taxonID	type	description	language	source
0388A953FF97FFC49A93F8F6FC953E15.taxon	description	Austroconophron Franz, 1971: 69 (as subgenus of Euconnus). Type specIes: Euconnus tindoui Franz, 1971 (des. orIg.). SynonymIzed wIth Sciacharis s. str. by Franz (1986 a); here removed from synonymy and resurrected as subgenus of Euconnus.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF97FFC49A93F8F6FC953E15.taxon	diagnosis	Revised diagnosis. Euconnus with the following combination of characters: antennae gradually thickened; pronotum suboval with weakly marked anterior and posterior corners, strongly rounded sides, broadest in front of middle, lacking antebasal pits, grooves and sublateral carinae; basal elytral foveae vestigial; procoxal rests on mesoventrite with diffuse posterior margins; metaventral intermetacoxal process narrower than 1 / 3 of metaventral width, deeply emarginate; bristles present on head and prothorax but on the latter restricted to small anteroventral area.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF97FFC49A93F8F6FC953E15.taxon	description	Redescription. General body shape (Fig. 1) strongly elongate and strongly convex, body deeply constricted between head and pronotum and between pronotum and elytra. Head (Figs 10 – 11) large in relation to pronotum; divided by occipital constriction into large anterior and small posterior part (' neck' region), posterior part retracted into pronotum. Occipital constriction slightly broader than half width of head. Anterior part of head capsule convex dorsally and flattened ventrally, rounded hexagonal; tempora longer than compound eyes; vertex transverse and not bulging posterodorsally, with posterior margin marked by rounded ridge, anteriorly vertex confluent with subtrapezoidal frons; supraantennal tubercles feebly marked; frontoclypeal groove distinct, deep. Gular plate (Fig. 11; gp) broad and with distinct gular sutures; posterior tentorial pits (Fig. 11; ptp) strongly elongate, slot-shaped and located in front of transverse impression separating ' neck' region from anterior part of head. Tempora, genae and postgenae with thick bristles. Submentum (Fig. 11; smn) subtriangular; hypostomal ridges (Fig. 11; hr) nearly complete, sinuate, not connected with posterior tentorial pits; mentum (Fig. 11; mn) subtrapezoidal; prementum small and transverse, largely membranous; labial palps short; maxillae generalized, as in all subgenera of Euconnus; mandibles only partly visible in the studied specimen, subtriangular and evenly curved, each with one preapical mesal tooth. Antennae (Fig. 1) long and slender, gradually and distinctly thickened. Prothorax (Figs 10 – 12) convex, elongate, broadest in front of middle, with strongly rounded sides and weakly marked, obtuse-angled anterior and posterior corners; pronotum lacking sublateral carinae, antebasal pits or grooves. Small anteroventral area on each side of prothorax covered with thick bristles. Prosternum with basisternal part (Fig. 12; bst) much shorter than coxal part; interprocoxal area with narrow and weakly elevated carina. Hypomeral ridges (Fig. 12; hyr) and notosternal sutures (Fig. 12; nss) complete. Mesothorax. Mesoscutellum in intact specimens entirely covered by posterior pronotal margin, its posterior tip not exceeding posterior margin of elytral articulating lobe. Mesoventrite (Figs 12 – 13) with a pair of transverse setose impressions functioning as procoxal rests (Figs 12 – 13; pcr) in anterior region, impressions separated at middle and with diffuse posterior margins; mesoventral intercoxal process (Fig. 13; msvp) carinate. Metathorax. Metaventrite (Fig. 13) much longer than mesoventrite, subtrapezoidal; intermetacoxal area weakly expanded posteriorly and forming short and relatively narrow metaventral intercoxal process (Fig. 13; mtvp) with strongly concave posteromedian margin. Elytra (Figs 1, 10) oval, with rounded apices and narrow base which is subequal in width to posterior pronotal margin; humeral calli and basal impressions absent; each elytron with two barely discernible and asetose rudiments of basal foveae. Metathoracic wings absent. Legs (Fig. 1) moderately long and slender, unmodified. Abdomen elongate, gradually narrowing posteriorly, suture between two terminal visible sternites (i. e., VII and VIII) less distinct than between remaining sternites. Aedeagus (Figs 30 – 31) large, with symmetrical median lobe, heavily sclerotized, with modified apical region, in lateral view median lobe curved and with distinct collar surrounding basal foramen; parameres slender, with apical setae.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF97FFC49A93F8F6FC953E15.taxon	discussion	Remarks. The type species of Austroconophron, Euconnus tindoui (Fig. 1), has the metaventral intermetacoxal process (Fig. 13; mtvp) relatively narrow, but it clearly separates metacoxae, whereas in Sciacharis s. str. the metacoxae are contiguous (Jałoszyński 2014 a). This character excludes Austroconophron from Sciacharis. Besides the separated metacoxae, the placement of Austroconophron in Euconnus is supported by strongly elevated and carinate mesoventral intercoxal process (Fig. 13; msvp); two (although vestigial) basal elytral foveae; and the prothorax with complete notosternal sutures, hypomeral ridges, short basisternal portion of prosternum and interprocoxal region with a barely discernible carina. Moreover, the aedeagus of E. tindoui (Figs 30 – 31) is not thin-walled and approximately drop- or bottle-shaped, as that in Sciacharis s. str., but it has a heavily sclerotized dorsal wall of the median lobe and a strongly modified apical region, both structures frequently found among species of Euconnus. Austroconophron resembles Tetramelus Motschulsky, 1870 in the general body shape, antennal structure and the elytra with reduced basal structures, even more so than in Tetramelus, which lacks humeral calli due to the loss of wings, but has a pair of distinct and deep basal elytral foveae; the latter are rudimentary in Austroconophron. These subgenera can be easily distinguished on the basis of cephalic, prothoracic and metaventral structures. Tetramelus has the vertex bulging posterodorsally; the sides of pronotum sinuate, distinctly concave behind the broadest site, distinct antebasal pits and sublateral carinae, and often also a median antebasal longitudinal carina; and the intermetacoxal process at least as broad as 1 / 4 of metaventral width (often broader).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF97FFC49A93F8F6FC953E15.taxon	description	Composition and distribution. Euconnus (Austroconophron) currently includes 21 valid species names; all species occur in New Caledonia. Lectotype designation. Franz (1971) mentioned two specimens included in the type series of Euconnus tindoui Franz, 1971, but did not fix one as holotype. Consequently, the two specimens preserved at NHMW are syntypes. The male (Fig. 1) with the aedeagus extracted (Figs 30 – 31) bearing the set of labels illustrated in Fig. 42, is here designated as the lectotype (des. P. Jałoszyński).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF93FFCA9A93F9BBFAF83C8D.taxon	diagnosis	Revised diagnosis. Antennae gradually thickened; head rounded, with long tempora and vertex not bulging posterodorsally; submentum lacking lateral sutures; hypomeral ridges reaching or nearly reaching posterior tentorial pits; pronotum broadest in front of middle, with sinuate lateral margins, indistinct anterior corners and distinct but obtuse-angled posterior corners; pronotal base with paired pits or transverse groove, often with median carina; basisternal part of prosternum only slightly longer than coxal part; procoxal cavities broadly open; notosternal sutures and hypomeral ridges nearly complete (obliterated in their posterior and anterior portions, respectively, but the obliterated part is so short that it can be seen only under SEM in cleaned specimens or in transparent mounts under compound microscope); interprocoxal region with faint and diffuse carina; head and prothorax with bristles; mesoventral intercoxal process narrowly subtriangular or carinate, developed in anterior portion of mesoventrite and reaching only to about middle of mesocoxal cavities, with distinctly defined tip; metaventrite with anterior metaventral process between or slightly behind posterior margins of mesocoxal cavities, and with subtriangular, short intermetacoxal process slightly notched at middle and not separating metacoxae; elytral base with one or two small asetose foveae; aedeagus weakly sclerotized, with symmetrical median lobe and asymmetrical endophallic structures; parameres free and slender.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF93FFCA9A93F9BBFAF83C8D.taxon	description	Redescription. General body shape (Fig. 5) strongly elongate and moderately convex, body deeply constricted between head and pronotum and between pronotum and elytra. Head (Figs 18 – 19) large in relation to pronotum; divided by occipital constriction into large anterior and small posterior part (' neck' region), posterior part retracted into pronotum. Occipital constriction about as broad as half width of head. Anterior part of head capsule convex dorsally and flattened ventrally, rounded; tempora longer than compound eyes; vertex transverse and not bulging posterodorsally, with posterior margin not demarcated, anteriorly vertex confluent with subtrapezoidal frons; supraantennal tubercles feebly marked; frontoclypeal groove distinct, deep. Gular plate (Fig. 19; gp) broad and with distinct gular sutures; posterior tentorial pits (Fig. 19; ptp) strongly elongate, slot-shaped and located far in front of transverse impression separating ' neck' region from anterior part of head. Tempora, genae and postgenae with thick bristles. Submentum (Fig. 19; smn) subtriangular; hypostomal ridges (Fig. 19; hr) nearly complete, behind posteromesal margin of each cardo nearly straight, not connected to but reaching posterior tentorial pits; mentum (Fig. 19; mn) subtrapezoidal; prementum small and transverse, largely membranous; labial palps short; maxillae unmodified; mandibles only partly visible in the studied specimen, subtriangular and evenly curved, mesal margins not visible. Antennae (Fig. 5) long and slender, gradually but weakly thickened. Prothorax (Figs 18, 20) convex, elongate, broadest in front of middle, with sides strongly rounded in anterior half and slightly concave behind the broadest site; anterior corners indistinct, posterior corners well-marked, obtuse-angled; base with short sublateral carinae (Fig. 18; slc), two pairs of antebasal pits (Fig. 18; abp) and short median antebasal carina (Fig. 18; mc). Prosternum (Fig. 20) with basisternal part (Fig. 20; bst) only slightly shorter than coxal part; notosternal sutures (Fig. 20; nss) and hypomeral ridges (Fig. 20; hyr) nearly complete; procoxal cavities open; interprocoxal region with faint carina; hypomera covered with bristles. Mesothorax. Mesoscutellum in intact specimens entirely covered by posterior pronotal margin, its posterior tip not exceeding posterior margin of elytral articulating lobe. Mesoventrite (Fig. 21) with a pair of transverse setose impressions functioning as procoxal rests (Fig. 21; pcr) in anterior region, impressions separated at middle and with diffuse posterior margins; mesoventral intercoxal process (Fig. 21; msvp) carinate, its posterior tip welldefined, subtriangular and located near middle of mesocoxal cavities. Metathorax. Metaventrite (Fig. 21) much longer than mesoventrite, subtrapezoidal; anterior metaventral process (Fig. 21; amvp) present; intermetacoxal area weakly expanded posteriorly and forming short subtriangular metaventral intercoxal process (Fig. 13; mtvp) indistinctly notched at middle and not separating metacoxae. Elytra (Figs 5, 18) oval, with rounded apices and narrow base which is subequal in width to posterior pronotal margin; humeral calli and basal impressions distinct; each elytron with two small asetose basal foveae (Fig. 18; bef). Metathoracic wings present. Legs (Fig. 5) moderately long and slender, unmodified. Abdomen elongate, gradually narrowing posteriorly, suture between two terminal visible sternites (i. e., VII and VIII) less distinct than between remaining sternites. Aedeagus (Figs 34 – 35) with symmetrical and weakly sclerotized median lobe; endophallic structures asymmetrical, parameres slender, with apical setae.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF93FFCA9A93F9BBFAF83C8D.taxon	discussion	Remarks. Magellanoconnus clearly differs from Sciacharis in open procoxal cavities, incomplete mesoventral intercoxal process not reaching posterior margins of mesocoxal cavities and in the presence of the anterior mesoventral process. The contiguous metacoxae differentiate this genus from Euconnus. Here the status of Magellanoconnus is restituted to the concept of Kuschel, 1990: 76, as a separate genus. Composition and distribution. Magellanoconnus currently includes 78 valid species-group names; 76 species and subspecies of Magellanoconnus s. str. occur in South Africa and Zimbabwe, Argentina, Bolivia, Chile, Colombia and Ecuador, and in New Zealand; two species of the other subgenus are treated in the next section.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF9FFFC89A93FF18FEAD3FD5.taxon	discussion	Remarks. Morphological characters of Anthicimimus were illustrated and described by Jałoszyński (2015 d, 2017 e). Open procoxal cavities were used as a primary character to distinguish Anthicimimus (Fig. 6) from Sciacharis (Figs 3 – 4) and similar genera (including Euconnus). As demonstrated here, the type species of Magellanoconnus (Fig. 5) has open procoxal cavities and it differs from the two species currently included in Anthicimimus only in minor characters of the pronotal and elytral base. These structures are insufficient to treat these taxa as belonging to separate genera, and therefore Anthicimimus is placed in Magellanoconnus as a subgenus. Magellanoconnus s. str. and Anthicimimus can be distinguished on the basis of the following diagnostic characters:	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF9FFFCE9A93FBFCFCF23A25.taxon	diagnosis	Revised diagnosis. Antennae gradually thickened; head subpentagonal with long tempora and occipital constriction slightly broader than half width of head; frontoclypeal groove indistinct; submentum lacking lateral sutures; hypostomal ridges not reaching posterior tentorial pits; pronotum with complete or nearly complete lateral carinae and transverse antebasal groove interrupted at middle; basisternal part of prosternum much shorter than coxal part; notosternal sutures complete; hypomeral ridges marked only posteriorly, but adcoxal part of hypomeron distinctly demarcated as asetose area contrasting with densely setose outer part of hypomeron; procoxal cavities broadly closed; interprocoxal region with faint trace of median carina; head and pronotum with bristles; each elytron with one barely discernible and asetose rudiment of basal fovea; mesoventral intercoxal process carinate but incomplete, separating mesocoxae only in their anterior half; metaventral intermetacoxal process subtriangular and not separating metacoxae; aedeagus with symmetrical and weakly sclerotized median lobe; endophallus asymmetrical; parameres free and slender.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF9FFFCE9A93FBFCFCF23A25.taxon	description	Redescription. General body shape (Fig. 7) elongate and moderately convex, pronotum flattened, body deeply constricted between head and pronotum and between pronotum and elytra. Head (Figs 22 – 23) large in relation to pronotum; divided by occipital constriction into large anterior and small posterior part (' neck' region), posterior part retracted into pronotum. Occipital constriction about as broad as half width of head. Anterior part of head capsule convex dorsally and flattened ventrally, subpentagonal; tempora longer than compound eyes; vertex transverse and not bulging posterodorsally, with posterior margin not demarcated, anteriorly vertex confluent with subtrapezoidal frons; supraantennal tubercles feebly marked; frontoclypeal groove indistinct. Gular plate (Fig. 23; gp) broad and with distinct gular sutures; posterior tentorial pits (Fig. 23; ptp) strongly elongate, slot-shaped and located in front of transverse impression separating ' neck' region from anterior part of head. Tempora, genae and postgenae with thick bristles. Submentum (Fig. 23; smn) subtriangular; hypostomal ridges (Fig. 23; hr) incomplete, nearly straight, not reaching posterior tentorial pits; mentum (Fig. 23; mn) subtrapezoidal; prementum small and transverse, largely membranous; labial palps short; maxillae unmodified; mandibles only partly visible in the studied specimen, subtriangular and evenly curved, mesal margins not visible. Antennae (Fig. 7) long and slender, gradually but weakly thickened. Prothorax (Figs 22, 24) flattened, elongate, broadest in front of middle, with sides weakly rounded in anterior half and slightly constricted in front of base; anterior corners indistinct, posterior corners well-marked, obtuseangled; base lacking sublateral carinae or pits, with transverse groove (Fig. 22; abg) interrupted at middle; sides of pronotum with nearly complete lateral carinae (Fig. 22; lc) or edges. Prosternum (Fig. 24) with basisternal part (Fig. 24; bst) much shorter than coxal part; notosternal sutures (Fig. 24; nss) complete; hypomeral ridges (Fig. 24; hyr) marked only posteriorly, but entire adcoxal part of hypomeron differentiated by its asetose surface, contrasting with densely setose outer part of hypomeron; procoxal cavities broadly closed; interprocoxal region with barely discernible trace of narrow carina; hypomera covered with bristles. Mesothorax. Mesoscutellum in intact specimens entirely covered by posterior pronotal margin, its posterior tip not exceeding posterior margin of elytral articulating lobe. Mesoventrite (Fig. 25) with a pair of transverse setose impressions functioning as procoxal rests (Fig. 21; pcr) in anterior region, impressions separated at middle and with all margins diffuse; mesoventral intercoxal process (Fig. 25; msvp) carinate, its posterior tip well-defined, subtriangular and located near middle of mesocoxal cavities. Metathorax. Metaventrite (Fig. 25) much longer than mesoventrite, subtrapezoidal; anterior metaventral process (Fig. 25; amvp) present; intermetacoxal area weakly expanded posteriorly and forming short subtriangular metaventral intercoxal process (Fig. 25; mtvp) not separating metacoxae. Elytra (Figs 7, 22) oval, with rounded apices and base distinctly broader than posterior pronotal margin; humeral calli and basal impressions distinct; each elytron with one vestigial asetose basal fovea (Fig. 22; bef) barely discernible in intact specimens. Metathoracic wings present. Legs (Fig. 7) moderately long and slender, unmodified. Abdomen elongate, gradually narrowing posteriorly, suture between two terminal visible sternites (i. e., VII and VIII) less distinct than between remaining sternites. Aedeagus (Figs 36 – 39) with symmetrical and weakly sclerotized median lobe; endophallic structures asymmetrical, parameres slender, with apical setae.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF9FFFCE9A93FBFCFCF23A25.taxon	discussion	Remarks. Neuraphoconnus clearly differs from Sciacharis in the incomplete mesoventral intercoxal process not reaching posterior margins of mesocoxal cavities and in the presence of the anterior mesoventral process. These characters are similar to those in Australian Spinosciacharis Jałoszyński, 2014 c; the latter differs from Neuraphoconnus in the pronotum lacking lateral carinae; the basisternal part of prosternum subequal in length to the coxal part; the hypomeral ridges complete; and each elytron with two deep and distinct foveae. Composition and distribution. Neuraphoconnus currently includes 16 valid species names; the genus is distributed in New Caledonia and New Zealand.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF9FFFCE9A93FBFCFCF23A25.taxon	description	Lectotype designation. Franz (1971) mentioned 17 specimens in his original description of Neuraphoconnus caledonicus Franz, 1971, but he did not fix one of them as holotype. Consequently, all specimens are syntypes. The male preserved at NHMW (Fig. 7) with the aedeagus extracted (Figs 38 – 39), bearing the set of labels illustrated in Fig. 45, is here designated as the lectotype (des. P. Jałoszyński).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF99FFD29A93F92CFC863D81.taxon	diagnosis	Revised diagnosis. Microscydmus with the following combination of characters: head lacking bristles; submentum much longer than half length of mentum; sides of pronotum with conspicuous bristles; each elytron with two small asetose basal foveae; procoxal rests on mesoventrite filled with setae, their posterior margin marked by fine carina, area behind the carina not impressed.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF99FFD29A93F92CFC863D81.taxon	description	Redescription. General body shape (Fig. 8) elongate and moderately convex, pronotum flattened, body deeply constricted between head and pronotum and between pronotum and elytra. Head (Figs 26 – 27) small in relation to pronotum; divided by occipital constriction into large anterior and small posterior part (' neck' region), posterior part retracted into pronotum. Occipital constriction slightly broader than half width of head. Anterior part of head capsule convex dorsally and flattened ventrally, subtrapezoidal; tempora slightly longer than compound eyes; vertex transverse and not bulging posterodorsally, posteriorly demarcated by stepwise rounded ridge, anteriorly vertex confluent with subtrapezoidal frons; supraantennal tubercles distinct but weakly elevated; frontoclypeal groove absent. Gular plate (Fig. 27; gp) broad and with indistinct gular sutures; posterior tentorial pits (Fig. 27; ptp) nearly circular and located slightly in front of deep and well-defined transverse impression separating ' neck' region from anterior part of head. Tempora, genae and postgenae lacking bristles. Submentum (Fig. 27; smn) subtrapezoidal, relatively long, much longer than half length of mentum; hypostomal ridges (Fig. 27; hr) incomplete, marked only just behind cardines, not reaching middle between each cardo and posterior tentorial pit; mentum (Fig. 27; mn) subtrapezoidal; prementum small and transverse, largely membranous; labial palps short; maxillae unmodified; mandibles only partly visible in the studied specimen, subtriangular and evenly curved, mesal margins not visible. Antennae (Fig. 8) long and slender, antennomeres IX – XI forming only slightly broadened club. Prothorax (Figs 26, 28) flattened, round, broadest in front of middle, with sides strongly rounded in anterior half and weakly rounded behind the widest site; anterior corners indistinct, posterior corners well-marked, obtuseangled; base with sublateral carinae (Fig. 26; slc) and two pairs of antebasal pits (Fig. 26; abp), of which inner pair connected by transverse groove. Prosternum (Fig. 28) with basisternal part (Fig. 28; bst) distinctly shorter than coxal part; notosternal sutures (Fig. 28; nss) incomplete, marked at the anterior margin of prothorax but internalized at middle, again visible posteriorly, in anterolateral margin of each procoxal cavity, as arcuate suture curved anterolaterally; hypomeral ridges (Fig. 28; hyr) complete, sinuate; procoxal cavities broadly closed; interprocoxal region with narrow, weakly elevated and diffuse carina; hypomera covered with bristles. Mesothorax. Mesoscutellum in intact specimens entirely covered by posterior pronotal margin, its posterior tip not exceeding posterior margin of elytral articulating lobe. Mesoventrite (Fig. 29) with a pair of transverse setose impressions functioning as procoxal rests (Fig. 29; pcr) in anterior region, impressions separated at middle and posteriorly demarcated by fine carina, area behind carina not impressed; mesoventral intercoxal process (Fig. 29; msvp) carinate, posteriorly fused with metaventrite. Metathorax. Metaventrite (Fig. 29) much longer than mesoventrite, subtrapezoidal; intermetacoxal area forming metaventral intercoxal process (Fig. 29; mtvp) composed of two long spinces separated by narrow notch. Elytra (Figs 8, 26) oval, with rounded apices and base nearly as wide as posterior pronotal margin; humeral calli and basal impressions distinct; each elytron with two very small asetose basal foveae (Fig. 26; bef). Metathoracic wings present. Legs (Fig. 8) moderately long and slender, unmodified (including protibial apices in male). Abdomen elongate, gradually narrowing posteriorly, suture between two terminal visible sternites (i. e., VII and VIII) less distinct than between remaining sternites. Aedeagus (Figs 40 – 41) stout, with symmetrical and weakly sclerotized median lobe; endophallic structures with asymmetrical distal sclerites, parameres slender, with apical setae.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF99FFD29A93F92CFC863D81.taxon	discussion	Remarks. The type species of Valdivioconnus has numerous characters that do not occur in the type species of Sciacharis: trimerous antennal club; head with strongly shortened area behind the mentum, with circular posterior tentorial pits and deep, narrow groove demarcating ventrally the ' neck' region from anterior part of head capsule; hypostomal ridges obliterated shortly behind cardines; short and round pronotum with the inner pair of antebasal pits connected by transverse groove; the posterior portion of notosternal suture extending from prosternal cavity anteriorly and distinctly curved laterally to disappear near middle between cavity and hypomeral ridge; and the metaventral intermetacoxal process composed of two elongate spines separated at middle by a narrow notch. These characters do not occur also in Euconnus, but are diagnostic for Microscydmus Saulcy & Croissandeau, 1893. Euconnus stenichniformis is also very small (body length 1.03 mm), as are all species of Microscydmus. FIGURES 30 – 41. Aedeagus In ventral (30, 32, 34, 36, 38, 40) and lateral (31, 33, 35, 37, 39, 41) vIews. Euconnus (Austroconophron) tindoui Franz, comb. rest. (type specIes of Austroconophron), lectotype (30 – 31); Sciacharis (s. str.) lanosa (Broun) (type specIes of Allomaoria), lectotype (32 – 33); Magellanoconnus (s. str.) laurisilvae (Franz), comb. rest. (type specIes of Magellanoconnus), holotype (34 – 35); Neuraphoconnus caledonicus Franz, comb. rest. (type specIes of Neuraphoconnus), lectotype (36 – 37) and paralectotype (38 – 39); Microscydmus (Valdivioconnus) stenichniformis (Franz), comb. n. (type specIes of Valdivioconnus), holotype (40 – 41).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF85FFD39A93FADAFAD23D80.taxon	materials_examined	Type species. Neuraphoconnus carinifrons Franz, 1975, here designated.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF85FFD39A93FADAFAD23D80.taxon	diagnosis	Diagnosis. Antennae gradually thickened; head subpentagonal with tempora longer than eyes; vertex not bulging posterodorsally; bristles present on head and prothorax; submentum lacking lateral sutures; hypomeral ridges nearly reaching posterior tentorial pits, the latter slightly elongate and located in front of transverse impression separating ' neck' region from anterior part of head capsule; occipital constriction slightly broader than half width of head; frontoclypeal groove absent; pronotum broadest in front of middle, with lateral margins concave in posterior third, with sharp lateral carinae in posterior half, long sublateral carinae, two pairs of antebasal pits (one located on outer margin of lateral carina, one in large submedian impression), and long median longitudinal carina; anterior and posterior pronotal corners well-defined but obtuse-angled; basisternal part of prosternum slightly shorter than coxal part; notosternal sutures complete; hypomeral ridges nearly complete (shortly disrupted in front of procoxal cavities); interprocoxal region lacking process or carina; procoxal cavities broadly closed; procoxal rests on mesoventrite short and broad, filled with setae, area behind them weakly convex; mesoventral intercoxal process carinate and completely separating mesocoxae, but only as long as mesocoxa; metaventral intermetacoxal process short but nearly as broad as 1 / 4 of metaventral width, with posterior margin concave; each elytron with two distinct, deep asetose foveae; aedeagus with weakly sclerotized, symmetrical median lobe and strongly sclerotized, strongly asymmetrical endophallic structures; parameres free, slender, with apical setae.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF85FFD39A93FADAFAD23D80.taxon	discussion	Remarks. The only Australian nominal species previously placed in Sciacharis (Magellanoconnus) was redescribed in detail by Jałoszyński (2014), who suggested that it should be removed from Sciacharis as possibly morphologically closer to Euconnus. Indeed, among other characters, the broadly separated metacoxae exclude this species from Sciacharis and are similar to the condition typical of Euconnus. However, this species differs from Euconnus in possessing sharp lateral pronotal carinae and a weakly elevated mesoventral intercoxal process, which is only as long as the mesocoxae and distant from the anterior ridge of the mesoventrite. Kangarooconnus carinifrons was originally placed in Neuraphoconnus (Franz 1975), from which it differs in the same characters as from Sciacharis. Later combined with Sciacharis (Magellanoconnus) as a result of the postulated synonymy between Neuraphoconnus and Magellanoconnus (Franz 1986 a), it certainly is not similar to the type species of Magellanoconnus, which has open procoxal cavities and contiguous metacoxae (among other differences).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF84FFD19A93FCD9FD073D4A.taxon	description	Magellanoconnus lifouensis (Franz); implied new stat. in Kuschel, 1990: 76.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF8BFFE39A93FEE1FCF03BCD.taxon	description	Euconnus (Maorinus) cilipes (Broun); Franz, 1980 a: 299.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF8BFFE39A93FEE1FCF03BCD.taxon	description	Euconnus (Maoria) erythonotus (Broun); Franz, 1975: 24 (misspelled).	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF8BFFE39A93FEE1FCF03BCD.taxon	description	Sciacharis (Maorinus) hunuae (Franz, 1980) NEW ZEALAND Euconnus (Maorinus) hunuae Franz, 1980: 291.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF8BFFE39A93FEE1FCF03BCD.taxon	description	Euconnus (Maorinus) monilifer (Broun); Franz, 1980 a: 296.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FF8BFFE39A93FEE1FCF03BCD.taxon	description	Euconnus (Maoria) setosus (Sharp); Franz, 1975: 35.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FFB6FFE59A93FF19FCB53BA9.taxon	description	Sciacharis (s. str.) angustatus (Broun); Franz 1980 a: 279. Sciacharis (Sciacharis) antennalis (Broun, 1893) NEW ZEALAND Phaganophana antennale Broun, 1893 a: 1064. Euconnus (Allomaoria) antennalis (Broun); Franz 1975: 99.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FFB6FFE59A93FF19FCB53BA9.taxon	description	Phagonophana fragilis (Broun); Lhoste, 1938: 101.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FFB6FFE59A93FF19FCB53BA9.taxon	description	Phaganophana munroi Broun, 1893 a: 1063. Synonymized by Kuschel, 1990: 76. Euconnus (Allomaoria) nelsonius Franz, 1975: 52. Synonymized by Kuschel, 1990: 76. Euconnus (Allomaoria) pseudolanosus Franz, 1975: 60. Synonymized by Kuschel, 1990: 76.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FFB6FFE59A93FF19FCB53BA9.taxon	description	Sciacharis (Sciacharis) sannio (Franz); Maddison, 2010: 418.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
0388A953FFB6FFE59A93FF19FCB53BA9.taxon	description	Sciacharis (Sciacharis) sinuata (Broun); Franz, 1980 a: 280.	en	Jałoszyński, Paweł, Newton, Alfred F. (2017): Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4344 (3): 401-443, DOI: 10.11646/zootaxa.4344.3.1
