taxonID	type	description	language	source
810E963FFFC0FFE7FF47FC18D633A93F.taxon	description	urn: lsid: zoobank. org: act: 3 C 8 C 7 D 6 D- 2834 - 4864 - AB 96 - 57 A 491 F 1 D 592 Figures 1 A – D, 2 A, 3	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC0FFE7FF47FC18D633A93F.taxon	type_taxon	Type species: Hathor spinosa Deschodt, new species.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC0FFE7FF47FC18D633A93F.taxon	diagnosis	Diagnosis. This new genus clearly belongs to the tribe Onthophagini by virtue of the following characters: scutellum not visible from above; body small and convex; short meso and metalegs second article of the labial palps longer than the first. It can be separated from all other Onthophagini genera by the following combination of characters: Antennae with eight segments; protibiae with four external teeth, fourth tooth reduced; distal margin of protibiae slanted and not obtuse. Hathor Deschodt, new genus shares some affinities with another known ant associated genus, Amietina Cambefort, 1981, as regards the eight antennal segments and oval body shape, but the ends of the front tibiae are truncated in Amietina while it also has only three teeth on the outside of the front tibiae. Hathor Deschodt, new genus also shares some superficial affinities with other onthophagine species form the subgenus Furconthophagus (Fig. 2 B) by virtue of the pronotal shape, the vertical prothoracic carina and having horns on the head. However, the different shaped horns and stockier legs and tarsi easily distinguish Hathor Deschodt, new genus from Furconthophagus.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC0FFE7FF47FC18D633A93F.taxon	description	Description. Female. Body: Black, shiny, oval, small, 4.8 mm long with head extended forward and 3.0 mm at the widest area over the elytra. Head: Antennae eight segmented; clypeus extended and narrowing anteriorly, punctate with associated setae; frontoclypeal suture distinct medially, effaced at the sides, curving anteriorly; vertex with distinct horns (Fig. 1 D). Prothorax: Punctate on horizontal surface with short associated setae; lateral vertical surface with small granules and short associated setae; anterior vertical surface smooth; triangular spine-like process present anteromedially. Ventrally the prothoracic carina is straight. Elytra: Surface of interstriae shagreened and with regular punctures; seventh interstriae clearly narrowed anteriorly to accommodate the markedly widened eighth interstriae. Protibiae: With four teeth on outside, fourth tooth reduced; ending at a slanted angle. Mesosternum: Punctate and very narrow medially. Metasternum: Surface smooth; punctate. Abdomen: surface smooth with short setae. Male. Unknown.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC0FFE7FF47FC18D633A93F.taxon	etymology	Etymology. Hathor was a principal deity in ancient Egyptian faith for music, dance, joy, love, sexuality, and maternal care. She was often portrayed as a woman wearing a headdress of cow horns. The clypeal armour of this new genus superficially resembles those horns. The generic name is feminine.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC0FFE7FF47FC18D633A93F.taxon	discussion	Remarks. The type specimen was collected walking amongst the ant species Anoplolepis custodiens with no dung present in the vicinity. Possession of eight antennal segments usually characterizes an ant-associated inquiline lifestyle (Davis et al. 2008) and as suggested by the field observations some association with those ants is anticipated.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC2FFE6FF47FF05D5EAA930.taxon	description	Figures 1 A – D, 2 A, 3	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC2FFE6FF47FF05D5EAA930.taxon	materials_examined	Type material. Holotype, female: Schurveberg | - 25.768 ° 27.964 ° | 07. i. 2023 | C. Deschodt (TMSA).	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC2FFE6FF47FF05D5EAA930.taxon	description	Description of the holotype (female): Head: (Fig. 1 A, C and D and 2 A) Clypeus extended and narrowing anteriorly, upturned at end, shallowly sinuate; anterior surface of frontoclypeal suture smooth and punctate with associated setae; frontoclypeal suture distinct medially, effaced at the sides, curving anteriorly; vertex with distinct and setate horns. Prothorax: Dorsal surface punctate with short associated setae; lateral surface with small granules and short associated setae; anterior vertical surface smooth, prothoracic carina straight.; triangular spine-like process present medially. Legs: Protibiae with four teeth on outside, fourth tooth reduced, small denticle present between second and third teeth and between third and fourth teeth; tibiae ending at a slanted angle; first article of the protarsus longer than the next three combined, last protarsus almost as long as the previous three segments combined; meso and metalegs widening distally. Venter. (Fig. 1 B) Mesometasternal suture distinct and very slightly sinuate; mesosternum narrow, punctate with small punctures that are closer together than their diameters; metasternum surface smooth with small punctures that are about three times their diameters apart. Abdomen: surface of ventrites smooth with short setae. Pygidium: Surface smooth with medium punctures about one puncture diameter apart; punctures with thin associated setae.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC2FFE6FF47FF05D5EAA930.taxon	etymology	Etymology. The specific epithet is feminine in gender and refers to the triangular spine-like process that is seen on the front of the prothorax.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
810E963FFFC2FFE6FF47FF05D5EAA930.taxon	discussion	Remarks. As it was found in putative association with the very widespread and commonly found ant species Anoplolepis custodiens, the potential distribution of Hathor spinosa Deschodt, new genus and species might be most of Southern Africa and further to the north east.	en	Deschodt, Christian M., Sole, Catherine L. (2023): A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa. Zootaxa 5375 (2): 279-284, DOI: 10.11646/zootaxa.5375.2.9, URL: https://www.mapress.com/zt/article/download/zootaxa.5375.2.9/52329
