identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A2C77C4668FFC7C79CB7201227DB1C.text	03A2C77C4668FFC7C79CB7201227DB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeoidea Balogh 1965	<div><p>Eutegaeoidea J. Balogh, 1965</p><p>Eutegaeoidea J. Balogh, 1965, p. 59.</p><p>Diagnosis. Monodactylous; hysterosoma distinctly wider than long, less than twice length of prodorsum; lamellae very broad, cusped, rounded or transverse and fused apically, typically overlapping lateral margins of prodorsum; bothridia long, cylindrical or horn-shaped or fused with lateral prodorsum, projecting anterolaterally and with internal spiral thickening; bothridial seta long, typically setiform or bacilliform. Prodorsum posterior of bothridia often conspicuously waisted and narrow in adults and immatures. Humeral processes short and triangular or long, pointed, projecting anteriorly beyond level of bothridia but typically not bearing setae (except  Porrhotegaeus catherinae sp. nov.; cf. below); dorsosejugal scissure transverse; cerotegument commonly present; notogastral integument smooth, never sculptured, with seven to ten pairs of setae. Circumpedal carina typically present (absent in  Cerocepheidae and  Bornebuschiidae fam. nov.). Perigenital carinae, enantiophyses E4 and complete or partial enantiophyses H, B and V present or absent. With five to seven pairs of genital setae and two or three pairs of adanal setae. Immatures eupheredermous, lacking central gastronotic setae of the d series in the nymphal stages; sometimes with flat, elaborate scales from which marginal setae emerge.</p><p>Remarks. A checklist and classification of the  Eutegaeoidea, based on the present work (Table 2), includes seven families, 14 genera and 84 species;  Eutegaeidae:  Eutegaeus (21 spp.);  Neseutegaeus (6 spp.);  Atalotegaeus (6 spp.);  Neoeutegaeidae fam. nov.:  Humerotegaeus gen. nov. (2 spp.);  Neoeutegaeus (7 spp.);  Compactozetidae:  Compactozetes (10 spp.);  Hamotegeus (4 spp.);  Sadocepheus (14 spp.);  Bornebuschiidae fam. nov.:  Bornebuschia (2 spp.);  Dicrotegaeus (3 spp.);  Pterozetidae:  Pterozetes (2 spp.);  Dudichella (1 sp.);  Porrhotegaeidae fam. nov.:  Porrhotegaeus (4 spp.);  Cerocepheidae:  Cerocepheus (2 spp.).</p><p>Luxton (1988a) used the smooth notogaster as one of the character states to differentiate  Eutegaeoidea from  Cepheoidea . No eutegaeoids have a sculptured notogaster, whereas all cepheoids do, except  Conoppia Berlese, 1908 and three species of  Sphodrocepheus Woolley &amp; Higgins, 1963 . Also, most cepheoids have the synapomorphy of a circumnotogastral ring or margin ornamented with different microsculpture from that of the centrodorsal region (present in all species of  Cepheus,  Hypocepheus,  Ommatocepheus,  Oribatodes,  Tereticepheus; some species of  Eupterotegaeus,  Sphodrocepheus and  Tritegeus; absent in  Conoppia,  Pilocepheus,  Protocepheus,  Reticulocepheus and  Roycepheus). The structure is similar to that in  Scapheremaeus ( Cymbaeremaeidae) but with the bases of the notogastral setae or, at most, a groove, rather than a scissure, marking the division between the two contrasting areas of cuticle. In  Tritegeus Berlese, 1913, some species of  Sphodrocepheus and  Hypocepheus Krivolutsky, in Krivolutsky and Tarba, 1971, this margin is positioned more medially and aligned with the bases of setae of the l and h series (cf. Bernini and Bernini, 1990, Figs. 12 and 16; Woolley and Higgins, 1963, Fig. 1; Krivolutsky and Tarba, 1971, Fig. 1). The circumnotogastral margin tends to be flattened (except in  Tritegeus spp.) and the centrodorsal region is flat or slightly convex (Bernini and Bernini, 1990, Fig. 1A therein), whereas in  Eutegaeoidea the notogaster is strongly and uniformly convex, except in  Porrhotegaeus where it is concave with raised margins.</p><p>Luxton (1988a) stated that the notogaster of  Cepheoidea is at least twice as long as the prodorsum and in  Eutegaeoidea it is markedly less so. The mean ratio of the length of the notogaster to the prodorsum in  Cepheoidea (n = 54 spp.) is 2.95 (range: 1.72–5.1) and for  Eutegaeoidea (n = 80 spp., not including  Hamotegeus spp.) 1.62 (range: 1.1–1.97).  Hamotegeus spp. are the only  Eutegaeoidea that markedly exceed the 2:1 ratio, with a mean of 2.27; range 1.98–2.57. The ratio of notogastral length to breadth in  Eutegaeoidea is 0.82 (range 0.71–1) and in  Cepheoidea 1.06 (range 0.92–1.27).</p><p>As well as the distinctly different biogeographical distribution of  Eutegaeoidea and  Cepheoidea (Fig. 1), the dorsosejugal scissure in eutegaeoids is transverse and straight (except in some species of  Hamotegeus), whereas in cepheoids it is convex and rounded. Humeral processes in  Eutegaeoidea are typically long, well-developed and pointed or rounded (except in  Cerocepheidae), whereas in  Cepheoidea they are small, short, ledge-shaped structures or absent. Other character states occur in a rather matrix-like pattern. For example, the bothridia of  Eutegaeoidea are either prominent, long, curved and corniculate, separate anteriorly from the margin of the lamella or, as in  Compactozetidae, large, but fused with the lamellae. In  Cepheoidea, the bothridia tend to be short and squat, except in  Eupterotegaeus where they are long and corniculate in some species. Bothridial setae of  Eutegaeoidea tend to be long, thin and setiform or bacilliform and those of  Cepheoidea are short and club-shaped, though in  Eupterotegaeus they tend to be long and club-shaped. In  Eutegaeoidea, the lamellae are generally parallel (except in  Sadocepheus) and the lamellar cusp is free and with the lamellar seta emerging from an indentation flanked laterally by teeth or, as in  Pterozetidae and some  Compactozetidae, the apices of the lamellae are fused. In  Cepheoidea, the lamellae are strongly convergent apically (except in  Eupterotegaeus and  Ommatocepheus) and the free lamellar cusp is typically triangular with the lamellar seta emerging from its apex, not flanked by teeth, except in  Eupterotegaeus and  Ommatocepheus where the lamellar cusps are curved and the lamellar setae emerge sub-apically.</p></div>	https://treatment.plazi.org/id/03A2C77C4668FFC7C79CB7201227DB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C466AFFC0C79CB6D514F6D960.text	03A2C77C466AFFC0C79CB6D514F6D960.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeidae J. Balogh 1965	<div><p>Eutegaeidae J. Balogh, 1965</p><p>Eutegaeidae J. Balogh, 1965, p. 59 .</p><p>Type genus:  Eutegaeus Berlese, 1916, p. 62 .</p><p>Diagnosis. the following diagnosis is modified from those of J. Balogh (1965), Woolley (1965) and Luxton (1988a). Rostrum acute, entire; lamellae very broad, covering lateral margins of prodorsum, with free cusps; lamellar seta emerging from excavation at apex of cusp, often between two teeth. Interlamellar setae positioned medial of bothridia; bothridium not fused with bases of lamellae, long, corniculate, opening anterolaterally, often with anterior condyle of enantiophysis H on posterior margin. Bothridial seta typically very long, setiform or bacilliform. Prodorsum waisted posterior of bothridium. Humeral processes prominent, ovoid, waisted basally, pointed apically, extending from anterolateral margin of notogaster to point at least halfway along length of lamella, but not reflexed ventrally. Notogaster circular to ovoid, convex, smooth, broader than long. With eight or nine pairs of notogastral setae, either all setae of the l and h series in the centrodorsal position ( Neseutegaeus) or at least one pair in the l series in centrodorsal position ( Eutegaeus and  Atalotegaeus); setae of l and h series (and occasionally p 1) longer and of different morphology from those of p series (or p 2 and p 3). Chelicerae chelate-dentate and of normal proportions or slim but not with needle-like modified digits. Pedotectum I (pd I) sub-rectangular in lateral outline; pd II triangular or rectangular; discidium pointed. Circumpedal carina present. Perigenital carinae and enantiophyses E4 absent. Five or six pairs of genital setae, three pairs of adanal setae. Anal plates lozenge-shaped. Pre-anal organ T-shaped. Nymphs with or without one or most gastronotic setae emerging from flat scales; lateral margin of lamellar region strongly incised.</p><p>Remarks. Since the publication by J. Balogh (1965), the  Eutegaeidae has contained between two and nine genera (Table 1). The diagnosis above is based on a concept of the  Eutegaeidae as containing only three genera (Table 2):  Eutegaeus,  Atalotegaeus and  Neseutegaeus, due to  Birotegaeus and  Pareutegaeus being newly synonymised with  Eutegaeus, as detailed below.</p><p>The main character states that differentiate  Eutegaeidae from the other families of  Eutegaeoidea are the long, pointed humeral processes that emerge from the anterolateral margin of the notogaster, directly posterior of the bothridia and have a narrow, waisted point of attachment to the notogaster; narrower than the anterior part of the humeral process and one or more of the setae of the l series in the centrodorsal position. These two character states are synapomorphies of  Eutegaeidae . In  Neoeutegaeidae fam. nov. (cf. below) the humeral process also emerges on the anterior notogaster but its base is broad and has a distinct bulge and incurve where it meets the notogaster. In  Compactozetidae the humeral process has its origin some distance posterior of that in  Eutegaeidae and  Neoeutegaeidae fam. nov., at the widest part of the notogaster. In  Pterozetidae and  Porrhotegaeidae fam. nov. the humeral process also originates in this position and is pointed apically and broad basally; its lateral margin is straight and lacks any prominent constriction or bulge at its base. In  Bornebuschiidae fam nov. (cf. below) the humeral process also originates in this posterior position but it is rounded apically and does not extend anteriorly beyond the bothridium. In  Cerocepheidae the humeral process is reduced to a short, triangular structure.</p><p>The lamellae of  Eutegaeidae,  Cerocepheidae and  Neoeutegaeidae fam. nov. and  Porrhotegaeidae fam. nov. have free cusps and are broad, not apically convergent, with convex or straight lateral margins and the lamellar seta emerges from an excavation at the apex of the cusp, typically between two teeth, or the cusp is incurved and the lamellar seta emerge sub-apically ( Atalotegaeus). In  Compactozetidae the lamellae are fused with each other apically and curved ventrally and fused with the rostrum in  Compactozetes or are slightly convergent apically and have a rounded or obliquely-angled free cusp ( Sadocepheus,  Hamotegeus). In  Pterozetidae the lamellae are fused with each other apically but not folded ventrally. In  Bornebuschiidae fam nov. the lamellar seta is positioned some distance posterolaterally, at the base of the lamellar cusp, which is either curved and blunt ( Bornebuschia) or very long and pointed, extending well beyond the rostrum ( Dicrotegaeus). In  Porrhotegaeidae fam. nov. the lamellae are similar to those of  Eutegaeidae, but the lamellar seta emerges from the ventral surface of the lamellar cusp.</p><p>The margins of the coxisternum, comprising pedotecta I and II and the discidium, are relatively uniform in morphology in  Eutegaeidae and  Cerocepheidae when viewed ventrally. The morphology of these structures in ventral aspect does not fully reveal their three-dimensional shape, which becomes more apparent in lateral aspect (compare Figs. 27b and 27c) but, as mentioned in the methods section above, lateral aspects were not illustrated because of the difficulty in making temporary mounts of specimens in which the hysterosoma is broader than long. However, the ventral aspects of these structures (Fig. 44) reveals useful similarities and differences between taxa. Pedoctectum I is sub-rectangular in all genera of  Eutegaeidae and  Cerocepheidae and the anterior margin is transverse or oblique. Pedotectum II is pointed and triangular/rectangular in  Atalotegaeus and  Cerocepheus, in those species of  Eutegaeus for which the ventral surface has been described and in  Neseutegaeus . The discidium is thin, triangular and pointed in  Cerocepheidae and all species of  Eutegaeidae for which the structure has been described. In  Neoeutegaeidae fam. nov. pd I is sub-rectangular, pd II is massive and either trapezoid or lobe-like and the discidium is lobe-like or sub-circular. In  Compactozetidae, pd I has a prominent anterior spine, pd II is very large and lobe-like or rectangular and the discidium is lobe-like or sub-triangular. In  Pterozetidae pd I is sub-rectangular, pd II is rectangular and the discidium is rectangular or trapezoid. In  Porrhotegaeidae fam. nov. pd I and II are sub-rectangular, pd I lacks a spine and the discidium is triangular and pointed.</p></div>	https://treatment.plazi.org/id/03A2C77C466AFFC0C79CB6D514F6D960	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C466DFFC2C79CB44F15AADDC0.text	03A2C77C466DFFC2C79CB44F15AADDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus Berlese 1916	<div><p>Eutegaeus Berlese, 1916</p><p>Eutegaeus Berlese, 1916, p. 62 .</p><p>Type species:  Oribata bostocki Michael, 1908, p. 136 .</p><p>Birotegaeus Luxton, 1988a, p. 82,  syn. nov.</p><p>Type species:  Eutegaeus biroi J. Balogh, 1970, p. 296 .</p><p>Pareutegaeus Woolley, 1965, p. 384,  syn. nov.</p><p>Type species:  Eutegaeus similis Trägårdh, 1931, p. 575 .</p><p>Diagnosis. The following diagnosis is modified from that of Luxton (1988a) and Ermilov (2020b) and includes a character state for immatures. Adults: large oribatid mites (&gt;800 μm); rostrum rounded; lamellar cusp about one-third length of lamella, projecting beyond rostrum, separated medially and joined by complete or partial translamella or translamella absent; lamellar setae inserted in notch on anterodorsal edge of lamellae, each seta flanked by one or two teeth or teeth absent. Bothridia often with anterior condyle of enantiophysis H on posterior margin but posterior condyle absent from anterior notogaster; with or without two ovoid structures posterior of interlamellar setae representing anterior condyles of enantiophysis B. Bothridial seta long and bacilliform or setiform, rarely short and club-shaped. Humeral process ovoid, pointed, waisted basally, projecting from anterior edge of notogaster, reaching beyond bothridia anteriorly but not extending posteriorly beyond anterior margin of notogaster nor broad basally. Notogaster with eight (rarely nine) pairs of setae; l series in centrodorsal position, the rest marginal or sub-marginal. Pedotectum I (pd I) sub-rectangular in ventral view, pd II and discidium triangular, pointed. Six pairs of genital setae with penultimate pair displaced laterally; three pairs of adanal setae; perigenital carina absent. Chelicerae chelate-dentate, of normal proportions. Pre-anal organ T-shaped. Nymphs: all gastronotal setae except c 1 and c</p><p>2 emerging from scales.</p><p>Remarks. Subías (2004) listed  Eutegaeus curviseta Hammer, 1966 as a junior synonym of  E. bostocki (Michael, 1908) although Luxton (1985) recognized them as distinct species. Both have short setae p 1 on squat tubercles and lamellar cusps with well-developed teeth and a pair of condyles on the prodorsum posterior of the interlamellar setae. This character state is also present in  E. biovatus Hammer, 1972 from Tahiti,  E. fueginus Arcidiacono, 1993 and  E. lagrecai Arcidiacono, 1993 from Tierra del Fuego,  E. nothofagi sp. nov. from Australia (cf. below) and  E. papuensis Aoki, 1964 from Papua New Guinea. However, the two species differ in that the bothridial setae of  E. bostocki are expanded apically and each has a sharp tip, described by Michael (1908, p. 136, Plate 17, Fig. 3) as ‘slightly fusiform heads upon thin peduncles’, whereas those of  E. curviseta are longer, slightly expanded apically but blunt. Also, the interlamellar setae and notogastral setae of the l and h series of  E. bostocki are much shorter than those of  E. curviseta and the l and h series are in the sub-marginal position, whereas in  E. curviseta setae la and lm are centrodorsal (cf. Fig. 13a for definition of locations of centrodorsal, sub-marginal and marginal notogastral setae). The lateral tooth of the lamellar cusp is larger than the medial one in  E. bostocki and the lamellar seta emerges from a deep cup-shaped cleft between them, whereas in  E. curviseta the medial tooth is larger and the lamellar seta emerges from a slight indentation of the apex of the lamellar cusp. Accordingly, I also consider  E. bostocki and  E. curviseta to be distinct species.</p><p>Woolley (1965) established  Pareutegaeus for  Eutegaeus similis Trägårdh, 1931 from the Juan Fernández Islands based on the rounded lamellar cusps and lamellar setae emerging from near their tips, reticulate lamellar microsculpture, long interlamellar setae and with a pair of small tubercles on the posterior margin of the notogaster. Luxton (1988a), who redescribed  E. similis based on the lectotype, found the lamellar cusp has a slight excavation and the lamellar seta emerges apically from the cusp in a shallow invagination between two short, blunt teeth. This character state is found in several  Eutegaeus spp. ( E. fueginus,  E. lagrecai and  E. papuensis). Likewise, the reticulate lamellar microsculpture is common in  Eutegaeus ( E. curviseta,  E. lagrecai,  E. paralagrecai,  E. parapapuensis,  E. pinnatus,  E. radiatus Hammer, 1966,  E. soror,  
E. stylesi Hammer, 1966 and 

E. woiwurrung 
sp. nov.) and in Atalotegaeus mensarosi. This pattern of microsculpture is part of the lamellae themselves, as illustrated in the scanning electron micrograph of E. lagrecai by Arcidiacono (1993, Fig. 2
 and 3 therein) and is not due to any overlying cerotegument as assumed by Ermilov (2020b, pp. 332, 340). The presence of long interlamellar setae, almost as long as the lamellae or longer, is also common in  Eutegaeus ( E. bidhawal sp. nov.,  E. curviseta,  E. paralagrecai,  E. pinnatus and  E. soror). Luxton (1988a) found no small tubercles on the posterior notogaster of  E. similis . He considered  Pareutegaeus a valid genus and differentiated it from  Eutegaeus based on the absence of prominent teeth on the lamellar cusps, but this character state also occurs in  Eutegaeus, as discussed above, as do the other character states in the diagnosis of  Pareutegaeus by Luxton (1988a). Accordingly, I do not consider  Pareutegaeus Woolley, 1965 is a valid genus and hereby designate it a junior synonym of  Eutegaeus Berlese, 1916 .</p><p>Luxton (1988a, p. 82) established  Birotegaeus for  Eutegaeus biroi J. Balogh, 1970 from Papua New Guinea and differentiated it from  Eutegaeus on the basis that it has nine pairs of notogastral setae rather than eight pairs, with la in the centrodorsal position, and a mound on the posterior notogaster. In other respects, the character states that Luxton (1988a) used in his diagnosis of  Birotegaeus do not differ from those of  Eutegaeus, including the shape of the lamellae and lamellar cusps, the humeral processes, the absence of condyles of enantiophyses H, B and E4 and the number and disposition of the notogastral setae.  Eutegaeus papuensis also has nine pairs of notogastral setae (with h 1 present) and several species have setae la (and le) displaced medially to the centrodorsal position, including  E. bostocki,  E. curviseta,  E. pinnatus Hammer, 1966,  E. membraniger Hammer, 1966,  E. woiwurrung sp. nov.,  E. nothofagi sp. nov. and  E. bidhawal sp. nov. A slight posterior notogastral mound is also present in  E. parapapuensis Ermilov, 2020 and  E. paralagrecai Ermilov, 2020 and several species have setae p 1 emerging from a pair of short, closely-adjacent tubercles on the posterior notogaster ( E. biovatus,  E. bostocki,  E. curviseta,  E. membraniger,  E. papuensis,  E. pinnatus,  E. radiatus and  E. stylesi). In the remaining species of  Eutegaeus, setae p 1 emerge from an alveolus. Accordingly, I do not consider the character states of Luxton (1988a) sufficient to differentiate  Birotegaeus from  Eutegaeus and hereby designate  Birotegaeus Luxton, 1988a as a junior synonym of  Eutegaeus Berlese, 1916 .</p><p>In his revised definition of the genus, Ermilov (2020b) included the morphology of the bothridial seta as clavate, lanceolate, setiform or bacilliform in his revised diagnosis of  Eutegaeus . The four species that have short, club-shaped bothridial setae ( E. fueginus,  E. lagrecai,  E. parapapuensis and  E. similis) are all from the southern Neotropical region.</p><p>Ermilov (2020b, p. 350) stated that  Eutegaeus has “eight pairs of notogastral setae (la, lm, lp, h 1, h 2, p 1, p 2, p 3)…rarely with nine pairs of setae (if h 3 present).” I consider that in  Eutegaeus it is not h 3 that is labile and subject to loss in the adult, but h 1, based on the occurrence in the deutonymph of  E. woiwurrung and  E. nothofagii of a scale corresponding with h 1 but the absence of a seta (arrowed in Figs. 3a, 6a). Neither the scale or seta h 1 is present in the deutonymph of  E. ptilosus sp. nov. (Fig. 9a).</p></div>	https://treatment.plazi.org/id/03A2C77C466DFFC2C79CB44F15AADDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C466FFFDDC79CB0E81639D932.text	03A2C77C466FFFDDC79CB0E81639D932.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus woiwurrung Colloff 2023	<div><p>Eutegaeus woiwurrung sp. nov.</p><p>(Figs. 2–4)</p><p>Dimensions. Holotype female: length 1,062 μm, breadth 858 μm. Paratype males (n = 4): mean length 976 μm (range 960–985 μm); mean breadth 774 μm (range 742–815 μm). Paratype females (n = 9): mean length 1,009 μm (range 983–1,036 μm); mean breadth 805 μm (range 750–819 μm). Ratio of prodorsum to total length: 0.38 (holotype). Paratype tritonymphs (n = 6): mean length 859 μm (range 823–908 μm); mean breadth 651 μm (range 602–719 μm).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) curved, well-developed, with spinose ornamentation, visible in dorsal view. Lamellae broad, separated medially, parallel; with partial translamella at base of lamellar cusps; lamellar setae (le) smooth, curved, extending well beyond apices of rostral setae, emerging from cup-shaped incisions, each seta flanked by two sharp teeth of equal size; cusps extending anteriorly just beyond rostrum (Fig. 2a). Interlamellar seta (in) about same length as seta le. Bothridia projecting laterally beyond margin of prodorsum, with anterior condyle of enantiophysis H in form of V-shaped posterior flange. Bothridial seta long, curved, bacilliform, smooth.</p><p>Notogaster: thin membranous cerotegument covering parts of notogaster; ratio of length to breadth: 0.78; notogaster rounded, convex. Humeral processes relatively narrow, with dorsal keel, slightly waisted basally, then expanded and pointed apically, extending to point level with middle of lamella (Fig. 2a). Short, sharp lateral projection bearing lyrifissure ia on notogaster posterolateral of humeral process. With eight pairs of notogastral setae, l series, h 2 and h 3 present, relatively long, curved, with fine spines along convex aspect; setae la and lm in sub-marginal position, lp, h 2 and h 3 and p series marginal (Fig. 2c); setae of p series very short, curved, smooth. Lyrifissures im present lateral of setae lm.</p><p>Ventral aspect: subcapitulum acute apically; mentum as long as broad; subcapitular setae short, smooth, sub-equal in length. Tutorium relatively short, apex blunt. Epimeral plates discrete, all broadly separated in midline, plates I sub-triangular to rectangular, others sub-rectangular, plates II shorter than others. Epimeral setation 3-1-4-2; setae 1b, 3b, 3d and 4c at least twice as long as others (Fig. 2b). Pedotectum I (pd I) well-developed, sub-rectangular; pd II small, pointed; discidium with sharp point. Ventral plate broader than long. Genital and anal plates separated by distance about same as length of genital plates; genital plates 115 µm long, with six pairs of setae, g 5 displaced laterally; lyrifissure iad in para-anal position, some distance from margin of anal plate, same as seta ad 3. Anal plates lozenge-shaped, 214 µm long. Pre-anal organ (po) broad, flat T-shaped.</p><p>Description of tritonymph, nymphal and larval scalps. Tritonymph prodorsum: rostrum acute; rostral setae with fine spines, on short tubercles; lamellar setae with spinose ornamentation, emerging from shallow depression on lamellar cusp; median apex of cusp with long, sharp tooth (Fig. 3a). Lamellae strongly incised laterally just posterior of setae le and again anterior of bothridia, with blocky, alveolate microsculpture. Lateral margin of prodorsum alveolate, posteromedian region smooth, porose; setae in minute. Bothridia with broad flanges posteriorly and anteriorly (Fig. 3b). Bothridial seta long, slim, expanded apically, with sharp tip.</p><p>Tritonymph gastronotum: U-shaped, broader (657 µm) than long (617 µm). Gastronotal setae emerging from elongate, cup-shaped or hemi-semicircular overlapping scales, each with pointed tip anteriorly and posteriorly and ornamented with fine, vein-like pattern from medial setal insertion canal (Fig. 3a). With 11 pairs of setae (c 1 absent): c 2 on short tubercles posterior of bothridium, smooth, setiform; setae c 3, the l and h series and p 1 straight, ornamented with fine spines; scale of seta h 1 smaller than those of other h and l series; setae p 1 on single rectangular scale; setae p 2 and p 3 minute, positioned ventrally. Cuticle alveolate between setal scales and with cerotegument of blunt spines.</p><p>Deutonymphal scalp: with eight pairs of setae: c 1 and c 3, l series and h 2, h 3 and p 1; scale of seta h 1 present but seta absent; setae p 1 without scale but on long apophyses (Fig. 3a).</p><p>Protonymphal scalp: with seven pairs of setae: c 3, l series, h 2, h 3 and p 1; p 1 on short tubercles (Fig. 3a).</p><p>Larval scalp: with alveolate microsculpture; scales absent, seven pairs of setae on short tubercles: l series, h 3 and centrodorsal d series. Setae dp on ovoid sclerite (Fig. 3a).</p><p>Deutonymph ventral aspect: epimeral setal formula 3-1-2-2; with three pairs of genital setae, one pair of aggenital setae, three pairs of adanal setae; anal setae absent (Fig. 4b).</p><p>Tritonymph ventral aspect: epimeral setal formula 3-1-3-3; with five pairs of genital setae, one pair of aggenital setae, two pairs of anal setae and three pairs of adanal setae (Fig. 4a).</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps): 8-(9,10,11)-8.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1027;  paratypes: six females, five males, eight nymphs, ANIC accession nos. 53-1028 &amp; 1029, ANIC 297, Moss and litter,  Nothofagus cunninghami rainforest, Cumberland Creek Valley, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Picnic Area</a> [Cumberland Memorial Scenic Reserve], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Yarra Ranges National Park</a>, Victoria, 37°34’S 145°52’E, 920 m, coll. R. W. Taylor and R. J. Bartell, 4.xi.1970  .   Paratype: female, ANIC accession no. 53-1030, Cumberland Creek Valley nr. Cambarville [Marysville-Woods Point Road], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.88333&amp;materialsCitation.latitude=-37.55" title="Search Plazi for locations around (long 145.88333/lat -37.55)">Yarra Ranges National Park</a>, Victoria, 37º33’S 145º53’E, coll. R. E. Roughley, 12.iii.1996  .   Paratypes: two females, one tritonymph, ANIC accession no. 53-1031, moss,  Nothofagus cunninghami rainforest, rainforest walk, Maits Rest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.55&amp;materialsCitation.latitude=-38.75" title="Search Plazi for locations around (long 143.55/lat -38.75)">Great Otway National Park</a>, Victoria, 38°45’S, 143°33’E, coll. M.J. Colloff, 19.vii.2011  .   Paratype female, ANIC accession no. 53-1032, TAS-012, pyrethrum knock-down,  Nothofagus cunninghamii rainforest, Site 3, Pipeline Road, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.33333&amp;materialsCitation.latitude=-41.5" title="Search Plazi for locations around (long 145.33333/lat -41.5)">Savage River</a>, Tasmania, 41º30'S 145º20'E, coll. J. Diggle, 20.iv.1989  .</p><p>Etymology. The specific name is a noun in apposition. The species is named for the Woiwurrung People of the Yarra Valley and the Victorian Great Dividing Range on whose traditional land this species was found.</p><p>Diagnosis.  Eutegaeus woiwurrung can be distinguished from other species in the genus by the following combination of characters: (1) the barbed rostral setae; (2) the presence of a projection posterior of the humeral process; (3) the epimeral setal formula 3-1-4-2; (4) some epimeral setae are twice as long as others; (5) lyrifissure iad is some distance from the margin of the anal plate; about the same as that of ad 3; (6) the lamellar seta emerges from a deep, cup-shaped incision of the lamellar cusp and is flanked by two sharp teeth of equal length; (7) the posterior margin of the bothridium bears the anterior condyle of enantiophysis H.</p><p>Remarks. The other  Eutegaeus spp. that have a sharp lateral projection posterior of the humeral process are  E. nothofagi sp. nov.,  E. bidhawal sp. nov.,  E. odontatus sp. nov.,  E. curviseta and  E. membraniger, the latter two from New Zealand.  E. nothofagi sp. nov. is morphologically most similar to  E. woiwurrung . The former species also has the epimeral setal formula 3-1-4-2 (also found in  E. papuensis from Papua New Guinea) and a V-shaped bothridial flange. However,  E. nothofagi sp. nov. is much larger, has small teeth on the lamellar cusp with a shallow incision and has condyles on the prodorsum.</p><p>The association of the immatures with the adult is based on their presence in the same samples and the absence of other adults of  Eutegaeus spp. The tritonymphs of  E. woiwurrung are 859 μm long, which accords with the mean length of the adults (females 1,062 μm, males 976 μm), whereas the tritonymphs of  E. nothofagi sp. nov. are longer than the adults of  E. woiwurrung (cf. below). Also, the shape and size of the large median tooth on the lamellar cusp of the tritonymph of  E. woiwurrung matches that of the adult, whereas the paired teeth on the lamellar cusp of the adult  E. nothofagi sp. nov. are smaller and of a different shape.</p><p>The tritonymph of this species has the epimeral setal formula 3-1-3-3, but in the adult, it is 3-1-3-2, with seta 4b absent as in  E. bhidawal sp. nov.,  E. ptilosus sp. nov.,  E. odontatus sp. nov.,  E. similis and  E. soror . Seta 4b is present in the adults of those other species of  Eutegaeus for which the epimeral setal formula can be clearly determined.</p></div>	https://treatment.plazi.org/id/03A2C77C466FFFDDC79CB0E81639D932	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4673FFD8C79CB388163ADB68.text	03A2C77C4673FFD8C79CB388163ADB68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus nothofagi Colloff 2023	<div><p>Eutegaeus nothofagi sp. nov.</p><p>(Figs. 5, 6)</p><p>Eutegaeus sp. nov. B: Colloff, 2022, p. 47.</p><p>Dimensions. Holotype: length 1,309 μm, breadth 980 μm; paratype females (n = 10): mean length 1,206 μm (range 1,106 –1,267 μm), mean breadth 874 μm (range 844–909 μm). Paratype males (n = 4): mean length 827 μm (range 792–901 μm), mean breadth 614 μm (range 614–620 μm). Tritonymph: mean length 995 μm, mean breadth 711 μm. Ratio of prodorsum to total length: 0.35 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) flagelliform, reflexed, smooth, on short tubercles, visible in dorsal aspect.Lamellae broad, separated medially, parallel, lateral margins conspicuously convex, translamella absent. Lamellar setae (le) smooth, curved, extending well beyond apices of rostral setae, emerging from shallow cup-shaped incisions, each flanked by two short sharp teeth of equal size; cusps extending anteriorly well beyond rostrum (Fig. 5a). Interlamellar setae (in) shorter than setae le; anterior condyles of enantiophyses B present. Bothridia projecting laterally to inner margins of humeral processes, openings angled anteriorly, with anterior condyle of enantiophysis H in form of V-shaped posterior flange. Bothridial setae long, curved, bacilliform, smooth.</p><p>Notogaster: thin cerotegument with sinuous interdigitate microsculpture covering parts of notogaster (Fig. 5c); ratio of length to breadth: 0.84; notogaster rounded, convex. Humeral processes very broad, with dorsal keel, slightly waisted at their bases, expanded then becoming pointed apically, extending to point level with middle of lamella (Fig. 5a). Short, sharp lateral projection bearing lyrifissure ia on notogaster posterolateral of humeral process. With eight pairs of notogastral setae; l series, h 2 and h 3 present, relatively long, curved, smooth but with sparse covering of cerotegument basally; setae la and lm in sub-marginal position, lp, h 2 and h 3 and p series marginal (Fig. 5a); setae of p series very short, straight, smooth. Lyrifissures im present lateral of setae lm.</p><p>Ventral aspect: subcapitulum broad apically; mentum broader than long; subcapitular setae short, smooth, setae m twice as long as a and h (Fig. 5b). Tutorium short with blunt apex. Epimeral plates discrete, all broadly separated in midline, sub-rectangular to ovoid, plates IV shorter than others; epimeral setation 3-1-3-3; setae 3b at least twice as long as others. Pedotectum I (pd I) well-developed, sub-rectangular, margins angled obliquely; pd II small, pointed; discidium with rounded point. Ventral plate broader than long. Genital and anal plates separated by distance same as length of genital plate (164 µm), with six pairs of setae, g 5 displaced laterally, g 1 longer than others; lyrifissure iad in para-anal position, same distance from anal plate as seta ad 3. Anal plates lozenge-shaped, 291 µm long; with U-shaped ridge posterior of anal plates bearing setae ad 1 and ad 2. Pre-anal organ (po) T-shaped.</p><p>Description of tritonymph, nymphal and larval scalps. Tritonymph prodorsum: rostrum acute; rostral setae with fine spines, setae on short tubercles; lamellar setae with spinose ornamentation, emerging from shallow depression on lamellar cusp; cusp lacking prominent tooth or teeth (Fig. 6a). Lamellae with blocky, alveolate microsculpture, strongly incised laterally just posterior of setae le and again anterior of bothridia. Posteromedian prodorsum smooth, porose; setae in minute, linked by faint horizontal ridge Bothridia relatively short, no longer than broad; bothridial seta long (166 µm) bacilliform, with pointed tip.</p><p>Tritonymph gastronotum: U-shaped, broader (855 µm) than long (777 µm). Gastronotal setae emerging from elongate, cup-shaped or hemi-semicircular overlapping scales, each with a pointed tip anteriorly and posteriorly and ornamented with diagonal ribs extending from their base and with a median setal insertion canal (Fig. 6a). With 12 pairs of setae ornamented with few fine spines: full complement of c, l, h and p series present, c 2 on margin of scale bearing seta c 3; setae c 3, l and h series and p 1 straight, spiniform; seta h 1 and associated scale smaller than other h and l series and their scales; setae p 1 on single sub-rectangular scale; setae p 2 and p 3 small, setiform, smooth, positioned ventrally (Fig. 6b). Cuticle sparingly alveolate between setal scales, with cerotegument of blunt spines.</p><p>Deutonymphal scalp: with eight pairs of setae: c 1 and c 2 (c 3 absent), l series and h 2, h 3 and p 1; c 2 on short tubercles posterior of bothridium, smooth, setiform; scale of seta h 1 present but seta absent; setae p 1 both on single rectangular scale (Fig. 6a).</p><p>Protonymphal scalp: with eight pairs of setae, same complement as deutonymphal scalp, scale of seta h 1 absent (Fig. 6a).</p><p>Larval scalp: microsculpture tuberculate; scales absent, eight pairs of setae on short tubercles: c 1, the l series, h 3 and centrodorsal series d. Setae dp on small sclerite (Fig. 6a).</p><p>Tritonymph ventral aspect: ventral plate with fine corrugations and tubercles; anal plate with fine tubercles; epimeral setal formula 3-1-3-3; with four pairs of genital setae, one pair of aggenital setae, two pairs of anal setae and three pairs of adanal setae (Fig. 6b).</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps): 9-(10,10,12)-8</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1033;  paratypes: ten females, four males, one tritonymph, ANIC accession no. 53-1034, litter,  Nothofagus cunninghami rainforest, Triplet Falls, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.5&amp;materialsCitation.latitude=-38.666668" title="Search Plazi for locations around (long 143.5/lat -38.666668)">Great Otway National Park</a>, Victoria, 38°40’S, 143°30’E, coll. M.J. Colloff, 19.vii.2011  .   Paratypes: ten females, one tritonymph, ANIC accession no. 53-1035, moss,  Nothofagus cunninghami rainforest, rainforest walk, Maits Rest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.55&amp;materialsCitation.latitude=-38.75" title="Search Plazi for locations around (long 143.55/lat -38.75)">Great Otway National Park</a>, Victoria, 38°45’S, 143°33’E, coll. M.J. Colloff, 19.vii.2011  .</p><p>Etymology. This species is named for the Myrtle Beech forest ( Nothofagus cunninghami) in which it was found.</p><p>Diagnosis.  Eutegaeus nothofagi can be distinguished from other species in the genus by the following combination of characters: (1) the very broad lamellar cusps with apical teeth pointing laterally; (2) the absence of a translamella; (3) the presence of anterior condyles of enantiophyses B; (4) the broad, sharply pointed humeral processes with a pronounced dorsal keel; (5) the bothridium with the anterior condyle of enantiophysis H; (6) the prominent, sharp projection posterolateral of the humeral process; (7) the prominent, sub-rectangular pedotecta I with their lateral margins angled obliquely; (8) the U-shaped ridge posterior of the anal plates bearing setae ad 1 and ad 2.</p><p>Remarks. The absence of a translamella in this species is shared with  E. bidhawal sp. nov.,  E. parapapuensis,  E. paralagrecai and  E. pulcher J. Balogh &amp; Csiszár, 1963 . The latter species also has broad, sharply pointed humeral processes and the anterior condyle of enantiophysis H on the posterior margin of the bothridium but differs in the shape of the lamellar cusps and the reticulate microsculpture on them, the barbed lamellar setae and very short interlamellar setae. The notogaster of  E. nothofagi is also much broader than long, whereas in  E. pulcher the notogaster is almost as long as broad.</p><p>The association of the immatures with the adult is based, in part, on their presence in the same samples. At Triplet Falls in the Otway National Park, adults of  E. woiwurrung were also present and the rationale for matching this species with its immatures is given above. In the material of from Maits Rest there were no adult eutegaeoid individuals present other than  E. nothofagi . The tritonymph, at 1,011 μm long, more closely matches the adult of this species, at 1,309 μm long, than that of  E. woiwurrung, at an average length of 1,009 μm for females and 976 μm for males. Also, the small paired teeth on the lamellar cusp of the tritonymph of  E. nothofagi matches the cusp of the adult.</p></div>	https://treatment.plazi.org/id/03A2C77C4673FFD8C79CB388163ADB68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4675FFDAC79CB6401671DFEC.text	03A2C77C4675FFDAC79CB6401671DFEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus bidhawal Colloff 2023	<div><p>Eutegaeus bidhawal sp. nov.</p><p>(Fig. 7)</p><p>Dimensions. Holotype female length 1,094 μm, breadth 896 μm. Paratype females (n = 4) mean length: 1093 μm (range: 1,077 –1,132 μm); mean breadth: 851 μm (range: 821–901 μm). Paratype male length: 1,012 μm, breadth 778 μm. Ratio of prodorsum to total length: 0.33 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) curved, smooth, on short tubercles, not visible in dorsal aspect. Lamellae broad, separated medially, parallel, lateral margins conspicuously convex, without a translamella. Lamellar setae (le) smooth, curved, extending well beyond apices of rostral setae, emerging from well-developed cup-shaped incisions, each flanked by two short sharp teeth of equal size; lamellar cusps extending well beyond rostrum (Fig. 7a). Interlamellar setae (in) on posteromedial margin of lamellae, at least four times length of setae le, extending well beyond rostrum and apices of setae le. Bothridia projecting laterally almost to inner margins of humeral processes, openings angled anterolaterally, posterior margin convex, lacking anterior condyle of enantiophysis H. Bothridial setae long, curved, bacilliform, smooth, pointed.</p><p>Notogaster: with thin granular cerotegument; ratio of length to breadth: 0.9; notogaster rounded, convex. Humeral processes narrow, with dorsal keel, waisted at their bases, expanded then pointed apically, extending anteriorly to point level with middle of lamella (Fig. 7a). Short, sharp lateral projection bearing lyrifissure ia on notogaster posterolateral of humeral process. With eight pairs of notogastral setae, l series, h 2, h 3 and p series present, l and h series in sub-marginal positions, flagelliform, smooth, very long (at least 250 μm); lm slightly shorter than others (Fig. 7c); bases of setae h 2 same distance apart as those of p 1; setae of p series much shorter than l and h series, p 1 longer than p 2 and p 3.</p><p>Ventral aspect: subcapitulum acute apically; mentum as long as broad; subcapitular setae short, smooth, setae m twice as long as a and h (Fig. 7b). Tutorium broad, right-angled. Epimeral plates discrete, all broadly separated in midline, sub-rectangular to trapezoid, plates II shorter than others; epimeral setation 3-1-3-2; setae 3a and 3b longer than others. Pedotectum I (pd I) well-developed, with large, conical anterior lobe and sub-rectangular posterior lobe; pd II small, acutely pointed; discidium with sharp point. Ventral plate broader than long. Genital and anal plates separated by distance about half length of genital plates; genital plates 173 µm long, with 6 pairs of setae, g 5 displaced laterally, g 1 longer than others; lyrifissure iad in para-anal position, some distance from margin of anal plate same as seta ad 3. Anal plates lozenge-shaped, 276 µm long; setae of the ad series very long; longer than setae p 2. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession nos. 53-1036; paratypes: one male, two females, ANIC accession no. 53-1037, moss on Sassafras ( Atherosperma moschatum), rainforest, 1,077 m., Errinundra Saddle, Errinundra National Park, Victoria, 37°19’S 148°51’E, 1,030 m., coll. M.J. Colloff, 29.ix.2009. Paratypes: two females, ANIC accession no. 53-1038, moss ( Dicranoloma billiardieri),  Nothofagus cunninghami rainforest, The Beeches, Lady Talbot Drive near Warburton, Yarra Ranges National Park, Victoria, 37°29’S, 145°50’E, 800 m., coll. G. Perdomo, May, 2009.</p><p>Etymology. The specific name is a noun in apposition. The species is named for the Bidhawal People of Gippsland and the Errinundra Plateau on whose traditional land this species was found.</p><p>Diagnosis.  Eutegaeus bidhawal can be distinguished from other species in the genus by the following combination of characters: (1) the absence of a translamella; (2) the prominent, sharp lateral projection on the notogaster; (3) the long, flagelliform interlamellar setae (4) the long, flagelliform setae of the l and h series; (5) the prominent, sub-rectangular pedotecta I with lobed apices; (6) the short, pointed pedotecta II; (7) relatively long setae p 1 but with setae p 2 and p 3 considerably shorter; (8) the very long adanal setae.</p><p>Remarks. This species shares with  E. soror (cf. below) the very long flagelliform interlamellar setae and notogastral setae of the l and h series. However, the l and h setae are shorter in  E. soror, which also has lamellae cusps with reticulate microsculpture, a V-shaped flange on the posterior margin of the bothridium, the bases of setae h 1 positioned further apart than those of setae p 1, the genital setae sub-equal in length, setae on epimeral plates III clustered medially instead of spaced along the length of the plate as in  E. bidhawal, with small tutoria and small pedotecta I with a short apical spur, rather than the broad, right-angled tutoria and prominent lobe-shaped pedotecta I of  E. bidhawal .</p></div>	https://treatment.plazi.org/id/03A2C77C4675FFDAC79CB6401671DFEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4677FFD4C79CB2C4162AD8CB.text	03A2C77C4677FFD4C79CB2C4162AD8CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus ptilosus Colloff 2023	<div><p>Eutegaeus ptilosus sp. nov.</p><p>(Figs. 8, 9)</p><p>Dimensions. Holotype length 1,074 μm, breadth 814 μm. Paratype females (n = 17): mean length 1,144 μm (range 1,101 –1,209 μm); mean breadth 844 μm (range 794–877 μm). Paratype males (n = 7): mean length 1016 μm (range 1,008 –1,023 μm); mean breadth 651 μm (range 605–690 μm). Paratype tritonymph length 893 μm, breadth 672 μm. Ratio of prodorsum to total length: 0.36 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) on short tubercles, setiform, curved, smooth, visible in dorsal aspect. Lamellae broad, separated medially, parallel, with reticulate microsculpture, lateral margins strongly convex, incurved basally, with a complete translamella. Lamellar setae (le) smooth, curved, extending well beyond apices of rostral setae, emerging from shallow cup-shaped incisions of lamellar cusp, each flanked laterally by short sharp tooth and medially by blunted projection; cusps extending well beyond rostrum, their medial margins convex (Fig. 8a). Interlamellar setae (in) very long, almost as long as bothridial setae, but not reaching to translamella, positioned medially of bases of lamellae. Bothridia projecting laterally beyond median margins of humeral processes, marked invagination anteriorly at base of lamella; with stepped, peg-like anterior projection; posterior bothridial margin with V-shaped anterior condyle of enantiophysis H. Bothridial setae long, straight, bacilliform, smooth, pointed.</p><p>Notogaster: with thin cerotegument covering sinuous, interdigitate microsculpture; notogaster rounded, convex, ratio of length to breadth: 0.84. Humeral processes very broad, without dorsal keel, waisted at bases, expanded then becoming pointed apically, extending to point level with middle of lamella (Fig. 8a). Short, broad projection on notogaster posterolateral of humeral process; lyrifissure ia not visible. With eight pairs of notogastral setae: l series, h 2 and h 3 present, short (sub-equal in length to setae in), slightly curved, smooth; setae la in centrodorsal position, lm in sub-marginal position, lp, h 2 and h 3 and p series marginal (Fig. 8a); setae p 1 almost as long as h 2 and h 3, closely-spaced and originating on squat tubercles. Setae p 2 and p 3 visible only in ventral aspect, one third of length of p 1. Lyrifissures im not visible.</p><p>Ventral aspect: subcapitulum broad apically; mentum same breadth as length; subcapitular setae smooth, setae h close to anterior margin of mentum, longer than a and m (Fig. 8b). Tutorium not visible in ventral aspect. Median margins of epimeral plates not evident, plates sub-rectangular to circular, plates IV almost circular, very large; epimeral setation 3-1-2-2; setae 3a and 3b markedly longer than others. Pedotectum I (pd I) well-developed, sub-rectangular, with median notch; pd II curved, horn-shaped; discidium small, with sharp point. Ventral plate broader than long. Genital and anal plates separated by distance about same as length of genital plates; genital plates 113 µm long, with six pairs of setae, g 5 displaced laterally, g 6 longer than others; lyrifissure iad in para-anal position, some distance from margin of anal plate, same as distance of bases of setae ad 3. Anal plates lozenge-shaped, 237 µm long. Pre-anal organ (po) T-shaped.</p><p>Description of tritonymph and nymphal scalps. Tritonymph prodorsum: rostrum acute; rostral setae curved, smooth, long, emerging from tubercles; lamellar setae well-developed, with three or four long, pectinate spines, emerging from tubercle on lamellar cusp; cusp ovoid, rounded apically, lacking prominent tooth or teeth (Fig. 9a). Lamellae smooth, strongly incised laterally posterior of setae le and again anterior of bothridia, with thin, amorphous cerotegument. Central region strongly reticulate, bounded by curved ridges between bases of interlamellar setae and anterior incision of lamella; setae in short, peg-like, on squat tubercles. Bothridia short, no longer than broad, projecting laterally, with rounded posterior flange; bothridial seta long (125 µm), curved, bacilliform, with pointed tip.</p><p>Tritonymph gastronotum: U-shaped, broader (712 µm) than long (634 µm). Gastronotal setae emerging from elongate, overlapping scales with long, horn-shaped lateral projections; scales smooth with median setal insertion canal (Fig. 9a). With 12 pairs of smooth setae: c, l, h and p series present, c 1 and c 2 shorter than l and h series, c 1 positioned medially on dorsosejugal scissure, c 2 emerging from tubercle on margin of scale bearing seta c 3; c 3, l and h series and p 1 long, straight, setiform, with rounded apex and three sharp tines (Fig. 9c); seta h 1 and associated scale smaller than other h and l series and their scales; both setae p 1 on single scale; setae p 2 and p 3 thin, setiform, smooth, positioned ventrally, posterolateral of anal plate, but not on scales (Fig. 9b). Cuticle with spines of cerotegument between scales.</p><p>Deutonymphal scalp: with eight pairs of setae: c 1 and c 3 (c 2 absent), l series and h 2, h 3 and p 1; c 3 thin, smooth, flagellate, emerging from short tubercles posterior of bothridium, tubercles with median apical spine; l series and h 2, h 3 and p 1 same morphology as tritonymphal setae but shorter; setae p 1 each on separate sub-rectangular scale (Fig. 9a). Cuticle with strongly reticulate microsculpture.</p><p>Protonymphal scalp: with six pairs of setae: l series, h 2, h 3 and p 1; setae of the c series and h 1 (and scale) absent. Cuticle with strongly reticulate microsculpture (Fig. 9a).</p><p>Tritonymph ventral aspect: ventral plate with fine striae with small tubercles of cerotegument; epimeral setal formula 3-1-2-2; four pairs of genital setae, one pair of aggenital setae, two pairs of anal setae and three pairs of adanal setae (Fig. 9b).</p><p>Ontogenetic formula of gastronotic setae (including two protonymphal and deutonymphal setae not present on scalps):?-(8,10,12)-8.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1039; paratypes: 12 females, one tritonymph, ANIC accession no. 53-1040, ANIC 754,  Nothofagus moorei and Sassafras ( Atherosperma moschatum) litter, rainforest, Cobark Forest Park, Barrington Tops, New South Wales, 31°59’S, 151°49’E, coll. T. Weir and A. Calder, 15.xi.1981. Paratypes: seven males, five females, ANIC accession no. 53-1041, ANIC 749, litter, closed  Eucalyptus forest, Barrington Tops National Park, Gloucester Road, 30°04’S, 151°41’E, coll. T. Weir and A. Calder, 12–14.ix.1981.</p><p>Etymology. The specific name is derived from the Greek, πτῐ́λον (ptilon), meaning a feather, referring to the feather-like lamellar setae of the tritonymph.</p><p>Diagnosis.  Eutegaeus ptilosus can be distinguished from other species in the genus by the following combination of characters: (1) the lamellar cusp with a lateral tooth and a blunt medial projection; (2) the complete translamella; (3) the elongated interlamellar setae; (4) the peg-like projection on the anterior of the bothridium; (5) the presence of the anterior condyle of enantiophysis H; (6) the pointed lateral projections posterior of the humeral processes; (7) the notogastral setae la in the centrodorsal position, lm in the sub-marginal position and the others in the marginal position; (8) setae p 1 about three times the length of p 2 and p 3, closely-spaced and on squat tubercles; (9) pedoctectum pd II curved, horn-shaped; (10) the large, sub-circular epimeral plates IV.</p><p>Remarks.  Eutegaeus ptilosus shares with  E. nothofagi,  E. bidhawal,  E. soror,  E. curviseta and  E. membraniger the lateral projections posterior of the humeral processes. The latter two species have setae p 1 on squat tubercles, but setae p 1 of  E. ptilosus are considerably longer. The squat tubercles, complete translamella, anterior condyle of enantiophysis H, notogastral setae la in the centrodorsal position and lm in the sub-marginal position place  E. ptilosus as morphologically closest to  E. curviseta . However,  E. curviseta has two teeth on the lamellar cusp, lamellar setae with spinose ornamentation and only five pairs of genital setae.</p><p>The epimeral setal formula of 3-1-2- 2 in the tritonymph and adult of  E. ptilosus is unusual. Seta 4b is absent as well as 3c.</p></div>	https://treatment.plazi.org/id/03A2C77C4677FFD4C79CB2C4162AD8CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4679FFD6C79CB41F15A7DA74.text	03A2C77C4679FFD6C79CB41F15A7DA74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus odontatus Colloff 2023	<div><p>Eutegaeus odontatus sp. nov.</p><p>(Fig. 10)</p><p>Dimensions. Holotype female length 843 μm, breadth 586 μm. Paratype females (n = 3): mean length 850 μm (range 837–861 μm); mean breadth 619 μm (range 608–639 μm). Paratype males: lengths 790, 805 μm, breadths 568, 576 μm. Ratio of prodorsum to total length: 0.36 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) setiform, curved, smooth, visible in dorsal aspect. Lamellae very broad apically, separated medially, with reticulate microsculpture, lateral margins conspicuously convex and heavily sclerotised, incurved basally, with partial translamella. Lamellar setae (le) very long, smooth, flagelliform, overlapping, emerging from cup-shaped incisions of broad lamellar cusp, each flanked laterally and medially by sharp tooth; cusps extending to same level as apex of rostrum, medial margins of lamellae concave; lateral margins heavily sclerotised; partial translamella present (Fig. 10a). Interlamellar setae (in) very long, setiform, smooth, extending beyond rostrum, positioned close together on central prodorsum on faint, inverted Vshaped ridge. Posterior prodorsum with anterior condyles of enantiophyses B. Bothridia projecting laterally beyond median margins of humeral processes, marked invagination at base of lamella; bothridia with peg-like anterior projection and bilobed anterior condyle of enantiophysis H on posterior margin. Bothridial seta short, extending just beyond lateral margin of humeral process, curved, with narrow, slightly expanded head with fine spines.</p><p>Notogaster: with thin cerotegument covering sinuous microsculpture (Fig. 10a); ratio of length to breadth: 0.87; notogaster rounded, convex. Humeral processes broad, without dorsal keel; waisted at their bases laterally, with a pronounced apical tooth, extending to point level with base of lamellar cusps. Short, sub-triangular projection on notogaster posterior of humeral process; lyrifissure ia not visible. With eight pairs of notogastral setae: l series, h 2 and h 3 curved, smooth relatively long: about same length as setae in; setae la and lm in centrodorsal position, all others sub-marginal (Fig. 10a); setae of p series short, sub-equal in length; p 1 on small flat tubercles. Lyrifissures im not visible.</p><p>Ventral aspect: subcapitulum broad; mentum about same breadth as length; subcapitular setae smooth, h sub-equal in length to a and m (Fig. 10b). Tutorium short, blunt, not visible in ventral aspect. Median margins of epimeral plates absent; plates sub-rectangular to sub-triangular, II and III about the same dimensions; epimeral setation 3-1-3- 2; seta 4c markedly longer than 4b. Pedotectum I (pd I) well-developed, sub-rectangular; pd II curved, horn-shaped; discidium triangular, with sharp point. Ventral plate broader than long, with slight invagination posterior of anal plate. Genital and anal plates separated by distance slightly less than length of genital plates; genital plates 94 µm long, with six pairs of setae, all aligned longitudinally, g 1 slightly longer than others; lyrifissure iad in para-anal position, close to margin of anal plate, same distance as its length. Anal plates lozenge-shaped, 171 µm long. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1042;  paratypes: three females, two males, ANIC accession no. 53-1043, ANIC 107, moist upland hardwood forest (cf. Invasive Species Council and TierraMar, 2021 for description), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.93333&amp;materialsCitation.latitude=-29.016666" title="Search Plazi for locations around (long 167.93333/lat -29.016666)">Mount Pitt</a>, Norfolk Island, 29°1’S, 167°56’E, 230 m, coll. M.S. Upton, 12.vii.1968.</p><p>Etymology. The specific name is derived from the Greek ὀδούς (odonts) meaning a tooth, referring to the tooth at the apex of the humeral process.</p><p>Diagnosis.  Eutegaeus odontatus can be distinguished from other species in the genus by the following combination of characters: (1) the long, thin, flagelliform lamellar setae; (2) the very broad lamellae, expanded apically; (3) the heavily sclerotised lateral margins of the lamellae; (4) the concave margins of the lamellar cusps; (5) the extremely long setae in; (6) the short bothridial seta with a slightly expanded head and fine spines; (7) the prominent bi-lobed anterior condyle of enantiophysis H; (8) the stepped, peg-like anterior projection of the bothridium; (9) the presence of anterior condyles of enantiophysis B; (10) the long notogastral setae of the l and h series, with la and lm located centrodorsally and lp and the h series located sub-marginally; (11) setae p 1 located on squat tubercles; (12) the genital setae all aligned longitudinally; (14) the invagination posterior of the anal plate.</p><p>Remarks.  Eutegaeus odontatus shares with  E. woiwurrung,  E. membraniger and  E. curviseta the short setae of the p series, with p 1 on squat tubercles, and with these species and  E. nothofagi the lamellar cusps with a pair of well-developed teeth. Like  E. nothofagi the species has the anterior condyles of enantiophysis B and, like this species,  E. woiwurrung and  E. soror, a prominent sub-triangular projection at the base of the humeral process, as well as notogastral setae la and lm in the centrodorsal position and lp and the h series in the sub-marginal position.  Eutegaeus odontatus and  E. woiwurrung both have a partial translamella but  E. odontatus differs from  E. woiwurrung and  E. nothofagi in that the lamellar setae, interlamellar setae, notogastral setae of the l and h series and setae 4c are much longer, the bothridial setae are much shorter, with slightly expanded heads covered with spinules and the humeral process lacks a dorsal keel.</p></div>	https://treatment.plazi.org/id/03A2C77C4679FFD6C79CB41F15A7DA74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C467BFFD0C79CB77C1470DEB0.text	03A2C77C467BFFD0C79CB77C1470DEB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeus soror P. Balogh 1985	<div><p>Eutegaeus soror P. Balogh, 1985</p><p>(Fig. 11)</p><p>Eutegaeus soror P. Balogh, 1985, p. 86 .</p><p>Dimensions. Holotype female length 1,178 μm, breadth 704 μm. Ratio of prodorsum to total length: 0.36 (holotype). Topotypes: females (n = 6): mean length 1,065 μm (range 1,027 –1,122 μm), mean length 774 μm (range 718–814 μm); males: length 940 μm and 963 μm, breadth 727 μm and 798 μm.</p><p>Redescription of adult. The holotype specimen of  Eutegaeus soror is badly damaged and missing the genital and anal plates as well as the notogastral and lamellar setae. Accordingly, the redescription below is supplemented by reference to topotypic material, collected within 18 km of the type locality from identical microhabitat (leaf litter) in the same vegetation community (sub-tropical rainforest).</p><p>Prodorsum: rostrum acute; rostral setae (ro) reflexed, smooth. Lamellae broad, separated medially, parallel, lateral margins conspicuously convex, translamella absent.Lamellar setae (le) smooth, curved, overlapping, emerging from well-developed cup-shaped incisions, each flanked by long sharp teeth of equal size; lamellar cusps extending anteriorly beyond rostrum (Fig. 11a). Interlamellar setae (in) at least twice as long as setae le, recurved; marked invagination laterally at base of lamella anterior of bothridium. Bothridia projecting laterally to inner margins of humeral processes, openings angled anterolaterally, posterior margin with anterior condyles of enantiophyses H in form of prominent V-shaped flange; anterior margin with rounded protuberance. Bothridial setae long, setiform, smooth.</p><p>Notogaster: with thin cerotegument; ratio of length to breadth: 0.81; notogaster rounded, convex. Humeral processes broad, without dorsal keel; waisted at their bases, then expanding and sharply pointed apically, extending to point level with middle of lamella (Fig. 11a). Short, sharp lateral projection on notogaster posterolateral of humeral process. With eight pairs of long, flagelliform, smooth notogastral setae, l series, h 2, h 3 and p series present: la and lm in centrodorsal position, lp and h series in sub-marginal position; h 2 longer than others, which are sub-equal in length; p 1 markedly longer than others in p series; notogastral margin with irregular flat tubercles.</p><p>Ventral aspect: subcapitulum acute apically; mentum as long as broad; subcapitular setae smooth, setae m and h each twice as long as a (Fig. 11b). Tutorium short, blunt, not well-developed, barely visible in ventral aspect. Epimeral plates discrete except IV, all broadly separated in midline, sub-rectangular, plates II slightly shorter than others; epimeral setation 3-1-3-2; setae on plates III clustered. Pedotectum I (pd I) slight, sub-rectangular in lateral outline, with short blunt spur apically; pd II small, pointed; discidium triangular with sharp point. Ventral plate broader than long. Genital and anal plates separated by distance about half length of genital plates; genital plates 148 µm long, with six pairs of setae sub-equal in length, g 4 displaced laterally; lyrifissure iad in para-anal position, some distance from anal plate, more than its length. Anal plates lozenge-shaped, 272 µm long; setae of the ad series very long, at least twice length of setae p 2. Pre-anal organ (po) T-shaped.</p><p>Material examined and locality data.   Holotype female, ANIC accession no. 53-638, leaf litter, rainforest, Lynchs Creek, Airdrop Road, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.05&amp;materialsCitation.latitude=-28.383333" title="Search Plazi for locations around (long 153.05/lat -28.383333)">Wiangaree State Forest</a>, New South Wales, 28°23’S, 153°03’E, 820 m, coll. V. J. Pattemore  .   Topotypes: six females, two males, ANIC 460, leaf litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m  ., coll. R. J. Kohout, 21.iii.1973 .</p><p>Diagnosis.  Eutegaeus soror can be distinguished from other species in the genus by the following combination of characters: (1) the absence of a translamella; (2) the presence of the anterior condyle of enantiophysis H; (3) the short tubercle-like projection on the anterior margin of the bothridium; (4) the sharp lateral projection on the notogaster posterolateral of the humeral process; (5) the extremely long, flagelliform interlamellar setae and notogastral setae of the l and h series; (6) the relatively small pedotectum I with a short apical spur; (7) the short, pointed pedotectum II; (8) the three setae on epimeral plate III clustered in a group on the medial part of the plate; (9) the relatively long setae p 1 but with setae p 2 and p 3 considerably shorter; (10) the very long adanal setae, at least twice as long as p 2 and p 3.</p><p>Remarks.  Eutegaeus soror is morphologically most similar to  E. bidhawal, as detailed above. P. Balogh (1985) considered  E. soror belongs to a species group containing  E. stylesi and  E. radiatus from New Zealand, characterised by very long setae p 1, very short setae p 2 and p 3 and deeply incised lamellar cusps.  Eutegaeus pinnatus from New Zealand, also has these character states. However, setae p 1 in the three New Zealand species are on short tubercles whereas such tubercles are absent in  E. bidhawal and  E. soror .</p></div>	https://treatment.plazi.org/id/03A2C77C467BFFD0C79CB77C1470DEB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C467DFFD0C79CB20512BDD923.text	03A2C77C467DFFD0C79CB20512BDD923.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neseutegaeus Woolley 1965	<div><p>Neseutegaeus Woolley, 1965</p><p>Neseutegaeus Woolley, 1965, p. 307 .</p><p>Neseutegaeus Woolley, 1965: Luxton, 1988a, p. 77.</p><p>Type species:  Neseutegaeus spinatus Woolley, 1965 .</p><p>Diagnosis. The following diagnosis is modified from those of Woolley (1965) and Luxton (1988a): small to medium-sized oribatid mites (330–550 μm); rostrum rounded; lamellar cusps about one-third length of lamellae, rounded or obtuse apically, not projecting markedly beyond rostrum, with long, narrow, incurved medial tooth, lamellae separated from each other and translamella complete, incomplete or absent; lamellar seta emerging dorsally from lateral margin of lamellar cusp. Interlamellar setae short, not overlapping; bothridial setae long, club-shaped apically. Posterior margin of bothridium with anterior condyles of enantiophyses H. Humeral process long, slim, waisted basally, originating on anterolateral margin of notogaster, reaching to point half-way along length of lamella; anterolateral margin of notogaster conspicuously incised at base of humeral process, but lacking Vshaped or rounded projection. Notogaster U-shaped; with eight pairs of thick, short setae, no longer than distance between them, those of l series in centrodorsal position; h series centrodorsal or sub-marginal; setae of l and h series differing in shape in some species: those of l series sharp, spiniform, those of h series blunt, club-shaped; setae of p series minute. Pedotectum I (pd I) sub-rectangular in ventral view, pd II rectangular, discidium triangular, pointed. Margins of epimeral plates IV incomplete. Five or six pairs of genital setae; three pairs of adanal setae. Perigenital carina and enantiophysis E4 absent. Chelicerae chelate-dentate, of normal proportions. Pre-anal organ oval.</p><p>Remarks.  Neseutegaeus spinatus Woolley, 1965 from Alex Knob Track, Westland, New Zealand was given a supplementary description by Hammer (1966, p. 72) based on non-type material from Fox Glacier and Milford. Woolley (1965) stated there are six pairs of genital setae, but Hammer reported five, present as alveoli: the setae could not be seen. Additional character states observed by Hammer (1966, Plate 32, Fig. 98c) of relevance to the generic definition include the lozenge-shaped anal plates; the circumpedal carinae (cc); the epimeral setal formula 3-1-3-2; the oval pre-anal organ and the morphology of the epimeral plates, with plates III smaller than others; lyrifissures iad in the para-anal position but not close to the anal plates and pedotectum I well-developed with margins semi-rectangular in outline.</p><p>Neseutegaeus denticulatus P. Balogh (1988) was described from Sri Lanka. But the species is not a  Neseutegaeus as the lamellae are fused anteriorly, not free and with a medially-directed tooth as is characteristic for the genus. Pedotecta I and II are large, broad and pointed which, if confirmed, is unique within  Eutegaeoidea . Coxisternal structures are uncertain from the description, including the morphology of the discidium, epimeral setal formula and the presence of enantiophyses E4 and perigenital carinae (P. Balogh, 1988, Figs. 16-18 therein). The species cannot be assigned to an existing genus within  Eutegaeoidea, but requires re-description before consideration whether the establishment of a new genus is warranted.</p><p>The redefinition of  Neseutegaeus above includes the four species described by Hammer (1966) ( N. angustus,  N. consimilis,  N. distendus and  N. latus) and  N. spinatus, all from New Zealand.  Neseutegaeus monteithi J. and P. Balogh, 1983b was described from Australia, but lacks the characteristic, thin elongated incurved tooth emerging from the centre of the apically transverse or obtuse lamellar cusp of the other species. Also, the interlamellar setae are long and recurved; the notogastral setae are not spiniform or club-shaped; there is a conspicuous pair of rounded projections at the base of the humeral process and the base of the process is not sharply incised.  Neseutegaeus monteithi has a lamellar cusp with a well-developed apical tooth and the lamellar setae emerge sub-apically from the dorsal surface of the lateral margin of the cusp and very long, recurved interlamellar setae, recurved setae of the l series and a thin, narrow humeral process, typical of  Atalotegaeus . On the basis of these character states,  N. monteithi is recombined as  Atalotegaeus monteithi (J. and P. Balogh, 1983b) comb. nov. and is redescribed below.</p></div>	https://treatment.plazi.org/id/03A2C77C467DFFD0C79CB20512BDD923	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C467CFFD2C79CB38815E6DA74.text	03A2C77C467CFFD2C79CB38815E6DA74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neseutegaeus wardi Colloff 2023	<div><p>Neseutegaeus wardi sp. nov.</p><p>(Fig. 12)</p><p>Dimensions. Holotype male length 306 μm, breadth 195 μm. Paratype females (n = 5): mean length 333 μm (range 331–337); mean breadth 222 μm (range 217–226). Paratype males (n = 4): mean length 313 μm (range 305–323); mean breadth 204 μm (range 195–212). Ratio of prodorsum to total length: 0.41 (holotype).</p><p>Description of adult. Prodorsum: rostrum rounded; rostral setae (ro) short, curved, smooth, visible in dorsal aspect. Lamellae broad, extending beyond lateral margin of prodorsum, separated medially, lateral margins convex, without translamella (Fig. 12a). Lamellar seta (le) long, smooth, incurved, emerging from apex of lamellar cusp just anterior of long, thin, medially-directed lamellar tooth. Interlamellar seta (in) thin, relatively short, flagelliform with thick cylinder of cerotegument basally, emerging from lateral margin of inverted V-shaped prodorsal ridge. Bothridia prominent, overlapping laterally with inner margins of humeral processes, openings angled anterolaterally, anterior margin with squat tubercle, posterior margin with anterior condyle of enantiophysis H in form of V-shaped projection. Bothridial seta long, recurved, apex slightly expanded with fine spines. Curved ridge with wavy margin on prodorsum between bothridia.</p><p>Notogaster: ratio of length to breadth: 0.89; notogaster U-shaped, convex. Humeral processes narrow, with few small scattered tubercles, lateral margin convex, medial margin concave, with V-shaped basal indentation laterally and slight notch medially, apex pointed extending anteriorly to point level with middle of lamella (Fig. 14a). With eight pairs of notogastral setae: l, h 2 and h 3 and p series present, setae of l and h series flat, leaf-shaped, slightly curved, with pectinate margins (Fig. 12c), in centrodorsal position; p series short, smooth, curved, visible in dorsal view.</p><p>Ventral aspect: subcapitulum acute apically; subcapitular setae short, smooth, seta a twice as long as m and h (Fig. 12b). Chelicerae chelate-dentate, of normal proportions. Tutorium not visible in ventral aspect. Epimeral plates I-III discrete, separated in midline, sub-rectangular to sub-triangular, plates I with a small ovoid depression between them, plates IV lacking posterior margin; epimeral setation 2-1-3-2; setae 4b slightly longer than others. Pedotectum I (pd I) sub-rectangular in lateral outline; pd II small, blunt, sub-rectangular, not projecting beyond lateral margin of pd I; discidium well-developed, blunt, pointed. Ventral plate U-shaped, broader than long, with cluster of short tubercles near circumpedal carina. Genital and anal plates separated by distance about same as length of genital plates; genital plates 35 µm long, with six pairs of setae aligned longitudinally, g 1 longer than others, distance between g 5 and g 6 markedly greater than between g 4 and g 5; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 49 µm long; setae of the ad series very short. Pre-anal organ (po) oval.</p><p>Type designation, material examined and locality data.   Holotype male, ANIC accession no. 53-1044, ANIC 578, moss forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=164.76666&amp;materialsCitation.latitude=-20.566668" title="Search Plazi for locations around (long 164.76666/lat -20.566668)">Mount Panie</a> [Mont Panié], New Caledonia, 20°34'S, 164°46'E, 750 m., coll. P.S. Ward, 14.ii.1977  .   Paratypes: seven females, five males, ANIC accession no. 53-1045, ANIC 580, litter, rainforest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.51666&amp;materialsCitation.latitude=-23.183332" title="Search Plazi for locations around (long 166.51666/lat -23.183332)">Mount Koghis</a> [Mont Koghi], New Caledonia, 23°11'S, 166°31'E, 749 m  . [summit], coll. P.S. Ward, 18.ii.1977.</p><p>Etymology. This species is named in honour of its collector, Professor Philip S. Ward, Department of Entomology and Nematology, University of California, Davis.</p><p>Diagnosis.  Neseutegaeus wardi can be distinguished from the other species in the genus by the following combination of characters: (1) the prominent bothridia, extending well beyond the lateral margin of the prodorsum; (2) the curved ridge between the bothridia; (3) the scattered tubercles on the humeral process; (4) the flattened, pectinate setae of the l and h series; (5) the epimeral setal formula 2-1-3-2; (6) epimeres I with a small ovoid depression between them; (7) the marked gap between genital setae g 5 and g 6 compared with the other genital setae; (8) the cluster of small tubercles near the circumpedal carina.</p><p>Remarks.  Neseutegaeus wardi is not particularly morphologically similar to other species of  Neseutegaeus . The setae of the l and h series are leaf-like, pectinate, and in the centrodorsal position, like those of  N. spinatus Woolley, 1965 (cf. Hammer, 1966, Fig. 98b therein), though the spines on the setae of  N. wardi are much longer. Unlike  N. wardi,  N. spinatus has heavily striated lamellae, a partial translamella and the interlamellar setae are setiform, not flagelliform.  Neseutegaeus wardi differs from  N. consimilis and  N. distentus in that their notogastral setae are thin, curved and smooth. The new species differs from  N. angustus and  N. latus in that these species have notogastral setae of the l series that differ in morphology from the h series.  Neseutegaeus wardi is the first species in the genus to be recorded outside New Zealand.</p></div>	https://treatment.plazi.org/id/03A2C77C467CFFD2C79CB38815E6DA74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C467FFFD3C79CB77D17C3DA74.text	03A2C77C467FFFD3C79CB77D17C3DA74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atalotegaeus Luxton 1988	<div><p>Atalotegaeus Luxton, 1988</p><p>Atalotegaeus Luxton, 1988a . p. 82.</p><p>Type species:  Eutegaeus mensarosi J. and P. Balogh, 1983a, p. 83.</p><p>Diagnosis. the following diagnosis is modified from that of Luxton (1988a) and includes new character states for immatures. Adults: medium sized oribatid mites (600–700 μm); rostrum broadly conical; lamellae delicate and transparent, lamellar cusps about one-third of lamellar length, extending just beyond rostrum, concave medially, with single tooth apically, lamellar setae typically positioned sub-apically (apically in  A. monteithi comb. nov.); lamellae separate medially, translamella incomplete or absent, lateral margin strongly convex, covering lateral margins of prodorsum. Interlamellar setae long, overlapping, recurved or flagelliform. Posterior margin of prodorsum generally lacking condyles, though anterior condyle of enantiophysis H present in  A. monteithi; humeral processes long, narrow and delicate, originating on anterior margin of notogaster, reaching to point level with middle of lamella, extremely narrow and waisted basally. With rounded or pointed projection at base of humeral process. Seven pairs of notogastral setae (lm absent) or eight pairs; l series in centrodorsal position, recurved. With five or six pairs of genital setae, penultimate pair displaced laterally. Perigenital carina and enantiophyses E4 absent. Pre-anal organ T-shaped. Chelicerae chelate-dentate and of normal proportions or narrow and elongated. Immatures: margin of prodorsum with two pairs of incisions between bothridia and apex of lamellae. Gastronotal setae emerging from tubercles, not scales, except tritonymphal seta c 3 emerging from a flat extension of the humeral region that is rounded anteriorly and pointed posteriorly; deutonymphal and protonymphal seta c 3 emerging from tubercle. Tritonymph with transverse groove posterior of dorsosejugal scissure.</p><p>Remarks. J. and P. Balogh (1983a) described  Eutegaeus mensarosi from Australia which was recombined with the newly-established genus,  Atalotegaeus, by Luxton (1988a) who differentiated it from other genera of  Eutegaeidae by its ‘peloptoid’ (or pelopsiform) chelicerae and narrow subcapitulum. Some  Atalotegaeus spp. have chelate-dentate chelicerae of normal proportions (e.g.  A. crobylus sp. nov.), whereas others have modified, slim chelicerae (cf. below) but they are not truly pelopsiform in that the basal part is not considerably broader than the rest of the chelicera as in  Eupelops acromios (Hermann, 1804) (cf. Grandjean, 1936, Fig. 11b therein). These chelicerae are in fact more like those of  Allosuctobelba and other  Suctobelbidae (cf. Grandjean, 1951): slim and tapering, or ‘attenuate-edentate’ (Norton and Behan-Pelletier (2009, Fig. 15.8.K therein). The subcapitulum is unlike that of the peloptoid form found in the  Phenopelopidae in which the rutella are considerably narrowed apically, lack teeth, and the genae are fused with the mentum and, hence, lack the labiogenal articulation. Some  Atalotegaeus spp. have a long, narrow subcapitulum and a slim, but toothed, rutellum (Fig. 15b) but the labiogenal articulation is present and the apices of the genae are toothed. For  A. mensarosi, J. and P. Balogh (1983a, p. 82) stated ‘mouth parts suctorial’ but without specifying which parts. Accordingly, I consider the character states of the mouthparts alone are insufficient to differentiate  Atalotegaeus from other  Eutegaeidae . Rather, the thin, narrow humeral processes (Luxton, 1988a, p. 82 described them as ‘delicate and transparent’), expanded lamella with a long, incurved tooth and the sub-apical emergence of the lamellar setae are more consistent character states for the adults. Also, there are morphological differences between immatures of  Eutegaeus and  Atalotegaeus, as detailed in the generic definitions above, most notably the tritonymphal seta c 3 of  Atalotegaeus emerging from a flat, humeral extension, pointed posteriorly, and from a tubercle in the deutonymph and protonymph, whereas in  Eutegaeus c 3 emerges from a scale in all the nymphal stages. These character states are consistent among immatures of the two genera, as detailed below.</p><p>Ermilov (2021) described adults and immatures of  Eutegaeus aysenensis and  E. queulatensis from Chile. The adults have an incurved lamellar tooth with a sub-apical lamellar seta and thin humeral processes, typical of species of  Atalotegaeus . The notogastral setae of the nymphs emerge from tubercles, as in the tritonymph of  A. mensarosi (cf. below) and lack elongate, hemi-semicircular scales found in immatures of all  Eutegaeus spp. described to date ( E. woiwurrung,  E. nothofagi,  E. parapapuensis Ermilov, 2020 and  E. paralagrecai Ermilov, 2020). Accordingly,  Eutegaeus aysenensis and  E. queulatensis are hereby recombined as  Atalotegaeus aysenensis (Ermilov, 2021) comb. nov. and  Atalotegaeus queulatensis (Ermilov, 2021) comb. nov. These, with the new species described below and the recombination of  N. monteithi to  Atalotegaeus, increases the total number of described species of  Atalotegaeus to five.</p></div>	https://treatment.plazi.org/id/03A2C77C467FFFD3C79CB77D17C3DA74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C467EFFECC79CB77D15A0D96E.text	03A2C77C467EFFECC79CB77D15A0D96E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atalotegaeus crobylus Colloff 2023	<div><p>Atalotegaeus crobylus sp. nov.</p><p>(Fig. 13)</p><p>Dimensions. Holotype female length 545 μm, breadth 378 μm. Paratype female length 540 μm, breadth 328 μm. Ratio of prodorsum to total length: 0.45 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) short, curved, smooth, visible in dorsal aspect. Lamellae broad, extending beyond lateral margin of prodorsum, separated medially, lateral margins convex, with an incomplete translamella (Fig. 13a). Lamellar seta (le) smooth, recurved, very short, emerging in sub-apical position on median edge of lamella; lamellar cusp with single tooth, extending just beyond rostrum. Interlamellar setae (in) long, smooth, strongly curved in almost circular configuration, emerging on posteromedian prodorsum. Bothridia relatively short, projecting as far as inner margins of humeral processes, openings angled anterolaterally, anterior margin with rounded, convex ledge. Bothridial setae long, curved, bacilliform, smooth, slightly expanded apically.</p><p>Notogaster: ratio of length to breadth: 0.79; notogaster rounded, convex. Humeral processes extending to point level with middle of lamella, thin, fine, waisted at base, with V-shaped projection pointing posterolaterally, then expanded and with fine, pointed apex, lateral margin slightly concave (Fig. 13a). With eight pairs of notogastral setae, l, h 2 and h 3 and p series present, setae la and lm smooth, strongly curved in almost circular configuration, in centrodorsal position, setae h 2 and h 3 straight, smooth, in sub-marginal position, short (65 μm); p series very short (15 μm).</p><p>Ventral aspect: subcapitulum acute apically; subcapitular setae short, smooth, setae a half length of m and h (Fig. 13b). Chelicerae chelate-dentate, of normal proportions. Tutorium short, blunt. Epimeral plates discrete, broadly separated in midline, sub-rectangular to sub-triangular, plates IV shorter than others; epimeral setation 3-1- 3-2; setae 3b and 3c longer than others. Pedotectum I (pd I) small, narrow; pd II small, with blunt point; discidium acutely pointed. Ventral plate ovoid, broader than long. Genital and anal plates close together at distance of less than half length of genital plate; genital plates 66 µm long, with five pairs of setae sub-equal in length, g 4 displaced laterally; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 98 µm long; setae of the ad series short, similar length to setae of the p series. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.  Holotype female ANIC accession no. 53-1047,  paratype female, ANIC accession no. 53-1048, ANIC 297, moss and litter,  Nothofagus cunninghami rainforest, Cumberland Creek Valley, near Picnic Area [Cumberland Memorial Scenic Reserve], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Yarra Ranges National Park</a>, Victoria, 37°34’S 145°52’E, 920 m., coll. R. W. Taylor and R. J. Bartell, 4.xi.1970.</p><p>Etymology. The prefix of the specific name, crobyl-, is derived from the Greek κρώβυλος (krobylos) meaning a curl, referring to the shape of the interlamellar setae.</p><p>Diagnosis.  Atalotegaeus crobylus can be distinguished from the other species in the genus by the following combination of characters: (1) the extremely short, thin rostral setae; (2) the mentum much broader than long; (3) the presence of an incomplete translamella; (4) the long, recurved and overlapping setae in; (5) the relatively short, broad bothridia with a markedly convex, rounded anterior border; (6) the apices of the humeral processes are flexed outwards and their lateral margins concave apically; (7) notogastral setae la and lm are strongly recurved; (8) the minute setae of the p series, sub-equal in length to the anal setae; (9) the small, blunt pedotectum II.</p><p>Remarks. This species is morphologically most similar to  A. mensarosi (cf. below), but differs in the outwardcurving humeral processes and strongly recurved setae la and lm and in lacking the reticulate microsculpture on the lamellae, the prominent pedotecta I and the extremely long, flagelliform setae in. Similarities between  A. crobylus and  A. deficiens sp. nov. are detailed below.</p></div>	https://treatment.plazi.org/id/03A2C77C467EFFECC79CB77D15A0D96E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4641FFEEC79CB44A16F4DC7C.text	03A2C77C4641FFEEC79CB44A16F4DC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atalotegaeus deficiens Colloff 2023	<div><p>Atalotegaeus deficiens sp. nov.</p><p>(Fig. 14)</p><p>Dimensions. Holotype female length 495 μm, breadth 343 μm. Paratype female length 503 μm, breadth 356 μm. Ratio of prodorsum to total length: 0.39 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute with small inverted V-shaped area of lighter cuticle; rostral setae (ro) short, curved, smooth, visible in dorsal aspect. Lamellae broad, extending well beyond lateral margin of prodorsum, separated medially, without translamella, lateral margins strongly convex (Fig. 14a). Lamellar seta (le) well-developed, with spinose ornamentation, curved, emerging sub-apically on median edge of lamella; lamellar cusp with single apical tooth, extending just beyond rostrum. Interlamellar setae (in) long, smooth, curved, on short inverted V-shaped ridge. Bothridia prominent, projecting laterally to inner margins of humeral processes, openings angled anterolaterally, posterior margin with anterior condyle of enantiophysis H in form of V-shaped projection. Bothridial setae long, curved, setiform, smooth.</p><p>Notogaster: ratio of length to breadth: 0.87; notogaster rounded, convex. Humeral processes thin, fine, waisted at base, extending anterior of apices of setae in, with two blunt lateral protuberances, apex pointed, lateral margin slightly concave (Fig. 14a). With seven pairs of notogastral setae: la, lp, h 2, h 3 and p series present, setae of l and h series with spinose ornamentation on one side, strongly curved, in sub-marginal position, p series very short, smooth, visible in dorsal view.</p><p>Ventral aspect: subcapitulum acute apically; subcapitular setae short, smooth, sub-equal in length (Fig. 14b). Chelicerae chelate-dentate and of normal proportions. Tutorium prominent, broad, with blunted apex. Epimeral plates I discrete, broadly separated in midline, sub-triangular, plates II only partially divided in midline, plates III with no median division, plates IV lacking posterior margin; epimeral setation 3-1-3-2; setae sub-equal in length. Pedotectum I (pd I) not prominent, sub-rectangular laterally in ventral view; pd II triangular, sharply pointed; discidium broad, pointed, with blunt apex. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of half length of genital plate; genital plates 65 µm long, with six pairs of setae, sub-equal in length, g 4 displaced laterally; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 87 µm long; setae of the ad series short, similar length to setae of the p series. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.   Holotype female, ANIC accession no. 53-1046, ANIC 580, litter, rainforest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.51666&amp;materialsCitation.latitude=-23.183332" title="Search Plazi for locations around (long 166.51666/lat -23.183332)">Mount Koghis</a> [Mont Koghi], New Caledonia, 23°11'S, 166°31'E, 749 m. [summit], coll. P.S. Ward, 18.ii.1977.</p><p>Etymology. The specific name,  deficiens, refers to the absence of notogastral seta lm.</p><p>Diagnosis.  Atalotegaeus deficiens can be distinguished from other species in the genus by the following combination of characters: (1) the small inverted V-shaped area of lighter cuticle on the rostrum; (2) the lamellar setae with spinose ornamentation; (3) the inverted V-shaped ridge on the prodorsum from which setae in emerge; (4) the long, smooth, curved setae in; (5) the two blunt protuberances at the base of the humeral process; (6) the absence of setae lm, thus seven pairs of notogastral setae; (7) setae of l and h series with spinose ornamentation on one side; (8) the prominent broad tutorium with a blunted apex.</p><p>Remarks. This species is morphologically most similar to  A. crobylus (cf. above) in that both have similar morphology of the lamellar cusp and chelate-dentate chelicerae of normal proportions, but  A. deficiens differs in having lamellar setae with spiniform ornamentation, the presence of the inverted V-shaped ridge on the prodorsum and the absence of setae lm.</p></div>	https://treatment.plazi.org/id/03A2C77C4641FFEEC79CB44A16F4DC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4643FFE8C79CB175165FDBB3.text	03A2C77C4643FFE8C79CB175165FDBB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atalotegaeus mensarosi (Balogh & P. Balogh 1983)	<div><p>Atalotegaeus mensarosi (J. &amp; P. Balogh, 1983)</p><p>(Figs. 15, 16)</p><p>Eutegaeus mensarosi J. &amp; P. Balogh, 1983a, p. 82 .</p><p>Atalotegaeus mensarosi (J. &amp; P. Balogh, 1983): Luxton, 1988a, p. 83.</p><p>Dimensions. Holotype male: length 577 μm, breadth 423 μm. Ratio of prodorsum to total length: 0.43. Non-type material: females (n = 5) mean length 611 (range 596–632) mean breadth 417 (range 395–435). Tritonymph: length 530 μm, breadth 394 μm.</p><p>Redescription of adult. Prodorsum: rostrum markedly acute; rostral setae (ro) long, recurved, smooth, visible in dorsal aspect. Lamellae broad, separated medially, lateral margins convex, extending beyond lateral margin of prodorsum, with microsculpture of scale-like plaques; translamella absent (Fig. 15a). Lamellar seta (le) smooth, long, flagelliform, emerging sub-apically on lateral edge of lamella; lamellar cusp with single tooth, extending just beyond rostrum. Interlamellar setae (in) extremely long (258 μm), smooth, flagelliform, emerging on posteromedian of prodorsum. Bothridia relatively short, angled laterally, projecting at least to inner margin of humeral processes, openings with small apical flange on lip. Bothridial setae very long, curved, bacilliform, smooth.</p><p>Notogaster: ratio of length to breadth: 0.82; notogaster rounded, convex. Humeral processes extending anteriorly to point level with third of lamellar length, narrow, waisted at base then expanded, with pointed apex, convex lateral margin, V-shaped projection at base (Fig. 15a). With eight pairs of notogastral setae; l, h 2 and h 3 and p series present, setae of l and h series long, curved, smooth, setae la and lm in centrodorsal position, others sub-marginal, setae lp markedly longer than others of l and h series (240 μm); p series markedly shorter than l and h series (47 μm).</p><p>Ventral aspect: subcapitulum narrow, acute apically, mentum much longer than broad; subcapitular setae short, smooth, sub-equal in length (Fig. 15b). Long, thin rutellum protruding beyond rostrum (Fig. 15c). Chelicerae thin, narrow but chelate-dentate apically, protruding beyond apex of subcapitulum. Tutorium short, rounded apically, not visible in ventral aspect. Epimeral plates discrete, all broadly separated in midline, sub-rectangular to sub-triangular, plates IV shorter than others; epimeral setation 3-1-3-2; setae 3c and 4b longer than others. Pedotectum I (pd I) well-developed, curved, pointed; pd II small, pointed; discidium sub-rectangular, bicornute. Ventral plate broader than long. Genital and anal plates close together; genital plates 70 µm long, with six pairs of setae, sub-equal in length, g 4 displaced laterally; lyrifissure iad in para-anal position, some distance from anal plate, more than length of iad. Anal plates lozenge-shaped, 98 µm long; setae of the ad series long, setiform, thin, similar length to setae p 1. Pre-anal organ (po) T-shaped.</p><p>Description of tritonymph, nymphal and larval scalps. Tritonymph prodorsum: with well-developed cerotegument of small columnar cells along prodorsal margins (Fig. 16a). Rostrum acute; rostral setae smooth, on short tubercles, extending beyond lamellar cusps; lamellar setae long, curved medially, with plumose cerotegument, emerging from short tubercle at base of lateral margin of lamellar cusp; cusps with small scattered tubercles, each cusp with prominent incurved tooth almost meeting opposing tooth. Lateral margin of lamella incised transversely posterior of seta le and again anterior of bothridium; central prodorsum with alveolate microsculpture. Setae in with leaf-shaped cerotegument, emerging from short tubercles. Bothridia long, cup-shaped, constricted basally, with prominent apical flange; bothridial seta long (210 µm), thin, straight, slightly expanded apically, with pointed tip.</p><p>Tritonymph gastronotum: U-shaped, broader (394 µm) than long (362 µm). Anterolateral margin with humeral process rounded anteriorly and bearing seta c 3, then incised posteriorly. Gastronotal setae emerging from well-developed tubercles (Fig. 16a). With 12 pairs of setae: full complement of c, l, h and p series present; c 3, l and h series and p 1 with broad, paddle-shaped cerotegumental covering, each with few spines basally; seta c 1 setiform, smooth; c</p><p>2 short, spinose.</p><p>Deutonymphal and protonymphal scalp: with eight pairs of setae: c 1 (c 2 and c 3 absent), l series and h 2, h 3 and p 1, all on short tubercles except c 1; deutonymphal scalp (Fig. 16a)</p><p>Larval scalp: with alveolate microsculpture; eight pairs of setae with leaf-like cerotegument, on short tubercles: l series, h 2 and h 3, p 1 and centrodorsal series d present. Setae dp on small ovoid sclerite (Fig. 16a).</p><p>Tritonymph ventral aspect: microsculpture of ventral plate with fine striations (Fig. 16b); anal plate with fine tubercles; epimeral setal formula 3-1-3-3; with four pairs of genital setae, one pair of aggenital setae, two pairs of anal setae and three pairs of adanal setae.</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps): 9-(10,10,12)-8.</p><p>Material examined and locality data.   Holotype, ANIC accession no. 53-637, litter,  Nothofagus moorei rainforest, Point Lookout, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.41667&amp;materialsCitation.latitude=-30.483334" title="Search Plazi for locations around (long 152.41667/lat -30.483334)">Ebor</a>, New South Wales, 30°29’S, 152°25’E, 1525 m, coll. I. Naumann, 25.iv.1973  .   Non-type material: five females, ANIC 460, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. R. J. Kohout, 21.iii.1973  .</p><p>Diagnosis.  Atalotegaeus mensarosi can be distinguished from the other species in the genus by the following combination of characters: (1) the elongated mentum; (2) the slim, fine chelicerae; (3) the lamellae with reticulate microsculpture; (4) the very long interlamellar setae (5) the long adanal setae; (6) the setae p 1 markedly longer than setae p 2 and p 3; (7) the very long epimeral setae 3c and 4c and (8) the small apical flange on the bothridium.</p><p>Remarks.  Atalotegaeus mensarosi is morphologically most similar to  A. crobylus, as detailed above.The nymphs of this species lack the intricate scales associated with the lateral gastronotal setae of the nymphs of  Eutegaeus spp. described herein. The association of the tritonymph with the adult is based, in part, on their presence in the same samples, but also the presence of the long, pointed, incurved lamellar cusp with the lamellar seta positioned sub-apically, similar to the morphology of the lamellar cusp in the adult.</p><p>The tritonymph of this species has the epimeral setal formula 3-1-3-3, but in the adult, it is 3-1-3-2 (as in  A. crobylus and  A. deficiens), with seta 4b absent in the adult. The seta is present in the adults of the other species of  Atalotegaeus . The loss of 4b in the adult also occurs in some species of  Eutegaeus (cf. remarks section for  E. woiwurrung above).</p></div>	https://treatment.plazi.org/id/03A2C77C4643FFE8C79CB175165FDBB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4645FFEAC79CB537130DDF78.text	03A2C77C4645FFEAC79CB537130DDF78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atalotegaeus monteithi (Colloff 2023) Colloff 2023	<div><p>Atalotegaeus monteithi (J. &amp; P. Balogh, 1983) comb. nov.</p><p>(Fig. 17)</p><p>Neseutegaeus monteithi J. &amp; P. Balogh, 1983b, p. 292.</p><p>Dimensions. Holotype female 469 μm, breadth 365 μm.</p><p>Redescription of adult. Prodorsum: rostrum acute; rostral setae (ro) short, straight, smooth, visible in dorsal aspect. Lamellae broad, smooth, separated medially, not extending as far as apex of rostrum, lateral margins convex, translamella incomplete (Fig. 17a); lamellar seta (le) very short, fine, flagelliform, emerging from shallow excavation on apex of cusp, with sharp tooth medially. Interlamellar setae (in) long, smooth, recurved, on posteromedian margins of lamellae. Bothridia broad, with anterior condyle of enantiophysis H in form of V-shaped flange; bothridium projecting beyond inner margin of humeral processes, openings angled laterally, lip with long cylindrical posterior projection. Bothridial setae very long, curved, bacilliform, smooth.</p><p>Notogaster: ratio of length to breadth: 0.83; notogaster rounded, convex. Humeral process narrow, fine, pointed, waisted basally, lateral margin convex, extending to point level with middle of lamella (Fig. 17a), with prominent mound at base and laterally and smaller one posteriorly. With eight pairs of notogastral setae: l, h 2 and h 3 and p series; setae of l and h series smooth, recurved, shorter than distance between them, sub-equal in length, setae la and lm in centrodorsal position, others sub-marginal; p series shorter than l and m series.</p><p>Ventral aspect: subcapitulum acute apically, mentum as long as broad; subcapitular setae short, smooth, sub-equal in length (Fig. 17b). Chelicerae chelate-dentate and of normal proportions. Tutorium short, rounded apically, not visible in ventral aspect. Epimeral plates discrete except IV, broadly separate in midline, sub-rectangular to sub-triangular, plates III shorter than others; large gap between plates II and III, epimeral setation 3-1-3-3; epimeral setae sub-equal in length: 3a-c clustered on medial part of plate. Pedotectum I (pd I) poorly-developed, with short curved point; pd II small, pointed; discidium acute, pointed. Ventral plate broader than long. Genital and anal plates about half length of genital plates apart; genital plates 62 µm long, with six pairs of minute setae, sub-equal in length, g 5 displaced laterally; lyrifissure iad in para-anal position, close to margin of anal plate, less than length of iad. Anal plates lozenge-shaped, 90 µm long; setae of the ad series very short, setiform, thin, on faint curved ridge posterior to anal plates. Pre-anal organ (po) T-shaped.</p><p>Material examined and locality data.   Holotype, ANIC accession no. 53-650, leaf litter,  Nothofagus moorei rainforest, Barrington Tops National Park, via  Salisbury, New South Wales, 1,524 m., coll. G.B. Monteith, 10.ii.1965.</p><p>Diagnosis.  Atalotegaeus monteithi can be distinguished from other species in the genus by the following combination of characters: (1) the gap between epimeral plates II and III; (2) the curved, transverse ridge posterior of the anal plates; (3) the short, flagelliform lamellar setae; (4) the long cylindrical projection on the lip of the bothridium; (5) the relatively short, strongly recurved setae la and lm; (6) the mound-like structures at the base of the humeral process; (7) the partial translamella; (8) the anterior condyle of enantiophysis H.</p><p>Remarks. The recombination of  Neseutegaeus monteithi to  Atalotegaeus is detailed above in the remarks section on  Neseutegaeus . The description of  N. monteithi by J. and P. Balogh (1983a) differs in certain respects from the redescription of the holotype above. I found the bothridial setae were smooth apically, not ciliate and the bothridium has a large apical projection. Also, the apex of the lamellar cusp has a medial tooth and the two mound-like structures at the base of humeral process are larger than originally illustrated.</p></div>	https://treatment.plazi.org/id/03A2C77C4645FFEAC79CB537130DDF78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4647FFEBC79CB7B91676DF00.text	03A2C77C4647FFEBC79CB7B91676DF00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Humerotegaeus Colloff 2023	<div><p>Humerotegaeus gen. nov.</p><p>Type species:  Humerotegaeus carinatus sp. nov.</p><p>Diagnosis. Medium-sized to large oribatid mites (ca. 650–900 μm long), lamellar cusps with cup-shaped incisions flanked either side by tooth; lamellae relatively narrow: only about as wide as prodorsum. Chelicerae chelate-dentate and of normal proportions. Humeral processes densely rugose, with pointed apices and humeral plates basally; humeral plate either separated from rest of humeral process by anterior invagination or wedge-shaped plate indicated by obtuse anterior and posterior sutures. Bothridial seta long, thin; posterior margin of bothridium lacking anterior condyle of enantiophyses H. With nine pairs of notogastral setae (setae h 1 present) distributed in marginal position. Pedotectum I rectangular in ventral aspect; pd II trapezoid; discidium lobe-like, pointed. Perigenital carina and enantiophysis E4 present. Pre-anal organ T-shaped. The genus can be distinguished from  Neoeutegaeus by the following combination of character states: (1) the presence of humeral plates posterior of the humeral processes; (2) the presence of enantiophyses E4 and perigenital carinae; (3) the absence of the anterior condyles of enantiophyses H; (4) the relatively narrow lamellae with pointed cusps; (5) the rugose ornamentation on the humeral processes and (5) the broad, rectangular pedotectum II.</p><p>Etymology. this genus is named for the presence of the humeral plates posteriolateral of the humeral processes in the type species. The gender of the genus-group name, with the Latin gender ending - us, latinised from the Greek ending - os, is masculine.</p><p>Remarks. Character states of note that distinguish  Humerotegaeus from  Eutegaeidae are the lamellae, which are relatively short and do not extend beyond the rostrum or cover the entire prodorsum.Their lateral margins are straight and not convex. Also, the fusion of the anteromedian part of the lamellae is not found in eutegaeids.  Humerotegaeus is morphologically most similar to  Neoeutegaeus, with which it shares the long, pointed, triangular humeral processes with broad bases, lamellar cusps with apical teeth, the presence of perigenital carinae and enantiophyses E4 (also present in  Bornebuschia and some species of  Porrhotegaeus,  Dicrotegaeus and  Compactozetes), short, narrow epimeral plates II, a lobe-like discidium, incomplete posterior margins of epimeral plates IV and notogastral setae of the l and h series in the marginal position.  Humerotegaeus spp. also differs from  Neoeutegaeus spp. by the rugose ornamentation on the humeral processes and the T-shaped pre-anal organ. The presence of humeral plates on the lateral margins of the notogaster posterior of the humeral processes is a generic autapomorphy in the  Neoeutegaeidae .</p></div>	https://treatment.plazi.org/id/03A2C77C4647FFEBC79CB7B91676DF00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4647FFEAC79CB27113BDDA36.text	03A2C77C4647FFEAC79CB27113BDDA36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeidae Colloff 2023	<div><p>Neoeutegaeidae fam. nov.</p><p>Type genus:  Neoeutegaeus Aoki, 1964, p. 307 .</p><p>Diagnosis. Small to large oribatid mites (280–850 μm long). Rostrum rounded, entire. Lamellae broad, covering lateral margins of prodorsum, with free cusps; lamellar seta emerging from excavation at apex of cusp, often between two teeth. Interlamellar setae typically minute. Bothridium long, corniculate, not fused with base of lamella, opening anterolaterally, with circular internal thickening. Bothridial seta long, setiform or bacilliform. Prodorsum markedly waisted posterior of bothridium. Humeral processes long, pointed apically, parallel, straight, broad basally, not waisted, extending from anterolateral margin of notogaster to point at least half the length of the lamella, with or without dorsal keel, posterolateral margin with protrusion where humeral process joins notogaster. Notogaster Ushaped, convex, smooth, with eight or nine pairs of setae, all positioned marginally or sub-marginally; setae of l and h series (and occasionally p 1) longer and of different morphology from those of p series (or p 2 and p 3). Chelicerae chelate-dentate, of normal proportions. Epimeral plates II short, narrow. Pedotectum I (pd I) sub-rectangular in lateral outline; pd II rectangular or lobed, leaf-shaped; discidium pointed or lobed. Circumpedal carinae, perigenital carinae and enantiophyses E4 present. Genital setae five or six pairs, g 4 pair displaced laterally, adanal setae two or three pairs. Anal plates lozenge-shaped. Pre-anal organ oval ( Neoeutegaeus) or T-shaped ( Humerotegaeus).</p><p>Remarks.  Neoeutegaeus was formerly placed in  Eutegaeidae (Table 1), but the humeral processes emerging from the anterolateral margin of the notogaster and with broad bases, the lobe-like discidium, the short and narrow epimeral plates III, the presence of perigenital carinae and enantiophyses E4 are character states shared with  Humerotegaeus gen nov. and are sufficient to justify the establishment of a separate family.</p></div>	https://treatment.plazi.org/id/03A2C77C4647FFEAC79CB27113BDDA36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4646FFE4C79CB2A81687DD5C.text	03A2C77C4646FFE4C79CB2A81687DD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Humerotegaeus carinatus Colloff 2023	<div><p>Humerotegaeus carinatus sp. nov.</p><p>(Fig. 18)</p><p>Dimensions. Holotype female: length 845 μm, breadth 608 μm. Paratype female: length 909 μm, breadth 624 μm; paratype males: lengths 814 and 791 μm, breadths 569 and 553 μm. Ratio of prodorsum to total length: 0.33 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) curved, smooth, on short tubercles, not visible in dorsal aspect. Lamellae extending slightly beyond anterolateral margin of prodorsum, lateral margins more-or-less straight; without translamella. Lamellar setae (le) smooth, almost straight, extending well beyond apices of rostral setae, emerging from small, shallow cup-shaped incisions, each flanked by short teeth; lamellar cusps not extending beyond rostrum, fused apically in midline (Fig. 18a). Interlamellar setae (in) same length as lamellar setae, on faint, inverted V-shaped ridge on posteromedian prodorsum. Bothridia strongly curved, projecting as far as inner margins of humeral processes, openings angled anterolaterally, posterior margin convex, lacking Vshaped anterior condyle of enantiophysis H. Bothridial setae long, curved, bacilliform, smooth, pointed.</p><p>Notogaster: ratio of length to breadth: 0.93; notogaster rounded, convex. Humeral processes with rugose microsculpture, lacking dorsal keel, slightly waisted at bases, pointed apically, extending anteriorly to point level with middle of lamella (Fig. 18a). With large, curved humeral plates (hp) posterolateral of humeral processes marked by indentations in margin of notogaster. With nine pairs of setae: complete l, h and p series present, marginally positioned, smooth, short (75-115 μm); l series slightly longer than h series (Fig. 12a).</p><p>Ventral aspect: subcapitulum acute apically; mentum as long as broad; subcapitular setae short, smooth, setae m twice as long as a and h (Fig. 18b). Tutorium broad, apex rounded with several short teeth. Epimeral plates broadly separated in midline, plates I triangular, with lateral ridges delineating margins of plates from pedotectum I, plates II-IV lacking median margins, plates III shorter than others with longitudinal ridge laterally with two setae medial of ridge and one lateral; epimeral setation 3-1-3-2; setae 3a and 3b longer than others. Pedotectum I (pd I) well-developed, sub-triangular, with incised region posterolaterally; pd II well developed, trapezoid, medial to and covered by large, curved humeral plates (hp) in ventral aspect; discidium with ovoid, pointed. Ventral plate broader than long. Genital and anal plates almost contiguous; perigenital carina and enantiophysis E4 present; genital plates 128 µm long, with five pairs of setae sub-equal in length, g 5 displaced laterally; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 213 µm long. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1049;  paratypes: one female, two males, ANIC accession no. 53-1050, ANIC 3751, sieved litter and mossy logs, rainforest with  Eucalyptus sp., Lyrebird Nature Walk, 3.3 km south-east of Mount Field, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.66667&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.66667/lat -42.683334)">Mount Field National Park</a>, Tasmania, 42°41'S, 146°40'E, 690 m., coll. T. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.66667&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.66667/lat -42.683334)">Weir</a> and C. Lemann.</p><p>Etymology. The specific name,  carinatus, refers to the presence of the perigenital carinae.</p><p>Diagnosis. Differences between  H. carinatus and  H. concentricus sp. nov. are detailed under the diagnosis for the latter species below.</p></div>	https://treatment.plazi.org/id/03A2C77C4646FFE4C79CB2A81687DD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4649FFE5C79CB09412B0D8FF.text	03A2C77C4649FFE5C79CB09412B0D8FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Humerotegaeus concentricus Colloff 2023	<div><p>Humerotegaeus concentricus sp. nov.</p><p>(Fig. 19)</p><p>Dimensions. Holotype female: length 699 μm, breadth 488 μm. Paratype females (n = 9): mean length 734 μm (range 717–763 μm), mean breadth 511 μm (range 480–530 μm); paratype males (n = 4): mean length 678 μm (range 667–688 μm), mean breadth 469 μm (range 444–493 μm). Ratio of prodorsum to total length: 0.4 (holotype).</p><p>Description of adult. Prodorsum: rostrum rounded; rostral setae (ro) curved, with sparse spinose ornamentation basally, visible in dorsal aspect, emerging from lateral margins of rostrum. Lamellae broad, extending well beyond lateral margin of prodorsum, separated medially, parallel, lateral margins curved, convex. Translamella absent. Lamellar setae (le) with sparse spinose ornamentation, strongly curved medially, extending beyond apices of rostral setae, emerging from shallow cup-shaped incisions, flanked by long medial tooth and shorter lateral tooth; lamellar cusps extending well beyond rostrum (Fig. 19a). Interlamellar setae (in) very short. Bothridia long, strongly curved, projecting beyond median margins of humeral processes, openings angled anterolaterally, posterior margin convex, with V-shaped anterior condyle of enantiophysis H. Bothridial setae long, bacilliform, smooth, pointed.</p><p>Notogaster: ratio of length to breadth: 0.87; notogaster rounded, convex. Humeral processes well-developed, with distinctive rugose microsculpture and lacking dorsal keel; pointed apically, extending anteriorly to point level with middle of lamella (Fig. 19a). Base of humeral process with humeral plate (hp) indicated by obtuse anterior suture and acute posterior suture giving plate wedge-shaped appearance. With nine pairs of notogastral setae, l, h and p series present, all marginally positioned, thin, setiform, smooth, short (30-45 μm); l series about same length as h series (Fig. 19a).</p><p>Ventral aspect: subcapitulum acute apically; mentum as long as broad; subcapitular setae short, smooth, setae m same length as a and h (Fig. 19b). Tutorium narrow, without teeth. Epimeral plates I sub-rectangular, partially separated in midline; plates II fused in midline, lacking medial margins, rectangular and shorter than plates I, plates III and IV separated in midline and very narrow; epimeral setation 3-1-3-3; setae sub-equal in length. Epimeral plates III with pair of small tubercles. Pedotectum I (pd I) well-developed, rectangular, with reticulate microsculpture laterally; pd II well developed, trapezoid; discidium short, curved, pointed. Ventral plate broader than long. Genital and anal plates separated by distance just less than length of genital plates; with perigenital carina and enantiophysis E4 present; genital plates 82 µm long, with six pairs of setae sub-equal in length, g 4 displaced laterally, g 1 not longer than others; lyrifissure iad in para-anal position, some distance from margin of anal plate, more than length of iad. Anal plates lozenge-shaped, 120 µm long; setae of the ad series short, no longer than anal setae. Pre-anal organ (po) T-shaped.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1051;  paratypes: eight females, three males, ANIC accession no. 53-1052, ANIC 669, moss and leaf litter,  Nothofagus cunninghamii and  Atherosperma moschatum rainforest, Mount Field, Mount Field National Park, Tasmania, 42°41'S, 146°43'E, 160–240 m., coll. J. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.71666/lat -42.683334)">Lawrence</a> and T. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.71666/lat -42.683334)">Weir</a>, 9.ii.1980  .   Paratype: male, ANIC accession no. 53-1053, ANIC 297, moss and litter,  Nothofagus cunninghami rainforest, Cumberland Creek Valley, near Picnic Area [Cumberland Memorial Scenic Reserve], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Yarra Ranges National Park</a>, Victoria. 37°34’S 145°52’E, 920 m., coll. R. W. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Taylor</a> and R. J. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Bartell</a>, 4.xi.1970  .   Paratype: female, ANIC accession no. 53-1054, ANIC 592, leaf litter and decaying log, Cement Creek, 5 km west of Warburton, Victoria, 37°3’3S 145°42’E, coll. J. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7&amp;materialsCitation.latitude=-37.05083" title="Search Plazi for locations around (long 145.7/lat -37.05083)">Lawrence</a> and T. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7&amp;materialsCitation.latitude=-37.05083" title="Search Plazi for locations around (long 145.7/lat -37.05083)">Weir</a>, 18.i.1977  .</p><p>Etymology. The specific name refers to the pair of circular structures at the base of the humeral process, with the cuticular ornamentation forming concentric ridges and furrows.</p><p>Diagnosis.  Humerotegaeus concentricus differs from  H. carinatus by the following combination of characters: (1) the long, curved lamellar setae with spinose ornamentation; (2) the free lamellar cusps with a large medial spine; (3) the minute interlamellar setae; (4) the wedge-shaped humeral plate at the base of the humeral process, marked by anterior and posterior sutures; (4) the markedly shorter notogastral setae; (5) the pair of circular structures positioned ventrally at the base of each humeral process; (6) the morphology of the epimeres; (7) the short, pointed discidium and (8) the genital and anal plates are separated by distance just less than the length of the genital plate.</p></div>	https://treatment.plazi.org/id/03A2C77C4649FFE5C79CB09412B0D8FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4648FFE6C79CB5F313C2DCE0.text	03A2C77C4648FFE6C79CB5F313C2DCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus Aoki 1964	<div><p>Neoeutegaeus Aoki, 1964</p><p>Neoeutegaeus Aoki, 1964, p. 307 .</p><p>Neoeutegaeus Hammer, 1962, p. 75, nomen nudum.</p><p>Type species:?  Eutegaeus silvicola Hammer, 1962, p. 75 .</p><p>Diagnosis. the following diagnosis is modified from those of Aoki (1964) and Luxton (1988a). Adults: small oribatid mites (&lt;350 μm), body with granular cerotegument; rostrum acute, rounded; lamellar cusps one-third length of lamellae, extending beyond rostrum, cusps close together medially, connected with complete or incomplete translamella; lamellar setae emerging from apex of cusp; apex rounded without teeth or pointed with two small teeth; posterior prodorsum and anterior notogaster with or without condyles of enantiophyses B; humeral processes long, broad, triangular, projecting from anterior edge of notogaster, reaching beyond bothridia, expanded at point level with dorsosejugal scissure; with eight or nine pairs of notogastral setae positioned sub-marginally. Six pairs of genital setae, two or three pairs of adanal setae; with perigenital carinae and associated enantiophysis E4. Chelicerae chelate-dentate, of normal proportions. With elongated strip of integument between median borders of epimeral plates I; pedotectum I rectangular in ventral aspect, pd II and discidium very large, rounded or pointed. Anal plate sub-rectangular, rounded anteriorly and laterally, not lozenge-shaped; pre-anal organ oval. Immatures: gastronotic setae expanded, ovoid, pointed, with fine spines, emerging from tubercles, not scale-like structures; lateral margin of prodorsum entire, not incised.</p><p>Remarks. Aoki (1964) recombined?  Eutegaeus silvicola Hammer, 1962 with  Neoeutegaeus and made it the type species of the genus. As well as this species and the new species described below, the genus contains  N. africanus Mahunka, 1974 from Cameroon and  N. phyllophorus from Australia, redescribed below.</p></div>	https://treatment.plazi.org/id/03A2C77C4648FFE6C79CB5F313C2DCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C464BFFE0C79CB1C816EDD9DE.text	03A2C77C464BFFE0C79CB1C816EDD9DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus torsteini Colloff 2023	<div><p>Neoeutegaeus torsteini sp. nov.</p><p>(Figs. 20, 21)</p><p>Neoeutegaeus sp.: Colloff, 2019, p. 306, Figs. 10c, 10d; 11f; 12f; 13f; 14f.</p><p>Dimensions. Holotype female length 301 μm, breadth 208 μm. Paratype males (n = 8): length 274 μm (range 261– 295 μm), breadth 182 μm (range 163–191 μm). Paratype females (n = 17): mean length 293 μm (range 287–308 μm); mean breadth 195 μm (range 188–209 μm). Ratio of prodorsum to total length: 0.48 (holotype). Tritonymphs (n = 5) mean length 317 μm (range 298–316 μm), mean breadth 202 μm (range 189–218 μm).</p><p>Description of adult. Prodorsum: rostrum acute, rostral setae (ro) short, curved, smooth, not visible in dorsal aspect. Lamellae broad, separated medially, lateral margins convex, extending beyond lateral margin of prodorsum and anteriorly beyond rostrum (Fig. 20a). Lamellar setae (le) with spinose ornamentation, curved medially, apices overlapping, emerging from apices of free, sub-triangular lamellar cusps; medial margins of cusps forming V-shape; with complete translamella. Interlamellar setae (in) minute, curved. Bothridia as broad as long, projecting beyond inner margins of humeral processes, openings angled anterolaterally, posterior margin lacking anterior condyle of enantiophysis H. Bothridial setae long, curved, bacilliform, head slightly expanded with small spines.</p><p>Notogaster: ratio of length to breadth: 0.81; notogaster rounded, convex with cerotegument of small, scattered tubercles. Humeral processes very long, straight, parallel, triangular, pointed, lacking dorsal keel, extending to point level with middle of lamella, lateral margin expanded where process joins margin of notogaster, then incurved (Fig. 20a). Lyrifissure ia long, positioned at base of humeral process lateral of posterior condyle of enantiophysis H. With eight pairs of notogastral setae, l series and h 2 and h 3 sub-marginal, smooth, thin, curved, covered in cerotegument thus appearing thickened (Fig. 20c); p series extremely short, setiform, not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum broad; subcapitular setae minute, smooth, setae h on anterior margin of mentum (Fig. 20b). Tutorium very broad, pointed. Epimeral plates discrete, separated in midline, sub-rectangular, plates III and IV shorter than I and II, with median strip of cuticle between plates I; epimeral setation 3-1-3-2; setae minute, sub-equal in length. Pedotectum I (pd I) well-developed, rectangular, with short lateral cusp; pd II long, broad, rounded apically; discidium lobe-like; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, much broader than long. Genital and anal plates separated by distance of just over half length of genital plates; genital plates 39 µm long, with six pairs of short setae sub-equal in length, g 4 displaced laterally; two pairs of minute adanal setae; lyrifissure iad in para-anal position, close to margin of anal plate, less than length of iad. Anal plates ovoid, 42 µm long. Pre-anal organ (po) oval.</p><p>Description of tritonymph, nymphal and larval scalps. Tritonymph prodorsum: with cerotegument of scattered tubercles; centrodorsal microsculpture tessellated, bounded by slight ridges (Fig. 21a). Rostrum broad, rounded; rostral setae straight, smooth, on short tubercles; lamellar seta broad, ovoid, with small spines, straight, emerging from short tubercle, joined by transverse ridge. Interlamellar seta very short, smooth, on squat tubercle. Bothridia cup-shaped, slightly constricted basally, with exobothridial seta (ex) on posterior margin; bothridial seta long (103 µm), curved, smooth, slightly expanded apically, pointed.</p><p>Tritonymph gastronotum: U-shaped, longer (229 µm) than broad (202 µm), with cerotegument of scattered tubercles. Anterolateral margin with rounded, squat humeral process bearing seta c 3 (Fig. 21a). With 12 pairs of gastronotal setae: full complement of c, l, h (h 1 present) and p series; c 3, l and h series and p 1 broad, ovoid, with fine spines; seta c 1 setiform, smooth; c 2 long, leaf, shaped, with fine spines.</p><p>Deutonymphal and protonymphal scalp: deutonymphal scalp with seven pairs of setae: c 3 (c 1 and c 2 absent), l series, h 2, h 3 and p 1, all on short tubercles; protonymphal scalp with six pairs of setae c 3 (c 1 and c 2 absent), l series, h 3 (h 2 absent) and p 1 (Fig. 21a).</p><p>Larval scalp: with scattered alveolate microsculpture; nine pairs of leaf-like setae on short tubercles: c 1 and c 3, la and lm, h 3 and p 1 and the centrodorsal d series. Setae dp on very long, pendulous tubercles (Fig. 21a).</p><p>Tritonymph ventral aspect: microsculpture of ventral surface with fine, widely-spaced striations (Fig. 21b); epimeral setal formula 2-1-2-2; with five pairs of genital setae, two pairs of adanal setae.</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps): 10-(8,9,12)-8.</p><p>Type designation, material examined and locality data.  Holotype female; ANIC accession no. 53-1058,  paratypes: 27 females, 12 males, five tritonymphs, ANIC accession no. 53-1059, moss on Sassafras ( Atherosperma moschatum), rainforest, Errinundra Saddle, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.85&amp;materialsCitation.latitude=-37.316666" title="Search Plazi for locations around (long 148.85/lat -37.316666)">Errinundra National Park</a>, Victoria, 37°19’S 148°51’E, 1,030 m., coll. M.J. Colloff, 29.ix.2009.</p><p>Etymology. This species is named in honour of my friend and colleague, the late Associate Professor Thorstein Solhøy, formerly of the Department of Biology, University of Bergen, in recognition of his contribution to the taxonomy and ecology of oribatid mites.</p><p>Diagnosis.  Neoeutegaeus torsteini can be distinguished from other species in the genus by the following combination of characters: (1) the sub-triangular apices of the lamellar cusps, without teeth; (2) the well-developed, overlapping lamellar setae with spinose ornamentation; (3) the posterior condyles of enantiophyses H; (4) the minute, stout interlamellar setae; (5) the very broad tutoria; (6) the very long lyrifissures ia; (7) the long bothridial setae with an expanded head bearing small spines.</p><p>Remarks.  Neoeutegaeus torsteini shares with  N. melipsilon sp. nov. (cf. below) the broad, rounded pedotecta II and short, thick, blunt notogastral setae, but the latter species has nine pairs of notogastral setae, two teeth on the lamellar cusp and much shorter lamellar setae.</p><p>The association of the tritonymph with the adult is based, in part, on their presence in the same samples, and the absence of other  Neoeutegaeus spp., but also the size range of the tritonymphs fits with that of the adults, being slightly shorter and narrower. However, there are no obvious morphological characteristics shared by the tritonymphs and the adults.</p></div>	https://treatment.plazi.org/id/03A2C77C464BFFE0C79CB1C816EDD9DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C464CFFE3C79CB38816BBD810.text	03A2C77C464CFFE3C79CB38816BBD810.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus melipsilon Colloff 2023	<div><p>Neoeutegaeus melipsilon sp. nov.</p><p>(Figs. 22, 23)</p><p>Dimensions. Holotype female 300 μm, breadth 194 μm. Paratype females (n = 14): mean length 308 μm (range 300–318 μm); mean breadth 192 μm (range 183–202 μm). Paratype males (n = 4): mean length 281 μm (range 279–283 μm); mean breadth 172 μm (range 169–176 μm). Tritonymph length 317 μm, breadth 202 μm. Ratio of prodorsum to total length (holotype): 0.38.</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) minute, smooth, not visible in dorsal aspect. Tutoria curved, projecting beyond margin of prodorsum. Lamellae broad, separated medially, lateral margins convex, covering prodorsum and extending well beyond rostrum (Fig. 22a). Lamellar seta (le) very short, curved, smooth, spiniform, emerging from slight concavity on apex of lamellar cusp between two small sharp teeth; medial margins of cusps parallel, with complete translamella basally. Interlamellar setae (in) well-developed, curved, smooth. Bothridia elongate, corniculate, not projecting as far as humeral processes, opening anterolaterally. Bothridial seta long, head expanded slightly then truncated, with few small spines.</p><p>Notogaster: cerotegument of small, scattered tubercles; ratio of length to breadth: 0.87; notogaster rounded, convex. Humeral processes long, straight, parallel, triangular, pointed, with dorsal keel, extending to point below middle of lamella, lateral margin expanded at anterolateral margin of notogaster forming slight conical projection (Fig. 22a). With eight pairs of notogastral setae, l series and h 2 and h 3 sub-marginal, smooth, setiform, curved, short, pointed, sub-equal in length; p series shorter, setiform, p 1 and p 2 visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute; subcapitular setae short, smooth, setae h on anterior margin of mentum (Fig. 22b). Tutorium very broad, apex right-angled. Epimeral plates discrete, separated in midline, sub-rectangular, plates III and IV much shorter than I and II; with strip of triangular cuticle between epimeral plates I; epimeral setation 3-1-3-2; setae very short, sub-equal in length. Pair of shallow epimeral foveolae (ef) present between plates II and III. Pedotectum I (pd I) well-developed, rectangular, with anterior cusp and alveolate microsculpture; pd II very long, broad, lobe-like; discidium long, broad, rounded apically; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance less than half length of genital plates; genital plates 38 µm long, with six pairs of short setae sub-equal in length, g 4 displaced laterally; two pairs of short adanal setae; lyrifissure iad in para-anal position, close to anal plate. Anal plates ovoid, 46 µm long. Pre-anal organ (po) oval.</p><p>Description of tritonymph, nymphal and larval scalps. Tritonymph prodorsum: with microsculpture of sub-hexagonal ridges on central prodorsum (Fig. 23a). Rostrum broad, rounded; rostral setae straight, smooth, short, on short tubercles; lamellar setae broad, ovoid, with small spines, straight, emerging from well-developed tubercles, joined by transverse ridge. Interlamellar setae very short, smooth, on squat tubercles. Bothridia cup-shaped, projecting laterally; bothridial seta long, curved, smooth, slightly expanded apically, pointed.</p><p>Tritonymph gastronotum: U-shaped, broader than long, smooth. Anterolateral margin with rounded, sub-rectangular humeral processes bearing setae c 3, indented posteriorly (Fig. 23a). With 12 pairs of gastronotal setae: full complement of c, l, h (h 1 present) and p series; c 3, l and h series and p 1 broad, ovoid, with fine spines, pointed; seta c 1 setiform, smooth; c 2 long, leaf, shaped, with fine spines.</p><p>Deutonymphal and protonymphal scalp: with eight pairs of setae: c 3 (c 1 and c 2 absent), l series, h series and p 1, all on short tubercles (Fig. 23a).</p><p>Larval scalp: with scattered alveolate microsculpture; seven pairs of leaf-like setae on short tubercles: la, lm, h 3 and p 1 and centrodorsal d series. Setae dp on long, pendulous tubercles (Fig. 23a).</p><p>Tritonymph ventral aspect: ventral plate with fine, widely-spaced ridges posteriorly (Fig. 23b); epimeral setal formula 3-1-3-2; with five pairs of minute genital setae, one pair of aggenital setae, two pairs of adanal setae.</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps): 8-(9,10,12)-8.</p><p>Type designation, material examined and locality data.   Holotype, ANIC accession no. 53-1060, wet moss, Cumberland Creek Valley, Marysville-Woods Point Road, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.55" title="Search Plazi for locations around (long 145.86667/lat -37.55)">Cambarville</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.55" title="Search Plazi for locations around (long 145.86667/lat -37.55)">Yarra Ranges National Park</a>, Victoria, 37º33’S 145º52’E, coll. R. E. Roughley, 12.iii.1996  .   Paratypes: 22 females, seven males, one tritonymph, ANIC accession no. 53-1061, ANIC 297, moss and litter,  Nothofagus cunninghami rainforest, Cumberland Creek Valley, near Picnic Area [Cumberland Memorial Scenic Reserve], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Yarra Ranges National Park</a>, Victoria. 37°34’S 145°52’E, 920 m. Coll. R. W. Taylor and R. J. Bartell, 4.xi.1970  .   Paratypes: 14 females, four males, ANIC accession no. 53- 1062, ANIC 592, leaf litter and decaying log, Cement Creek, 5 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7&amp;materialsCitation.latitude=-37.05083" title="Search Plazi for locations around (long 145.7/lat -37.05083)">Warburton</a>, Victoria, 37°3’3S 145°42’E, coll. J. Lawrence and T. Weir, 18.i.1977  .</p><p>Etymology. The prefix of the specific name is from the Latin mel (honey), referring to the colour of the cuticle. The suffix - ipsilon is an invented word, chosen for its Greek phonaesthetic.</p><p>Diagnosis.  Neoeutegaeus melipsilon can be distinguished from other species in the genus by the following combination of characters: (1) the short, smooth, spiniform lamellar setae each emerging from between two teeth; (2) the relatively long, curved, stout interlamellar setae; (3) the bothridial setae with club-shaped heads, truncated apically and with a few spines; (4) the prominent median strip of triangular cuticle between epimeral plates I; (5) the large tutoria with blunt apices extending beyond the lateral margins of the lamellae; (6) the alveolate microsculpture on the lateral margin of pedotectum I; (7) the very large, lobe-shaped pedotectum II; (8) the pair of epimeral foveolae between epimeral plates II and II.</p><p>Remarks.  Neoeutegaeus melipsilon shares with  N. africanus the relatively long interlamellar setae, lamellar cusps each with two small apical teeth and the large discidia with rounded apices, but it lacks the anterior condyles of enantiophyses H. Also, in  N. africanus the head of the bothridial seta is club-shaped with small spines, but is bilobed.</p><p>The association of the tritonymph with the adult is based on their presence in the same samples in the absence of other  Neoeutegaeus spp., but also the size range of the tritonymphs fits with that of the adults, being slightly shorter and narrower. However, as for  N. torsteini, there are no obvious morphological characteristics shared by the tritonymphs and adults.</p></div>	https://treatment.plazi.org/id/03A2C77C464CFFE3C79CB38816BBD810	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C464EFFFDC79CB5D81505DD70.text	03A2C77C464EFFFDC79CB5D81505DD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus malcolmi Colloff 2023	<div><p>Neoeutegaeus malcolmi sp. nov.</p><p>(Fig. 24)</p><p>Dimensions. Holotype female length 280 μm, breadth 186 μm, paratype females (n = 16) length 278 μm (range: 271–292 μm), breadth 175 μm (range: 166–193 μm); paratype males (n = 4) length 265 μm (range: 263–267 μm), breadth 162 μm (range: 160–164 μm). Ratio of prodorsum to total length (holotype): 0.48.</p><p>Description of adult. Prodorsum: rostrum acute; rostral setae (ro) short, straight, smooth, not visible in dorsal aspect. Lamellae broad, very long, almost half body length, extending well beyond rostrum, separated medially, lateral margins convex (Fig. 24a). Lamellar seta (le) short, curved medially, smooth, emerging from alveolus on anterior margin of cone-shaped lamellar cusp; medial margins of cusps convex, with complete translamella. Interlamellar setae (in) minute, close together. Bothridia elongate, corniculate, projecting laterally beyond inner margins of humeral processes, openings angled anterolaterally. Bothridial seta long, head club-shaped, apex rounded, smooth.</p><p>Notogaster: ratio of length to breadth: 0.82; notogaster rounded, convex. Humeral processes very long, as long as lamellae, straight, parallel, triangular, pointed, extending to point just posterior of translamella, lateral margin expanded where process joins anterolateral margin of notogaster forming slight projection (Fig. 24a). Lyrifissures ia and im short. With eight pairs of notogastral setae, l series and h 2 and h 3 sub-marginal, smooth, short, sub-equal in length, stout, covered in cerotegument; p series very short, not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute; subcapitular setae extremely short, sub-equal in length, smooth (Fig. 24b). Tutorium very broad, pointed. Epimeral plates discrete, broadly separated in midline, sub-rectangular to ovoid, plates III and IV much shorter than I and II; with median strip of dumbbell-shaped cuticle between plates I and II; epimeral setation 3-1-2-2; setae very short, sub-equal in length. Pedotectum I (pd I) well-developed, rectangular, with blunt anterior apex; pd II very long, broad, lateral margin sloped obliquely; discidium broad, pointed apically; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of half length of genital plates; genital plates 36 µm long, with five pairs of short setae sub-equal in length, aligned longitudinally; two pairs of extremely short adanal setae; lyrifissure iad in para-anal position, some distance from margin of anal plate, about same as distance from plate as bases of setae ad 2. Anal plates ovoid, 41 µm long. Pre-anal organ (po) oval.</p><p>Type designation, material examined and locality data.   Holotype female, ANIC accession no. 53-1063, ANIC 3751, sieved litter and mossy logs, rainforest with  Eucalyptus sp., Lyrebird Nature Walk, 3.3 km south-east of Mount Field, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.66667&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.66667/lat -42.683334)">Mount Field National Park</a>, Tasmania, 42°41'S, 146°40'E, 690 m., coll. T. Weir and C. Lemann, 8.iii.2008  .   Paratypes: 12 females, three males, ANIC accession no. 53-10064, ANIC 3757, sifted litter and mossy logs, rainforest with  Eucalyptus sp., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.09721&amp;materialsCitation.latitude=-41.64528" title="Search Plazi for locations around (long 145.09721/lat -41.64528)">State Forest</a>, 2.2 km NE Corinna, Tasmania, 41°38’43”S 145°05’50”E, 45 m  ., coll. T. Weir &amp; C. Lemann, 14.iii.2008 .   Paratype female, ANIC accession no. 53-1065, ANIC 666, litter and rotting log debris,  Nothofagus cunninghami rainforest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-42.683334" title="Search Plazi for locations around (long 146.71666/lat -42.683334)">Mount Field National Park</a>, Tasmania, 42°41'S, 146°43'E, 160–240 m., coll. J. Lawrence &amp; T. Weir, 30.i.1980  .   Paratypes: four females, one male, ANIC accession no. 53-1066, TAS-183, litter,  Eucalyptus forest, Old Farm Road, Mount Wellington, Tasmania, 42°54'S, 147°14'E, coll. P. Greenslade, 20.v.1989  .</p><p>Etymology. This species is named in honour and in memory of my friend and colleague, the late Dr Malcolm Luxton, formerly of the Department of Biology, Liverpool John Moores University and National Museum of Wales, Cardiff, in recognition of his substantial contribution to the systematics, taxonomy and ecology of oribatid mites.</p><p>Diagnosis.  Neoeutegaeus malcolmi can be distinguished from other species in the genus by the following combination of characters: (1) the narrow, acute, cone-shaped lamellar cusps lacking teeth,; (2) the short, smooth lamellar setae; (3) the minute interlamellar setae; (4) the prodorsum almost as long as the notogaster; (5) the long, club-shaped bothridial setae with smooth head; (6) the large, curved pedotecta II with straight, oblique margins folded dorsally; (7) the broad, pointed discidium; (8) the epimeral setal formula 3-1-2-2.</p><p>Remarks.  Neoeutegaeus malcolmi shares with  N. torsteini the lamellar seta emerging from the apex of the free lamellar cusp, but in  N. malcolmi the cusps are cone-shaped and relatively long, whereas in  N. torsteini they are sub-triangular, shorter and broader.</p></div>	https://treatment.plazi.org/id/03A2C77C464EFFFDC79CB5D81505DD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4650FFFEC79CB078158AD8DE.text	03A2C77C4650FFFEC79CB078158AD8DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus corniculatus Colloff 2023	<div><p>Neoeutegaeus corniculatus sp. nov.</p><p>(Fig. 25)</p><p>Dimensions. Holotype male length 275 μm, breadth 180 μm. Paratype males (n = 4) mean length 280 μm (range 275–287 μm), mean breadth 181 μm (range 176–183 μm). Paratype females (n = 8) mean length 292 μm (range 283–300 μm), mean breadth 190 μm (range 187–193 μm). Ratio of prodorsum to total length: 0.46 (holotype)</p><p>Description of adult. Prodorsum: rostrum truncated apically; rostral setae (ro) extremely short, smooth, not visible in dorsal aspect. Lamellae broad, very long, almost half length of body, extending well beyond rostrum, separated medially, lateral margins convex (Fig. 25a). Lamellar seta (le) very short, smooth, emerging from alveolus on anterior margin of lamellar cusp; medial margins of cusps convex, almost meeting in mid-line, with partial translamella; lamella with small pointed cusp just anterior of bothridium. Interlamellar setae (in) minute, broadly separated. Bothridia elongate, corniculate, projecting laterally beyond inner margins of humeral processes, angled anterolaterally, posterior margin of opening with curved projection. Bothridial seta long, head club-shaped, broad, with fine spines.</p><p>Notogaster: ratio of length to breadth: 0.77; notogaster rounded, convex. Humeral processes very long, as long as lamellae, straight, parallel, triangular, with dorsal keel, extending to point level with translamella; lateral margin expanded where process joins anterolateral margin of notogaster (Fig. 25a). Lyrifissures ia and im not observed. With eight pairs of notogastral setae, l series and h 2 and h 3 sub-marginal, smooth, curved, relatively short, sub-equal in length, stout, covered in cerotegument, rounded apically; p series very short, setiform, not visible in dorsal aspect. Cerotegument of coarse granules.</p><p>Ventral aspect: subcapitulum as broad as long; subcapitular setae short, sub-equal in length, smooth (Fig. 25b). Tutoria prominent, stout, curved, pointed, projecting well beyond lateral margins of prodorsum. Epimeral plates discrete, broadly separated in midline, sub-rectangular to ovoid, plates III and IV much shorter than I and II; with median strip of dumbbell-shaped cuticle between plates I and II, plates IV the smallest, ovoid; pair of epimeral foveolae (ef) present between plates II and III. Epimeral setation 3-1-2-2; setae very short, sub-equal in length. Pedotectum I (pd I) well-developed, rectangular, with blunt anterior apex; pd II long, broad, lateral margin sloped obliquely, indented; discidium broad, rounded; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of half length of genital plates; genital plates 31 µm long, with five pairs of short, spinose setae sub-equal in length, g 4 displaced laterally; two pairs of short adanal setae; lyrifissure iad long, in para-anal position, some distance from margin of anal plate, about same as length of seta ad 2. Anal plates ovoid, 38 µm long. Pre-anal organ (po) oval.</p><p>Type designation, material examined and locality data.  Holotype male, ANIC accession no. 53-1067,  paratypes: four males, eight females, ANIC accession no. 53-1068, litter,  Nothofagus cunninghami rainforest, Triplet Falls, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.5&amp;materialsCitation.latitude=-38.666668" title="Search Plazi for locations around (long 143.5/lat -38.666668)">Great Otway National Park</a>, Victoria, 38°40’S, 143°30’E, coll. M.J. Colloff, 17.vii.2011.</p><p>Etymology. This specific name,  corniculatus, is a Latin diminutive for ‘with horn-shaped projections’, referring to the prominent tutoria projecting from the lateral margins of the prodorsum.</p><p>Diagnosis.  Neoeutegaeus corniculatus can be distinguished from other species in the genus by the following combination of characters: (1) the small pointed cusp on the lateral margin of the lamellar just anterior of the bothridium; (2) the prominent horn-shaped tutoria; (3) the incomplete translamella; (4) the posterior margin of the bothridial opening with a curved projection; (5) the minute interlamellar setae, broadly separated; (6) the presence of epimeral foveolae between epimeral plates II and III.</p><p>Remarks.  Neoeutegaeus corniculatus shares with  N. phyllophorus the pointed lamellar cusps, each with a single, short tooth positioned medially, the club-shaped heads of the bothridial setae with spinose ornamentation, the cusp anterior of the bothridium at the base of the lamella, the keeled humeral processes and the presence of epimeral foveolae. However,  N. corniculatus has very short and smooth lamellar setae, not long barbed ones; the notogastral setae are blunt apically due to the covering of cerotegument, not pointed and it lacks the anterior condyle of enantiophyses H on the posterior prodorsum.</p></div>	https://treatment.plazi.org/id/03A2C77C4650FFFEC79CB078158AD8DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4653FFF9C79CB41F1536DC94.text	03A2C77C4653FFF9C79CB41F1536DC94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoeutegaeus phyllophorus Balogh & P. Balogh 1983	<div><p>Neoeutegaeus phyllophorus J. &amp; P. Balogh, 1983</p><p>(Figs. 26, 27)</p><p>Neoeutegaeus phyllophorus J. &amp; P. Balogh, 1983b, p. 291.</p><p>Neoeutegaeus sp.: Hunt et al., 1998.</p><p>Dimensions. Holotype male: length 281 μm, breadth 196 μm. Ratio of prodorsum to total length: 0.42.</p><p>Redescription of adult. The holotype specimen of  Neoeutegaeus phyllophorus is badly damaged and missing the anal plates, the bothridial, notogastral and lamellar setae. Accordingly, the redescription below is supplemented by reference to topotypic material, collected within 20 km of the type locality from identical microhabitat (leaf litter) in the same vegetation community (sub-tropical rainforest).</p><p>Prodorsum: rostrum acute apically; rostral setae (ro) extremely short, smooth, not visible in dorsal aspect. Lamellae broad, extending anteriorly well beyond rostrum, separated medially, lateral margins convex with a small invagination anterior of bothridium (Fig. 26a, 27a). Lamellar seta (le) short, curved medially, with fine spines, emerging on short apophysis, with shallow invagination and small tooth medially; medial margins of cusps parallel, with complete translamella. Interlamellar setae (in) minute, close together. Bothridia elongate, corniculate, projecting laterally beyond inner margins of humeral processes, openings angled anterolaterally. Bothridial seta long, head club-shaped, with fine spines. Anterior condyle of post-bothridial enantiophyses B on posterior margin of prodorsum.</p><p>Notogaster: ratio of length to breadth: 0.86; notogaster rounded, convex. Humeral processes long, straight, parallel, triangular, pointed, with dorsal keel, extending to point level with translamella; lateral margin expanded where process joins anterior margin of notogaster forming slight projection (Fig. 26a). Lyrifissures not visible. With eight pairs of sub-marginal notogastral setae, l series and h 2 and h 3 smooth, curved, short, pointed, sub-equal in length (Fig. 26a), covered in cerotegument; p series very short, setiform, not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum broad; subcapitular setae h extremely short, on anterior margin of mentum; setae a at least twice their length; mentum much broader than long, with fine, granular microsculpture (Fig. 26b, 27b). Tutorium very broad, pointed. Epimeral plates I-II discrete, broadly separated in midline, plates I massive, as broad as long; plates III shorter and narrower than others; with triangular median strip of cuticle between plates I and pair of epimeral foveolae (ef) at its base and second pair between plates II and III; epimeral setation 3-1-3-2; setae very short, sub-equal in length. Pedotectum I (pd I) well-developed, rectangular in lateral outline; pd II very long and broad; lateral margins sloped obliquely; discidium broad, rounded apically; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of less than half length of genital plates; genital plates longer than anal plates: 36 µm long, with five pairs of short setae sub-equal in length, g 3 displaced laterally; three pairs of short adanal setae; lyrifissure iad in para-anal position, some distance from margin of anal plate, about same as distance of bases of seta ad 2 from plate. Anal plates sub-rectangular, 33 µm long. Pre-anal organ (po) oval.</p><p>Material examined and locality data. Holotype male, ANIC accession no. 53-00637, litter,  Nothofagus moorei rainforest, Point Lookout, near Ebor, New England National Park, New South Wales, 30°29'S, 152°25'E, 1,525 m, coll. I. Naumann, 25.iv.1973. Non-type material: three males, four females, ANIC 836, litter, closed forest, Gibraltar Range National Park, New South Wales, 29°31'S, 152°22'E, 920 m, coll. L. Hill, 14.vi.1982. One adult female, ANIC 754,  Nothofagus and Sassafras ( Atherosperma moschatum) litter,  Nothofagus moorei forest, Cobark Forest Park, Barrington Tops, New South Wales, 31°59’S, 151°49’E, coll. T. Weir and A. Calder, 15.xi.1981. Eight females, two males, ANIC 459, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland, 28°14’S 153°08’E, 920 m, coll. R.W. Taylor, 21.iii.1973. Six females, two males, ANIC 460, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland, 28°14’S 153°08’E, 920 m, coll. R.J. Kohout, 21.iii.1973.</p><p>Diagnosis.  Neoeutegaeus phyllophorus can be distinguished from other species in the genus by the following combination of characters: (1) the long lamellar seta with spinose ornamentation; (2) the lamellar seta emerging from a short apophysis, with a shallow invagination and a small tooth medially; (3) the long bothridial setae with a club-shaped head bearing spinules; (4) the presence of the anterior tubercles of enantiophyses B on the posterior prodorsum; (5) the pronounced notch anterior of the bothridium; (6) the pointed notogastral setae; (7) the presence of two pairs of epimeral foveolae.</p><p>Remarks.  Neoeutegaeus phyllophorus is morphologically most similar to  N. corniculatus, as detailed above. The  Neoeutegaeus sp. recorded by Hunt et al. (1998) from rainforest litter at O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland (28°14’S 153°08’E, coll. R. Kitching), and illustrated in Fig. 27, is identified here as  N. phyllophorus: it has well-developed epimeral foveolae, a character state that does not occur in other  Neoeutegaeus species. Also,  Neoeutegaeus sp. shares with  N. phyllophorus similar dimensions, the invagination of the lateral lamella anterior of the bothridium, the long bothridial seta with a dilated head and short spines and the notogastral setae are the same size and shape.</p></div>	https://treatment.plazi.org/id/03A2C77C4653FFF9C79CB41F1536DC94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4654FFFAC79CB05D1246D8FC.text	03A2C77C4654FFFAC79CB05D1246D8FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetidae Luxton 1988	<div><p>Compactozetidae Luxton, 1988</p><p>Compactozetidae Luxton, 1988a, p. 86 .</p><p>Type genus:  Compactozetes Hammer, 1966, p. 76 .</p><p>Diagnosis. The following diagnosis is modified from that of Luxton (1988a). Lamellae well developed, fused with each other apically ( Compactozetes) or with short, broad, rounded or truncated free cusps ( Sadocepheus and  Hamotegeus). Anterior projections of humeral processes short and blunt; humeral processes originating on lateral margin at widest part of notogaster; posterior region of humeral process with two to four distinct lobes evident from lighter areas of cuticle and wavy lateral margin. Bothridia fused with bases of lamellae, not free. Notogaster circular, broader than long, with nine or ten pairs of marginal setae originating on alveoli or tubercles. Chelicerae chelate-dentate, of normal proportions. Pedotectum I with large curved point or spine, discidium lobe-like, not triangular. Anterior margin of pedotectum I and epimere I forming straight or curved transverse tectum covering posterior mentum. With six pairs of genital setae, three pairs of adanal setae. Circumpedal carina present. Perigenital carinae and enantiophyses E4 present or absent. Anal plates rectangular. Pre-anal organ oval or circular, occasionally with cylindrical apex.</p><p>Remarks. Luxton (1988b) included the genera  Bornebuschia,  Compactozetes,  Dicrotegaeus,  Hamotegeus and  Sadocepheus in his definition of  Compactozetidae . Norton and Behan-Pelletier (2009) made  Compactozetidae a junior synonym of  Cepheidae because of what they considered a lack of family-level diagnostic characters. The genera included in  Compactozetidae by Luxton (1988b) initially appear to be morphologically diverse, with an inconsistent, mosaic-like, distribution of shared characters, attributable largely to the inclusion of  Bornebuschia and  Dicrotegaeus . For example, in  Bornebuschia and  Dicrotegaeus the notogastral setae are arranged in three ranks (centrodorsal, sub-marginal and marginal; cf. Fig. 13a for distribution pattern), whereas in the other genera the setae are all marginal.  Dicrotegaeus has extremely long, pointed lamellae that extend well beyond the rostrum, unlike those of  Compactozetes,  Bornebuschia and  Hamotegeus which generally do not extend beyond the rostrum, have blunt, transverse or oblique free cusps or the cusps are fused with each other and the rostrum (in  Compactozetes).  Bornebuschia,  Dicrotegaeus and some  Compactozetes spp. have perigenital carinae and enantiophyses E4 (as does  Porrhotegaeus herminae J. and P. Balogh, 1983c in the Porrhotegaeidae  fam. nov.,  Humerotegaeus and  Neoeutegaeus spp. in the  Neoeutegaeidae and most species of  Pterozetidae), whereas the compactozetid genera  Hamotegeus and  Sadocepheus lack these structures.  Bornebuschia and  Dicrotegaeus have well-developed enantiophyses H on the posterior prodorsum and anterior notogaster, as well as anterior condyles of enantiophyses B, not found in other  Compactozetidae but present in some species of  Neoeutegaeus and  Sadocepheus spp. Bornebuschia and  Dicrotegaeus also have only two pairs of adanal setae, whereas all  Compactozetidae have three pairs, though most species of  Neoeutegaeus have two pairs ( N. phyllophorus has three pairs; cf. above).  Hamotegeus,  Sadocepheus spp. and  Compactozetes have elongated humeral processes, extending from a point level with notogastral setae lm and with two to four lobes of lighter cuticle separated by indentations and a square or trapezoid anterior lobe with a pointed or rounded apex. This latter character state is a family-level synapomorphy within the  Eutegaeoidea, as is the rectangular (rather than lozenge-shaped) anal plates. Finally,  Compactozetes zeugus Luxton 1988b,  Sadocepheus foveolatus Luxton, 1988b and  Hamotegeus spp. have the marginal notogastral setae (l and h series) emerging from tubercles rather than alveoli and  Sadocepheus elevatus Mahunka, 1987 has seta la only on a tubercle, the others emerging from alveoli. Notogastral setae emerging from tubercles is a synapomorphic generic character state within the  Eutegaeoidea, though clearly not all  Compactozetidae share this state.</p><p>J. Balogh and Mahunka (1969, p. 2) defined  Hamotegeus has having ten pairs of notogastral setae, of which seven pairs are on tubercles (‘basally incrassate’) on the notogastral margin, and with six pairs of genital setae, with humeral processes and cylindrical bothridia projecting anteriorly and laterally (‘pro- and exclinate’). They stated (p. 4) that  Hamotegeus “…takes an intermediate position between the genus  Cepheus,  Compactozetes and  Eutegaeus . Owing to the combination of characters given above, the new genus cannot be relegated to either one.” Subías (2004, p. 84) made  Hamotegeus the junior synonym of  Sadocepheus, though without giving reasons. Ermilov (2020a, p. 877) agreed, stating of  Hamotegeus: “It is very similar to  Sadocepheus distinguishing [sic] from the latter by such important traits as, e.g. the location of dorsal notogastral setae on tubular tubercles and the position of notogastral setae h 1 very close to each other. However, these traits were not unique. For example,  Sadocepheus foveolatus Luxton, 1988 also has dorsal notogastral setae inserted on tubular tubercles;  Sadocepheus makarchevae Sitnikova, 1975 and  Sadocepheus elevatus Mahunka, 1987 also have notogastral setae h 1 inserted close to each other.”</p><p>Luxton (1988a) differentiated  Hamotegeus from  Sadocepheus on the basis that  Hamotegeus has ten pairs of notogastral setae on apophyses or tubercles, whereas  Sadocepheus has nine pairs on alveoli. However, as noted above,  S. foveolatus Luxton, 1988b has the l and h series on tubercles. P. Balogh (1986) noted that in  H. longiseta and  H. breviseta seta la is positioned in a ventro-lateral position between the humeral process and the bothridum, but la is absent in  H. franzi and  H. granulatus J. Balogh &amp; Mahunka, 1969 . Thus  Hamotegeus has either nine or ten pairs of notogastral setae. Regarding the positioning of setae h 1, in  H. breviseta,  H. franzi and  H. granulatus both setae emerge from a single tubercle, whereas in  H. longiseta there are two tubercles that are adjacent but not connected.</p><p>Other than the arrangement of the notogastral setae, there are additional character states that differentiate  Hamotegeus from  Sadocepheus . Like  Sadocepheus and  Compactozetes, the humeral processes in  Hamotegeus originate on the margin of the notogaster at its widest part but their apices are short and strongly bilobed, projecting obliquely from the anterior margin of the notogaster, whereas in  Sadocepheus they are similar to those of  Compactozetes: elongated, with several lateral lobes and a square or trapezoid anterior lobe with a broad, pointed apex. In  Hamotegeus the dorsosejugal scissure is convex, whereas in  Sadocepheus it is transverse. In  Hamotegeus pedotectum I is short with a pointed apex and pedotectum II consists of a short, curved lobe, whereas in  Sadocepheus pedotectum I is prominent, often considerably expanded laterally and with a large, curved, horn-like apex and pedotectum II is large, laterally projecting and either square or trapezoid or with long, diagonally-sloping anterior margins. In  Hamotegeus the genital setae are extremely long, much longer than the width of each genital plate, representing an autapomorphy within the  Eutegaeoidea (and  H. breviseta exhibits genital neotrichy, with 7-9 pairs of setae), whereas in  Sadocepheus the genital setae are shorter than the width of each genital plate. For these reasons I reject the proposition of Subías (2004) that  Hamotegeus is a junior synonym of  Sadocepheus .</p><p>In the generic composition of the  Compactozetidae, I retain  Compactozetes,  Sadocepheus and  Hamotegeus but establish a new family,  Bornebuschiidae fam. nov. (cf. below) for  Bornebuschia and  Dicrotegaeus . Differences between  Compactozetidae and  Pterozetidae are detailed under the remarks section for the latter family below.</p></div>	https://treatment.plazi.org/id/03A2C77C4654FFFAC79CB05D1246D8FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4657FFFBC79CB5F416D9DD5C.text	03A2C77C4657FFFBC79CB5F416D9DD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes Hammer 1966	<div><p>Compactozetes Hammer, 1966</p><p>Compactozetes Hammer, 1966, p. 76 .</p><p>Type species:  Compactozetes rotoruensis Hammer, 1966</p><p>Diagnosis: the following diagnosis is modified from that of Luxton (1988a). Medium-sized to large oribatid mites (450–1,300 μm); rostrum rounded or acute; apices of lamellae fused with each other and rostrum, forming large paired foramina between lamellae and prodorsum. Lamellar setae located on anterior margin of lamellae or just ventral of it; prodorsum lacking condyles of enantiophyses. Humeral processes with ornate microsculpture of reticulations and striae; apical portion of humeral processes short, not reaching beyond bothridia, medial edge projecting obliquely from dorsosejugal scissure; posterior portion of humeral processes consisting of 2-4 lobes, indicated by thin, semi-transparent cuticle and wavy margin. Notogaster ovoid, with nine pairs of short, smooth setae positioned marginally. Epimeral plates II short and narrow. Pedotectum I with long, stout apical spine; pd II trapezoid or very large, lobe-shaped; discidium lobe-shaped. With six pairs of genital setae, penultimate pair displaced laterally; one pair aggenital setae, two pairs anal setae, three pairs adanal setae. With or without perigenital carina and/or enantiophyses E4. Chelicerae chelate-dentate, of normal proportions. Pre-anal organ oval.</p><p>Remarks.  Compactozetes can be differentiated from  Sadocepheus and  Hamotegeus by the different structure of the lamellae. In  Compactozetes the anterior lamellae are fused medially with each other, folded ventrally and fused with the rostrum, with large foramina either side of the point of fusion. In  Sadocepheus and  Hamotegeus the lamellar cusps are free.</p><p>Two species groups have been identified: ‘rotoruensis’ which includes  C. bundjalung sp. nov.,  C. pirumorpha sp. nov.,  C. duonodulus sp. nov. (cf. below),  C. niger Hammer, 1966 and  C. rotoruensis Hammer, 1966 . In this group, pedotectum II is rectangular with a median indentation. Other members of the genus belong to the ‘zeugus’ species group, which includes  C. zeugus Luxton, 1988b,  C. calderi sp. nov. (cf. below) and  C. goongerah sp. nov. (cf. below). These species have a large blade-like pedotectum I with a prominent, straight spine and pedotectum II is in the form of a large lobed or trapezoid process. Pedotectum II of  C. hastatus Hammer, 1973 was not illustrated or mentioned in the description.</p></div>	https://treatment.plazi.org/id/03A2C77C4657FFFBC79CB5F416D9DD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4656FFF4C79CB094122ED808.text	03A2C77C4656FFF4C79CB094122ED808.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes bundjalung Colloff 2023	<div><p>Compactozetes bundjalung sp. nov.</p><p>(Fig. 28)</p><p>Dimensions. Holotype female length 697 μm, breadth 591 μm. Paratype males (n = 5) mean length 623 μm (range: 566–645 μm); mean breadth 489 μm (range: 480–506 μm). Paratype females (n = 17) mean length 694 μm (range: 656–735 μm); mean breadth 566 μm (range 535–621 μm). Ratio of prodorsum to total length: 0.38 (holotype).</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) only visible in ventral aspect, short, smooth, curved, on short tubercles. Lamellae very broad, with reticulate apical microsculpture, posteromedian margins extending as far as dorsosejugal scissure (Fig. 28a), lacking lamellar cusp, anterior margin truncated, forming curved, transverse apex, sloping ventrally and fused with each other and rostrum adjacent to tutorium, with large foramina between lamellae and prodorsum, visible in ventral view (Fig. 28b). Lamellar seta (le) well-developed, curved medially, smooth, emerging from alveolus on anterior margin of fused lamellae. Interlamellar setae (in) minute, close together lateral to posteromedian mound or bulge. Bothridia corniculate, fused with lamellae. Bothridial seta long, head expanded slightly, with few blunt apical spines.</p><p>Notogaster: ratio of length to breadth: 0.75; notogaster rounded, convex, with reticulate cerotegument laterally, becoming granular in centrodorsal region, with faint curved ridges posteriorly. Humeral processes with short anterior section extending as far as bothridia, lateral margin sub-rectangular, extending posteriorly as far as bases of setae lm, with region of four indented, ovoid and lighter areas of cuticle (Fig. 28a). Lyrifissures ia and im not visible. With nine pairs of extremely short, smooth setiform notogastral setae in marginal position, sub-equal in length; p series not visible in dorsal aspect. Notogastral margin medial of setae p 1 with median slit.</p><p>Ventral aspect: subcapitulum acute; subcapitular setae h extremely long, thin, flagelliform. Tutorium very broad, with two spur-like lobes. Epimeral plates discrete, broadly separated in midline, sub-rectangular to trapezoid, plates I extremely large, plates IV much shorter than I-III; epimeral setation 3-1-3-3 setae very short, sub-equal in length (Fig. 28b). Pedotectum I (pd I) massive, curved, with horn-shaped apex; pd II short, rectangular, with a median indentation; discidium broad, lobe-like. Perigenital carina and enantiophysis E4 present; apical portion of perigenital carina reflexed. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of half length of genital plates; genital plates 85 µm long, with six pairs of short setae, g 1 longer than others, aligned longitudinally; three pairs of very short adanal setae; anal setae barely visible; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates rectangular, 121 µm long. Pre-anal organ (po) oval.</p><p>Etymology. The specific name is a noun in apposition. This species is named for the Bundjalung People of the Richmond Range on whose ancestral land this species is found.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1069,  paratypes: five males, 17 females, ANIC accession no. 53-1071, ANIC 775, litter, closed forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.58333&amp;materialsCitation.latitude=-28.483334" title="Search Plazi for locations around (long 152.58333/lat -28.483334)">Richmond Range State Forest</a>, New South Wales, 28°29’S 152°35’E, 600 m, coll. T. Weir and A. Calder, 13-14.ii. 1983  .   Paratypes: two females, ANIC accession no. 53-1070, ANIC 655 A, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m   ., coll. J. Lawrence and T. Weir, 22- 27.xi.1978.  Lamington National Park .</p><p>Diagnosis.  Compactozetes bundjalung can be distinguished from other species in the genus by the following combination of characters: (1) the relatively long, recurved lamellar setae; (2) the concave transverse apex of the fused lamellae; (3) the posteromedian mound on the prodorsum; (4) the massive, curved pedotectum I with a sharply pointed apex; (5) the rectangular pedotectum II with a median indentation; (6) the extremely long, thin, flagelliform subcapitular setae h; (7) the sinuous, recurved perigenital carina; (8) the presence of enantiophyses E4; (9) the posterior notogastral margin with a median slit.</p><p>Remarks.  Compactozetes bundjalung is morphologically most similar to  C. niger Hammer, 1966 from New Zealand in the shape of the anterior part of the lamellae, the humeral processes and the lateral margin of the notogaster. It differs in the presence of the mound on the posterior part of the prodorsum, the perigenital carinae and enantiophyses E4 and in its far smaller size: 687 μm long, compared with 1,008 μm long for  C. niger .</p></div>	https://treatment.plazi.org/id/03A2C77C4656FFF4C79CB094122ED808	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4659FFF6C79CB5A112A0DCE0.text	03A2C77C4659FFF6C79CB5A112A0DCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes pirumorpha Colloff 2023	<div><p>Compactozetes pirumorpha sp. nov.</p><p>(Fig. 29)</p><p>Dimensions. Holotype female length 880 μm, breadth 721 μm. Paratype males (n = 4) mean length 833 μm (range 829–838 μm), mean breadth 672 μm (range 656–695 μm). Paratype females (n = 9) mean length 912 μm (range 885–928 μm), mean breadth 732 μm (range 695–758 μm). Ratio of prodorsum to total length: 0.41 (holotype).</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) only visible in ventral aspect, short, smooth, curved. Lamellae very broad, with reticulate apical microsculpture, apices fused; anterior lamellar margin truncated, forming concave transverse apex with small sharp points laterally (Fig. 29a); lamellae curving ventrally and fused with rostrum; with large lateral foramina between lamellae and prodorsum, visible in ventral view (Fig. 29b) and corresponding with lighter, ovoid areas of cuticle on dorsal surfaces of lamellae. Tutorium poorly-developed with short, pointed apex. Massive, curved pedotectum I projecting laterally well beyond margin of prodorsum and as far as level of apices of tutorium. Lamellar seta (le) short, thin, smooth, curved laterally on anterolateral margin of lamella. Interlamellar setae (in) setiform, on lamellae close to median margins. Bothridia corniculate, fused with lamellae, projecting slightly beyond posterolateral lamellar margins. Bothridial seta long, head slightly expanded with few apical spines.</p><p>Notogaster: ratio of length to breadth: 0.74; notogaster rounded, convex, with reticulate cerotegument laterally. Humeral processes short, broad, anterior region sub-triangular extending to bothridia, lateral margin extending posteriorly as far as bases of setae lp, with four indented, ovoid lobes indicated by lighter areas of cuticle (Fig. 29a). Lyrifissures ia and im not visible. With nine pairs of short, smooth, setiform notogastral setae in marginal position, l and h series longer than p series which are visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongated; subcapitular setae h thin, longer than a and m. Margins of epimeral plates diffuse, plates broadly separated in midline, sub-rectangular to trapezoid, plates III much smaller than others; epimeral setation 3-1-2-2; epimeral setae short, 4c longer than others (Fig. 29b). Pedotectum I (pd I) massive, with prominent horn-shaped apex, lateral margins strongly convex with small lateral cusp; pd II rectangular, with median indentation; discidium broad with blunt apex; perigenital carina present, enantiophysis E4 absent. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of two-thirds length of genital plates; genital plates 86 µm long, with six pairs of short setae, g 1 slightly longer than others, aligned longitudinally; three pairs of extremely short adanal setae; lyrifissure iad in para-anal position, some distance from margin of anal plate, more than length of iad. Anal plates sub-rectangular, 112 µm long. With shallow, curved ridge posterior of anal plates. Pre-anal organ (po) oval.</p><p>Etymology. The specific name is derived from the Latin pirum, meaning ‘a pear’, referring to the shape of the species in dorsal view.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1072,  paratypes: eight females, three males, ANIC accession no. 53-1073, ANIC 107, moist upland hardwood forest (cf. Invasive Species Council and TierraMar, 2021 for description), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.93333&amp;materialsCitation.latitude=-29.016666" title="Search Plazi for locations around (long 167.93333/lat -29.016666)">Mount Pitt</a>, Norfolk Island, 29°1’S, 167°56’E, 230 m, coll. M.S. Upton, 12.vii.1968  .   Paratype: male, ANIC accession no. 53-1074, ANIC 1025, litter under  Araucaria heterophylla, Red <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.95&amp;materialsCitation.latitude=-29.016666" title="Search Plazi for locations around (long 167.95/lat -29.016666)">Road Track</a>, Norfolk Island National Park, Norfolk Island, 29°01'S 167°57'E, coll. T. A. Weir, 14.xi.1984  .</p><p>Diagnosis.  Compactozetes pirumorpha can be distinguished from other species in the genus by the following combination of characters: (1) the very short, curved rostral setae; (2) the small sharp points on the anterolateral margin of the fused lamellae; (3) the humeral process with a sub-triangular anterior lobe and a slight indentation posterior of it; (4) the notogastral setae of the l, h and p series all short, subequal in length; (5) the very large, broad, curved pedotectum I with a pointed apex; (6) the small cusp on the lateral margin of pedotectum I; (7) the rectangular pedotectum II with a median indentation; (8) the presence of faint perigenital carinae; (9) the shallow, curved ridge posterior of the anal plates.</p><p>Remarks.  Compactozetes pirumorpha belongs to the species group, ‘rotoruensis’, which includes  C. duonodulus sp. nov. (cf. below) to which it is morphologically most similar, including the shape of pedotectum I and pd II; the broad, pointed discidium and the short notogastral setae, sub-equal in length.  Compactozetes pirumorpha differs from  C. duonodulus sp. nov. in the sharply pointed (rather than rounded, tuberculate) lateral apices of the lamellae; the absence of enantiophyses E4 and the shorter humeral processes, extending only to just posterior of the bothridia.</p></div>	https://treatment.plazi.org/id/03A2C77C4659FFF6C79CB5A112A0DCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C465BFFF7C79CB1C81568D86C.text	03A2C77C465BFFF7C79CB1C81568D86C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes calderi Colloff 2023	<div><p>Compactozetes calderi sp. nov.</p><p>(Fig. 30)</p><p>Dimensions. Holotype female length 1,051 μm, breadth 830 μm; paratype female length 1,072 μm, breadth 857 μm. Ratio of prodorsum to total length: 0.4 (holotype).</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) relatively long, smooth, curved, only visible in ventral aspect. Lamellae very broad, with reticulate apical microsculpture, anterior margin truncated, concave (Fig. 30a), curving ventrally, fused with each other and rostrum; with large paired lateral foramina between lamellae and prodorsum visible in ventral view (Fig. 30b), corresponding with lighter, ovoid areas of cuticle on dorsal surface of lamellae. Tutorium with pointed apex, extending beyond margin of lamellae. Massive, curved pedotectum I projecting laterally well beyond margin of prodorsum and anteriorly as far as level of apices of tutorium. Lamellar seta (le) relatively long, stout basally, flagelliform apically. Interlamellar setae (in) short, setiform, smooth, on median margins of lamellae. Bothridia fused with lamellae, projecting slightly beyond posterolateral lamellar margins. Bothridial seta long, bacilliform, with few apical spines. Central region of prodorsum between median lamellar margins and anterior of setae in with area of lighter cuticle, marked by inverted U-shaped margin.</p><p>Notogaster: ratio of length to breadth: 0.77; notogaster rounded, convex, smooth. Humeral process with triangular anterior lobe with blunt apex, microsculpture of wavy ridges, extending to level of bothridia, lateral margin undulating and with shorter secondary lobe and small posterior lobe just anterior of setae lm (Fig. 30a). Lyrifissures ia and im not visible. With nine pairs of notogastral setae in marginal position; l and h series long, curved, smooth, at least four times longer than p series which are not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongated; subcapitular setae h thin, longer than a and m, positioned close to anterior margin of mentum. Epimeral plates diffuse, broadly separated in midline, sub-rectangular to trapezoid, plates III smaller than others; epimeral setation 3-1-2-3; setae relatively short, 4c slightly longer than others (Fig. 30b). Pedotectum I (pd I) massive, curved laterally, with sharp, horn-shaped apex extending as far as apex of tutorium; pd II elongated, lobed, extending beyond lateral notogastral margin; discidium broad, lobed, with blunt apex, larger than pd II; perigenital carina and enantiophysis E4 absent. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of same length as genital plates; genital plates 130 µm long, with six pairs of short setae, g 1 same length as others, g 5 displaced laterally; three pairs of short adanal setae; lyrifissure iad in para-anal position, moderately close to margin of anal plate. Anal plates lozenge-shaped, 170 µm long. Pre-anal organ (po) oval.</p><p>Etymology. This species is named for its co-collector, Dr. Andrew A. Calder, coleopterist and my former colleague at the Australian National Insect Collection, Canberra, who collected many of the Berlesate samples from which I have described new oribatid species.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1075, ANIC 754,  Nothofagus and Sassafras ( Atherosperma moschatum) litter,  Nothofagus moorei forest, Cobark Forest Park, Barrington Tops, New South Wales, 31°59’S, 151°49’E, coll. T. Weir and A. Calder, 15.xi.1981. Paratype: female, ANIC accession no. 53-1076, ANIC 749, litter, closed  Eucalyptus forest, Barrington Tops National Park, Gloucester Road., 30°04’S, 151°41’E, T. Weir and A. Calder, 12-14.ix.1981.</p><p>Diagnosis.  Compactozetes calderi can be distinguished from other species in the genus by the following combination of characters: (1) the flagelliform lamellar setae; (2) the bacilliform bothridial seta with apical spinules; (3) the lighter area of cuticle on the central prodorsum marked anteriorly by an inverted U-shaped margin; (4) the humeral process consisting of a blunt, triangular anterior lobe with a posterior indentation, a second protruding lobe and then a third, smaller, lobe; (5) the relatively long, curved notogastral setae of the l and h series; (6) the very large, broad, curved pedotectum I with a sharp, pointed apex.</p><p>Remarks.  Compactozetes calderi belongs to the ‘zeugus’ species group. It shares with  C. goongerah sp. nov. (cf. below) the relatively long notogastral setae of the l and h series (though those of  C. calderi are longer), the large, blade-shaped pedotectum II and discidium and the triangular blunt anterior lobe of the humeral process. However, the humeral process of the latter species has four lobes rather than three (of which the hindmost is longer and more prominent than the second), a smaller pedotectum I with markedly concave lateral margins, expanded apices to the bothridial setae and very long setae 4c.</p></div>	https://treatment.plazi.org/id/03A2C77C465BFFF7C79CB1C81568D86C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C465AFFF1C79CB5441488DF24.text	03A2C77C465AFFF1C79CB5441488DF24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes goongerah Colloff 2023	<div><p>Compactozetes goongerah sp. nov.</p><p>(Fig. 31)</p><p>Dimensions. Holotype female length 916 μm, breadth 735 μm. Paratype female length 893 μm, breadth 711 μm; paratype male length 865 μm, breadth 664 μm. Ratio of prodorsum to total length: 0.34 (holotype).</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) relatively short, smooth, curved, only visible in ventral aspect. Lamellae very broad, with reticulate microsculpture, anterior margin truncated, with flat, triangular lateral lobes forming curved, concave, transverse apex (Fig. 31a), curving ventrally and fused with each other and rostrum; with large paired lateral foramina, visible in ventral view (Fig. 31b), corresponding with lighter, ovoid areas of cuticle on dorsal surfaces of lamellae. Tutorium well-developed with pointed apex. Curved pedotectum I projecting laterally well beyond margin of prodorsum and anteriorly as far as mid-level of lateral lamellar margins. Lamellar seta (le) short, stout basally, curved. Interlamellar setae (in) short, setiform, smooth, on median margins of lamellae. Bothridia fused with lamellae, projecting slightly beyond posterolateral lamellar margins. Bothridial seta long, curved, with slightly expanded apex with a few spines.</p><p>Notogaster: ratio of length to breadth: 0.82; notogaster rounded, convex, smooth. Anterior lobe of humeral process sub-triangular with blunt apex and reticulate microsculpture, extending to level of bothridia, lateral margin undulating, with shorter secondary lobe, smaller tertiary lobe and terminal projection, with sinuous microsculpture at level of seta la (Fig. 31a). Lyrifissures ia and im not visible. With nine pairs of notogastral setae in marginal position; l and h series relatively long (for  Compactozetes spp.), curved, smooth, setiform, about twice as long as setae of the p series, of which only p 1 are visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongated; setae h thin, longer than setae a and m, positioned near anterior margin of mentum. Epimeral plates diffuse, broadly separated in midline, sub-rectangular to trapezoid, plates III smaller than others; epimeral setation 3-1-3-3; setae short, 4c very long (Fig. 31b). Pedotectum I (pd I) with reticulate microsculpture, concave lateral margins, apex acute; pd II massive, elongated, lobed, extending well beyond notogastral margin; discidium broad, lobed, with blunt apex, smaller than pd II; perigenital carina and enantiophysis E4 absent. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of same length as genital plates; genital plates 103 µm long, with six pairs of short setae, g 1 same length as others, g 5 displaced laterally; three pairs of short adanal setae; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 155 µm long. Pre-anal organ (po) oval.</p><p>Etymology. This species is named for the community of Goongerah on the Brodribb River, adjacent to Errinundra Plateau, in East Gippsland. Goongerah means ‘egg rock’ in the language of the Bidhawal People.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1077,  paratypes: two females, ANIC accession no. 53-1078, moss on Sassafras ( Atherosperma moschatum), rainforest, 1,077 m., Errinundra Saddle, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.85&amp;materialsCitation.latitude=-37.316666" title="Search Plazi for locations around (long 148.85/lat -37.316666)">Errinundra National Park</a>, Victoria, 37°19’S 148°51’E, 1030 m., coll. M.J. Colloff, 29.ix.2009.</p><p>Diagnosis.  Compactozetes goongerah can be distinguished from other species in the genus by the following combination of characters: (1) the flat, triangular lateral lobes on the apex of the lamellae; (2) the humeral process consisting of a blunt, triangular anterior lobe with a posterior indentation, a second protruding lobe, a third smaller lobe and a fourth rectangular lobe (3) the sinuous microsculpture on the fourth humeral lobe; (4) the relatively long, stout, curved notogastral setae; (5) pedotectum I with sharply pointed apices and concave lateral margins; (6) pedotectum II massive, blade-like, extending beyond lateral margin of notogaster.</p><p>Remarks.  Compactozetes goongerah belongs to the ‘zeugus’ species group and is morphologically most similar to  C. calderi, as outlined in the remarks section for that species above.</p></div>	https://treatment.plazi.org/id/03A2C77C465AFFF1C79CB5441488DF24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C465CFFF2C79CB28C14FBDA50.text	03A2C77C465CFFF2C79CB28C14FBDA50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes duonodulus Colloff 2023	<div><p>Compactozetes duonodulus sp. nov.</p><p>(Fig. 32)</p><p>Dimensions. Holotype female length 742 μm, breadth 606 μm. Length of paratype females: 732 and 707 μm, breadth 586 and 561 μm. Ratio of prodorsum to total length: 0.35.</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) short, smooth, curved, only visible in ventral aspect. Lamellae very broad, anterior margin truncated and forming straight, transverse apex with two short blunt protuberances laterally and flat, triangular lateral lobes bearing setae le (Fig. 32a). Lamellae curving ventrally, fused with each other and rostrum, with large paired lateral foramina (Fig. 32b) corresponding with lighter, ovoid areas of cuticle on dorsal surface of lamellae; cuticle on ventral lamellar surface surrounding foramina with series of concentric whorls. Pedotectum I well-developed, lateral margins straight, projecting laterally well beyond margin of prodorsum and anteriorly as far as level of apices of median lamellar margins. Lamellar seta (le) relatively long, flagelliform apically. Interlamellar setae (in) short, setiform, smooth, close to median margins of lamellae. Bothridia fused with lamellae, projecting slightly beyond posterolateral lamellar margins. Bothridial seta long, slightly expanded apically, with few spines</p><p>Notogaster: ratio of length to breadth: 0.8; notogaster rounded, convex, smooth. Anterior lobe of humeral processes triangular with sharp apex and reticulate microsculpture, extending to level just anterior of bothridia; lateral margin of humeral process concave, with two shorter concavities and three small ovoid lobes posteriorly, each with sinuous microsculpture; humeral process terminating posteriorly at level of seta lp (Fig. 32a). Lyrifissures ia not visible. With nine pairs of notogastral setae in marginal position; l and h series short, slightly spiniform, same length as setae of p series, of which p 1 is visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongated; subcapitular setae h thin, longer than a and m, positioned close to anterior margin of mentum. Epimeral plates diffuse, broadly separated in midline, sub-rectangular to trapezoid, plates III much smaller than others; epimeral setation 3-1-3-2; setae relatively short, 4c slightly longer than others (Fig. 32b). Pedotectum I (pd I) large, with reticulate microsculpture and straight lateral margins, with sharply pointed apex, indented at its base; pd II relatively short, sub-rectangular, with median indentation; discidium broad, lobed, with blunt apex; perigenital carina and enantiophysis E4 present. Ventral plate ovoid, markedly broader than long, with marked indentation or ‘shoulder’ at lateral margin of circumpedal carina. Genital and anal plates separated by distance of half length of genital plates; genital plates 97 µm long, with six pairs of short setae aligned longitudinally, g 1 same length as others; three pairs of short adanal setae; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 136 µm long. Pre-anal organ (po) oval.</p><p>Etymology. The specific name, duo nodulus, is Latin for ‘two small knobs’, referring to the pair of small, blunt protuberances on the apices of the lamellae.</p><p>Type designation, material examined and locality data.  Holotype female, ANIC accession no. 53-1079,  paratypes: two females, ANIC accession no. 53-1080, ANIC 460, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. R. J. Kohout, 21.iii.1973.</p><p>Diagnosis.  Compactozetes duonodulus can be distinguished from other species in the genus by the following combination of characters: (1) the pair of small protuberances on the apices of the lamellae; (2) the long, pointed anterior lobe of the humeral process with concave lateral margins; (3) the three smaller humeral lobes, each with wavy microsculpture; (4) the notogastral setae of the l, h and p series short, subequal in length; (5) the cuticle around the foramina on the ventral aspect of the lamellae ornamented with a series of concentric whorls; (6) the long, thin point on pedotectum I with an invagination at its base; (7) the rectangular pedotectum II with a median indentation; (8) the presence of perigenital carinae and enantiophyses E4; (9) the genital setae are all in alignment longitudinally.</p><p>Remarks.  Compactozetes duonodulus is a member of the ‘rotoruensis’ group. It is morphologically most similar to  C. pirumorpha, as detailed in the remarks section for that species above.</p></div>	https://treatment.plazi.org/id/03A2C77C465CFFF2C79CB28C14FBDA50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C465FFFF3C79CB7981439D961.text	03A2C77C465FFFF3C79CB7981439D961.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Compactozetes crenellatus Colloff 2023	<div><p>Compactozetes crenellatus sp. nov.</p><p>(Fig. 33)</p><p>Dimensions. Holotype male length 837 μm, breadth 672 μm. Paratype females (n = 5) mean length: 927 μm (range 916–971 μm); mean breadth 781 μm (range 758–822 μm). Paratype males (n = 3) mean length: 833 μm (range 818–845 μm); mean breadth 677 μm (range 664–695 μm). Ratio of prodorsum to total length: 0.37.</p><p>Description of adult. Prodorsum: rostrum entirely covered by lamellae; rostral setae (ro) relatively long, smooth, curved, only visible in ventral aspect. Lamellae very broad, anterior margin truncated, forming convex apex with two short lateral lobes (Fig. 33a). Lamellae curving ventrally, fused with each other and with prodorsum, with large paired lateral foramina between lamellae and rostrum, visible in ventral view (Fig. 33b), corresponding with lighter, ovoid areas of cuticle on dorsal surface. Lamellar seta (le) short, thin, smooth. Interlamellar setae (in) minute, on lamellae close to median margins, connected by faint transverse line. Bothridia fused with lamellae. Bothridial seta shorter than distance between setae in, with sparse apical spines.</p><p>Notogaster: ratio of length to breadth: 0.85; notogaster rounded, convex, smooth. Humeral process with reticulate microsculpture and rounded anterior lobe with three ridges and crenellated margin, extending anteriorly to level with opening of bothridium; lateral margin of humeral process indented posterior of anterior lobe, with one short lobe and one longer posterior one; humeral process terminating posteriorly at point level with seta lm (Fig. 33a). Lyrifissure ia not visible. With nine pairs of notogastral setae in marginal position; l and h series short, curved, smooth, slightly longer than p series.</p><p>Ventral aspect: subcapitulum acute, elongated; subcapitular setae thin, short, sub-equal in length. Epimeral plates I-III well-defined, plates IV lacking posterior margins, plates III longer (in transverse plane) but much narrower than plates II; epimeral setation 3-1-3-3; setae relatively short, sub-equal in length (Fig. 33b). Pedotectum I (pd I) with very large spine, extending to point level with tutorium, projecting laterally well beyond margin of prodorsum; pd II broad, trapezoid in lateral outline with slight median indentation; discidium lobed, rounded; perigenital carina and enantiophysis E4 absent. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of two-thirds length of genital plates; genital plates 90 µm long, with indentations on anterolateral margins and six pairs of short setae, aligned longitudinally, g 1 same length as others; three pairs of short adanal setae; lyrifissure iad in para-anal position, close to margin of anal plate. Anal plates lozenge-shaped, 135 µm long. Pre-anal organ (po) oval.</p><p>Etymology. The specific name refers to the crenellated margin of the anterior lobe of the humeral process.</p><p>Type designation, material examined and locality data.  Holotype male, ANIC accession no. 53-1081,  paratype: female, ANIC accession no. 53-1082, ANIC 299, wet sclerophyll forest, Mount Donna Buang,  Yarra Ranges National Park, Victoria, 1,060 m, coll. R. W. Taylor and R. J. Bartell, 5.xi.1970  .   Paratypes: four females, three males, ANIC accession no. 53-1083, ANIC 592, leaf litter and decaying log, Cement Creek, 5 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7&amp;materialsCitation.latitude=-37.05083" title="Search Plazi for locations around (long 145.7/lat -37.05083)">Warburton</a>, Victoria, 37°3’3S 145°42’E, coll. J. Lawrence and T. Weir, 18.i.1977  .</p><p>Diagnosis.  Compactozetes crenellatus can be distinguished from other species in the genus by the following combination of characters: (1) the lamellar apex consisting of two short trapezoid lateral lobes; (2) the relatively short bothridial seta with apical spines; (3) pedotectum I with a very large anterior spine extending to a point level with the tutorium; (4) the crenellated anterior lobes of the humeral processes; (5) the broad pedotectum II, trapezoid in lateral outline; (6) the absence of perigenital carinae and enantiophyses E4; (7) the indentations on the anterolateral margins of the genital plates.</p><p>Remarks.  Compactozetes crenellatus belongs to the ‘zeugus’ species group. It is morphologically most similar to  C. goongerah, though in the former species the anterolateral margin of pedotectum II is trapezoid, not lobed and rounded. The anterior margin of the fused lamellae is similar in both species, as is the lateral margin of the humeral process, though lacking the crenelations in  C. goongerah .</p></div>	https://treatment.plazi.org/id/03A2C77C465FFFF3C79CB7981439D961	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C465EFF8CC79CB44A17F8D810.text	03A2C77C465EFF8CC79CB44A17F8D810.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sadocepheus Aoki 1965	<div><p>Sadocepheus Aoki, 1965</p><p>Sadocepheus Aoki, 1965, p. 4 .</p><p>Type species:  Sadocepheus undulatus Aoki, 1965</p><p>Diagnosis. The following diagnosis is modified from those of Luxton (1988a) and Ermilov and Corpuz-Raros (2017).</p><p>Small to large oribatid mites (390–900 μm); rostrum rounded; free lamellar cusps about one-quarter of length of lamellae, frequently projecting beyond rostrum, not fused with rostrum, with or without translamella; lamellar setae emerging from anterior margin of cusps, sometimes flanked by two small, sharp teeth. Bothridia fused with lamellae, projecting laterally. Posterior margin of prodorsum lacking condyles of enantiophyses H and B. Humeral process elongated, with reticulations and striae, originating from lateral margin at widest part of notogaster, with several undulating lobes and trapezoid anterior lobe with broad, pointed apex not reaching beyond bothridia. Concave curve formed between apex of anterior lobe of each humeral process and dorsosejugal scissure. Notogaster ovoid; with nine pairs of marginal or sub-marginal setae emerging from alveoli or tubercles ( S. foveolatus). Epimeral plates II short, narrow. Pedotectum I prominent, often expanded laterally with large, curved spine; pd II large, square, trapezoid or lobed; discidium spine-like or lobed. Genital setae six pairs, shorter than width of genital plate, aligned longitudinally, aggenital setae one pair, anal setae two pairs, adanal setae three pairs; enantiophyses E4 and perigenital carinae absent; posterior margin of hysterosoma lacking mounds; chelicerae chelate-dentate, of normal proportions.</p><p>Remarks. Aoki (1965) defined  Sadocepheus as having a smooth, rounded notogaster; broad, rugose lamellae and a translamella; six pairs of genital setae and five pairs of notogastral setae, later modified to six pairs (Fujita and Fujikawa, 1986, p. 12), referring to the l and h series, i.e. with nine pairs including the p series.</p><p>Compactozetes serratus J. Balogh, 1970 was newly combined to  Sadocepheus by Subías (2020) in the 15 th online update of his list of oribatids of the world. It is important to note this was a taxonomic opinion, not a nomenclatorial act, so is not covered under the International Code of Zoological Nomenclature or the 2012 amendment on electronic publication (International Commission on Zoological Nomenclature, 2012). Subias (2020) is free to propose a new combination in his electronic catalogue and I agree with his opinion.  Sadocepheus serratus has free lamellar cusps; they are not fused anteriorly with each other or with the rostrum and this character state differentiates  Sadocepheus from  Compactozetes which has fused cusps and paired, lateral prodorsal foramina.</p><p>In  S. foveolatus Luxton, 1988b,  S. makarcevae Sitnikova 1975,  S. nortonroyi Ermilov, 2020a,  S. remus sp. nov.,  S. sausai Ermilov and Kalúz, 2021,  S. serratus,  S. tohokuensis Fujikawa, 2003,  S. yakuensis Aoki, 2006 and  S. undulatus the lamellae are broad, and more-or-less parallel and with a free cusp which is straight and transverse apically and may be indented where the lamellar setae emerge. In  S. dubius Hammer, 1979,  S. tohokuensis Fujikawa, 2003,  S. subniger (Ewing, 1917) and  S. dhatiwalalensis Ermilov &amp; Rybalov, 2019 the anterior lamellar cusp is in the form of a convex curve rather than a straight, transverse margin and the lamellae are not parallel but strongly convergent apically. In  S. elevatus Mahunka, 1987 and  S. donvictorianoensis Ermilov &amp; Corpuz-Raros, 2017 the lamellae are strongly convergent and the cusps have indentations where the lamellar setae emerge.</p><p>In their definition of  Sadocepheus, Ermilov and Corpuz-Raros (2017) state there are nine or ten pairs of notogastral setae. Of the 14 species in the genus described to date, all have nine pairs, except for  S. dubius and  S. subniger (Ewing, 1917) in which the number cannot be determined from the descriptions. Ermilov and Corpuz-Raros (2017) described their new species,  S. donvictorianoensis, as having ten pairs (p. 32) but only nine are illustrated. In the definition of  Sadocepheus, these authors also stated there are six to nine pairs of genital setae (p. 29). I determined all species have six pairs except for  S. subniger, for which the number is not stated or figured in the original description by Ewing (1917), though Woolley (1957) illustrated five pairs in his re-description. However, he comments the single specimen is ‘almost black’ (p. 116) and has a heavy cerotegument, so he may have missed a pair.</p></div>	https://treatment.plazi.org/id/03A2C77C465EFF8CC79CB44A17F8D810	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4621FF8EC79CB5D9151FDD38.text	03A2C77C4621FF8EC79CB5D9151FDD38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sadocepheus remus Colloff 2023	<div><p>Sadocepheus remus sp. nov.</p><p>(Fig. 34)</p><p>Dimensions. Holotype male length 667 μm, breadth 576 μm. Paratype male length 667 μm, breadth 576 μm. Paratype females (n = 5) mean length 737 μm (range 712–793 μm), mean breadth 575 μm (range 551–602 μm). Ratio of prodorsum to total length: 0.37 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute, apex not covered by lamellae; rostral setae (ro) straight, thin, visible in dorsal aspect (Fig. 34a). Lamellae broad, more lightly sclerotised oval region centrally and anteriorly, with reticulate cerotegument, apically with a translamella. Lamellar cusps broad, sub-rectangular, extending well beyond rostrum. Lamellar setae short, stout, recurved, smooth. Large, curved pedotectum I projecting laterally well beyond margin of prodorsum and anteriorly almost level with translamella. Interlamellar setae (in) short, straight, smooth, on median margins of lamellae anterior of faint, transverse ridge. Bothridia corniculate, fused with lamellae. Bothridial seta long, head expanded, with sparse apical spines.</p><p>Notogaster: ratio of length to breadth: 0.72; notogaster rounded, convex, with reticulate cerotegument laterally, becoming smooth in centrodorsal region. Humeral processes with short, broad anterior section, heavily notched at base, extending as far anteriorly as bothridia, lateral margin sub-rectangular, extending posteriorly to between bases of setae lm and lp, posterior part with three indented, ovoid, lighter areas of cuticle bearing sinuous microsculpture (Fig. 34a). Lyrifissure ia visible. With nine pairs of thin, relatively long, smooth, setiform notogastral setae, sub-equal in length; p series shorter than others, not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongate; subcapitular setae h slightly longer than a and m. Tutorium broad, well-developed, connected with ventral surface of lamellar cusp. Epimeral plates diffuse, broadly separated in midline, sub-rectangular to trapezoid, plates I extremely large, plates III much smaller than others; epimeral setation 3-1-2-2; setae very short, sub-equal in length (Fig. 34b). Pedotectum I (pd I) massive, with prominent horn-shaped apex; pd II very long, paddle-shaped, with convex anterior margin; discidium lobe-like, rounded. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance of one third of length of genital plates; genital plates 95 µm long, with six pairs of short setae, g 1 slightly longer than others; aligned longitudinally; three pairs of very short adanal setae; lyrifissure iad in para-anal position, close to margin of anal plate, about same as length of iad. Anal plates rectangular, 122 µm long. Pre-anal organ (po) oval.</p><p>Etymology. The specific name  remus is Latin for oar or paddle, referring to the long, paddle-shaped pedotectum II.</p><p>Type designation, material examined and locality data.   Holotype male, ANIC accession no. 53-1084, ANIC 459, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14’S 153°08’E, 920 m, coll. R. W. Taylor, 21.iii.1973  .   Paratypes: one male, one female, ANIC accession no. 53-1085, ANIC 460, litter,  Nothofagus moorei rainforest O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m  ., coll. R. J. Kohout, 21.iii.1973 .   Paratype: female, ANIC accession no. 53- 1086, ANIC 655 A, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. J. Lawrence and T. Weir, 22-27.xi.1978  .   Paratypes: three females, ANIC accession no. 53-1087, ANIC 838, litter, closed forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.73334&amp;materialsCitation.latitude=-30.383333" title="Search Plazi for locations around (long 152.73334/lat -30.383333)">Dorrigo National Park</a>, New South Wales, 30°23'S, 152°44'E, 600 m, coll. L. Hill, 14.vi.1982  .   One paratype female, ANIC accession no. 53-1088, ANIC 775, litter, closed forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.58333&amp;materialsCitation.latitude=-28.483334" title="Search Plazi for locations around (long 152.58333/lat -28.483334)">Richmond Range State Forest</a>, New South Wales, 28°29’S 152°35’E, 600 m, coll. T. Weir and A. Calder, 13-14.ii. 1983  .</p><p>Diagnosis.  Sadocepheus remus can be distinguished from other species in the genus by the following combination of characters: (1) the short, blunt lamellar cusps; (2) the recurved lamellar setae; (3) the apex of the rostrum visible in dorsal view; (4) the light areas of cuticle with wavy microsculpture on the humeral processes; (5) the prominent notch at the base of the anterior lobe of the humeral process; (6) the relatively long notogastral setae of the l and h series; (7) the prominent, wide tutorium connected with the ventral surface of lamellar cusps; (8) the massive pedotectum I with a prominent sharp spine; (9) the elongated, paddle-shaped pedotectum II with a curved anterior margin; (10) the large, lobe-shaped discidium.</p><p>Remarks.  Sadocepheus remus is morphologically quite different from other members of the genus. In  S. dhatiwalalensis,  S. donvictorianoensis Ermilov and Corpuz-Raros, 2017,  S. dubius,  S. foveolatus,  S. makarcevae,  S. sausai,  S. serratus and  S. undulatus pedotectum II is sub-rectangular. Of these species, the discidium is pointed and spine-like in  S. donvictorianoensis,  S. elevatus and  S. sausai . In  S. nortonroyi pedotectum II is sub-circular. In  S. yakuensis Aoki, 2006 and  S. subniger the coxisternal region has not been described but  S. yakuensis has much longer notogastral setae than  S. remus and  S. subniger has much more strongly convergent lamellae and the humeral processes have different morphology.</p></div>	https://treatment.plazi.org/id/03A2C77C4621FF8EC79CB5D9151FDD38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4623FF8FC79CB0B1124CDFC8.text	03A2C77C4623FF8FC79CB0B1124CDFC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bornebuschiidae Colloff 2023	<div><p>Bornebuschiidae fam. nov.</p><p>Type genus:  Bornebuschia Hammer, 1966, p. 75 .</p><p>Diagnosis. Rostrum complex, modified: with one or two incisions medially or with median ridge or tongue-like protuberance and associated lateral lobes; anterior margin transverse, not rounded. Lamellae either very long with pointed free cusps extending considerably beyond rostrum or with blunt free cusps extending just beyond rostrum; lamellar setae positioned markedly sub-apically and laterally. Bothridia free, not fused with lamellae. With well-developed condyles of enantiophyses H on posterior prodorsum and anterior notogaster; anterior condyle, or both condyles, of prodorsal enantiophysis B present, positioned medially of enantiophyses H. Humeral process short, not reaching further than bothridium, rounded apically, emerging from anterolateral margin of notogaster. Subcapitulum narrow, much longer than broad. Notogaster U-shaped; with eight ( Bornebuschia) or nine ( Dicrotegaeus) pairs of setae arranged in three ranks: marginal (p series); sub-marginal (h 2 and h 3), and centrodorsal (la and lm); setae of p series typically same size and shape as other notogastral setae, visible in dorsal aspect: only p 1 occasionally slightly smaller, positioned on ventral notogastral margin. Chelicerae modified: either thin with chelicerate digits but with reduced number of teeth or needle-like, with mobile digit elongated, longer than fixed digit, bearing two minute apical teeth. Perigenital carinae and/or enantiophyses E4 present, enantiophyses V on lateral margins of epimeral plates III and IV. Pedotectum I trapezoid/rectangular in ventral aspect; pd II rectangular; discidium lobed, rounded. Circumpedal carina absent. Six pairs of genital setae, penultimate pair displaced laterally; two or three pairs of adanal setae. Anal plates lozenge-shaped. Pre-anal organ T-shaped ( Bornebuschia) or oval ( Dicrotegaeus).</p><p>Remarks. Luxton (1988b) included  Bornebuschia and  Dicrotegaeus, both confined to New Zealand, within the  Compactozetidae, but highlighted the modified rostrum and fine, needle-like chelicerae in  Bornebuschia nodosa Luxton1988 and  Dicrotegaeus mirabilis Luxton,1988,a synapomorphy,and quite unlike the slim, chelate chelicerae of  Atalotegaeus mensarosi, though this type of modification is found in  Dicrotegaeus mariehammerae Ermilov &amp; Minor 2015 and  D. incurvus Ermilov &amp; Minor 2015 .  Bornebuschia and  Dicrotegaeus are unlike other  Compactozetidae, as detailed above, particularly regarding the disposition of the notogastral setae in three ranks and the morphology of the lamellae, rostrum and subcapitulum and are hereby placed in the new family,  Bornebuschiidae . There are several other synapomorphies for the family, including the lobed or incised rostrum; presence of enantiophyses V; setae le emerging sub-apically, at or near the base of the lamellar cusp and the arrangement of the notogastral setae in three ranks. Luxton (1988b, p. 310 and Fig. 1B) described the subcapitulum of  Bornebuschia nodosa Luxton, 1988b as ‘oval’ with 3-4 pairs of ‘hypostomal setae’ and that of  Dicrotegaeus mirabilis Luxton 1988b as ‘urnlike’ (p. 311, Fig. 3B), i.e. slim and acute, broadening basally and not arched as in other  Eutegaeoidea, and with 4-5 pairs of ‘fine hypostomal setae’. These setae were not mentioned in the original description of  B. peculiaris Hammer, 1966, which did not include a diagram of the ventral aspect. Nor were they mentioned in the descriptions of  D. mariehammerae Ermilov &amp; Minor, 2015 and  D. incurvus Ermilov &amp; Minor, 2015 . If confirmed, subcapitular neotrichy would represent a further synapomorphy of  Bornebuschia and  Dicrotegaeus within the  Eutegaeoidea .  Bornebuschia and  Dicrotegaeus both have the lamellar setae positioned markedly sub-apically and laterally of the apices of the lamellar cusps and are the only known genera in the  Eutegaeoidea with enantiophyses V present.</p></div>	https://treatment.plazi.org/id/03A2C77C4623FF8FC79CB0B1124CDFC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4622FF88C79CB2E11495DC58.text	03A2C77C4622FF88C79CB2E11495DC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterozetidae Luxton 1988	<div><p>Pterozetidae Luxton, 1988</p><p>Pterozetidae Luxton, 1988a, p. 83 .</p><p>Type genus:  Pterozetes Hammer, 1966, p. 78 .</p><p>Diagnosis. The following diagnosis is based on that of Luxton (1988a), modified to accommodate the transfer of  Porrhotegaeus to  Porrhotegaeidae fam. nov. (cf. below).</p><p>Lamellae long, well developed, fused with each other apically, their anterior margin concave and curved or Vshaped, with lateral points, but anterior margin not folded ventrally and fused with rostrum. Bothridia fused with bases of lamellae and prodorsum. With long, blade-like humeral processes extending from lateral margin at widest part of notogaster to point at least level with mid-length of lamellae, reflexed ventrally ( Dudichella), but lacking marginal lobes posterior of main anterior lobe. Interlamellar setae present ( Pterozetes novazealandicus Hammer, 1966) or absent. Notogaster U-shaped, with eight or nine pairs of short, thin, smooth setae; those of l and h series positioned marginally. Pedotectum I large, broad, sub-rectangular in ventral outline; pd II rectangular or pointed; discidium sub-rectangular or ovoid. Anterior margin of pedotectum I and epimere I forming straight or curved transverse tectum covering posterior mentum. Perigenital carinae present ( Dudichella) or absent ( Pterozetes). Enantiophysis E4 present as posterior tubercle only ( Pterozetes lawrencei sp. nov.) or absent ( Dudichella and  Pterozetes novazealandicus). Six pairs of genital setae: aligned longitudinally in  Pterozetes or with g 2 and g 5 displaced laterally in  Dudichella . Three pairs of adanal setae. Anal plates lozenge-shaped.</p><p>Remarks. Norton and Behan-Pelletier (2009) considered  Pterozetidae a junior synonym of  Eutegaeidae due to the apparent lack of diagnostic characters at the family-level. However, the humeral processes of  Pterozetidae originate on the lateral margin of the notogaster at its widest part, well posterior of the dorsosejugal scissure, and are curved and convex along their external margins, forming triangular, blade-like structures extending almost to the rostrum, covering most or all of legs I and II, and quite different from those of the  Eutegaeidae . In her definition of  Pterozetes, Hammer (1966, p. 78) pointed out that the humeral process ‘is different from that of  Eutegaeus ’, in that it is more like a pteromorph than the cuticular projection from the anterior margin of the notogaster found in the latter genus. In  Dudichella, the ventrally curved median margins of the humeral processes overlap with the lateral margins of epimeres I and II, which are broadened and excavated, covering legs I and II (J. Balogh, 1970).</p><p>The main character state that differentiates  Pterozetidae from other families of  Eutegaeoidea is that the anterior parts of the lamellae are fused with each other and their anterior margin is concave and curved or V-shaped, but is not folded ventrally and fused with the rostrum and with lateral foramina as in  Compactozetes . Luxton (1988a, pp. 84, 85) considered the lamellar cusps of  Dudichella are free and not fused, but J. Balogh (1970b, p. 36) described them as ‘medially confluent and thus X-shaped’ and his Fig. 1 clearly shows the cusps as fused in the midline, with a deep V-shaped indentation between the lateral apices. This arrangement is completely different from the free lamellar cusps of the  Eutegaeidae,  Neoeutegaeidae and  Porrhotegaeidae fam. nov. I consider the morphology of the anterior margin of the lamellae a synapomorphy of  Pterozetidae .</p><p>Pterozetidae is morphologically most similar to  Compactozetidae in that the margins of the bothridia are fused with the prodorsum and bases of lamellae and the humeral processes originate on the margin of the notogaster at its widest part. However, they differ from those of the  Compactozetidae in lacking the 2-4 lobes posterior of the apical humeral projection. As mentioned above, the lamellae of  Pterozetidae are quite different in morphology from those of  Compactozetes . Also, the anal plates of  Pterozetidae are lozenge-shaped while those of  Compactozetidae are rectangular.</p><p>Additional diagnostic characters for  Pterozetidae are that epimeres I and II are very broad and long, whereas epimeres III and IV are much narrower and shorter. Pedotectum I is very large with sub-rectangular lateral margins, with an anterolateral point in  Dudichella, and extend to the medial margin of the humeral process ( Pterozetes) or overlap it ( Dudichella). The anterior margin of pedotectum I and epimere I forms a straight or curved transverse tectum covering the posterior part of the mentum and the ventro-lateral margins of the gnathosoma, as in most  Compactozetidae, but this structure is not present in the  Eutegaeidae or  Neoeutegaeidae . I consider there are sufficient character states of diagnostic value to retain  Pterozetidae as a valid family. The interlamellar setae are absent in  Pterozetes lawrencei sp. nov. and  Dudichella but present in  Pterozetes novazealandicus Hammer, 1966 (cf. Luxton, 1988c, Fig. 1A therein).</p><p>In  Dudichella the lamellar setae emerge from the ventral surface of the lamellae, as in  Porrhotegaeus (cf. below).  Dudichella also differs from  Pterozetes in having what J. Balogh (1970b) described as ‘minute, peloptoide’ [sic] chelicerae. It is unclear whether the chelicerae are truly pelopsiform (i.e. expanded basally) or are simply thin, attenuate and chelate-dentate as in  Atalotegaeus mensarosi (cf. above).</p></div>	https://treatment.plazi.org/id/03A2C77C4622FF88C79CB2E11495DC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4625FF88C79CB19012ACDA86.text	03A2C77C4625FF88C79CB19012ACDA86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterozetes , Hammer 1966	<div><p>Pterozetes Hammer, 1966</p><p>Type species:  Pterozetes novazealandicus Hammer, 1966, p. 78 .</p><p>Diagnosis. The following diagnosis is based on those of Hammer (1966) and Luxton (1988a). Small to large oribatid mites (330–950 μm long). Lamellae fused with each other anteriorly in midline, with curved, concave anterior lamellar margin with short lateral spine-like cusps, reticulate microsculpture and short, smooth lamellar setae; lateral margins of lamellae slightly convex, almost parallel; median margins parallel. Tutorium prominent, broad, extending beyond lateral margin of lamella. Bothridia fused with lamellae; bothridial seta long, thin, smooth, pointed. Humeral process long, very broad at level of dorsosejugal scissure, pointed apically, extending from lateral margin of notogaster at widest part to level of tutorium, with extensive microsculpture of striations and reticulations, lacking posterior lateral lobes. Notogaster U-shaped, with nine pairs of short setae positioned marginally. Pedotectum I large, sub-rectangular; pedotectum II rectangular; discidium lobe-shaped or sub-rectangular. Circumpedal carina present. With five or six pairs of genital setae aligned longitudinally; three pairs of adanal setae. Anal plates lozenge-shaped.</p><p>Remarks. Details of the taxonomic status of  Pterozetes are given in the remarks section on  Pterozetidae above. The genus contains two species,  P. novazealandicus from New Zealand and  P. lawrencei sp. nov. from Australia.</p></div>	https://treatment.plazi.org/id/03A2C77C4625FF88C79CB19012ACDA86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4625FF8AC79CB629167FDF24.text	03A2C77C4625FF8AC79CB629167FDF24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterozetes lawrencei Colloff 2023	<div><p>Pterozetes lawrencei sp. nov.</p><p>(Fig. 35)</p><p>Dimensions. Holotype female length 838 μm breadth 677 μm. Paratype female lengths 808 μm, 830 μm; breadths 667 μm, 656 μm. Ratio of prodorsum to total length: 0.41 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute, apex almost covered by lamellae; rostral setae (ro) curved, thin, smooth, not visible in dorsal aspect (Fig. 35b). Lamellae narrow, fused anteriorly in midline, concave anterior margin bearing reticulate microsculpture and two sharply-pointed lateral apices, lateral margins almost straight, parallel (Fig. 35a). Lamellar setae short, thin, smooth, on anterior margin. Median area of lighter cuticle and two long, thin lighter oval regions lateral of median area. Interlamellar setae absent. Bothridia rounded projections, fused with lamellae. Bothridial seta long, thin, smooth, pointed.</p><p>Notogaster: ratio of length to breadth: 0.75; notogaster U-shaped, convex. Humeral processes long, extending as far as apex of tutorium, very broad at level of dorsosejugal scissure, with slight notch just anterior of seta la, anterior section broad, curved apically, with short, blunt terminal tooth and striated and reticulate microsculpture (Fig. 35a). Lyrifissures ia not visible. With nine pairs of thin, short, smooth, setiform setae positioned marginally, l and h series sub-equal in length, p series shorter than others, not visible in dorsal aspect.</p><p>Ventral aspect: subcapitulum acute, elongate, covered by tectum basally; subcapitular setae h slightly longer than a and m. Chelicerae chelate-dentate, of normal proportions. Tutorium very broad, apex blunt. Epimeral plates I very broad, trapezoid, lateral margins diffuse, broadly separated in midline; plates II and III very narrow, plates IV lacking posterior margin and setae; epimeral setation 3-1-3-2; setae very short, sub-equal in length (Fig. 35b). Pedotectum I (pd I) well-developed, sub-rectangular in ventral outline, with short, blunt apex; pd II rectangular in outline with medial indentation; discidium curved, sub-rectangular with posterior tooth; perigenital carina absent, posterior tubercle of enantiophysis E4 present. Ventral plate ovoid, markedly broader than long. Genital and anal plates separated by distance same as length of genital plates; genital plates 84 µm long, with five pairs of short, curved setae, g 1 positioned markedly posterior of anterior margin; setae aligned longitudinally; three pairs of extremely short adanal setae; lyrifissure iad in para-anal position, some distance from margin of anal plate, more than length of iad. Anal plates rectangular, 145 µm long. Pre-anal organ (po) oval with cylindrical apex.</p><p>Etymology. This species is named in honour of its co-collector, Dr John F. Lawrence, my former colleague and curator of  Coleoptera at the Australian National Insect Collection, Canberra.</p><p>Type designation, material examined and locality data.   Holotype female, ANIC accession no. 53-1055, ANIC 592, leaf litter and decaying log, Cement Creek, 5 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7&amp;materialsCitation.latitude=-37.05083" title="Search Plazi for locations around (long 145.7/lat -37.05083)">Warburton</a>, Victoria, 37°3’3S 145°42’E, coll. J. Lawrence and T. Weir, 18.i.1977  .   Paratypes: two females, ANIC accession no. 53-1056 &amp; 1057, ANIC 297, Moss and litter,  Nothofagus cunninghami rainforest, Cumberland Creek Valley, nr. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Picnic Area</a> [Cumberland Memorial Scenic Reserve], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.86667&amp;materialsCitation.latitude=-37.566666" title="Search Plazi for locations around (long 145.86667/lat -37.566666)">Yarra Ranges National Park</a>, Victoria. 37°34’S 145°52’E, 920 m  ., coll. R. W. Taylor and R. J. Bartell, 4.xi.1970 .</p><p>Diagnosis.  Pterozetes lawrencei has the following distinctive character states: (1) the median area of lighter prodorsal cuticle and two long, thin lighter oval regions lateral of the median area; (2) the absence of interlamellar setae; (3) the apically curved humeral processes with a short terminal tooth and striated and reticulate microsculpture; (4) the broad tutorium with blunt apices; (5) the broad, trapezoid epimeral plates I; (6) the very narrow epimeral plates II and III; (7) the rectangular discidium with a posterior tooth; (8) five pairs of genital setae with the first pair well posterior of the anterior margin of the genital plate; (9) the oval pre-anal organ with a cylindrical apex.</p><p>Remarks. The humeral processes of  P. lawrencei are different from those of  P. novazealandicus which are broad basally with a triangular extension that is folded ventrally (Hammer, 1966, Plate XXXV, Fig. 106a). However, both species have extensive reticulate and striate microsculpture on the humeral processes. The anterior margins of the lamellae of  P. lawrencei are much more strongly concave, the lateral teeth are more prominent and the species has five pairs of genital setae rather than six. The internal margins of the anterior parts of the humeral processes of  P. novazealandicus are concave and the apex is sharply pointed, not convex with a terminal tooth as in  P. lawrencei . Hammer did not illustrate the full complement of notogastral, epimeral and anal setae or the lateral margins of the coxisternum in  P. novazealandicus because the specimen she examined was heavily sclerotised, so it cannot be confirmed whether  P. novazealandicus shares character states related to these structures without re-examination of the type material.</p></div>	https://treatment.plazi.org/id/03A2C77C4625FF8AC79CB629167FDF24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4627FF8AC79CB28D1447DB2A.text	03A2C77C4627FF8AC79CB28D1447DB2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porrhotegaeidae Colloff 2023	<div><p>Porrhotegaeidae fam. nov.</p><p>Type genus:  Porrhotegaeus J. Balogh &amp; Mahunka, 1966, p. 557 .</p><p>Diagnosis. Lamellae broad, well developed, ovoid, with free cusps bearing one or two teeth, lacking translamella. Bothridia free, not fused with lamellae. With long blade-like humeral processes extending from lateral margin of notogaster at widest part to point mid-way along lamellae, lateral margin reflexed ventrally; humeral processes lacking lobes posterior of the main anterior structure. Tutorium broad, with a well-developed, curved point. Prodorsum waisted posterior of bothridium. Notogaster U-shaped, surface flat or concave, never convex; margins elevated above level of central notogaster; with eight or nine pairs of long notogastral setae positioned marginally and pointed dorsally. Pedotectum I sub-rectangular; pedotectum II sub-rectangular or lobe-like; discidium pointed. Anterior margin of pedotectum I and epimere I forming straight or curved transverse tectum covering posterior mentum. Circumpedal carina and enantiophyses E4 present; with or without perigenital carinae. Six pairs of genital setae aligned longitudinally, three pairs of adanal setae. Anal plates lozenge-shaped. Pre-anal organ T-shaped. Nymphs with well-developed free lamellar cusps and lateral notogastral setae emerging from long, fine apophyses fused with gastronotum; lateral setae flanked by elongated, flat scales with strongly serrated margins.</p><p>Remarks.  Porrhotegaeidae is established for  Porrhotegaeus because the genus shares few character states with  Dudichella and  Pterozetes which Luxton (1988a) placed with  Porrhotegaeus in the  Pterozetidae . The genus clearly does not fit within  Eutegaeidae, where it was placed by J. and P. Balogh (1992) and Subías (2004) (Table 1) although it shares with  Eutegaeidae the prodorsum markedly waisted posterior of the bothridia. The establishment of a separate family for  Porrhotegaeus is justified based on a series of autapomorphies including the broad, ovoid lamellae with pointed, free cusps; the flat or concave surface of the notogaster with a raised margin; the reflexed humeral processes and the long notogastral setae positioned marginally that are curved at their bases and pointed dorsally. Furthermore, the immatures are unique among those known in the  Eutegaeoidea in having strongly-developed lamellar cusps projecting beyond the rostrum, and the notogastral setae c 3 and those of the l, h and p series in the nymphs emerge from extremely long, fine apophyses that are part of the gastronotum and which are flanked by elongated, flat scales with strongly serrated margins.</p></div>	https://treatment.plazi.org/id/03A2C77C4627FF8AC79CB28D1447DB2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4627FF8BC79CB6BE1349DF00.text	03A2C77C4627FF8BC79CB6BE1349DF00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porrhotegaeus J. Balogh & Mahunka 1966	<div><p>Porrhotegaeus J. Balogh &amp; Mahunka, 1966</p><p>Porrhotegaeus J. Balogh &amp; Mahunka, 1966, p. 557 .</p><p>Type species:  Porrhotegaeus ornatus J. Balogh &amp; Mahunka, 1966</p><p>Diagnosis. The following diagnosis is modified from that of Luxton (1988a): large oribatid mites (&gt;950 μm long); rostrum rounded; lamellar cusps one-fifth length of lamellae, widely separated from each other, with an apical tooth or pair of teeth, lacking a translamella; lamellar setae emerging from ventral surfaces of lamellar cusps. Interlamellar setae absent; posterior margin of prodorsum lacking enantiophyses H, though anterior condyles of enantiophyses B present in  P. ornatus . Bothridia long, curved; bothridial setae long, setiform or bacilliform, smooth. Humeral processes reflexed ventrally, with fine striae, originating from lateral regions of notogaster at same level as posterior margin of genital plates, extending beyond bothridia but not as far as bases of lamellar cusps. With nine pairs of notogastral setae (complete l, h and p series), eight in  P. herminae, originating marginally and directed vertically. Central part of notogaster concave, flat or with slight medial mound, notogastral margin forming upturned lip giving entire notogaster concave appearance. Chelicerae chelate-dentate, not thin or modified. Pedotectum II well-developed, broad, pointed or nearly square. With six pairs of genital setae, three pairs of adanal setae; with or without enantiophyses E4 and perigenital carinae. Adanal fissurae (iad) some distance from anal plates.</p><p>Remarks. J. Balogh and Mahunka (1966) mentioned the presence of nine pairs of notogastral setae in the marginal position. There is a full complement of setae in the h, l and p series except in  P. herminae which has only eight pairs, though nine may well be present. Two important character states of  Porrhotegaeus that were overlooked in the definitions by J. Balogh and Mahunka (1966) and Luxton (1988a) are that the central part of the notogaster is generally flat or concave (saucer shaped), with the margins forming a distinct elevated rim and the lamellar setae arise from the ventral surfaces of the lamellae, a character state shared with  Dudichella .</p></div>	https://treatment.plazi.org/id/03A2C77C4627FF8BC79CB6BE1349DF00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C4626FF86C79CB2A9131DDCA8.text	03A2C77C4626FF86C79CB2A9131DDCA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porrhotegaeus githabul Colloff 2023	<div><p>Porrhotegaeus githabul sp. nov.</p><p>(Figs. 36–38)</p><p>Dimensions. Holotype male: length 1,019 μm, breadth 788 μm; paratype male: length 1,021 μm, breadth 790 μm; paratype females: length 1,098, 1,058 μm, breadth 814, 802 μm; paratype tritonymph: length 922 μm, breadth 704 μm; paratype deutonymph: length 581 μm, breadth 455 μm.</p><p>Description of adult. Prodorsum: rostrum acute, rounded; rostral setae (ro) short, straight, smooth. Lamellae broad, with fine striae, lateral margins strongly convex, serrated; cusps broadly separated medially, with a single apical tooth; lamellar setae (le) long, curved, with spinose ornamentation, originating on ventral surface of lamellar cusp close to margin of rostrum (Fig. 36a). Bothridia long, corniculate, extending beyond lateral margins of lamellae, with marked indentation anterior of bothridium; bothridial seta long, smooth, pointed.</p><p>Notogaster: ratio of length to breadth: 0.76; notogaster rounded, flat, smooth, without obvious cerotegument. Humeral process long, with striate microsculpture becoming reticulate basally, pointed, reflexed ventrally, serrated, lateral margin slightly concave basally; medial margins convex: incised and folded laterally at their bases (Fig. 36a). Lyrifissures ia and im not visible. With nine pairs of long, stout, setiform, marginal setae (with complete l, h and p series) bearing fine spines, projecting vertically; setae p 2 and p 3 shorter than l and h series, but not markedly so; setae p 1 positioned dorsally and same length as l and h series.</p><p>Ventral aspect: subcapitulum acute, narrow; subcapitular setae h extremely long, thin; a and m shorter. Chelicerae chelate-dentate and of normal proportions. Tutorium broad, curved, with sharp apex (Fig. 36b). Epimeral plates not discrete in midline, sub-rectangular to trapezoid, plates I and II large, plate I with medial incision, plates III and IV fused, much smaller than others; epimeral setation 3-1-3-2; setae very short, sub-equal in length except much longer 4c. Pedotectum I (pd I) well-developed, apex sub-rectangular pd II with longitudinal margin convex, inflated, posterior margin concave; pd I and pd II with reticulate microsculpture; discidium curved, blunt apically; and enantiophysis E4 present, perigenital carina absent. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of third of length of genital plates; genital plates 140 µm long, with six pairs of short setae sub-equal in length, g 5 displaced laterally; three pairs of long, thin adanal setae; lyrifissure iad in para-anal position, some distance from margin of anal plate, about same as length of iad. Anal plates lozenge-shaped, 183 µm long. Pre-anal organ (po) T-shaped.</p><p>Description of tritonymph, deutonymph and nymphal scalps. Tritonymph prodorsum: Rostrum acute; rostral setae with spinose ornamentation, on short tubercles adjacent to medial margin of lamellar cusps, extending well beyond cusps; lamellar setae long, curved medially, with spinose ornamentation, emerging from ventral surface of lamellar cusp close to margin of subcapitulum; anterolateral margin of lamellar cusps with small spinose projections, posterolateral margin smooth (Fig. 37). Central prodorsum smooth, porose; setae in small, thick, spinose. Bothridia long, curved, tubular, projecting laterally well beyond prodorsal margin. Bothridial seta long, thin, bacilliform.</p><p>Tritonymph gastronotum: U-shaped, broader than long. Dorsosejugal scissure transverse, with short, smooth setae c 1 and c 2 on short tubercles. With 12 pairs of notogastral setae: setae c 3, the l and h series and p 1 long, straight, with spinose ornamentation, emerging from elongated apophyses of gastronotum and bear elongate lateral scales with strongly serrated margins and fine venation (Fig. 37); setae p 2 and p 3 short, curved, positioned ventrally, p 2 with spinose ornamentation.</p><p>Tritonymph ventral aspect: microsculpture of ventral plate with small sparse tubercles (Fig. 38a); epimeral setal formula 3-1-3-3; with four pairs of genital setae, one pair of aggenital setae, two pairs of anal setae and three pairs of adanal setae.</p><p>Deutonymph gastronotum: with 11 pairs of setae: complete c and l series, h 2 and h 3; h 1 absent; c 1 and c 2 short, smooth, others long, with spinose ornamentation, on long apophyses with lateral serrated scales (Fig. 38a); p series short smooth positioned ventrally; p 3 represented only by their alveoli (Fig. 38b).</p><p>Deutonymph ventral aspect: ventral plate smooth; epimeral setal formula 3-1-2-2; with three pairs of genital setae, one pair of aggenital setae, two pairs of adanal setae; anal setae absent (Fig. 38b).</p><p>Ontogenetic formula of gastronotic setae (including one larval and two protonymphal and deutonymphal setae not present on scalps):?-(11,11,12)-9.</p><p>Etymology. The specific name is a noun in apposition. The species is named for the Githabul People of the Kyogle, Woodenbong and Tenterfield areas of north-eastern New South Wales on whose traditional land the species occurs.</p><p>Type designation, material examined and locality data.  holotype male, ANIC accession no. 53-1089,   paratypes: one male, two females, ANIC accession no. 53-1090, ANIC 775, litter, closed forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.58333&amp;materialsCitation.latitude=-28.483334" title="Search Plazi for locations around (long 152.58333/lat -28.483334)">Richmond Range State Forest</a>, New South Wales, 28°29’S 152°35’E, 600 m., coll. T. Weir and A. Calder, 13-14.ii.1983  .   Paratype tritonymph, ANIC accession no. 53-1091, ANIC 655 A, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. J. Lawrence and T. Weir, 22- 27.xi.1978  .   Paratype deutonymph, ANIC accession no. 53-1092, ANIC 459, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.13333&amp;materialsCitation.latitude=-28.233334" title="Search Plazi for locations around (long 153.13333/lat -28.233334)">Lamington National Park</a>, Queensland, 28°14’S 153°08’E, 920 m, coll. R. W. Taylor, 21.iii.1973  .</p><p>Diagnosis.  Porrhotegaeus githabul can be distinguished from other species in the genus by the following combination of characters: (1) the striate lamellae with serrated margins; (2) the lamellar cusp with a single tooth; (3) the marked indentation anterior of bothridium at the base of the lamella; (4) the humeral processes with serrated medial margins; (5) the long notogastral setae p 2 and p 3; at least half the length of p 1; (6) the relatively small pedotectum II, not projecting markedly beyond the margin of the ventral plate; (7) the very long, smooth epimeral setae 4c; (8) the curved, pointed discidium with a blunt apex.</p><p>Remarks.  Porrhotegaeus githabul is morphologically most similar to  P. ornatus in having striated lamellae, a single apical tooth on the lamellar cusp and long setae p 2 and p 3. However, in  P. ornatus pedotectum II is large and rectangular, the lamellae reticulate, the lamellar setae smooth and the lateral margins of the tutoria serrated. Also, the discidium of  P. githabul is curved, not straight, the anterior condyles of enantiophyses B and the perigenital carinae are absent, the lateral margins of the lamellae are toothed, not smooth, and the medial margin of the lamellar cusp is convex, not concave.</p><p>The association of the tritonymph and deutonymph with the adult is based, in part, on their presence in the same samples. Adults of  P. githabul and  P. ornatus occur at Lamington National Park where the tritonymph and deutonymph of the former species were also found and it is possible they belong to  P. ornatus . However, the tritonymph in Fig. 38 shares with the adults of  P. githabul the sharply pointed lamellar cusps with strongly toothed borders and convex medial and lateral margins. The morphology of the lamellae is quite different from those of the adults of  P. ornatus .</p><p>A tritonymph of a  Porrhotegaeus sp., from Lamington National Park is illustrated in the poster, Mites of the Rainforest (Walter et al., n.d.) and by Walter (2004, p. 230). The morphology of the lamellae is different from the tritonymph of  P. githabul, being broader, in a more posterior position on the prodorsum and the margins have short, sparse serrations. It is probable the illustrations are of the tritonymph of  P. ornatus .</p></div>	https://treatment.plazi.org/id/03A2C77C4626FF86C79CB2A9131DDCA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C462BFF87C79CB00116BAD848.text	03A2C77C462BFF87C79CB00116BAD848.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porrhotegaeus catherinae Colloff 2023	<div><p>Porrhotegaeus catherinae sp. nov.</p><p>(Fig. 39)</p><p>Dimensions. Holotype male length 980 μm, breadth 718 μm. Paratype male length 896 μm, breadth 678 μm. Paratype females (n = 3): mean length 1,097 μm (range 1,088 –1,103 μm); mean breadth 753 μm (range 735–781 μm). Ratio of prodorsum to total length: 0.39 (holotype).</p><p>Description of adult. Prodorsum: rostrum acute, prominent, anterior margin transverse; rostral setae (ro) long, with spinose ornamentation, on short tubercles on lateral margin of rostrum. Lamellae broad, with reticulate microsculpture, lateral margins strongly convex, smooth; cusps broadly separated medially, with two broadly-separated apical teeth, bases incised medially; lamellar setae (le), long, curved, with spinose ornamentation, emerging from ventral surface of lamellar cusp (Fig. 39a). Bothridia long, corniculate, extending beyond lateral margins of lamellae, with slight fold anterior of bothridium; bothridial seta long, smooth, bacilliform.</p><p>Notogaster: ratio of length to breadth: 0.85; notogaster rounded, flat, smooth, without obvious cerotegument; dorsosejugal scissure transverse, incomplete medially. Humeral process long, with reticulate microsculpture, pointed, reflexed ventrally, margins smooth, lateral margin slightly concave basally; medial margins convex: incised and folded over basal part of humeral process (Fig. 39a). Lyrifissures ia and im not visible. With nine pairs of long, stout, marginally-positioned setae with fine spines, their bases projecting vertically; with complete l, h and p series (h 1 is present); setae la positioned on central part and lm on basal part of humeral process; setae p 2 and p 3 markedly shorter than other setae; setae p 1 positioned dorsally and same length as l and h series.</p><p>Ventral aspect: subcapitulum acute, narrow; subcapitular setae h extremely long, thin; a and m shorter. Chelicerae chelate-dentate and of normal proportions. Tutorium broad, curved, with sharp apex, margin serrated (Fig. 39b). Epimeral plates not discrete in midline, sub-rectangular to trapezoid, plates I and II large, plates III and IV fused, much smaller than others; epimeral setation 3-1-3-3; setae well-developed, 1b longer than others. Apex of pedotectum I (pd I) right-angled; pd II broad, lobed, with blunt point, protruding laterally; discidium curved, acutely pointed apically; posterior tubercles of enantiophysis E4 present, perigenital carina absent. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of third of length of genital plate; genital plates 97 µm long, with six pairs of short setae sub-equal in length, g 4 displaced laterally; three pairs of short thin adanal setae; lyrifissure iad in para-anal position, some distance from anal plate, more than length of iad. Anal plates lozenge-shaped, 120 µm long. Pre-anal organ (po) T-shaped.</p><p>Etymology. This species is named in honour of my friend and former colleague, Cate Lemann, in honour and recognition of her many years of dedicated service to the Australian National Insect Collection.</p><p>Type designation, material examined and locality data.  Holotype male, ANIC accession no. 53-1093,  paratypes: one male, four females, ANIC accession no. 53-1094, leaf litter, under  Syzygium fullagarii (scalybark), closed forest, 100 m east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=159.08112&amp;materialsCitation.latitude=-31.552776" title="Search Plazi for locations around (long 159.08112/lat -31.552776)">Soldier Creek</a>, Lord Howe Island, 31°33’10”S, 159° 4’52”E, coll. I. Hutton and K. Lees, 8.vi.2003.</p><p>Diagnosis.  Porrhotegaeus catherinae can be distinguished from other species in the genus by the following combination of characters: (1) the transverse rostrum; (2) the very long rostral setae; (3) the bases of the lamellar cusps with medial incisions; (4) the lamellar cusps with two broadly-separated apical teeth; (5) the reticulate microsculpture on the lamellae; (6) the reticulate microsculpture on the humeral process; (7) the notogastral setae la and lm positioned near the centre and at the base of the humeral process, respectively, with a large gap between lm and lp; (8) the very short setae p 2 and p 3; (9) the broad tutoria with serrated margins; (10) the large, broad pedotectum II with a blunt, pointed apex.</p><p>Remarks.  Porrhotegaeus catherinae is not particularly morphologically similar to  P. githabul,  P. ornatus or  P. herminae, particularly in the morphology of the lamellae and lamellar cusp, though it shares with  P. herminae the large, leaf-shaped pedotectum II, though lacks the faint perigenital carinae and posterior tubercles of enantiophyses E4 of the latter species.</p></div>	https://treatment.plazi.org/id/03A2C77C462BFF87C79CB00116BAD848	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C462AFF81C79CB561159CDC8B.text	03A2C77C462AFF81C79CB561159CDC8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Porrhotegaeus ornatus J. Balogh & Mahunka 1966	<div><p>Porrhotegaeus ornatus J. Balogh &amp; Mahunka, 1966</p><p>(Fig. 40)</p><p>Porrhotegaeus ornatus J. Balogh &amp; Mahunka, 1966, p. 557 .</p><p>Dimensions. Holotype female length 1,020 μm, breadth 806 μm. Non-type material: females (n = 8): mean length 1,111 μm (range 1,082 –1,178 μm); mean breadth 867 μm (range 806–909 μm); males (n = 3): mean length 1,034 μm (range 1,024 –1,049 μm); mean breadth 849 μm (range 844–858 μm).</p><p>Redescription of adult. Prodorsum: rostrum acute, rounded; rostral setae (ro) long, thin, smooth, on short tubercles. Lamellae broad, with fine striate microsculpture dorsally, becoming reticulate ventrally, lateral margins strongly convex; cusps broadly separated medially, medial edge concave, with single sharp apical tooth; lamellar setae (le), relatively short, thin, smooth, emerging from ventral surface of lamellar cusp near apex (Fig. 40a). Bothridia long, corniculate, extending beyond lateral margins of lamellae, with slight fold anterior of bothridium; bothridial seta long, smooth, bacilliform, pointed. Posterior margin of prodorsum with anterior condyle of postbothridial enantiophysis B.</p><p>Notogaster: ratio of length to breadth: 0.79; notogaster flat, smooth, without obvious cerotegument. Humeral process long, with striate microsculpture, pointed, reflexed ventrally, lateral margin straight; medial margin strongly serrated, convex: incised and folded over cuticle of basal part of process (Fig. 40a). Lyrifissures ia and im not visible. With nine pairs of long, marginally-positioned setae with fine spines, projecting vertically; with complete l, h and p series (h 1 is present); setae p 2 and p 3 smooth, p 2 as long as h series, p 3 shorter; p 1 positioned dorsally and same length as l and h series.</p><p>Ventral aspect: subcapitulum elongated, narrow; subcapitular setae h extremely long, thin; a and m shorter. Chelicerae chelate-dentate and of normal proportions. Tutorium broad, curved, with sharp apex, margin serrated, microsculpture reticulate (Fig. 40b). Epimeral plates not discrete in midline, sub-rectangular to trapezoid, plates I and II large, plates III extremely narrow, plates IV sub-ovoid, smaller than others; epimeral setation 3-1-3-2; setae well-developed, 1b longer than others. Apex of pedotectum I (pd I) horn-shaped; pd II broad, rectangular, protruding laterally; pd I and II with reticulate microsculpture; discidium straight, acutely pointed apically; faint perigenital carina (pc) and posterior tubercles of enantiophysis E4 present. Ventral plate ovoid, broader than long. Genital and anal plates separated by distance of a third of length of genital plates; genital plates 119 µm long, with six pairs of short setae sub-equal in length, aligned longitudinally; three pairs of long, thin adanal setae; lyrifissure iad in para-anal position, some distance from anal plate, more than length of iad. Anal plates lozenge-shaped, 136 µm long. Pre-anal organ (po) T-shaped.</p><p>Material examined and locality data.  Holotype female, ANIC accession no. 53-000665, Murwillumbah, New South Wales, coll. J. Balogh, 7.iii.1965 .   Non-type material: three females, ANIC 460, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.06667&amp;materialsCitation.latitude=-28.383333" title="Search Plazi for locations around (long 153.06667/lat -28.383333)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. R. J. Kohout, 21.iii.1973.   Four females, ANIC 655 A, litter,  Nothofagus moorei rainforest, O’Reilly’s Rainforest Retreat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.06667&amp;materialsCitation.latitude=-28.383333" title="Search Plazi for locations around (long 153.06667/lat -28.383333)">Lamington National Park</a>, Queensland, 28°14'S, 153°08'E, 920 m., coll. J. Lawrence and T. Weir, 22- 27.xi.1978.   Two females, one male, ANIC 771, litter under  Nothofagus moorei, Wiangaree <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.06667&amp;materialsCitation.latitude=-28.383333" title="Search Plazi for locations around (long 153.06667/lat -28.383333)">State Forest</a>, New South Wales, 28°23’S, 153° 4’E, 1,050 m., coll. T. Weir and A. Calder, 10-12.ii.1983.  One female, ANIC 772, same data as ANIC 771.</p><p>Remarks:  Porrhotegaeus ornatus has some morphological similarities with  P. githabul, as outlined above. However, it and  P. herminae share the faint perigenital carinae which are absent in the two other  Porrhotegaeus species. Porrhotegaeus ornatus is the only member of the genus with the anterior condyles of enantiophyses B. Further,  P. ornatus differs from  P. herminae in that pedotectum II in the latter species is extremely large, leaf-shaped and striated, whereas that in  P. ornatus is rectangular and reticulate. Also, in  P. herminae the adanal setae are much shorter than those of  P. ornatus, the lamellar cusps lack an apical tooth and the humeral processes are not striated.</p><p>J. Balogh and Mahunka (1966, Fig. 5 therein) illustrated  P. ornatus with a translamella. However, the holotype lacks a translamella, consistent with the other three species in the genus. Also, the serrated medial margins of the humeral processes, the anterior condyles of enantiophyses B and the faint perigenital carinae were not shown in the original description and the setae of the l and h series were illustrated as emerging from the ventral surface of the notogastral margin instead of the dorsal surface.</p><p>Porrhotegaeus ornatus was recorded at Murwillumbah and Barrington Tops, New South Wales, and at Beechmont, Queensland, by Hunt et al. (1998b).</p></div>	https://treatment.plazi.org/id/03A2C77C462AFF81C79CB561159CDC8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
03A2C77C462CFF82C79CB02512B8D908.text	03A2C77C462CFF82C79CB02512B8D908.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutegaeoidea Balogh 1965	<div><p>A key to the families and genera of  Eutegaeoidea</p><p>1 Anterior region of humeral process in the form of a short, shallow triangle not extending as far as bothridia (Figs. 41n)................................................................................  Cerocepheidae …  Cerocepheus</p><p>- Anterior region of humeral process long, pointed apically, extending anteriorly as far as bothridium (Figs. 41f–j) or to point between bothridia and lamellar apices (Figs. 41 a-e, k-m)...................................................... 2</p><p>2 Humeral process originating on anterolateral margin of notogaster (Figs. 41 a-e).................................... 3</p><p>- Humeral process originating on margin of notogaster at its widest part (Figs. 41 k-m)................................ 7</p><p>3 Humeral process waisted basally (Fig. 41 a-c); pedotectum I sub-rectangular in lateral outline; pedotectum II pointed or sub-rectangular; discidium pointed (Figs. 43 a-c)....................................................  Eutegaeidae …4</p><p>- Humeral process broad basally (Fig. 41d, e); pedotectum I sub-rectangular in lateral outline; pedotectum II lobed or sub-rectangular; discidium lobed (Fig. 43d, e)...........................................  Neoeutegaeidae fam. nov. ...6</p><p>4 Humeral process strongly curved, with distinct basal notch (Fig. 41c); lamellar cusp with long, incurved spine (Fig. 42c); notogastral setae of l and h series in centrodorsal position (Fig. 12a)...................................  Neseutegaeus</p><p>- Humeral process more-or-less straight, with basal protrusion (Figs. 41a, b) or V-shaped plate (Fig. 41d); lamellar cusp pointed (Fig. 42b) or with two teeth (Figs. 42a, d).................................................................. 5</p><p>5 Lamellar cusp with single point (Fig. 42d); humeral processes narrow, thin, transparent, delicate (Fig. 41b).....  Atalotegaeus</p><p>- Lamellar cusp with two teeth (Fig. 42a); humeral processes not narrow or transparent (Fig. 41a)................  Eutegaeus</p><p>6 Lage mites (&gt;700 μm long); notogastral setae of l and h series thin, setiform (Fig. 18a); humeral processes with rugose ornamentation (Fig. 41d)....................................................................  Humerotegaeus</p><p>- Small mites (&lt;350 μm long); notogastral setae of l and h series stout, curved, spiniform (Fig. 20a); humeral processes smooth or with globular cerotegument (Fig. 41e).........................................................  Neoeutegaeus</p><p>7 Humeral processes with large, pointed anterior lobe and with 2-4 smaller lobes of lighter cuticle positioned posteriorly........................................................................................  Compactozetidae …8</p><p>- Humeral process lacking posterior lobes.................................................................. 10</p><p>8 Lamellae fused with each other apically, folded ventrally and fused with rostrum; with ovoid foramina lateral of fusion with rostrum (Figs. 42f).........................................................................  Compactozetes</p><p>- Lamellae with short, blunt, free cusps with or without a translamella (Figs. 42g, h)................................. 9</p><p>9 Apices of humeral processes triangular (Fig. 41h); notogastral setae emerging from alveoli; dorsosejugal scissure transverse; pedotectum I with long apical spine, pd II large lobe (Fig. 43h)........................................  Sadocepheus</p><p>- Apices of humeral processes bilobed (Fig. 41g); notogastral setae on tubercles; dorsosejugal scissure convex; pedotectum I lacking long apical spine; pd II short spur (Fig. 43g).................................................  Hamotegeus</p><p>10 Humeral process short, rounded apically, extending only as far as bothridium; rostrum lobed or incised (Figs. 411, j)................................................................................  Bornebuschiidae fam. nov …..11</p><p>- Humeral process long, pointed apically, extending well anterior of bothridium; rostrum rounded, entire (Figs. 411, j)..... 12</p><p>11 Lamellae very long with sharp apices, extending well beyond rostrum (Fig. 42j).........................  Dicrotegaeus</p><p>- Lamellae not extending beyond rostrum (Fig. 42i).................................................  Bornebuschia</p><p>12 Lamellae fused apically (Figs. 42k,l, m); notogastral setae short, thin, positioned marginally or sub-marginally on convex notogaster (Figs. 35a, 36a)................................................................  Pterozetidae …13</p><p>- Lamellae separate, with free cusps (Fig. 43m); notogastral setae long, with spinose ornamentation, positioned marginally and projecting vertically from concave notogaster (Figs. 37a)..................  Porrhotegaeidae fam. nov …..  Porrhotegaeus</p><p>13 Anterior of lamellae strongly concave (Fig. 42l); pedotectum I massive, with apical spine, lateral margin straight, longitudinal; pd II thin, pointed (Fig. 43l).....................................................................  Dudichella</p><p>- Anterior of lamellae convex or slightly concave (Fig. 42k, l); pedotectum I and pd II rectangular (Fig. 43k) or pd I with curved anterolateral projection and pd II narrow, pointed (Fig. 43l)............................................  Pterozetes</p></div>	https://treatment.plazi.org/id/03A2C77C462CFF82C79CB02512B8D908	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colloff, Matthew J.	Colloff, Matthew J. (2023): The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families. Zootaxa 5365 (1): 1-93, DOI: 10.11646/zootaxa.5365.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220
