former Senior Development Officer, and Deputy Director for Advancement at the Smithsonian’s National Museum of Natural History. Loretta’s dedication to museum research is unparalleled, and her tireless efforts to raise awareness and secure funding for the conservation and study of the natural world make her a true champion for both people and the environment. Loretta’s support for the Area de Conservación Guanacaste is particularly noteworthy and greatly appreciated. </p>
            <p> Molecular characterization.  Achagua cooperae is represented in BOLD by the BIN: BOLD:AAM6724 (n = 5). The pairwise distance to the nearest neighbor,  Achagua velata (n = 1, French Guiana), is about 3.8%. </p>
            <p> Remarks. I tentatively regard two individuals from the Sierra de los Tuxtlas of Veracruz, Mexico, as  A. cooperae . However, I have opted not to include these individuals in the type series. While the genitalia of both sexes in this Mexican population are consistent with those in Costa Rica, the Sierra de los Tuxtlas is a region characterized by biological endemism (Sánchez-González et al. 2008), and further, this population is disjunct from the remaining known distribution of  A. cooperae . </p>
            <p>The male phallus depicted in Figure 7b may give the impression of a spinate cornutus. However, this is an artifact of the preparation and the structure in question is rather a sclerotized fold.</p>
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	https://treatment.plazi.org/id/039987ADFFC1E020D398FA98FB8CFD25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matson, Tanner A.	Matson, Tanner A. (2023): The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae). Zootaxa 5352 (4): 565-576, DOI: 10.11646/zootaxa.5352.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5352.4.7
039987ADFFCFE022D398FD71FB75FD08.text	039987ADFFCFE022D398FD71FB75FD08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Achagua velata Matson 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Achagua velata Matson ,  n. sp.</p>
            <p>(Figs. 4, 8, 11, 12)</p>
            <p>LSID: 0E4A82F4-6BDF-43BE-86F8-572966809E46</p>
            <p> Diagnosis. The following diagnosis is based on information from a single individual of  A. velata , and caution should be exercised when interpreting it. As of now,  Achagua velata is currently the only member of the genus known to inhabit the Guianan moist forests of northeastern South America. This species bears a conspicuous antemedial black spot along the forewing inner margin and a weakly gray, discal, reniform spot that are absent in both  A. magna and  A. obsoleta . The terminal band on both wings is thicker and more complete than in other congeners. The male genitalia of  A. velata lack a spatulate dorsal process of the uncus, a postmedial digitate protuberance on the costa, and cornuti on the vesica, all of which are found in  A. magna and  A. obsoleta . While male  A. velata shares many genitalic similarities with  A. cooperae , the dorsal process of the uncus appears knob-like in  A. cooperae (Fig. 7a), whereas in  A. velata (Fig. 8a), it is more uniform in shape.  Achagua velata and  A. cooperae may also be separated by their COI barcode (see Molecular characterization). </p>
            <p>Description. MALE. Forewing length, 17 mm (n = 1).</p>
            <p>Head. Antenna mostly bipectinate, but with gradually diminishing rami that are absent in distal third of antenna; scales above white and light gray, rami dark gray to black. Vertex white; frons mostly light gray. Labial palpus short, decumbent, 1.5x diameter of eye, light gray and white. Chaetosemata in transverse row.</p>
            <p>Thorax. Patagium, tegula, and mesothorax admixture of gray and white scales. Legs mostly white and mottled with gray; epiphysis well-developed; hind tibia with hair pencil (not easily visualized in holotype); tibial spur formula 0–2–4.</p>
            <p>Forewing. Pearly-white; widely scattered with inconspicuous light gray scales. Basal and costal areas lightly maculated with gray to brown scales. Subtle, light gray, transverse antemedial line, and weakly gray, discal, reniform spot. Inner margin with antemedial black spot. Terminal area broadly maculated with brown scales. Underside patterned as in upperside, but darkened areas more diffuse and given more toward black.</p>
            <p>Hindwing. Pearly-white except for complete, broadly dark brown outer margin. Underside patterned as in upperside.</p>
            <p>Abdomen. Admixture of gray and white. Third sternite of male abdomen with comb of setae.</p>
            <p>Genitalia (Fig. 8). Uncus abruptly widened at base; dorsal and ventral processes thumb-like. Base of gnathos subquadrangular, with upcurved, heavily sclerotized, pointed apical projection; projection lightly papillated. Valve elongate and large with heavily sclerotized costa; apex strongly falcate. Anellar processes large and triangulate, directed inward, and with apical recurved hooks. Juxta with medial, long cylindrical process with acuminate apex. Vesica with small medial sclerotized patch; cornuti absent.</p>
            <p>FEMALE. Unknown.</p>
            <p>Type Material.</p>
            <p>Holotype</p>
            <p> FRENCH GUIANA [FRANCE] • ♁;  
Régina, 
Nouragues Nature Reserve; (4.096°, -52.683°); elev. 419 m; 09 Jul. 2010;  Carlos Lopez-Vaamonde leg.; Genitalia: TAM-2023-289; BOLD Process ID: LNOUD2050 -12; MNHN. </p>
            <p> Distribution (Fig. 11). Presently,  A. velata is only known from the Guianan moist forests at the type locality in French Guiana. </p>
            <p>Biology. Immature stages and host associations remain unknown. Adults are known to fly in July at the type locality.</p>
            <p>Etymology. The specific epithet velata is derived from the Latin “velatus,” meaning “veiled.” The name was chosen because of the rarely seen nature of this species—thus far only known from a single individual—and for its white, wedding-veil-like ground color.</p>
            <p> Molecular characterization.  Achagua velata is represented in BOLD by the BIN: BOLD:ABV2454 (n = 1). The pairwise distance to the nearest neighbor,  Achagua cooperae (n = 5, Costa Rica), is about 3.8%. </p>
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	https://treatment.plazi.org/id/039987ADFFCFE022D398FD71FB75FD08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matson, Tanner A.	Matson, Tanner A. (2023): The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae). Zootaxa 5352 (4): 565-576, DOI: 10.11646/zootaxa.5352.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5352.4.7
