taxonID	type	description	language	source
A587CF9A10795BD69A1248318951032E.taxon	description	Figs 17, 18	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	diagnosis	Diagnosis. Hosta longipedicellata is similar to H. scabrinervia, but distinguished by its flower bracts 2 - 2.2 cm long, withering during flowering (compared to (2 -) 2.2 - 4.6 cm long, fresh during flowering), smaller and narrower leaf blades (compared to up to 25 x 15 cm), and the occurrence at elevations at 1700 - 1750 m (in contrast to at 90 - 550 m).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	materials_examined	Type. Japan. Kochi Pref.: Tosa-gun, Hongawa-mura, Mt. Tsutsujo, 1750 m elev., 1 Aug. 2006, S. Kobayashi & Y. Kurahashi FOK- 077809 with flowers (holotype MBK 0164413!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 3 per ramet; blades ovate or oblong-ovate, 13.4 - 30.2 cm long, 5.4 - 11.8 cm, 2.2 - 3 time longer than wide, thinly papery, glabrous on both surfaces, base rounded to subcordate, apex acute to short acuminate, margin entire, veins in 7 - 10 pairs, papillose on the lower surface; petioles 15 - 34.3 cm long, narrowly winged, wing 0.1 - 0.2 cm wide, glabrous. Scape 26.5 - 32.5 cm long, terete. Raceme 7.6 - 9 cm long, 7 - 15 - flowered; flower bracts withering in anthesis, erect, purplish light green in the upper part, light green in the lower part, oblong-lanceolate, boat-shaped, 2 - 2.5 cm long, 0.3 - 0.4 cm wide, membranous, glabrous, apex acute. Flowers not fragrant; pedicels 2.5 - 3.3 cm long, glabrous; perianth 4.4 - 6.4 cm long, funnel-form, glabrous, 6 - lobed, whitish purple outside in flower buds, almost white outside and inside when flowering, midveins purplish adaxially; tube dilated from apical 2 / 3, lobes triangular, 1.0 - 1.5 cm long, 0.6 - 0.9 cm wide, apex obtuse. Stamens 6, slightly shorter than perianth, not exerted; filaments 4.6 - 4.8 cm long, upwardly curved at the distal part, white, free, glabrous, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Ovary ellipsoid, 0.6 - 0.7 cm long, style 5.1 - 5.6 cm long, upwardly curved at the distal part, 1.1 cm exerted from perianths, glabrous, stigma capitate. Capsules or seeds not observed.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	distribution	Distribution and habitat. Japan (Kochi Prefecture). This species grows on rock cliffs at Mt. Tsutsujo, Mt. Tebako, Yasui Valley, and its vicinity in Kochi Prefecture on Shikoku Island.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	etymology	Etymology. The specific epithet is derived from the long pedicel.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A587CF9A10795BD69A1248318951032E.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Tosa-gun. Hongawa-mura, Mt. Tsutsujo, 28 Jun. 2022, K. Fuse et al. JPN 12637 - 12639, 12647, 12654 sterile (FU!); Agawa-gun, Niyodogawa-cho, 612 m elev., 3 May 2022, Se. Fujii JPN 12425, 12426 sterile (FU!); Agawa-gun, Niyodogawa-cho, Yasui Valley, 470 m elev., 30 Jun. 2022, K. Fuse et al. JPN 12674, 12676, 12680 sterile, JPN 12675 with flower buds (FU!); Agawa-gun, Niyodogawa-cho, Miyagahira, 612 m elev., Jul. 22, 2023, Se. Fujii FJW 48 - 1, 2, 3 with flowers (MBK!); cultivated at Makino Botanical Garden, derived from Mt. Tsutsujo, 1750 m elev., T. Yahara et al. JPN 3927 sterile (FU!); cultivated at Makino Botanical Garden, derived from Mt. Tebako, 1700 m elev., T. Yahara et al. JPN 3936 sterile (FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	description	Figs 12, 13	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	diagnosis	Diagnosis. Hosta minazukiflora is similar to H. longipedicellata in having leaves shorter than 20 cm, straight scapes shorter than 43 cm, and perianth lobes 0.6 - 0.8 cm wide. However, H. minazukiflora is distinguished from H. longipedicellata by its lavender flowers (vs. whitish), shorter pedicels (1.1 - 1.2 cm vs. 2.5 - 3.3 cm), narrower leaves (2.9 - 6.2 cm wide vs. (5.4 -) 9 - 12 cm wide) smooth on the lower surface (vs. papillose), and the occurrence at elevations of 270 - 280 m (vs. 1700 - 1750 m).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	materials_examined	Type. Japan. Kochi Pref.: Nagaoka-gun, Motoyama-cho, Asemi River, 25 Jun. 2017, Se. Fujii & K. Yabe FOS- 012016 with flowers (holotype MBK 0318414!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 4 - 6 per ramet; blades oblong-ovate to oblong-lanceolate, 11.8 - 17.0 cm long, 2.9 - 6.2 cm wide, 2.4 - 4.3 times longer than width, thinly papery, glabrous on both surfaces, base cuneate, decurrent, slightly folded, apex acuminate, margin entire, veins in 6 - 9 pairs, smooth on the lower surface; petioles 8 - 19 cm long, narrowly winged, wing 0.1 - 0.3 cm wide, glabrous. Scape 18 - 43 cm long, terete. Raceme 10 - 15 cm long, 3 - 6 - flowered; flower bracts vivid (not withering) in anthesis, erect, whitish purple, oblong-lanceolate, boat-shaped, 1.3 - 1.6 cm long, 0.2 - 0.4 cm wide, membranous, glabrous, apex acute. Flowers not fragrant, 5.2 - 5.9 cm long; pedicels 1.1 - 1.2 cm long, glabrous. Perianth whitish purple, funnel-form, 4.1 - 4.7 cm long, glabrous, 6 - lobed; abruptly dilated from apical 2 / 3, lobes narrowly triangular, 1.2 - 1.4 cm long, 0.6 - 0.8 cm wide, apex acute. Stamens 6; filaments 4.6 - 5.4 cm long, white, free, glabrous, 0.5 - 0.7 cm exerted from perianth, upwardly curved at the distal part, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Style 5.2 - 6.1 cm long, upwardly curved at the distal part, 1.1 - 1.4 cm exerted from the perianth, glabrous, stigma capitate. Ovary ellipsoid, 0.6 cm long, glabrous. Capsule green, cylindrical, 1.8 - 2.3 cm long, 0.3 - 0.4 cm wide, shallowly 3 - angled. Seeds ellipsoid-ovoid, 2 mm long, with wings 4 mm long, black when dry.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	distribution	Distribution and habitat. Japan (Kochi Prefecture). This species grows on rock cliffs along the Asemi River, a branch of the Yoshino River, in the central part of the Kochi Prefecture on Shikoku Island.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	etymology	Etymology. The specific epithet was derived from the flowering season in June. Minazuki refers to June in Japanese.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
9CA812E2FA2E5ADBAEE73135FB2643FE.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Nagaoka-gun, Motoyama-cho, Asemi River, 280 m elev., 19 Aug. 2020, T. Yahara et al. JPN 1308 sterile, JPN 1309 sterile, and JPN 1310 with fruit (FU!); ditto, 13 Jun. 2004, Y. Yamashita et al. FOK- 066868 with flowers (MBK 0083845!); ditto, 23 May 2002, Y. Kokami et al. FOK- 055767 with flower buds (MBK 0146148!); ditto, 270 m elev., 25 Jun. 2017, Se. Fujii & K. Yabe FOS- 012017 with flowers (MBK 0318415!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	description	Fig. 9	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	diagnosis	Diagnosis. Hosta polyneuronoides is distinguished from H. tardiva subsp. densinervia in having 3 - 10 flowers per inflorescence (in contrast to 15 - 25 in subsp. Hosta tardiva densinervia) and anthers 3.5 - 4 mm long (compared to 5 mm long).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	materials_examined	Type. Japan. Kochi Pref.: Agawa-gun, Ino-cho, near Nitaki Bridge, 237 m elev., 20 Aug. 2020, T. Yahara, K. Fuse & H. Sato JPN 1378 with flowers (holotype FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 3 - 15 per ramet; blades ovate or oblong-ovate, 7.5 - 28.5 cm long, 3.3 - 15.3 cm wide, 0.9 - 2.9 times longer than width, thinly papery, glabrous on both surfaces, base cuneate to subcordate, apex acute to short acuminate, margin entire, veins 8 - 9 pairs, smooth on the lower surface; petioles 5 - 40 cm long, narrowly winged, 0.3 - 0.5 cm wide, glabrous. Scape 15 - 51 cm long, rachis terete, bract lanceolate, 4 cm long, 0.3 - 0.5 cm wide, light green, glabrous. Raceme 4.3 - 9 cm long, 3 - 10 - flowered; flower bracts vivid (not withering) in anthesis, erect or diagonally spreading, whitish green, oblong-lanceolate, boat-shaped, 1.2 - 3.1 cm long, 0.1 - 0.8 cm wide, membranous, glabrous, apex acuminate. Flowers not fragrant, 5.7 - 6.0 cm long; pedicels 0.6 - 1.3 cm long, glabrous. Perianths white or very pale purple-white, funnel-form, 2.1 - 4.9 cm long, glabrous, 6 - lobed; tube abruptly dilated from apical 2 / 3, lobes narrowly triangular, 0.7 - 1.2 cm long, 0.4 - 1 cm wide, apex acute. Stamens 6, 0.1 - 0.4 cm exerted from perianth; filaments white, free, 4.5 - 4.8 cm long, glabrous, anthers purple to yellow when fresh, dark blue-grey to light yellow when dried, 3.5 - 4 mm long. Ovary ellipsoid, 0.5 cm long, glabrous style 5.5 - 5.7 cm long, upwardly curved at the distal part, subequal to 1.5 cm exerted from perianth, glabrous, stigma capitate. Capsules or seeds not observed.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	distribution	Distribution and habitat. Japan (Kochi and Ehime Prefectures). Hosta polyneuronoides grows on open or shaded rocks and rock cliffs along rivers in the upper reaches and headwaters of the Yoshino River, and on the rocky ridgeline from Mt. Ishizuchi to Mt. Komochi-gongen and its vicinity.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	etymology	Etymology. This subspecies was named for its resemblance to H. tardiva subsp. densinervia identified as Hosta kikutii var. polyneuron by Fujita (1976).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0454E2E817A253E9A31BE0057C723C8A.taxon	materials_examined	Additional specimens examined. Japan. Ehime Pref.: Niihama City, Mt. Higashi-akaishi, 970 m elev., 21 Jun. 2021, T. Yahara et al. JPN 6269 - 6271 sterile (FU!); ditto, 1040 m elev., 21 Jun. 2021, T. Yahara et al. JPN 6153 - 6156 sterile (FU!); Mt. Ishizuchi, 28 Jul. 1972, S. Takafuji 799 with flowers (KYO!); Kamiukena-gun, Mt. Ishizuchi, 1600 m elev., 23 Jun. 2021, T. Yahara et al. JPN 6390 sterile (FU!); ditto, 1980 m elev., 23 Jun. 2021, T. Yahara et al. JPN 6407 sterile (FU!); Saijyo City, Mts. Ishizuchi, from Yoake-toge to Mt. Nishinokanmuri-dake, 1600 m elev., 7 Aug. 1971, H. Takahashi & N. Fujita 226 with flowers (KYO!). Kochi Pref.: Nagaoka-gun, Motoyama-cho, Along Asemi River, 480 m elev., 19 Aug. 2020, T. Yahara et al. JPN 1327 - 1329 with flower buds (FU!); Tosa-gun, Okawa-mura, 570 m elev., 24 Jun. 2021, T. Yahara et al. JPN 6493 - 6495 sterile (FU!); Tosa-gun, Ohkawa-mura, Kawasaki, 400 m elev., 24 Jun. 2021, T. Yahara et al. JPN 6501 - 6503 with flower buds (FU!); Tosa-gun, Ohkawa-mura, Takano, 24 Jun. 2022, K. Fuse et al. JPN 12529, 12530, 12532, 12535 sterile (FU!); Tosa-gun, Ohkawa-mura, near Okina Waterfall, 443 m elev., Jul. 25, 2023, Se. Fujii, FJW 49 - 1, 2 with flowers (MBK!); Tosa-gun, Tosa-cho, 24 Jun. 2022, K. Fuse et al. JPN 12506 - 12508 sterile (FU!); Agawa-gun, Ino-cho, 750 m elevation, 23 Jun. 2021, T. Yahara et al. JPN 6449 - 6451 sterile (FU!); Agawa-gun, Ino-cho, 900 m elevation, 23 Jun. 2021, T. Yahara et al. JPN 6446 - 6448 sterile (FU!); Agawa-gun, Ino-cho, Kamegamori Forest Road, 1650 m elev., 22 Jun. 2021, T. Yahara et al. JPN 6342, 6361 sterile (FU!); ditto, Mt. Komochi-gongen, 29 Jun. 2022, K. Fuse et al. JPN 12663, 12664 sterile (FU!); Agawa-gun, Ino-cho, Ohmori-gawa dam, 755 m elev., 3 May 2022, Se. Fujii JPN 12427 sterile (FU!); Agawa-gun, Ino-cho, Terakawa-Ohdaru, 29 Jun. 2022, K. Fuse et al. JPN 12666 - 12668 sterile (FU!). The sterile specimens were identified using MIG-seq.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	description	Figs 19, 20	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	diagnosis	Diagnosis. Hosta samukazemontana is distinguished from H. polyneuronoides by anther size (3 mm vs. 3.5 - 4 mm), and inflorescences (deflected and curved upward at the top in contrast to erect and straight). Hosta samukazemontana is also similar to H. tosana in having inflorescences deflected and curved upward at the top, but distinguished from H. tosana by whitish perianths 4.5 - 5 cm long (in contrast to purple perianths 6.1 - 6.9 cm long) and whitish purple flower bracts 1.7 - 2.1 cm long (as opposed to whitish green peraiths 2.2 - 5.2 cm long).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	materials_examined	Type. Japan. Kochi Pref.: Tosa County, Hongawa Village, Mt. Kanpu, 25 Jul. 1983, Y. Kokami M 83 - 255 with flowers (holotype MBK 0175579!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 2 - 6 per ramet; blades oblong-ovate, elliptic-ovate, ovate, 7 - 22 cm long, 3.6 - 10.7 cm wide, 1.8 - 2.9 time longer than wide, thinly papery, glabrous on both surfaces, base cuneate, rounded, cordate, decurrent, slightly folded, apex acuminate, margin entire, veins in 6 - 12 pairs, smooth on the lower surface; petioles 3.2 - 14.5 cm long, narrowly winged, wings 0.1 - 0.2 cm wide, glabrous. Scape 20 - 43 cm long, terete, deflected and curved upward at the top. Raceme 7 - 9 cm long, 5 - 18 - flowered; flower bracts vivid (not withering) in anthesis, erect, whitish purple, oblong-lanceolate, boat-shaped, 1.7 - 2.1 long, 0.3 - 0.5 cm wide, membranous, glabrous, apex acuminate. Flowers 5 - 6 cm long; pedicels 1 - 1.8 cm long, glabrous. Perianths 4.5 - 5 cm long, purple-white outside, almost white inside, funnel-form, glabrous, 6 - lobed, tube abruptly dilated from apical 2 / 3, lobes narrowly triangular, 1 - 1.3 cm long, 0.7 - 0.8 cm wide, apex acute, three veins distinct inside. Stamens 6; filaments 4.8 cm long, white, free, glabrous, almost as long as perianth, upwardly curved at the distal part, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Ovary ellipsoid, 1 cm long, style 5 cm long, upwardly curved at the distal part, 0.5 cm exerted from perianth, glabrous, stigma capitate. Capsules green, cylindrical, 2.3 - 2.4 cm long, 0.4 cm wide, 3 - angled. Seeds 2.2 mm long, with wings ca. 4 mm long, blackish brown.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	distribution	Distribution and habitat. Japan (Ehime and Kochi Prefectures) (endemic). This species grows on rock cliffs in the vicinity of Mt. Kanpu in the central part of the Kochi Prefecture on Shikoku Island.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	etymology	Etymology. The specific epithet was derived from the old name of the type locality (Mt. Samukaze).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
8FF0D1002B84539C94449363650EA8C1.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Agawa-gun, Ino-cho, along Kamegamori Forest Road, 1660 m elev., 22 Jun. 2021, T. Yahara et al. JPN 6325 with flower buds (FU!). Ehime Pref.: Saijyo City, W slope of Mt. Kanpu, 1050 m elev., 20 Aug. 2020, T. Yahara et al. JPN 1380 - 1382 with fruit (FU!); Saijyo City, Mt. Kanpu 1600 m, 24 Jul. 2022, Se. Fujii JPN 15235 with flowers (FU!); Mt. Kanpu, June 19, 1943, S. Yamawaki s. n. with flowers (TI!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
7D582A1FC4935BD9AFDEA13B3EEC40D8.taxon	description	Fig. 10	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
7D582A1FC4935BD9AFDEA13B3EEC40D8.taxon	distribution	Distribution and habitat. Japan (Tokushima and Kochi Prefectures). The typical lineage of this species grows in rock crevices in the open habitats of riverbanks along the middle reach of Yoshino River. The upstream lineage (JPN 12523 - 12525, FJS 00006, FJS 00007) grows on the wet cliffs in the upper reaches of the Yoshino River.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
7D582A1FC4935BD9AFDEA13B3EEC40D8.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Nagaoka-gun, Ohtoyo-mura, Isodani, along Yoshino River, 8 Sep. 1958, T. Yamanaka 26222 with fruit (KYO!); Nagaoka-gun, Ohtoyo-cho, Higashidoi, Yoshino River, 4 Jul. 2002, H. Sasaoka FOK- 603540 with flowers (MBK 0170549!); left bank of Yoshino River, Okubo, Ohtoyo-cho, Nagaoka-gun, 26 Nov. 2017, A. Sakamoto et al. FOS- 017749 with fruit (MBK 0319724!); ditto, 18 Jul. 2004, M. Matsumoto et al. FOK- 067597 with flowers (MBK 0087490!); Tosa-gun, Ohkawa-mura, Takano, 24 Jun. 2022, K. Fuse et al. JPN 12523 - 12525 (FU!); Tosa-gun, Ohkawa-mura, Kawasaki, Jul. 25, 2023, Se. Fujii FJW- 50 - 1, 2 with flowers (MBK 0342373!, MBK 0342374!); Tosa-gun, Ohkawa-mura, Kogane Waterfall, 23 Jul. 2022, Se. Fujii JPN 15231, 15232, with flowers (FU!); ditto, Jul. 25, 2023, Se. Fujii FJW- 51 - 1 - 4 with flowers (MBK!); Nagaoka-gun, Motoyama-cho, in front of Kizenzan Park, 8 Aug. 1971, H. Takahashi & N. Fujita 214 with flowers and fruit (KYO!);. Tokushima Pref.: Miyoshi-gun, Oboke, 11 Apr. 2011, T. Yahara et al. JPN 4080, 4082 - 4087 sterile (FU!); ditto, 22 Jul. 1967, C. Abe 33157 with fruits (KYO!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0A67C87C4E2F5C1CB939FBFC8D14D1C6.taxon	description	Fig. 11	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0A67C87C4E2F5C1CB939FBFC8D14D1C6.taxon	materials_examined	Type. Japan. Ehime Pref., Mt. Higashi-akaishi, 7 Aug. 1957, T. Yamanaka 22475 (holotype KYO!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0A67C87C4E2F5C1CB939FBFC8D14D1C6.taxon	distribution	Distribution and habitat. Japan (Ehime and Kochi Prefectures) (endemic). This species grows in open habitats on the rocky ridgeline of Mt. Higashi-akaishi and its surrounding areas on Shikoku Island.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
0A67C87C4E2F5C1CB939FBFC8D14D1C6.taxon	materials_examined	Additional specimens examined. Japan. Ehime Pref.: Mt. Higashi-akaishi, 15 Jul. 1952, T. Yamanaka 8901 with flowers (KYO!); ditto (as Mt. Akaishi), 9 Jul. 1928, G. Koidzumi s. n. with flower buds (KYO!); ditto, 1650 m elev., 21 Jun. 2021, T. Yahara et al. JPN 6197, 6228, 6239, 6249, and 6252 with flower buds (FU!); ditto, 1640 m elev., 14 Jul. 1960, K. Tsuchiya 500 with flowers (KYO!); ditto, Mt. Hachimaki (a peak of Mts. Higashi-akaishi), 22 Jul. 2022, Se. Fujii JPN 9953, 31 Jul. 2022, Se. Fujii JPN 9954 with flowers (FU!); derived from Mt. Higashi-akaishi, cultivated in Makino Botanical Garden, 8 Apr. 2021, T. Yahara & Se. Fujii JPN 3939 - 3942 sterile (FU!); between Mt. Higashiakaishi and Mt. Hutatsudake, 1600 m elev., 8 Sep. 1961, G. Murata 14981 with fruit (KYO!); ditto, cultivated stock of G. Murata 14981, G. Murata s. n. with flowers (KYO!). Kochi Pref.: Mt. Shiraga (cultivated stock), T. Yahara & Se. Fujii JPN 3937, 8 Apr. 2021, sterile (FU!); ditto, Se. Fujii JPN 9955, 23 Jul. 2021 with flowers (FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	description	Figs 14, 15	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	diagnosis	Diagnosis. Hosta takiminazukiflora is similar to H. longipedicellata and H. minazukiflora. It is distinguished from H. longipedicellata by leaves smooth on the lower surface (vs. papillose), pedicels 1.4 - 2.5 cm long (compared to 2.5 - 3.3 cm long), and flower bracts being fresh during flowering (in contrast to being withering), and from H. minazukiflora by perianths with a distinct midvein (as opposed to three distinct veins) and pedicels 1.4 - 2.5 cm long (compared to 1.1 - 1.2 cm long).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	materials_examined	Type. Japan. Kochi Pref.: Tosa County, Ookawa village, Mt. Higashikado-yama, 22 Jul. 2007, with flowers, N. Inagaki et al. FOK- 080097 with flowers (holotype MBK 0189902!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 3 - 12 per ramet; blades oblong-ovate, 11 - 26.5 cm long, 3.6 - 10.5 cm, 2.1 - 3.3 times longer than wide, thinly papery, glabrous on both surfaces, base cuneate to obtuse, often decurrent, apex long acuminate, margin entire, veins in 6 - 10 pairs, smooth on the lower surface; petioles 3.4 - 22 cm long, narrowly winged, wing 0.1 - 0.6 cm wide, glabrous. Scape 8.6 - 28.5 cm long, terete. Raceme 5.9 - 18.5 cm long, 4 - 13 - flowered; flower bracts vivid (not withering) in anthesis, erect, purplish light green, light bluish purple, ovate oblong-lanceolate, boat-shaped, 2 - 4.7 cm long, 0.2 - 0.7 cm wide, papery, glabrous, apex acute to acuminate. Flowers not fragrant; pedicels 1.4 - 2.5 cm long, glabrous; perianths 4 - 5.7 cm long, funnel-form, pale white-purple to light bluish-purple outside, almost white to light pale purple inside, midveins more or less purplish, glabrous, 6 - lobed; tube dilated from apical 1 / 2, lobes triangular, 1.2 - 1.8 cm long, 0.5 - 1 cm wide, apex acute. Stamens 6, same or ca. 0.5 cm exerted from perianths; filaments 3.9 - 5.8 cm long, upwardly curved at the distal part, white, free, glabrous, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Ovary ellipsoid, 0.6 - 0.7 cm long, style 4.5 - 6.5 cm long, upwardly curved at the distal part, up to 1 cm exerted from perianth, glabrous, stigma capitate. Young capsules 2.8 cm long (for MBK 0087737).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	distribution	Distribution and habitat. Japan (Kochi Prefecture: Tosa County, endemic to Mt. Inamura, Mt. Higashikado, and the surrounding area). It grows on cliffs.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	etymology	Etymology. A specific epithet is derived from its habit of growing on rock cliffs (called ' taki' in Kochi dialect) and flowering in June (Minazuki).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
78121A0F0DA55E8A81A3FDD7731470E3.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Tosa County, Ookawa village, 900 m, 27 Jul. 2004, N. Inagaki et al. FOK- 067742 with flowers and young fruits (MBK 0087737!); Tosa County, Tosa Town, 620 m elev., 24 Jun. 2021, T. Yahara et al. JPN 6478 - 6486, 6490 with flowers (FU!); Agawa-gun, Ino-cho, Mt. Inamura, 1390 m elev., 2 Aug. 2019, Y. Oohira 14695 with flowers (MBK 0314655!); Agawa-gun, Ino-cho, north cliff of Mt. Inamura, 23 Jul. 2022, Se. Fujii JPN 15229, 15230 with flowers (FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	description	Fig. 16	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	diagnosis	Diagnosis. Hosta takiminazukiflora subsp. grandis is distinguished from subsp. Hosta takiminazukiflora takiminazukiflora in having scapes 31 - 36 cm long (compared to 8.6 - 28.5 cm long in subsp. Hosta takiminazukiflora takiminazukiflora), broader leaf blades (13 - 14 cm wide compared to 3.6 - 10.5 cm wide), cordate leaf base (vs. cuneate to obtuse), glaucous lower leaf surface, lateral veins 12 - 13 pairs (as opposed to 6 - 10 pairs), and petioles green, not dotted with purple.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	materials_examined	Type. Japan. Kochi Pref.: Tosa-gun, Tosa-cho, along Seto River, 24 Jun. 2021, T. Yahara et al. JPN 6487 with flowers (holotype FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	description	Description. Herbs perennial. Leaves basal, spiral, long petiolate, 3 - 5 per ramet; blades oblong-ovate, 22 - 25 cm long, 13 - 14 cm wide, 1.7 - 1.8 times longer than width, thinly papery, glabrous on both surfaces, base usually cordate, often decurrent, apex acuminate, margin entire, veins in 12 - 13 pairs, smooth or slightly papillose on the lower surface; petioles 35 - 38 cm long, narrowly winged, wing 0.3 cm wide, glabrous. Scape 31 - 36 cm long, terete. Raceme 12 - 15.5 cm long, 4 - 18 - flowered; flower bracts vivid (not withering) in anthesis, erect, purplish light green, oblong-lanceolate, boat-shaped, 2.8 - 4.3 cm long, 0.4 - 0.5 cm wide, papery, glabrous, apex acuminate. Flowers not fragrant; pedicels 1.9 - 2 cm long, pale purple, glabrous; perianths 3.8 - 4.9 cm long, funnel-form, glabrous, 6 - lobed, pale blue-purple outside, almost white to pale purple inside, midveins faintly purplish adaxially; tube dilated from apical 2 / 3, lobes oblong-triangular, 1.2 - 1.3 cm long, 0.7 - 0.8 cm wide, apex acute. Stamens 6, ca. 0.5 cm exerted from perianth; filaments 4.3 - 4.5 cm long, upwardly curved at the distal part, white, free, glabrous, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Ovary ellipsoid, 0.6 - 0.7 cm long, style 4.5 - 5.1 cm long, upwardly curved at the distal part, ca. 1 cm exerted from perianth, glabrous, stigma capitate. Capsules or seeds not observed.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	distribution	Distribution and habitat. Japan (Kochi Prefecture) (endemic). This species grows on soil near waterfalls, whereas subsp. Hosta takiminazukiflora takiminazukiflora grows on rock cliffs.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	etymology	Etymology. The specific epithet is derived from its larger plant size rather than that of the typical subspecies.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
55F71D71E399529B8E88A852E6AF67CE.taxon	materials_examined	Additional specimens examined. Japan. Kochi Pref.: Tosa-gun, Tosa-cho, along Seto River, 24 Jun. 2021, T. Yahara et al. JPN 6488 - 6489 with flowers (FU!); ditto, 24 Jun. 2022, K. Fuse et al. JPN 12518 - 12521 with flowers (FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
A398401389DD5AFFAE7BEB7404B6A7BA.taxon	materials_examined	Type. Japan. Prov. Awa (Ehime Pref.): tractu Myōtō, in oppido Kamomyō, n. d., J. Nikai 1377 (holotype TI, n. v.).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
1E8A6E780F305B34A986512789246DDF.taxon	description	Fig. 8	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
1E8A6E780F305B34A986512789246DDF.taxon	distribution	Distribution and habitat. Japan (Ehime, Kagawa, Kochi, and Tokushima Prefectures). This subspecies is distributed in the lower reaches of the Yoshino River, the middle and lower reaches of the Niyodo River, the headwater of the Shinjyo River (near the headwaters of the Shimanto River), and the lower reaches of the Shimanto River (Fig. 7). It grows in open habitats on rocks and soil along the riverbanks.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
1E8A6E780F305B34A986512789246DDF.taxon	materials_examined	Specimens examined. Japan. Ehime Pref.: Kamiukena-gun, Omogokei, 13 Aug. 1970, N. Fujita 269, with flowers (KYO!); Saijyo City, 450 m elev., 6 Aug. 1971, H. Takahashi & N. Fujita 273 with young flower buds (KYO!); Nii-gun, 500 m elev., 30 Aug. 1955, G. Murata 9233 with flowers (KYO!). Kagawa Pref.: Mt. Otaki, 24 Sep. 1961, S. Sakaguchi s. n. with flowers (KYO!). Kochi Pref.: Agawa-gun, Niyodogawa-cho, Kuki, 80 m elev., 18 Aug. 2020, T. Yahara et al. JPN 1255, 1258, 1263 with flowers (FU!); ditto, in the headwater of a tributary of Shinjyo River, 900 m elev., cultivated in Makino Botanical Garden, 8 Apr. 2021, T. Yahara et al. JPN 3929 sterile (FU!); Agawa-gun, Ino-cho, Kashiki, 40 m elev., 20 Aug. 2020, T. Yahara et al. JPN 1331 - 1333 with flowers (FU!); Takaoka-gun, Ochi-cho, Kataoka, 50 m elev., 20 Aug. 2020, T. Yahara et al. JPN 1375 - 1377 with flowers (FU!); ditto, Tokoroyama, 13 Sep. 1962, G. Murata 17106 and 17107 with flowers (KYO!); Hata-gun, Nakamura-cho, along the Shimanto River, 20 Aug. 1913, Z. Tashiro s. n. with flowers (KYO!). Tokushima Pref.: Miyoshi City, Shikino-kami, 90 m elev., 11 Apr. 2021, sterile, T. Yahara et al. JPN 4071, 4072, and 4075 (FU!); Ikeda-cho, along the Yoshino River, 16 Sep. 1971, C. Abe 43049 with flowers (KYO!); Miyoshi-gun, Minawa-mura, Kawasaki, along the Iya River, 8 Sep. 1958, T. Yamanaka 26270 with flowers (KYO!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
C4D4FDE9DB8E5426A02E07DD85B03A40.taxon	distribution	Distribution and habitat. Japan (Shikoku Island). This species is widely distributed in Shikoku and typically thrives on herbaceous slopes near villages. It is cultivated in Honshu but often naturalized when it escapes cultivation.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
C4D4FDE9DB8E5426A02E07DD85B03A40.taxon	materials_examined	Specimens examined. Japan. Kochi Pref.: Takaoka-gun, Kubokawa-cho, 8 Aug. 2001, Se. Fujii 8725 (JPN 10003, FU!), ditto, 23 Aug. 2002, Se. Fujii 9271 (JPN 10002, FU!); Cultivated at Makino Botanical Garden (32 specimens listed in the Suppl. material 1; FU!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
938D172DFD1C5E249B5166AD01316F33.taxon	description	Fig. 21, 22, 23, 24	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
938D172DFD1C5E249B5166AD01316F33.taxon	distribution	Distribution and habitat. Japan (Kochi and Tokushima Prefectures). This species grows on wet slopes, rocky cliffs, rocky riverbanks, and tree trunks in the elevations from 77 m to 1420 m.	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
938D172DFD1C5E249B5166AD01316F33.taxon	materials_examined	Specimens examined. Morphs corresponding to " var. Hosta tosana tosana ": Kochi Pref.: Locality not specified, cultivated in Tokyo, 1 Aug. 1943, Maekawa 7043 with flowers (TI; the cultivated plant may have been collected by T. Yoshinaga from the type locality, as this is the only specimen of this taxon deposited in TI, where Maekawa studied Hosta); Nagaoka-gun, Otoyo-cho, Mt. Kajigamori, 30 Apr. 2022, Se. Fujii JPN 12424 sterile (FU!); Kami City, Monobe Town, O-dochi, 500 m elev., 25 Jun. 2021, T. Yahara et al. JPN 6619 - 6620 with flowers (FU!); ditto, Go-o-do, 553 m elev., cultivated at Makino Botanical Garden, 8 Apr. 2021, T. Yahara & Se. Fujii JPN 3931 - 3933 sterile (FU!); ditto, 600 m elev., 25 Jun. 2021, T. Yahara et al. JPN 6614 & 6616 with flowers, JPN 6611 - 6613, 6615 sterile (FU!); Miyanose, 400 m elev., T. Yahara et al. JPN 6603 sterile (FU!); Befu, 550 m elev., cultivated at Makino Botanical Garden, 8 Apr. 2021, T. Yahara & Se. Fujii JPN 3934 sterile (FU!); Befu Valley, 700 m elev., 25 Jun. 2021, T. Yahara et al. JPN 6553 - 6555 with flowers (FU!); Aki City, 77 m elev., cultivated at Makino Botanical Garden, 8 Apr. 2021, T. Yahara & Se. Fujii JPN 3935 sterile (FU!); Aki-gun, Kitagawa-mura, Shima, along the Nahari River, 28 May 2022, K. Fuse et al. JPN 12653 sterile (FU!). Tokushima Pref.: Mt. Tsurugi, 14 Aug. 1931, Z. Tashiro s. n. with flowers (KYO!); ditto, below Minokoshi, 1400 m elev., 14 Aug. 1954, G. Murata 7977 with flowers (KYO!); ditto, 1420 m elev., 23 Jun. 2022, T. Yahara et al. JPN 12821 - 12823 sterile (FU!); Higashiiyayama-mura, Inter Sugeoi et Minokoshi, 800 m elev., 12 Aug. 1954, G. Murata 7814 & 7816 with flowers and young fruits (KYO!); Naka County, Wajiki Town, 14 Jul. 1965, S. Takafuji 216 with flowers (KYO!); ditto, Kizawa Village, 24 Jul. 1980, S. Takafuji 1442 with flowers (KYO!); ditto, Riu-toge in Miyahama-mura, 2 Jul. 1952, G. Murata 5752 with flowers (KYO!); ditto, Kizu Village, 24 Jul. 1974, S. Takafuji 950 with flowers (KYO!). Morphs corresponding to " var. Hosta tosana caput-avis ": Kochi Pref.: Aki-gun, Yanase, 25 Jun. 2020, K. Fuse et al. 12536 - 12539 sterile (FU!); Kami City, Monobe Town, Befu Valley, 25 Jun. 2021, T. Yahara et al. JPN 6524 - 6526 with flower buds, 6571 with flowers, 6572 with flower buds (FU!); Mt. Ishidate, 25 Jun. 2021, T. Yahara et al. JPN 6585 - 6587, 6593 - 6596 with flower buds (FU!); on the river bank of Yanase, 1000 m elev., 20 Jul. 1958, S. Hatusima 22009 A with wilted flowers (KAG 056576!); transplanted from Yanase, 29 Jun. 1960, S. Hatusima s. n. with flowers (KAG 056577!); ditto, 10 Jun. 1967, Hatusima s. n. with flowers (KAG 056578!). The Kagami River lineage: Kochi Pref.: Kochi City, Tosayama, Hirose, 85 m elev., 3 Oct. 2008, Hamaguchi & N. Shintani PRC- 00118 with fruits (MBK 0208327!); ditto, 14 Jul. 2001, K. Hosokawa et al. FOK- 001722 with flowers (MBK 0104375!); Kuwao, 130 m elev., 14 Jul. 2013, A. Sakamoto FOS- 004959 with flowers (MBK 0247214!, MBK 0247215!); from Kuwao to Tsuami, 27 Jul. 1968, N. Naruhashi & M. Wakabayashi 223 with flowers (KYO!); Namekawa, 30 m elev., 19 Aug. 2020, T. Yahara et al. JPN 1265 with fruit, JPN 1266 sterile, JPN 1267 sterile, JPN 1269 with fruit (FU!); Kajitani, 160 m elev., 29 July 2013, A. Sakamoto FOS- 005031 with young fruits (MBK 0247386!) and FOS- 005032 with young fruits (MBK 0247387!); Miyanokubo, 160 m elev., 14 Jul. 2013, A. Sakamoto FOS- 004958 with flowers (MBK 0247213!); ditto, 170 m elev., 14 Jul. 2013, A. Sakamoto FOS- 004957 with fruits (MBK 0247212!); Oh-ana Valley, 110 m elev., 19 Aug. 2020, T. Yahara et al. JPN 1298 sterile, JPN 1299 sterile, JPN 1307 with flowers (FU!); Kagami-mura, Kawaguchi, along the Kagami River, 9 Jul. 1958, T. Yamanaka 25548 with flowers (KYO!).	en	Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro, Suyama, Yoshihisa (2023): Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments. PhytoKeys 235: 137-187, DOI: http://dx.doi.org/10.3897/phytokeys.235.99140, URL: http://dx.doi.org/10.3897/phytokeys.235.99140
