taxonID	type	description	language	source
0396878DF55C6C7E3EA3A208F5F5FFCF.taxon	materials_examined	Type species Acanthoctenus variatus Thorell, 1890, from Nias Island.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55C6C7E3EA3A208F5F5FFCF.taxon	discussion	Note. The genus Acantheis was described in a brief footnote, with the three species being assigned to it (Thorell, 1891): viz., A. variatus, A. dimidiatus and A. laetus. The type species was not indicated in that paper. Simon (1897) was the first who considered the genus and selected the generotype (A. variatus), although it was described from a juvenile specimen (probably lost according to Lehtinen [1967]), while two others were known from the males. It is worth noticing that Simon (1897) considered the genus in Acantheae, which later were called the subfamily Acantheinae by Simon (1897). Initially, we identified our specimens as Acantheis based on similarity of their male palps with that of A. laetus, which was depicted by Lehtinen (1967). The following features, highlighted by Simon (1897), support their belonging to this genus: high clypeus, long palpal tibia in the males (length / width ratio 4 +) and 8 – 9 pairs of ventral spines on tibiae I (cf. Simon 1897: figs 109 – 111). Our specimens from Sumatra possess all these characters. Acantheis nipponicus from the heavily isolated South Iwo Jima Island has the cymbium with a shortened tip and a slightly modified proximal part and the embolus originating from the proximal part of bulb, which casts doubt on the accuracy of the generic assignment of this geographically isolated species. It could belong to another, yet undescribed genus. Africactenus unumus Sankaran & Sebastian, 2018, described based on males from the south of India (Sankaran & Sebastian 2018), is another species with doubtful placement. This species possesses several features which we have identified as distinctive for Acantheis, namely: 1) presence of clusters of stiff setae on the dorso-posterior part of opisthosoma; 2) elongated cymbial tip as long as the bulb; 3) the bifurcated embolic tip is divided into dorsal and ventral branches pointing in the same direction (cf. Sankaran & Sebastian 2018: figs 1 A, 3). Due to this there is a possibility that Africactenus unumus actually belongs to Acantheis. However, Africactenus unumus has features not typical for Acantheis (except A. nipponicus): modified cymbium with lamella-like outgrowth retro-proximally (vs. cymbium with no modifications) and straight and thick embolus (vs. thin and strongly curved). So, we refrain from transferring of this species as long as Acantheis remains unrevised. A series of photographs of four Acantheis morphospecies, which are most probably new to science, were published in the photographic field guide (Koh & Bay 2019). This account, as well as our findings, suggests that the real species diversity of Acantheis in South-East Asia is much higher than the currently known. We are confident that a large number of Acantheis species will be discovered and described in the near future.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55C6C7E3EA3A208F5F5FFCF.taxon	diagnosis	Diagnosis. Acantheis differs from all the ctenid genera known from South-East Asia (Amauropelma Raven, Stumkat et Gray, 2001; Anahita Karsch, 1879; Bowie Jäger, 2022 and Sinoctenus Marusik, Zhang et Omelko, 2012) by the presence of clusters of stiff setae (CS) on the dorso-posterior part of opisthosoma (Fig. 7). Males of Acantheis differ in having an elongated cymbial tip, which is as long as the bulb (vs. the cymbial tip significantly shorter than the bulb, or even absent, cf. Figs 14, 19 and Polotow & Brescovit 2014: fig. 6 b; Jäger 2012: fig. 32; Jäger 2022: fig. 338; Marusik et al. 2012: figs 9 – 11), and the bifurcated embolic tip, which is divided into dorsal and ventral branches pointing in the same direction (Figs 30, 31; vs. undivided apical part of embolus or divided into prolateral and retrolateral branches). Additionally, Acantheis males can be distinguished from all south-east Asian ctenids but Anahita by the proximal part of the cymbium with no modifications (vs. the presence of prolateral cymbial bulge or retro-proximal cymbial outgrowth). Males of Acantheis can be separated from those of Anahita by the embolus starting from the prolateral side of the bulb (vs. the retrolateral or posterior side, cf. Figs 23, 27 and Jäger 2012: fig. 32). Females of Acantheis differ from those of other Oriental ctenid genera, except Bowie, in having the receptacles subdivided into two chambers (vs. undivided, cf. Fig. 34 and Polotow & Brescovit 2014: fig. 15 b; Jäger 2012: fig. 26). The endogynes of Bowie are polymorphic, representing both subdivided and undivided receptacles (cf. Jäger 2022: figs 6, 529, 532). However, the Bowie species with subdivided receptacles always have the receptacle chambers of different sizes: one could be 2 – 4 times (or even more) larger than the second one. In Acantheis, such receptacle chambers are about equal in size.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	description	Figs 1 – 2, 7 – 9, 12 – 16, 22 – 25, 30, 46 – 48	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	materials_examined	Type. INDONESIA: Sumatra Island: Aceh Prov.: holotype: ♁ (ISEA, 001.9531), Ketambe Vil. [03 ° 41 ′ N, 97 ° 39 ′ E], 400 – 500 m, 1988 (precise date unknown), unknown collector.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	etymology	Etymology. The species name is a patronym given in honor of the teacher and friend of Alexander A. Fomichev, Sergei I. Mishenin (Novosibirsk, Russia), on the occasion of his 60 th anniversary. S. I. Mishenin is the first mentor of Alexander A. Fomichev, who inspired his interest in arachnology.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	diagnosis	Diagnosis. This new species is similar to A. laetus from Kalimantan and to A. dimidiatus and A. andreimishenini sp. n. from Sumatra in having a slender embolus (Em), a medially located tegular apophysis (TA) and an elongated cymbial tip. The male can be easily distinguished from that of A. laetus by the strongly curved embolus (vs. smoothly rounded, cf. Fig. 23 and Lehtinen 1967: fig. 410). The male of A. sergeimishenini sp. n. differs from that of A. andreimishenini sp. n. in having a larger body size (carapace length 7.7 vs. 5.8), a longer bulb (1.32 mm vs. 1.03 mm), a bifurcated retrolateral tibial apophysis (RTA) (vs. non-bifurcated, cf. Figs 15 and 20), the comma-shaped tegular apophysis (in ventral view; vs. round, cf. Figs 23 and 27), and the ventral embolic branch (VB) twice as wide as the dorsal one (DB) (vs. the same width, cf. Figs 30 and 31). The male of A. sergeimishenini sp. n. differs from that of A. dimidiatus by the bifurcated retrolateral tibial apophysis (vs. non-bifurcated, cf. Figs 15 and 41), the comma-shaped tegular apophysis (vs. triangular, cf. Figs 23 and 43), and the ventral embolic branch twice as wide as the dorsal one (vs. ventral embolic branch twice as thin as the dorsal one, cf. Figs 30 and 41). For a list of the differences between the males of A. sergeimishenini sp. n., A. andreimishenini sp. n. and A. dimidiatus, see Table 7.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	description	Description. Male. Total length 15.1, carapace 7.7 long, 6.3 wide. Opisthosoma 7.0 long, 4.0 wide. Eye sizes and interdistances: AME 0.46, ALE 0.24, PME 0.5, PLE 0.49, AME – AME 0.2, AME – ALE 0.33, PME – PME 0.21, PME – PLE 0.31, AME – PME 0.21, ALE – PLE 0.2. Clypeus height at AME 0.7, at ALE 1.17. For palp and legs measurements see Table 1. For palp and legs spination see Table 2. Coloration (Figs 1 – 2, 7 – 9), carapace brown, with a thin longitudinal light stripe. Clypeus, chelicerae, endites and labium dark brown. Sternum and coxae yellow. Palps yellow with brown-grey annulations. Legs yellow-brown, with very vague brown-grey annulations. Opisthosoma yellow-grey with thin longitudinal light stripe dorsally. Posterior part of opisthosoma with two pairs of clusters of stiff yellow setae (CS). Venter of opisthosoma dirty yellow. Spinnerets dark grey. Male palp as shown in Figs 12 – 16, 22 – 25, 30. Tibia ca. 4.5 longer than wide, with 4 very long spines, longest ca. 0.9 of tibia length. Retrolateral tibial apophysis (RTA) small, bifurcated. Ventral RTA branch longer than dorsal one. Cymbium length / width ratio ca. 2.7. Cymbial tip about 0.43 of its length, as long as bulb. Subtegulum (St) large, oval. Tegular length / width ratio ca. 1.5. Sperm duct (SD) clearly visible only in retrolateral view. Retrolateral margin of tegular process (TP) convex. Tegular apophysis (TA) comma-shaped. Conductor (Cn) five times longer than wide, not covering embolic tip. Embolic base with basal process (BP) located near subtegular notch (SN). Membranous extension of embolus (ME) wider than its adjoining part. Embolus (Em) strongly curved, with a small spike (ES) ventrally. Ventral branch of embolus (VB) twice as wide as dorsal one (DB). Female. Unknown.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55F6C7A3EA3A27EF4FBFF55.taxon	distribution	Distribution. Only known from the type locality (Figs 46 – 48).	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	description	Figs 3 – 6, 10 – 11, 17 – 21, 26 – 29, 31 – 35, 46 – 48	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	materials_examined	Types. INDONESIA: Sumatra Island: Aceh Prov.: holotype ♁ (ISEA, 001.9532), Kedah Vil. [03 ° 59 ′ N, 97 ° 15 ′ E], 1300 – 1500 m, 1988 (precise date unknown), unknown collector. Paratypes: 3 ♁ (ISEA, 001.9533), 1 ♀ (ISEA, 001.9534), together with the holotype.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	etymology	Etymology. The species name is a patronym given in honor of the teacher and friend of Alexander A. Fomichev, Andrei I. Mishenin (Novosibirsk, Russia), brother of Sergei I. Mishenin, on the occasion of his 60 th anniversary.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	diagnosis	Diagnosis. This new species is similar to A. laetus from Kalimantan and to A. sergeimishenini sp. n. and A. dimidiatus from Sumatra in having a slender embolus (Em), a medially located tegular apophysis (TA) and an elongated cymbial tip. The male can be easily distinguished from that of A. laetus by the strongly curved embolus (vs. smoothly rounded, cf. Fig. 27 and Lehtinen 1967: fig. 410), and from that of A. sergeimishenini sp. n. by the smaller body size (carapace length 5.8 vs. 7.7), the shorter bulb (1.03 mm vs. 1.32 mm), the non-bifurcated retrolateral tibial apophysis (RTA) (vs. bifurcated, cf. Figs 20 and 15), the round tegular apophysis (seen in ventral view; vs. comma-shaped, cf. Figs 27 and 23) and the ventral (VB) and dorsal (DB) branches of the embolus with the same width (vs. the ventral branch twice as wide as the dorsal one, cf. Figs 31 and 30). The male of A. andreimishenini sp. n. differs from that of A. dimidiatus by the wide cardiac mark reaching middle part of opisthosoma (vs. thin stripe reaching spinnerets, cf. Figs 3 and 36), the median band on the carapace as wide as the distance between PLE (vs. as wide as distance between PME), the circular tegular apophysis (vs. triangular, cf. Figs 27 and 43), and the dorsal and ventral embolic branches equal in size (vs. ventral embolic branch twice thinner than dorsal one, cf. Figs 31 and 41). For a complete list of the differences between the males of A. andreimishenini sp. n., A. sergeimishenini sp. n. and A. dimidiatus, see Table 7. The female of A. andreimishenini sp. n. is similar to that of A. indicus Gravely, 1931 from India, in having the rectangular septal base (SB), but differs in having parallel epigynal teeth (ET) (vs. converging), the posterior part of septal base with rounded angles (vs. sharp), and a wide gap between epigynal teeth and septal base (vs. narrow, cf. Figs 32 – 33 and Gravely 1931: fig. 3 D).	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	description	Description. Male (holotype). Total length 11.5, carapace 5.8 long, 5.2 wide. Opisthosoma 6.1 long, 3.5 wide. Eye sizes and interdistances: AME 0.39, ALE 0.2, PME 0.46, PLE 0.43, AME – AME 0.17, AME – ALE 0.26, PME – PME 0.21, PME – PLE 0.29, AME – PME 0.14, ALE – PLE 0.23. Clypeus height at AME 0.49, at ALE 0.93. Labium as long as wide. For palp and legs measurements see Table 3. For palp and leg spination see Table 4. Coloration (Figs 3 – 4, 10), carapace brown, with yellow median band. Clypeus and chelicerae dark brown. Labium and endites brown. Sternum and coxae yellow. Palps and legs yellow-brown, with grey annulations. Opisthosoma dorsally yellow-gray, with yellow cardiac mark. Posterior part of opisthosoma bears several clusters of stiff yellow setae. The venter dirty yellow. Spinnerets yellow-grey. Male palp as shown in Figs 17 – 21, 26 – 29, 31. Tibia ca. 4.5 longer than wide, with five very long spines, the longest is about 0.7 times of tibia length. Retrolateral tibial apophysis (RTA) small, non-bifurcated. Cymbium length / width ratio ca. 2.4. Tip of cymbium about 0.39 of cymbial length, sligthly shorter than bulb. Bulb 1.25 times longer than cymbial apex. Subtegulum (St) oval. Tegular length / width ratio ca. 1.4. Sperm duct (SD) clearly visible only in retrolateral view. Retrolateral margin of tegular process (TP) concave. Tegular apophysis almost round. Conductor (Cn) 2.4 times longer than wide, covering embolic tip. Embolic base with basal process (BP) located near subtegular notch (SN). Membranous extension of embolus (ME) thinner than its adjoining part. Embolus (Em) strongly curved, with a small spike (ES) ventrally. Dorsal branch (DB) of embolus as wide as ventral one (VB). Female. Total length 14.86, carapace 6.38 long, 5.42 wide. Opisthosoma 8.32 long, 5.66 wide. Eye sizes and interdistances: AME 0.37, ALE 0.23, PME 0.46, PLE 0.48, AME – AME 0.22, AME – ALE 0.34, PME – PME 0.23, PME – PLE 0.35, AME – PME 0.20, ALE – PLE 0.25. Clypeus height at AME 0.53, at ALE 1.05. For palp and legs measurements see Table 5. For palp and legs spination see Table 6. Coloration (Figs 5 – 6, 11), carapace, clypeus, chelicerae, labium and endites as in the male. Sternum and coxae brown. Palps and legs as in the male, but with more contrasting annulations. Opisthosoma as in the male, but somewhat lighter, its posterior part bears several clusters of stiff yellow setae. Female genitalia as shown in Figs 32 – 35. Epigyne slightly wider than long. Septal base (SB) almost square, with heavily sclerotized septal humps (SH). Septal stem (SS) 1.5 times shorter than septal base. Epigynal teeth (ET) parallel, as long as septal stem. Receptacles (Re) subdivided into two chambers: chamber I (RI) and chamber II (RII), both are about of the same size.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF55B6C7B3EA3A5C1F4FBFB9B.taxon	distribution	Distribution. Only known from the type locality (Figs 46 – 48).	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF5566C773EA3A793F4FBF9AE.taxon	description	Figs 36 – 48	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF5566C773EA3A793F4FBF9AE.taxon	materials_examined	Material examined. 1 ♁ (holotype) (MSNG), INDONESIA: Sumatra Island: Singalong, O. Beccari. Examined by photo.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF5566C773EA3A793F4FBF9AE.taxon	diagnosis	Diagnosis. The species is similar to A. laetus from Kalimantan and to A. sergeimishenini sp. n. and A. andreimishenini sp. n. from Sumatra in having the slender embolus (Em), the medially located tegular apophysis (TA) and the elongated cymbial tip. The male of A. dimidiatus differs from that of A. laetus by by the strongly curved embolus (vs. smoothly rounded, cf. Fig 43 and Lehtinen 1967: fig. 410). The male of A. dimidiatus can be distinguished from that of A. sergeimishenini sp. n. by the non-bifurcated retrolateral tibial apophsis (RTA) (vs. bifurcated, cf. Figs 41 and 15), the triangular tegular apophysis (vs. comma-shaped, cf. Figs 43 and 23), and the ventral embolic branch twice thinner than dorsal one (vs. ventral embolic branch twice wider than dorsal one, cf. Figs 41 and 30). From the males of A. andreimishenini sp. n. those of A. dimidiatus differs by the dorsal surface of opisthosoma with a thin stripe reaching spinnerets (vs. wide cardiac mark reaching middle part of opisthosoma, cf. Figs 36 and 3), the median band on the carapace as wide as distance between PME (vs. as wide as the distance between PLE), the triangular tegular apophysis (TA) (vs. circular, cf. Figs 43 and 27), and the ventral embolic branch twice thinner than dorsal one (vs. dorsal and ventral embolic branches equal in size, cf. Figs 41 and 31). For a complete list of the differences between the males of A. dimidiatus, A. sergeimishenini sp. n. and A. andreimeshenini sp. n. see Table 7.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF5566C773EA3A793F4FBF9AE.taxon	description	Description. Male palp as shown in Figs 37 – 43. Tibia ca. 4.6 longer than wide, 4 very long spines, longest ca. 0.7 of tibia length. Retrolateral tibial apophysis small, non-bifurcated. Cymbial tip about 0.42 of its length, slightly shorter than bulb. Sperm duct clearly visible only in retrolateral view. Tegular apophysis triangular. Embolus strongly curved. Ventral branch of embolus twice thinner than dorsal one. Female. Unknown.	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
0396878DF5566C773EA3A793F4FBF9AE.taxon	distribution	Distribution. Only known from the type locality (Figs 46 – 48).	en	Fomichev, Alexander A., Omelko, Mikhail M., Marusik, Yuri M. (2023): A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species. Zootaxa 5353 (2): 117-130, DOI: 10.11646/zootaxa.5353.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5353.2.2
