identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E511CE49FFEEFFF17FC7972A709CF904.text	E511CE49FFEEFFF17FC7972A709CF904.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atlantor Borkenhagen & Freyhof 2023	<div><p>Atlantor,  new genus</p><p>Figs 1–2</p><p>Material examined.   BMNH 1874.1.30.22-24, 3, syntypes, 96–155 mm SL; Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.344167&amp;materialsCitation.latitude=32.032776" title="Search Plazi for locations around (long -9.344167/lat 32.032776)">Oued Tensift</a>, [32°1′58″N, 9°20′39″W]; K. v. Fritsch &amp; J. Rein, 1872  .—   BMNH 1901.7.26.2-3, 2, 125–146 mm SL; Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.333889&amp;materialsCitation.latitude=33.327778" title="Search Plazi for locations around (long -8.333889/lat 33.327778)">Oued Oum</a> er Rbia [33°19′40″N, 8°20′2″W]; E. Hartert  .—  MNHN 1912-94, 1, 120 mm SL;  MNHN 1912-95, 1, 124 mm SL;   MNHN 1912-96, 1, 92 mm SL; Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.3425&amp;materialsCitation.latitude=33.289444" title="Search Plazi for locations around (long -8.3425/lat 33.289444)">Oued Oum</a> er Rbia near Azemmour [33°17′22″N, 8°20′33″W], C. du Gast, 1912  .—   MNHN 1927-98, 1, 111 mm SL; Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.668611&amp;materialsCitation.latitude=32.939167" title="Search Plazi for locations around (long -5.668611/lat 32.939167)">Oued Oum</a> er Rbia near Khenifra [32°56′21″N, 5°40′7″W], A. Gruvel  &amp;   R.  Dollfus, 1927  .—  SMF 636, 1, 82 mm SL;   SMF 952, 2, 49–83 mm SL; Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.7675&amp;materialsCitation.latitude=31.483055" title="Search Plazi for locations around (long -9.7675/lat 31.483055)">Oued Ksob</a> [31°28′59″N, W 9°46′3″W], K. v. Fritsch &amp; J. Rein, 1872  .</p><p>Diagnosis.  Atlantor is distinguished from  Carasobarbus by having 5½ branched anal-fin rays (vs. 6½), and modally 8½ branched dorsal-fin ray (vs. 9½–10½), and from  Mesopotamichthys and  Caecocypris by having two pairs of long barbels (vs. no barbels). There is no consistent morphological character that clearly distinguishes  Atlantor from all species of  Arabibarbus and  Labeobarbus, and the new genus can only be distinguished from  Arabibarbus and  Labeobarbus by its distinct phylogenetic position and genetic distance (p-distance in cytochrome b of 5.3% and 6.7% respectively; Kai Borkenhagen, unpublished).  Atlantor differs from  Pterocapoeta by having two pairs of long barbels (vs. one pair of minute maxillary barbels hidden in a skin fold), dorsal fin with 8½–9½ branched rays (vs. 10½–11½), 25–32 total lateral line scales (vs. 43–46), well developed, fleshy lips and a small median lobe on the lower lip (vs. lower lip with cornified sheath), mouth arched (vs. straight). Current knowledge indicates that all Western Asian, and African Torinae are evolutionarily hexaploid, and as  Atlantor belongs to this phylogenetic group of species, it is assumed, that they are also hexaploid. As a consequence,  Atlantor is distinguished from the Indomalayan genera  Hypselobarbus,  Lepidopygopsis,  Naziritor,  Neolissochilus,  Osteochilichthys, and from  Tor by being evolutionarily hexaploid (vs. tetraploid (2n≈100) (Yang et al. 2015).</p><p>Type species.  Barbus reinii Günther, 1874: 231</p><p>Included species.  Barbus reinii Günther, 1874:231</p><p>Etymology. The name is derived from the Atlas Mountains in Morocco and  Tor, the generic name of the tribe that includes  Atlantor (neutral).</p><p>Distribution and conservation status. The type locality of  A. reinii is the Tennsift River in Morocco (Günther 1874). Records from the Oum er Rbia (Günther 1902, Pellegrin 1921) are confirmed by vouchers (MNHN 1912-94– 96, MNHN 1927-98). In recent years, the species was only found in the Ksob River, a tributary to Oued Igrounzar, by Machordom &amp; Doadrio (2001) (I. Doadrio, pers. communication, 2019).  Atlantor reinii has not been found since 2001, despite a number of attempts to confirm its continued existence. Freyhof &amp; Ford (2022) treat  A. reinii as extinct.</p><p>Remarks. There are only two cytochrome b sequences available, which were published by Machordom &amp; Doadrio (2001) and Durand et al. (2002), and later included in phylogenetic studies by Tsigenopoulos et al. (2010), Berrebi et al. (2014), Borkenhagen (2014), Yang et al. (2015). All these studies show  Atlantor as a distinct phylogenetic lineage from related genera such as  Arabibarbus,  Carasobarbus,  Labeobarbus,  Mesopotamichthys, and  Ptercapoeta . We have not reproduced the phylogenetic trees here, as they are available open-access from their original sources, which indicate the unique phylogenetic position of  Atlantor .</p><p>Comparative material. See Borkenhagen &amp; Krupp (2013) and Borkenhagen (2014) for materials of additional species and genera examined.</p></div>	https://treatment.plazi.org/id/E511CE49FFEEFFF17FC7972A709CF904	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Borkenhagen, Kai;Freyhof, Jörg	Borkenhagen, Kai, Freyhof, Jörg (2023): Atlantor, a new generic name for Barbus reinii Günther, 1874 from Morocco (Teleostei: Cyprinidae). Zootaxa 5319 (3): 429-434, DOI: 10.11646/zootaxa.5319.3.9, URL: http://dx.doi.org/10.11646/zootaxa.5319.3.9
