identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A22887D1561BDA55FF57F8F0FCEBF2B8.text	A22887D1561BDA55FF57F8F0FCEBF2B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus ecolab Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus ecolab sp. n.</p>
            <p>(Figs. 2a–e, 11a, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, Itabirito, Córrego prox. a  Fazenda Velha , long - 43.8091, lat -20.2991, 862 m a.s.l.,rock-face seepages 20.vi.2019, leg. J.B. R.Alencar” (INPA,  Coleoptera , in alcohol). </p>
            <p>Diagnosis. Medium-sized species. Pronotum with rounded posterolateral margins. Elytra surface even, with rows of fine systematic punctures bearing short setae, interrupted in some regions along their length; lateral rows very indistinct, blending with surrounding ground punctation. Elytral sutural interval not raised. Prosternum with two anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites, in strong contrast to yellow to light brown legs and pseudoepipleura.</p>
            <p>Adult. Body length 4.2 mm (n= 1); TL/GW 1.76 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 2a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending ca. 4/5 along lateral margin. Elytra with irregular patches of bronze or green iridescence; with row of rounded black [=without sheen] spots along suture. Each elytron with two small circular pale spots: anterior pale spot on distal third, about twice as far from suture than posterior spot; posterior spot situated on distal quarter. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum yellowish brown; pseudoepipleura and epipleura yellowish brown. Thorax dark brown to black ventrally. Legs light brown to yellow, except for dark brown coxae and base of femora. Abdominal ventrites dark brown, with slightly lighter posterior margins on ventrites 1–4, sometimes with pale patches laterally on ventrites 2–3.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 2d). Ground punctation on labrum denser than on clypeus, distance between punctures 0.5– 1.5× width of one puncture; with few distinct systematic punctures medially. Labrum bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 2a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming irregular field. Lateral margins of pronotum with fine setae distributed along its length, being more densely pubescent in anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra not raised. Elytral interval with weak or non-existent pubescence in ground punctation (e.g. Figs. 12a, c, d). Elytra with five rows of systematic punctures: rows 1 and 2 each represented by punctures not larger that surrounding punctuation and at about same spacing; rows 3–5 with punctures not larger than surrounding ground punctuation and as such hardly distinguishable. Elytral surface without any depression. Humeral region (in frontal view) without antero-medial elevation. Elytral margins with short spine-like setae equally spaced along its length. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 2e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 3–4 coarse spine-like setae. Metaventrite with oval glabrous area posteromedially, slightly longer than wide; length of glabrous area about two-thirds length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with 6–7 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with setae of varying lengths; ventrites 1–4 with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 2b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, sinuous, gradually tapering towards apex (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at fourfifths length of median lobe. Basal piece rounded (Fig. 11a).</p>
            <p> Etymology. The specific epithet  ecolab is derived from the acronym of the Community Ecology Lab of the Universidade Federal do Amazonas, in Brazil. It is a tribute to all the support received for the development of this work. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species shares several diagnostic features with  O. caparao Clarkson &amp; Short, 2012 : non-pale abdominal ventrites, elytral suture not raised, elytra surface with rows of systematic punctures bearing short setae. However,  O. caparao differs in that its elytral surface has faint, very shallowly impressed longitudinal grooves near the suture and the prosternum usually set with six spine-like setae, while longitudinal grooves are absent and the prosternum bears two spine-like setae in  O. ecolab sp. n.</p>
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	https://treatment.plazi.org/id/A22887D1561BDA55FF57F8F0FCEBF2B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D1561EDA51FF57FF3EFAB6F09D.text	A22887D1561EDA51FF57FF3EFAB6F09D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus espinhacu Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus espinhacu sp. n.</p>
            <p>(Figs. 3a–e, 11b, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS,  Parque Estadual Grão Mogol , stream near the archaeological site, long -42.943556 lat -16.618111, 719 m a.s.l., rock-face seepages, 29.iv.2019, leg. J.B. R. Alencar” (INPA,  Coleoptera , in alcohol)  .   Paratypes (339 exs.): BRAZIL: Minas Gerias: Same data as holotype (24 exs.  INPA ; 10 exs. SEMC, in alcohol); except: “Waterfall of Mirante, long -42.899694, lat -16.576639, 821 m a.s.l., rock-face seepages, 27.iv.2019, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol); “ Waterfall da Bocaina, long -42.837083, lat -16.301167, 669 ma.s.l., rock-face seepages, 28.iv.2019, leg. J.B. R. Alencar” (21 exs.  INPA , in alcohol); “  Waterfall of Almas , long -42.931194, lat -16.618139, 691 m a.s.l., rock-face seepages, 29.iv.2019, leg. J.B. R. Alencar” (16 exs.  INPA , in alcohol); “  Waterfall of Coronhas , long -42.885194 lat -16.522417, 792 m a.s.l., rock-face seepages, 30.iv.2019, leg. J.B. R. Alencar” (71 exs.  INPA , in alcohol); “ PARNA Sempre Vivas,  Waterfall of Felipes , long -43.703639, lat -17.907472, 1090 m a.s.l., rock-face seepages, 24.iv.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Stream southwest of camp, long -43.773861, lat -17.924444, 1259 m a.s.l., rock-face seepages, 23.iv.2019, leg. J.B. R. Alencar” (15 exs.  INPA , in alcohol); “Curimataí,  Waterfall of Rio Curimataí , long -43.946806, lat -17.87825, 625 m a.s.l., vertical surfaces with water, 23.vi.2019, leg. J.B. R. Alencar” (9 exs.  INPA , in alcohol); “ PARNA  Serra do Cipó ,  Waterfall of Andorinhas , long -43.491472, lat -19.237611, 1028 m a.s.l., vertical surfaces with water, 10.iv.2019, leg. J.B. R. Alencar” (24 exs.  INPA , in alcohol); “  Waterfall of Tombador , long -43.539833, lat -19.335639, 867 m a.s.l., vertical surfaces with water, 12.iv.2019, leg. J.B. R. Alencar” (24 exs.  INPA , in alcohol); “  Waterfall of Gavião , long -43.555806, lat -19.335639, 844 m a.s.l., vertical surfaces with water, 12.iv.2019, leg. J.B. R. Alencar” (24 exs.  INPA , in alcohol); “  Conceição do Mato Dentro ,  Waterfall of Três Barras , long -43.539833, lat -19.335639, 701 m a.s.l., rock-face seepages, 21.vi.2019, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); “  Santo Antônio do Itambé ,  Rio Guanhães , long -43.307167, lat -18.454611, 731 m a.s.l., margin of  Preto river , 21.vi.2019, leg. J.B. R. Alencar” (31 exs.  INPA , in alcohol); “  Santo Antônio do Itambé ,  Rio Preto , long -43.320778, lat -18.461417, 702 m a.s.l., vertical surfaces with water, 30.iii.2020, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Serro,  Waterfall of Amaral , long -43.406944, lat -18.388528, 1043 m a.s.l., vertical surfaces with water, 22.vi.2019, leg. J.B. R. Alencar” (17 exs.  INPA , in alcohol); “Itacambira,  Waterfall of Curiango , long - 43.324861, lat -17.147556, 1051 m a.s.l., vertical surfaces with water, 24.vi.2019, leg. J.B. R. Alencar” (6 exs.  INPA , in alcohol); “Mato Verde, Waterfall Maria Rosa, long -42.795444, lat -15.432167, 725 m a.s.l., vertical surfaces with water and green algae, 25.iii.2020, leg. J.B. R. Alencar” (19 exs.  INPA , in alcohol); “  Serranópolis de Minas ,  Waterfall of Serrado , long -42.843986, lat -15.677725, 720 m a.s.l., spray zones of waterfalls, 25.iii.2020, leg. J.B. R. Alencar” (12 exs.  INPA , in alcohol); “Botumirim, Waterfall Quatro Oitavas, long -43.014047, lat -16.883492, 862 m a.s.l., vertical surfaces with water, 27.iii.2020, leg. J.B. R. Alencar” (6 exs.  INPA , in alcohol); “Botumirim,  Waterfall of Bananal , long -43.037042, lat -16.839917, 908 m a.s.l., vertical surfaces with water, 28.iii.2020, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); “Serro,  Waterfall of Malheiros , long -43.445917, lat -18.654139, 738 m a.s.l., tiver margin, 31.iii.2020, leg. J.B. R. Alencar ” (9 exs.  INPA , in alcohol)  . </p>
            <p>Diagnosis. Large-sized species. Pronotum with rounded posterolateral margins. Elytra surface with shallowly impressed longitudinal grooves, with sparse short setae. Elytra with sutural interval raised (Fig. 12d) and margins explanate (Fig. 13b). Prosternum with 4–5 anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites, in strong contrast to legs and pseudoepipleura, which are from yellow to light brown.</p>
            <p>Adult. Body length 5.0–6.0 mm (n= 10, mean= 5.72, SD= ±0.33); TL/GW 1.65± 0.09 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 3a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. four fifths of lateral margin length. Elytra with irregular patches of bronze or green iridescence; with row of rounded black [=without sheen] spots along suture. Each elytron with rounded pale spot in distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum; pseudoepipleura and epipleura yellowish brown. Thorax dark brown to black ventrally. Legs light brown to yellow, except for dark brown coxae and basal region of femora. Abdominal ventrites dark brown, with slightly lighter posterior margins on ventrites 1–4, sometimes with pale patches laterally on ventrites 2–3.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 3d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× twidth of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 3a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming irregular field. Lateral margins of pronotum with fine setae distributed along its length, being more densely pubescent on anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra slightly raised along its length. Elytral interval with weak or non-existent pubescence on ground punctation (e.g. Figs. 12a, c, d). Rows of systematic punctures on elytra present but blending with surrounding fine ground punctation and as such hardly distinguishable (e.g. Fig. 12d). Elytral surface with shallow longitudinal depressions medially in posterior half. Humeral region (in frontal view) without antero-medial elevation. Elytral margins with short spine-like setae situated along its length and explanate on posterior half. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 3e); elevated region with 4–5 spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 6–7 coarse spine-like setae. Metaventrite with posteromedial oval glabrous area ca. twice as long as wide; length of glabrous area about two-thirds length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with 9–12 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with scattered, sparsely distributed setae of varying lengths; ventrites 1–4 with setae shorter to subequal than longest setae around margin of metasternal glabrous area (Fig. 3b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at 4/5 length of median lobe. Basal piece rounded (Fig. 11b).</p>
            <p> Etymology. The specific epithet  espinhacu is derived from “Serra do Espinhaço ”, the name of the Brazilian mountain range where this species is widely distributed. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species shares diagnostic features with  O. itapecum Clarkson &amp; Short, 2012 and  O. fryanus Balfour-Browne, 1939 : non-pale abdominal ventrites, elytral suture raised and four prosternal spines. However, in  O. itapecum , the body shape is elongated (TL/GW =1.80), the elytra are smooth, without undulating grooves near the suture interval, and the legs are light brown to dark brown, while in  O. espinhacu sp. n. the body is less elongated (TL/GW = 1.65), the elytra have undulating grooves near the suture interval and the legs are paler in color (light brown to yellow).  O. fryanus , for its part, has multiple pale elytral spots, a more distinctive lateral depression on humeral region and a distinct medial indentation on the elytra easily distinguish it from  O. espinhacu sp. n.</p>
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	https://treatment.plazi.org/id/A22887D1561EDA51FF57FF3EFAB6F09D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D1561CDA5FFF57FA47FAEAF3BD.text	A22887D1561CDA5FFF57FA47FAEAF3BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus giganteus Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus giganteus sp. n.</p>
            <p>(Figs. 4a–e, 11c, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: BAHIA, Abaíra,  Waterfall Guarda Mor , long -41.8606, lat -13.3127, 1272 m a.s.l., vertical surfaces with water, 13.iii.2020, leg. J.B. R. Alencar” (INPA,  Coleoptera , in alcohol)  .   Paratypes (173 exs.): BRAZIL: Bahia:  Same data as holotype (94 exs.  INPA ; 10 exs. SEMC, in alcohol); except: “BA-564 Piatã-Mucugê, near the road, long -41.5931, lat -13.1546, 844 m a.s.l., vertical surfaces with water and green algae, 26.vi.2019, leg. J.B. R. Alencar” (31 exs.  INPA , in alcohol); “Rio de Contas, Waterfall of Fraga, long -41.5931, lat -13.1546, 844 m a.s.l., vertical surfaces with water, 23.iii.2020, leg. J.B. R. Alencar” (38 exs.  INPA , in alcohol)  . </p>
            <p>Diagnosis. Large-sized species. Pronotum with angulate posterolateral margins. Elytra surface even, with very fine ground punctation, with faint rows of systematic punctures (some of these punctures can be detected by presence of setae). Elytral sutural interval raised (e.g. Fig. 12d) margins explanate (e.g. Fig. 13b). Prosternum with two anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 5.7–6.3 mm (n= 10, mean= 6.02, SD= ±0.15); TL/GW 1.60± 0.04 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 4a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending slightly beyond the half of margin length. Elytra with irregular patches of bronze or green iridescence; with row of rounded black [=without sheen] spots along suture. Each elytron with rounded pale spot on distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum; pseudoepipleura and epipleura yellowish brown. Thorax dark brown to black ventrally. Legs dark brown, except for light brown tarsi. Abdominal ventrites dark brown with posterior margin of each ventrite light to brown.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 1.5–2.0× width of one puncture (Fig. 4d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; sometimes bearing short setae. Frons with irregular row of systematic punctures toward median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 4a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming irregular field. Lateral margins of pronotum with fine setae distributed along of its length, being more densely pubescent on anterior half. Pronotal posterolateral angles angulate. Sutural interval on elytra slightly raised along its length. Elytral interval with weak or non-existent pubescence in ground punctation (e.g. Fig 12a, c, d). Rows of systematic punctures on elytra present but not larger than surrounding ground punctuation and as such hardly distinguishable, blending uniformly with ground punctuation. Elytral surface with shallow longitudinal depressions medially on posterior half. Humeral region (in frontal view) without antero-medial elevation. Elytral margins with short spine-like setae situated along its length and explanate on posterior half. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 4e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 6–7 coarse spine-like setae. Metaventrite with oval glabrous area posteromedially, slightly longer than wide; length of glabrous area about half length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal. Protibiae each with 6–7 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with setae of varying lengths; pubescence usually with suberect to recumbent setae; longest setae shorter to subequal in length to longest setae around margin of metasternal glabrous area (Fig. 4b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, straight, tapering from the distal half (dorsal view); median lobe subequal in length to the parameres, without hook in apex. Gonopore located at four-fifths the length of median lobe. Basal piece constricted (Fig. 11c).</p>
            <p> Etymology. The specific epithet  giganteus derived from giga- (G.), meaning giant, refers to the large size of the body. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Bahia) (Fig. 14).</p>
            <p> Remarks. This species shares several diagnostic features with  O. flexus Clarkson &amp; Short, 2012 : non-pale abdominal ventrites, elytral suture raised, and the presence of two prosternal spines. However, in  O. flexus the posterolateral margins of the pronotum are broadly rounded and shorter body length (4.4–5.2 mm), while the posterolateral margins of the pronotum are angulate and the body size varies from 5.7–6.3 mm in  O. giganteus sp. n.</p>
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	https://treatment.plazi.org/id/A22887D1561CDA5FFF57FA47FAEAF3BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15612DA5CFF57F8AAFEF3F17E.text	A22887D15612DA5CFF57F8AAFEF3F17E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus humboldti Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus humboldti sp. n.</p>
            <p>(Figs. 5a–e, 11d, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, Parque Estadual Serra do Cabral,  Waterfall of Demir , long -44.211975 lat -17.920439, 833 m a.s.l., rock-face seepages, 14.iv.2019, leg. J.B. R. Alencar” (  INPA , in alcohol)  .   Paratypes (266 exs.): BRAZIL: Minas Gerais: Same data as holotype (20 exs.  INPA ; 10 exs. SEMC, in alcohol); except: “  Waterfall of Boqueirão , long -44.178528, lat -17.758333, 680 m a.s.l., rock-face seepages  ,   13.iv.2019,leg. J.B. R.Alencar” (32 exs.  INPA , in alcohol); “  Waterfall of Fervedouro , long -44.247978,lat -17.917764, 1071 m a.s.l., rock-face seepages  ,   14.iv.2019, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol); “Curimataí,  Waterfall of Simão , long -43.955333, lat -17.896139, 650 m a.s.l., rock-face seepages  ,   16.iv.2019, leg. J.B. R. Alencar” (28 exs.  INPA , in alcohol); “ PARNA Sempre Vivas,  Waterfall of Felipes , long -43.703639, lat -17.907472, 1090 m a.s.l., rock-face seepages  ,   24.iv.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Parque Estadual Grão Mogol,  Waterfall of Almas , long -42.931194, lat -16.618139, 691 m a.s.l., rock-face seepages  ,   29.iv.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Parque Estadual Grão Mogol, Stream of the Dé, long -42.963833, lat -16.592361, 746 m a.s.l., rock-face seepages  ,   29.iv.2019, leg. J.B. R. Alencar” (10 exs.  INPA , in alcohol); “Parque Estadual Grão Mogol,  Waterfall of Almas , long -42.870111, lat -16.511139, 624 m a.s.l., rock-face seepages  ,   30.iv.2019, leg. J.B. R. Alencar” (27 exs.  INPA , in alcohol); “Lassance,  Waterfall of Palmeiras , long -44.504167, lat -17.834333, 520 m a.s.l., vertical surfaces with water  ,   1.v.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “ PARNA  Serra do Cipó , Waterfall Capão dos Palmitos, long -43.609556, lat -19.377472, 830 m a.s.l., rock-face seepage  ,   8.vi.2019, leg. J.B. R. Alencar” (9 exs.  INPA , in alcohol); “Waterfall Taioba, long -43.571806 lat -19.391806, 863 m a.s.l., vertical surfaces with water  ,   9.vi.2019, leg. J.B. R. Alencar” (7 exs.  INPA , in alcohol); “Canyon of the Bandeirinhas, long -43.562722, lat -19.423917, 880 m a.s.l., seepage habitat  ,   11.vi.2019, leg. J.B. R. Alencar” (8 exs.  INPA , in alcohol); “Itabirito, Corrego Prox a Fazenda Velha, long -43.809139, lat -20.299111, 862 m a.s.l., rock-face seepages  ,   20.vi.2019, leg. J.B. R. Alencar” (82 exs.  INPA , in alcohol); “Serro,  Waterfall of Amaral , long -43.406944, lat -18.388528, 1043 m a.s.l., vertical surfaces with water  ,   22.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Gameleiras,  Waterfall of Andorinhas , long -43.079419, lat -14.981272, 931 m a.s.l., vertical surfaces with water and green algae  ,   24.iii.2019, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); “Mato Verde, Waterfall Maria Rosa, long -42.795444, lat -15.432167, 725 m a.s.l., vertical surfaces with water and green algae  ,   25.iii.2020, leg. J.B. R. Alencar” (3 exs.  INPA , in alcohol); Bahia: “ BAHIA, BA-564 Piatã-Mucugê, near the road, long -41.593139, lat -13.154611, 844 m a.s.l., vertical surfaces with water and green algae  ,   26.vi.2019, leg. J.B. R. Alencar” (16 exs.  INPA , in alcohol)  . </p>
            <p>Diagnosis. Small to medium-sized species. Pronotum with rounded posterolateral margins. Elytra surface even, with rows of fine systematic punctures bearing short setae, interrupted in some regions along their length; lateral rows very indistinct, blending with surrounding ground punctation. Elytral sutural interval not raised (Figs. 12a–c). Prosternum with two anteromedial spine-like setae. Abdominal ventrites pale, all ventrites usually with distinct medial dark macula (varying in width).</p>
            <p>Adult. Body length 3.5–4.2 mm (n= 10, mean= 3.91, SD= ±0.24); TL/GW 1.65± 0.10 mm. General body shape broadly oval, strongly convex.</p>
            <p>Color. Dorsum black (Fig. 5a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with wide pale strip, extending to ca. 4/5 of lateral margin length. Elytra with irregular patches of bronze or green iridescence; without row of rounded black [=without sheen] spots along suture. Each elytron with rounded pale spot on distal quarter. Antennae with antennomeres 1–5 and 3-segmented club yellowish brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum black to dark brown; pseudoepipleura and epipleura yellowish brown. Thorax, ventrally brown to black, with lighter posterior margins in metaventrite, sometimes extending laterally along entire length of anepisternum. Legs light brown, except for dark brown coxae and basal region of femora. Abdominal ventrites brownish yellow, usually with distinct medial dark macula, sometimes completely covering ventrite.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 5d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; and sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 5a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming an irregular field. Lateral margins of pronotum with fine setae distributed along its length, being more densely pubescent on anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra not raised. Elytral interval composed of short setae forming inconspicuous rows. Five rows of systematic punctures: rows 1 and 2 each represented by regular series of punctures; punctures not larger that surrounding punctuation and at about the same spacing. Rows 3–5 with punctures not larger than surrounding ground punctuation and as such hardly distinguishable. Elytral surface without shallow longitudinal depressions medially on posterior half. Humeral region (in frontal view) without anteromedial elevation. Elytral margins with short spine-like setae situated along its length. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 5e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 5–6 coarse spine-like setae. Metaventrite with oval glabrous area posteromedially, slightly longer than wide; length of glabrous area about two-thirds of length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with eight spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with scattered, long setae; ventrites 1–4 with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 5b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, straight; inner margin, straight, gradually tapering towards apex (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at four-fifths of median lobe. Basal piece tapered (Fig. 11d).</p>
            <p> Etymology. The specific epithet  humboldti is in honor to Friedrich Wilhelm Heinrich Alexander von Humboldt, he was one of the pioneers to propose relationships between South American mountain environments and biodiversity. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species was found in rock-face seepages along streams, as well as in marginal areas that simulate infiltrations due to low flow. This species shares distinguishable diagnostic features with  O. rotundus Clarkson &amp; Short, 2012 such as pale abdominal ventrites (in most specimens), absence of black circular spots along the elytral suture and a nearly circular, strongly convex body. However, in  O. rotundus the median lobe is distinctly shorter than the parameres and it has a tapered basal piece in contrast to  O. humboldti sp. n. which has short setae on the elytral surface and the median lobe is slightly shorter than the length of the parameres, and the basal piece is rounded (Fig. 11d). </p>
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	https://treatment.plazi.org/id/A22887D15612DA5CFF57F8AAFEF3F17E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15611DA5AFF57FB67FBD6F031.text	A22887D15611DA5AFF57FB67FBD6F031.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus lacia Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus lacia sp. n.</p>
            <p>(Figs. 6a–e, 11e, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, Serro,  Waterfall of Amaral , long -43.4069, lat -18.3885, 1043 m a.s.l., vertical surfaces with water, 22.vi.2019, leg. J.B. R. Alencar” (INPA,  Coleoptera , in alcohol). </p>
            <p>Diagnosis. Large-sized species. Pronotum with rounded posterolateral margins. Elytra surface with shallowly impressed longitudinal grooves and sparse short setae. Elytral sutural interval raised (e.g. Fig. 12d). Prosternum with six anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 4.9 mm (n= 1); TL/GW 1.73 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 6a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. 4/5 of lateral margin length. Elytra with faint, often indistinct yellowish or bronze maculae, especially near the suture; with row of rounded black [=without sheen] spots along suture. Each elytron with two to three small circular pale spots: two anterior pale spot very close together in distal third, about twice as far from suture than posterior spot; posterior spot situated in distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum and pseudoepipleura yellowish brown; and epipleura dark brown to black. Thorax dark brown to black ventrally. Legs brown to dark brown, except for light brown tarsi. Abdominal ventrites dark brown, with slightly lighter posterior margins in ventrites 1–4, sometimes with pale patches laterally in ventrites 2–3.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 6d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; and sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 6a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming irregular field. Lateral margins of pronotum with fine setae distributed along of its length, being more densely pubescent on anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra slightly raised along its length. Elytral interval with weak or non-existent pubescence in ground punctation (e.g. Figs. 12a, c, d). Rows of systematic punctures on elytra present but blending with surrounding ground punctation and as such hardly distinguishable. Elytral surface without shallow longitudinal depressions medially on posterior half. Humeral region (in frontal view) without anteromedial elevation. Elytral margins with short spine-like setae situated along its length. Prosternum with clearly defined median carina; strongly elevated anteromedially (Fig. 6e); elevated region with six spine-like setae. Elevated process of mesoventrite wide and elongate, about three times as long as wide; slightly raised anteriorly; with five coarse spine-like setae. Metaventrite with posteromedial oval glabrous area ca. twice as long as wide; length of glabrous area about three-quarters length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with 7–9 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with setae of varying lengths; ventrites 1–4 rather sparsely pubescent with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 6b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, not curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at four-fifths of median lobe. Basal piece tapered (Fig. 11e).</p>
            <p> Etymology. The specific epithet  lacia derived from the acronym of the Laboratório de Citotaxonomia e Insetos Aquáticos, within the Instituto Nacional de Pesquisas da Amazônia, in Brazil. It is a tribute to all the support received for the development of this work. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species shares diagnostics feature with  O. fryanus , such as non-pale abdominal ventrites, large size (&gt;5.0 mm), and six prosternal spines. However,  O. fryanus has a lateral indentation on the elytra, the shape of the body is very elongated (TL/GW = 1.85) and does not have elytra grooves, in  O. lacia sp. n. a lateral elytral indentation is absent, body is more oval (TL/GW =1.73) and elytra have grooves. </p>
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	https://treatment.plazi.org/id/A22887D15611DA5AFF57FB67FBD6F031	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15617DA58FF57FA2BFDE0F375.text	A22887D15617DA58FF57FA2BFDE0F375.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus ovalis Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus ovalis sp. n.</p>
            <p>(Figs. 7a–e, 11f, 12c, 13a, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, Serro,  Waterfall of Malheiros , long -43.4459, lat -18.6541, 738 m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R. Alencar” (INPA,  Coleoptera , in alcohol)  .   Paratypes (14 exs.): BRAZIL: Minas Gerais:  Same data as holotype (10 exs.  INPA ; 4 exs. SEMC, in alcohol)  . </p>
            <p>Diagnosis. Medium-sized species. Pronotum with rounded posterolateral margins. Elytra surface even, with rows of fine systematic punctures bearing short setae, interrupted in some regions along their length; lateral rows very indistinct, blending with surrounding ground punctation. Elytral sutural interval not raised (Fig. 12c). Prosternum with two or four anteromedial spine-like setae. Abdominal ventrites brown to dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 4.0– 4.4 mm (n= 10, mean= 4.21, SD= ±0.21); TL/GW 1.55 ± 0.08 mm. General body shape broadly oval, strongly convex.</p>
            <p>Color. Dorsum black (Fig. 7a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. 4/5 of lateral margin length. Elytra with irregular patches of bronze or green iridescence; without row of rounded black [=without sheen] spots along suture. Each elytron with one rounded pale spot on distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum; pseudoepipleura and epipleura yellowish brown. Thorax, ventrally brown to black. Legs brown with bronze or green iridescence, except for light brown tarsi. Abdominal ventrites brown to dark brown.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0×width of one puncture (Fig. 7d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5×width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with few distinct systematic punctures along anterolateral margins, larger than surrounding punctation and sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps subequal in length to width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 7a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; anterior and posterior series each forming an irregular field. Lateral margins of pronotum with fine setae distributed along of its length, being more densely pubescent on anterior half. Pronotal posterolateral angles broadly rounded. Sutural interval on elytra not raised (Fig. 12c). Elytral interval with weak or non-existent pubescence in ground punctation. Five rows of systematic punctures: rows 1 and 2 each bearing one small recumbent setae; punctures not larger than surrounding punctuation and at about the same spacing. Rows 3–5 each represented by punctures not larger than surrounding ground punctuation; bearing short setae and as such hardly distinguishable (Fig. 12c). Elytral surface without shallow longitudinal depressions medially on posterior half (Fig. 13a). Humeral region (in frontal view) without anteromedial elevation. Elytral margins with short spine-like setae situated along their length. Prosternum with clearly defined median carina, slightly elevated anteromedially (Fig. 7e); elevated region with two or four spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 4–6 coarse spine-like setae. Metaventrite with posteromedial oval glabrous area ca. twice as long as wide; length of glabrous area about two-thirds length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with scattered, sparsely distributed setae of varying lengths; ventrites 1–4 with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 7b). Aedeagus: outer margin of parameres sinuous, gradually tapering to apex, not curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at four-fifths of median lobe. Basal piece tapered (Fig. 11f).</p>
            <p> Etymology. The specific epithet  ovalis (L.), refers to the "egg-shaped" body. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species shares distinguishable diagnostic features with  O. caparao and  O. ecolab sp. n. : nonpale abdominal ventrites, elytral suture not raised, elytra surface with rows of systematic punctures bearing short setae. However,  O. caparao differs from  O. ovalis sp. n. by elytral surface with very shallowly impressed longitudinal grooves near the suture and by the prosternum set with six spine-like setae, in most specimens.  O. ecolab sp. n. differs from  O. ovalis sp. n. by a row of rounded black [=without sheen] spots along elytral suture and two small circular pale spots on each elytron. </p>
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	https://treatment.plazi.org/id/A22887D15617DA58FF57FA2BFDE0F375	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15615DA47FF57F912FC0DF2DD.text	A22887D15615DA47FF57F912FC0DF2DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus sulcatus Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus sulcatus sp. n.</p>
            <p>(Figs. 8a–d, 11g, 12d, 13b, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: BAHIA, PARNA  Chapada Diamantina, Waterfall da Fumaça , long -41.456278, lat -12.600333, 1286 m a.s.l., vertical surfaces with water, 19.iii.2020, leg. J.B. R. Alencar” (  INPA , in alcohol)  .   Paratypes (41 exs.): BRAIL: Bahia: Same data as holotype (4 exs.  INPA , in alcohol); except “Waterfall of Mosquito, long -41.3705 lat -12.3706, 517 m a.s.l., vertical surfaces with water, 29.vi.2019, leg. J.B. R. Alencar” (3 exs.  INPA , in alcohol); “Waterfall da Matinha, long -41.3358 lat -13.0765 1021 m a.s.l., vertical surfaces with water, 21.iii.2020, leg. J.B. R. Alencar” (21 exs.  INPA ; 10 exs. SEMC, in alcohol); “ Palmeiras,  Waterfall of Riachinho ,   long -41.5157, lat -12.5721, 935 m a.s.l., vertical surfaces with water, 18.iii.2020, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol)  . </p>
            <p>Diagnosis. Large-sized species. Pronotum with rounded posterolateral margins. Elytra surface with impressed longitudinal grooves and shallow longitudinal depressions medially in posterior half, rows of systematic punctures present, but blending with surrounding coarse ground punctation and as such hardly distinguishable (Fig. 12d). Elytral sutural interval raised (Fig. 12d) and margins explanate (Fig. 13b). Prosternum with two anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 5.2–6.2 mm (n= 10, mean= 5.88, SD= ±0.31); TL/GW 1.60± 0.08 mm. General body shape broadly oval, strongly convex.</p>
            <p>Color. Dorsum black (Fig. 8a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. 4/5 of lateral margin length. Elytra with faint, often indistinct yellowish or bronze maculae, especially near suture; with row of rounded black [=without sheen] spots along suture. Each elytron with one rounded pale spot on distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum and pseudoepipleura yellowish brown; and epipleura brown. Thorax dark brown to black ventrally. Legs dark brown, except for light brown tarsi. Abdominal ventrites dark brown with posterior margin of each ventrite light to brown.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 8d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; and sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra unevenly distributed and moderately coarse; elytra with distinctly larger punctures and with shallowly impressed longitudinal grooves (Figs. 8a, c, d). Pronotal systematic punctures similar in size to surrounding ground punctures; anterior and posterior series each forming an irregular field. Lateral margins of pronotum with fine setae distributed along its length, more densely pubescent on anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra (Fig. 12d) slightly raised. Elytral interval with weak or non-existent pubescence in ground punctation. Rows of systematic punctures on elytra present but blending with surrounding coarse ground punctation and as such hardly distinguishable (Fig. 12d). Elytral surface with shallow longitudinal depressions medially on posterior half (Fig. 13b). Humeral region (in frontal view) distinctly elevated anteromedially. Elytra margins with short spine-like setae spaced along its length and explanate on posterior half. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 8e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with six coarse spine-like setae. Metaventrite with posteromedial oval glabrous area ca. twice as long as wide; length of glabrous area about three-quarters of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with 7–9 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with setae of varying lengths; ventrites 1–4 with scattered long erect setae, distinctly longer than longest setae on metaventrite (Fig. 8b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe subequal in length to parameres, without apical hook. Basal piece rounded (Fig. 11g).</p>
            <p> Etymology. The specific epithet  sulcatus (L.), refers to the grooved elytra. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Bahia) (Fig. 14).</p>
            <p> Remarks. This species shares some diagnostic features with  O. iguazu (Oliva, 1996) : such as non-pale abdominal ventrites, large-size (&gt;5.0 mm) and two prosternal spines. However,  O. sulcatus sp. n. can be easily distinguished by its impressed longitudinal grooves and shallow longitudinal depressions medially on the posterior part of elytra (Fig. 13b). In some specimens, the visualization of the diagnostic spots and green or bronze sheen (iridescence) depends on the view and lighting, sometimes they are not captured in the photographs, visible by moving the specimen or by changing the lighting system of the photographic equipment. </p>
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	https://treatment.plazi.org/id/A22887D15615DA47FF57F912FC0DF2DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15608DA43FF57FF3EFE84F681.text	A22887D15608DA43FF57FF3EFE84F681.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus thrixdiastematus Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus thrixdiastematus sp. n.</p>
            <p>(Figs. 8a–d, 11h, 12b, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, PARNA Serra do Cipó, Waterfall Capão dos Palmitos, long -43.6096, lat -19.3775, 830 m a.s.l., rock-face seepage, 8.vi.2019, leg. J.B. R.  Alencar ” (INPA,  Coleoptera , in alcohol)  .   Paratypes (30 exs.): BRAZIL: Minas Gerais: Same data as holotype (5 exs.  INPA , in alcohol); except: “Waterfall of Andorinhas, long -43.4915, lat -19.2376, 577 m a.s.l., rock-face seepages, 10.vi.2019, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol);”Canyon of the Bandeirinhas, long -43.5627, lat -19.4239, 880 m a.s.l., seepage habitat, 11.vi.2019, leg. J.B. R. Alencar” (3 exs.  INPA , in alcohol); “ PARNA Serra do Gandarela, Waterfall of Mingu, long -43.7596, lat -20.0622, 1006 m a.s.l., rock-face seepages, 15.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “  Parque Estadual Serra do Cabral ,  Waterfall of Demir , long -44.212 lat -17.9204, 833 m a.s.l., rock-face seepages, 14.iv.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “  Diamantina, Waterfall of Andorinhas, long -43.5949 lat -18.2309, 1168 m a.s.l., 11.iv.2019, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol); “Lassance,  Waterfall of Palmeiras , long -44.5042, lat -17.8343, 520 m a.s.l., vertical surfaces with water, 1.v.2019, leg. J.B. R. Alencar” (8 exs.  INPA , in alcohol); Goias: “Alto Paraíso de Goiás, Waterfall da muralha, long -47.7507, lat -14.2774, 957 m a.s.l., 2.VII.2019, leg. J.B. R. Alencar” (4 exs.  INPA ; 4 exs. SEMC, in alcohol)  . </p>
            <p>Diagnosis. Medium-sized species. Pronotum with rounded posterolateral margins. Elytra surface even, with rows of systematic punctures present, but merging with ground punctation, both (ground punctures and systematic punctures) bearing fine setae, appearing pubescent in dorsal view and each elytron with two small pale circular spots. Elytra sutural interval not raised (Fig. 12b). Prosternum with two anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 4.0– 4.7 mm (n= 10, mean= 4.55, SD= ±0.23); TL/GW 1.65± 0.09 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 9a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. 2/3 of lateral margin length. Elytra with irregular patches of bronze or green iridescence; with row of rounded black [=without sheen] spots along suture. Each elytron with two small circular pale spots: anterior spot on distal third, about twice as far from suture than posterior spot; posterior spot situated on distal quarter. Antennae yellowish brown. Maxillary palps and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum; pseudoepipleura and epipleura yellowish brown. Thorax dark brown to black ventrally. Legs light brown to yellow, except for dark brown coxae and basal region of femora. Abdominal ventrites dark brown, with slightly lighter posterior margins on ventrites1–4, sometimes with pale patches laterally on ventrites 2–3.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 9d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; and sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2; apical segment longer than penultimate segment. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra evenly distributed and moderately coarse (Figs. 9a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; the anterior and posterior series each forming an irregular field. Lateral margins of pronotum with fine setae distributed along its length, being more densely pubescent in the anterior half. Pronotal posterolateral angles rounded. Sutural interval on elytra not raised. Elytral interval with prominent pubescence on ground punctation (Fig. 12b). Rows of systematic punctures on elytra present but not larger than surrounding ground punctuation and as such hardly distinguishable, blending uniformly with ground punctuation. Elytral surface without shallow longitudinal depressions medially on posterior half. Humeral region (in frontal view) without anteromedial elevation. Elytral margins with short spinelike setae situated along their length. Prosternum with a clearly defined median carina; slightly elevated anteromedially (Fig. 9e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 5–7 coarse spine-like setae. Metaventrite with posteromedial oval glabrous area ca. twice as long as wide; length of glabrous area about three-quarters length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with 6–7 spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with scattered, sparsely distributed setae of varying lengths; ventrites 1–4 with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 9b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe shorter than parameres, without apical hook. Gonopore located at medial half of median lobe. Basal piece tapered (Fig. 11h).</p>
            <p> Etymology. The specific epithet  thrixdiastematus is a combination of the adjective thrix (G.), meaning “hairy” referring to the prominent pubescence on elytral intervals and the noun diastema (G.), meaning “space” or “interval” which refers to the location of pubescence on elytra. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Bahia, Minas Gerais and Goias) (Fig. 14).</p>
            <p> Remarks. This species shares with  O. ecolab sp. n. the non-pale abdominal ventrites, elytral surface with rows of systematic punctures bearing short setae, each elytron with two small circular spots and two prosternal spines. However, both species can be distinguished by the following characters:  O. ecolab sp. n. has the elytral interval with weak or non-existent pubescence in ground punctation; aedeagus with gonopore located at 4/5 the length on median lobe and the basal piece is rounded and short, while  O. thrixdiastematus sp. n. has the elytral interval with dense pubescence, aedeagus with gonopore located at half on median lobe and the basal piece is tapered. The aedeagus of  O. thrixdiastematus sp. n. is similar to that of  O. pilosus Jordão, Clarkon &amp; Ferreira-Jr, 2018 and co-occurs in at least some localities the same locality, but  O. pilosus is easily distinguishable by the absence of the two spines-like setae in prosternum. </p>
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	https://treatment.plazi.org/id/A22887D15608DA43FF57FF3EFE84F681	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D1560EDA40FF57FC7BFE08F2E4.text	A22887D1560EDA40FF57FC7BFE08F2E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus thysanus Alencar & Short & Hamada 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus thysanus sp. n.</p>
            <p>(Figs. 10a–e, 11i, 12a, 14)</p>
            <p> Type material.   Holotype (male): “ BRASIL: MINAS GERAIS, PARNA Serra do Cipó,  Waterfall da Farofa , long -43.5761, lat -19.3794, 866 m a.s.l., vertical surfaces with water, 8.vi.2019, leg. J.B. R. Alencar” (INPA,  Coleoptera , in alcohol)  .   Paratypes (10 exs.): BRAZIL: Minas Gerais: Same data as holotype (2 exs.  INPA , in alcohol); except: “ PARNA Serra do Gandarela, Waterfall of Mingu, long -43.7596, lat -20.0622, 1006 m a.s.l., rock-face seepages, 15.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA ; 1 ex. SEMC, in alcohol); “Parque Estadual Rola-Moça,  Waterfall of Sazona , long-43.9899, lat -20.0966, 1335 m a.s.l., rock-face seepages, 18.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA ; 1 ex. SEMC, in alcohol). Goias  :   “ GOIÁS: Alto Paraíso de Goiás,  Waterfall of Muralha , long -47.7507, lat -14.2774, 957 m a.s.l., 2.VII.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol)  . </p>
            <p>Diagnosis. Large-sized species. Pronotum with angulate posterolateral margins. Elytral surface even, with rows of fine systematic punctures, row 1 very distinctly impressed and bearing erect setae, interrupted in some regions along their length; lateral rows very indistinct, blending with surrounding ground punctation. Elytral sutural interval not raised (Fig. 12a). Prosternum with two anteromedial spine-like setae. Abdominal ventrites dark brown, not paler than thoracic ventrites.</p>
            <p>Adult. Body length 4.7–5.4 mm (n= 10, mean= 5.11, SD= ±0.22); TL/GW 1.60± 0.07 mm. General body shape broadly oval, slightly convex.</p>
            <p>Color. Dorsum black (Fig. 10a), covered with dense, irregular green or bronze sheen. Anterolateral margins of pronotum with pale strip, extending to ca. 4/5 of lateral margin length. Elytra with irregular patches of bronze or green iridescence; without row of rounded black [=without sheen] spots along suture. Each elytron with rounded pale spot on distal quarter. Antennomeres 1–5 yellowish brown, club dark brown. Maxillary and labial palps yellow. Ventral face of head dark brown to black. Lateral margins of prosternum black to dark brown; pseudoepipleura yellowish brown; epipleura brown. Thorax, ventrally brown to black. Legs brown to dark brown, except for the light brown tarsi. Abdominal ventrites dark brown with posterior margin of each ventrite light to brown.</p>
            <p>Head. Ground punctation on clypeus and frons moderately coarse, distance between punctures 2.0–3.0× width of one puncture (Fig. 10d). Ground punctation on labrum denser than punctation of clypeus, distance between punctures 0.5–1.5× width of one puncture; with few distinct systematic punctures medially. Labrum sometimes bearing short setae. Clypeus with several scattered indistinct systematic punctures laterally; sometimes bearing short setae. Frons with irregular row of systematic punctures toward the median plane of each eye and few fine setae around ocular suture. Maxillary palps shorter than width of labrum; segment 3 shorter than segment 2. Labial palps about two-thirds as long as width of mentum. Mentum smooth, with scattered coarse punctures; subquadrate, anterior margin slightly convex and depressed.</p>
            <p>Thorax. Ground punctation on pronotum and elytra unevenly distributed and moderately coarse; elytra punctures slightly larger (Figs. 10a, c, d). Pronotal systematic punctures with short fine setae, at least 2× size of surrounding ground punctures; the anterior and posterior series each forming one irregular field. Lateral margins of pronotum with fine setae distributed along its length, more densely pubescent on anterior half. Pronotal posterolateral angles angulate. Sutural interval on elytra (Fig. 12a) not raised. Elytral interval with weak or non-existent pubescence in ground punctation (Fig 12a). Five rows of systematic punctures: row 1 represented as nearly continuous row of densely distributed, small punctures bearing long, erect setae, (Fig. 12a). Rows 2–5 each represented by punctures only slightly larger than surrounding ground punctuation and hardly distinguishable, some set with short setae. Elytral surface with shallow longitudinal depressions medially on posterior half. Humeral region (in frontal view) without anteromedial elevation. Elytral margins with short spine-like setae situated along their length and explanate on posterior half. Prosternum with clearly defined median carina; slightly elevated anteromedially (Fig. 2e); elevated region with two spine-like setae. Elevated process of mesoventrite wide and elongate, about twice as long as wide; slightly raised anteriorly; with 5–6 coarse spine-like setae. Metaventrite with oval glabrous area posteromedially, slightly longer than wide; length of glabrous area about one-third length of metaventrite. Procoxae with fine short pubescence and set with coarse, short spine-like setae; grouped on basal half. Protibiae each with eight spine-like setae on dorsal face. Meso- and metafemora with scattered punctures bearing short thickened, spine-like setae.</p>
            <p>Abdomen. Ventrites covered with setae of varying lengths; ventrites 1–4 with scattered, long setae, distinctly longer than longest setae on metaventrite, ventrite 5 more densely pubescent (Fig. 10b). Aedeagus: outer margin of parameres weakly sinuous, gradually tapering to apex, curving inward; inner margin, straight, tapering from distal half (dorsal view); median lobe shorter than parameres, with hooked apex. Gonopore apically on median lobe. Basal piece rounded (Fig. 11i).</p>
            <p> Etymology. The specific epithet is derived from  thysanus = fringe (G) and refers to the first elytral row densely set with erected setae. To be treated as a noun in apposition. </p>
            <p>Distribution. Brazil (Minas Gerais) (Fig. 14).</p>
            <p> Remarks. This species shares several diagnostic features with  O. decorus (Kuwert, 1890) : non-pale abdominal ventrites, two prosternal spines, and elytra surface with distinct rows of systematic punctures bearing erect setae. However,  O. decorus has a weakly spinose posterolateral pronotal angle and brightly iridescent blue elytral maculae, while  O. thysanus sp. n. has the posterolateral margins of pronotum angulate, but not spinose and lacks brightly iridescent blue elytral maculae. </p>
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	https://treatment.plazi.org/id/A22887D1560EDA40FF57FC7BFE08F2E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15602DA4FFF57FA44FC16F228.text	A22887D15602DA4FFF57FA44FC16F228.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus pilosus Jordao, Clarkson & Ferreira-Jr 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus pilosus Jordão, Clarkson &amp; Ferreira-Jr, 2018</p>
            <p> Material examined. (4 exs.): “   BRASIL: MINAS GERAIS, Parque Estadual Serra do Cabral, Waterfall of Demir, long -44.211975 lat -17.920439, 833 m a.s.l., rock-face seepages, 14.iv.2019, leg. J.B. R.  Alencar ” (1 ex.  INPA , in alcohol); “ PARNA  Serra do Gandarela ,  Waterfall of Dona Chica , long -43.688806, lat -20.138139, 1165 m a.s.l., rock-face seepages, 14.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “Serro,  Waterfall of Esteira , long -43.492306, lat -18.571306, 1037m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol)  . </p>
            <p> Remarks. First record of the  O. pilosus from the state of Minas Gerais. </p>
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	https://treatment.plazi.org/id/A22887D15602DA4FFF57FA44FC16F228	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15602DA4EFF57F8D0FEA1F3D6.text	A22887D15602DA4EFF57F8D0FEA1F3D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus rotundus Clarkson & Short 2012	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus rotundus Clarkson &amp; Short, 2012</p>
            <p> Material examined. (380 exs.): “  BRASIL: MINAS GERAIS, Diamantina, spring, long -43.743694, lat -18.077278 ,   1350 m a.s.l., seepage habitat, 19.iv.2019, leg. J.B. R. Alencar ” (1 ex.  INPA , in alcohol); except: “long -43.6395, lat -18.216528  ,   1246 m a.s.l., seepage habitat, 20.iv.2019, leg. J.B. R. Alencar ” (1 ex.  INPA , in alcohol); “long - 43.599111, lat -18.231333  ,   1221 m a.s.l., seepage habitat, 21.iv.2019, leg. J.B. R. Alencar ” (2 exs.  INPA , in alcohol); “long -43.744639, lat -18.091639  ,   1359 m a.s.l., seepage habitat, 22.iv.2019, leg. J.B. R. Alencar ” (9 exs.  INPA , in alcohol); “ São João de Chapada,  Estação Córrego próx. Copanor, long -43.731139, lat -18.089972  ,   1284 m a.s.l., seepage habitat, 22.iv.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “ PARNA Sempre Vivas, Stream southeast of the camp, long -43.773861, lat -17.924444  ,   1259 m a.s.l., seepage habitat, 23.iv.2019, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); “ PARNA Sempre Vivas,  Waterfall of Felipes , long -43.703639, lat -17.907472  ,   1090 m a.s.l., rock-face seepages, 24.iv.2019, leg. J.B. R. Alencar ” (5 exs.  INPA , in alcohol); “ Parque Estadual Grão Mogol,  Waterfall Véu de Noiva , long -42.954639, lat -16.595667  ,   793 m a.s.l., vertical surfaces with water, 27.iv.2019, leg. J.B. R. Alencar ” (59 exs.  INPA , in alcohol); “ Parque Estadual Grão Mogol,  Waterfall of Mirante , long -42.899694, lat -16.576639  ,   821 m a.s.l., vertical surfaces with water, 27.iv.2019, leg. J.B. R. Alencar ” (2 exs.  INPA , in alcohol); “ Parque Estadual Grão Mogol,  Lapa da Água Fria , long -42.899556, lat -16.557083  ,   845 m a.s.l., vertical surfaces with water, 27.iv.2019, leg. J.B. R. Alencar ” (20 exs.  INPA , in alcohol); “ Parque Estadual Grão Mogol, stream near the archaeological site, long -42.943556 lat -16.618111  ,   719 m a.s.l., rock-face seepages, 29.iv.2019, leg. J.B. R. Alencar” (87 exs.  INPA , in alcohol); “ PARNA  Serra do Cipó , Waterfall Capão dos Palmitos, long -43.609556, lat - 19.377472  ,   830 m a.s.l., rock-face seepage, 8.vi.2019, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); “ PARNA Serra do Cipó,  Waterfall da Farofa , long -43.576139, lat -19.379389  ,   866 m a.s.l., vertical surfaces with water, 8.vi.2019, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol); “ PARNA Serra do Cipó,  Waterfall of Andorinhas , long -43.491472, lat -19.237611  ,   1028 m a.s.l., vertical surfaces with water, 10.iv.2019, leg. J.B. R. Alencar” (25 exs.  INPA , in alcohol); “ PARNA  Serra do Cipó , Canyon of the Bandeirinhas, long -43.562722, lat -19.423917  ,   880 m a.s.l., seepage habitat, 11.vi.2019, leg. J.B. R. Alencar” (8 exs.  INPA , in alcohol); “ PARNA Serra do Gandarela,  Waterfall of Dona Chica , long -43.688806, lat -20.138139  ,   1165 m a.s.l., rock-face seepages, 14.vi.2019, leg. J.B. R. Alencar” (59 exs.  INPA , in alcohol); “ PARNA Serra do Gandarela,  Waterfall of Capivari , long -43.581639, lat -20.132639  ,   1257 m a.s.l., seepage habitat, 17.vi.2019, leg. J.B. R. Alencar” (8 exs.  INPA , in alcohol); “ PARNA Serra do Gandarela,  Waterfall of Capivari , long -43.989917, lat -20.096611  ,   1335 m a.s.l., seepage habitat, 18.vi.2019, leg. J.B. R. Alencar ” (4 exs.  INPA , in alcohol); “ BRASIL: Minas Gerais, Serro,  Waterfall of Amaral , long -43.406944, lat -18.388528  ,   1043 m a.s.l., vertical surfaces with water, 22.vi.2019, leg. J.B. R. Alencar ” (52 exs.  INPA , in alcohol); “  Parque Estadual Serra do Cabral , Córrego, long -44.182778, lat -17.781111  ,   683 m a.s.l., rock-face seepages, 23.vi.2019, leg. J.B. R. Alencar ” (3 exs.  INPA , in alcohol); “ Itacambira,  Waterfall of Curiango , long -43.324861, lat -17.147556  ,   1051 m a.s.l., rock-face seepages, 24.vi.2019, leg. J.B. R. Alencar ” (6 exs.  INPA , in alcohol); “ Rio de Contas,  Waterfall of Fraga , long -41.8295, lat -13.598194  ,   617 m a.s.l., 23.iii.2020, leg. J.B. R. Alencar ” (4 exs.  INPA , in alcohol); “ Serranópolis de Minas, Canyon near the  Waterfall of Talhado , long -42.806389, lat -15.8175  ,   795 m a.s.l., 27.iii.2020, leg. J.B. R. Alencar ” (14 exs.  INPA , in alcohol); “ Botumirim,  Cahoeira do Bananal , long -43.037042, lat -16.839917  ,   908 m a.s.l., 28.iii.2020, leg. J.B. R. Alencar ” (14 exs.  INPA , in alcohol); “ Serro,  Waterfall de Esteira , long -43.492306, lat -18.571306  ,   1037 m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R. Alencar” (26 exs.  INPA , in alcohol); “ BAHIA: PARNA Chapada  Diamantina, Ribeirão do Meio, long -41.389194, lat -12.587639  ,   383 m a.s.l., 28.vi.2019, leg. J.B. R. Alencar ” (50 exs.  INPA , in alcohol); “ Piatã, Guarda-Mor,  Waterfall Saia de Noiva , long -41.867528, lat -13.311333  ,   1364 m a.s.l., 13.iii.2020, leg. J.B. R. Alencar ” (2 exs.  INPA , in alcohol); “ GOIÁS: Alto Paraíso de Goiás, Waterfall of muralha, long -47.7507, lat -14.2774, 957 m a.s.l., 2.VII.2019, leg. J.B. R. Alencar ” (4 exs.  INPA , in alcohol)  . </p>
            <p> Remarks. First record of  O. rotundus for Goiás state. The specimens most recently collected are not dorsally glabrous as illustrated by Clarkson &amp; Short (2012), the elytral surface has rows of fine systematic punctures bearing short setae similar to  O. humboldti sp. n. but can be easily differentiated by the aedeagus (see diagnosis of  O. humboldti sp. n. ). </p>
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	https://treatment.plazi.org/id/A22887D15602DA4EFF57F8D0FEA1F3D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15603DA4DFF57F88FFB06F5B0.text	A22887D15603DA4DFF57F88FFB06F5B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus xanthus Clarkson & Short 2012	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus xanthus Clarkson &amp; Short, 2012</p>
            <p> Material examined (7 exs.):   “ BRASIL: MINAS GERAIS, Serro,  Waterfall de Esteira , long -43.492306, lat - 18.571306, 1037 m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R. Alencar” (1 ex.  INPA , in alcohol); “ Pernambuco, Bonito, long -35.726722, lat -8.516972, 770 m a.s.l., 03.v.2014, leg. J.O. Silva; J.M.C. Nascimento; N. Hamada,” (6 exs.  INPA , in alcohol)  . </p>
            <p> Remarks. First record of the  O. xanthus Clarkson &amp; Short, 2012 from the state of Pernambuco. </p>
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	https://treatment.plazi.org/id/A22887D15603DA4DFF57F88FFB06F5B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15600DA4BFF57FA57FAAAF489.text	A22887D15600DA4BFF57FA57FAAAF489.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus yubai Clarkson & Short 2012	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oocyclus yubai Clarkson &amp; Short, 2012</p>
            <p>
                 Material examined (1649 exs.):   BRASIL: MINAS GERAIS, PARNA Serra do Cipó, Waterfall of Andorinhas, long -43.491472, lat -19.237611, 1028 m a.s.l., vertical surfaces with water, 10.iv.2019, leg. J.B. R. Alencar " (15 exs.  INPA ; 10 exs. SEMC, in alcohol); except: Waterfall Capão dos Palmitos, long -43.609556, lat -19.377472, 830 m a.s.l., rock-face seepage, 8.vi.2019, leg. J.B. R. Alencar ” (5 exs.  INPA , in alcohol); " Waterfall Grande, long -43.634333, lat -19.347056, 779 m a.s.l., vertical surfaces with water, 13.vi.2019, leg. J.B. R. Alencar ” (10 exs.  INPA , in alcohol); “  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Parque Estadual Serra do Cabral</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Boqueirão</a>
                 , long -44.178528, lat -17.758333, 680 m a.s.l., habitat higropétrico, 13.iv.2019, leg. J.B. R. Alencar ” (41 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Demir</a>
                 , long -44.211975 lat -17.920439, 833 m a.s.l., rock-face seepages, 14.iv.2019, leg. J.B. R. Alencar ” (118 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Fervedouro</a>
                 , long -44.247978, lat -17.917764, 1071 m a.s.l., rock-face seepages, 14.iv.2019, leg. J.B. R. Alencar ” (9 exs.  INPA , in alcohol); " Augusto Lima, Waterfall Santa Bárbara, long -44.065583, lat - 18.031556, 616 m a.s.l., vertical surfaces with water, 15.vi.2019, leg. J.B. R. Alencar ” (36 exs.  INPA , in alcohol); " Curimataí,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Simão</a>
                 , long -43.955333, lat -17.896139, 650 m a.s.l., rock-face seepages, 16.iv.2019, leg. J.B. R. Alencar ” (125 exs.  INPA , in alcohol); " Caetemirim,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Moinho</a>
                 , long -43.987889, lat -17.919472, 594 m a.s.l., vertical surfaces with water, 17.iv.2019, leg. J.B. R. Alencar ” (2 exs.  INPA , in alcohol); " Caetemirim, tributary of the  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Preto river</a>
                 , long -43.981217, -17.919472, 662 m a.s.l., vertical surfaces with water, 17.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (153 exs.  INPA , in alcohol); "  Diamantina, Stream of fazenda, long -43.769472, lat -18.062528, 1127 m a.s.l., vertical surfaces with water, 18.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (12 exs.  INPA , in alcohol); "  Diamantina, Stream of the  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Luiz Carlos</a>
                 , long -43.728417, lat -18.028611, 990 m a.s.l., seepage habitat, 19.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (1 ex.  INPA , in alcohol); “  Diamantina, São João de Chapada, Estação Córrego próx.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Copanor</a>
                 , long -43.731139, lat -18.089972, 1284 m a.s.l., seepage habitat, 22.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (2 exs.  INPA , in alcohol); "  Diamantina, Waterfall of  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Bocaina</a>
                 , long -42.837083, lat -16.301167, 669 m a.s.l., rock-face seepages, 28.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (80 exs.  INPA , in alcohol); " PARNA  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Sempre Vivas</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Stream</a>
                 southeast of the camp, long -43.773861, lat -17.924444, 1259 m a.s.l., seepage habitat, 23.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (5 exs.  INPA , in alcohol); " PARNA  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Sempre Vivas</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Felipes</a>
                 , long -43.703639, lat -17.907472, 1090 m a.s.l., rock-face seepages, 24.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (5 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Parque Estadual Grão Mogol</a>
                 , stream near the archaeological site, long -42.943556 lat -16.618111, 719 m a.s.l., rock-face seepages 29.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (87 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Parque Estadual Grão Mogol</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Almas</a>
                 , long -42.931194, lat -16.618139, 691 m a.s.l., rock-face seepages, 29.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (16 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Parque Estadual Grão Mogol</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Stream of Dé</a>
                 , long -42.963833, lat -16.592361, 746 m a.s.l., rock-face seepages, 29.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (17 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Parque Estadual Grão Mogol</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Almas</a>
                 , long -42.870111, lat -16.511139, 624 m a.s.l., rock-face seepages, 30.iv.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (29 exs.  INPA , in alcohol); "Lassance, Waterfall of Palmeiras, long -44.504167, lat -17.834333, 520 m a.s.l., vertical surfaces with water, 1.v.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); "" PARNA  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serra do Gandarela</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Viana</a>
                 , long -43.751472, lat -20.0975, 1078 m a.s.l., rock-face seepages, 14.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (8 exs.  INPA , in alcohol); "" PARNA Serra of Gandarela, Waterfall ofna Chica, long -43.688806, lat -20.138139, 1165 m a.s.l., rock-face seepages, 14.vi.2019, leg. J.B. R. Alencar” (59 exs.  INPA , in alcohol); "" PARNA  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serra do Gandarela</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Mingu</a>
                 , long -43.7596, lat -20.0622, 1006 m a.s.l., rock-face seepages, 15.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (44 exs.  INPA , in alcohol); “  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Itabirito</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Corrego Prox</a>
                 a  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Fazenda Velha</a>
                 , long -43.809139, lat -20.299111, 862 m a.s.l., rock-face seepages, 20.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (25 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Conceição of Mato Dentro</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Três Barras</a>
                 , long -43.539833, lat -19.335639, 701 m a.s.l., rock-face seepages, 21.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (31 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serro</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Amaral</a>
                 , long -43.406944, lat -18.388528, 1043 m a.s.l., vertical surfaces with water, 22.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (52 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serro</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Malheiros</a>
                 , long -43.4459, lat -18.6541, 738 m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (8 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serro</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Esteira</a>
                 , long -43.492306, lat -18.571306, 1037 m a.s.l., rock-face seepages, 31.iii.2020, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (26 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Curimataí</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Rio Curimataí</a>
                 , long -43.946806, lat -17.87825, 625 m a.s.l., vertical surfaces with water, 23.vi.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (7 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Gameleiras</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall of Andorinhas</a>
                 , long -43.079419, lat -14.981272, 931 m a.s.l., vertical surfaces with water and green algae, 24.iii.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (1 ex.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Mato Verde</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall Maria Rosa</a>
                 , long -42.795444, lat -15.432167, 725 m a.s.l., vertical surfaces with water and green algae, 25.iii.2019, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (46 exs.  INPA , in alcohol); “  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Serranópolis de Minas</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall Sete Quedas</a>
                 , vertical surfaces with water, 26.iii.2020, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (6 exs.  INPA , in alcohol); "  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Botumirim</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Waterfall Quatro Oitavas</a>
                 , long -43.014047, lat -16.883492, 862 m a.s.l., vertical surfaces with water, 27.iii.2020, leg. J.B. R.  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Alencar</a>
                 ” (9 exs.  INPA , in alcohol); “  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Santo Antônio do Itambé</a>
                 ,  
                <a title="Search Plazi for locations around (long -17.919472/lat -43.981216)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-17.919472&amp;materialsCitation.latitude=-43.981216">Rio Preto</a>
                 , long -43.320778, lat -18.461417, 702 m a.s.l., vertical surfaces with water, 30.iii.2020, leg. J.B. R. Alencar” (5 exs.  INPA , in alcohol); "   BAHIA: BA-564 Piatã-Mucugê, near the road, long -41.593139, lat -13.154611, 844 m a.s.l., vertical surfaces with water and green algae, 26.vi.2019, leg. J.B. R. Alencar ” (149 exs.  INPA , in alcohol); "Mucugê Ponte-BA-142, near the road, long -41.351444, lat -12.989583, 881 m a.s.l., seepage habitat, 27.vi.2019, leg. J.B. R. Alencar” (2 exs.  INPA , in alcohol); “ PARNA Chapada Diamantina, long -41.276139, lat -12.948639, 806 m a.s.l., seepage habitat, 27.vi.2019, leg. J.B. R. Alencar” (53 exs.  INPA , in alcohol); “ PARNA Chapada Diamantina, long -41.358306, lat -12.700667, 388 m a.s.l., seepage habitat, 27.vi.2019, leg. J.B. R. Alencar” (66 exs.  INPA , in alcohol); “ PARNA Chapada  Diamantina, Andaraí , long -41.348639, lat -12.746972, 314 m a.s.l., seepage habitat, 27.vi.2019, leg. J.B. R. Alencar” (31 exs.  INPA , in alcohol); “ PARNA Chapada  Diamantina, Lençóis , long -41.397444, lat -12.564167, 435 m al, 28.vi.2019, leg. J.B. R. Alencar” (139 exs.  INPA , in alcohol); “ PARNA Chapada  Diamantina, Ribeirão do Meio, long -41.389194, lat -12.587639, 383 m a.s.l., 28.vi.2019, leg. J.B. R. Alencar” (50 exs.  INPA , in alcohol); "Palmeiras,  Waterfall of Riachinho , long -41.515694, lat -12.572111, 935 m a.s.l., vertical surfaces with water, 18.iii.2020, leg. J.B. R. Alencar ” (53 exs.  INPA , in alcohol)  ;   “ MATO GROSSO DO SUL: Anastácio,  Aquidauana (c. 27 km S) on MS-170, long -55.69127, lat -20.72281, 225 m elev., 27.vi.2018, seepage on rock, leg. Hamada and team (10 exs. SEMC,  INPA ; pinned)”; “Aquidauana, Aquidauana (c. 15 km E) on plateau, long -55.6218, lat -20.4509, 380 m elev., 22.vi.2019, detritus and roots at margin of rock, leg. Hamada and team” (5 exs., SEMC,  INPA ; pinned)  . 
            </p>
            <p> Remarks. We expand the known distribution of this species to the states of Bahia, Minas Gerais and Mato Grosso do Sul; this is also the first record of  Oocyclus Sharp for the state of Mato Grosso do Sul. All adults examined are few punctures bear setae on the pronotum and elytra, thus the dorsal condition may appear glabrous </p>
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	https://treatment.plazi.org/id/A22887D15600DA4BFF57FA57FAAAF489	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
A22887D15606DA4AFF57F9BAFA62F2E5.text	A22887D15606DA4AFF57F9BAFA62F2E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oocyclus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Updated key to the Brazilian species of  Oocyclus Sharp</p>
            <p>This key is adapted from Clarkson &amp; Short (2012) and includes also an additional species from Jordão et al. (2018).</p>
            <p>1 Prosternal carina with two or more spine-like setae (Figs. 3e, 9e)................................................ 2</p>
            <p> - Prosternal carina without spine-like setae (see Jordão et al., (2018): Fig. 2a................................  O . pilosus </p>
            <p>2 Posterolateral angles of pronotum usually rounded (very rarely angulate) (Figs. 2, 4); prosternum with two or more spine-like setae; elytra with or without iridescent maculae, if present, maculae never blue; elytra with or without rows of erect setae.. 3</p>
            <p> - Posterolateral angles of pronotum weakly spinose (see Clarkson &amp; Short, 2012: Fig. 5); prosternum always with two spine-like setae; elytra with distinctive blue maculae and a dense row of erect setae (Fig. 5)...........................  O. decorus</p>
            <p>3 Prosternal carina set with 2 spine-like setae anteriorly (Figs. 2, 7).............................................. 12</p>
            <p>- Prosternal carina set with more than 2 spine-like setae anteriorly (usually 4–6; Figs. 3, 6)............................ 4</p>
            <p>4 Elytral surface heavily sculptured to appear bumpy or strongly undulating; glabrous (see Clarkson &amp; Short, 2012: Figs. 1, 3) ................................................................................................... 5</p>
            <p>- Elytral surface smooth, without bumps or strong undulations; glabrous or set with setae............................. 6</p>
            <p> 5 Elytral surface appearing blistered (see Clarkson &amp; Short, 2012: Fig. 3)..................................  O. alterosa</p>
            <p> - Elytral surface with irregular sculpturing, but without distinctly raised bumps (see Clarkson &amp; Short, 2012: Fig. 1)....................................................................................................  O. acuru</p>
            <p>6 Elytra with setae between rows of systematic punctures, appearing pubescent (Fig. 12b)............................. 7</p>
            <p>- Elytra with weak or non-existent pubescence between rows of systematic punctures (Figs. 12a, c, d)................... 8</p>
            <p> 7 Posterolateral angles of pronotum rounded. Elytra pubescence decumbent laterally and on elytral intervals, but with at least two erect rows of setae medially. Posterior margin of the elytra darkly coloured, except for the pale spot on posterior quarter (see Clarkson &amp; Short, 2012: Fig. 9).................................................................  O. itapercori</p>
            <p> - Posterolateral angles of pronotum angulate. Elytra pubescence decumbent; not forming distinct rows medially. Posterior margin of elytra broadly depigmented medially (see Clarkson &amp; Short, 2012: Fig. 13).............................  O. xanthus</p>
            <p> 8 Posterolateral margins of pronotum angulate (see Clarkson &amp; Short, 2012: Fig. 4); elytral surface with very shallowly impressed longitudinal grooves near suture..........................................................  O. caparao</p>
            <p>- Posterolateral margins of pronotum broadly rounded; elytral surface with or without very shallowly impressed longitudinal grooves near suture.................................................................................... 9</p>
            <p> 9 Elytra with strong indentation mediolaterally (see Clarkson &amp; Short, 2012: Fig. 7); body attenuated posteriorly..  O. fryanus</p>
            <p>- Elytra without or with only weak indentation mediolaterally; body not attenuated posteriorly........................ 10</p>
            <p> 10 Elytra surface with shallowly impressed longitudinal grooves; (e.g. Figs. 6a–e); each elytron with two or more posterior, small rounded pale spots..........................................................................  O. lacia sp. n.</p>
            <p>- Elytra surface without shallowly impressed longitudinal grooves or with weak undulating grooves near sutural interval; each elytron with a single posterior, rounded pale spot........................................................... 11</p>
            <p> 11. Elytra with undulating grooves near the sutural interval; TL/GW ratio = 1.65; abdominal ventrites with color strongly contrasting to the legs (e.g. Figs. 3a–e)............................................................  O. espinhacu sp. n.</p>
            <p> - Elytra without undulating grooves near the sutural interval; TL/GW ratio = 1.80; abdominal ventrites never in strong contrast to the legs..................................................................................  O. itapecum</p>
            <p>12 Elytra surface with fine punctures (see Clarkson &amp; Short, 2012: Figs. 2, 6); elytral suture raised or not................. 13</p>
            <p>- Elytra surface coarsely punctures (see Clarkson &amp; Short, 2012: Fig. 8); elytral suture raised......................... 14</p>
            <p>13 Posterolateral margins of pronotum angulate, but never spinose (Figs. 4c, 10c).................................... 15</p>
            <p>- Posterolateral margins of pronotum rounded............................................................... 16</p>
            <p> 14 Elytra with large, coarse punctures coarsely punctuate and without undulating grooves; posterolateral margins of pronotum angulate......................................................................................  O. iguazu</p>
            <p> - Elytra with moderately sized, coarse punctures and with undulating grooves; posterolateral margins of pronotum rounded (e.g. Figs. 8a–e)............................................................................  O. sulcatus sp. n.</p>
            <p> 15 Elytra with conspicuous rows of systematic punctures, row 1 bearing long, erect setae in contrast to row 2 (Figs. 10a–e, 12a); body size 4.7–5.4 mm; aedeagus as in Fig. 11i................................................  O . thysanus  sp. n.</p>
            <p> - Elytra with hardly distinguishable rows of systematic punctures, rows 1 and 2 with similar setae; body size 5.7–6.3 mm; aedeagus as in Fig. 11c (e.g. Figs. 4a–e)......................................................  O. giganteus sp. n.</p>
            <p>16 Abdominal ventrites dark brown to black, similar in color to thoracic venter (see Clarkson &amp; Short, 2012: Figs. 2, 6); elytral suture raised or not on posterior half; elytral margins slightly reflexed or not..................................... 17</p>
            <p>- Abdominal ventrites uniformly pale yellow or orange, distinctly darker than thoracic venter (see Clarkson &amp; Short, 2012: Fig. 14) and in many cases with distinct medial macula present on one or more ventrites (e.g. Fig. 5b); elytral suture not raised on posterior half; elytral margins not reflexed on posterior half.................................................. 18</p>
            <p>17 Each elytron with one rounded pale spot on distal quarter.................................................... 21</p>
            <p>- Each elytron with two rounded pale spots on distal quarter.................................................... 22</p>
            <p>18 Elytra without a row of large black spots alongside the suture (e.g. Fig. 5a)...................................... 19</p>
            <p>- Elytra with a row of large black spots alongside the suture (e.g. Fig. 6a)......................................... 20</p>
            <p> 19 Elytra surface without setae; abdominal ventrites uniformly pale from yellow to orange (see Clarkson &amp; Short, 2012: Fig. 11)..........................................................................................  O. rotundus</p>
            <p> - Elytra surface with setae; one or more abdominal ventrites in many cases with distinct medial macula as wide as or narrower than respective ventrites (e.g. Figs. 5a–e)...................................................  O. humboldti sp. n.</p>
            <p> 20 Thoracic venter and legs brown and slightly darker than abdominal ventrites (see Clarkson &amp; Short, 2012: Fig. 12).................................................................................................  O. schubarti</p>
            <p> - Thoracic venter and legs dark brown to black and strongly contrast to the yellow abdominal ventrites (see Clarkson &amp; Short, 2012: Fig. 14).................................................................................  O. yubai</p>
            <p>21 Elytral suture not raised on posterior half; elytral margins not reflexed on posterior half............................ 23</p>
            <p> - Elytral suture raised on posterior half; elytral margins weakly reflexed on posterior half.......................  O. flexus</p>
            <p> 22 Elytral intervals with dense pubescence inserted in ground punctation (Fig 12b); aedeagus with gonopore located on medial half of median lobe, basal piece tapered (Fig. 11h)...........................................  O. thrixdiastematus sp. n.</p>
            <p> - Elytral interval with weak or non-existent pubescence inserted in ground punctation (Figs. 12a, c, d); aedeagus with gonopore located at four-fifths the length of median lobe, basal piece rounded (Fig. 11a).........................  O. ecolab sp. n.</p>
            <p> 23 Metaventrite with oval, glabrous posteromedial area about half the length of metaventrite; aedeagus as in Figs. 2, 15b, in Clarkson &amp; Short (2012) ............................................................................  O. aeneus</p>
            <p> - Metaventrite with oval, glabrous posteromedial area about two-thirds the length of metaventrite; aedeagus as in Figs. 11f (Fig. 7a–e)..................................................................................  O. ovalis sp. n.</p>
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	https://treatment.plazi.org/id/A22887D15606DA4AFF57F9BAFA62F2E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alencar, Janderson Batista R.;Short, Andrew Edward Z.;Hamada, Neusa	Alencar, Janderson Batista R., Short, Andrew Edward Z., Hamada, Neusa (2022): New species and new distributional records of the hygropetric water scavenger beetle genus Oocyclus Sharp (Coleoptera, Hydrophilidae) from the Brazilian Shield. Zootaxa 5087 (2): 275-305, DOI: 10.11646/zootaxa.5087.2.3
