identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
1D10BFF986365B86B74598934F7F1600.text	1D10BFF986365B86B74598934F7F1600.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichosetodes carmelae Laudee & Malicky 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Trichosetodes carmelae Laudee &amp; Malicky sp. nov.</p>
            <p>Fig. 1</p>
            <p>Type material.</p>
            <p>Holotype. Male. Cambodia: Ratanakiri Province: Banlung, Katieng Waterfall, 13°40'38"N, 106°58'33"E, elev. 203 m, 13.v.2022, Pongsak Laudee. (PSUNHM). Paratypes: Same data as holotype, 6 males: 2 males (PSUNHM), 2 males (CHM), 2 males (CUAC).</p>
            <p>Diagnosis.</p>
            <p> The male genitalia of the new species are moderately similar to those of  T. sotet Oláh , 2013 described in Vietnam in the form of segment IX and inferior appendages, as well as of  T. insularis Schmid, 1987 in the form of segment IX and segment X; but the phallicata or aedeagus is clearly different. The phallicata of  T. sotet and  T. insularis are divided into subbasodorsal branch and subbasoventral branch, whereas such features are missing in  T. carmelae . The phallicata in  T. carmelae bears tuft of long hairs in the middle of dorsal edge, which does not occur in  T. sotet . The phallicata in  T. carmelae is slightly bent upward subapically, while it is curved downward in  T. sotet . </p>
            <p>Description.</p>
            <p>Length of each male forewing 4.5 mm (N = 5); specimens in alcohol with head, thorax, abdomen, legs, forewings light brown.</p>
            <p>Male genitalia (Fig. 1A-D). Segment IX right trapezoid, anterior margin convex dorsally and truncated ventrally, posterior margin truncated in lateral view (Fig. 1A); rectangular with pair of notches anteriorly in ventral view (Fig. 1C). Preanal appendages thumb-like covered with hairs (Fig. 1A, B). Segment X produced in pair of javelin-like processes each with acute apex (Fig. 1A, B). Inferior appendages each with dorsal and ventral lobes, dorsal lobe broad rectangular, serrated dorsally with small triangular process, ventral lobe slender with acute apex directed dorsad in lateral view (Fig. 1A); in ventral view, claw-like, bent inward, each with inner broad tooth sub-basally (Fig. 1C). phallicata in dorsal view, slender, lancet-like, acute apex, with isolated bunch of long hair in the middle; ejaculatory duct tube-like, about half as long as phallicata length (Fig. 1B); in lateral view, phallus typically large, axe-like, broad basally, strongly curved backward subbasally, with isolated bunch of long hairs dorsally, bent subapically, sharp apex (Fig. 1D).</p>
            <p>Etymology.</p>
            <p>The species name is dedicated to Dr Carmela R. Centrino who works for United Nations Industrial Development, Vienna International Centre for Southeast Asian Countries.</p>
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	https://treatment.plazi.org/id/1D10BFF986365B86B74598934F7F1600	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Laudee, Pongsak;Malicky, Hans;Kong, Chamroeun;Takenaka, Masaki;Tojo, Koji	Laudee, Pongsak, Malicky, Hans, Kong, Chamroeun, Takenaka, Masaki, Tojo, Koji (2023): New species of the genus Trichosetodes Ulmer, 1915 (Trichoptera, Leptoceridae) from Ratanakiri province, Cambodia, based on morphological and molecular data. ZooKeys 1182: 153-164, DOI: http://dx.doi.org/10.3897/zookeys.1182.105716, URL: http://dx.doi.org/10.3897/zookeys.1182.105716
5AA2A611A1AD588AA1BA8FA849DBCF26.text	5AA2A611A1AD588AA1BA8FA849DBCF26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichosetodes katiengensis Laudee & Malicky 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Trichosetodes katiengensis Laudee &amp; Malicky sp. nov.</p>
            <p>Fig. 2</p>
            <p>Type material.</p>
            <p>Holotype. Male. Cambodia: Ratanakiri Province: Banlung, Katieng Waterfall, 13°40'38"N, 106°58'33"E, elev. 203 m, 13.v.2022, Pongsak Laudee. (CHM). Paratypes: same data as holotype. 2 males: 1 male (PSUNHM), 1 male (CHM).</p>
            <p>Diagnosis.</p>
            <p> The male genitalia of the new species are moderately similar to those of  Trichosetodes pales Malicky &amp; Chaibu, 2006 described in Thailand, in the form of segment IX and inferior appendages, however, the phallus is clearly different. The phallicata of  T. pales is divided into a dorsal branch, median branch and ventral branch, however, these features are missing in  T. katiengensis . In addition, ventral lobes of inferior appendages are truncated and pointed in  T. pales in both ventral and lateral views. </p>
            <p>Description.</p>
            <p>Length of each male forewing 3.5-4.0 mm (N = 3); specimens in alcohol with head, thorax, abdomen, legs, forewings dark brown.</p>
            <p>Male genitalia (Fig. 2A-D). Segment IX with pair of thumb-like lobes posteriorly in dorsal view (Fig. 2B); right trapezoid, anterior margins convex with small lobe anterodorsally, posterior margin slightly truncated in lateral view (Fig. 2A); square with shallow notches anteriorly in ventral view (Fig. 2C). Preanal appendages thumb-like covered with hairs (Fig. 2A, B). Segment X not evident. Inferior appendages with dorsal and ventral lobes, dorsal lobe triangular in lateral view with sharp process basoposteriorly, ventral lobe tubular and truncated apically in lateral view (Fig. 2A); in ventral view, horn-like, bent inward, truncated apically (Fig. 2C). In dorsal view, phallus complex, phallicata divided into dorsal and ventral branches; dorsal branch with outer edge denticulated and ventral branch undulated with acute apex; ejaculatory duct short and thin (Fig. 2B). In lateral view, phallicata sickle-like with dorsal and ventral branches, dorsal branch of phallicata straight, covered with numerous short protrusions subapically, apex with small spines; ventral branch of phallicata claw-like, curved downward, acute apex; ejaculatory duct curved tube-like (Fig. 2D).</p>
            <p>Etymology.</p>
            <p>The species is named for the type locality, Katieng Waterfall.</p>
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	https://treatment.plazi.org/id/5AA2A611A1AD588AA1BA8FA849DBCF26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Laudee, Pongsak;Malicky, Hans;Kong, Chamroeun;Takenaka, Masaki;Tojo, Koji	Laudee, Pongsak, Malicky, Hans, Kong, Chamroeun, Takenaka, Masaki, Tojo, Koji (2023): New species of the genus Trichosetodes Ulmer, 1915 (Trichoptera, Leptoceridae) from Ratanakiri province, Cambodia, based on morphological and molecular data. ZooKeys 1182: 153-164, DOI: http://dx.doi.org/10.3897/zookeys.1182.105716, URL: http://dx.doi.org/10.3897/zookeys.1182.105716
F36027ACD7C45B559F2FC7D8531DCB25.text	F36027ACD7C45B559F2FC7D8531DCB25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichosetodes ratanakiriensis Laudee & Malicky 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Trichosetodes ratanakiriensis Laudee &amp; Malicky sp. nov.</p>
            <p>Fig. 3</p>
            <p>Type material.</p>
            <p>Holotype. Male. Cambodia: Ratanakiri Province: Banlung, Katieng Waterfall, 13°40'38"N, 106°58'33"E, elev. 203 m, 13.v.2022, Pongsak Laudee. (CHM). Paratypes: same data as holotype. 33 males: 18 males (PSUNHM), 5 males (CHM), 5 males (CUAC), 5 males (NMPC).</p>
            <p>Diagnosis.</p>
            <p> The male genitalia of  T. ratanakiriensis are moderately similar to those of  T. pandareos Malicky, 2006 described in Laos and  T. kampongspeuensis Malicky &amp; Kong, 2020 (Fig. 4), however it can be differentiated by the shape of left inferior appendage and the shape of segment IX. The left inferior appendage is forked in  T. pandareos whereas this feature is missing in  T. ratanakiriensis . In addition, the prolongations of the subapicoventral part of segment IX are symmetric in  T. pandareos but asymmetric in  T. ratanakiriensis in ventral view. Compared to  T. kampongspeuensis ,  T. ratanakiriensis exhibits a longer, cylindrical right inferior appendage with a sharp apex in lateral view, whereas in  T. kampongspeuensis , this appendage is oval and splits into two at the tip. The prolongations of subapical part of segment IX in ventral view are thin in  T. kampongspeuensis but prominent in  T. ratanakiriensis . </p>
            <p>Description.</p>
            <p>Length of each male forewing 5 mm (N = 12); specimens in alcohol with head, thorax, abdomen, legs, forewings dark brown.</p>
            <p>Male genitalia (Fig. 3A-D). Segment IX square with U-shaped incision anteriorly in dorsal view (Fig. 3C); in left lateral view, complicated shape, strongly convex anteriorly, undulated edge dorsally, curved downward ventrally with pimple sub-anteroventrally, long cylindrical subapicoventrally, rounded apically (Fig. 3A); in right lateral view the same as left lateral view but without pimple (Fig. 3B); in ventral view, vertical profile rectangular with 1/3 of height in U-shaped incision apically, rounded apex (Fig. 3D). Preanal appendages slender covered with hairs (Fig. 3A, B). Segment X not evident. Inferior appendages asymmetrical; circular basally, conical apically in left lateral view (Fig. 3A); in right lateral view, cylindrical with expanded basally, curved downward with tooth at dorsal edge subapically, pointed apically (Fig. 3B); in dorsal and ventral view, left inferior appendage claw-like, right inferior appendage long claw-like with an inner tooth. Phallicata large, tubular, curved and bent subapically, pointed apically in dorsal view; in lateral view, upside down U-shaped, pointed apically.</p>
            <p>Etymology.</p>
            <p>The species is named for the type locality, Ratanakiri Province.</p>
            <p>Molecular analysis</p>
            <p> The molecular diversity of the new  Trichosetodes species was analyzed using the mitochondrial large subunit ribosomal rRNA gene region (16S rRNA). Based on genetic distance (p -distance) of this gene fragment,  T. ratanakiriensis sp. nov. and  T. kampongspeuensis have a close genetic relationship, whereas  T. carmelae sp. nov. showed greater genetic divergence from both  T. ratanakiriensis sp. nov. and  T. kampongspeuensis (Table 1). The estimated phylogenetic relationships based on the mtDNA 16S rRNA are shown in Fig. 5, where all three  Trichosetodes sp. nov. were genetically differentiated. The monophyly of  T. ratanakiriensis sp. nov. and  T. kampongspeuensis was highly supported by the bootstrap value (NJ BP). In addition,  T. carmelae sp. nov. was differentiated from the monophyletic clade of  T. ratanakiriensis sp. nov. and  T. kampongspeuensis (Fig. 5). </p>
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	https://treatment.plazi.org/id/F36027ACD7C45B559F2FC7D8531DCB25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Laudee, Pongsak;Malicky, Hans;Kong, Chamroeun;Takenaka, Masaki;Tojo, Koji	Laudee, Pongsak, Malicky, Hans, Kong, Chamroeun, Takenaka, Masaki, Tojo, Koji (2023): New species of the genus Trichosetodes Ulmer, 1915 (Trichoptera, Leptoceridae) from Ratanakiri province, Cambodia, based on morphological and molecular data. ZooKeys 1182: 153-164, DOI: http://dx.doi.org/10.3897/zookeys.1182.105716, URL: http://dx.doi.org/10.3897/zookeys.1182.105716
