taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825898/files/figure.png	http://doi.org/10.5281/zenodo.5825898	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825904/files/figure.png	http://doi.org/10.5281/zenodo.5825904	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825910/files/figure.png	http://doi.org/10.5281/zenodo.5825910	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825914/files/figure.png	http://doi.org/10.5281/zenodo.5825914	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825916/files/figure.png	http://doi.org/10.5281/zenodo.5825916	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825918/files/figure.png	http://doi.org/10.5281/zenodo.5825918	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825920/files/figure.png	http://doi.org/10.5281/zenodo.5825920	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825922/files/figure.png	http://doi.org/10.5281/zenodo.5825922	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825924/files/figure.png	http://doi.org/10.5281/zenodo.5825924	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825926/files/figure.png	http://doi.org/10.5281/zenodo.5825926	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825928/files/figure.png	http://doi.org/10.5281/zenodo.5825928	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825930/files/figure.png	http://doi.org/10.5281/zenodo.5825930	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825932/files/figure.png	http://doi.org/10.5281/zenodo.5825932	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825934/files/figure.png	http://doi.org/10.5281/zenodo.5825934	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825956/files/figure.png	http://doi.org/10.5281/zenodo.5825956	FIGURE 26. Centruroides rileyi Sissom, 1995, ♂ (CNAN SC4002), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 26. Centruroides rileyi Sissom, 1995, ♂ (CNAN SC4002), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825958/files/figure.png	http://doi.org/10.5281/zenodo.5825958	FIGURE 27. Centruroides rileyi Sissom, 1995, ♀ (CNAN SC4003), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 27. Centruroides rileyi Sissom, 1995, ♀ (CNAN SC4003), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825960/files/figure.png	http://doi.org/10.5281/zenodo.5825960	FIGURE 28. Centruroides cuauhmapan, sp. nov., ♂ (CNAN T01396), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 28. Centruroides cuauhmapan, sp. nov., ♂ (CNAN T01396), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825962/files/figure.png	http://doi.org/10.5281/zenodo.5825962	FIGURE 29. Centruroides cuauhmapan, sp. nov., ♀ (CNAN T01399), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 29. Centruroides cuauhmapan, sp. nov., ♀ (CNAN T01399), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825964/files/figure.png	http://doi.org/10.5281/zenodo.5825964	FIGURE 30. Centruroides catemacoensis, sp. nov., ♂ (CNAN T01424), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 30. Centruroides catemacoensis, sp. nov., ♂ (CNAN T01424), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825966/files/figure.png	http://doi.org/10.5281/zenodo.5825966	FIGURE 31. Centruroides catemacoensis, sp. nov., ♀ (CNAN T01423), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 31. Centruroides catemacoensis, sp. nov., ♀ (CNAN T01423), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825968/files/figure.png	http://doi.org/10.5281/zenodo.5825968	FIGURE 32. Centruroides hamadryas, sp. nov., ♂ (CNAN T01408), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 32. Centruroides hamadryas, sp. nov., ♂ (CNAN T01408), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825970/files/figure.png	http://doi.org/10.5281/zenodo.5825970	FIGURE 33. Centruroides hamadryas, sp. nov., ♀ (CNAN T01415), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 33. Centruroides hamadryas, sp. nov., ♀ (CNAN T01415), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825972/files/figure.png	http://doi.org/10.5281/zenodo.5825972	FIGURE 35. Centruroides berstoni, sp. nov., ♀ (CASENT 9073313), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 35. Centruroides berstoni, sp. nov., ♀ (CASENT 9073313), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825978/files/figure.png	http://doi.org/10.5281/zenodo.5825978	FIGURE 37. Centruroides schmidti Sissom 1995, ♀ (CASENT 9073317), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm	FIGURE 37. Centruroides schmidti Sissom 1995, ♀ (CASENT 9073317), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825980/files/figure.png	http://doi.org/10.5281/zenodo.5825980	FIGURE 38. Centruroides hoffmanni Armas 1996, ♂ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 38. Centruroides hoffmanni Armas 1996, ♂ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825982/files/figure.png	http://doi.org/10.5281/zenodo.5825982	FIGURE 39. Centruroides hoffmanni Armas 1996, ♀ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 39. Centruroides hoffmanni Armas 1996, ♀ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825986/files/figure.png	http://doi.org/10.5281/zenodo.5825986	FIGURE 41. Centruroides chanae, sp. nov., ♀ (CNAN T01405), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 41. Centruroides chanae, sp. nov., ♀ (CNAN T01405), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825988/files/figure.png	http://doi.org/10.5281/zenodo.5825988	FIGURE 42. Centruroides yucatanensis, sp. nov., ♂ (CNAN T01416), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	FIGURE 42. Centruroides yucatanensis, sp. nov., ♂ (CNAN T01416), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D54FF992410FD71FB05FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825990/files/figure.png	http://doi.org/10.5281/zenodo.5825990	FIGURE 43. Centruroides yucatanensis, sp. nov., ♀ (CNAN T01417), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	FIGURE 43. Centruroides yucatanensis, sp. nov., ♀ (CNAN T01417), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825898/files/figure.png	http://doi.org/10.5281/zenodo.5825898	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825914/files/figure.png	http://doi.org/10.5281/zenodo.5825914	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825920/files/figure.png	http://doi.org/10.5281/zenodo.5825920	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825928/files/figure.png	http://doi.org/10.5281/zenodo.5825928	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825930/files/figure.png	http://doi.org/10.5281/zenodo.5825930	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5AFF9E2436FBA1FF74FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825904/files/figure.png	http://doi.org/10.5281/zenodo.5825904	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825910/files/figure.png	http://doi.org/10.5281/zenodo.5825910	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825918/files/figure.png	http://doi.org/10.5281/zenodo.5825918	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825924/files/figure.png	http://doi.org/10.5281/zenodo.5825924	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825926/files/figure.png	http://doi.org/10.5281/zenodo.5825926	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825964/files/figure.png	http://doi.org/10.5281/zenodo.5825964	FIGURE 30. Centruroides catemacoensis, sp. nov., ♂ (CNAN T01424), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 30. Centruroides catemacoensis, sp. nov., ♂ (CNAN T01424), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D5DFF872795FB10FCCAFC01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825966/files/figure.png	http://doi.org/10.5281/zenodo.5825966	FIGURE 31. Centruroides catemacoensis, sp. nov., ♀ (CNAN T01423), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 31. Centruroides catemacoensis, sp. nov., ♀ (CNAN T01423), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825904/files/figure.png	http://doi.org/10.5281/zenodo.5825904	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825916/files/figure.png	http://doi.org/10.5281/zenodo.5825916	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825922/files/figure.png	http://doi.org/10.5281/zenodo.5825922	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825932/files/figure.png	http://doi.org/10.5281/zenodo.5825932	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825934/files/figure.png	http://doi.org/10.5281/zenodo.5825934	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D44FFB0242DFBF5FC7FFBA2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825904/files/figure.png	http://doi.org/10.5281/zenodo.5825904	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825910/files/figure.png	http://doi.org/10.5281/zenodo.5825910	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825918/files/figure.png	http://doi.org/10.5281/zenodo.5825918	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825924/files/figure.png	http://doi.org/10.5281/zenodo.5825924	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825926/files/figure.png	http://doi.org/10.5281/zenodo.5825926	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825960/files/figure.png	http://doi.org/10.5281/zenodo.5825960	FIGURE 28. Centruroides cuauhmapan, sp. nov., ♂ (CNAN T01396), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 28. Centruroides cuauhmapan, sp. nov., ♂ (CNAN T01396), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D73FFB825D3FB59FBDDFDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825962/files/figure.png	http://doi.org/10.5281/zenodo.5825962	FIGURE 29. Centruroides cuauhmapan, sp. nov., ♀ (CNAN T01399), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 29. Centruroides cuauhmapan, sp. nov., ♀ (CNAN T01399), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825914/files/figure.png	http://doi.org/10.5281/zenodo.5825914	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825920/files/figure.png	http://doi.org/10.5281/zenodo.5825920	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825928/files/figure.png	http://doi.org/10.5281/zenodo.5825928	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825930/files/figure.png	http://doi.org/10.5281/zenodo.5825930	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825968/files/figure.png	http://doi.org/10.5281/zenodo.5825968	FIGURE 32. Centruroides hamadryas, sp. nov., ♂ (CNAN T01408), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 32. Centruroides hamadryas, sp. nov., ♂ (CNAN T01408), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D7BFFA025C5FDA5FCA8F937.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825970/files/figure.png	http://doi.org/10.5281/zenodo.5825970	FIGURE 33. Centruroides hamadryas, sp. nov., ♀ (CNAN T01415), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 33. Centruroides hamadryas, sp. nov., ♀ (CNAN T01415), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825916/files/figure.png	http://doi.org/10.5281/zenodo.5825916	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825922/files/figure.png	http://doi.org/10.5281/zenodo.5825922	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825932/files/figure.png	http://doi.org/10.5281/zenodo.5825932	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825980/files/figure.png	http://doi.org/10.5281/zenodo.5825980	FIGURE 38. Centruroides hoffmanni Armas 1996, ♂ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 38. Centruroides hoffmanni Armas 1996, ♂ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D67FFA62796FF2BFC4AF93A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825982/files/figure.png	http://doi.org/10.5281/zenodo.5825982	FIGURE 39. Centruroides hoffmanni Armas 1996, ♀ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 39. Centruroides hoffmanni Armas 1996, ♀ (CNAN SC3996), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825904/files/figure.png	http://doi.org/10.5281/zenodo.5825904	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. catemacoensis, sp. nov. (triangles); C. chanae, sp. nov. (circles); C. cuauhmapan, sp. nov. (diamonds); C. rileyi Sissom, 1995 (squares).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825910/files/figure.png	http://doi.org/10.5281/zenodo.5825910	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 5. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825918/files/figure.png	http://doi.org/10.5281/zenodo.5825918	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	FIGURE 8. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), B. ♀ (CNAN SC4003). C, D. C. cuauhmapan, sp. nov., C. holotype ♂ (CNAN T01396), D. paratype ♀ (CNAN T01399). E, F. C. catemacoensis, sp. nov., E. holotype ♂ (CNAN T01424), F. paratype ♀ (CNAN T01423). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825924/files/figure.png	http://doi.org/10.5281/zenodo.5825924	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	FIGURE 11. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♂ (CNAN SC4002). B. C. cuauhmapan, sp. nov., holotype ♂ (CNAN T01396). C. C. catemacoensis, sp. nov., holotype ♂ (CNAN T01424). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825926/files/figure.png	http://doi.org/10.5281/zenodo.5825926	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 12. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. rileyi Sissom, 1995, ♀ (CNAN SC4003). B. C. cuauhmapan, sp. nov., paratype ♀ (CNAN T01399). C. C. catemacoensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825956/files/figure.png	http://doi.org/10.5281/zenodo.5825956	FIGURE 26. Centruroides rileyi Sissom, 1995, ♂ (CNAN SC4002), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 26. Centruroides rileyi Sissom, 1995, ♂ (CNAN SC4002), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6AFFAD27FBFAEFFC74F947.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825958/files/figure.png	http://doi.org/10.5281/zenodo.5825958	FIGURE 27. Centruroides rileyi Sissom, 1995, ♀ (CNAN SC4003), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	FIGURE 27. Centruroides rileyi Sissom, 1995, ♀ (CNAN SC4003), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 5 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825898/files/figure.png	http://doi.org/10.5281/zenodo.5825898	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825914/files/figure.png	http://doi.org/10.5281/zenodo.5825914	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825920/files/figure.png	http://doi.org/10.5281/zenodo.5825920	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825928/files/figure.png	http://doi.org/10.5281/zenodo.5825928	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825930/files/figure.png	http://doi.org/10.5281/zenodo.5825930	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D6FFFD72788FBACFDB5FAEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825902/files/figure.png	http://doi.org/10.5281/zenodo.5825902	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825908/files/figure.png	http://doi.org/10.5281/zenodo.5825908	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses).	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825916/files/figure.png	http://doi.org/10.5281/zenodo.5825916	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 7. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825922/files/figure.png	http://doi.org/10.5281/zenodo.5825922	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	FIGURE 10. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hoffmanni Armas, 1996, A. ♂ (CNAN SC3996), B. ♀ (CNAN SC3996). C, D. C. chanae, sp. nov., C. holotype ♂ (CNAN T01403), D. paratype ♀ (CNAN T01405). E, F. C. yucatanensis, sp. nov., E. holotype ♂ (CNAN T01416), F. paratype ♀ (CNAN T01417). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825932/files/figure.png	http://doi.org/10.5281/zenodo.5825932	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	FIGURE 15. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♂ (CNAN SC3996). B. C. chanae, sp. nov., holotype ♂ (CNAN T01403). C. C. yucatanensis, sp. nov., holotype ♂ (CNAN T01416). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825934/files/figure.png	http://doi.org/10.5281/zenodo.5825934	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 16. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hoffmanni Armas, 1996, ♀ (CNAN SC3996). B. C. chanae, sp. nov., paratype ♀ (CNAN T01405). C. C. yucatanensis, sp. nov., paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825936/files/figure.png	http://doi.org/10.5281/zenodo.5825936	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825938/files/figure.png	http://doi.org/10.5281/zenodo.5825938	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825942/files/figure.png	http://doi.org/10.5281/zenodo.5825942	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825944/files/figure.png	http://doi.org/10.5281/zenodo.5825944	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825946/files/figure.png	http://doi.org/10.5281/zenodo.5825946	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825948/files/figure.png	http://doi.org/10.5281/zenodo.5825948	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825950/files/figure.png	http://doi.org/10.5281/zenodo.5825950	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825952/files/figure.png	http://doi.org/10.5281/zenodo.5825952	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825954/files/figure.png	http://doi.org/10.5281/zenodo.5825954	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825988/files/figure.png	http://doi.org/10.5281/zenodo.5825988	FIGURE 42. Centruroides yucatanensis, sp. nov., ♂ (CNAN T01416), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	FIGURE 42. Centruroides yucatanensis, sp. nov., ♂ (CNAN T01416), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
510E87A56D14FFDD278DFA9CFC77FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5825990/files/figure.png	http://doi.org/10.5281/zenodo.5825990	FIGURE 43. Centruroides yucatanensis, sp. nov., ♀ (CNAN T01417), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	FIGURE 43. Centruroides yucatanensis, sp. nov., ♀ (CNAN T01417), habitus. A. Dorsal aspect. B. Ventral aspect. Scale bars = 10 mm.	2021-09-16	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.		Zenodo	biologists	Goodman, Aaron M.;Prendini, Lorenzo;Francke, Oscar F.;Esposito, Lauren A.			
