taxonID	type	description	language	source
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	description	(Figs 1 – 15, 24, 27)	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	materials_examined	Material examined Holotype female: SW Slovakia, Podunajská Rovina Flatland, Svätý Jur Town, Panónsky Háj Forest, oak forest (Quercus spp.), under the bark of an old oak, in galleries with larvae of Cerambyx cerdo Linnaeus (Coleoptera: Cerambycidae), elevation 130 m, 21 March 2023. Paratypes: one female, with the same data as the holotype; one female, SW Slovakia, Borská Nížina Lowland, Šaštín-Stráže Town, oak forest with admixed pines (Pinus sylvestris), in dying oak and galleries with larvae of C. cerdo, elevation 170 m, 5 June 2021; 25 females, SW Slovakia, Podunajská Rovina Flatland, Bratislava Capital, Rusovce Settlement, park, under the bark of an old oak, in galleries with larvae of C. cerdo, elevation 130 m, 26 June 2023. All type specimens were collected by the author, and deposited in the Institute of Zoology of the Slovak Academy of Sciences, Bratislava, Slovakia.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	description	Description (Female)	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	diagnosis	Diagnosis. Podonotal and opisthonotal regions of dorsal shield with 21 and 15 pairs of setae, respectively. Dorsolateral soft cuticle with eight pairs of marginal setae and four pairs of submarginal setae. Setae z 1, s 2, and J 5 shortest and needle-like, other setae of dorsal shield slightly thickened, finely tricarinate, and about as long as distance to each succeeding seta. Presternal area with a pair of platelets widely connected to anterior margin of sternal shield. Epigynal shield relatively large, longer than sternal or ventrianal shield. Ventrianal shield with four pairs of pre-anal setae (JV 1 – JV 3 and ZV 2). Opisthogaster with complete series of pre-anal setae (JV 1 – JV 5 and ZV 1 – ZV 5); JV 5, ZV 4, and ZV 5 slightly thickened and tricarinate, other ventral setae short and needle-like. One pair of metasternal platelets and two pairs of metapodal platelets present. Spermathecal apparatus formed as pear-shaped structure. Epistome triramous. Fixed and movable digits of chelicerae with 8 – 9 and three teeth, respectively. Legs I and IV longer than dorsal shield; tarsi II and III with two flattened and curved macrosetae (ad 2 and pl 2 on tarsus II, ad 2 and pd 2 on tarsus III), tarsus IV with one such macroseta (ad 2). Dorsal idiosoma (Figs 1, 7). Idiosoma 670 ‒ 695 long and 410 ‒ 455 wide at level of r 5 (three specimens measured), almost regularly oval, not completely covered by shield at lateral margins. Dorsal shield elongate and gradually narrowing behind humeral setae (r 3), 665 ‒ 690 long and 390 ‒ 405 wide at widest point at level of r 3, with 36 pairs of setae and conspicuous reticulate pattern on entire surface. Dorsal shield setae (Fig. 14) relatively well developed and thick, almost spine-like, usually slightly inflated basally, and tricarinate (setal carinae smooth, without serration or denticulation); most dorsal setae similarly long, reaching base of following setae in longitudinal row or slightly beyond; only z 1, s 2, and J 5 thin and distinctly shorter, z 1 and s 2 smooth or only imperceptibly tricarinate, J 5 finely serrate; r 3 longest. Podonotum of dorsal shield with 21 pairs of setae (j 1 – j 6, z 1 – z 6, s 1 – s 6, and r 2 – r 4), opisthonotum with 15 pairs of setae (J 1 – J 5, Z 1 – Z 5, and S 1 – S 5); marginal setae r 5, r 6, R 1 – R 6 and four (often asymmetrically three) pairs of submarginal setae (UR) placed on soft cuticle adjacent to shield margin, each with base on small suboval and well-sclerotised platelet. The lengths of some selected dorsal setae are as follows: j 1 54 – 59, j 2 59 – 65, j 3 66 – 72, j 4 63 – 71, j 5 64 – 68, j 6 66 – 73, z 1 30 – 37, s 2 30 – 33, r 2 51 – 55, r 3 80 – 87, r 4 63 – 67, r 5 58 – 65, r 6 49 – 57, other podonotal setae 64 – 72, J 1 65 – 72, J 2 61 – 68, J 3 59 – 64, J 4 57 – 62, J 5 18 – 21, Z 1 65 – 74, Z 2 60 – 66, Z 3 58 – 67, Z 4 59 – 66, Z 5 70 – 80, S 1 61 – 66, S 2 60 – 66, S 3 61 – 69, S 4 61 – 65, S 5 53 – 59, R 1 ‒ R 6 40 – 50, and UR - setae 30 – 36. Ventral idiosoma (Figs 2, 12, 13). Tritosternum with short columnar base and two laciniae; laciniae separated along their entire length, each with sparse, fine and short pilosity. Presternal area with a pair of relatively small and partially to completely fused platelets widely connected to anterior margin of sternal shield; each platelet with usually three fine transverse lines; area of fusion of the two platelets weakly sclerotised. Sternal shield subquadrate, 160 ‒ 180 long and 130 ‒ 145 wide at narrowest point between coxae II, with smooth surface, antrolateral angles well developed and laterally directed, posterior margin almost straight, three pairs of subequal and apically attenuated setae (st 1 ‒ st 3), and two pairs of poroids (iv 1 and iv 2). Each metasternal seta (st 4) and associated poroid (iv 3) on a small and suboval or drop-like metasternal platelet posterior to posterolateral margins of sternal shield. Endopodal platelets reduced and not well-sclerotised, indistinct. Epigynal shield relatively large, elongate, 255 ‒ 275 long, 108 ‒ 115 wide at widest point posterior to setae st 5, 92 ‒ 97 wide at level of st 5, and 126 ‒ 138 wide at widest anterior part, with narrowly convex anterior margin reaching level between setae st 2 and poroids iv 2, moderate medial constriction, and nearly straight posterior margin; surface largely smooth, except for a pattern of fine lines in anterior part and two diagonal lines converging anteriorly in posterior part; setae st 5 on shield surface near posterolateral margins; genital poroids (iv 5) on soft cuticle posterior to st 5. Four postgenital sclerites present, weakly sclerotised. Exopodal platelets fused into long and narrow plates, each parallel to peritrematal shield; parapodal part of exopodal platelets IV narrowly connected to short poststigmatic part of peritrematal shields. Peritremes relatively broad, with anterior ends reaching about level of paravertical setae (z 1); peritrematal shields narrow, developed along outer side of peritremes, and fused with podonotal part of dorsal shield at level of setae s 1. Two pairs of suboval metapodal platelets posterior to coxae IV, anteromedial pair conspicuously smaller than posterolateral pair (23 – 28 long). Ventrianal shield relatively small, inversely pear-shaped, slightly wider than long (192 ‒ 212 long and 209 ‒ 222 wide at widest anterior part), abruptly narrowed behind setae JV 3, almost straight anteriorly and convex anterolaterally and posteriorly; with four pairs of pre-anal setae (JV 1 ‒ JV 3 and ZV 2) in addition to three circum-anal setae and gland pores gv 3 near posterolateral margins at level between setae ad and pa; postanal seta longer than adanal setae (pa 44 ‒ 47, ad 25 – 31), JV 2 distinctly longer than other pre-anal setae; anus small (25 ‒ 30 x 21 ‒ 24 in size), placed in posterior part of shield; pre-anal surface with reticulate pattern dominated by transverse lines, remaining surface without conspicuous ornamentation and smooth, except for narrow cribrum along posterior margin. Soft opisthogastric cuticle with six pairs of pre-anal setae (JV 4, JV 5, ZV 1, and ZV 3 ‒ ZV 5). All ventral setae smooth and needle-like, except for the more robust tricarinate setae JV 5, ZV 4, and ZV 5, with the following lengths: st 1 and st 2 45 – 50, st 3 46 – 54, st 4 44 – 52, st 5 38 – 44, JV 1 40 – 45, JV 2 46 – 58, JV 3 34 – 39, JV 4 24 – 28, JV 5 50 – 57, ZV 1 25 – 30, ZV 2 36 – 39, ZV 3 29 – 37, ZV 4 33 – 37, and ZV 5 41 – 45. Sperm induction structures. Spermathecal apparatus (Fig. 15) well developed, unusually large (total length 75 ‒ 80), and pear-shaped, with a well-sclerotised neck, an almost regularly spherical vesicle or saccule (33 ‒ 36 in diameter), and with a long, thin, and convoluted duct leading from an indistinct atrium to a slightly thickened terminus. Gnathosomal structures (Figs 6, 8 ‒ 11). Corniculi horn-like, normal in size and spacing; internal malae each formed as a pointed projection with imperceptibly serrated lateral margins, projecting far beyond corniculi; hypognathal groove with seven rows of denticles, rows similar in width, with many denticles and connected laterally by longitudinal lines; subcapitular setae (Fig. 8) smooth and needle-like, h 2 shortest, the others similar in length, but pc slightly longer than h 1 and h 3 (h 1 56 – 59, h 2 21 – 23, h 3 55 – 58, pc 58 – 62). Anterior margin of epistome triramous, rami with denticulate distal or apical margins; lateral rami about half as long as median ramus but much wider than median ramus (Fig. 9). Middle article of chelicerae unusually long (240 ‒ 250; cheliceral digits only about 1 / 4 length of middle article); cheliceral digits similar in length and size, dentate; movable digit 63 ‒ 68 long, tridentate (all teeth in distal part of digit); fixed digit with eight or nine teeth, including two small subterminal teeth near terminal hook, 4 ‒ 5 medial teeth slightly larger than a pair of most proximal teeth; pilus dentilis short and small, spine-like; dorsal seta relatively long and thin (Fig. 6). Palp trochanter with two long setae, both with thickened base and attenuated distal part; palptarsal claw two-tined. Legs. Each with well-developed pretarsus and ambulacral apparatus, including pulvillus with three rounded lobules and two claws (claws of tarsi I distinctly smaller than those of tarsi II ‒ IV); legs I and IV longer and legs II and III shorter than idiosoma: legs I 690 – 710, legs II and III 575 – 590, and legs IV 740 – 815 long. Leg setae smooth and acicular, mostly similar in shape and length, except modified setae formed as elongate, flattened, and distally curved macrosetae (ad 2 on tarsi II ‒ IV, pl 2 on tarsi II, and pd 2 on tarsi III). Chaetotactic formulae for each leg segment as follows: leg I – coxa (2), trochanter (6), femur 2 - 3 / 1, 2 / 2 - 2 (12), genu 2 - 3 / 2, 3 / 1 - 2 (13), tibia 2 - 3 / 2, 3 / 1 - 2 (13); leg II – coxa (2), trochanter (5), femur 2 - 3 / 1, 2 / 2 - 1 (11), genu 2 - 3 / 1, 2 / 1 - 2 (11), tibia 2 - 2 / 1, 2 / 1 - 2 (10); leg III – coxa (2), trochanter (5), femur 1 - 2 / 1, 1 / 0 - 1 (6), genu 2 - 2 / 1, 2 / 1 - 1 (9), tibia 2 - 1 / 1, 2 / 1 - 1 (8); leg IV – coxa (1), trochanter (5), femur 1 - 2 / 1, 1 / 0 - 1 (6), genu 2 - 2 / 1, 3 / 0 - 1 (9), tibia 2 - 1 / 1, 3 / 1 - 2 (10).	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	etymology	Etymology The specific name is derived from the generic name “ Cheiroseius ” and the Greek suffix “ - oides ” (resembling), referring to the fact that this new species resembles some species of the genus Cheiroseius in general appearance (well-sculptured dorsal shield, elongated legs) and in some specific characters (e. g., presence of specialised macrosetae on the tarsi II ‒ IV, shape of the epistome).	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D661FFCCFF0CC68FFE6AFDE0.taxon	discussion	Taxonomic notes Lasioseius cheiroseioides is easily recognised, especially by the presence of specifically formed macrosetae (ad 2 on tarsi II ‒ IV, pl 2 on tarsi II, and pd 2 on tarsi III), the conspicuously elongate middle article of the chelicerae (the cheliceral digits are about 1 / 4 as long as the middle article), the elongate legs (of which the first and fourth pairs are longer than the idiosoma), and the three pairs of marginal setae on the dorsal shield (r 2 ‒ r 4). Of the more than 200 valid species of the genus Lasioseius, only a few species have been described that have developed modified macrosetae on the tarsi II ‒ IV, but most often it is only an unpaired macroseta on tarsus IV. The homology of the specialised macrosetae in individual species of Lasioseius was studied and reviewed by Lindquist & Moraza (2016). These authors pointed out a certain general pattern in the chaetotaxy of the elongate and usually more or less strap-like tarsal macrosetae: tarsus II with two modified macrosetae (ad 2 and pl 2), tarsus III with one macroseta (ad 2), tarsus IV with 0 ‒ 1 macroseta (either al 2 or pl 2, rarely pd 2). However, the pattern described above varies in several respects, and only some of the macrosetae listed may be present. Lasioseius cheiroseioides is unusual in the presence of two macrosetae, namely pd 2 on the tarsus III and ad 2 on the tarsus IV. Also, the flattened (strap-like) and distally whip-like form of the macrosetae in the new species is rarely expressed in members of the genus [L. epicrioides (Krantz, 1962), L. americanellus (De Leon, 1964), L. elegans Fain, Hyland & Aitken, 1977, and L. chelaserratus Naeem, Dobkin & OConnor, 1985]. Lasioseius cheiroseioides is most similar to L. ningxiaensis Bai, Ma & Yan, 2014 [in Bai et al. (2014); = L. multisetus Ma & Bai, 2006], a species found in China in the same habitat as the new species (galleries of cerambycid larvae). Both species can be easily distinguished from each other, especially by the placement of the anteromarginal setae r 5 and r 6, which may be located on the dorsal shield in L. ningxiaensis or on the soft cuticle in L. cheiroseioides. Some important morphological structures of L. ningxiaensis are not described in sufficient detail, but undoubtedly it can be considered as the closest relative of L. cheiroseioides and together with L. lacunosus Westerboer, 1963, it could be placed in a complex of related species that share atypical or little specific characters for the genus, at least like the chaetotaxy of the tarsi II ‒ IV and the form and dentition of the chelicerae. Using my own available specimens of L. lacunosus from Slovakia, it is possible to shed more light on the individual similarities between members of this species complex. Lasioseius lacunosus has homologous macrosetae on the tarsi II ‒ IV like the new species, but they are relatively shorter, which might be related to the relatively shorter legs compared to the body. The middle article of the chelicerae is analogous in the conspicuously elongated form, the dentition of both digits, and some other structures: the movable digit is tridentate, with teeth on the distal part of the masticatory surface, and the fixed digit is oligodentate, with a group of 4 ‒ 5 small teeth on the medial masticatory surface; the pilus dentilis is short and stout, but the dorsal seta is long and thin. In addition, the distal part of the middle article of the chelicerae has a conspicuous, peculiar, and longitudinally oriented slit-like furrow found in both species (Fig. 20). The epistome is very similar in both species, triramous, with apical denticulation or strong serration, and the medial ramus is widened apically and narrower than the lateral ones. The species have the full number of j / J, z / Z, s / S, and r / R rows of setae, with different arrangement of anteromarginal setae on / off the dorsal shield: the shield with r 2 ‒ r 4 in L. cheiroseioides and r 2 ‒ r 5 in L. lacunosus (r 2 ‒ r 6 in L. ningxiaensis). The same number of four pairs of submarginal setae (UR) are located on the soft cuticle adjacent to the shield margin (UR setae are not shown or annotated in L. ningxiaensis). The most important difference between L. cheiroseioides and L. lacunosus is the number of pre-anal setae on the ventrianal shield in addition to the three circum-anal setae, four pairs in L. cheiroseioides (JV 1 ‒ JV 3 and ZV 2) and six pairs in L. lacunosus (JV 1 ‒ JV 4, ZV 2, and ZV 3). Another feature of significant differential value that would also support the validity of this group as a separate taxon within the genus (in a new or revised taxonomic concept of the genus) is the unusual presence of slightly enlarged and spurred legs II in males (Figs 21, 22, 30). In males, the anteroventral seta (av) on the femur II is modified into a relatively conspicuous and well-sclerotised spur (Fig. 21), as is av 2 on the tarsus II (Fig. 22). A similar spur-like av 2 is described only in males of the two unrelated species L. furcisetus Athias-Henriot, 1959 and L. corticeus Lindquist, 1971. Male legs II are well spurred only in L. kshamae Bhattacharyya, 2003, but without modified seta av 2 on the tarsus.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D669FFCCFF0CC633FD22F85D.taxon	description	(Figs 16 ‒ 23, 25, 30, 31)	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D669FFCCFF0CC633FD22F85D.taxon	materials_examined	Material examined One female, three males: SW Slovakia, Little Carpathians Mts., Bratislava Capital, Záhorská Bystrica Settlement, Kačín Forest, elevation 335 m, decidouous forest, in bark crevices of a maple (Acer sp.) soaked with xylem sap, 27 June 2019 (leg. author). All specimens were deposited in the Institute of Zoology of the Slovak Academy of Sciences, Bratislava, Slovakia.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D669FFCCFF0CC633FD22F85D.taxon	discussion	Taxonomic notes This is the first record of L. lacunosus for Slovakia. Some specific morphological structures of a female from Slovakia, which are important for verifying its identity, can be seen on the micrographs (Figs 16, 17, 20, 23, 25, 31). I also examined a fairly well preserved female, clearly conspecific with the female from Slovakia, on a slide deposited in the Bavarian State Collection of Zoology in Munich and labeled ʻ Lasioseius lacunosus n. sp. ʼ (No. W 34 / 14). Unfortunately, this available specimen (slide) does not bear a type designation or specific collection data, but according to Gwiazdowicz et al. (2008) it belongs to the original series of Westerboer. As also noted in the taxonomic notes on the new species above, L. lacunosus has specially modified macrosetae on the tarsi II ‒ IV, and this fact was not illustrated or mentioned in the original description by Westerboer (1963). Similar to the legs, the males have not been described before, so I include the following description of some of the most important morphological structures:	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D669FFCCFF0CC633FD22F85D.taxon	description	Description (Male) ‒ (Figs 18, 19, 21, 22, 30) Dorsal shield 500 ‒ 555 long, shorter than legs IV (525 ‒ 575), with coarse reticulate pattern and setae similar to those of female; all anteromarginal setae (r 2 ‒ r 6) on dorsal shield, but all posteromarginal setae (R 1 ‒ R 6) on soft cuticle adjacent to shield margin; UR setae of submarginal cuticle completely reduced, absent (four pairs in female); opisthogastric region normally with six pairs of pre-anal setae on ventrianal shield (JV 1 ‒ JV 3, JV 5, ZV 1, ZV 2, and asymmetrically sometimes ZV 4) and two pairs on soft cuticle (ZV 4 and ZV 5), but setae JV 4 and ZV 3 absent (present in female); chelicerae with elongate middle article, fixed digit with one small subapical tooth and a row of five teeth, movable digit with one distal tooth and an intricately formed spermatodactyl, with club-shaped proximal and medial parts and mushroom-shaped distal part (Fig. 18); epistome as in female (Fig. 19); legs II slightly thickened and stouter than the others, and partially armed ‒ some anteroventral setae, namely av on femur II (Fig. 21) and av 2 on tarsus II (Fig. 22), modified into a stout spur.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66DFFC8FF0CC487FCC6FB39.taxon	materials_examined	Type material examined Holotype female: SW Slovakia, Borská Nížina Lowland, Šaštín-Stráže Town, Gazárka Settlement, Jubilejný Les Forest, elevation 175 m, in bark crevices of an old oak (Quercus sp.) heavily soaked with xylem sap, in a mixed forest with pines (Pinus sylvestris) and oaks, 10 August 2019 (leg. author); paratype female, with the same data as the holotype; type specimens were deposited in the Institute of Zoology, Slovak Academy of Sciences, Bratislava, Slovakia.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66DFFC8FF0CC487FCC6FB39.taxon	diagnosis	Diagnosis (Female) Dorsal shield with 37 pairs of setae, including four pairs of marginal setae (r 2 ‒ r 5); r 6 and R 1 ‒ R 6 and two pairs of submarginal setae (UR) on soft cuticle adjacent to lateral margins of dorsal shield; dorsal setae needle-like and long, usually projecting far beyond bases of following setae in longitudinal row; vertical setae (j 1) straight, blunt, rod-shaped, and arranged on regularly curved vertex. Pre-anal setae incomplete, opisthogaster with only eight pairs of setae (JV 1 ‒ JV 5, ZV 1, ZV 4, ZV 5), of which only one pair (JV 2) on ventrianal shield. Peritremes thin, with free anterior and posterior ends, each terminating in a stigma between coxae III and IV. Peritrematal shields narrow, each with short poststigmatic part connected to parapodal part of exopodal platelet. Anterior margin of epistome tricuspid, median cusp conspicuously larger than lateral cusps. Middle article of chelicerae long and thin. Pretarsi and claws of legs I distinctly smaller than those of legs II ‒ IV; tarsi I longer than tibiae I; legs shorter than idiosoma except well elongated legs IV; trochanter I with four setae. Minor duct relatively short and broad (not filiform).	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66DFFC8FF0CC487FCC6FB39.taxon	description	Description (Female) As in Mašán (2023 a), pages 462 – 466, figures 1 – 8.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66DFFC8FF0CC487FCC6FB39.taxon	discussion	Note This species was described by Mašán (2023 a), but its name is not available because the author did not indicate where the type specimens are deposited (ICZN Article 16.4.2).	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66DFFCBFF0CC16EFD22FA38.taxon	discussion	The genus Hyattella was established as a monotypic genus by Krantz (1962: 6), based on Hyattella epicrioides Krantz, 1962 as the type species, collected from Central Africa (Democratic Republic of the Congo), with the following brief diagnosis for the female: “ Dorsal shield entire, without incisions; posterior part of shield with fifteen pairs of pectinate, distally expanded setae; vertical setae well developed; presternal shields present; pulvillar lobes rounded. ” All of these characters are too general and fit several genera. Krantz (1962) placed Hyattella in the subfamily Platyseiinae and the family Aceosejidae (now Blattisociidae) along with Sejus C. L. Koch (now Cheiroseius Berlese), Plesiosejus Evans & Hyatt, and Platyseius Berlese. Hyattella was placed in Platyseiinae primarily because of the elongated setae on the hypostome (which are actually normal) and palp trochanter, and the presence of the specialised strap-like macrosetae on the telotarsi II ‒ IV. Krantz considered the placement of his new genus within the subfamily as tentative and pointed out the absence of some typical features of Platyseiinae with respect to some structures of the leg ambulacrum and cheliceral dentition. Important apomorphic features of Platyseiinae include an acuminate median lobe of the pulvillus and elongate and acuminate paradactyli on the ambulacrum of the tarsi II ‒ IV (as an adaptation to movement in moist substrates), as well as some structures on the fixed cheliceral digit such as a deep subapical receptacle for the tip of the movable digit and a row of small to fine teeth confined to a ridge in the apical third of the masticatory surface (Lindquist & Moraza 2016). Krantz (1962) also proposed that Hyattella represents an aberrant member of the aceosejine genus Lasioseius (based on the rounded pulvillar lobes, dorsal chaetotaxy, and cheliceral dentition) or a taxon with a position between Platyseiinae and Aceosejinae, and he systematically listed the genus Zerconopsis Hull as the most closely related group with respect to the modified strap-like macrosetae on the tarsi II ‒ IV. Although the presence of tarsal macrosetae did not enter into the genus diagnosis, it was the main reason for Krantz to establish his new genus. However, Lindquist & Moraza (2016) evaluated the presence of these macrosetae as an expression of convergence rather than apomorphy. Currently, the genus is recognised by several authors as a synonym of the genus Lasioseius Berlese, 1916 (Lindquist & Evans 1965; Walter & Lindquist 1997; Halliday et al. 1998). In addition, the generic name Hyattella Krantz, 1962 is a primary junior homonym and is occupied by Hyattella Lendenfeld, 1888: 233 (Porifera: Dictyoceratida: Spongiidae), so this name is not available in the systematic nomenclature of Acari. Since Hyattella Krantz, 1962 has an available synonym (Lasioseius Berlese, 1916), it is not necessary to establish the replacement name for this junior homonym (ICZN Article 60.1). Since Lasioseius cheiroseioides (and its related species from the Palaearctic) also possesses specifically modified tarsal macrosetae on the legs II – IV as originally described in Hyattella epicrioides (actually Lasioseius epicrioides), the type specimens of this type species of Hyattella were also examined to more accurately assess the mutual systematic affiliation of the two species. The examination and comparisons revealed some minor inconsistencies in the original description and illustrations of L. epicrioides, as well as significant morphological differences in some of the characters considered, despite the unusual presence of specialised tarsal macrosetae in both species (Figs 26, 28, 32, 33). Krantz (1962) illustrated L. epicrioides with 38 pairs of setae on the dorsal shield and ten pairs of setae on the lateroventral soft cuticle, making no special comments on the number of setae in his description, and indicating and illustrating only five pairs of pre-anal setae on the ventrianal shield. In the holotype and the six paratypes available for my study (all females), I found 37 pairs of dorsal setae (all j / J, z / Z, and s / S setae, and r 2 ‒ r 5), including the rather short and needle-like setae s 1, s 2, r 2, and r 4 (these setae are not modified and serrated like the other short but somewhat longer setae j 2, z 2, and J 5); the ventrianal shield (Fig. 33) has six pairs of pre-anal setae (JV 1 ‒ JV 4, ZV 2, and ZV 3) in addition to three circum-anal setae, and the soft cuticle has JV 5, ZV 4, ZV 5, and 4 ‒ 5 pairs that could be considered both UR setae and R setae (one pair shown by Krantz on the soft cuticle at the level of the coxae III is actually on the dorsal shield). A similar species described from Angola by Aswegen & Loots (1969), L. dundoensis, appears to be identical to L. epicrioides, and its validity should be verified in a further study of the genus. From my comparison of L. cheiroseioides and L. epicrioides, it appears that both species cannot be placed together in one species group in some further taxonomic concepts of the genus (if they are developed in the future). In contrast to L. cheiroseioides, the dorsal shield of H. epicrioides has a reticulate ornamentation with spherical tubercles (Fig. 26) and covers the entire dorsum and is unusually lateroventrally expanded to cover also the narrow lateral and posteroventral margins of the ventral idiosoma, the rows of marginal setae are incomplete, the presternal platelets are free from the sternal shield (Fig. 32), the metapodal region has only one pair of platelets, setae JV 5 are short and needle-like, legs I are conspicuously long (including the tarsus) and significantly longer than legs IV, the chelicerae do not have a conspicuously elongated middle article, but the fixed digit is densely toothed along the entire masticatory surface, the epistome is tricuspid rather than triramous, the strap-like macrosetae of the tarsi II ‒ IV are straight (not curved and whip-like in the distal part), the tarsus III has only one macroseta (pd 2 absent), and the macroseta on the tarsus IV is al 2 instead of ad 2.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D66EFFD4FF0CC271FE25F9D4.taxon	discussion	Lasioseius minor was described as one of the smallest species native to Europe (Fig. 29), based on females from a soil sample from a thermophilic oak forest in southwestern Slovakia (Kalúz 2009: 1157). The dorsal shield of this species (Fig. 34) was originally described and illustrated with the correct number of 36 pairs of setae, but as a revision of the type specimens and my own material showed, the idiosomal setation was misinterpreted in some details: (1) the paravertical setae (z 1) are shown in the original illustration of the dorsal shield, although they were absent from all four type specimens available for my study; (2) setae s 2 (labeled r 2) are considered to be located outside the shield, but are clearly part of the number of 36 pairs of dorsal shield setae, as in other species of the genus; (3) setae ZV 5 are omitted and absent from the original illustrations of the idiosoma, although they are normally located behind R 6. In addition, Kalúz (2009) misinterpreted the chaetotaxy of several leg segments (trochanter II, femur II, and tarsi II ‒ IV), which are in fact characterised by the standard number of setae, as is the case in most species of the genus. Christian (1990: 31) described an equally small and similar Lasioseius species, namely L. kargi, from the leaf litter of a deciduous forest in Germany [the holotype was examined via ʻThe Virtual Microslide Collectionʼ of the ʻSenckenberg Museum für Naturkunde Görlitzʼ; see Christian et al. (2018)]. Like L. minor, the species is also characterised by the absence of paravertical setae [this feature is found exclusively in morphologically distinct members of the L. phytoseioides species group, as defined by Moraes et al. (2015 b)], the placement of marginal setae r 5 on the dorsal shield, the presence of six pairs of pre-anal setae on the ventrianal shield, the relatively large outer pair of metapodal platelets, and the unusual four teeth on the movable digit of the chelicerae. Just as Kalúz (2009) overlooked the existence of L. kargi Christian, 1990, Christian (1990) also overlooked the existence of L. kargi Kandil, 1980. Only later Christian & Karg (2006: 125) assigned a new replacement name for the homonymous species established by Christian (1990), namely L. diffindatus Christian & Karg, 2006. Based on my comparisons of the type specimens of L. minor and L. diffindatus as well as an examination of my own numerous specimens from Slovakia, I consider both species to be identical. Therefore, I relegate L. minor into synonymy with L. diffindatus. There is only one minor inconsistency in the morphological concepts of L. minor and L. diffindatus that could be commented on. In contrast to the illustration of the metapodal areas in Kalúz (2009), Christian‘s illustration of these areas lacks the anteromedial pair of platelets, which are indeed very small and suboval and resemble sclerites rather than platelets. According to Christian and his photographs (personal communication), these structures are abnormally and asymmetrically developed in the holotype (the only type specimen available). On the right side, this platelet is divided into two smaller ones, and on the left side, it is not free and well separated, as it usually is, but lies close to the edge of the adjacent larger platelet. In this context, Lindquist (1964) and Moraza & Lindquist (2011) pointed out the variability of metapodal scutal elements in some species of Lasioseius. They gave the following two examples: in L. confusus Evans, 1958, the metapodals vary from entire to partly subdivided, to entirely subdivided but contiguous, while in L. allii Chant, 1958 they are consistently subdivided, although the smaller platelet may be very small in some specimens. It is interesting to note that these two identical species proposed here for synonymisation were placed in two different subgenera by their original describers, namely L. minor in Crinidens Karg, 1980 [synonymised by Moraza & Lindquist (2011) under the nominate subgenus] and L. diffindatus in Lasioseius s. str. This difference in placement was mainly a consequence of Christian (1990) describing and illustrating the metapodal areas of this species with only one pair of metapodal platelets instead of two. The undescribed male stage is unusual by almost complete reduction of R and UR rows of setae (only R 3 present and placed on soft cuticle; compared to female, five pairs of R setae and two pairs of UR setae are absent); setae r 6 on dorsal shield, not on soft cuticle as in female; ventrianal region with only JV 1 ‒ JV 3, JV 5, ZV 1 and ZV 2 on ventrianal shield, and ZV 5 on soft integument (compared to female, JV 4, ZV 3, and ZV 4 are absent). This thermophilic and saproxylic species is not rare in Slovakia, and I collected it on the vast territory of southern Slovakia in the following habitats: on wood-destroying fungi on deciduous trees (eight finds), under the bark of deciduous trees (seven finds), in decomposing wood of old oaks inhabited by a colony of ants (five finds), in leaf litter of oak forests (two finds), in nests of Micromammalia and in decaying plant remains (one find each). In Slovakia I have not yet found this species in higher and colder areas in the central and northern regions or in association with conifers.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
03A72A36D671FFD6FF0CC203FB74FD2C.taxon	discussion	Willmann (1954: 225) described Lasioseius zerconoides based on 13 females he had collected from under the bark of trees in the former Czechoslovakia (now the Czech Republic). He illustrated the dorsal and ventral idiosoma, some dorsal setae, and the epistome. The poor condition of the specimens he examined undoubtedly did not allow him to describe some important morphological structures in greater detail or accuracy. For example, he described and illustrated the sternal shield with four pairs of setae (st 1 ‒ st 4) and the ventrianal shield with seven pairs of pre-anal setae (JV 1 ‒ JV 5, ZV 2, and ZV 3), and these misleading statements were later adopted by Karg (1971) and Bregetova (1977) for their then modern and still widely used identification keys. Later, Karg (1980) and then also Gwiazdowicz (2007), examining and illustrating the same type specimens of Willmann, found only three pairs of setae on the sternal shield (st 1 ‒ st 3) and four pairs of pre-anal setae on the ventrianal shield (JV 1 ‒ JV 3 and ZV 2). But unfortunately, like Willmann (1954), they also neglected the specific concave formation of the posterior margin of the sternal shield with its deep notch (desclerotisation), the most important diagnostic feature of several species with reduced sclerotisation of the sternal shield (Mašán & Halliday 2023). The above circumstances lead to misinterpretation of some morphological characters and misidentification of easily recognisable Willmann species. The sternal areas of all four examined females of the original Willmann type series currently deposited in the the Bavarian State Collection of Zoology in Munich (the slides No. W 35 / 21 ‒ 24) are shown in the micrographs in Figs 35 ‒ 38. In all photographed specimens, a conspicuous incision on the posterior margin of the sternal shield is more or less pronounced. Such a specific, deeply incised shape of the sternal shield, together with four pairs of pre-anal setae on the ventrianal shield, is typical for only two species of the genus, namely L. ometes (Oudemans, 1903) and L. verruciger Mašán & Halliday, 2023 [for reliable distinguishing characters between them see the identification key of Mašán & Halliday (2023)]. I have not examined the type specimens of L. ometes, described by Oudemans (1903: 100) from the Netherlands, but the available specimens of L. zerconoides are clearly conspecific with the specimens that are commonly and abundantly collected in Europe and that are widely identified as L. ometes. Since I could not find any reliable distinguishing characters in Willmann‘s specimens that would provide a useful basis for establishing a separate species, and all important characters of the type specimens of L. zerconoides are within the range of variation that I found in my own specimens of L. ometes, I relegate L. zerconoides into synonymy with L. ometes.	en	Mašán, Peter (2023): On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962. Zootaxa 5361 (2): 159-180, DOI: 10.11646/zootaxa.5361.2.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.2.2/52177
