taxonID	type	description	language	source
2D1AF3A9C35C3A893BB6B73042902779.taxon	materials_examined	Material examined. Neotype (CSR SASA 37013 / 37013 a) (Fig. 8 A−E). Conspecifics (BOLD-ID: BARCA 121 - 10, ACCESSION NR: HQ 171594) (Weigand et al. 2011 )), SMF 341636 (Fig. 2 B−F), NMBE 532009 (Fig. 8 F): Turjeva jama, Robic, Kobarid, Julian Alps, Slovenia, 46.2435 ° N, 13.5046 ° E, alt. ca. 253 m, air temp. in cave 9.3 ° C, June 1, 2013. Leg. Rajko Slapnik.	en	Jochum, Adrienne, Slapnik, Rajko, Klussmann-Kolb, Annette, Pall-Gergely, Barna, Kampschulte, Marian, Martels, Gunhild, Vrabec, Marko, Claudia Nesselhauf,, Weigand, Alexander M. (2015): Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeumisselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165, DOI: http://dx.doi.org/10.3897/subtbiol.16.5758, URL: http://dx.doi.org/10.3897/subtbiol.16.5758
2D1AF3A9C35C3A893BB6B73042902779.taxon	diagnosis	Diagnosis. Shell minute, transparent when fresh, conical with entire, half-roundish and more or less thickened peristome; parietal lamella; columella non-introrse, secondary columellar dilatation just above umbilical indentation; columellar lamella inclinate.	en	Jochum, Adrienne, Slapnik, Rajko, Klussmann-Kolb, Annette, Pall-Gergely, Barna, Kampschulte, Marian, Martels, Gunhild, Vrabec, Marko, Claudia Nesselhauf,, Weigand, Alexander M. (2015): Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeumisselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165, DOI: http://dx.doi.org/10.3897/subtbiol.16.5758, URL: http://dx.doi.org/10.3897/subtbiol.16.5758
2D1AF3A9C35C3A893BB6B73042902779.taxon	description	Description. Measurements of neotype specimen CSR SASA 37013 (Fig. 8 A−E): shell height 1.56 mm, shell diameter 1.07 mm, peristome width 0.75 mm, peristome height 0.84 mm. Measurements of conspecifics from Turjeva jama are provided in Table 3 (based on neotype and conspecifics of neotype). Shells minute (Table 5), variable in height (1.05 - 1.63 mm), conical with about 5 whorls, regularly coiled, suture deep, whorls convex, more or less shouldered; aperture semicircular, peristome closely adhering to spire (Fig. 2 D), thickened, higher than wide, taking up to ca. 35 % of shell height; umbilicus closed, shallow depression; contact between the columellar edge of peristome and last whorl moderately long (0.34 - 0.59 mm); non-expressed parietal lamella in aperture continues only in traces within shell; parietal plica present or absent (Fig. 8 F); penultimate whorl " sinks " into ultimate whorl at upper junction of peristome and penultimate whorl in profile perspective (Fig. 2 D); columella twisted but non-introrse, slender, secondary columellar dilatation at base; columellar lamella inclinate, extended (Fig. 8 A−D); secondary columellar dilatation equal to 1 / 2 distance of maximal upper columellar lamella extension; protoconch with pattern of spiral interconnected pits separated by zones of non-pitted bands (Fig. 9 D−E); teleoconch with tightly spaced irregular spiral striae of densely interconnected pits (Fig. 9 B, C, F); broader spiral bands interrupt finer rows of pitted striae (Fig. 9 B).	en	Jochum, Adrienne, Slapnik, Rajko, Klussmann-Kolb, Annette, Pall-Gergely, Barna, Kampschulte, Marian, Martels, Gunhild, Vrabec, Marko, Claudia Nesselhauf,, Weigand, Alexander M. (2015): Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeumisselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165, DOI: http://dx.doi.org/10.3897/subtbiol.16.5758, URL: http://dx.doi.org/10.3897/subtbiol.16.5758
2D1AF3A9C35C3A893BB6B73042902779.taxon	distribution	Distribution. Although we question past distribution records (Fig. 1, Table 1), Zospeum isselianum likely occurs within a narrow radius encompassing the Southeastern Alpine region of contiguous cave and river systems of northeastern Italy, southern Austria, Slovenia and northwest Croatia (Bole 1974, Maier and Bole 1975, Mildner 1976, Slapnik 1991, 1994, Slapnik and Ozimec 2004).	en	Jochum, Adrienne, Slapnik, Rajko, Klussmann-Kolb, Annette, Pall-Gergely, Barna, Kampschulte, Marian, Martels, Gunhild, Vrabec, Marko, Claudia Nesselhauf,, Weigand, Alexander M. (2015): Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeumisselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165, DOI: http://dx.doi.org/10.3897/subtbiol.16.5758, URL: http://dx.doi.org/10.3897/subtbiol.16.5758
2D1AF3A9C35C3A893BB6B73042902779.taxon	conservation	Conservation. In the caves sampled for this study, empty shells were sparsely found in sediment and live individuals sparsely populated certain cave walls during the summer season. These findings suggest that Zospeum isselianum occurred there for more than one season, and that these populations are / were not immediately threatened. Still, on a global scale, its distribution is fragmented and likely limited to far less than 312 caves within a radius of 20,000 km 2. In conjunction with the categories for the IUCN Red List (IUCN Standards and Petitions Subcommittee 2014), it can be considered a taxon of Least Concern (LC) since Zospeum isselianum is (potentially) known from many sites and these populations seem to be rather stable. Habitat disturbance through human agency poses the greatest threat.	en	Jochum, Adrienne, Slapnik, Rajko, Klussmann-Kolb, Annette, Pall-Gergely, Barna, Kampschulte, Marian, Martels, Gunhild, Vrabec, Marko, Claudia Nesselhauf,, Weigand, Alexander M. (2015): Groping through the black box of variability: An integrative taxonomic and nomenclatural re-evaluation of Zospeumisselianum Pollonera, 1887 and allied species using new imaging technology (Nano-CT, SEM), conchological, histological and molecular data (Ellobioidea, Carychiidae). Subterranean Biology 16: 123-165, DOI: http://dx.doi.org/10.3897/subtbiol.16.5758, URL: http://dx.doi.org/10.3897/subtbiol.16.5758
