taxonID	type	description	language	source
038C87C4FFF0FFD6BDAEFC3CFB916180.taxon	materials_examined	Material. Indonesia: Lombok, Gn. Rinjani N. P., c 1250 m, Mal. trap 13, 1 ♀ 13 – 26. iii. 1991, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD6BDAEFC3CFB916180.taxon	discussion	Remarks. Flagellum with 33 flagellomeres; 1 st flagellomere 5.0 × longer than wide. Hind femur red, black in distal third. It is otherwise a typical specimen.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD6BDAEFC3CFB916180.taxon	distribution	Distribution. Widespread in the Oriental region, new record for Indonesia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD5BDAEFB78FEB3662F.taxon	materials_examined	Material. Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, 1 ♂ x. 1995, Y. van Nierop & C. van Achterberg; Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 2, 1 ♂ 7. ii. – 28. iii. 1995, c 750 m, C. van Achterberg & Y. Yasir; Malaysia: SW Sabah, nr Long Pa Sia, Payaka-laba, 2 ♂♂ 25. xi. – 1. xii. 1987, c 1010 m, Mal. trap 16, C. van Achterberg (all in Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD5BDAEFB78FEB3662F.taxon	description	Description. ♂. Body length 5.5 – 7.0 mm. Head. Flagellum with 29 – 32 flagellomeres, bristle-shaped; 1 st flagellomere 3.8 – 4.6 × longer than wide. Tyloids indistinct. Temple short, strongly and roundly narrowed behind eye. OED 0.9 – 1.0 × and OOD 1.6 – 1.7 × ocellar diameter. Frons smooth ventrally, with microsculpture and fine sparse punctures dorsally. Face densely punctate, with transverse striation medially. Clypeus moderately convex, with sparse punctures; apical margin sharp and slightly impressed medially. Mandible long and narrow, with two teeth, ventral tooth much smaller than dorsal one and distinctly turned inwards (by 45 °). Malar space 1.1 × as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base, ± indented sub-basally. Mesosoma. stout. Antero-ventral margin of pronotum with distinct edge. Notaulus strongly impressed in frontal 3 / 4 of mesoscutum, with transverse ribs. Mesoscutum coarsely rugose-punctate; rugose apico-medially. Sternaulus distinctly impressed in frontal 0.7 of mesopleuron, with transverse ribs. Epicnemial carina low, dorsally reaching subtegular ridge. Mesopleuron and metapleuron coarsely rugose, sometimes ± striate. Scutellum moderately elevated, coarsely rugose, with complete lateral and apical carina, lateral carina with distinct denticular elevation at scutellar base. Propodeum completely carinated, coarse reticular rugose with strong blunt apophysis. Spiracle small and roundish. Area superomedia halfmoon-shaped, 1.4 × wider than long; costula reaching its middle. Area petiolaris transversally striate. Legs stout. Hind coxa densely punctate. Hind femur densely punctate, 2.7 – 2.9 × longer than wide. Claws simple. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1 × their diameter. Vein 1 cu-a interstitial or slightly postfurcal by 2 × its width. Metasoma. Petiole slightly wider than high. Postpetiole moderately widened, with microsculpture medially, punctate laterally; median field not separated. Gastrocoelus impressed, with fine longitudinal ridges. Thyridium transverse, about as wide as the interval between thyridia. 2 nd tergite about as long as wide. 2 nd and 3 rd tergite densely rugose-punctate; 4 th tergite densely punctate; following tergites with superficial punctures. Hypopygium unmodified; apical margin straight. Colour. black. Scape and pedicel red ventrally; flagellum black, 1 st flagellomere ± yellowish basally. Palps and mandible except teeth yellowish; clypeus reddish in ♂♂ from Sabah, black in ♂ from Sulawesi. Tegula reddish. Hind edge of pronotum and sometimes subtegular ridge ivory. In ♂ from Sulawesi ventral edge of pronotum and apophyses with ivory spots. Apical spot of scutellum reddish or ivory. Tergites reddish-brown to black, petiole reddish-yellow in ♂♂ from Sabah; apical margin of 6 th tergite and large median spot of 7 th tergite ivory. Legs red; in the ♂ from Sulawesi hind leg entirely blackish; in ♂♂ from Sabah hind femur brown in apical 0.2; hind tibia brown, with diffuse subbasal reddish ring and hind tarsus brown. Wings hyaline; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD5BDAEFB78FEB3662F.taxon	discussion	Remarks. There are slight colour differences between the available ♂♂ from Borneo and from Sulawesi, but their morphology is similar. They might represent different subspecies of this widespread species, but the ♀♀ from these isles are still unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF0FFD5BDAEFB78FEB3662F.taxon	distribution	Distribution. Widespread in the Oriental region, known from Sumatra (Riedel 2011), new records for Borneo and Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFD5BDAEFC15FE3763CD.taxon	type_taxon	Type species. Platylaboides javanicus nov. sp.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFD5BDAEFC15FE3763CD.taxon	diagnosis	Diagnosis. ♀. Small, body length 6.1 mm. Head. Flagellum slender, slightly lanceolate. Temple very short and strongly narrowed behind eye. Clypeus large, flat; apical margin slightly bulging medially (fig. 1 A). Mandible long, upper tooth blunt, lower tooth very small and slightly turned inwards (fig. 1 E). Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Stout. Pronotum without frontal angular prominence. Notaulus impressed at frontal 0.3 of mesoscutum. Sternaulus distinctly impressed. Epicnemial carina low, reaching the subtegular ridge dorsally. Scutellum moderately elevated, wider than long, with distinct lateral carina in frontal 0.7. Propodeum (fig. 1 B) completely carinated, with small but distinct apophysis; spiracle short-oval. Area superomedia wider than long, smooth. Legs rather stout. Claws simple. Areolet pentagonal. Vein 1 cu-a slightly antefurcal (fig. 1 C). Nervellus of hind wing slightly inclivous, intercepted in apical 0.8. Fore wing with sparse pilosity in basal half. Metasoma. Strongly amblypygous. Petiole slightly wider than high. Postpetiole moderately widened, coarsely rugose-punctate. Gastrocoelus large, almost square. Thyridium transverse, about as wide as the interval between thyridia (fig. 1 C). 2 nd and 3 rd tergites densely rugose-punctate. Ovipositor sheath slender, not extending behind metasomal apex. Ovipositor very thin, slightly bent upwards. Colour. Black with extensive ivory colour pattern.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFD5BDAEFC15FE3763CD.taxon	biology_ecology	♂ and hosts unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFD5BDAEFC15FE3763CD.taxon	discussion	Remarks. This genus belongs to the tribe Platylabini and runs to the genus Cyclolabus Heinrich in the key to World genera of Platylabini (Tereshkin 2009: 1432 – 1437). It is characterized by the unique form of the mandible with a long blunt dorsal tooth and very small ventral tooth, and the form of clypeus, completely carinated propodeum with distinct apophysis, strongly impressed large gastrocoelus and transverse thyridium which is as wide as the interval between thyridia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFDBBDAEF8B2FDF0606F.taxon	description	(Figs 1 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFDBBDAEF8B2FDF0606F.taxon	materials_examined	Holotype. (♀) Indonesia: Jawa Barat, Halimun Salak NP, Gunung Botol, 6.7266 ° S 106.484 ° E, 1700 m, 27 – Apr- 2016, A. Floren, 45 / 4469 ec, COI 658 [0 n] INDOBIOSYS-CCDB 25286 - C 05 (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFDBBDAEF8B2FDF0606F.taxon	description	Description. ♀. Body length 6.1 mm. Head. Flagellum with 27 flagellomeres, slender and slightly lanceolate; 1 st flagellomere 5.7 × longer than wide, about 12 th flagellomere square, widest flagellomeres 1.3 × wider than long. Temple very short, strongly and roundly narrowed behind eye, dorsally 0.2 × as long as eye. OED 1.2 × and OOD 1.2 × ocellar diameter. Frons smooth and shining, with few very fine punctures. Face rather densely punctate, smooth; inner eye margins divergent ventrally. Clypeus flat, large, 2.5 × wider than long, with sparse punctures; apical margin sharp and slightly bulging medially (fig. 1 A). Malar space 0.7 × as long as width of mandibular base. Gena with very sparse punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Side of pronotum smooth and shining, with few fine punctures at dorsal margin. Notaulus impressed in frontal 1 / 4 of mesoscutum. Mesoscutum with dense superficial punctures, shining. Sternaulus distinctly impressed in frontal 0.4 of mesopleuron. Epicnemial carina low, dorsally reaching subtegular ridge. Mesopleuron with rather dense punctures ventrally, smooth and shining dorsally, with few fine punctures below subtegular ridge. Metapleuron with rather dense punctures, shining; juxtacoxal carina present. Scutellum moderately elevated, 1.2 × wider than long, with rather dense punctures, strong lateral carina in basal 0.7. Propodeum (fig. 1 B) completely carinated, with small but distinct apophysis; spiracle small and short-oval. Area basalis trapezoid, wider than long. Area superomedia 1.3 × wider than long, almost smooth; costula reaching at 4 / 10 of its length. Area petiolaris and area posteroexterna transversally striated. Hind coxa densely punctate, without scopa. Hind femur densely punctate, 3.6 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2 × their diameter. Vein 1 cu-a slightly antefurcal by 1.0 × its width (fig. 1 D). Nervellus of hind wing slightly inclivous, intercepted in its apical 0.8. Fore wing covered with sparse setae in basal half, setae shorter than the interval between them. Metasoma. Strongly amblypygous. Petiole 1.3 × wider than high. Postpetiole moderately widened, coarse rugose-punctate, without latero-median carina; median field not separated. Gastrocoelus strongly impressed, large and almost square, with some longitudinal ridges. Thyridium transverse, about as wide as the interval between thyridia (fig. 1 C). 2 nd tergite 1.2 × wider than long. 2 nd to 3 rd tergites densely rugose-punctate, dull. Following tergites with superficial punctures, ± shining. 2 nd to 4 th sternites with median folds. Ovipositor sheath and ovipositor narrow. Colour. Black with extensive ivory marks. Scape and pedicel brownish, with ivory ventral spots; flagellum black, flagellomeres 5 – 13 with ivory rings. Palps, mandible except teeth, clypeus, face except black median stripe, wide orbits completely, frontal and upper margins of pronotum, tegula, subtegular ridge, large spot of ventral mesopleuron, scutellum and postscutellum, upper division of metapleuron, apical half of metapleuron and propodeum (except black area petiolaris), base of petiole and apical margin of postpetiole, basal half and apical margin of 2 nd tergite, basal band (widened laterally) and apical median spot of 3 rd tergite, 5 th to 7 th tergite almost completely, folds of 2 nd to 4 th sternites and apical margin of 5 th sternite ivory. Fore and mid coxae, all trochanters and fore and mid trochantelli ivory; hind coxa black, ivory dorsally and apically. Femora, tibiae and tarsi reddish; hind femur diffusely brownish in apical third; hind tibia and hind metatarsus ochreous, infuscate apically; hind tarsus otherwise black. Wings hyaline, pterostigma brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF3FFDBBDAEF8B2FDF0606F.taxon	distribution	Distribution. Only known from Java.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFDFFD9BDAEF98DFD11672B.taxon	description	(Figs 2 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFDFFD9BDAEF98DFD11672B.taxon	materials_examined	Holotype. (♀) Malaysia: Sabah, Kinabalu Park Head Q., 8 – 11. iii. 1987, c 1600 m, Malaise trap 1, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFDFFD9BDAEF98DFD11672B.taxon	description	Description. ♀. Body length 10.5 mm. Head. Flagellum with 40 flagellomeres, slender, almost filiform; 1 st flagellomere 3.7 × longer than wide, widest flagellomeres 1.2 × longer than wider. Temple rather long, distinctly and linearly narrowed behind eye (fig. 2 B). OED 1.0 × and OOD 1.1 × ocellar diameter. Frons smooth ventrally, transversely rugose-punctate dorsally. Face densely punctate, shining. Clypeus almost flat, with sparse punctures; apical margin sharp and slightly concave (fig. 2 A). Mandible long, with two teeth, ventral tooth smaller than dorsal one. Malar space 1.2 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Side of pronotum densely punctate dorsally, with irregular rugae ventrally. Notaulus indistinct. Mesoscutum finely and densely punctate, slightly shining. Mesopleuron densely punctate and finely striate ventrally, densely punctate dorsally; speculum smooth. Metapleuron rugose-punctate; juxtacoxal carina present. Scutellum moderately elevated, about as long as wide, with fine sparse punctures; without lateral carina. Propodeum (fig. 2 C) rugose, with distinct blunt apophysis; spiracle slit-shaped. Area basalis not carinated, without median tubercle. Area superomedia almost rectangular, 1.4 × longer than wide; costula absent. Hind coxa densely punctate, without scopa. Hind femur slender, densely punctate, 5.1 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 5 × their diameter. Vein 1 cu-a postfurcal by 3.0 × its width. Metasoma. Strongly amblypygous; hypopygium unmodified, with fine setae, reaching metasomal apex. 2 nd and 3 rd sternites with median folds. Postpetiole moderately widened, without latero-median carina; median field moderately elevated, finely rugose; lateral field punctate. Gastrocoelus impressed, without ridges. Thyridium very small, oblique, 0.2 × as wide as the interval between thyridia (fig. 2 D). 2 nd tergite as long as wide, finely rugose and slightly aciculate in basal half, granulate apically, ± dull. Following tergites finely granulate, shining. Ovipositor sheath not reaching behind metasomal apex. Colour. Black. Flagellum black; flagellomeres 8 – 12 brownish dorsally. Face and gena with dark reddish suffusion; frontal orbit with reddish-yellow stripe. Hind edge of pronotum, tegula, subtegular ridge, central spot of mesoscutum and scutellum reddish. Tergites dark reddish-brown; 7 th tergite with ivory median spot. Legs black; fore and mid femora and tibia reddish-yellow stripes frontally; hind tibia diffusely reddish-brown sub-basally. Wings hyaline; pterostigma blackish. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFDFFD9BDAEF98DFD11672B.taxon	discussion	Remarks. This new species runs to Achaius flavorufus Heinrich, 1965 in the key of the Oriental Achaius species (Heinrich 1965: 111 – 112), but differs from all known Oriental species by the strong reduction of pale coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFDFFD9BDAEF98DFD11672B.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFD9BDAEFD11FCD7662F.taxon	materials_examined	Material. Indonesia: W Java, Mt. Halimun-Salak NP, Cikaniki, 6.7486 ° S 106.536 ° E, 1121 m, 1 ♀ iv. 2016, leg. S. Schmidt (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFD9BDAEFD11FCD7662F.taxon	distribution	Distribution. Known from Sulawesi, new record for Java.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	description	(Figs 3 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	materials_examined	Holotype. (♀) Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, xi. 1994, Y. v. Nierop & C. van Achterberg, RMNH´ 95 (Leiden). Paratypes. (♂) Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, vii. 1994, Y. van Nierop & C. van Achterberg, RMNH´ 95; (2 ♀♀) Indonesia: N Sumatra, Aceh, Ketambe, 400 m, viii. 1995, Mal. trap, nr edge rainforest, leg. Y. van Nierop & C. van Achterberg, RMNH´ 99; (♀) Indonesia: N Sumatra, Aceh, Ketambe, 400 m, x. 1995, Mal. trap, nr edge rainforest, leg. Y. van Nierop & C. van Achterberg, RMNH´ 99; (♂) Indonesia: N Sumatra, Aceh, Ketambe, 400 m, v. 1995, Mal. trap, nr edge rainforest, leg. Y. van Nierop & C. van Achterberg, RMNH´ 99; (♂) Indonesia: N Sumatra, Aceh, Ketambe, 400 m, vi. 1995, Mal. trap, nr edge rainforest, leg. Y. van Nierop & C. van Achterberg, RMNH´ 99 (all in Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	etymology	Etymology. The species name refers to the largely reddish metasoma.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	description	Description. ♀: Body length 10.0 – 12.5 mm. Head. Flagellum with 38 – 41 flagellomeres, slender and slightly lanceolate; 1 st flagellomere 4.1 – 4.2 × longer than wide, widest flagellomeres 1.1 – 1.3 × wider than long. Temple short, strongly and roundly narrowed behind eye (fig. 3 B). OED 1.6 × and OOD 0.7 × ocellar diameter. Temple with fine sparse punctures. Frons with microsculpture and fine very sparse punctures. Face with sparse superficial punctures. Clypeus large, moderately swollen baso-medially, slightly concave centrally, with sparse punctures; apical margin sharp, slightly rounded (fig. 3 A). Mandible sickle-shaped, ventral tooth slightly smaller than dorsal tooth and completely turned inwards. Malar space 0.7 × as long as width of mandibular base. Gena with fine rather dense punctures. Genal carina reaching mandibular base. Mesosoma. Notaulus impressed in frontal 0.3 of mesoscutum. Mesoscutum finely rugose-punctate, dull. Mesopleuron and metapleuron with fine superficial punctures, shining. Juxtacoxal carina present. Scutellum moderately elevated, longer than wide, with complete lateral carina. Propodeum completely carinated (fig. 3 C), with blunt short apophysis. Area basalis with median ridge. Area superomedia hexagonal, c 1.5 × longer than wide, usually with median ridge; costula reaching at 4 / 10 of its length. Legs slender. Hind coxa densely finely punctate and pilose, without distinct scopa. Hind femur 4.1 – 4.2 × longer than wide. Claws simple. Areolet pentagonal; frontal distance of veins 2 rs-m and 3 rs-m 1 - 2 × their width. Vein 1 cu-a interstitial or antefurcal by 1.0 × its width. Metasoma. Slender, oxypygous. Postpetiole slightly widened, finely granulate but shining; median field not separated. Gastrocoelus indicated by longitudinal ridges. Thyridium transverse, separated from basal margin of 2 nd tergite by 2.0 × its length, 2.5 – 3.0 × wider than the interval between thyridia (fig. 3 D). 2 nd tergite 1.2 – 1.4 × longer than wide. All tergites very finely rugose-punctate and dull; 2 nd tergite rugose-punctate basally. Ovipositor sheath long and slender, reaching behind metasomal apex by length of 7 th tergite. Colour. Red. Scape black dorsally, red ventrally; flagellum black, flagellomeres 7 – 15 / 16 ivory. Clypeus and face yellowish or cream-yellow. Frons dorsally and temple frontally reddish-brown. Tergites reddish-yellow; postpetiole reddish-brown; 2 nd tergite basally and apically reddish-brown. Thyridium black. 3 rd tergite blackish basally, reddish-brown apically. Following tergites with diffuse reddish-brown suffusion. 7 th tergite with small yellowish or ivory apical spot. Legs red; mid tarsus brown; hind femur black, narrowly red basally; hind tibia black, sometimes with reddish basal spot; hind tarsus black. Wings hyaline; pterostigma brown. ♂. Body length 9.5 – 10.0 mm. Head: Flagellum with 36 flagellomeres, bristle-shaped; 1 st flagellomere 4.0 × longer than wide. Tyloids indistinct, on flagellomeres 10 – 16, long-oval, maximally 0.7 × as long as their flagellomeres. Malar space 0.7 × as long as width of mandibular base. Hind femur 4.0 – 4.3 × longer than wide. 2 nd tergite 1.4 – 1.6 × longer than wide. Apical margin of hypopygium slightly rounded. Structure otherwise as described for the ♀. Colour reddish. Flagellomeres 10 – 16 ivory dorsally. 2 nd tergite with black thyridia and black apical margin. Coloration otherwise as described for the ♀.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	discussion	Remarks. This new species is closely related to Allonotus rufus Cameron. It differs by wider thyridium, stouter 2 nd tergite, paler face and clypeus and a different coloration of metasoma.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFFFFDFBDAEFC15FD1C672B.taxon	distribution	Distribution. Only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF9FFDFBDAEFD11FCE76283.taxon	description	(Figs 4 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF9FFDFBDAEFD11FCE76283.taxon	materials_examined	Holotype. (♀) Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, iv. 1994, Y. van Nierop & C. van Achterberg, RMNH´ 95 (Leiden). Paratype. (♀) Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, viii. 1994, Y. van Nierop & C. van Achterberg, RMNH´ 95 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF9FFDFBDAEFD11FCE76283.taxon	description	Description. ♀: Body length 12 mm. Head. Flagellum with 40 flagellomeres, slender and almost bristle-shaped; 1 st flagellomere 4.8 × longer than wide, widest flagellomeres about square. Temple short, strongly and roundly narrowed behind eye (fig. 4 B). OED 1.5 × and OOD 0.7 × ocellar diameter. Temple with fine dense punctures. Frons with microsculpture and fine sparse punctures. Face with sparse superficial punctures. Clypeus large, moderately swollen baso-medially, slightly concave centrally, with sparse punctures; apical margin sharp, slightly rounded (fig. 4 A). Mandible sickle-shaped, ventral tooth slightly smaller than dorsal tooth and completely turned inwards. Malar space 0.7 × as long as width of mandibular base. Gena with fine rather dense punctures. Genal carina reaching mandibular base. Mesosoma. Notaulus impressed in frontal 0.4 of mesoscutum. Mesoscutum finely rugose-punctate, dull. Mesopleuron and metapleuron with fine superficial punctures, shining. Juxtacoxal carina present. Scutellum moderately elevated, longer than wide, with complete lateral carina. Propodeum completely carinated (fig. 4 C), with blunt short apophysis. Area basalis with median ridge. Area superomedia hexagonal, 1.5 × longer than wide, sometimes with median ridge; costula reaching at 3 / 10 of its length. Legs slender. Hind coxa densely finely punctate and pilose, without distinct scopa. Hind femur 4.1 – 4.5 × longer than wide. Claws simple. Areolet pentagonal; frontal distance of veins 2 rs-m and 3 rs-m 1.0 – 2.0 × their width. Vein 1 cu-a interstitial or postfurcal by 1.0 × its width. Metasoma. Slender, oxypygous. Postpetiole slightly widened, with some fine sparse punctures; median field not separated. Gastrocoelus indicated by some longitudinal ridges. Thyridium transverse, separated from basal margin of 2 nd tergite by c 2 × its length, 1.4 – 1.7 × wider than the interval between thyridia (fig. 4 D). 2 nd tergite 1.7 – 1.8 × longer than wide. All tergites very finely rugose-punctate and dull; 2 nd tergite rugose basally. Ovipositor sheath long and slender, reaching behind metasomal apex by length of 7 th tergite. Colour. Red. Scape black dorsally, red ventrally; flagellum black, flagellomeres 8 – 15 ivory dorsally. Frons dorsally and temple frontally blackish. Petiole black, postpetiole reddish in apical half. Tergites blackish; 2 nd and 3 rd tergites with apical yellowish bands; 6 th and 7 th tergites with large ivory median spots. Legs red; fore and mid tarsi ± black; hind femur black, narrowly red basally; hind tibia and tarsus black. Wings slightly yellowish; pterostigma brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF9FFDFBDAEFD11FCE76283.taxon	discussion	Remarks. This subspecies differs from Allonotus rufus rufus Cameron, 1907 from Borneo by its entirely red fore and mid femora and the mainly black hind femur.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF9FFDFBDAEFD11FCE76283.taxon	distribution	Distribution. This subspecies is only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF8FFDDBDAEF8AAFB70672B.taxon	materials_examined	Material. Indonesia: W Sumatra, Lahan basah, Andalas University, Padang, 0.933 ° S 100.455 ° E, 156 m, 2 ♂♂ 23. i. 2016, leg. H. Hamid, S 16036, BOLD: ACD 4550, INDOBIOSYS-CCDB 26088 - G 08 (ZSM)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF8FFDDBDAEF8AAFB70672B.taxon	discussion	Remarks. ♂. Body length 13 – 14 mm. Flagellum with 36 – 37 flagellomeres; 1 st flagellomere 2.3 × longer than wide, distal flagellomeres ± nodose. Tyloids on flagellomeres 8 / 9 – 18 / 19, long-oval, maximally almost as long as the flagellomere. OED 1.1 × and OOD 0.8 – 1.0 × ocellar diameter. Malar space 0.5 × as long as width of mandibular base. Hind femur 4.3 – 4.5 × longer than wide. Colour. Black. Flagellum black dorsally, brownish or reddish ventrally. Palps, mandible except teeth, clypeus, face, wide frontal orbit, narrow vertical orbit, wide outer orbit and gena, collar and upper margin of pronotum, tegula, subtegular ridge, two paramedian stripes of mesoscutum, large spot or stripe of ventral mesopleuron, prescutellar ridge, scutellum and postscutellum, spot on metapleuron, basal 2 / 3 of propodeum (sometimes except area basalis and area superomedia), apical bands on 1 st and 2 nd tergites and sometimes diffuse lateral spots of 3 rd tergite ivory. Fore and mid coxae except brown bases, all trochanters and fore and mid trochantelli ivory. Hind coxa brown, with dorsal ivory spot. Legs otherwise brown; fore and mid legs with ivory stripes frontally; hind femur with ivory stripe externally; hind tibia and sometimes hind metatarsus ivory in basal 2 / 3. Wings hyaline, pterostigma reddish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFF8FFDDBDAEF8AAFB70672B.taxon	distribution	Distribution. Widely distributed in the Oriental region, new record for Sumatra and Indonesia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	description	(Figs 5 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	materials_examined	Holotype. (♀) Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 225 m, 15. x. – 5. xi. 1989, C. van Achterberg, RMNH´ 89 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	etymology	Etymology. The species name refers to the mainly black coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	description	Description. ♀. Body length 20 mm. Length of fore wing 14 mm. Head. Flagellum with 44 flagellomeres, slightly lanceolate; 1 st flagellomere 2.5 × longer than wide, 12 th flagellomere square, widest flagellomeres 1.5 × wider than long. Temple strongly and roundly narrowed behind eye, punctate and granulate. OED 1.5 × and OOD 1.0 × ocellar diameter. Frons smooth ventrally, punctate dorsally. Face densely punctate and granulate, ± shining. Clypeus slightly convex, densely punctate, shining; apical margin blunt and straight (fig. 5 A). Mandible with two teeth, ventral tooth much smaller than dorsal one, slightly bent inwards. Malar space 0.7 × as long as width of mandibular base. Gena with coarse dense punctures. Genal carina reaching hypostomal carina far away from mandibular base. Mesosoma. Rather stout. Epomia present. Side of pronotum densely punctate dorsally, coarsely rugose ventrally. Notaulus shortly impressed at frontal margin of mesoscutum. Mesoscutum densely punctate and finely granulate, slightly shining. Mesopleuron densely and coarsely punctate, shining; speculum smooth. Epicnemial carina low, reaching frontal margin of mesopleuron dorsally. Sternaulus not impressed. Metapleuron rugose-punctate; juxtacoxal carina present only in apical half. Scutellum about as long as wide, with dense punctures; without lateral carina. Propodeum incompletely carinated (fig. 5 B), continuously sloping down apically, without apophysis; spiracle slit-shaped. Area basalis much wider than long, without median tubercle. Area superomedia about heart-shaped, slightly longer than wide, finely aciculate, apical carina weak; costula absent. Hind coxa densely punctate and pilose, with weak scopa apico-ventrally. Hind femur densely punctate, 3.9 × longer than wide. Claws simple, strongly bent apically (by 90 °). Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their diameter. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Stout and blunt apically, semi-amblypygous; hypopygium unmodified, with fine setae, almost as long as the distance of its apical margin to metasomal apex. 2 nd to 4 th sternites with median folds. Postpetiole strongly widened, basally with latero-median carina; median field moderately elevated, aciculate and apically punctate; lateral field punctate (fig. 5 C). Gastrocoelus impressed, almost smooth. Thyridium almost transverse, 0.8 × as wide as the interval between thyridia. 2 nd tergite 1.4 × wider than long. 2 nd and 3 rd tergites densely rugose-punctate; 2 nd tergite aciculate medially. 4 th tergite densely punctate, following tergites with superficial punctures, shining. Ovipositor sheath short, not extending behind metasomal apex. Colour. Black. Flagellum black, basal six flagellomeres ochreous, ventrally, flagellomeres 7 – 12 ivory dorsally. Palps, apical margin of clypeus, facial and frontal orbits and two paramedian spot below antennal sockets yellowish; spot on outer orbit and mandible basally red (fig. 5 A). 1 st to 3 rd tergites with transverse reddish-yellow stripes postero-laterally (fig. 5 C); 6 th and 7 th tergites reddish-brown. Coxae black; fore and mid coxae with apical yellowish spots; fore and mid legs otherwise reddish-yellow; fore and mid femur with brown stripes posteriorly; mid tarsus mainly blackish; hind femur brown dorsally, reddish ventrally; hind tibia reddish, with black external stripe; hind tarsus black. Wings strongly brownish infuscate; pterostigma dark reddish-brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	discussion	Remarks. This new species is characterized by a semi-amblypygous metasoma, stout metasomal apex and incompletely carinated propodeum. Its generic position is questionable. The area superomedia is heart-shaped and not distinctly elevated above the surrounding areas as in typical members of Amblyjoppa Cameron; it resembles the structure found in Coelichneumon Thomson. Due to the rather stout and blunt metasomal apex, the species runs to Uchidia Heinrich (synonymized with Naenaria Cameron by Uchida 1942) in the key of Heinrich (1934), but from typical Naenaria species it can be differentiated by its spindle-shaped metasoma and shorter hypopygium. I have included this species in the genus Amblyjoppa here, but the generic arrangement of the Oriental members of Ichneumoninae with continuously slanting propodeum, the Protichneumonini sensu Heinrich (1967), needs further investigation.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFFBFFC3BDAEFD16FD2464BB.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC3BDAEFE61FCE761D3.taxon	materials_examined	Holotype. (♂) Sumatera Barat, Lahan basah, Andalas University, Padang, 0.933 ° S 100.455 ° E, 156 m, H. Hamid, S 16042, COI 658 [0 n], BOLD: ACD 4550, INDOBIOSYS-CCDB 26088 - C 06 (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC3BDAEFE61FCE761D3.taxon	description	Description. ♂. Body length 15.5 mm. Flagellum with 38 flagellomeres; 1 st flagellomere 2.3 × longer than wide. Tyloids on flagellomeres 9 – 19, long-oval, maximally 0.7 × as long as the flagellomeres. Temple moderately and roundly narrowed behind eye. OED 1.1 × and OOD 1.2 × ocellar diameter. Malar space 0.6 × as long as width of mandibular base. Hind femur 4.2 × longer than wide. 2 nd tergite 1.3 × wider than long. Gastrocoelus strongly impressed, with longitudinal ridges. Thyridium transverse, 0.9 × as wide as the interval between thyridia. Colour. Black. Flagellum dark brown, with ivory ring. Ivory are palps, mandible except teeth, clypeus, face, wide frontal orbit, narrow vertical orbit, wide outer orbit and gena, collar and upper margin of pronotum, tegula, subtegular ridge, two short paramedian stripes of mesoscutum, large spot of ventral mesopleuron, scutellum and postscutellum, area dentipara of propodeum and wide apical band of postpetiole. Petiole black; 2 nd to 5 th tergites reddish; 6 th and 7 th tergites brownish. Legs brown; fore and mid coxae except brown bases, all trochanters and trochantelli and dorsal spot of hind coxa ivory. Legs otherwise brown; frontal stripes of all femora and fore and mid tibiae and tarsi ivory; hind tibia externally and hind metatarsus basal 2 / 3 ivory. Wings hyaline; pterostigma ochreous.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC3BDAEFE61FCE761D3.taxon	discussion	Remarks. This new subspecies differs from A. rufobalteata amoena Heinrich, 1968 from Myanmar by its smaller size, absence of ivory flagellar ring, and red 2 nd tergite without posterior ivory spots.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC3BDAEFE61FCE761D3.taxon	distribution	Distribution. This subspecies is only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	description	(Figs 6 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	materials_examined	Holotype. (♂) Malaysia: SW Sabah, nr. Long Pa Sia (East), 25. xi. – 8. xii. 1987, c. 1000 m, Mal. trap 4, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	etymology	Etymology: The species name refers to the strongly striated basal tergites.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	description	Description: ♂. Body length 15.5 mm. Head. Flagellum with 38 flagellomeres, bristle-shaped; 1 st flagellomere 2.6 × longer than wide. Tyloids on flagellomeres 7 – 17, long-oval, maximally 0.5 × as long as the flagellomeres. Temple rather short, moderately and roundly narrowed behind eye. OED 1.0 × and OOD 0.9 × ocellar diameter. Frons finely granulated, with rather dense punctures dorsally. Face rather densely punctate, medially and laterally slightly bulging. Clypeus almost flat, with sparse punctures; apical margin blunt, straight (fig. 6 A). Malar space 0.5 × as long as width of mandibular base. Mandible with two large teeth, ventral tooth moderately shorter than dorsal tooth, not turned inwards. Gena densely punctate. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus distinctly impressed in frontal 0.4 of mesoscutum. Mesoscutum coarsely punctate and smooth and shining between punctures. Mesopleuron densely punctate, shining, speculum smooth. Metapleuron coarsely punctate, partly rugose; juxtacoxal carina present. Scutellum moderately elevated, with highest point baso-medially and continuously slanting apically, about as long as wide, densely punctate, lateral carina in basal 0.3. Propodeum complete carinated (fig. 6 B), continuously slanting towards apex (as in Coelichneumon), without apophysis, covered with long setae laterally. Area basalis 1.7 × wider than long, without median tubercle. Area superomedia hexagonal, slightly wider than long; costula reaching at 7 / 10 of its length. Hind coxa densely punctate. Fore metatarsus without apical tooth. Hind femur with sparse punctures in ventral half, 4.2 × longer than wide. Claws simple. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their width. Vein 1 cu-a interstitial. Metasoma. Postpetiole moderately widened, punctate and finely aciculate, median field not separated. 2 nd tergite 1.2 × longer than wide. Gastrocoelus large, strongly impressed, smooth. Thyridium large, transverse, 2.0 × wider than the interval between thyridia (fig. 6 C). 2 nd and 3 rd tergites coarsely punctate and strongly striate medially; 4 th tergite with superficial punctures, distinctly striate medially. 2 nd to 4 th sternites with median folds. Apical margin of hypopygium distinctly elongate and almost pointed medially. Colour. Black. Ivory are palps, mandible basally, clypeus except brown apical margin, face, gena and outer orbit, frontal orbit, small spot of vertex, scape and pedicel ventrally, collar and upper margin of pronotum, spot of tegula, and subtegular ridge. 1 st to 3 rd tergites red (fig. 6 C), apical margin of 3 rd tergite narrowly brown. Following tergites blackish. Fore and mid coxae and trochanters ivory frontally, ± brown dorsally; hind coxa and trochanter black, ± ivory apically; fore and mid legs ivory frontally, black dorsally; hind femur reddish, infuscate basally and ± ivory apically; hind tibia brown, ivory externally; hind tarsus black. Wings with brownish suffusion in distal half; pterostigma black. ♀ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	discussion	Remarks. The generical position of this new species in the genus Amblyjoppa Cameron is questionable, since the ♀ is unknown. The structure of the propodeum resembles members of the genus Coelichneumon. The new ♂ is mainly characterized by the large and smooth gastrocoelus, very wide thyridium and strongly striated basal tergites.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE5FFC1BDAEFA89FD116603.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC1BDAEFCF9FC776123.taxon	materials_examined	Material. Malaysia: SW Sabah, nr Long Pa Sia, Payaka-laba, 1 ♀ 25. xi. – 1. xii. 1987, c 1010 m, Mal. trap 16, C. van Achterberg; Malaysia: SW Sabah, nr Long Pa Sia (West), Mal. trap 2, c 1020 m, 1 ♂ 25. xi. – 8. xii. 1987, Mal. trap 3, c 1050 m, 1 ♂ 25. xi. – 8. xii. 1987, Mal. trap 7, c 1200 m, 1 ♂ 25. xi. – 8. xii. 1987, C. van Achterberg (all Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC1BDAEFCF9FC776123.taxon	distribution	Distribution. Known from China and Myanmar, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC1BDAEFB19FCDC633B.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr. Danum Valley Field C, W 12, Malaise trap 12, c 200 m, 14 – 26. iii. 1987, C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC1BDAEFB19FCDC633B.taxon	discussion	Remarks. This specimen from Borneo differs slightly from the material described from Vietnam (Riedel 2023: 627 – 628) by its more extended cream yellow coloration: ventral spot of mesopleuron, small lateral stripe of mesoscutum beside wing base and posterolateral spot of propodeum yellow. Pterostigma reddish, not brown. Body length 8.5 mm. Flagellum with 25 flagellomeres. Malar space 0.7 × as long as width of mandibular base. Hind femur 3.1 × longer than wide. Vein 1 cu-a of fore wing slightly postfurcal by 1.0 × its width. Structure and coloration are otherwise as described for A. vietnamensis s. str. Riedel. In my opinion, it represents a new subspecies of this species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC1BDAEFB19FCDC633B.taxon	distribution	Distribution. This subspecies is only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC0BDAEF9E1FD24640B.taxon	materials_examined	Material. Indonesia: N. Sulawesi, Duoga-Bone N. P. ca 220 m, 0 ° 34´N 123 ° 54´E, 1 ♀ 10 – 18. xi. 1985, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE7FFC0BDAEF9E1FD24640B.taxon	discussion	Remarks. Flagellum with 29 flagellomeres, lanceolate; 1 st flagellomere 1.7 × longer than wide, 9 th flagellomere square, widest flagellomeres 2.2 × wider than long. Temple almost parallel behind eye, moderately narrowed apically. OED 1.7 × and OOD 1.5 × ocellar diameter. Pronotum medially with a weak longitudinal ridge. Hind femur very stout, 2.2 × longer than wide. Postpetiole smooth, with some sparse punctures. 2 nd tergite 1.4 × wider than long. Gastrocoelus moderately impressed, with some concentric ridges. Thyridium transverse, 0.8 × as wide as the interval between thyridia. Colour. Black with rich ivory coloration. Petiole basally, apical bands of postpetiole and 2 nd to 6 th tergites ivory, 7 th tergite mainly ivory medially. Flagellomeres 6 – 20 with ivory stripes. Pterostigma black. Coloration otherwise as described by Heinrich (1934: 220 – 221). Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFEF0FD2467DB.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 225 m, 1 ♀ 15. x. – 5. xi. 1989, Mal. trap 12, c 200 m, 1 ♂ 19. x. – 5. xi. 1989, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFEF0FD2467DB.taxon	discussion	Remarks. ♀: Body length 16 mm. Flagellum with 38 flagellomeres, moderately lanceolate; 1 st flagellomere 3.9 × longer than wide; widest flagellomeres 1.6 × wider than long. Flagellomeres 6 – 13 dorsally ivory. ♂: Body length 15.5 mm. Flagellum with 37 flagellomeres, bristle-shaped; 1 st flagellomere 3.7 × longer than wide. Tyloids absent. Hind femur 4.1 × longer than wide. 2 nd tergite 1.1 × longer than wide. Collar, small spot of subtegular ridge and apical stripe of postpetiole ivory; mesosoma and metasoma otherwise entirely black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFEF0FD2467DB.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFC80FC11606F.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 200 m, 1 ♂ 10 – 15. x. 1989, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFC80FC11606F.taxon	discussion	Remarks. The studied ♂ differs slightly from the description of Heinrich (1965: 222 – 223). Body length 17 mm. Flagellum with 39 flagellomeres, bristle-shaped; 1 st flagellomere 4.1 × longer than wide. Tyloids on flagellomeres 10 – 16, long-oval, central, maximally 0.5 × as long as the flagellomeres. Hind femur 4.7 × longer than wide. 2 nd tergite 1.2 × longer than wide. Colour. Black with yellow coloration. Yellow are palps, mandible except teeth, clypeus, face, gena, frontal orbit, scape ventrally, apical margin of propleuron, frontal and upper margins of pronotum, spot of tegula, subtegular ridge, large stripe on ventral mesopleuron, apical spot on metapleuron, two short paramedian stripes of mesoscutum, scutellum laterally and apically, postscutellum, petiole, basal bands of 2 nd to 4 th tergites and apical margins of 2 nd and 3 rd tergites, 7 th tergite mainly, 2 nd to 4 th sternites except brown lateral spots, hind margins of 5 th and 6 th sternites, parameres, fore and mid coxae and trochanters, apical spots on hind coxa and hind trochanter. Legs otherwise black with some reddish-brown suffusion; fore and mid femora, tibiae and tarsi with yellow stripes frontally, hind tibia with subbasal yellow ring. Wings slightly infuscate; pterostigma ochreous centrally, margins blackish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFC80FC11606F.taxon	distribution	Distribution. Known from Myanmar, new record for Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFA54FC9063AB.taxon	materials_examined	Material. Malaysia: SE Sabah, nr. Danum Valley Field C, c 150 m, WONO, Mal. trap 5, 1 1 ♂ 5 – 26. v. 1987, 1 ♀ 26. v. – 20. vi. 1987, leg. C. van Achterberg & D. Kennedy, 1 ♀ 1 ♂ 20. i. – 20. ii. 1988, 1 ♂ 19. iii. – 19. iv. 1988, 3 ♀♀ 1 ♂ 19. iv. – 22. v. 1988, leg. C. van Achterberg & T. Burghouts; Malaysia: Sabah, Kinabalu Park, Poring, Hot Spring, c 475 m, 1 ♂ 9 – 12. iii. 1987, Mal. trap 8, leg. C. van Achterberg (all Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC0BDAEFA54FC9063AB.taxon	distribution	Distribution. Known from Indonesia and Malaysia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC7BDAEF891FC3D66DF.taxon	type_taxon	Type species. Atanyjoppoides mandibulator nov. sp.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC7BDAEF891FC3D66DF.taxon	diagnosis	Diagnosis. Medium-sized. Head. Flagellum bristle-shaped, without distinct tyloids; distal flagellomeres ± nodose. Temple strongly and linearly narrowed behind eye (fig. 7 B). Clypeus slightly convex, much wider than long; apical margin sharp and moderately rounded (fig. 7 A). Mandible long and slender, strongly narrowed behind base (as in Benyllus Cameron), ventral tooth much smaller than dorsal one (fig. 7 C). Genal carina reaching hypostomal carina far from mandibular base, both carinae low. Mesosoma. Epomia present. Notaulus impressed in frontal 1 / 5 of mesoscutum. Mesoscutum densely rugose-punctate, ± dull. Epicnemial carina low, dorsal end not reaching frontal margin of mesopleuron or subtegular ridge. Juxtacoxal carina present. Scutellum pyramidal, about as long as wide, with dense coarse punctation and with lamelliform lateral carina in frontal 0.7 (fig. 7 D). Propodeum (fig. 7 E) short and gradually slanting toward apex (as in Coelichneumon Thomson); spiracle slit-shaped. Area basalis indicated as slightly elevated smooth area. Area superomedia small, about 2.0 × longer than wide. Anterior transverse carina present; posterior transverse carina absent. Apophysis absent. Legs slender. Fore metatarsus without apical tooth. Claws with some short basal teeth. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a interstitial. Metasoma. Petiole about as wide as high. Postpetiole moderately widened; median field roundly and ± hump-like raised (as in Cobunus Uchida) (fig. 7 F). Gastrocoelus strongly impressed, with fine longitudinal ridges. Thyridium slightly oblique, 1.1 – 1.5 × wider than the interval between thyridia. 2 nd to 5 th tergites with distinct denticular spines at posterolateral edges (fig. 7 G). 2 nd to 4 th sternites with median folds. Hypopygium unmodified, apical margin moderately rounded. Color. Black with extensive ivory coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE6FFC7BDAEF891FC3D66DF.taxon	discussion	Remarks. Due to the rounded and slanting structure of the propodeum, this new genus belongs to the Protichneumonini sensu Heinrich (1967) and Tereshkin (2009). It runs to Atanyjoppa Cameron in the key of Heinrich (1934) but differs by the genal carina reaching the hypostomal carina far away from mandibular base, different form of scutellum, hump-like median field of postpetiole and carination of the propodeum. In the key of Townes et al. (1961) it runs to Hedyjoppa Cameron but differs by the dentiform posterolateral extensions of the basal tergites, form of postpetiole and different carination of its propodeum. Although this genus seems to be widespread in the Oriental region, the corresponding ♀ and host records are unknown yet.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE1FFC5BDAEFB85FDAE6793.taxon	description	(Figs 7 A – G)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE1FFC5BDAEFB85FDAE6793.taxon	materials_examined	Holotype. (♂) Malaysia: SW Sabah, nr Long Pa Sia (West), c. 1020 m, 25. xi. – 8. xii. 1987, Mal. trap 2, C. van Achterbeg, RMNH´ 87 (Leiden). Paratypes. (♂) Malaysia: SE Sabah, nr. Danum Valley Field C, WON 1, Mal. trap 5, c 150 m, 23. viii. – 13. xi. 1987, C. van Achterberg, RMNH´ 87 (Leiden); (♂) Laos, Prov. Hua Phan, Phon Pan, Umg. Ort Ban Saleui, 20 ° 13´30 ´´ N 103 ° 59´26 ´´ E, 1350 – 1900 m, 27.06.2013, KJa, leg. C. Holzschuh & locals (Linz).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE1FFC5BDAEFB85FDAE6793.taxon	etymology	Etymology. The species name refers to the specialized form of mandible.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE1FFC5BDAEFB85FDAE6793.taxon	description	Description. ♂. Body length 11 – 13 mm. Head. Flagellum with 37 flagellomeres, slender, bristle-shaped; 1 st flagellomeres 3.8 – 5.0 × longer than wide, distal flagellomeres ± nodose. Temple short, strongly and almost linearly narrowed behind eye (fig. 7 B). OED 1.4 × and OOD 1.0 × ocellar diameter. Frons smooth ventrally, with distinct longitudinal or oblique striae dorsally. Face and clypeus densely punctate, ± shining. Face moderately bulging medially. Clypeus slightly convex, 1.7 – 1.8 × wider than long; apical margin sharp and moderately rounded (fig. 7 C). Mandible long and slender, with complete ventral lamella, strongly narrowed behind base, ventral tooth much smaller than dorsal one and slightly bent inwards. Gena densely punctate. Malar space 0.8 – 1.1 × longer than width of mandibular base. Mesosoma. Side of pronotum with rather superficial punctures dorsally, almost smooth ventrally. Epomia present. Notaulus impressed in frontal 1 / 5 of mesoscutum. Mesoscutum densely rugose-punctate, ± dull. Mesopleuron densely punctate; speculum large, smooth. Metapleuron densely punctate; juxtacoxal carina present. Scutellum pyramidal, about as long as wide, with dense coarse punctation and with lamelliform lateral carina in frontal 0.7 (fig. 7 D). Propodeum coarsely rugose-punctate; spiracle slit-shaped. Area basalis indicated as slightly elevated smooth area. Area superomedia small, about 2.0 × longer than wide; costula reaching its frontal 0.4. Area dentipara and area posteroexterna confluent (fig. 7 E). Apophysis absent. Area petiolaris with transverse striae. Legs slender. Fore metatarsus without apical tooth. Hind femur 5.1 – 5.4 × longer than wide. Claws with some short basal teeth. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a interstitial. Nervellus of hind wing reclivous, intercepted in apical 0.8. Metasoma. Postpetiole moderately widened; median field almost hump-like raised, coarsely rugose-punctate; lateral field densely punctate. 2 nd tergite 1.3 – 1.5 × wider than long. Gastrocoelus strongly impressed, with fine longitudinal ridges. Thyridium slightly oblique, 1.1 – 1.5 × wider than the interval between thyridia (fig. 7 F). 2 nd to 5 th tergites with distinct denticular spines at posterolateral edges (fig. 7 G). 2 nd to 6 th tergites coarsely rugose-punctate; 2 nd to 4 th tergites ± striate medially. 2 nd to 4 th sternites with median folds. Hypopygium unmodified, apical margin moderately rounded. Colour. Black with extensive ivory coloration. Flagellum black, central flagellomeres ± reddish-brown ventrally. Ivory are palps, mandible except teeth, labrum, clypeus, face, gena, outer orbit and frontal orbit up to lateral ocellus, propleuron, frontal and upper margins of pronotum, central round spot of mesoscutum, spot on tegula, subtegular ridge, epicnemium, wide ventral stripe and small dorso-apical spot of mesopleuron, upper division and central spot of metapleuron, apical 2 / 3 of scutellum, postscutellum, area posteroexterna and area dentipara of propodeum mainly, petiole, posterolateral spots on 1 st to 5 th tergites (medially confluent on 2 nd tergite), entire 7 th tergite and parameres. 1 st to 3 rd tergite sometimes with diffuse reddish-brown suffusion medially. Fore and mid coxae and trochanters ivory; hind coxa black or brown, with dorsal and ventral ivory spots. Fore and mid femora, tibiae and tarsi yellowish, with long brown stripes; hind femur reddish, sometimes diffusely yellow basally, blackish in apical 0.3 – 0.6; hind tibia blackish, with wide yellow subbasal ring; hind tarsus black. Wings hyaline; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE1FFC5BDAEFB85FDAE6793.taxon	distribution	Distribution. Borneo and Laos.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFD49FB4A6011.taxon	materials_examined	Material. Indonesia: C Sulawesi, nr Batui, Seseba, c 375 m, 3 ♂♂ 6 – 9. xi. 1989, Mal. trap 19, leg. C. van Achterberg; C Sulawesi, Lore-Lindu N. P., nr. Dongi-Dongi shelter, c 1100 m, 1 ° 15´S 120 ° 20´E, Mal. trap 2, 1 ♂ 4 – 9. xii. 1985, leg. C. van Achterberg; SW Sulawesi, Ciaker Alam Laiya, c 15 km S of Camba, c 400 m, 1 ♂ 28. xi. 1991, C. van Acternberg; Sula Islands, Mangole, near Buya, Mal. Trap 9, c 480 m, 1 ♀ 1 ♂ 12. x. – 2. xi. 1993, Mal. trap 11, 24 ♂♂ 12. x. – 2. xi. 1993, Mal. trap 12, c 495 m, 6 ♂♂ 12. x. – 2. xi. 1993, Mal. Trap 13, c 465 m, 1 ♀ 10 ♂♂ 13. x. – 2. xi. 1993, Mal. Trap 14, c 450 m, 1 ♂ 13. x. – 2. xi. 1993, Mal. trap 15, c 445 m, 3 ♂♂ 13. x. 2. xi. 1993, Mal. Trap 16, c 500 m, 5 ♂♂ 12. x. – 2. xi. 1993, C. van Achterberg; Malaysia: SE Sabah, nr. Danum Valley Field C, WO, c 150 m, Mal. trap 5, 1 ♂ 12. vii. – 2. viii. 1987, leg. C. van Achterberg & D. Kennedy (all Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFD49FB4A6011.taxon	discussion	Remarks. Heinrich (1934: 128 – 129) separated 4 different “ races ” from Sulawesi due to different colour pattern. In my opinion, all his subspecies are colour variants only. The specimen from Borneo represents the dark nominal form; the material from Sulawesi belongs to the var. centralis and makassariensis, the material from Sula Islands has a red ground colour, with only head and mesoscutum ± black (var. maculutus).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFD49FB4A6011.taxon	distribution	Distribution. Known from Borneo, Sulawesi and Sumatra, new record for the Sula Islands.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFAC6FB526398.taxon	materials_examined	Material. Indonesia: W Sumatra, Lahan basah, Andalas University, Padang, 0.933 ° S 100.455 ° W, 156 m, 1 ♀ 16. i. 2016, leg. H. Hamid, S 16042, COI 631 [0 n], BOLD ADE 1162, INDOBIOSYS- CCDB 26088 - D 09 (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFAC6FB526398.taxon	discussion	Remarks. ♀. Body length 8.5 mm. Flagellum with 39 flagellomeres, semi-bristle-shaped; 1 st flagellomere 2.4 × longer than wide, widest flagellomeres about square, apical flagellomere blunt. OED 1.1 × and OOD 1.4 × ocellar diameter. Hind femur 4.8 × longer than wide. 2 nd tergite 1.2 × wider than long. Gastrocoelus moderately impressed. Thyridium slightly oblique, 0.6 × as wide as the interval between thyridia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC5BDAEFAC6FB526398.taxon	distribution	Distribution. Widespread in the Oriental region, known from Java, new record for Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC4BDAEF940FD24649F.taxon	materials_examined	Material. Indonesia: C Sulawesi, Lore-Lindu N. P., nr. Dongi-Dongi shelter, c 1100 m, 1 ° 15´S 120 ° 20´E, Mal. trap 6, 1 ♂ 6 – 9. xii. 1985, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC4BDAEF940FD24649F.taxon	discussion	Remarks. ♂. Body length 10.5 mm. Body with dense rather long pilosity. Flagellum with 40 flagellomeres; 1 st flagellomere 3.0 × longer than wide. Tyloids on flagellomeres 9 – 18, long-oval, maximally 0.8 × as long as their flagellomeres. Head dense punctate. OED 1.6 × and OOD 1.5 × ocellar diameter. MI 0.7. Mesoscutum, mesopleuron and metapleuron densely punctate. Hind femur 4.1 × longer than wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rsm 2.0 × their width. Vein 1 cu-a postfurcal by 3.0 × its width. 2 nd tergite as long as wide. Gastrocoelus distinctly impressed. Thyridium 0.6 × as wide as the interval between thyridia. Colour. Black. Ivory are palps partly, side of clypeus, facial and frontal orbits, collar and upper margin of pronotum, spot of tegula, subtegular ridge, apical 1 / 3 of scutellum, apical band on 6 th tergite, entire 7 th tergite and paramere. Legs black; apical spot on fore and mid femora, frontal stripes on fore and mid tibiae and basal spot of hind tibia ivory. Wings almost hyaline; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE3FFC4BDAEF940FD24649F.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFF08FD246027.taxon	description	(Figs 8 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFF08FD246027.taxon	materials_examined	Material. Indonesia: N Sulawesi, Dumoga-Bone N. P., c 220 m, Mal. trap: Maze Toraut R., 0 ° 34´N 123 ° 54´E, 2 ♂♂ 1 – 9. xi. 1985, leg. C. van Achterberg; Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 2, c 750 m, 1 ♀ 7. ii. – 28. iii. 1995, Mal. trap 3, c 750 m, 2 ♂♂ 7. ii. – 28. iii. 1991, leg. C. van Achterberg & Y. Yasir (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFF08FD246027.taxon	description	Description. ♂. Body length 9.5 mm. Head. Flagellum with 37 flagellomeres, bristle-shaped; 1 st flagellomere 2.4 × longer than wide. Tyloids on flagellomeres 10 – 17, long-oval, maximally 0.8 × as long as their flagellomeres. Temple long, almost parallel behind eye, slightly narrowed apically (fig. 8 B). OED 1.1 × and OOD 1.2 × ocellar diameter. Temple coarsely punctate. Frons rugose-punctate dorsally, with median longitudinal ridge ventrally. Face and clypeus densely punctate. Clypeus slightly convex; apical margin blunt and almost straight (fig. 8 A). Mandible with two teeth, ventral tooth slightly smaller than dorsal one, not turned inwards. Malar space 0.7 × as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Collar of pronotum with small longitudinal median ridge. Notaulus impressed in frontal 1 / 6 of mesoscutum. Mesoscutum densely punctate, partly rugose-punctate. Mesopleuron densely punctate, partly rugose ventrally; speculum smooth. Metapleuron with finer dense punctures; juxtacoxal carina present. Scutellum moderately and roundly elevated, slightly wider than long, with lateral carina in basal 0.3 – 0.4. Propodeum completely carinated, with small apophysis (fig. 8 C). Area basalis with median tubercle. Area superomedia hexagonal, slightly wider than long; costula reaching its middle. Hind femur densely punctate, 3.9 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Postpetiole moderately widened, coarsely rugose-punctate, with blunt latero-median carina basally; median field elevated and distinctly separated (fig. 8 D). Gastrocoelus impressed, with longitudinal ridges. Thyridium small, 0.3 × as wide as the interval between thyridia. 2 nd tergite 1.2 × longer than wide. 2 nd to 4 th tergites coarsely rugose-punctate; following tergites with superficial punctures. Colour. Black. Flagellum black, basal flagellomeres ± reddish-brown. Ivory are palps, basal spot of mandible, clypeus, face except small brownish median spot, wide frontal orbit, scape and pedicel ventrally, collar and upper margin of pronotum, tegula, subtegular ridge, scutellum, apical spot of 6 th tergite, large median spot of 7 th tergite, and fore and mid trochanters. Coxae black, sometimes fore and mid coxae with apical yellowish spots; legs otherwise reddish-yellow; hind tibia slightly and diffusely infuscate apically, hind tarsus black. Wings almost hyaline; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFF08FD246027.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFA1CFC3B62CB.taxon	materials_examined	Material. Indonesia: N Sulawesi, Dumoga-Bone N. P. c 220 m, 0 ° 34´N 123 ° 54´E, Mal. trap: Maze Toraut R., 4 ♂♂ 1 – 9. xi. 1985, 1 ♂ 16 – 23. xi. 1985, leg. C. van Achterberg; SE Sulawesi, nr Sanggona, Base Camp, Gn. Watuwila, Mal. trap 5, c 200 m, 1 ♂ 12 – 15. x. 1989, 3 ♂♂ 15. x. – 5. xi. 1989, leg. C. van Achterberg; C Sulawesi, Lore-Lindu N. P., nr. Dongi-Dongi, c 1010 m, 1 ° 15´S 120 ° 20´E, Mal. trap 2, 1 ♀ 4 – 9. xii. 1985, leg. C. van Achterberg; Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 3 ♂♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong; C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♂ 21 – 31. x. 1989, Mal. trap 15, leg. C. van Achterberg; Sula Islands, Mangole, near Buya, Mal. trap 10, c 480 m, 1 ♂ 12. x. – 2. xi. 1993, Mal. trap 11, c 460 m, 1 ♂ 12. x. – 2. xi. 1993, Mal. trap 13, c 465 m, 1 ♂ 13. x. – 2. xi. 1993, leg. C. van Achterberg; Sula Islands, Mangole, Buluhaya, Mal. trap 18, c 90 m, 1 ♀ 6 – 23. iii. 1995, leg. C. van Achterberg & Y. Yasir; Sula Islands, Mangole, behind Mandafuhi Camp, 1 ♂ 4. iii. 1995, leg. Y. Yasir (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFA1CFC3B62CB.taxon	discussion	Remarks. ♂: Small, body length 6.0 – 6.5 mm. Flagellum with 31 flagellomeres. Tyloids on flagellomeres 11 – 16. Postpetiole smooth, without punctures. Meso- and metasoma including mesoscutum red, mesoscutum often ± infuscate. Head blackish, face red; side of clypeus, facial and frontal orbits yellow. Flagellomeres 9 – 15 with ivory rings. Legs red; hind tibia reddish or ± brown, with reddish subbasal spot; hind tarsus blackish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEDFFCBBDAEFA1CFC3B62CB.taxon	distribution	Distribution. Known from Sulawesi, new record for the Moluccas.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFF08FD2467E7.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 2, 2 ♂♂ 7. ii. – 28. iii. 1995, Mal. trap 3, c 750 m, 1 ♀ 6 ♂♂ 7. ii. – 28. iii. 1991, leg. C. van Achterberg & Y. Yasir (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFF08FD2467E7.taxon	discussion	Remarks. ♀. Body length 12 mm. Flagellum with 39 flagellomeres, moderately lanceolate; 1 st flagellomeres 3.8 × longer than wide, 12 th flagellomere square, widest flagellomere 1.4 × wider than long. OED 1.0 × and OOD 1.2 × ocellar diameter. Hind femur 4.4 × longer than wide. Areolet pentagonal, distance between veins 2 rs-m and 3 rs-m 2.0 × their width; vein 2 m-cu reaching areolet distad its middle. Vein 1 cu-a slightly antefurcal by 1.0 × its width. Postpetiole and 2 nd tergite densely and finely punctate; 2 nd tergite 1.0 × as long as wide. Gastrocoelus very small; thyridium indistinct. Colour red; mesoscutum blackish. Flagellum black, flagellomeres 5 – 16 with ivory dorsal stripes. Side of clypeus, facial and frontal orbits, gena, upper margin of pronotum, subtegular ridge, stripe on prescutellar ridge yellow. ♂. Smaller, body length 7 – 9 mm. Flagellum with 31 – 32 flagellomeres. Tyloids on flagellomeres 11 – 14 / 16, oval, maximally 0.5 × as long as their flagellomeres. Hind femur 4.5 × longer than wide. Colour as in ♀, but ivory stripes on flagellomeres 8 – 15, and additionally frontal margin of pronotum, face ± and ventral stripe on mesopleuron ivory.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFF08FD2467E7.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFCDCFC7460D7.taxon	materials_examined	Material. Indonesia: Sulawesi, nr Bantaeng, Borong Rappoa, Gn. Daulu, Mal. trap 19, c 690 m, 1 ♀ 4 – 24. iv. 1991, leg. C. van Achterberg; SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 225 m, 1 ♀ 15. x. – 5. xi. 1989, Mal. trap 12, c 200 m, 1 ♂ 19. x. – 5. xi. 1989, leg. C. van Achterberg; C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♂ 1 – 14. xi. 1989, Mal. trap 16, leg. C. van Achterberg; C Sulawesi, nr Batui, Seseba, c 375 m, 1 ♂ 6 – 9. xi. 1989, leg. C. van Achterberg; Sula Islands, Mangole, Buluhaya, Mal. trap 18, c 90 m, 6 – 23. iii. 1995, leg. C. van Achterberg & Y. Yasir; Sula Islands, Mangole, near Buya, Mal. trap 14, c 490 m, 2 – 22. iii. 1995, C. van Achterberg & Y. Yasir (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFCDCFC7460D7.taxon	discussion	Remarks. The ♂♂ differ slightly from the original description (Heinrich 1934: 159). Body length 9.5 – 11.0 mm. Flagellum with 34 – 37 flagellomeres; 1 st flagellomere 3.6 – 3.8 × longer than wide. Tyloids on flagellomeres 10 / 11 – 18 / 19, long-oval, maximally 0.7 × as long as their flagellomeres. OED 1.1 × and OOD 1.3 × ocellar diameter. Hind femur 3.9 × longer than wide. Colour as described for the ♀ (Heinrich 1934: 158), but flagellum black, ochreous ventrally; mesoscutum black in ♂♂ from Sulawesi, but with short yellow paramedian stripes in ♂♂ from Sula islands. Hind coxa red, yellowish dorsally; hind femur red, often slightly infuscate apically; hind tibia red, blackish in basal 0.1 and apical 0.2 – 0.3; hind tarsus black. Tergites red; postpetiole and 2 nd and 3 rd tergites with yellow apical bands or posterolateral stripes; 7 th tergite yellow; paramere ivory.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFCABDAEFCDCFC7460D7.taxon	distribution	Distribution. Only known from Sulawesi, new record for the Moluccas.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	description	(Figs 9 A – E)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	materials_examined	Holotype. (♀) Malaysia: SW Sabah, nr Long PA Sia (West), c 1010 m, 25. xi. – 8. xii. 1987, Mal. trap 1 b, RMNH´ 87, C. van Achterberg (Leiden). Paratypes. (♂) similar label (coll. Riedel); (♂) Malaysia: SE Sabah, nr. Danum Valley Field C, Malaise trap 1, c 150 m, 14 – 20. iii. 1987, C. van Achterberg, RMNH´ 87 (Leiden); (♂) Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, 7. iii – 15. iv. 1994, Y. van Nierop & C. van Achterberg, RMNH´ 95 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	etymology	Etymology. The species name refers to the mesoscutum having two paramedian ivory stripes.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	description	Description. ♀. Body length 9.3 mm. Head. Flagellum with 40 flagellomeres, moderately lanceolate; 1 st flagellomere 3.4 × longer than wide, about 14 th flagellomere square, widest flagellomeres 1.6 × wider than long. Temple very short, strongly and roundly narrowed behind eye (fig. 9 B). OED 1.1 × and OOD 1.2 × ocellar diameter. Frons transversely rugose-punctate dorsally. Face densely punctate, shining, slightly impressed above anterior tentorial pits and slightly swollen ventro-laterally. Clypeus almost flat, with sparse punctures apically; apical margin sharp and straight (fig. 9 A). Mandible long and distinctly narrowed behind base, with two teeth, ventral tooth smaller than dorsal tooth. Malar space 1.2 × as long as width of mandibular base. Gena with dense punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus indistinct. Mesoscutum densely rugose-punctate, dull. Mesopleuron and metapleuron rugose-punctate; speculum smooth; juxtacoxal carina present. Scutellum slightly elevated and slightly longer than wide, rugose-punctate; lateral carina complete, not extended or raised apically. Propodeum completely carinated (fig. 9 C), with distinct apophysis; spiracle slit-shaped. Area superomedia heart-shaped, 1.2 × longer than wide; costula reaching at apical 0.6 of its length. Hind coxa with fine dense punctures, without scopa. Hind femur densely punctate, 4.4 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 4.0 × their diameter. Vein 1 cu-a slightly antefurcal by 1.0 × its width. Metasoma. Oxypygous. Petiole about as wide as high. Postpetiole distinctly widened, without latero-median carina, densely punctate; median field not separated. Gastrocoelus weakly impressed, with fine ridges. Thyridium small, slightly transverse, 0.2 × as wide as the interval between thyridia (fig. 9 D). 2 nd tergite 1.2 × wider than long, densely rugose-punctate. 3 rd tergite finely rugose-punctate; 4 th tergite with superficial punctures apically. Ovipositor sheath narrow, reaching widely behind metasomal apex by combined length of 6 th and 7 th tergite (fig. 9 E). Colour. Black. Flagellum with ivory ring on flagellomeres 5 – 17. Ivory are palps, base of mandible, clypeus except brownish spot medially, face except M-shaped black median spot, gena and outer orbit, frontal orbit up to lateral ocellus, frontal and upper margins of pronotum, subtegular ridge, two short paramedian stripes on mesoscutum, spot on ventral mesopleuron, prescutellar ridge, apical scutellum and postscutellum, upper division and apical spot on metapleuron, spot in front of propodeal spiracles, posterolateral stripes on propodeum, apical bands of all tergites (shortly interrupted medially on 3 rd and 4 th tergites), fore and mid coxae and trochanters, dorsal spots on hind coxa and trochanter and entire hind tarsus. Legs reddish; hind femur black in apical 0.3; hind tibia yellowish, black in basal 0.1 and apical 0.3. Wings hyaline; pterostigma dark brown. ♂. Body length 7.2 – 9.4 mm. Flagellum with 35 – 38 flagellomeres; 1 st flagellomere 3.1 – 3.3 × longer than wide. Tyloids on flagellomeres 10 / 11 – 17 / 18, long-oval to linear, maximally 0.8 × as long as their flagellomeres. OED 1.0 × and OOD 1.1 × ocellar diameter. Scutellum ± pyramidal, rugose-punctate, with complete, slightly lamelliform lateral carina. Hind femur 4.1 – 4.7 × longer than wide. 2 nd tergite 1.3 – 1.4 × wider than long. Hypopygium unmodified; its apical margin rounded. Structure otherwise as described for the ♀. Colour. As in ♀, but differs by: ivory rings on flagellomeres 8 – 15; tyloids yellowish. Face entirely ivory; gena sometimes black; scape ivory ventrally. Mesoscutum with paramedian stripes in the ♂ from Sumatra, but with weak reddish paramedian spot in the ♂♂ from Borneo. Area spiracularis of propodeum black. Postpetiole with apical ivory band; 2 nd and 3 rd tergites with posterolateral ivory spots; 6 th and 7 th tergites and paramere entirely ivory. Hind femur red basally, ± infuscate centrally and apically.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	discussion	Remarks. This new species resembles Benyllus egregiscutellatus Heinrich, but differs by the ivory paramedian stripes on mesoscutum, entirely black area petiolaris and black hind tarsus in ♀. The scutellum is rounded in the ♀, but ± pyramidal in the available ♂♂. It might represent a subspecies of B. egregiscutellarus.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFECFFC8BDAEF98CFDD56043.taxon	distribution	Distribution. Borneo and Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	description	(Figs 10 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	materials_examined	Holotype. (♀) West-Sumatra, Maninjau lake, 0 ° 16´13.76 ´´ S 100 ° 13´57.96 ´´ E, 19. xi. 2015, leg. V. Gromenko (coll. Riedel). Paratype. (♀) similar label (coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	etymology	Etymology. The species name refers to the red-banded basal tergites.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	description	Description. ♀. Body length 8.8 – 9.6 mm. Head. Flagellum with 34 – 36 flagellomeres, slightly lanceolate; 1 st flagellomere 4.4 × longer than wide, about 14 th flagellomere square, widest flagellomeres 1.2 – 1.3 × wider than long. Temple very short, strongly and linearly narrowed behind eye, dorsally 0.1 × as long as eye (fig. 10 B). OED 1.0 × and OOD 0.8 × ocellar diameter. Frons shining, with some fine superficial punctures. Face rather densely punctate, shining, slightly impressed above anterior tentorial pits, laterally as side of clypeus slightly swollen (fig. 10 A). Clypeus with sparse punctures, apical margin sharp and straight. Mandible long, and distinctly narrowed behind base, with two teeth, lower tooth smaller. Malar space 1.0 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus indistinct. Mesoscutum finely and densely rugose-punctate, posterior margin ± with longitudinal striae, dull. Mesopleuron with fine superficial punctures, shining; speculum smooth. Epicnemial carina low, dorsally reaching frontal margin of mesopleuron. Metapleuron with rather dense superficial punctures; juxtacoxal carina present. Scutellum moderately elevated, slightly longer than wide, densely punctate and with complete lateral carina. Propodeum completely carinated, with distinct apophysis (fig. 10 C); spiracle slit-shaped. Area basalis trapezoid, wider than long and with median tubercle. Area superomedia dome-shaped, slightly longer than wide; costula reaching at apical 0.8 of its length. Hind coxa with dense punctures, without scopa; hind femur densely punctate, 4.3 – 4.5 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rsm and 3 rs-m 3.0 × their diameter. Vein 1 cu-a slightly antefurcal by 1.0 × its width. Metasoma. Oxypygous. Petiole about as wide as high. Postpetiole slightly widened, without latero-median carina; median field not separated, finely aciculate; lateral field striate and punctate. Gastrocoelus small, much longer than wide. Thyridium very small, pit-shaped (fig. 10 D). 2 nd tergite 1.1 × wider than long, with fine very dense punctures, dull. Following tergites with superficial punctures, ± shining. 2 nd to 5 th sternites with median folds. Ovipositor sheath narrow, reaching widely behind metasomal apex by length of 2 nd hind tarsomere. Colour. Head black; palp, clypeus, face, frontal orbit up to vertex (widened dorsally) ivory. Mandible reddish. Antenna black; scape ventrally and basal flagellomere red, flagellomeres 7 – 15 with ivory stripes. Mesosoma red; collar and spot of subtegular ridge ivory. Mesoscutum except reddish margins and spots on area externa of propodeum blackish. Petiole red; postpetiole blackish, with wide red apical band. 2 nd to 3 rd tergite black, with wide apical red bands; 4 th tergite red, with black basal spot; following tergites reddish. Ovipositor sheath black. Legs red; hind femur black in apical 0.4; hind tibia reddish-yellow, black in apical half; mid tarsus brown, hind tarsus black. Wings hyaline; pterostigma blackish, with paler margins. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	discussion	Remarks. This new species runs to Benyllus rufopictus Heinrich in his key (Heinrich 1934: 159 – 160), but differs from this (and other Indonesian) species by its reduced ivory coloration and black / red banded basal tergites.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEEFFCEBDAEFA39FD2A6697.taxon	distribution	Distribution. Only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE8FFCDBDAEFC4DFCA06457.taxon	materials_examined	Material. Indonesia: W Java, Gunung Botol, 6.729 ° S 106.491 ° E, 1490 m, 2 ♂♂ 24. iv. 2016, leg. X. Mengal & T. Burton (ZSM); Indonesia: W Bali, nr Negara, rainforest above Batuagung, c 575 m, 1 ♂ 4 – 13. xii. 1991, Mal. trap 1, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE8FFCDBDAEFC4DFCA06457.taxon	description	Description. ♂ (from Bali). Body length 8.5 mm. Head. Flagellum with 36 flagellomeres; 1 st flagellomere 3.6 × longer than wide. Tyloids on flagellomeres 11 – 19, long-oval, maximally 0.8 × as long as their flagellomeres. Temple very short, strongly and linearly narrowed behind eye, dorsally 0.1 × as long as eye. OED 0.9 × and OOD 1.3 × ocellar diameter. Frons densely punctate, ± dull. Face densely punctate, shining. Clypeus with dense punctures, impressed around anterior tentorial pit; apical margin sharp and slightly concave. Mandible long and distinctly narrowed behind base, with two teeth, lower tooth smaller. Malar space 1.2 × as long as width of mandibular base. Gena with fine punctures. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus indistinctly impressed frontally. Mesoscutum densely punctate, dull. Ventral half of mesopleuron densely rugose-punctate, with dense superficial punctures dorsally; speculum smooth. Metapleuron densely rugose-punctate; juxtacoxal carina present. Scutellum moderately elevated, slightly longer than wide, densely punctate and with complete lateral and apical carina. Propodeum completely carinated, with distinct apophysis; spiracle slit-shaped. Area basalis with median tubercle. Area superomedia heart-shaped, 1.4 × wider than long; costula reaching at apical 0.8 of its length. Hind coxa with dense punctures. Hind femur 4.6 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 3.0 × their diameter; vein 2 m-cu slightly distad its middle. Vein 1 cu-a interstitial. Metasoma. Postpetiole moderately widened, without latero-median carina, densely punctate but with reduced punctuation medially; median field not separated. Gastrocoelus small, much longer than wide. Thyridium indistinct. 2 nd tergite 1.1 × wider than long, densely punctate, dull. 3 rd tergite densely punctate, but ± shining. Following tergites with superficial punctures, shining. Hypopygium unmodified, apical margin straight. Colour. Black. Flagellum black, with ivory ring of flagellomeres 10 – 16; tyloids yellowish. Ivory are palps, mandible except teeth, clypeus and face except median brown stripe, frontal orbit up to vertex, ventral spot of scape, frontal margin and hind half of upper margin of pronotum, tegula, subtegular ridge, large spot of ventral mesopleuron, apical 2 / 3 of scutellum, posterolateral stripe of propodeum around apophysis, posterolateral spots of 1 st to 3 rd tergites, narrow apical margins of 2 nd to 5 th tergites, large median spots of 6 th and 7 th tergites and parameres. Legs reddish; fore and mid coxae ivory; hind coxa brownish dorsally; hind femur red, diffusely infuscate in apical third; hind tibia black, with ochreous basal spot; mid and hind tarsi ivory, with brown distal tarsomeres. Wings slightly infuscate; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFE8FFCDBDAEFC4DFCA06457.taxon	distribution	Distribution. Known from Java, new record for Bali.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEBFFCDBDAEFE0DFD32625B.taxon	description	(Figs 11 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEBFFCDBDAEFE0DFD32625B.taxon	materials_examined	Holotype. (♀) Indonesia: Sumbawa, nr Pancasila, Gn. Tambora, Mal. trap 4, c 1090 m, 22. ix. – 2. x. 1993, C. van Achterberg, RMNH´ 93 (Leiden). Paratype. (♀) Indonesia: Sumbawa, nr Pancasila, Gn. Tambora, Mal. trap 8, c 500 m, 27. ix. – 2. x. 1993, C. van Achterberg, RMNH´ 93 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEBFFCDBDAEFE0DFD32625B.taxon	description	Description. ♀. Body length 7.0 – 8.0 mm. Head. Flagellum with 36 – 38 flagellomeres, slightly lanceolate; 1 st flagellomere 3.6 – 3.7 × longer than wide, about 12 th flagellomere square, widest flagellomeres 1.4 × wider than long. Temple very short, strongly and roundly narrowed behind eye. OED 0.9 × and OOD 0.9 × ocellar diameter. Frons finely rugose, with some superficial punctures. Face densely punctate, shining, weakly impressed above anterior tentorial pits, latero-ventrally slightly swollen. Clypeus almost flat, with sparse punctures; apical margin sharp and slightly concave (fig. 11 A). Mandible long and distinctly narrowed behind base, with two teeth, ventral tooth smaller than dorsal tooth. Malar space 0.9 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus indistinct. Mesoscutum finely and densely rugose-punctate, dull. Mesopleuron and metapleuron with fine dense striated punctuation; speculum partly smooth; juxtacoxal carina present (fig. 11 B). Scutellum slightly elevated, about as long as wide, with dense punctures and complete lateral carina. Propodeum completely carinated, with distinct apophysis (fig. 11 C); spiracle slit-shaped. Area superomedia heart-shaped, about as long as wide; costula reaching at apical 0.7 of its length. Hind coxa with fine dense punctures, without scopa. Hind femur densely punctate, 4.4 – 4.5 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 4 – 5 × their diameter; vein 2 m-cu distad its middle. Vein 1 cu-a slightly antefurcal by 1.0 × its width. Metasoma (fig. 11 D). Oxypygous. Petiole about as wide as high. Postpetiole slightly widened, without latero-median carina, densely punctate; median field not separated. Gastrocoelus very small, indicated as longitudinal ridge. Thyridium very small, pit-shaped. 2 nd tergite 1.1 × wider than long, finely and densely rugose-punctate, surface ± smoothened in apical 1 / 4. 3 rd tergite with fine punctures in basal half, with very superficial punctures apically. Ovipositor sheath narrow, reaching widely behind metasomal apex by length of 7 th tergite. Colour. Black. Flagellomeres 5 – 14 ivory dorsally. Palps, base of mandible, sometimes side of clypeus, facial and frontal orbits up to vertex, gena and outer orbit, frontal margin and hind edge of pronotum, tegula, subtegular ridge, spot on ventral mesopleuron, scutellum and postscutellum, apical spot on metapleuron confluent with spot around apophysis, triangular spots on postpetiole and 2 nd to 3 rd (in HT) or 4 th (in PT) tergites, narrow apical margins of 4 th and 5 th tergites and apical spots on 6 th and 7 th tergites ivory (fig. 11 D). Legs mainly red; fore and mid coxae with dorsal ivory spots; hind leg entirely red in HT; hind tibia blackish with reddish base and hind tarsus black in PT. Wings hyaline; pterostigma dark brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEBFFCDBDAEFE0DFD32625B.taxon	discussion	Remarks. This species differs from Benyllus satageus (Tosquinet) from Java by its red or black hind tarsus (ivory in B. satageus) and ivory posterolateral spots on 3 rd tergite (black in B. satageus). It might represent a subspecies of B. satageus.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEBFFCDBDAEFE0DFD32625B.taxon	distribution	Distribution. Only known from Sumbawa.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEAFFCCBDAEF890FD2462AF.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Ciaker Alam Laiya, c 15 km S of Camba, c 400 m, 1 ♀ 18. xi. 1991, leg. C. van Achterberg (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFEAFFCCBDAEF890FD2462AF.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD5FFF2BDAEFA3EFD1167E7.taxon	description	(Figs 12 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD5FFF2BDAEFA3EFD1167E7.taxon	materials_examined	Holotype. (♂) Malaysia: SW Sabah, nr. Long Pa Sia (West), 25. xi. – 8. xii. 1987, c. 1020 m, Mal. trap 2, C. van Achterberg, RMNH´ 87 (Leiden). Paratypes. (2 ♂♂) similar labels (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD5FFF2BDAEFA3EFD1167E7.taxon	etymology	Etymology. The species name refers to the largely ivory frons.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD5FFF2BDAEFA3EFD1167E7.taxon	description	Description. ♂. Body length 11 mm. Head. Flagellum with 36 – 38 flagellomeres, very slender and bristle-shaped; 1 st flagellomere 3.8 × longer than wide. Tyloids absent. Temple moderately and roundly narrowed behind eye (fig. 12 B). OED 1.4 × and OOD 0.9 × ocellar diameter. Frons with sparse punctures, shining. Face and clypeus densely punctate. Clypeus large, almost flat; apical margin sharp, almost straight (fig. 12 A). Mandible sickle-shaped, ventral tooth absent. Malar space 0.6 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Notaulus impressed in frontal 1 / 3 of mesoscutum. Mesoscutum densely punctate, ± dull. Mesopleuron dense punctures and some striae posteriorly, speculum large and smooth. Metapleuron rugose-punctate; juxtacoxal carina present. Scutellum pyramidal, slightly longer than wide, densely rugose-punctate, lateral carina strong in basal 0.7 – 0.8. Propodeum completely carinated, rugose, with short tooth-like apophysis (fig. 12 C); spiracle slit-shaped. Area basalis wider than long. Area superomedia 1.3 × wider than long; costula reaching its middle. Hind coxa with fine dense punctures. Hind femur 4.5 × longer than wide. Claws simple. Areolet almost pointed frontally. Vein 1 cu-a slightly antefurcal, interstitial or slightly postfurcal. Metasoma. Postpetiole slightly widened, without latero-median carina; median field not separated, finely rugose. 2 nd tergite 1.1 × wider than long. Gastrocoelus slightly impressed, with fine ridges. Thyridium indistinct, 0.3 × as wide as the interval between thyridia. 2 nd and 3 rd tergites finely and densely rugose-punctate, dull; following tergites with superficial punctures, ± shining. Hypopygium unmodified; apical margin straight. Colour. Black. Ivory are dorsal stripes on flagellomeres 7 – 14 / 15, palps, mandible basally, spot of gena, clypeus and face, frons except black median stripe, scape ventrally, collar and upper margin of pronotum, tegula mainly, subtegular ridge, apical 2 / 3 of scutellum and postscutellum, usually narrow apical margin of 6 th tergite, entire 7 tergite, hypopygium and paramere. 1 st and 2 nd tergites reddish-yellow. Coxae black, fore and mid coxae with ivory spots; fore and mid trochanters ivory; fore and mid femora and tibiae reddish-yellow; fore tarsus distally and mid tarsus entirely black; hind trochanter and femur brown or black; hind tibia yellowish, black in basal 0.1 and apical 0.2 – 0.3; hind tarsus black, 4 th and 5 th hind tarsomeres ivory. Wings slightly yellowish; pterostigma brown. ♀ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD5FFF2BDAEFA3EFD1167E7.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD4FFF1BDAEFCDDFD1165C3.taxon	description	(Figs 13 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD4FFF1BDAEFCDDFD1165C3.taxon	materials_examined	Holotype. (♀) Malaysia: SW Sabah, nr. Long Pa Sia (East), 25. xi. – 7. xii. 1987, c. 1000 m, Mal. trap 5, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD4FFF1BDAEFCDDFD1165C3.taxon	etymology	Etymology. The species name refers to the mainly black coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD4FFF1BDAEFCDDFD1165C3.taxon	description	Description. ♀. Body length 13 mm. Head. Flagellum with 40 flagellomeres, very long and slender; 1 st flagellomere 4.6 × longer than wide; all flagellomeres much longer than wide. Head covered with dense grey setae. Temple rather long, moderately and roundly narrowed behind eye (fig. 13 B). OED 1.4 × and OOD 0.3 × ocellar diameter. Frons rather densely punctate, ± dull. Face punctate, centrally with some longitudinal striae. Clypeus large, punctate, almost flat, but ± impressed around anterior tentorial grooves; apical margin sharp, slightly bulging medially (fig. 13 A). Mandible sickle-shaped, ventral tooth absent. Malar space 0.5 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Covered with dense setae. Side of pronotum punctate dorsally, with some coarse striae at posterior margin. Notaulus strongly impressed in frontal 1 / 4 of mesoscutum, with some transverse ribs. Mesoscutum densely punctate, ± dull. Sternaulus strongly impressed in frontal half of mesopleuron. Mesopleuron with coarse longitudinal striae in dorsal 2 / 3, coarsely rugose ventrally, speculum finely punctate. Metapleuron densely rugose-punctate; juxtacoxal carina present. Scutellum strongly elevated (as in Eccoptosage Kriechbaumer), about as long as wide, densely rugose-punctate, lateral carina strong and slightly lamelliform widened apically. Propodeum completely carinated, with strong tooth-like apophysis (fig. 13 C); spiracle slit-shaped. Area basalis wider than long. Area superomedia hexagonal, as long as wide; costula reaching its middle. Hind coxa with fine dense punctures, without scopa. Legs slender. Hind femur 5.1 × longer than wide. Claws simple. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their diameter. Vein 1 cu-a slightly antefurcal by 1 × its width. Metasoma. Strongly amblypygous, slender. Postpetiole slightly widened, without latero-median carina; median field not separated, with sparse fine punctures; lateral field with larger dense punctures (fig. 13 D). 2 nd tergite about as long as wide. Gastrocoelus not impressed. Thyridium very small, 0.2 × as wide as the interval between thyridia (fig. 13 D). 2 nd and 3 rd tergites finely and densely rugose-punctate, dull; following tergites with superficial punctures, ± shining. Ovipositor sheath slightly extending behind metasomal apex. Colour. Black with weak metallic shine. Ivory are dorsal stripes on flagellomeres 9 – 12 / 13, palps, tips of apophysis, small apical spot of 5 th tergite and wide apical bands of 6 th and 7 th tergites, apical 0.2 of hind metatarsus and all following hind tarsomeres. Frontal orbit with small reddish-yellow spot. Mandible reddish medially. Legs black; fore and mid legs ± brownish frontally. Wings slightly infuscate; pterostigma black. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD4FFF1BDAEFCDDFD1165C3.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF7BDAEFF08FD116088.taxon	description	(Figs 14 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF7BDAEFF08FD116088.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr. Danum Valley Field C, c 150 m, Mal. trap 6, 15 – 19. iii. 1987, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF7BDAEFF08FD116088.taxon	description	Description. ♀. Body length 10.5 mm. Head. Flagellum with 37 flagellomeres, very long and slender; 1 st flagellomere 3.5 × longer than wide; all flagellomeres much longer than wide. Temple rather long, shortly parallel behind eye, moderately and roundly narrowed apically (fig. 14 B). OED 1.7 × and OOD 1.4 × ocellar diameter. Frons smooth, with some lateral punctures. Face rather densely punctate. Clypeus large, punctate, almost flat; apical margin blunt, slightly bulging medially (fig. 14 A). Mandible sickle-shaped, ventral tooth absent. Malar space 0.8 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Covered with short setae. Side of pronotum almost smooth, with some coarse striae ventrally. Notaulus strongly impressed in frontal 0.3 of mesoscutum, with some transverse ribs. Mesoscutum densely punctate, ± dull. Mesopleuron rugose-punctate, speculum large, smooth and shining. Metapleuron densely rugose-punctate; juxtacoxal carina present. Scutellum pyramidal, slightly longer than wide, coarsely rugose, lateral carina complete. Propodeum completely carinated, but costula obsolete laterally, with rather short tooth-like apophysis (about as long as wide at base); spiracle slit-shaped. Area basalis present, distinctly impressed. Area superomedia sickle-shaped, 1.6 × wider than long; anterior transverse carina reaching widely behind its middle. Hind coxa with fine dense punctures, without scopa. Hind femur 4.1 × longer than wide. Claws simple. Areolet pointed frontally. Vein 1 cu-a antefurcal by 2.0 × its width. Metasoma (fig. 14 C). Strongly amblypygous, slender. Postpetiole moderately widened, without latero-median carina; median field slightly elevated, with fine striae and some punctures; lateral field with dense punctures. 2 nd tergite 1.3 × wider than long. Gastrocoelus slightly impressed. Thyridium transverse, 0.7 × as wide as the interval between thyridia. 2 nd to 4 th tergites densely rugose-punctate, dull; following tergites with superficial punctures, ± shining. Ovipositor sheath slightly extending behind metasomal apex. Colour. Black with ivory coloration. Ivory are dorsal stripes on flagellomeres 6 – 18, palps, base of mandible, clypeus except black margins, face except black M-shaped central spot (fig. 14 A), gena, dorsally widened stripe of frontal orbit, collar and upper margin of pronotum, tegula, subtegular ridge, scutellum and postscutellum, upper division of metapleuron, spot of apical half of area dentipara and entire area posteroexterna of propodeum, apical bands of all tergites (fig. 14 C), 2 nd to 5 th sternites except brown lateral spots, apical half of hypopygium and ovipositor sheath. Fore and mid coxae and trochanters and hind tarsus ivory; fore and mid legs brownish, femora and tibiae with frontal yellowish stripes; hind coxa and trochanter black; hind femur and tibia reddish, basally and apically blackish. Wings hyaline; pterostigma black. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF7BDAEFF08FD116088.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF5BDAEFA70FD2464E3.taxon	description	(Figs 15 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF5BDAEFA70FD2464E3.taxon	materials_examined	Holotype. (♀) Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF5BDAEFA70FD2464E3.taxon	etymology	Etymology. The species name refers to the tri-colored metasoma.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF5BDAEFA70FD2464E3.taxon	description	Description. ♀. Body length 10.8 mm. Head. Flagellum with 34 flagellomeres, very long and slender; 1 st flagellomere 5.6 × longer than wide; all flagellomeres much longer than wide. Temple rather long, moderately and roundly narrowed behind eye (fig. 15 B). OED 1.4 × and OOD 0.7 × ocellar diameter. Temple finely punctate. Frons with microsculpture and sparse fine punctures. Face densely but superficially punctate, ± shining. Clypeus large, superficially punctate, almost flat; apical margin blunt, slightly bilobed (fig. 15 A). Mandible sickle-shaped, ventral tooth absent. Malar space 0.8 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Covered with short setae. Notaulus impressed in frontal 0.3 of mesoscutum. Mesoscutum densely but superficially punctate, ± shining. Mesopleuron finely punctate, ventrally part striate in its frontal half; speculum large, smooth and shining. Metapleuron with smoothened surface; juxtacoxal carina present. Scutellum strongly pyramidal, about as long as wide, coarsely rugose, lateral carina in its basal 0.6. Propodeum completely carinated, with strong apophysis (fig. 15 C); spiracle slit-shaped. Area superomedia small, hexagonal, about as wide as long; anterior transverse carina reaching at 4 / 10 of its length. Hind coxa with fine dense punctures, without scopa. Hind femur 4.6 × longer than wide. Claws simple. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their width. Vein 1 cu-a interstitial. Metasoma (fig. 15 C). Strongly amblypygous, slender. Postpetiole moderately widened, without latero-median carina; median field not separated, smooth; lateral field punctate. 2 nd tergite about as long as wide. Gastrocoelus indistinct. Thyridium small, indistinctly impressed. 2 nd tergite with dense fine punctures, ± dull; following tergites with superficial punctures, ± shining. Ovipositor sheath not extending behind metasomal apex. Colour. Black. Ivory are rings on flagellomeres 6 – 12, narrow apical margin of 6 th tergite and entire 7 th tergite. Palps, tegula, subtegular ridge, scutellum and postscutellum yellowish. Mandible except tooth, propodeum and metapleuron, 1 st to 3 rd tergites red; 3 rd tergite with black triangular spot apically (fig. 15 C). 2 nd and 3 rd sternites red; following sternites black. Legs completely red. Wings weakly infuscate; pterostigma red. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD1FFF5BDAEFA70FD2464E3.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD3FFF5BDAEFDD9FA1C62FB.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 1, c 750 m, 1 ♂ 7. ii. – 28. iii. 1995, Mal. trap 2, c 750 m, 3 ♂♂ 7. ii. – 28. iii. 1995, Mal. trap 3, c 750 m, 2 ♀♀ 4 ♂♂ 7. ii. – 28. iii. 1991, 2 ♀♀ 7. ii. – 28. viii. 1995, C. van Achterberg & Y. Yasir, c. 800 m, 1 ♀ 21. ii. – 25. iii. 1997, leg. C. van Achterberg; SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 12, c 200 m, 2 ♂♂ 19. x. – 5. xi. 1989, leg. C. van Achterberg; C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♀ 1 – 14. xi. 1989, 2 ♀♀ 21 – 31. x. 1989, Mal. trap 16, leg. C. van Achterberg; N Sulawesi, Dumoga-Bone N. P. c 220 m, 0 ° 34´N 123 ° 54´E, Mal. trap: Maze Toraut R., 1 ♂♂ 1 – 9. xi. 1985, C. van Achterberg (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD3FFF5BDAEFDD9FA1C62FB.taxon	description	Description. ♂. Body length 11 – 12 mm. Flagellum with 36 flagellomeres, bristle-shaped; 1 st flagellomere 4.0 × longer than wide. Tyloids absent. Mandible with one large tooth. Genal carina reaching mandibular base. Scutellum with lateral carina in basal half. Hind femur 4.1 × longer than wide. 2 nd tergite 1.2 × wider than long. Otherwise as described for the ♀ (Heinrich 1934: 162 – 163). Colour. Black. Rings on flagellomeres 8 – 15, palps, clypeus, face, large triangular spot of frontal orbit, wide frontal margin and spot of hind edge of pronotum, tegula, subtegular ridge, scutellum and postscutellum, 7 th tergite and parameres, fore and mid coxae and trochanters ivory. Mandible, propodeum and apical half of metapleuron red. 1 st to 3 rd tergites red; 4 th tergite red basally, black apically; 5 th and 6 th tergites black. Legs red; hind tarsus black. Wing with yellowish suffusion; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD3FFF5BDAEFDD9FA1C62FB.taxon	discussion	Remarks. The scutellum is usually not carinated, but one otherwise typical ♀ has a lateral carina in basal half of scutellum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD3FFF5BDAEFDD9FA1C62FB.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFF4BDAEFF08FD24608B.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 6 ♂♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFF4BDAEFF08FD24608B.taxon	description	Description. ♂. Body length 16 mm. Head. Flagellum with 43 – 44 flagellomeres, bristle-shaped; 1 st flagellomeres 2.7 × longer than wide. Tyloids on flagellomeres 8 / 9 – 15, long-oval, maximally 0.6 × as long as their flagellomeres. Temple rather short, moderately and roundly narrowed behind eye. OED 1.1 × and OOD 1.0 × ocellar diameter. Face densely punctate, medially and laterally with longitudinal bulging (almost as in Hiorada Cameron). Clypeus moderately convex, densely punctate, apical margin blunt, slightly rounded. Mandible with two teeth, ventral tooth smaller, not distinctly bent inwards. Malar space 0.8 × as long as width of mandibular base. Genal carina reaching mandibular base. Mesosoma. Mesoscutum densely punctate and granulate, dull. Mesopleuron and metapleuron densely punctate, shining; speculum finely punctate; juxtacoxal carina present. Scutellum moderately elevated, slightly longer than wide, punctate, with complete lateral carina. Propodeum without apophysis, spiracle slit-shaped. Area basalis with strong median tubercle. Area superomedia heart-shaped; costula reaching at 7 / 10 of its length. Area dentipara long, roundly sloping apically. Area petiolaris without distinct lateral carina. Hind femur 4.0 – 4.1 × longer than wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 1 – 3 × their width. Vein 1 cu-a interstitial or postfurcal by 1 – 2 × its width. Metasoma. Postpetiole moderately widened, almost smooth with some fine punctures; median field not separated. 2 nd tergite 1.2 × longer than wide. Gastrocoelus distinctly impressed, with longitudinal ridges. Thyridium slightly oblique, 0.7 × as wide as the interval between thyridia. 2 nd and 3 rd tergites finely and densely punctate, dull; 2 nd tergite finely aciculate between thyridia. Apical margin of hypopygium moderately and roundly elongate, almost pointed medially. Colour. Black with extensive ivory and reddish coloration. Flagellum black, flagellomeres 14 – 19 ivory dorsally. Ivory are palps, mandible except teeth, clypeus, face, gena, complete orbits except short interruption at vertex, scape and pedicel ventrally, propleuron, frontal and upper margins of pronotum, two central paramedian stripes on mesoscutum, subtegular ridge, mesosternum and ventral part of mesopleuron, fore and mid coxae and trochanters, 7 th tergite and parameres. Dorsal part of mesopleuron red, sometimes with black spot. Tegula, scutellum except black central spot, postscutellum, metapleuron and propodeum except black transverse band in basal third yellowish-red. 1 st to 4 th tergites reddish, with diffuse yellowish apical bands; 4 th tergite with black spots sub-basally and apically. 5 th and 6 th tergites black. Legs except fore and mid coxae and trochanters reddish; hind tibia apically and hind tarsus entirely blackish. Wings slightly yellowish; pterostigma reddish-brown or brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFF4BDAEFF08FD24608B.taxon	discussion	Remarks. The genus Celebichneumon Heinrich cannot be identified in the key of Townes et al. (1961) unequivocally since the genal carina reaches the mandibular base and not the hypostomal carina. In this key, Celebichneumon runs to Coelojoppa Cameron but has a different carination of propodeum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFF4BDAEFF08FD24608B.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFFBBDAEFA70FD2467BF.taxon	description	(Figs 16 A – B)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFFBBDAEFA70FD2467BF.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 1 ♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFFBBDAEFA70FD2467BF.taxon	description	Description. ♂. Body length 13 mm. Head. Flagellum with 40 flagellomeres, bristle-shaped; 1 st flagellomeres 3.1 × longer than wide. Tyloids on flagellomeres 7 – 17, long-oval, maximally 0.8 × as long as their flagellomeres. Temple short, strongly and roundly narrowed behind eye. OED 1.0 × and OOD 0.9 × ocellar diameter. Face densely punctate. Clypeus moderately convex, densely punctate; apical margin blunt, slightly rounded. Mandible with two teeth, ventral tooth smaller, not distinctly bent inwards. Malar space 0.5 × as long as width of mandibular base. Genal carina reaching mandibular base. Mesosoma. Mesopleuron and metapleuron densely punctate, shining; speculum finely punctate; juxtacoxal carina present. Scutellum moderately elevated, slightly longer than wide, punctate, with lateral carina in basal 0.7. Propodeum without apophysis, spiracle slit-shaped (fig. 16 A). Area basalis without lateral carina, with strong median tubercle. Area superomedia heart-shaped, 1.3 × wider than long; costula reaching at 8 / 10 of its length. Hind femur 4.6 × longer than wide. Metasoma. Postpetiole slightly widened, finely granulate; median field slightly elevated, not separated. 2 nd tergite 1.2 × longer than wide. Gastrocoelus distinctly impressed, with longitudinal ridges. Thyridium oblique, 0.7 × as wide as the interval between thyridia (fig. 16 B). 2 nd and 3 rd tergites densely rugose-punctate, dull; 2 nd tergite aciculate medially. Apical margin of hypopygium moderately and roundly elongate medially. Colour. Black with extensive ivory coloration. Flagellum black; tyloids ivory. Ivory are palps, mandible except teeth, clypeus, face, gena, complete orbits except interruption at vertex, scape ventrally, frontal and upper margins of pronotum, spot of tegula, subtegular ridge, mesosternum and ventral part of mesopleuron, upper division of metapleuron, prescutellar ridge, side of scutellum, postscutellum posterolateral stripe of propodeum, wide apical bands of 1 st to 3 rd tergite, narrow apical band of 6 th tergite, entire 7 th tergite and paramere, 2 nd and 3 rd sternites, apical margin of following sternites. Legs brown; fore and mid coxae and all trochanters and ventral side of hind coxa ivory. Fore and mid femora and tibiae and fore tarsus with ivory stripes frontally. Wings slightly infuscate; pterostigma blackish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD2FFFBBDAEFA70FD2467BF.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDDFFFBBDAEF940FD24629C.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 3 ♂♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDDFFFBBDAEF940FD24629C.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFF08FB2D6457.taxon	materials_examined	Material. Indonesia: Lombok, Batukok, c 500 m, nr Hulu Kalimuntur, Mal. trap 9, 1 ♀ 13 – 26. iii. 1991, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFF08FB2D6457.taxon	distribution	Distribution. Widespread in the Oriental region including Java, new record for Lombok.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFE14FB9F67B4.taxon	materials_examined	Material. Indonesia: N Sumatra, Ketambe, c. 400 m, near N. P. Gn. Leuser, Mal. trap, 1 ♂ ix. 1994, leg. Y. van Nierop & C. van Achterberg (Leiden); N Sumatra, Sibolangit, 500 m, 1 ♂ 19. xii 1954, leg. J. van der Vecht (Leiden); Malaysia: SW Sabah, nr. Long Pa Sia (East), 3 ♂♂ 25. xi. – 7. xii. 1987, c 1000 m, Mal. trap 5, leg. C. van Achterberg; SW Sabah, nr Long Pa Sia (West), c. 1020 m, 1 ♂ 1 – 14. iv. 1987, leg. C. van Achterberg (Leiden); W Sabah, Crocker Range E, W of Apin Apin, 1 ♀ v. 1999, leg. Sreek (Linz).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFE14FB9F67B4.taxon	distribution	Distribution. Known from India, Java and Sumatra, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFD74FC3B615F.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 2, c 200 m, 1 ♀ 1 ♂ 15. x. – 5. xi. 1989, 1 ♀ Mal. trap 12, 1 ♂ 19. v. – 5. vi. 1989, 1 ♀ 1 ♂ 12 – 15. x. 1989, 1 ♂ 15 – 19. x. 1989, 1 ♀ 19. x. – 5. xi. 1989, Mal. trap 10, c 225 m, 12 – 15. x. 1989, leg. C. van Achterberg; SE Sulawesi, Gn. Bulusaraung, n. Camba, Malawa, Mal. trap 1, c 750 m, 2 ♂♂ 7. ii. – 28. iii. 1995, leg. C. van Achterberg & Y. Yasir; Sula Islands, Taliabu, near Tubang, Mal. Trap 20, c 40 m, 1 ♂ 9 – 20. iii. 1995, leg. C. van Achterberg & Y. Yasir (all in Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFD74FC3B615F.taxon	discussion	Remarks. Flagellum of ♀ with 36 flagellomeres, 1 st flagellomere 4.5 × longer than wide. Flagellomeres 5 – 14 ivory dorsally.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFD74FC3B615F.taxon	distribution	Distribution. Known from Sulawesi, new record for the Moluccas.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFB0CFC6E6074.taxon	materials_examined	Material. Malaysia: Sabah, Kinabalu Park, on Aporusa sp., 1 ♂ 26. ii. 1996, leg. A. Floren (ZSM); Sabah, Keningu district, Trus Medi Mt., 1160 m, 1 ♂ 20. vi. 2011, leg. Chuj (coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFB0CFC6E6074.taxon	discussion	Remarks. The specimens from Borneo have completely black flagella, they are otherwise typical.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFFABDAEFB0CFC6E6074.taxon	distribution	Distribution. Known from India and Myanmar, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFF9BDAEFA34FB85640B.taxon	description	(Figs 17 A – B)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFF9BDAEFA34FB85640B.taxon	materials_examined	Material. Indonesia: Sumbawa, nr Pancasila, Gn. Tambora, Mal. trap 1, c 1250 m, 2 ♂♂ 22. ix. – 2. x. 1993, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFF9BDAEFA34FB85640B.taxon	description	Description. ♂. Body length 16 mm. Flagellum with 39 flagellomeres, bristle-shaped; 1 st flagellomeres 3.0 × longer than wide. Tyloids on flagellomeres 9 – 20, long-oval, slightly apically located, maximally 0.6 × as long as their flagellomeres. Temple rather short, distinctly and roundly narrowed behind eye (fig. 17 A). Fore metatarsus with apical tooth. Hind femur 4.3 × longer than wide. Areolet pointed frontally. Vein 1 cu-a postfurcal by 2.0 × its width. 2 nd tergite 1.3 × longer than wide. Thyridium slightly oblique, 1.0 × as wide as the interval between thyridia. Colour. Black with extensive ivory coloration. Flagellomeres 10 – 15 dorsally, palps, mandible except teeth, clypeus, face, gena, complete orbits, scape ventrally, frontal and upper margins of pronotum, two central paramedian stripes on mesoscutum (fig. 17 B), prescutellar ridge, scutellum except black median stripe, postscutellum, large stripe of ventral mesopleuron, posterior spot of mesosternum, upper division of metapleuron, area spiracularis frontal of spiracles, apical half of area dentipara and area posteroexterna, apical margin of postpetiole, posterolateral spots of 2 nd to 5 th tergites (smaller on 5 th tergite), 2 nd to 4 th sternites except lateral brown spots and apical margins of 5 th and 6 th sternites ivory. Legs black; fore and mid coxae and trochantelli, all trochanters, fore and mid legs frontally, subbasal ring of hind tibia and hind tarsus ivory; hind femur black, reddish-brown in basal 1 / 3 – 2 / 3. Wings hyaline, pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDCFFF9BDAEFA34FB85640B.taxon	distribution	Distribution. Known from Laos and Myanmar, new record for Indonesia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEFF08FD1160F0.taxon	description	(Figs 18 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEFF08FD1160F0.taxon	materials_examined	Holotype. (♀) Malaysia: SW Sabah, nr. Long Pa Sia (East), 25. xi. – 7. xii. 1987, c. 1000 m, Mal. trap 5, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEFF08FD1160F0.taxon	description	Description. ♀. Body length 15.3 mm. Head. Flagellum with 37 flagellomeres, distinctly lanceolate; 1 st flagellomere 2.4 × longer than wide, about 11 th flagellomere square, widest flagellomeres 1.9 × wider than long. Temple rather short, distinctly and linearly narrowed behind eye, dorsally 0.5 × as long as eye. OED 1.4 × and OOD 1.0 × ocellar diameter. Frons almost smooth, shining. Face and clypeus rather densely punctate (fig. 18 A). Malar space 0.7 × as long as width of mandibular base. Gena in profile 0.8 × as wide as eye, with sparse punctures. Genal carina reaching hypostomal carina, both carinae low. Mesosoma. Side of pronotum punctate dorsally, with coarse striae ventrally. Notaulus impressed in frontal 1 / 5 of mesoscutum. Mesoscutum coarsely and densely punctate, finely granulate, posterior margin with sparse punctures. Mesopleuron densely punctate, partly rugose, speculum smooth. Metapleuron coarsely rugose-punctate; juxtacoxal carina present. Scutellum slightly elevated, slightly wider than long, with rather dense punctures, without lateral carina. Area basalis and area superomedia confluent; area superomedia slightly wider than long (fig. 18 B); costula reaching at apical 0.7 of its length. Hind coxa with dense punctures and small low pale scopa ventro-apically. Hind femur densely punctate, 3.9 × longer than wide. Tarsi moderately widened, 3 rd mid tarsomere 1.2 × longer than apically wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 3.0 × their diameter. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Oxypygous and rather slender. Postpetiole strongly widened, median field moderately elevated, with fine aciculation and some coarse punctures; lateral field 0.8 × as wide as median field, with coarse punctures. 2 nd tergite 1.1 × wider than long. Gastrocoelus strongly impressed and with ridges. Thyridium almost transverse, 1.0 × as wide as the interval between thyridia. 2 nd and 3 rd tergites coarsely rugose-punctate, with strong median striae (fig. 18 C). 4 th tergite with superficial punctures and fine striae in basal third. Following tergites almost smooth. Ovipositor sheath slightly extending behind metasomal apex by 0.5 × length of 7 th tergite. Colour. Black. Ivory are rings of flagellomeres 4 – 18, palps, base of mandible, clypeus, face except roundish central black spot, complete orbit except narrow interruption at lateral ocellus, ventral spot of scape, collar and upper margin of pronotum, spot of tegula, subtegular ridge, wide stripe on ventral mesopleuron, side of scutellum, posterolateral spots of 1 st to 3 rd tergites, narrow apical margins of 2 nd and 3 rd tergite, and apical bands on 4 th to 7 th tergites. Legs reddish; fore and mid coxae, all trochanters and small ventral and dorsal spots on hind coxa ivory. Frontal stripes of fore and mid tibiae and all tarsi ivory. Hind femur red, black in apical 0.4; hind tibia black, with wide subbasal ivory ring. Wings hyaline, pterostigma black. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEFF08FD1160F0.taxon	discussion	Remarks. This new species belongs to the subgenus Exephanides and resembles Coelichneumon (Exephanides) pseudoneotypus Riedel, 2013 from India. It differs by stouter tarsomeres, slenderer hind femur, stouter metasoma, ivory fore and mid tarsi and slightly different colour patterns of face and tergites.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEFF08FD1160F0.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEF9A8FBD66213.taxon	materials_examined	Material. Indonesia: C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♀ 3 ♂♂ 21 – 31. x. 1989, 2 ♂♂ 1 – 14. xi. 1989, Mal. trap 17, leg. C. van Achterberg; SW Java, Malang, 1 ♀ (no date); W Java, Bogor, 1 ♂ viii. 1977, leg. P. H. van Doesburg (all Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFFBDAEF9A8FBD66213.taxon	distribution	Distribution. Widespread in the Oriental region, known from Java and Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFEBDAEF8C8FBC365EF.taxon	materials_examined	Material. Malaysia: SW Sabah, nr. Long Pa Sia (East), 2 ♂♂ 25. xi. – 7. xii. 1987, c. 1000 m, Mal. trap 5, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFEBDAEF8C8FBC365EF.taxon	discussion	Remarks. Prescutellar ridge black or slightly ivory in the specimens from Borneo. The ♂♂ are otherwise typical.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD9FFFEBDAEF8C8FBC365EF.taxon	distribution	Distribution. Known from China, Myanmar, and Laos, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFEBDAEFED5FD2464E3.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 200 m, 1 ♀ 10 – 15. x. 1989, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFEBDAEFED5FD2464E3.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFEBDAEFDD9FBC16107.taxon	materials_examined	Material. Indonesia: C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♂ 1 – 14. xi. 1989, Mal. trap 17, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFEBDAEFDD9FBC16107.taxon	discussion	Remarks. The colour pattern of this ♂ differs partly from the description of the ♀ of C. maculiceps given by Heinrich (1975: 202 – 203), it may represent a new subspecies of the widespread species. Body length 14 – 15 mm. Flagellum with 41 flagellomeres. Hind femur 4.5 × longer than wide. Colour metallic-blue. Flagellum reddish, black dorsally. Palps, mandible except teeth, lateral spots of clypeus, wide facial orbit, narrow frontal obit, gena, tegula, spot of subtegular ridge, scutellum apical band of postpetiole, apical half of 2 nd tergite, narrow apical margin of 3 rd tergite, apical transverse spots on 5 th to 7 tergites, fore and mid coxae and trochanters cream-yellow. Petiole and area of gastrocoelus reddish. Hind coxa metallic-blue; fore and mid femora, tibiae and tarsi reddish; hind femur brown, hind tibia ochreous, black in apical 0.2; hind tarsus black. Wings hyaline; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFEBDAEFDD9FBC16107.taxon	distribution	Distribution. Known from India, Myanmar and Java, new record for Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	description	(Figs 19 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	materials_examined	Holotype. (♂) Indonesia: Sulawesi, nr Bantaeng, Borong Rappoa, Gn. Daulu, Mal. trap 16, c 700 m, 4 – 24. iv. 1991, C. van Achterberg, RMNH´ 91 (Leiden). Paratype. (♂) similar label (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	etymology	Etymology. The species name refers to the four ivory spots on the mesoscutum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	description	Description. ♂. Body length 14 – 15 mm. Head. Flagellum with 37 – 39 flagellomeres, bristle-shaped, with distinctly nodose flagellomeres. 1 st flagellomere 2.0 × and 2 nd flagellomere 2.4 × longer than wide. Tyloids not visible. Head large. Temple long, slightly widened behind eye and moderately narrowed apically (fig. 19 B). OED 1.2 – 1.6 × and OOD 0.7 – 1.0 × ocellar diameter. Vertex and frons with sparse punctures, strongly shining. Face and clypeus densely punctate, face dull. Clypeus flat, c 1.8 × wider than long; apical margin sharp and slightly bilobed (fig. 19 A). Mandible large; ventral tooth large and only slightly shorter than dorsal tooth. Malar space 0.8 × as long as width of mandibular base. Gena with rather dense punctures. Genal carina reaching mandibular base. Mesosoma. Mesoscutum with coarse vermiform rugae. Notaulus indistinctly but widely impressed. Mesopleuron rugose-punctate ventrally, smooth and with sparse punctures dorsally; speculum large, smooth. Metapleuron with superficial punctures, shining; juxtacoxal carina present. Scutellum distinctly elevated, almost square, coarsely punctate or rugose, with complete lateral carina. Propodeum short and strongly sloping downwards to apex, with reticular-rugose surface (fig. 19 C), without apophysis; spiracle slit-shaped. Area superomedia small, oval, c 2.0 × longer than wide; costula in its middle. Legs slender. Hind coxa densely punctate. Hind femur 5.0 – 5.3 × longer than wide. Claws simple. Areolet pointed or shortly stalked frontally. Vein 1 cu-a strongly postfurcal by 3 – 4 × its width. Nervellus of hknd wing distinctly reclivous, intercepted in apical 0.7. Metasoma. Slender. Petiole with lateral carina in distal half. Postpetiole distinctly widened, without latero-median carina, finely rugose or aciculate, median field not separated. 2 nd tergite 1.0 – 1.2 × as long as wide. Gastrocoelus distinctly impressed, with fine ridges. Thyridium large, slightly oblique, 1.7 – 2.3 × wider than the interval between thyridia (fig. 19 D). 2 nd tergite finely and densely rugose-punctate, finely aciculate medially, dull. Following tergites finely punctate, ± shining. Only 2 nd sternite with median fold. Hypopygium unmodified, apical margin straight. Colour. Black with extended ivory coloration. Flagellum black. Head and mesosoma black with ivory colour pattern. Ivory are palps, mandible except teeth, clypeus, face, gena, frontal orbit, scape ventrally, collar, upper margin and ventral third of pronotum, tegula, subtegular ridge, epicnemium, ventral stripe and dorsal spot of mesopleuron, median stripe of mesosternum, short anterolateral and paramedian spots of mesoscutum, lateral spots of scutellum, postscutellum, upper division and central spot of metapleuron. Propodeum mainly ivory, area superomedia brownish, subbasal transverse band of propodeum and spot on area petiolaris blackish (fig. 19 C). In holotype, 1 st to 4 th tergites except black base of petiole red, with yellow apical bands. 5 th to 7 th tergites black, with narrow ivory apical margins. In paratype, 1 st and 2 nd tergites reddish-brown, following tergites blackish, apical bands of 1 st to 4 th tergites yellow, following tergites with narrow yellow apical margins. Legs reddish; fore and mid coxae and trochanters ivory; hind coxa reddish-brown, with ivory dorsal spot; fore tibia with yellow stripe frontally; hind tibia infuscate in apical 0.2; hind tarsus ochreous or black. Wings with weak yellowish suffusion; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	discussion	Remarks. This new species belongs to Xenojoppa Cameron, 1902 which has been synonymized with Compsophorus by Townes & Townes (1973) but separated by Heinrich (1975). Xenojoppa is preliminarily included as a subgenus in the genus Composphorus here. In his key of the known Xenojoppa species (Heinrich 1975: 487 – 489) it runs to Xenojoppa parvidens Heinrich, 1975 from Sulawesi but differs by yellowish anterolateral and paramedian spots of mesoscutum, apical yellow bands or margins of all tergites, entirely black flagellum and extended ivory coloration of the propodeum. Since many Xenojoppa species show a considerable colour variability and the ♀ of the new species is still unknown, it is possible that the new species represents a colour variant of Xenojoppa parvidens only. It seems necessary that the separation of different Xenojoppa species as shortly presented by Heinrich (1975: 487 – 488) needs further confirmation by additional material and genomic studies.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFD8FFFCBDAEFBFDFD24662F.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFC1CFD24618F.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Ciaker Alam Laiya, c 15 km S of Camba, c 400 m, 1 ♀ 28. xi. 1991, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFC1CFD24618F.taxon	discussion	Remarks. This specimen belongs to the var. bonthaina Heinrich, 1934 with its slightly reduced ivory coloration. Body length 15 mm. Flagellum with 42 flagellomeres; moderately lanceolate; 1 st flagellomere 2.7 × longer than wide, widest flagellomere 2.1 × wider than long. Hind femur 3.8 × longer than wide, ventral 1 / 3 with sparse punctures.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFC1CFD24618F.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFB79FCF7607B.taxon	materials_examined	Material. Malaysia: Sabah, Kinabalu Park, 6 ° 5´N 116 ° 33´E, Sorinsim, 1 ♂ 11. iii. 1997, leg. A. Floren (ZSM). Distribution. Known from Sulawesi, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFA28FCF762FB.taxon	materials_examined	Material. Indonesia: C Sulawesi, Lore Lindu N. P., Marena Shelter, 600 m, 1 ♀ 14 – 17. xii. 1985, leg. J. van Tol & J. Krikken (Leiden); Malaysia: SE Sabah, nr Danum Valley Field C, c 150 m, Mal. trap 10, 1 ♀ 15 – 19. iii. 1987, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFA28FCF762FB.taxon	discussion	Remarks. ♀. Body length 12 mm. Flagellum with 35 flagellomeres, slightly lanceolate; 1 st flagellomeres 5.0 × longer than wide; widest flagellomere 1.2 × wider than long. Notaulus on frontal 0.6 of mesoscutum. Mesoscutum with fine very sparse punctures and with coarse rugae apico-medially. Scutellum moderately elevated, slightly wider than long, with complete carina laterally and apically. Area superomedia heart-shaped; about as long as wide; costula reaching at 7 / 10 of its length. Postpetiole moderately widened, densely punctate; median field not separated. 2 nd tergite 1.1 × longer than wide. Gastrocoelus impressed, almost smooth. Thyridium large, transverse, 2.0 × wider than the interval between thyridia. Colour. As described by Heinrich (1934: 164 – 165), but flagellomeres 5 – 17 ivory; 1 st to 3 rd tergites red, with diffuse yellow apical bands; 4 th to 7 th tergites black; 4 th tergite with triangular posterolateral cream-yellow stripes; 5 th to 7 th tergites largely ivory; hind femur apically and hind tibia basally and apically narrowly black; hind tarsus brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFDAFFFCBDAEFA28FCF762FB.taxon	distribution	Distribution. Known from Sulawesi, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC5FFE2BDAEF98DFD116123.taxon	description	(Figs 20 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC5FFE2BDAEF98DFD116123.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr Danum Valley Field C, WON 1, c 150 m, Mal. trap 5, 14 – 20. iii. 1987, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC5FFE2BDAEF98DFD116123.taxon	description	Description. ♀. Body length 13 mm. Head. Flagellum with> 37 flagellomeres (tips broken off), very slender and long, bristle-shaped; 1 st flagellomere 5.7 × longer than wide, all flagellomeres much longer than wide. Temple very short, strongly and linearly narrowed behind eye (fig. 20 B). OED 1.3 × and OOD 1.2 × ocellar diameter. Frons almost smooth, with small dorsal tentorial pit above antennal groove. Face densely punctate, smooth, medially moderately bulging. Clypeus moderately convex, 1.8 × wider than long, with sparse punctures; apical margin sharp and slightly rounded (fig. 20 A). Mandible long, continuously narrowed towards apex, with two teeth, ventral tooth much smaller than dorsal one. Malar space 0.5 × as long as width of mandibular base. Gena punctate. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Upper margin of pronotum distinctly and almost lamelliform widened (fig. 20 C). Notaulus strongly impressed and crenulate, reaching apical 0.8 of mesoscutum (fig. 20 C). Mesoscutum with superficial punctures, shining. Speculum smooth. Metapleuron rugose, dull; juxtacoxal carina absent. Scutellum moderately elevated, wider than long, almost rectangular, with coarse rugae and complete carina laterally and apically (fig. 20 D). Postscutellum with coarse longitudinal ridges. Propodeum completely carinated (fig. 20 D), without distinct apophysis but apical transverse carina at outer edge of area dentipara slightly lamelliform widened. Area basalis trapezoid, wider than long. Area superomedia almost rectangular, 1.3 × longer than wide; costula reaching at 6 / 10 of its length. Legs slender. Hind coxa densely punctate, without scopa. Hind femur 5.6 × longer than wide. Fore and mid claws with short basal teeth. Hind claw simple. Areolet quadrangular, pointed frontally; vein 2 m-cu slightly proximad its middle. Vein 1 cu-a interstitial. Nervellus of hind wing slightly reclivous, intercepted in its apical 0.7. Metasoma. Strongly oxypygous; 2 nd to 5 th sternites with median folds. Petiole as high as wide. Postpetiole moderately widened; median field slightly elevated, with longitudinal rugae; lateral field punctate. Gastrocoelus strongly impressed, with longitudinal ridges. Thyridium transverse, 1.5 × wider than the interval between thyridia. 2 nd tergite 1.1 × wider than long. 2 nd to 5 th tergites densely and finely rugose-punctate, dull. Following tergites with superficial punctures, ± shining. Ovipositor sheath long and slender, reaching behind metasomal apex by length of 3 rd hind tarsomere. Colour. Black with extended ivory coloration. Ivory are flagellomeres 6 – 15, palps, mandible except teeth, clypeus (blackish around anterior tentorial pits), face except dorso-median black stripe, frontal orbit up to vertex, gena, frontal and upper margins of pronotum, rectangular central spot of mesoscutum, subtegular ridge, large ventral stripe and posterior margin of mesopleuron, scutellum and postscutellum, posterolateral spot of propodeum, base of petiole, gastrocoelus, basal margins of 3 rd to 5 th tergites, posterolateral spot of postpetiole and apical bands of all following tergites. Foe and mid coxae and trochanters and dorsal spot of hind coxa ivory. Femora reddish, hind femur black in apical 0.3; tibiae ivory, hind tibia black basally and in apical 0.2; tarsi ivory, mid tarsus partly brownish. Wings hyaline; pterostigma dark brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC5FFE2BDAEF98DFD116123.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC4FFE2BDAEFBE1FA8C605B.taxon	materials_examined	Material. Malaysia, SW Sabah, nr Long Pa Sia, Payakalaba, 1 ♀ 25. xi. – 1. xii. 1987, c. 1010 m, Mal. trap 14, 1 ♀ 12 – 13. iv. 1987, Mal. trap 12, leg. C. van Achterberg; SW Sabah, nr. Long Pa Sia (East), 1 ♂ 25. xi. – 7. xii. 1987, c. 1000 m, Mal. trap 5, leg. C. van Achterberg (all Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC4FFE2BDAEFBE1FA8C605B.taxon	distribution	Distribution. Known from China, India, Indonesia (Sulawesi) and Japan, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC4FFE2BDAEFA08FBF362E7.taxon	materials_examined	Material. Indonesia: N Sumatra, Ketambe, c. 400 m, near N. P. Gn. Leuser, Mal. trap, 1 ♂ 7. iii. – 15. iv. 1994, 2 ♂♂ vi. 1994, 1 ♂ viii. 1994, 1 ♂ ix. 1994, 1 ♀ ii. 1995, 1 ♂ vi. 1995, Y. van Nierop & C. van Achterberg (Leiden); W Sumatra, Lahan basah, Andalas University, Padang, 0.933 ° S 100.455 ° E, 156 m, 1 ♀ 1 ♂ 23. i. 2016, leg. H. Hamid (ZSM); Java, Cisama, 900 m, 1 ♀ 9. viii. 1986, leg. M. Riedel (coll. Riedel); Malaysia: SW Sabah, nr Long Pa Sia (East), c 1000 m, 1 ♀ 1 ♀ 1 – 13. iv. 1987, 1 ♂ 25. xi. – 7. xii. 1987, Mal. trap 5, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC4FFE2BDAEFA08FBF362E7.taxon	discussion	Remarks. ♀. Flagellum with 36 – 37 flagellomeres. Colour: head black, frontal orbit with large ivory spot. Flagellum with ivory rings on flagellomeres 8 – 12. Mesosoma black; scutellum, metapleuron and propodeum red; collar with yellow band. Legs red, fore coxa yellowish; hind tibia apically and hind tarsus black. 1 st to 3 rd tergites red, following tergites black, 6 th and 7 th tergites with large apical ivory spots. The ♀ from Sabah has a yellow scutellum, black metapleuron, brown petiole and reddish propodeum with brownish sides. ♂: Flagellum with 36 flagellomeres, bristle-shaped; 1 st flagellomere 2.7 × longer than wide. Tyloids absent. Mesopleuron except speculum entirely striate. Colour reddish. Flagellum black, basal flagellomeres ± red. Frons medially, stemmaticum, vertex, pronotum medially and dorsally, mesoscutum entirely and mesopleuron in dorsal 2 / 3 black. Hind tarsus black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC4FFE2BDAEFA08FBF362E7.taxon	distribution	Distribution. Known from Java and Sulawesi, new record for Borneo and Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	description	(Figs 21 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr Danum Valley Field C, Mal. trap 7, c 150 m, 21 – 25. iii. 1987, C. van Achterberg, RMNH´ 1987 (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	etymology	Etymology. The species name refers to the black hind femur.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	description	Description. ♀. Body length 12.3 mm. Head. Flagellum with 40 flagellomeres, slightly lanceolate; 1 st flagellomere 4.0 × longer than wide, widest flagellomere 1.4 × wider than long. Temple rather short, strongly and roundly narrowed behind eye. OED 1.6 × and OOD 0.8 × ocellar diameter. Frons concave, with microsculpture, dorsally with some superficial punctures. Face densely but superficially punctate medially, almost smooth laterally. Clypeus large, flat, with sparse superficial punctures; apical margin sharp, moderately rounded (fig. 21 A). Mandible seen from frontal sickle-shaped, with two teeth, ventral tooth smaller and strongly bent inwards (fig. 21 A). Malar space 0.9 × as long as width of mandibular base. Gena with fine dense punctures. Genal carina reaching mandibular base. Mesosoma. Densely covered with setae; setae blackish on mesoscutum and scutellum, otherwise pale. Mesoscutum finely punctate, dull. Notaulus impressed in frontal 1 / 4 of mesoscutum. Mesopleuron rugose- punctate and striate ventrally, punctate dorsally; speculum with fine punctures. Metapleuron ± striate. Juxtacoxal carina present. Scutellum pyramidal, slightly longer than wide, rugosely punctate, with strong lateral carina forming slightly raised lamella apically. Propodeum completely carinated, punctate, with strong apophysis (fig. 21 B); spiracle slit-shaped. Area superomedia slightly longer than wide; costula reaching at its basal 0.25. Legs slender; hind coxa punctate, without scopa. Hind femur 5.9 × longer than wide. Claws simple, strongly bent (by 90 °) apically. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Postpetiole moderately widened, almost smooth. 2 nd tergite about as long as wide. Gastrocoelus narrow and elongate, with some fine ridges. Thyridium slightly oblique, 0.7 × as wide as interval between thyridia (fig. 21 C). 2 nd and 3 rd tergites finely and densely punctate, ± dull; following tergites with superficial punctures and shining. Ovipositor sheath reaching behind metasomal apex by length of 2 nd hind tarsomere. Colour. Red. Flagellum black, flagellomeres 7 – 12 ivory dorsally. Palps pale yellow. Mandible except teeth, clypeus and face reddish-yellow; facial orbit yellowish. Mid tarsus ± brownish; hind leg except red hind coxa blackish. Wings yellowish; pterostigma ochreous. ♀ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	discussion	Remarks. This new species differs from other Eccoptosage species with entirely red mesosoma and metasoma by its black hind legs.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE0BDAEFF08FD116040.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE7BDAEFA38FBD6670F.taxon	materials_examined	Material. Indonesia: S Sulawesi, Toraya-land, 1000 m, Rantepao, Puncak, 1 ♀ 21 – 24. ii. 1994, leg. J. Oehlke (Linz); C Sulawesi, nr Luwuk, Bunga, c 300 m, 1 ♀ 1 – 14. xi. 1989, Mal. trap 17, leg. C. van Achterberg (Leiden); C Sulawesi, Salodik, 400 m, 1 ♂ 27 – 30. x. 1989, leg. M. Tuhung (Leiden); SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 3, c 750 m, 1 ♀ 2 ♂♂ 7. ii. – 28. iii. 1991, leg. C. van Achterberg & Y. Yasir (Leiden); N Sulawesi, Dumoga-Bone N. P. c 245 m, Mal. trap, Toraut, 1 ♂ 26. vi. – 3. vii. 1985, leg. J. Huijbregts ET AL. (Leiden); SE Sulawesi, nr Sanggona, 1 km W base camp, Gn. Watuwila, Mal. trap 12, c 200 m, 3 ♂♂ 12 – 15. x. 1989, 1 ♂ 15 – 19. x. 1989, 1 ♀ 4 ♂♂ 19. x. – 5. xi. 1989, Mal. trap 1, c 225 m, 1 ♀ 15. x. – 5. xi. 1989, Mal. trap 6, c 200 m, 1 ♂ 15. x. – 5. xi. 1989, leg. C. van Achterberg (Leiden); N Sumatra, Ketambe, c. 400 m, near N. P. Gn. Leuser, Mal. trap, xi. 1994, leg. Y. van Nierop & C. van Achterberg (Leiden); W Bali, nr Negara, rainforest above Batuagung, c 575 m, 1 ♀ 4 – 13. xii. 1991, Mal. trap 1, leg. C. van Achterberg (Leiden); W Java, Mt. Halimun Salak NP, Cikaniki, 6.7445 ° S 106.538 ° E, 1052 m, 1 ♂ 21. iv. 2016, X. Mengal & T. Burton (ZSM); W Sumatra, Bukittingi env., Sianok, 0 ° 17,5946´S 100 ° 20,5449´E, 1 ♀ 15. xi. 2015, leg. V. Gromenko (coll. Riedel); Malaysia: SE Sabah, nr. Danum Valley Field C, WON 1, c 150 m, Mal. trap 5, 1 ♀ 15. iv. – 15. v. 1987, leg. C. van Achterberg (Leiden); SW Sabah, nr. Long Pa Sia, Payakalaba, 1 ♂ 12 – 13. iv. 1987, c. 1010 m, Mal. trap 13, leg. C. van Achterberg (Leiden); SW Sabah, nr Long Pa Sia (East), c 1000 m, 1 ♂ 1 – 13. iv. 1987, 1 ♂ 25. xi. – 7. xii. 1987, Mal. trap 5, leg. C. van Achterberg (Leiden); N Sabah, Kinabalu Park Poring, c 475 m, Mal. trap 6, 1 ♂ 9 – 12. iii. 1987, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE7BDAEFA38FBD6670F.taxon	discussion	Remarks. ♀: Body length 9 – 11 mm. Flagellum with 33 – 37 flagellomeres, slightly lanceolate; 1 st flagellomere 3.6 × longer than wide, widest flagellomeres 1.4 × wider than long. OED 1.7 × and OOD 0.7 × ocellar diameter. Mesopleuron superficially punctate and sometimes with weak striation in ventral half. Apophysis long. ♂: Body length 11 – 14 mm. Flagellum with 36 – 37 flagellomeres, bristle-shaped, 1 st flagellomere 3.3 × longer than wide, all flagellomeres longer than wide. Tyloids on flagellomeres 9 – 13 / 14, long-oval, maximally 0.8 × as long as their flagellomeres. OED 1.8 × and OOD 0.7 × ocellar diameter. Mesopleuron punctate, not striate. Apophysis short. Colour reddish. Flagellum reddish basally, mainly black, in ♂ flagellomeres 12 – 17 with ivory stripes. Hind tarsus blackish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC6FFE7BDAEFA38FBD6670F.taxon	distribution	Distribution. Known from Sulawesi, new records for Borneo, Java and Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC1FFE7BDAEFDF4FD1162AF.taxon	description	(Figs 22 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC1FFE7BDAEFDF4FD1162AF.taxon	materials_examined	Material. Malaysia: SE Sabah, nr. Danum Valley Field C, c 150 m, Mal. trap 4, over rivelet, 1 ♀ 20 – 22. xi. 1987, Mal. trap 10, 1 ♂ 20 – 22. iii. 1987, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC1FFE7BDAEFDF4FD1162AF.taxon	description	Description. ♀. Body length 16 mm. Flagellum with 43 flagellomeres, slightly lanceolate; 1 st flagellomere 4.3 × longer than wide, 13 th flagellomere square, widest flagellomeres 1.3 × wider than wide. OED 1.6 × and OOD 0.6 × ocellar diameter. Mesoscutum finely rugose-punctate, dull. Pronotum, mesopleuron and metapleuron striate. Hind femur 5.4 × longer than wide. 2 nd tergite 1.1 × wider than long. Thyridium small, oval, 0.3 × as wide as the interval between thyridia. Colour. Flagellum black, without ivory ring. Tegula ivory. Hind femur red, black in apical 0.3, apex with small yellowish spot. 1 st to 3 rd tergites red, 3 rd tergite infuscate in apical 0.3. Petiole mainly and hind margins of all tergites with narrow ivory bands. Colour otherwise as described by Cameron (1905: 154 – 155). ♂: Body length 14 mm. Head. Flagellum with 42 flagellomeres, slender and bristle-shaped; 1 st flagellomere 3.5 × longer than wide. Tyloids on flagellomeres 10 – 14, long-oval, maximally 0.7 × as long as their flagellomeres. Temple rather short, moderately and roundly narrowed behind eye. OED 1.3 × and OOD 0.6 × ocellar diameter. Frons with microsculpture, ± shining. Face and clypeus densely punctate. Clypeus large, flat; apical margin sharp, slightly rounded (fig. 22 A). Mandible sickle-shaped, ventral tooth small and strongly turned inwards. Malar space 0.5 × as long as width of mandibular base. Genal carina reaching mandibular base. Mesosoma. Mesoscutum finely punctate, dull. Mesopleuron densely punctate, ventrally partly rugose-punctate; speculum with fine punctures. Metapleuron punctate and ± striate. Scutellum pyramidal, with strong lateral carina forming a distinct denticular spine apically. Propodeum with strong apophysis (fig. 22 B). Area superomedia hexagonal, 1.3 × longer than wide; costula reaching its basal 0.3. Hind femur 5.1 × longer than wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 3.0 × their width. Vein 1 cu-a postfurcal by 3.0 × its width. Metasoma. Postpetiole moderately widened, almost smooth centrally and finely punctate laterally and apically. 2 nd tergite 1.1 × wider than long. Gastrocoelus slightly impressed, smooth. Thyridium small, 0.3 × as wide as interval between thyridia (fig. 22 C). 2 nd and 3 rd tergites finely and densely punctate; following tergites with superficial punctures. Hypopygium unmodified, apical margin straight. Colour. Black. Flagellum black, brownish ventrally. Ivory are palps, mandible except teeth, clypeus, face, wide frontal orbit, gena up to vertex, scape ventrally, frontal margin and apical half of upper margin of pronotum, spot of tegula, subtegular ridge, spot of ventral mesopleuron, apical tip of lateral carina of scutellum, postscutellum, apophysis, apical spot of postpetiole and apical margins of 2 nd to 5 th tergites. 1 st to 3 rd tergites reddish. Legs black; fore and mid coxae and trochanters and frontal stripes on fore and mid femora, tibiae and tarsi ivory; hind coxa black, red dorsally; hind femur red dorsally, black ventrally, with ivory spot apically; hind tibia black, with ivory stripe externally; hind tarsus black. Fore wing distinctly infuscate, somewhat paler basally; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC1FFE7BDAEFDF4FD1162AF.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFE4BDAEFF08FD116088.taxon	description	(Figs 23 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFE4BDAEFF08FD116088.taxon	materials_examined	Holotype. (♂) Malaysia: SE Sabah, nr Danum Valley Field C, Malaise trap 15, c 150 m, 20. x. – 22. xii. 1987, C. van Achterberg, RMNH´ 87 (Leiden). Paratype. (♂) Malaysia: SW Sabah, nr Long Pa Sia (West), c. 1050 m, 1 – 14. iv. 1987, Mal. trap 3, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFE4BDAEFF08FD116088.taxon	description	Description. ♂. Body length 10.0 – 10.5 mm. Head. Flagellum with 35 flagellomeres, bristle-shaped; 1 st flagellomere 2.8 × longer than wide. Tyloids on flagellomeres 10 / 11 – 13 / 14, long-oval, maximally 0.5 × as long as their flagellomeres. Temple short, strongly and roundly narrowed behind eye. OED 1.6 × and OOD 0.4 × ocellar diameter. Frons smooth, with some sparse punctures dorsally. Face densely punctate. Clypeus large, flat, with rather dense punctuation; apical margin sharp, slightly rounded (fig. 23 A). Mandible seen from frontal sickle-shaped, ventral tooth small and strongly turned inwards (fig. 23 A). Malar space 0.6 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Mesoscutum finely and densely punctate, dull. Mesopleuron with rather dense superficial punctures, shining; speculum with fine punctures. Metapleuron punctate (in HT) or ± striate (in PT). Scutellum pyramidal, with strong lateral carina forming a distinct lamelliform extension which is slightly bent outwards. Propodeum with strong apophysis (fig. 23 B). Area superomedia hexagonal, slightly wider than long (confluent with area basalis in HT); costula reaching its basal 0.4. Hind femur 4.7 × longer than wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their width. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Postpetiole slightly widened, with microsculpture and few fine punctures, shining. 2 nd tergite 1.1 × longer than wide. Gastrocoelus slightly impressed. Thyridium small, transverse, 0.3 × as wide as interval between thyridia (fig. 23 C). 2 nd tergite finely and densely punctate, finely rugose and aciculate basally, dull. Following tergites with superficial punctures, ± shining. Hypopygium unmodified, apical margin rounded. Colour. Flagellum black, flagellomeres 8 / 9 – 16 ivory dorsally. Head and mesosoma red. Palps, mandible except teeth, clypeus, face and frontal orbit ivory; gena yellowish. Frons and temples black. Scape red ventrally. 1 st tergite reddish. 2 nd and 3 rd tergites mainly black, yellowish laterally and apically (in PT) or yellowish, with black basal spots (in HT) (fig. 23 C). 4 th to 6 th tergites blackish or brownish, yellowish laterally and apically. 7 th tergite and paramere yellowish. Fore and mid legs including coxae reddish-yellow; mid coxa basally and mid tarsus mainly brown; hind leg black, hind coxa red dorsally. Wings slightly yellowish; pterostigma yellow.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFE4BDAEFF08FD116088.taxon	discussion	Remarks. I was not able to correlate these ♂♂ with any ♀♀ of the known Eccoptosage species from the Oriental region and describe them as a new species here, but the taxonomy of the Oriental Eccoptosage species is fragmentary to date. More studies of the available type material, additional collection activities and genomic studies are necessary in the future for a comprehensive revision of this rather large genus.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFE4BDAEFF08FD116088.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFEBBDAEFA70FC6E6457.taxon	materials_examined	Material. Indonesia: Sulawesi, nr Sanggona, 1 km W base camp, Gn. Watuwila, Mal. trap 12, c 200 m, 1 ♂ 12 – 15. x. 1989, 1 ♀ 19. x. – 5. xi. 1989, leg. C. van Achterberg; C Sulawesi, nr Luwuk, Bunga, Mal. trap 15, c 300 m, 1 ♀ 1 – 14. xi. 1989, leg. C. van Achterberg; SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 3, c 750 m, 1 ♀ 4 ♂♂ 7. ii. – 28. iii. 1995, leg. C. van Achterberg & Y. Yasir; N Sulawesi, Dumoga-Bone N. P. c 220 m, 0 ° 34´N 123 ° 54´E, Mal. trap: Maze Toraut R., 1 ♂ 1 – 9. xi. 1985, leg. C. van Achterberg (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFEBBDAEFA70FC6E6457.taxon	discussion	Remarks. ♀. Body length 13 mm. Flagellum with 38 – 45 flagellomeres, moderately lanceolate; 1 st flagellomeres 4.0 × longer than wide, widest flagellomere 1.6 × wider than long. OED 1.7 × and OOD 0.8 × ocellar diameter. Mesopleuron striate ventrally and punctate dorsally, metapleuron striate-punctate. Apophysis and apical extension of lateral carina of scutellum long. Hind femur 5.0 × longer than wide. Colour red. Flagellum black, basal flagellomeres ± red, flagellomeres 7 – 14 ivory dorsally. Mandible, clypeus and face ± reddish-yellow. Hind tarsus red or ± brownish. Wings slightly yellowish; pterostigma ochreous. ♂: Body length 12 – 16 mm. Flagellum with 40 – 43 flagellomeres, bristle-shaped. 1 st flagellomere 3.7 × longer than wide. Tyloids on flagellomeres 9 / 10 – 13 / 15, long-oval, maximally 0.8 × as long as their flagellomeres. OED 1.8 × and OOD 0.7 × ocellar diameter. Mesopleuron and metapleuron finely rugose-punctate. Hind femur 4.5 × longer than wide. Colour red. Flagellum black, ± brownish ventrally. Mandible, clypeus, face and gena reddish-yellow. Hind tibia ± infuscate apically; hind tarsus black. Pterostigma reddish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC2FFEBBDAEFA70FC6E6457.taxon	distribution	Distribution. Widespread in the Oriental region, known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFE0DFCDA67E7.taxon	materials_examined	Material. Indonesia: W Bali, nr Negara, rainforest above Batuagung, c 575 m, 4 ♂♂ 4 – 13. xii. 1991, Mal. trap 1, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFE0DFCDA67E7.taxon	discussion	Remarks. Small, body length 8.0 – 10.5 mm. Flagellum with 31 flagellomeres. Hind femur reddish basally, black apically. Apical margin of postpetiole and 2 nd to 4 th tergites reddish, following tergites black; 1 st to 2 nd or 3 rd tergites with ivory posterolateral spots, 5 th or 6 th to 7 th tergites with apical ivory margins. The structure and colour pattern of these ♂♂ largely resemble Heresiarches daedalus (Tosquinet, 1903) from Java. Probably, H. flavomaculatus represents only a colour variant of the latter species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFE0DFCDA67E7.taxon	distribution	Distribution. Known from Myanmar, new record for Bali.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	materials_examined	Material. Indonesia: W Java, Gunung Botol, 6.729 ° S 106.491 ° E, 1490 m, 2 ♂♂ 24. iv. 2016, leg. X. Mengal & T. Burton (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	discussion	Remarks. The Javanese ♂♂ slightly differ from the original description (Heinrich 1970: 77 – 78) by: mesoscutum entirely black; tegula black with small ivory spots; hind femur black, narrowly reddish at base; 2 nd tergite with diffuse yellowish posterolateral spot. The ♂♂ are otherwise typical.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	distribution	Distribution. Known from Laos and Myanmar, new record for Java. Heresiarches secundus Heinrich, 1970	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	materials_examined	Material. Indonesia: W Bali, nr Negara, rainforest above Batuagung, Mal. trap 2, c 550 m, 1 ♂ 4 – 13. xii. 1991, Mal. trap 1, c 575 m, 1 ♂ 4 – 13. xii. 1991, leg. C. van Achterberg; N Sumatra, Ketambe, 400 m, Mal. trap, near edge rainforest, xi. 1995, leg. Y. van Nierop & C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	discussion	Remarks. Body length 9 mm. Flagellum with 32 – 34 flagellomeres; 1 st flagellomere 3.2 × longer than wide. OED 0.9 × and OOD 0.8 × ocellar diameter. Hind femur 4.1 × longer than wide.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFEBBDAEFCDDFAF060D7.taxon	distribution	Distribution. Widespread in the Oriental region, known from Sumatra (Riedel 2013), new record for Bali.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	description	(Figs 24 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	materials_examined	Holotype. (♀) Malaysia: SW Sabah, nr Long Pa Sia (West), c 1050 m, 1 – 14. iv. 1987, Mal. trap 3, C. van Achterberg, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	etymology	Etymology. The species name refers to the similarity of this new species with the type species of this genus, Heresiarches eudoxius (Wesmael, 1845).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	description	Description. ♀. Body length 11 mm. Head. Flagellum with 33 flagellomeres, almost filiform; 1 st flagellomere 5.0 × and 2 nd flagellomere 4.0 × longer than wide, widest flagellomeres slightly longer than wide. Temple short, strongly and roundly narrowed behind eye, dorsally 0.3 × as long as eye. OED 1.0 × and OOD 1.4 × ocellar diameter. Frons shining, with microsculpture and very sparse fine punctures. Face and clypeus with fine sparse punctures (fig. 24 A). Malar space 0.8 × as long as width of mandibular base. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus indistinct. Mesoscutum with fine dense punctures and granulate, ± dull. Mesopleuron with dense punctures, shining; speculum large and smooth. Metapleuron densely punctate; juxtacoxal carina present. Scutellum moderately elevated, about as long as wide, with fine sparse punctures, with lateral carina in basal 0.8. Area basalis and area superomedia confluent, slightly elevated, smooth, frontal margin bulging towards postscutellum medially, without lateral carina (fig. 24 B). Hind coxa with dense superficial punctures, without scopa. Hind femur 4.3 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rsm 2.0 × their diameter. Distal part of vein Rs distinctly sinusoid. Vein 1 cu-a postfurcal by 3.0 × its width. Nervellus of hind wing reclivous, intercepted in its apical 0.7. Metasoma (fig. 24 C). Strongly oxypygous. Postpetiole slightly widened, without latero-median carina, finely aciculate basally and smooth posteriorly; median field not separated. Gastrocoelus distinctly impressed, almost triangular, with strong ridges. Thyridium small, oblique, 0.5 × as wide as the interval between thyridia. 2 nd tergite 1.3 × wider than long, densely punctate and distinctly striate medially. 3 rd and 4 th tergites with superficial punctures, shining; following tergites almost smooth. Colour. Black, tergites with brownish suffusion. Ivory are rings on flagellomeres 7 – 13, palps, lateral margin of clypeus, wide facial orbit (widened and almost confluent below antennal grooves) (fig. 24 A), gena, complete orbit (except narrow interception at malar space), scape ventrally, frontal and upper margins of pronotum, apical spot of tegula, subtegular ridge, stripe of ventral mesopleuron, upper division of metapleuron, area posteroexterna of propodeum, posterolateral spots of 1 st to 3 rd tergites, narrow apical margins of 4 th and 5 th tergites, wide apical bands of 6 th and 7 th tergites (fig. 24 C). Legs reddish; fore coxa entirely, mid coxa apically, dorsal spot of hind coxa and all trochanters ivory; hind trochantellus and hind femur brown, hind femur with narrow red spots basally; fore tibia with yellow stripe frontally; hind tibia yellow, infuscate in apical 0.3; mid tarsus brown; hind tarsus black. Wings hyaline; pterostigma brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	discussion	Remarks. This new species is similar to the European Heresiarches eudoxius (Wesmael) and runs to this species in the key to known Heresiarches species (Riedel 2013: 2061 – 2063). It differs mainly by a slenderer flagellum (1 st flagellomere 4.0 × longer than wide and widest flagellomere 1.2 × wider than long in H. eudoxius) and might represent an Oriental subspecies of H. eudoxius.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCDFFE9BDAEF98DFD116107.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE9BDAEFBFCFDF0606F.taxon	materials_examined	Material. Indonesia: Java, Cibodas, 1600 m, 2 ♀♀ 2 ♂♂ 3. viii. 1986, leg. M. Riedel (coll. Riedel); W Java, Gunung Botol, 6.7257 ° S 106.488 ° E, 1638 m, 2 ♂♂ 24. iv. 2016, leg. X. Mengal & T. Burton (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE9BDAEFBFCFDF0606F.taxon	discussion	Remarks. The ♂♂ differ from the description of Heinrich (1980: 165 – 166) by their black hind coxa with large ventral ivory mark and small ivory spot dorsally. They differ from L. abdominalis (Uchida) by their ivory malar spaces.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE9BDAEFBFCFDF0606F.taxon	distribution	Distribution. Only known from Java.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE8BDAEFA54FD3A626F.taxon	description	(Figs 25 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE8BDAEFA54FD3A626F.taxon	materials_examined	Holotype. (♀) Indonesia: S Halmahera, 20 km S Payahe, Sagutora, Mal. trap 12, c 125 m, 18. ii. – 18. iii. 1995, C. van Achterberg, R. de Vries & Y. Yasir, RMNH´ 95 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE8BDAEFA54FD3A626F.taxon	etymology	Etymology. The species name refers to the mainly blackish coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE8BDAEFA54FD3A626F.taxon	description	Description. ♀. Body length 9.2 mm. Head. Flagellum with 31 flagellomeres, almost filiform; 1 st flagellomere 2.7 × longer than wide, about 10 th flagellomere square, widest flagellomeres square. Temple very short, strongly and linearly narrowed behind eye, dorsally 0.25 × as long as eye (fig. 25 B). OED 1.6 × and OOD 0.7 × ocellar diameter. Head smooth and shining; face, clypeus and gena with very sparse fine punctures. Clypeus almost flat; apical margin sharp and straight (fig. 25 A). Mandible long, with two teeth, lower tooth small and slightly bent inwards (by 30 °). Malar space 0.5 × as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base, both carinae low. Mesosoma. Covered with short setae. Notaulus not impressed. Mesoscutum with rather dense fine punctures in frontal half and sparse punctures posteriorly, shining. Pronotum almost smooth, with few rugae at ventral edge; epomia distinct. Mesopleuron with fine sparse punctures dorsally and rather dense punctures ventrally, with some fine ridges at posterior margin. Epicnemial carina low. Speculum smooth and shining. Metapleuron with sparse punctures in frontal half and transverse striae posteriorly; juxtacoxal carina present. Scutellum moderately elevated, slightly longer than wide, with fine sparse punctures, with lateral carina in frontal half. Propodeum (fig. 25 C) completely carinated, spiracle slit-shaped, with short but distinct apophysis. Area basalis trapezoid, wider than long. Area superomedia hexagonal, 1.3 × longer than wide; costula in its frontal third. Hind coxa with fine dense punctures, without scopa; hind femur densely punctate, 4.1 × longer than wide. Hind tibia without denticular spurs externally. Claws not pectinate. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 4.0 × their diameter. Vein 1 cu-a slightly postfurcal by 1.0 × its width. Metasoma. Oxypygous. Petiole about as wide as high. Postpetiole moderately widened, without latero-median carina, smooth and with very sparse fine punctures; median field not separated. 2 nd tergite 1.3 × wider than long. Gastrocoelus not impressed. Thyridium small, transverse, 0.4 × as wide as the interval between thyridia. Tergites with fine superficial punctures, shining. 2 nd to 5 th sternites with median folds. Ovipositor sheath narrow, reaching behind metasomal apex by the length of 3 rd hind tarsomere. Colour. Black with slightly metallic shine. Rings of flagellomeres 6 – 17, upper frons and vertex except black stemmaticum, small spot of mandibular base and large median spot of 7 th tergite ivory. Palps reddish-brown. Legs black, fore and mid legs with reddish-brown suffusion. Wings slightly infuscate; pterostigma black. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCFFFE8BDAEFA54FD3A626F.taxon	distribution	Distribution. Only known from Halmahera.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC9FFEEBDAEF8DFFD24608B.taxon	description	(Figs 26 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC9FFEEBDAEF8DFFD24608B.taxon	materials_examined	Holotype. (♀) Indonesia, Südsulawesi, Bantimurung, 18. / 19.2.94, Oehlke (SDEI). Etymology. The species name refers to the wide ivory frontal orbits.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC9FFEEBDAEF8DFFD24608B.taxon	description	Description. ♀. Body length 12.0 mm. Head. Flagellum with 33 flagellomeres, almost filiform; 1 st flagellomere 2.4 × longer than wide, about 9 th flagellomere square, widest flagellomeres 1.2 × wider than long. Temple rather short, distinctly and roundly behind eye, with fine sparse punctures (fig. 26 B). OED 1.1 × and OOD 1.3 × ocellar diameter. Frons almost smooth. Face densely punctate medially, with sparse punctuation laterally. Clypeus almost flat, with sparse punctures; apical margin blunt, straight (fig. 26 A). Mandible with two teeth, ventral tooth much smaller than dorsal tooth, not bent inwards. Malar space 0.4 × as long as width of mandibular base. Gena with sparse punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus impressed in frontal 0.25 of mesoscutum. Mesoscutum with sparse, ± superficial punctures, strongly shining. Mesopleuron with dense punctures ventrally, with fine very sparse punctures dorsally. Speculum large and smooth. Metapleuron with rather dense punctures; juxtacoxal carina present. Scutellum slightly elevated, about as long as wide, with rather dense punctures, without lateral carina. Propodeum (fig. 26 C) completely carinated, without apophysis; spiracle slit-shaped. Area basalis 1.5 × wider than long, without median tubercle, ± confluent with area superomedia. Area superomedia heart-shaped, 1.2 × longer than wide; costula in its middle. Hind coxa with dense punctures and distinct scopa apico-ventrally. Hind femur densely but superficially punctate, 3.5 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a slightly postfurcal by 1.0 × its width. Metasoma (fig. 26 D). Oxypygous. Postpetiole moderately widened, without latero-median carina; median field somewhat elevated, with fine aciculation; lateral field coarsely punctate. 2 nd tergite 1.1 × wider than long. Gastrocoelus distinctly impressed, almost smooth. Thyridium slightly oblique, 0.8 × as wide as the interval between thyridia. 2 nd and 3 rd tergites densely rugose-punctate, dull, with longitudinal rugae medially. 4 th tergite with superficial puncture, shining. 2 nd to 4 th sternites with median folds. Ovipositor sheath slightly reaching behind metasomal apex. Colour. Black. Ivory are dorsal stripes of flagellomeres 7 – 17, palps, mandible except teeth, apical margin of clypeus, spot on gena, wide frontal orbit up to vertex (fig. 27 B), upper margin of pronotum, subtegular ridge, posteroventral spot of mesopleuron, prescutellar ridge, postscutellum, spot in front of propodeal spiracle and posterolateral stripe on propodeum (fig. 26 C). Tergites black; postpetiole, 2 nd and 3 rd tergites with posterolateral ivory spots; 6 th and 7 th tergites mainly ivory (fig. 26 D). Fore and mid coxae brown, with apical ivory spots; hind coxa black, with dorsal ivory spot; fore and mid trochanters ivory; fore and mid legs otherwise reddish-brown; hind leg black; hind tibia with diffuse subbasal dark reddish ring. Wings hyaline; pterostigma black. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC9FFEEBDAEF8DFFD24608B.taxon	discussion	Remarks. This new species runs to Lissosculpta celebensis Heinrich in his key of Lissosculpta species from Sulawesi (Heinrich 1934: 195 – 196). It differs by its smaller size and reduced ivory colour pattern: face, collar, mesoscutum and scutellum are entirely black. It also resembles L. irregularis Heinrich but differs by the surface structure of its postpetiole, black face and mainly dark fore and mid coxae. L. irregularis seems to be rather variable (Heinrich 1934: 202), and L. albifrons might represent a variant of this species only.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC9FFEEBDAEF8DFFD24608B.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC8FFEEBDAEFA71FE036283.taxon	materials_examined	Material. Indonesia: S Sulawesi, N of Rappang, 7 km E of Carekang, 1 ♀ and 1 ♂ 1 – 2. v. 1999, leg. Becvar & Zabransky (Linz); N Sumatra, Ketambe, 400 m, Mal. trap, near edge rainforest, 1 ♂ ii. 1995, leg. Y. van Nierop & C. van Achterberg (Leiden); W Sumatra, Jorong Akar Gadang, Simpang Tanjung Nam IV, Danau Kembar, Solok, 1.0579 ° N 100.697 ° E, 1412 m, 2 ♀♀ 13. v. 2016, leg. H. Hamid (ZSM); Sumbawa, nr Pancasila, Gn. Tambora, Mal. trap 8, c 500 m, 3 ♂♂ 27. ix – 2. x. 1993, leg. C. van Achterberg (Leiden); Lombok, Suranadi N. R., nr. Mataram, c 185 m, Mal. trap 3, 1 ♂ 9 – 29. iii. 1991, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC8FFEEBDAEFA71FE036283.taxon	discussion	Remarks. One ♀ from Sumatra has no posterolateral ivory spots on 5 th tergite. The ♂ from Lombok has no ivory flagellar ring. These specimens are otherwise typical.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFC8FFEEBDAEFA71FE036283.taxon	distribution	Distribution. Known from Bangladesh, India and Indonesia including Lombok, Sulawesi and Sumatra, new record for Sumbawa.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCBFF93BDAEF8DDFBA76473.taxon	description	(Figs 27 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCBFF93BDAEF8DDFBA76473.taxon	materials_examined	Holotype. (♀) Indonesia: S Halmahera, nr Payahe, Tayawi, 22. ii. 1995, Y. Yasir, RMNH´ 95 (Leiden). Paratypes. (♀) Indonesia: S Halmahera, nr Payahe, road to Weda, 25. ii. 1995, c 300 m, C. van Achterberg, RMNH´ 95; (5 ♂♂) Indonesia: S Halmahera, between Payahe & Gita Woda, Mal. trap 5, c 25 m, 17. ii. – 17. iii. 1995, C. van Achterberg, R. de Vries & Y. Yasir, RMNH´ 95 (Leiden and coll. Riedel); (♂) Indonesia: S Halmahera, between Payahe & Gita Woda, Mal. trap 8, c 25 m, 17. ii. – 17. iii. 1995, C. van Achterberg, R. de Vries & Y. Yasir, RMNH´ 95; (♂) Indonesia: S Halmahera, between Payahe & Gita Woda, Mal. trap 7, c 25 m, 17. ii. – 17. iii. 1995, C. van Achterberg, R. de Vries & Y. Yasir, RMNH´ 95; (♂) Indonesia: Sula Isl., Mangole, Buluhaya, Mal. trap 18, c 90 m, 6 – 23. iii. 1995, C. van Achterberg & Y. Yasir, RMNH´ 95; (♂) Indonesia: Sula Isl., Mangole, 2 km W Mandafuhi camp, Mal. trap 19, c 70 m, 7 – 24. iii. 1995, C. van Achterberg & Y. Yasir, RMNH´ 95; (♂) Indonesia: N Ceram, 20 km E Wahai, nr Pasahari, rainforest, 3 – 20. iii. 1987, Mal. trap 19, C. van Achterberg & R. de Vries (Leiden and coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCBFF93BDAEF8DDFBA76473.taxon	description	Description. ♀. Body length 13.5 – 15.5 mm. Head. Flagellum with 36 – 39 flagellomeres, moderately lanceolate; 1 st flagellomere 1.9 × longer than wide, about 9 th flagellomere square, widest flagellomeres 2.0 × wider than long. Temple rather short, moderately and roundly behind eye, with fine punctures (fig. 27 B). OED 1.2 × and OOD 1.2 × ocellar diameter. Frons almost smooth. Face densely punctate medially, with sparse punctuation laterally. Clypeus almost flat, with sparse punctures; apical margin blunt, slightly bulging medially (fig. 27 A). Mandible with two teeth, ventral tooth smaller than dorsal tooth, not bent inwards. Malar space 0.4 – 0.5 × as long as width of mandibular base. Gena with sparse punctures. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus impressed in frontal 0.4 of mesoscutum. Mesoscutum with dense fine punctures on median lobe, with fine sparse or very sparse punctures laterally, shining. Mesopleuron with dense punctures. Speculum smooth and shining. Metapleuron with dense punctures; juxtacoxal carina present. Scutellum moderately elevated, 1.2 × wider than long, densely punctate, without lateral carina. Propodeum (fig. 27 C) completely carinated, without apophysis; spiracle slit-shaped. Area basalis wider than long, sometimes confluent with area superomedia. Area superomedia dome- or drum-shaped, 1.2 – 1.5 × longer than wide; costula in its middle. Hind coxa with dense punctures and distinct scopa apico-ventrally. Hind femur densely punctate, 3.3 – 3.4 × longer than wide. Claws simple. Areolet pointed or almost pointed frontally. Vein 1 cu-a postfurcal by 3.0 × its width. Metasoma. Oxypygous. Postpetiole moderately widened, without latero-median carina; median field not separated, with some punctures and fine partly obsolete aciculation. 2 nd tergite 1.2 × wider than long. Gastrocoelus distinctly impressed, almost smooth. Thyridium transverse, 0.7 × as wide as the interval between thyridia (fig. 27 D). 2 nd and 3 rd tergites densely rugose-punctate, dull; 2 nd tergite ± striate medially. Following tergites with superficial punctures, shining. 2 nd to 4 th sternites with median folds. Ovipositor sheath slightly reaching behind metasomal apex. Colour. Black. Apical margin of clypeus and tegula reddish. Scape brownish. Ivory are dorsal stripes of flagellomeres 7 – 18 / 19, palps, mandible basally, clypeus laterally, face except black median spot, complete orbits, collar and upper margin of pronotum, two paramedian stripes of mesoscutum, subtegular ridge, longitudinal band of ventral mesopleuron, prescutellar ridge, side of scutellum, postscutellum, spot in front of propodeal spiracle and posterolateral stripe on propodeum (fig. 27 C). Tergites black; postpetiole, 2 nd and 3 rd tergites with posterolateral ivory spots, sometimes medially confluent on 2 nd tergite (fig. 27 D); 6 th and 7 th tergites mainly ivory. Legs red; coxae with yellow spots dorsally, hind tarsus reddish-brown. Wings hyaline; pterostigma black. ♂: Body length 13 – 15 mm. Flagellum with 37 – 39 flagellomeres, bristle-shaped; 1 st flagellomere 2.5 × longer than wide. Tyloids on flagellomeres 8 – 19 / 21, oval, maximally 0.5 × as long as their flagellomeres. OED and OOD 1.0 × ocellar diameter. Face with moderate paramedian impressions, also around anterior tentorial pits. Hind femur 3.5 – 3.6 × longer than wide. 2 nd tergite 1.1 × longer than wide. Apical margin of hypopygium tongue-like elongate medially. Structure otherwise as in ♀. Colour. As in ♀, but flagellum black, reddish ventrally, without ivory ring. Face entirely, scape ventrally, propleuron, mesosternum, fore and mid coxae and trochanters, and hind trochantellus ivory. Tergites black; petiole ± reddish; basal tergites sometimes with partly reddish suffusion; postpetiole with posterolateral ivory spots; 2 nd and 3 rd tergites with apical ivory bands (medially narrowed), 4 th tergite sometimes with reddish or yellowish posterolateral spot; 6 th and 7 th tergite with large apico-median ivory spots. Hind tibia sometimes basally and apically narrowly infuscate; hind tarsus blackish. Variation. Two ♂♂ from Sula Islands have a slightly different colour pattern: 1 st to 4 th tergites black, with distinct ivory posterolateral spots; ivory orbital stripes shortly interrupted between frontal and vertical orbits and between vertical and outer orbits; hind femur red, black in apical 0.2 – 0.4; hind tibia black, with red subbasal ring. Structure and colour pattern are otherwise typical. One ♂ from Seram differs from the ♂♂ from Halmahera by: flagellomeres 12 – 19 ivory dorsally; mesosternum ivory, partly reddish and blackish; legs entirely red; 1 st to 4 th tergites mainly reddish-brown, 1 st to 3 rd tergites with ivory posterolateral spots; 6 th and 7 th tergites and paramere ivory. Structure and coloration are otherwise typical.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCBFF93BDAEF8DDFBA76473.taxon	discussion	Remarks. This species is closely related to Lissosculpta albopicta Horstmann and might represent a variable subspecies of this widely distributed species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFCBFF93BDAEF8DDFBA76473.taxon	distribution	Distribution. Only known from the Moluccas (Halmahera, Seram and Sula).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF93BDAEFE29FCD367BF.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 1, c 750 m, 1 ♂ 7. ii. – 29. iii. 1995, leg. C. van Achterberg; C Sulawesi, nr Luwuk, Salodik, c 400 m, 1 ♂ 21 – 31. x. 1989, Mal. trap 14, leg. C. van Achterberg; W Bali, nr. Negara, rainforest above Batuagung, c 575 m, 1 ♂ 4 – 13. xii. 1991, Mal. trap 1, leg. C. van Achterberg (all in Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF93BDAEFE29FCD367BF.taxon	distribution	Distribution. Known from Sulawesi, new record for Bali.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF93BDAEFD6DFD2463EB.taxon	materials_examined	Material. Indonesia: SW Sulawesi, Gn. Bulusaraung, nr. Camba, Malawa, Mal. trap 1, c 800 m, 1 ♂ 21. ii. – 25. iii. 1997, Mal. trap 2, c 750 m, 1 ♂ 7. ii. – 28. iii. 1995, Mal. trap 3, c 750 m, 1 ♂ 7. ii. – 28. iii. 1995, leg. C. van Achterberg & R. de Vries (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF93BDAEFD6DFD2463EB.taxon	discussion	Remarks. ♂. Body length 10.5 mm. Head. Flagellum with 34 flagellomeres, bristle-shaped; 1 st flagellomere 3.4 × longer than wide. Tyloids on flagellomeres 9 – 20, long-oval, maximally 0.5 × as long as their flagellomeres. Temple short, strongly and roundly narrowed behind eye. OED 1.1 × and OOD 1.0 × ocellar diameter. Face densely punctate. Clypeus almost flat, densely punctate; apical margin blunt, almost straight. Mandible with two teeth, ventral tooth much smaller and slightly bent inwards. Malar space 0.6 × as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus distinct in frontal 0.3 of mesoscutum. Mesoscutum with fine rather dense punctures, shining between punctures. Mesopleuron densely punctate, speculum smooth. Metapleuron densely punctate, partly smooth; juxtacoxal carina present. Scutellum moderately elevated, 1.4 × wider than long, with sparse punctures, without lateral carina. Propodeum completely carinated, without apophysis, spiracle slit-shaped. Area basalis smooth, without median tubercle. Area superomedia heart-shaped, about as long as wide; costula reaching at 7 / 10 of its length. Hind femur 3.8 × longer than wide. Areolet pentagonal, frontal distance of veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Postpetiole moderately widened, almost smooth; median field not separated. 2 nd tergite 1.1 × longer than wide. Gastrocoelus distinctly impressed, with some fine ridges. Thyridium almost transverse, 0.8 × as wide as the interval between thyridia. 2 nd to 4 th tergites coarsely punctate, shining. Apical margin of hypopygium tongue-like elongate medially. Colour. Black. Flagellum black, ± reddish-brown ventrally. Mandible except teeth, clypeus, face, gena and orbits (with a short interruption at vertex), scape ventrally and tegula cream yellow. Ivory are palps, frontal and upper margins of pronotum, subtegular ridge ventral mesopleuron (dorsal margin ± reddish) and complete mesosternum, prescutellar ridge and side of scutellum. Postscutellum, metapleuron and propodeum red. All tergites red. Legs red; fore and mid coxae and trochanters ivory; hind tarsus black. Wings hyaline, pterostigma reddish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF93BDAEFD6DFD2463EB.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF91BDAEF8D5FD06616B.taxon	description	(Figs 28 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF91BDAEF8D5FD06616B.taxon	materials_examined	Holotype. (♂) Indonesia: N Ceram, 20 km E Wahai, nr. Pasahari, rainforest, 25 m, 3 – 20. iii. 1997, Mal. trap 18, C. van Achterberg & R. de Vries, RMNH´ 97 (Leiden). Paratype. (♂) Indonesia: N Ceram, 11 km E Wahai, nr. PHPA-Q, coastal rainforest, 2 – 20. iii. 1997, Mal. trap 11, C. van Achterberg & R. de Vries, RMNH´ 97 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF91BDAEF8D5FD06616B.taxon	description	Description. ♂. Body length 12.0 – 12.5 mm. Head. Flagellum with 36 – 37 flagellomeres, bristle-shaped; 1 st flagellomere 2.6 × longer than wide. Tyloids on flagellomeres 7 – 20, oval, maximally 0.4 × as long as their flagellomeres. Temple rather long, moderately and roundly narrowed behind eye (fig. 28 B). OED 1.2 × and OOD 1.5 × ocellar diameter. Frons smooth, with few punctures dorso-laterally. Face densely punctate. Clypeus almost flat, with rather dense punctures; apical margin blunt, almost straight but slightly bulging medially (fig. 28 A). Mandible with two teeth, ventral tooth much smaller and slightly bent inwards. Malar space 0.4 × as long as width of mandibular base. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus distinct in frontal 0.25 of mesoscutum. Mesoscutum with dense punctures, shining between punctures. Mesopleuron coarsely punctate ventrally, with finer punctures dorsally; speculum smooth. Metapleuron densely punctate dorsally, partly smooth above juxtacoxal carina. Scutellum moderately elevated, 1.2 × wider than long, with dense punctures, without lateral carina. Propodeum (fig. 28 C) completely carinated, without apophysis, spiracle slit-shaped. Area basalis smooth, without median tubercle. Area superomedia heart-shaped, about as long as wide, frontal carina ± obsolete; costula reaching at 6 / 10 of its length. Hind femur 3.7 – 3.8 × longer than wide. Areolet pentagonal, frontal distance of veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a postfurcal by 1 – 3 × its width. Metasoma. Postpetiole moderately widened; median field slightly elevated, finely aciculate; lateral field punctate. 2 nd tergite 1.1 × wider than long. Gastrocoelus distinctly impressed, almost smooth. Thyridium transverse, 0.7 × as wide as the interval between thyridia (fig. 28 D). 2 nd and 3 rd tergites coarsely rugose-punctate, striate medially, dull. Following tergites with superficial punctures, ± shining. Apical margin of hypopygium rounded medially. Colour. Black. Flagellum black, reddish ventrally, flagellomeres 8 / 9 – 18 / 20 ivory dorsally. Ivory are palps, mandible except teeth, clypeus, face, gena, stripes on outer, frontal and vertical orbits, scape ventrally, propleuron, collar and upper margin of pronotum, subtegular ridge, two short paramedian stripes of mesoscutum, ventral mesopleuron (dorsal margin ± reddish) and mesosternum, prescutellar ridge and side of scutellum, spot in front of propodeal spiracle and posterolateral stripe of propodeum (fig. 28 C). Tegula and postscutellum red. 1 st to 5 th tergites red, sometimes petiole ± blackish; postpetiole with diffuse yellowish posterolateral spot. 6 th and 7 th tergites black, apical band of 6 th tergite and median spot of 7 th tergite ivory. Legs red; fore and mid coxae and trochanters ± ivory. Wings hyaline; pterostigma blackish. ♀ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF91BDAEF8D5FD06616B.taxon	discussion	Remarks. These new ♂♂ resemble Lissosculpta silvarum Heinrich, 1934 from Sulawesi but differ by finely aciculate postpetiole, shorter area superomedia, paramedian ivory stripes of mesoscutum, and reddish instead of black 4 th and 5 th tergites. They might represent a subspecies of Lissosculpta silvarum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB5FF91BDAEF8D5FD06616B.taxon	distribution	Distribution. Only known from Seram.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF91BDAEFB51FD2460F3.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 1 ♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF91BDAEFB51FD2460F3.taxon	discussion	Remarks. The available ♂ differs from the original description (Heinrich 1934: 202 – 203) by its entirely black flagellum without ivory ring, narrowly blackish apex of hind tibia and black hind tarsus. There seem to be several closely related Lissosculpta species in Sulawesi. Therefore, the determination of this ♂ is not unequivocal.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF91BDAEFB51FD2460F3.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF97BDAEF9A8FB9766B3.taxon	description	(Figs 29 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF97BDAEF9A8FB9766B3.taxon	materials_examined	Holotype. (♀) Indonesia: W Bali, nr. Negara, rainforest above Batuagung, c 575 m, 4 – 13. xii. 1991, Mal. trap 1, C. v. Achterberg, RMNH´ 91 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF97BDAEF9A8FB9766B3.taxon	description	Description. ♀. Body length 10.2 mm. Head. Flagellum with 45 flagellomeres, distinctly lanceolate; 1 st flagellomere 2.0 × longer than wide, about 8 th flagellomere square, widest flagellomeres 2.3 × wider than long. Temple very short, strongly and roundly narrowed behind eye. OED 1.2 × and OOD 0.7 × ocellar diameter. Frons smooth and shining, with some punctures laterally. Face densely punctate. Clypeus with dense punctures medially and sparse punctures laterally; apical margin blunt, slightly bulging medially (fig. 29 A). Malar space 0.7 × as long as width of mandibular base. Gena with sparse punctures. Genal carina reaching mandibular base. Mesosoma. Notaulus distinct in frontal 0.5 of mesoscutum. Mesoscutum densely punctate, shining. Mesopleuron densely and finely punctate, dorsal and posterior margins with some coarse rugae. Metapleuron coarsely rugose-punctate; juxtacoxal carina present. Scutellum slightly elevated, slightly longer than wide, with sparse punctures and complete lateral carina. Propodeum (fig. 29 B) completely carinated, without apophysis. Area superomedia large, slightly longer than wide, with coarse rugae; costula reaching at 4 / 10 of its length. Hind coxa with dense punctures, without scopa. Hind femur 4.3 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 3.0 × their diameter; vein 2 cu-m slightly distad its middle. Vein 1 cu-a antefurcal by 2.0 × its width. Nervellus of hind wing distinctly reclivous, intercepted at apical 0.7. Metasoma. Oxypygous; 2 nd to 4 th sternites with median folds. Postpetiole distinctly widened; median field not separated, almost smooth; lateral field punctate. Gastrocoelus slightly impressed, with some ridges. Thyridium small, slightly oblique, 0.3 × as wide as the interval between thyridia (fig. 29 C). 2 nd tergite about as wide as long, with distinct lateral impressions. 2 nd and 3 rd tergites densely rugose-punctate, dull; 4 th tergite with fine punctures, shining; following tergites almost smooth. Ovipositor sheath slightly extending behind metasomal apex. Colour. Black with extended ivory coloration. Scape and tegula reddish. Ivory are stripes on flagellomeres 9 – 17, palps, mandible except teeth, clypeus, face except brown central spot (fig. 29 A), gena, complete orbits, collar, fronto-ventral stripes and upper margin of pronotum, two short paramedian stripes of mesoscutum, subtegular ridge, wide ventral band of mesopleuron, apical spot on metapleuron, prescutellar ridge, scutellum laterally and apically, postscutellum, spot in front of propodeal spiracle and posterolateral spot on propodeum (fig. 29 B), posterolateral spot of postpetiole, apical bands of 2 nd and 3 rd tergites (narrowed medially), large posterolateral stripe of 4 th tergite and small posterolateral spot on 5 th tergite; apical band of 6 th tergites and entire 7 th tergite. Legs reddish; fore and mid coxae and trochanters and hind coxa ventrally and internally ivory; hind tibia and tarsus black. Wings hyaline; pterostigma blackish.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB7FF97BDAEF9A8FB9766B3.taxon	discussion	Remarks. This new species runs to Losgna bambusicola Heinrich in the key for Oriental Losgna species (Heinrich 1965: 138 – 139), but differs by its stouter 1 st flagellomere, less strongly lanceolate flagellum, shorter malar space, slightly different coloration of metasoma and entirely blackish hind tibia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF97BDAEFC68FC5F601B.taxon	materials_examined	Material. Indonesia: Sumbawa, nr Pancasila, Gn. Tambora, Mal. trap 5, c 1025 m, 1 ♂ 22. ix. – 2. x. 1993, leg. C. van Achterberg (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF97BDAEFC68FC5F601B.taxon	discussion	Remarks. The available ♂ has extended ivory marks. In addition to the ivory coloration mentioned by Heinrich (1965: 140), the following parts are ivory: paramedian stripes on mesoscutum, longitudinal band on mesopleuron, prescutellar ridge, apical spot on metapleuron and area spiracularis in front of the spiracle.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF97BDAEFC68FC5F601B.taxon	distribution	Distribution. Known from Borneo (Sabah), new record for Sumbawa.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF97BDAEFAC0FD24631F.taxon	materials_examined	Material. Indonesia: SE Sulawesi, Watuwila, Base Camp c 200, Mal. trap 11, 1 ♂ 12 – 15. x. 1989, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF97BDAEFAC0FD24631F.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF96BDAEF9C4FC5A6457.taxon	materials_examined	Material. Malaysia: SE Sabah, nr Danum Valley Field C, Mal. trap 12, W 12, c 150 m, 1 ♂ 20. x. – 22. xii. 1987, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF96BDAEF9C4FC5A6457.taxon	discussion	Remarks. ♂. Body length 11 mm. Flagellum with 40 flagellomeres; 1 st flagellomere 3.8 × longer than wide. Tyloids on flagellomeres 9 – 20, long-oval, maximally 0.5 × as long as their flagellomeres. Hind femur 4.2 × longer than wide. 2 nd tergite 1.2 × longer than wide. Colour. Black. Ivory are flagellomeres 9 – 18, palps, mandible except teeth, clypeus, face, gena, narrow stripes of frontal orbit, scape ventrally, spot on tegula, apical margins of scutellum and postscutellum, weak posterolateral spot of propodeum around apophysis, 6 th tergite mainly, 7 th tergite and paramere entirely, fore and mid coxae and trochanters. Fore and mid legs otherwise pale yellow, their tarsi ± infuscate; hind coxa and trochanter and hind femur yellow; hind tibia and tarsus black. Postpetiole brownish apically, 2 nd tergite with an apical reddish band. Pterostigma black. This ♂ differs from the original description of Losgna nigrita (Heinrich 1965: 140 – 141) by the apical reddish band of 2 nd tergite. It is otherwise typical and might represent a colour variant or subspecies of that species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB1FF96BDAEF9C4FC5A6457.taxon	distribution	Distribution. Known from Malaysia and Indonesia including Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	description	(Figs 30 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	materials_examined	Holotype. (♀) Indonesia: N Sumatra, Ketambe, 400 m, Mal. trap, near edge rainforest, xi. 1995, Y. van Nierop & C. van Achterberg, RMNH´ 99 (Leiden). Paratype. (♂) Indonesia: N Sumatra, Ketambe, 400 m, Mal. trap, near edge rainforest, iii. 1995, Y. van Nierop & C. van Achterberg, RMNH´ 99 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	etymology	Etymology. The species name refers to the reddish area basalis.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	description	Description. ♀. Body length 9.5 mm. Head. Flagellum with 44 flagellomeres, moderately lanceolate; 1 st flagellomere 2.4 × longer than wide, about 12 th flagellomere square, widest flagellomeres 1.8 × wider than long. Temple rather short, strongly and roundly narrowed behind eye, dorsally 0.4 × as long as eye (fig. 30 B). OED 1.3 × and OOD 0.8 × ocellar diameter. Frons smooth and shining, with very sparse punctures. Face rather densely punctate. Clypeus almost flat, with sparse punctures; apical margin blunt, slightly bulging medially (fig. 30 A). Malar space 0.8 × as long as width of mandibular base. Gena with very sparse punctures. Genal carina reaching hypostomal carina at mandibular base. Mesosoma. Notaulus distinct in frontal 0.4 of mesoscutum. Mesoscutum densely but superficially punctate, shining. Mesopleuron with superficial punctures, shining, with short coarse striae at posterior margin. Metapleuron rugose-punctate; juxtacoxal carina present. Scutellum almost flat, slightly longer than wide, with sparse punctures and complete lateral carina. Propodeum (fig. 30 C) completely carinated, without apophysis. Area basalis with median tubercle. Area superomedia large, about as long as wide, with coarse irregular striae; costula reaching at 3 / 10 of its length. Hind coxa with dense punctures, without scopa. Hind femur 4.0 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their diameter. Vein 1 cu-a antefurcal by 1.0 × its width. Metasoma. Oxypygous; 2 nd to 4 th sternites with median folds. Postpetiole distinctly widened, almost smooth with fine very sparse punctures; median field not separated. Gastrocoelus small, longer than wide, with some ridges. Thyridium very small, strongly oblique, 0.2 × as wide as the interval between thyridia. 2 nd tergite about as wide as long, with distinct paramedian impressions (fig. 30 D). 2 nd tergite with coarse longitudinal rugose-punctate striation. 3 rd tergites rugose-punctate in basal half; with fine dense punctures and shining apically; following tergites with fine punctures, shining. Ovipositor sheath rather long, extending distinctly behind metasomal apex. Colour. Black. Ivory are flagellomeres 8 – 17 dorsally, palps, mandible except teeth, clypeus, face, gena, narrow frontal orbits, spot on vertex, scape ventrally, spot of subtegular ridge, prescutellar ridge, scutellum laterally and apically, spot of postscutellum, spot in front of propodeal spiracle and posterolateral spot of propodeum around outer edge of area dentipara, petiole, small posterolateral spots of postpetiole and 2 nd tergite (fig. 30 D), wide apical band of 6 th tergites and entire 7 th and 8 th tergites. Collar and upper margin of pronotum, tegula, ventral spot of mesopleuron and area basalis ± reddish. Legs red; fore and mid coxae and trochanters ivory; hind tibia and tarsus black. Wings hyaline; pterostigma blackish, paler basally. ♂: Body length 10.0 mm. Flagellum with> 34 flagellomeres (tips broken off); 1 st flagellomere 2.6 × longer than wide. Tyloids on flagellomeres 11 – 19, long-oval, maximally 0.6 × as long as their flagellomeres. OED 1.5 × and OOD 1.0 × ocellar diameter. Malar space 0.8 × as long as width of mandibular base. Genal carina reaching mandibular base. Notaulus short, impressed in frontal 0.2 of mesoscutum. Area superomedia 2.0 × wider than long, with longitudinal striation. Hind femur 4.5 × longer than wide. 2 nd tergite 1.1 × longer than wide. Apical margin of hypopygium rounded. Otherwise as described for the ♀. Colour. As in ♀. Different are: flagellum reddish-brown proximally, infuscate distally; flagellomeres 11 – 19 with ivory rings. Frontal orbit up to vertex, spot of collar, upper margin and ventral edge of pronotum, spot on tegula and wide stripe on ventral mesopleuron, 6 th tergite mainly and 7 th tergite and paramere entirely ivory. 1 st to 3 rd tergites with small posterolateral ivory spots; 2 nd tergite with wide reddish-yellow apical band. Hind trochantellus brownish; hind tibia brownish, hind tarsus black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	discussion	Remarks. This new species is closely related to Losgna cariniscutis (Cameron, 1905) and probably represents a subspecies of that species. The ♂ of this species differs from the ♂ of Losgna cariniscutis by the ivory ring of flagellum, apical reddish-yellow band of 2 nd tergite and wide ivory stripe on ventral mesopleuron.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB0FF95BDAEFE0CFD1C6473.taxon	distribution	Distribution. Only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF94BDAEFF08FBF364BB.taxon	materials_examined	Material. Malaysia: SE Sabah, nr Danum Valley Field c., WON 1, c 150 m, Mal. trap 5, 1 ♀ 20. iv. – 12. vii. 1987, 1 ♂ 23. viii. – 13. xi. 1987, leg. C. van Achterberg & D. Kennedy; Indonesia: N Sumatra, Ketambe, c. 400 m, near N. P. Gn. Leuser, Mal. trap, iv. 1994, leg. Y. van Nierop & C. van Achterberg; Sulawesi, nr Bantaeng, Borong Rappoa, Gn. Daulu, Mal trap 16. c 700 m, 2 ♂♂ 4 – 24. iv. 1991, leg. C. van Achterberg; SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 2, c 200 m, 1 ♀ 10 – 15. x. 1989, leg. C. van Achterberg (all Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF94BDAEFF08FBF364BB.taxon	distribution	Distribution. Known from Java and Sulawesi, new record for Borneo and Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF94BDAEFE60FD246603.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 1 ♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF94BDAEFE60FD246603.taxon	discussion	Remarks. Body length 16 mm. Flagellum with 37 flagellomeres, bristle-shaped; 1 st flagellomeres 3.1 × longer than wide. Tyloids on flagellomeres 7 – 19, long-oval, 0.7 × as long as the flagellomeres. OED 1.4 × and OOD 0.8 × ocellar diameter.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF94BDAEFE60FD246603.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF9BBDAEF8A9FD2467DB.taxon	description	(Figs 31 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF9BBDAEF8A9FD2467DB.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 1 ♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF9BBDAEF8A9FD2467DB.taxon	description	Description. ♂. Body length 12.5 mm. Head. Flagellum with 34 flagellomeres, bristle-shaped; 1 st flagellomeres 3.1 × longer than wide. Tyloids on flagellomeres 8 – 15, small (0.5 × as long as their flagellomeres), long-oval, centrally located. Temple short, strongly and roundly narrowed behind eye. OED 1.0 × and OOD 1.1 × ocellar diameter. Mesosoma. Scutellum moderately elevated, with strong and complete lateral carina. Propodeum (fig. 31 A) completely carinated; are basalis with strong median tubercle. Area superomedia horseshoe-shaped, slightly longer than wide. Hind femur densely punctate, 3.5 × longer than wide. Metasoma. Postpetiole with fine aciculation medially, with coarse punctures laterally. 2 nd tergite about as long as wide. Gastrocoelus narrow, with 2 oblique ridges. Thyridium oblique, 0.7 × as long as the interval between thyridia (fig. 31 B). Colour. Scape reddish-brown dorsally, yellow ventrally. Seven basal flagellomeres reddish, following flagellomeres blackish. Fore and mid coxae and all trochanters and trochantelli cream-yellow. Hind coxa red, brownish baso-ventrally, cream yellow dorsally. Petiole black; postpetiole reddish-brown basally, yellowish apically. Tergites reddish; 2 nd to 4 th tergites with diffuse yellowish posterolateral spots; median spot of 7 th tergite and paramere yellowish. Coloration otherwise as described for the ♀ (Heinrich 1934: 211).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB2FF9BBDAEF8A9FD2467DB.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9BBDAEFC81FC6E616B.taxon	materials_examined	Material. Malaysia, SW Sabah, nr. Long Pa Sia, Payakalaba, 1 ♀ 25. xi. – 1. xii. 1987, c 1010 m, Mal. trap 15, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9BBDAEFC81FC6E616B.taxon	discussion	Remarks. Body length 18 mm. Flagellum with 46 flagellomeres, distinctly lanceolate; 1 st flagellomere 1.7 × longer than wide, widest flagellomere 2.2 × wider than long. Hind coxa densely punctate, without scopa, dorsally with distinct oblique striae. Hind femur 3.8 × longer than wide. 2 nd tergite about as long as wide. Gastrocoelus not impressed. Thyridium small, slightly oblique, 0.3 × as wide as the interval between thyridia. Otherwise as described by Heinrich (1980: 134 – 136).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9BBDAEFC81FC6E616B.taxon	distribution	Distribution. Known from India and Myanmar, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9ABDAEFB51FD6D65C3.taxon	type_taxon	Type species. Moluccarches albimaculatus nov. sp.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9ABDAEFB51FD6D65C3.taxon	diagnosis	Diagnosis. Small, body length 7.0 mm. Head. Flagellum long, slightly lanceolate; widest flagellomeres square. Temple rather short, slightly narrowed behind eye. Clypeus large, almost flat; apical margin sharp, slightly bulging apically, rounded medially (fig. 32 A). Mandible sickle-shaped, ventral tooth absent. Genal carina reaching mandibular base. Mesosoma (figs 32 C, 32 D). Rather stout, mesoscutum 1.1 × wider than long. Epomia very long. Notaulus shortly impressed at frontal margin of mesoscutum. Epicnemial carina low, dorsally reaching subtegular ridge. Sternaulus impressed in frontal half of mesopleuron. Juxtacoxal carina present. Scutellum distinctly but roundly elevated, wider than long, with complete and slightly lamelliform lateral carina. Propodeum (fig. 32 D) completely carinated; without distinct apophysis. Spiracle oval. Area basalis without median tubercle. Area superomedia hexagonal; anterior transverse carina reaching at 7 / 10 of its length. Hind coxa without scopa. Hind femur 3.2 - 3.4 × longer than wide. Claws simple. Areolet pentagonal; frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a slightly postfurcal. Metasoma (fig. 32 E). Oxypygous, stout. Petiole as wide as high. Postpetiole distinctly widened; without latero-medial carina, median field not separated. Gastrocoelus impressed, ± smooth. Thyridium small, transverse. Ovipositor sheath long and slender. Colour. Black with extensive ivory coloration. ♂ and hosts unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBDFF9ABDAEFB51FD6D65C3.taxon	discussion	Remarks. Due to the sickle-shaped and unidentate mandible, this new genus resembles Celebarches Heinrich and runs to that genus in the keys of Heinrich (1934) and Townes et al. (1961). It differs by the stout habitus, oxypygous metasoma with thin and long ovipositor sheath, the epicnemial carina which reaches the subtegular ridge dorsally, and a different carination of the propodeum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBCFF99BDAEF925FD4561FF.taxon	description	(Figs 32 A – E)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBCFF99BDAEF925FD4561FF.taxon	materials_examined	Holotype. (♀) Indonesia: Sula Isl., Mangole, near Buya, Mal. trap 14, c 490 m, 2 – 22. iii. 1995, C. van Achterberg & Y. Yasir, RMNH (Leiden). Paratype. (♀) similar label (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBCFF99BDAEF925FD4561FF.taxon	etymology	Etymology. The species name refers to the extended ivory coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBCFF99BDAEF925FD4561FF.taxon	description	Description. ♀. Body length 7.0 mm. Head. Flagellum with 33 flagellomeres, long, slightly lanceolate; 1 st flagellomere 5.0 × longer than wide; widest flagellomeres square. Temple rather short, slightly narrowed behind eye, moderately and roundly narrowed apically (fig. 32 B). OED 1.5 × and OOD 1.3 × ocellar diameter. Temple and frons smooth, with fine sparse punctures. Face with rather dense fine punctures (fig. 32 A). Clypeus with rather dense punctures; apical margin sharp, slightly bulging apically, rounded medially. Mandible sickle-shaped, ventral tooth absent. Palps very long and slender; distal maxillary palp about 2 × longer than 4 th palp. Malar space 0.8 × as long as width of mandibular base. Gena with sparse punctures. Mesosoma. Side of pronotum almost smooth; epomia almost reaching the ventral margin of pronotum. Notaulus shortly impressed at frontal margin of mesoscutum. Mesoscutum densely punctate, shining. Ventral half of mesopleuron densely punctate frontally, coarsely striate posteriorly; dorsal part with fine punctures; speculum smooth and shining (fig. 32 C). Metapleuron densely punctate frontally, coarsely striate in apical 2 / 3. Scutellum distinctly but roundly elevated, 1.3 × wider than long, punctate, with complete and slightly lamelliform lateral carina. Propodeum (fig. 32 D) completely carinated; without distinct apophysis, but apical transverse carina slightly lamelliform at laterally. Spiracle oval, 2.0 × longer than wide. Area basalis trapezoid, wider than long, without median tubercle. Area superomedia hexagonal, 1.4 × wider than long; anterior transverse carina reaching at 7 / 10 of its length. Hind coxa densely punctate. Hind femur 3.2 – 3.4 × longer than wide. Areolet pentagonal; frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their width. Vein 1 cu-a postfurcal by 1.0 × its width. Nervellus of hind wing vertical, intercepted at apical 0.7. Metasoma (fig. 32 E). Oxypygous, stout. Postpetiole distinctly widened, with fine sparse punctures, smooth and shining. 2 nd tergite 1.5 × wider than long. Thyridium small, transverse, 0.3 × as wide as the interval between thyridia. 2 nd tergite densely punctate, shining; 3 rd tergite with superficial punctures; following tergites almost smooth. Ovipositor sheath extending behind metasomal apex by combined length of 6 th and 7 th tergites. Colour. Black with extensive ivory coloration. Scape and pedicel brownish, red ventrally. Flagellum black, flagellomeres 5 – 17 / 18 ivory. Ivory are palps, mandible except tooth, clypeus except black narrow margins, face, gena, wide complete orbits, upper margin of pronotum, tegula, subtegular ridge, scutellum and postscutellum, short and wide paramedian spots on mesoscutum, dorso-frontal and ventral spots of mesopleuron, posterolateral stripe of propodeum, entire 1 st tergite, and wide apical bands of all tergites (widely interrupted medially on 3 rd tergite and strongly narrowed medially on 4 th tergite). Legs reddish; fore and mid coxae and trochanters ivory; hind coxa ivory dorsally; hind trochanter blackish; fore and mid tarsi distally ± blackish; hind femur black in apical 0.2; hind tibia ± yellowish basally, infuscate in apical 0.2; hind tarsus except brown distal tarsomere ivory. Wings hyaline; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBCFF99BDAEF925FD4561FF.taxon	distribution	Distribution. Only known from Sula islands.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBFFF9FBDAEFAA5FBAC6603.taxon	description	(Figs 33 A – B)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBFFF9FBDAEFAA5FBAC6603.taxon	materials_examined	Material. Indonesia: Lombok, Pusuk, c 450 m, nr Mataram, Mal. Trap 7, 21 – 27. iii. 1991, C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBFFF9FBDAEFAA5FBAC6603.taxon	description	Description. ♂. Body length 5.0 mm. Head. Flagellum with 33 flagellomeres, bristle-shaped; 1 st flagellomere 3.7 × longer than wide. Tyloids on flagellomeres 10 – 15, long-oval, maximally 0.7 × as long as their flagellomeres. Temple long, slightly widened behind eye, moderately narrowed apically (fig. 33 B). Occiput strongly concave. OED 2.1 × and OOD 1.3 × ocellar diameter. Vertex around stemmaticum and frons densely rugose-punctate. Face densely punctate, rugose-punctate centrally. Clypeus slightly convex; apical margin sharp and with 6 strong denticles (fig. 33 A). Mandible unidentate, sickle-shaped. Malar space 0.4 × as long as width of mandibular base. Genal carina reaching mandibular base. Mesosoma. Upper margin of pronotum bulging. Notaulus shortly impressed on frontal mesoscutum. Mesoscutum densely punctate, shining. Mesopleuron densely punctate; speculum large, smooth. Epicnemial carina low, dorsally reaching subtegular ridge. Sternaulus impressed in frontal 0.4 of mesopleuron. Metapleuron with fine sparse punctures; juxtacoxal carina absent. Scutellum moderately and roundly elevated, 1.5 × wider than long, without lateral carina. Propodeum completely carinated, without apophysis; spiracle oval. Area basalis indicated as fine median ridge (as in Imeria). Area superomedia pentagonal, about as long as wide; costula reaching at 4 / 10 of its length. Area petiolaris with transverse rugae. Hind femur densely punctate, 4. 0 × longer than wide. Claws simple. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their diameter. Vein 1 cu-a interstitial. Metasoma. Petiole long and slender. Postpetiole slightly widened, with fine sparse punctures, shining. Gastrocoelus and thyridium not impressed. 2 nd tergite 1.1 × longer than wide. 2 nd tergite densely punctate, with striated rugae in basal 2 / 3. 3 rd tergite with coarse dense punctures in basal half, with finer rather dense punctures apically. Following tergites with sparse fine setiferous punctures. Apical margin of hypopygium strongly rounded. Colour. Black. Scape dorsally ivory; basal 4 flagellomeres laterally and flagellomeres 7 – 18 dorsally ivory. Ivory are palps, mandible except tooth, clypeus, face except brown median band, frontal orbit, outer orbit reaching up to occiput (fig. 33 B), collar and hind half of upper margin of pronotum, tegula, subtegular ridge, postero-dorsal spot of mesopleuron, scutellum, metapleuron, area dentipara, petiole, apical bands of postpetiole and 2 nd tergite; posterolateral spots and narrow apical margins of 3 rd to 5 th tergites, large median spots of 6 th and 7 th tergites and paramere. Fore and mid legs except brownish distal tarsomeres ivory; hind coxa reddish, with ivory spot dorsally; hind trochanter brown; hind femur reddish, basally and apically infuscate and with dorsal ivory stripe; hind tibia reddish-yellow, basally and apically infuscate; hind tarsus ivory, base of metatarsus and distal tarsomere infuscate. Wings hyaline; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBFFF9FBDAEFAA5FBAC6603.taxon	discussion	Remark. I have studied the holotype of Nesostendontus formosanus from SDEI to confirm the correlation of both sexes. The species is closely related to N. bakeri Cushman but differs mainly by smaller size, less distinctly widened upper margin of pronotum and completely carinated propodeum.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBFFF9FBDAEFAA5FBAC6603.taxon	distribution	Distribution. Known from Taiwan and Myanmar, new record for Lombok.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB9FF9FBDAEFCF9FAF46251.taxon	type_taxon	Type species. Orientodromus thyridialis nov. sp.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB9FF9FBDAEFCF9FAF46251.taxon	diagnosis	Diagnosis. Small, body length 5.2 mm. Head. Flagellum slender, almost bristle-shaped; widest flagellomeres about square. Temple very short, almost absent (fig. 34 B). Clypeus slightly convex, slightly impressed apico-medially; apical margin blunt, slightly bilobed (fig. 34 A). Mandible distinctly narrowed behind base, with two teeth, ventral tooth much smaller and strongly bent inwards (by 80 °), mandible — seen from frontal — almost sickle-shaped. Genal carina reaching mandibular base. Mesosoma. Stout, mesoscutum 1.2 × wider than long. Upper margin of pronotum ± widened and flattened (as in Deniya Cameron). Epomia present. Notaulus not impressed. Mesopleuron coarsely and densely punctate ventrally, almost smooth dorsally; speculum smooth. Epicnemial carina reaching frontal margin of mesopleuron dorsally. Sternaulus not impressed. Juxtacoxal carina absent. Scutellum moderately elevated, almost vertically slanting at apical margin, with complete lateral carina. Propodeum (fig. 34 C) completely carinated; without apophysis; spiracle slit-shaped. Area basalis without median tubercle. Area superomedia hexagonal; anterior transverse carina reaching its middle. Hind coxa with scopa apico-ventrally. Claws simple. Areolet almost pointed frontally. Vein 1 cu-a postfurcal. Metasoma. Oxypygous, stout. Petiole as wide as high. Postpetiole moderately widened, with sparse punctures, apico-medially smooth. Gastrocoelus impressed, ± smooth. Thyridium large, transverse (fig. 34 D). 2 nd to 4 th sternites with median folds. Ovipositor sheath slender, extending behind metasomal apex by length of 7 th tergite. Colour. Black with extensive ivory coloration.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB9FF9FBDAEFCF9FAF46251.taxon	discussion	Remarks. The general habitus of this new genus resembles members of the genus Clypeodromus Tereshkin, but the new species differs distinctly by: claws simple, juxtacoxal carina absent, mandible almost sickle-shaped with strongly bent lower tooth, and genal carina reaching mandibular base. In the key of Heinrich (1934), the new genus runs to Stenichneumonpsis Heinrich and in the key of Townes et al. (1961) to Stenarches Heinrich, but can be differentiated by its very short and almost absent temples, stout habitus, different form of mandible, different carination of propodeum and absence of the juxtacoxal carina.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB8FF9DBDAEF897FD11601B.taxon	description	(Figs 34 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB8FF9DBDAEF897FD11601B.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr Danum Valley Field C, WO, Malaise trap 5, c 150 m, 12. vii. – 2. viii. 1987, C. van Achterberg & D. Kennedy, RMNH´ 87 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB8FF9DBDAEF897FD11601B.taxon	etymology	Etymology. The species name refers to the very wide thyridia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB8FF9DBDAEF897FD11601B.taxon	description	Description. ♀. Body length 5.2 mm. Head. Flagellum with 33 flagellomeres; 1 st flagellomere 4.7 × longer than wide; widest flagellomeres about square. Temple very short, strongly and almost linearly narrowed behind eye (fig. 35 B). OED 1.5 × and OOD 1.4 × ocellar diameter. Frons smooth and shining. Face with rather dense fine punctures laterally, with transverse striae medially (fig. 34 A). Clypeus slightly convex, with sparse punctures; apical margin blunt, slightly bilobed. Mandible with two teeth, ventral tooth much smaller and strongly bent inwards (by 80 °). Malar space 0.7 × as long as width of mandibular base. Gena with coarse sparse punctures. Genal carina reaching mandibular base. Mesosoma. Very stout. Upper margin of pronotum ± widened and flattened. Side of pronotum smooth; epomia present. Notaulus not impressed. Mesoscutum 1.2 × wider than long, with rather dense punctures, shining. Mesopleuron coarsely and densely punctate ventrally, almost smooth dorsally; speculum smooth. Metapleuron with very sparse fine punctures; juxtacoxal carina absent. Scutellum moderately elevated, almost vertically slanting at apical margin, 1.2 × wider than long, transversely rugose, with complete lateral carina (fig. 34 C). Propodeum completely carinated; without apophysis; spiracle slit-shaped (fig. 34 C). Area basalis c 3.0 × wider than long, without median tubercle. Area superomedia hexagonal, slightly wider than long; anterior transverse carina reaching its middle. Hind coxa with dense punctures and large scopa apico-ventrally. Hind femur finely punctate, 3.8 × longer than wide. Claws simple. Areolet almost pointed frontally. Vein 1 cu-a postfurcal by 2.0 × its width. Nervellus of hind wing vertical, intercepted at apical 0.7. Metasoma. Oxypygous, stout. Postpetiole moderately widened, with sparse punctures, apico-medially smooth. 2 nd tergite 1.3 × wider than long. Thyridium large, slightly oblique, 2 × wider than the interval between thyridia (fig. 34 D). 2 nd and 3 rd tergites coarsely rugose-punctures basally, with sparse punctures apically. Following tergites with fine sparse setiferous punctures. 2 nd to 4 th sternites with median folds. Ovipositor sheath slender, extending behind metasomal apex by length of 7 th tergite. Colour. Black. Antenna black; scape, pedicel and basal three flagellomeres reddish ventrally; flagellomeres 7 – 12 ivory dorsally. Ivory are palps, mandible except tooth, clypeus, face, gena, orbits (except interruption on outer orbit), frontal and upper margins of pronotum, subtegular ridge, short oval paramedian spots on mesoscutum, scutellum and postscutellum, wide ventral stripe and posterior margin of mesopleuron, upper division and apical spot on metapleuron, area dentipara of propodeum, petiole, apical bands on 1 st to 4 th tergites, posterolateral spots on 5 th tergite and small apical spot of 7 th tergite. Fore and mid legs including coxae and trochanters ivory, fore and mid femora and tibiae with reddish stripes dorsally; hind coxa red, black apically and with ivory spot dorsally; hind trochanter brown; hind femur red, black basally and apically; hind tibia brown, with ivory subbasal spot; hind tarsus except black distal tarsomere ivory. Wings hyaline; pterostigma brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFB8FF9DBDAEF897FD11601B.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEF92EFC1862F0.taxon	materials_examined	Material. Indonesia: N Sumatra, Aceh, Ketambe, 400 m, 1 ♀ ii. 1995, Mal. trap, nr edge rainforest, leg. Y. van Nierop & C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEF92EFC1862F0.taxon	discussion	Remarks. Body length 9.5 mm. Flagellum with 35 flagellomeres, moderately lanceolate; 1 st flagellomere 3.6 × longer than wide, widest flagellomere 1.4 × wider than long. OED 1.5 × and OOD 1.5 × ocellar diameter. Hind femur 4.0 × longer than wide. Flagellomeres 6 – 13 with ivory ring. Structure and coloration otherwise as described by Heinrich (1975: 493 – 494).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEF92EFC1862F0.taxon	distribution	Distribution. Known from Myanmar, new record for Indonesia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEFAC9FBFA6371.taxon	materials_examined	Material. Indonesia: N Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, 1 ♀ x. 1994, leg. Y. van Nierop & C. van Achterberg (Leiden)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEFAC9FBFA6371.taxon	discussion	Remarks. The available ♀ differs from the original description (Heinrich 1975: 70 – 71) by: flagellum with 35 flagellomeres; 1 st flagellomere reddish, following flagellomeres black, flagellomeres 6 – 11 with ivory rings. The hind tibia has a small roundish smoothened area apico-externally, not a longitudinal pit as in Maraces species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBBFF9DBDAEFAC9FBFA6371.taxon	distribution	Distribution. Known from Laos, Malaysia and Myanmar, new record for Indonesia.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFF08FB56649C.taxon	materials_examined	Material. Malaysia: SW Sabah, nr Long Pa Sia (West), 1 ♀ 25. xi. – 8. xii. 1987, c 1200 m, Mal. trap 7, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFF08FB56649C.taxon	discussion	Remarks. Flagellum with 39 flagellomeres. 5 th tergite with posterolateral ivory spots; hind tarsus mainly ivory.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFF08FB56649C.taxon	distribution	Distribution. Known from India, Peninsula Malaysia and Myanmar, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFE4FFCEE67D4.taxon	materials_examined	Material. Indonesia: C Sulawesi, Lore Lindu N. P., Marena Shelter, 600 m, 1 ♀ 14 – 17. xii. 1985, leg. J. van Tol & J. Krikken (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFE4FFCEE67D4.taxon	discussion	Remarks. Flagellum with 38 flagellomeres; 1 st flagellomere 5.1 × longer than wide, widest flagellomere 1.1 × wider than long. 5 th tergite black, without ivory spots; hind tarsus except ivory metatarsus black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFE4FFCEE67D4.taxon	distribution	Distribution. This subspecies is only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFC8BFD1C62E7.taxon	description	(Figs 35 A – D) Holotypus. (♀) West Sumatra, Maninjau lake, 0 ° 16´13.76 S 100 ° 13´57.96 E, 19. xi. 2015, leg. V. Gromenko (coll. Riedel).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFC8BFD1C62E7.taxon	etymology	Etymology. The species name refers to the red and black metasoma.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFC8BFD1C62E7.taxon	description	Description. ♀. Body length 10.1 mm. Head. Flagellum with 30 flagellomeres, slightly lanceolate; 1 st flagellomere 3.3 × longer than wide; 9 th flagellomere square; widest flagellomere 1.4 × wider than long. Head densely punctate. Temple long, dorsally 0.8 × as long as eye, slightly and roundly narrowed behind eye (fig. 35 B). OED 2.2 × and OOD 1.7 × ocellar diameter. Frons and face rugose-punctate, dull. Clypeus almost flat, sides slightly bulging, moderately blade-like apically, with sparse punctures; apical margin thin, slightly concave (fig. 35 A). Mandible with two teeth, lower tooth small and strongly bent inwards; mandible — seen from frontal — sickle-shaped. Malar space 1.5 × as long as width of mandibular base. Genal carina reaching mandibular base. Mesosoma. Mesoscutum finely and densely punctate and granulate, dull. Side of pronotum rugose-punctate dorsally, coarse rugose ventrally. Mesopleuron and metapleuron densely rugose-punctate; speculum punctate; juxtacoxal carina present. Scutellum distinctly elevated and almost pyramidal, about as long as wide, coarsely punctate, with lateral carina in basal 0.8. Propodeum completely carinated (fig. 35 C), spiracle slit-shaped. Area basalis and area superomedia confluent. Area basalis with median tubercle. Area superomedia hexagonal, 1.8 × longer than wide; costula reaching at its frontal 0.4. Apical transverse carina slightly lamelliform at outer edge of area dentipara, but without apophysis. Hind coxa densely punctate, with distinct scopa in its apico-ventral third. Hind femur 4.2 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 3.0 × their width; vein 1 cu-a slightly antefurcal by 1.0 × its width. Metasoma. Oxypygous; 2 nd to 4 th sternites with median folds. Postpetiole moderately widened, without latero-median carina; median field not separated, finely granulate; lateral filed punctate. 2 nd tergite 1.1 × wider than long. Gastrocoelus strongly impressed, with some ridges. Thyridium transverse, 2.1 × wider than the interval between thyridia. 2 nd to 4 th tergites densely rugose-punctate and dull; 2 nd tergite finely striate between thyridia. Ovipositor sheath slightly reaching behind metasomal apex. Colour. Black. Flagellum black, flagellomeres 7 – 10 with ivory rings. Hind edge of pronotum, collar and apical 2 / 3 of scutellum yellow. Mesopleuron red, black fronto-dorsally. Mesosternum, metapleuron and propodeum red. 1 st to 3 rd tergites red, following tergites black (fig. 35 D). Fore and mid legs red; mid tibia externally and fore and mid tarsi entirely blackish; hind coxa red, diffusely brownish apically; hind trochanter red; hind femur, tibia, and tarsus black. Wings hyaline; pterostigma brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFBAFF9CBDAEFC8BFD1C62E7.taxon	distribution	Distribution. Only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	description	(Figs 36 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	materials_examined	Holotype. (♀) Indonesia: N Sumatra, Ketambe, 400 m, Mal. trap, near edge rainforest, iii. 1995, Y. van Nierop & C. van Achterberg, RMNH´ 99 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	etymology	Etymology. The species name refers to the ivory colour pattern.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	description	Description. ♀. Body length 10.2 mm. Head. Flagellum with 37 flagellomeres, almost filiform; 1 st flagellomere 4.5 × longer than wide, widest flagellomeres about as long as wide, distal flagellomere blunt. Temple rather long, moderately and roundly narrowed behind eye. OED 1.0 × and OOD 1.2 × ocellar diameter. Frons densely punctate dorsally. Face densely punctate, shining. Clypeus almost flat, wide, 3.1 × wider than long, densely punctate; apical margin sharp, slightly bilobed (fig. 36 A). Mandible long and slender, dorsal tooth pointed, ventral tooth very small and blunt. Malar space 0.6 × as long as width of mandibular base. Gena with dense punctures. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus slightly impressed at frontal margin of mesoscutum. Mesoscutum very densely punctate, slightly shining. Mesopleuron finely rugose-punctate, partly with longitudinal rugae; speculum punctate. Metapleuron rugose-punctate; juxtacoxal carina absent. Scutellum slightly elevated, about as long as wide, with dense punctures; without lateral carina. Propodeum coarsely rugose-punctate, with very blunt short apophysis (fig. 36 C); spiracle slit-shaped. Area basalis trapezoid, wider than long, without median tubercle. Area superomedia rectangular, 1.1 × longer than wide; costula absent. Area petiolaris with transverse rugae. Hind coxa densely punctate, without scopa. Hind femur slender, densely punctate, 5.3 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 4.0 × their diameter. Vein 1 cu-a postfurcal by 3.0 × its width. Metasoma. Amblypygous; hypopygium with an apical brush of fine but elongate setae, not reaching metasomal apex (fig. 36 B). 2 nd and 3 rd sternites with median folds. Postpetiole moderately widened, without latero-median carina; median field moderately elevated and separated, finely punctate and weakly striate; lateral field striate. Gastrocoelus triangularly impressed, confluent medially, without multiple longitudinal ridges. Thyridium small, oblique, 0.4 × as wide as the interval between thyridia (fig. 36 D). 2 nd tergite as long as wide. 2 nd and 3 rd tergites densely rugose-punctate, dull; 2 nd tergite with longitudinal ridges basally. Following tergites finely punctate, shining. Ovipositor sheath not reaching behind metasomal apex. Colour. Black. Ivory are rings on flagellomeres 8 – 15, palps, mandible except teeth, clypeus face except ventral brownish spot, narrow stripes of frontal orbit, gena, scape ventrally, collar, apical half of upper margin of pronotum, spot on tegula, subtegular ridge, apical spot of scutellum, postscutellum, upper division of metapleuron, posterolateral spot of propodeum around apophysis, posterolateral spot of postpetiole, wide posterolateral stripes and narrow apical margins of all tergites. Legs reddish-yellow; fore and mid coxae and trochanters and dorsal spot of hind coxa ivory; fore and mid femora with infuscate dorsal stripes; hind femur except reddish base blackish; mid tibia in apical 0.3 and hind tibia basally and in apical 0.3 black; hind tarsus ivory, 4 th and 5 th hind tarsomeres black. Wings hyaline; pterostigma dark brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	discussion	Remarks. The generical position of this new ♀ is somewhat questionable, since the associated ♂ is unknown. The habitus and the morphology of clypeus and gastrocoelus resemble members of the genus Setanta Cameron, but the form of mandible and the brush of setae of apical hypopygium would be more characteristic for Eutanyacra Cameron. I provisionally include this ♀ in the genus Setanta here. In the key of Oriental Setanta species (Heinrich 1974 a: 439 – 440), this species runs to Setanta albitarsis Heinrich, 1974 but differs by black mesoscutum and mesopleuron without ivory coloration and the mainly black hind femur.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA5FF81BDAEF8CAFD1C61FF.taxon	distribution	Distribution. Only known from Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF81BDAEFAA4FC4763AB.taxon	materials_examined	Material. Indonesia: C Sulawesi, Lore-Lindu N. P., nr. Dongi-Dongi, c 1010 m, 1 ° 15´S 120 ° 20´E, Mal. trap 2, 6 ♂♂ 4 – 9. xii. 1985, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF81BDAEFAA4FC4763AB.taxon	discussion	Remarks. Body length 12 – 13 mm. Flagellum with 37 flagellomeres. Tyloids on flagellomeres 11 – 19. Legs mainly black; fore and mid coxae and trochanters ivory; hind coxa black, with ivory dorsal spot; fore femur, tibia and tarsus ivory frontally; mid femur apically and mid tibia basally ± ivory; hind tibia with subbasal ivory spot or ring. The colour of legs is much darker in the specimens from Java and Sulawesi compared to the Indian types of Setanta rufipes (see Heinrich 1974 a: 441 – 443). It is not clear yet, if the Indonesian material might represent a different subspecies or separate species.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF81BDAEFAA4FC4763AB.taxon	distribution	Distribution. Known from India and Java, new record for Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF80BDAEF891FCD366FB.taxon	materials_examined	Material. Indonesia: W Bali, nr. Negara, rainforest above Batuagung, c 575 m, 1 ♂ 4 – 13. xii. 1991, Mal. trap 1, leg. C. van Achterberg, RMNH´ 91 (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF80BDAEF891FCD366FB.taxon	description	Description. ♂. Body length 12.6 mm. Head. Flagellum with 36 flagellomeres, bristle-shaped; 1 st flagellomere 3.2 × longer than wide, all flagellomeres longer than wide. Tyloids absent. Temple short, strongly and roundly narrowed behind eye. OED 1.1 × and OOD 1.2 × ocellar diameter. Frons rugose-punctate. Face and clypeus densely punctate, ± shining. Clypeus almost flat, 2.3 × wider than long; apical margin blunt, slightly concave. Mandible with two teeth, lower tooth small and moderately bent inwards (by 45 °). Malar space 0.7 × as long as width of mandibular base. Gena densely punctate. Genal carina reaching mandibular base. Mesosoma. Mesoscutum very densely punctate, ± dull. Notaulus impressed in frontal 0.3. Mesopleuron and metapleuron densely punctate, partly rugose-punctate; speculum large, smooth. Juxtacoxal carina present. Scutellum moderately elevated, about as long as wide, densely punctate, with complete lateral carina. Propodeum completely carinated, spiracles slit-shaped. Carination see figure. Legs rather slender; hind femur 4.0 × longer than wide. Claws not pectinate. Areolet almost quadrangular, frontal distance between veins 2 rs-m and 3 rs-m 1.0 × their diameter. Vein 1 cu-a slightly antefurcal by 1.0 × its width. Metasoma. Petiole about as wide as high. Postpetiole distinctly widened, without latero-median carina; median field moderately elevated, finely aciculate and with some coarse punctures; lateral field punctate. 2 nd tergite 1.2 × wider than long. Gastrocoelus strongly impressed, with oblique ridges. Thyridium very large, transverse, 1.9 × wider than the interval between thyridia. 2 nd to 4 th tergites densely rugose-punctate. Following tergites with superficial punctuation. 2 nd to 4 th sternites with median folds. Apical margin of hypopygium straight. Colour. Black. Flagellomeres 9 – 18, palps, base of mandible, labrum, clypeus and face, wide orbits (except interruption at malar space), scape ventrally. Ivory are collar, upper margin and ventral edge of pronotum, apical spot of tegula, subtegular ridge, wide band on ventral mesopleuron, scutellum except black median band, postscutellum, upper division of metapleuron, large posterolateral spots of propodeum, hind margins of all tergites, fore and coxae and trochanters, and dorsal spot of hind coxa. Hind coxa and trochanter otherwise black; legs otherwise reddish; hind femur black in apical 0.3; hind tibia black, with ivory subbasal band; mid and hind tarsi black. Wings hyaline; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA7FF80BDAEF891FCD366FB.taxon	distribution	Distribution. Known from Sulawesi, new record for Bali.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	description	(Figs 37 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	materials_examined	Holotype. (♀) Malaysia: SE Sabah, nr. Danum Valley Field C, c 150 m, WONO, Mal. trap 5, 19. iv. – 22. v. 1988, RMNH´ 89, C. van Achterberg & T. Burghouts (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	etymology	Etymology. The species name refers to the completely ivory coxae.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	description	Description. ♀. Body length 11.0 mm. Head. Flagellum with 32 flagellomeres, slightly lanceolate; 1 st flagellomere 3.4 × longer than wide, 9 th flagellomere square, widest flagellomeres 1.3 × wider than long, apical flagellomere blunt. Temple very short, strongly and linearly narrowed behind eye. OED 1.1 × and OOD 0.9 × ocellar diameter. Frons shining, finely granulate and with superficial punctures dorsally. Face rather densely but superficially punctate, shining. Clypeus large, almost flat, with sparse punctures; apical margin blunt and slightly rounded (fig. 37 A). Mandible long and rather slender, with two teeth; ventral tooth much smaller and slightly bent inwards. Malar space 0.8 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus distinctly impressed in frontal third of mesoscutum. Mesoscutum finely and densely punctate, dull. Mesopleuron with dense superficial punctures, with some coarse longitudinal rugae ventrally; speculum superficially punctate (fig. 37 B). Metapleuron punctate and finely striate; juxtacoxal carina present. Scutellum slightly elevated and laterally carinate in basal 0.8, slightly longer than wide, with rather dense punctures. Propodeum completely carinated, with blunt short apophysis (fig. 37 C); spiracle slit-shaped. Area basalis trapezoid, about as wide as long, without median tubercle. Area superomedia horseshoe-shaped, 1.3 × longer than wide; costula reaching at basal 0.35 of its length. Hind coxa with fine dense punctures, without scopa; hind femur densely punctate, 4.8 × longer than wide. Claws simple, strongly bent apically. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 1 – 2 × their diameter. Vein 1 cu-a interstitial. Metasoma. Oxypygous. Petiole about as wide as high. Postpetiole slightly widened, without latero-median carina; median field not separated, finely granulate; lateral field punctate. Gastrocoelus small, much longer than wide, with fine ridges. Thyridium very small, oblique, 0.2 × as wide as the interval between thyridia (fig. 37 D). 2 nd tergite 1.1 × longer than wide, entirely granulate and with fine dense punctures in basal half, dull. Following tergites finely granulate, slightly shining. 2 nd to 5 th sternites with median folds. Ovipositor sheath narrow, reaching widely behind metasomal apex by length of 7 th tergite. Colour. Black. Flagellum black, flagellomeres 1 – 4 reddish-brown, flagellomeres 5 – 15 ivory dorsally. Palps, all coxae and trochanters and 6 th and 7 th tergites mainly ivory. Spot of mandible, lateral margin of clypeus, scape ventrally and two small apical spots of scutellum reddish. Fore and mid femora and fore tibia and tarsus yellow; mid tibia yellow, apically blackish; mid and hind tarsi black; hind femur black, narrowly red at base; hind tibia blackish, diffusely reddish-brown in basal half. 2 nd and 3 rd tergites with narrow reddish apical margins. Wings slightly infuscate; pterostigma reddish-brown. ♂ unknown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	discussion	Remarks. This new species differs from the known mainly black Oriental species of Stirexephanes by the completely ivory coxae and trochanters.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA6FF86BDAEFBA1FD11640B.taxon	distribution	Distribution. Only known from Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF85BDAEFF08FCF761D0.taxon	description	(Figs 38 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF85BDAEFF08FCF761D0.taxon	materials_examined	Material. Malaysia: SW Sabah, nr Long Pa Sia, Payaka-laba, 1 ♀ 25. xi. – 1. xii. 1987, c 1010 m, Mal. trap 14, leg. C. van Achterberg; Sabah, Kinabalu Park Head Q., 1 ♂ 8 – 11. iii. 1987, c 1525 m, Mal. trap 2, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF85BDAEFF08FCF761D0.taxon	description	Description. ♀. Body length 10.5 mm. Flagellum with 29 flagellomeres, almost filiform; 1 st flagellomere 3.6 × longer than wide. OED 0.9 × and OOD 0.9 × ocellar diameter. Hind coxa ventrally with distinct scopa. Hind femur 4.5 × longer than wide. 2 nd tergite 1.0 × as long as wide; thyridium transverse, 0.3 × as wide as the interval between thyridia. Colour. black. Flagellomeres 6 – 12 and 7 th tergite ivory. Wings slightly infuscate; pterostigma black. The available ♀ differs from the material from Sulawesi by its completely black 6 th tergite. It is otherwise typical. ♂. Body length 15 mm. Head (figs 38 A, 38 B). Flagellum with 41 flagellomeres, bristle-shaped; 1 st flagellomere 4.1 × longer than wide. Tyloids on flagellomeres 10 – 15, linear, maximally 0.6 × as long as their flagellomeres. Temple short, strongly and linearly narrowed behind eye (fig. 38 B). OED 1.3 × and OOD 0.8 × ocellar diameter. Mesosoma. Mesoscutum with fine dense punctures, dull. Mesopleuron with superficial punctures, ventrally with superficial striation. Metapleuron with longitudinal striation. Scutellum moderately elevated and strongly declining apically, slightly longer than wide, with complete lateral carina. Propodeum completely carinated, with strong apophysis (fig. 38 D). Area superomedia hexagonal, about as long as wide; costula reaching its middle. Legs slender; hind femur 5.7 × longer than wide. Metasoma. Slender. Postpetiole slightly widened, almost smooth; median field not separated. 2 nd tergite 1.4 × longer than wide. Gastrocoelus slightly impressed. Thyridium small, oblique, 0.4 × as wide as interval between thyridia (fig. 38 C). Apical margin of hypopygium slightly rounded. Colour. Black. Ivory are palps, clypeus, face, stripes of gena and of frontal orbit, small ventral spot of scape, hind edge of pronotum, subtegular ridge, scutellum apically, postscutellum, posterolateral stripe of propodeum posterior to apophysis, apical spot on postpetiole, narrow apical margins of 2 nd to 4 th tergites, large median spot of 7 th tergite, parameres, fore and mid coxae and trochanters frontally, frontal stripes of fore femur and tibia and fore and mid tarsi entirely, and hind tarsus except black basal 2 / 3 of metatarsus. Wings moderately infuscate; pterostigma black.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF85BDAEFF08FCF761D0.taxon	distribution	Distribution. Known from Sulawesi, new record for Borneo.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	description	(Figs 39 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	materials_examined	Holotype. (♀) Indonesia: Sumatera Barat, Joron Akar Gadang, Simpang Tanjung Nam IV, Danau Kembar, Solok, 1.0579 ° S 100.697 ° E, 1412 m, 13 - May- 2016, H. Hamid, COI 0, INDOBIOSYS-CCDB 23977 - G 04 (ZSM). Paratype. (♂) Malaysia: SW Sabah, nr Long Pa Sia (West), c. 1010 m, 25. xi. – 8. xii. 1987, Mal. trap 1 b, C. van Achterberg (Leiden); Indonesia: Sumatera Barat, Lahan basah, Andalas University, Bukit, Karamuntiang, Padang, 0.9087 ° S 100.478 ° E, 400 m, 23 – Jan – 2016, H. Hamid, INDOBIOSYS-CCDB 23977 - A 05 (ZSM).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	etymology	Etymology. The species name refers to the reddish 1 st tergite.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	description	Description. ♀. Body length 8.1 mm. Head. Flagellum with 30 flagellomeres, slightly lanceolate; 1 st flagellomere 3.2 × longer than wide, 9 th flagellomere square, widest flagellomeres 1.4 × wider than long, apical flagellomere very blunt. Temple very short, strongly and linearly narrowed behind eye. OED 1.0 × and OOD 0.6 × ocellar diameter. Vertex finely granulated. Frons shining, finely granulate and with superficial punctures dorsally. Face densely punctate, with some transverse rugae medially. Clypeus large, almost flat, with sparse punctures; apical margin blunt and almost straight (fig. 39 A). Mandible long and rather slender, with two teeth; ventral tooth much smaller and slightly bent inwards. Malar space 1.0 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina away from mandibular base. Mesosoma. Notaulus distinctly impressed in frontal fourth of mesoscutum. Mesoscutum finely and densely rugose-punctate, dull. Sternaulus distinctly impressed in frontal half of mesopleuron. Mesopleuron finely rugose-punctate and ± striate; speculum partly smooth (fig. 39 B). Metapleuron finely rugose and striate; juxtacoxal carina present. Scutellum slightly elevated, with lateral carina in basal 0.8, slightly longer than wide, with sparse punctures. Propodeum completely carinated, with blunt short apophysis (fig. 39 C); spiracle slit-shaped. Area basalis trapezoid, wider than long, without median tubercle. Area superomedia hexagonal, 1.3 × longer than wide; costula reaching at basal 0.35 of its length. Hind coxa with fine dense punctures, without scopa; hind femur densely punctate, 4.3 × longer than wide. Claws simple, strongly bent apically. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their diameter. Vein 1 cu-a interstitial. Metasoma (fig. 39 D). Oxypygous. Petiole about as wide as high. Postpetiole slightly widened, without latero-median carina; median field not separated, finely granulate; lateral field finely punctate. Gastrocoelus small, longer than wide, with fine ridges. Thyridium small, comma-shaped, 0.3 × as wide as the interval between thyridia. 2 nd tergite 1.1 × longer than wide. 2 nd and 3 rd tergites finely and densely rugose-punctate, dull. Following tergites finely granulate, slightly shining. 2 nd to 5 th sternites with median folds. Ovipositor sheath narrow, reaching behind metasomal apex by length of 7 th tergite. Colour. Black. Flagellum black, flagellomeres 6 – 13 ivory dorsally. Palps ivory. Median spot of mandible, lateral margin of clypeus and scape ventrally reddish. Mesosoma black; scutellum, postscutellum, posterior half of mesopleuron, metapleuron and propodeum red. 1 st tergite red; following tergites black. 2 nd tergite diffusely reddish in posterior half and with narrow yellow apical margin. Apical band of 6 th tergite and 7 th tergite mainly ivory (fig. 40 D). Fore and mid coxae and all trochanters ivory; hind coxa black ventrally, red dorsally. Fore and mid femora and tibiae and fore tarsus reddish; mid tarsus brown. Hind femur, tibia and tarsus black; hind tibia with small yellowish spot basally. Wings almost hyaline; pterostigma ochreous. ♂. Body length 10.8 – 11.6 mm. Flagellum with 34 – 35 flagellomeres, bristle-shaped; 1 st flagellomere 3.2 – 3.3 × longer than wide, distal flagellomeres slightly nodose. Tyloids on flagellomeres 6 – 13, linear, maximally 0.8 × as long as their flagellomeres. OED 1.1 × and OOD 0.9 × ocellar diameter. Malar space 0.6 × as long as width of mandibular base. Notaulus impressed in frontal 0.2 of mesoscutum. Mesoscutum with fine dense and ± superficial punctures, dull. Mesopleuron and metapleuron densely and superficially punctate, not striate. Hind femur 4.2 – 4.6 × longer than wide. 2 nd tergite 1.1 × longer than wide. Hypopygium unmodified, apical margin slightly rounded. Structure otherwise as described for the ♀. Colour. Black. Flagellum black, basal flagellomeres ± reddish, flagellomeres 11 / 12 – 18 / 19 ivory dorsally. Palps, mandible except teeth, clypeus, face, gena, frontal orbit, sometimes outer orbit and scape ventrally ivory. Mesosoma black; propleuron, and sometimes collar and spot of tegula ivory. Scutellum ±, postscutellum, tegula, sometimes posterior half of mesopleuron, metapleuron and propodeum red. 1 st tergite red; 2 nd tergite black, reddish or yellowish in apical 0.4 – 0.5. Following tergites black; sometimes 3 rd and 4 th tergites with narrow yellowish apical bands; apical band of 6 th tergite, 7 th tergite and paramere ivory. Fore coxa and fore and mid trochanters ivory; mid coxa reddish-yellow, with apical ivory spot; hind coxa red, diffusely infuscate ventrally. Fore and mid femora and tibiae and fore tarsus reddish or ochreous; mid tarsus brown; hind leg black; hind tibia with external ivory stripe, sometimes with basal ivory spot. Pterostigma ochreous.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	discussion	Remarks. This new species differs from all other known Stirexephanes species by the bicolored mesosoma and red 1 st tergite.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFA3FF8BBDAEFA88FD6C601B.taxon	distribution	Distribution. Known from Borneo and Sumatra.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFADFF89BDAEFACCFD24672B.taxon	description	(Figs 40 A – D)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFADFF89BDAEFACCFD24672B.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Mosquito Camp, Gn. Watuwila, Malaise trap, c 1200 m, 9 ♂♂ 13 – 17. x. 1989, leg. J. Huisman & R. de Jong; Sulawesi, nr Bantaeng, Borong Rappoa, Gn. Daulu, Mal. trap 16, c 700 m, 1 ♂ 4 – 24. iv. 1991, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFADFF89BDAEFACCFD24672B.taxon	description	Description. ♂. Body length 13 mm. Head. Flagellum with 35 – 36 flagellomeres, bristle-shaped; 1 st flagellomere 2.8 × longer than wide. Tyloids on flagellomeres 10 – 17 / 18, long-oval, maximally 0.7 × as long as their flagellomeres. Temple parallel behind eye and moderately narrowed apically (fig. 40 B). OED 1.4 × and OOD 0.9 × ocellar diameter. Face and clypeus punctate, head otherwise almost smooth and shining (fig. 40 A). Mesosoma. Mesoscutum with rather dense punctures, punctuation becoming very sparse in posterior third. Scutellum moderately elevated, wide than long, with lateral carina in basal half. Propodeum completely carinated, without apophysis (fig. 40 C). Area superomedia heart-shaped, about as long as wide; costula reaching its middle. Hind femur 3.5 × longer than wide. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2.0 × their diameter. Vein 1 cu-a interstitial. Metasoma. Stout. Postpetiole strongly widened and coarsely punctate. 2 nd tergite 1.1 × wider than long. Gastrocoelus strongly impressed. Thyridium large, transverse, 2.1 – 2.4 × wider than the interval between thyridia (fig. 40 D). Apical margin of hypopygium distinctly elongate and pointed medially. Colour. Black. Flagellum reddish-yellow, infuscate in distal 1 / 2 – 2 / 3. Ivory are palps, mandible except teeth, clypeus, face, gena, wide orbits (with narrow interception at lateral ocellus), ventral spots of scape and pedicel, spot of propleuron, frontal and upper margins of pronotum, tegula, subtegular ridge, large stripe of ventral mesopleuron, rather wide paramedian stripes on mesoscutum, prescutellar ridge, scutellum (sometimes bis black median stripe), postscutellum, upper division and apical spot of metapleuron, posterolateral stripe on propodeum, small posterolateral spot on postpetiole. Propodeum sometimes ± reddish-brown basally. Metasoma red. Legs red; fore and mid coxae and trochanters, and dorsal spot of hind coxa ivory. Fore and mid femora and tibiae ± yellowish frontally; hind tarsus brown. Wings hyaline; pterostigma brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFADFF89BDAEFACCFD24672B.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFAFFF89BDAEFD11FD2462AF.taxon	description	(Figs 41 A – C)	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFAFFF89BDAEFD11FD2462AF.taxon	materials_examined	Material. Indonesia: SE Sulawesi, nr Sanggona, Base camp, Gn. Watuwila, Mal. trap 1, c 225 m, 1 ♂ 15. x. – 5. xi. 1989, leg. C. van Achterberg; N Sulawesi, Dumoga-Bone N. P., c 220 m, Mal. trap: Maze Toraut R., 0 ° 34´N 123 ° 54´E, 1 ♂ 1 – 9. xi. 1985, leg. C. van Achterberg (Leiden).	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFAFFF89BDAEFD11FD2462AF.taxon	description	Description. ♂. Body length 9 – 10 mm. Head. Flagellum with 32 – 34 flagellomeres, bristle-shaped; 1 st flagellomere 1.9 × longer than wide, distal flagellomeres ± nodose. Tyloids on flagellomeres 9 – 14, long-oval, maximally 0.6 × as long as their flagellomeres. Temple short, strongly and roundly narrowed behind eye. OED 1.2 × and OOD 1.2 × ocellar diameter. Face wide and short, densely punctate (fig. 41 A). Clypeus slightly convex, densely punctate; apical margin blunt, slightly rounded. Mandible with two teeth, ventral tooth much smaller and slightly bent inwards. Malar space 0.5 × as long as width of mandibular base. Genal carina reaching hypostomal carina close to mandibular base. Mesosoma. Notaulus distinct in frontal 0.2 of mesoscutum. Mesoscutum densely punctate, shining. Mesopleuron and metapleuron densely punctate, speculum smooth; juxtacoxal carina present. Scutellum moderately elevated, slightly wider than long, with rather dense punctures, without lateral carina. Propodeum completely carinated, without apophysis (fig. 41 B); spiracle slit-shaped. Area basalis 3.0 × wider than long, moderately bulging forward medially. Area superomedia hexagonal, c 1.5 – 2.0 × wider than long; costula reaching at 6 / 10 of its length. Hind femur 4.0 × longer than wide. Areolet almost quadrangular, frontal distance of veins 2 rs-m and 3 rs-m 1.0 × their width. Vein 1 cu-a postfurcal by 2.0 × its width. Metasoma. Postpetiole moderately widened, with strong latero-median carina; median field moderately elevated, with coarse and dense punctuation (fig. 41 C). 2 nd tergite 1.1 × wider than long. Gastrocoelus distinctly impressed, with lateral ridge. Thyridium slightly oblique, 1.1 × wider than the interval between thyridia. 2 nd to 4 th tergites coarsely rugose-punctate; 2 nd and 3 rd tergite ± striate medially. Colour. Black. Flagellum black, reddish ventrally. Palps ivory. Yellow are mandible except teeth, clypeus, face, gena, stripes on outer, frontal and vertical orbits, scape ventrally, propleuron ventrally, collar and upper margins of pronotum, tegula, subtegular ridge, short paramedian stripes on mesoscutum, wide band at ventral mesopleuron, scutellum except reddish central stripe, area dentipara, and posterolateral spot on postpetiole. 1 st to 5 th tergites red, sometimes 5 th tergite partly brownish; 6 th and 7 th tergites blackish, sometimes with narrow yellowish apical margins, paramere brown. Legs red; fore and mid coxae and trochanters and dorsal spot of hind coxa ivory; hind femur sometimes narrowly black apically. Wings hyaline; pterostigma ochreous or brown.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFAFFF89BDAEFD11FD2462AF.taxon	discussion	Remarks. The species was described as a member of Intermedichneumon Heinrich by Heinrich (1934) and catalogued in that genus by Gupta (1988) but transferred to Vulgichneumon Heinrich by Heinrich (1967). It differs from typical members of Intermedichneumon by the coarse punctuation and distinct latero-median carina of the postpetiole, and from Vulgichneumon ♂♂ by the large and subbasally located thyridia. Therefore, this species is transferred to Virgichneumon Heinrich here, a genus which is new for the Oriental region.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
038C87C4FFAFFF89BDAEFD11FD2462AF.taxon	distribution	Distribution. Only known from Sulawesi.	en	Riedel, Matthias (2023): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia. Zootaxa 5363 (1): 1-94, DOI: 10.11646/zootaxa.5363.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218
