taxonID	type	description	language	source
4701EEAA8A72A7234E62335499BA7090.taxon	description	Description. Female. Body (Fig. 2 A) subcylindrical, 610 - 635 µm (mean 620 µm; n = 3) long (measured from tip of rostrum to posterior margin of caudal rami) and 148 - 155 µm (mean 153 µm; n = 3) wide (at posterolateral margin of cephalothorax). Prosome composed of cephalothorax and 3 free pedigerous somites; tergite of first two pedigerous somites each with elliptical integumental window. Urosome comprised of fifth pedigerous somite, genital double-somite, and 3 free abdominal somites. Fifth pedigerous somite with short, dorsolateral row of spinules and numerous short rows of minute denticles (not drawn) and minute surface pits (not drawn) on dorsal and ventral surfaces. Components of genital double-somite (Fig. 2 A, B) partially fused dorsally but completely fused ventrally, ornamented with short, anterolateral row of spinules on ventral surface, minute surface pits and numerous short rows of minute denticles on dorsal and ventral surfaces (only minute denticles on ventral surface are shown), and row of large spinules and serrated hyaline frill encircling posterior margin; genital field with large median copulatory pore, chitinized copulatory duct leading anteriorly to pair of bilobate seminal receptacles, and median genital pore covered by operculiform leg 6. First free abdominal somite with large integumental window on ventral surface, minute surface pits and numerous short rows of minute denticles on dorsal and ventral surfaces (only minute denticles on ventral surface are shown), and row of large spinules and serrated hyaline frill ringing posterior margin. Second free abdominal somite similar to preceding somite, but without integumental window. Anal somite (Fig. 2 B, C) with minute surface pits (not drawn) and numerous short rows of minute denticles on dorsal and ventral surfaces and row of large spinules along posterior border and along posterior margin of anal operculum. Caudal ramus (Fig. 2 B, C) short, about 1.25 times longer than wide, bears minute surface pits (not drawn) and 7 setae. Insertion point of setae III, VI and VII flanked by spinules. Setae IV and V spinulate, with proximal breaking planes; other setae naked. Seta VII basally tri-articulate. Rostrum (Fig. 2 D) subtrianglar, not demarcated at base, with 2 dorsal sensilla. Antennule (Fig. 2 E) 8 - segmented, with armature as follows: 1, 9, 8, 4 + ae, 2, 3, 4, and 8. Segment 1 proximally with additional spinular row and large tubular pore. Two (of 4) and 4 (of 8) setae basally biarticulate on segments 7 and 8, respectively. Two (of 4) anterodistal setae on segment 8 fused at base. Antenna (Fig. 2 F), comprising coxa, basis, exopod, and 2 - segmented endopod. Coxa naked and unarmed. Basis with 2 small proximal spinules and 2 large inner distal spinules. Exopod 1 - segmented, cylindrical, and armed with 2 pinnate setae. Proximal endopodal segment naked and unarmed. Distal endopodal segment as long as basis and proximal endopodal segment combined; ornamented with 2 distolateral hyaline frills, 3 proximomedial spinules, and 2 distomedial spinules; armed with 2 spines (1 spine with minute spinules along inner margin; other with subapical flagellum) plus 2 naked setae along inner subdistal margin and 1 pilose and 5 geniculate setae along apical margin (lateralmost geniculate seta with 2 spinules at mid-point and fused basally with pilose seta; shortest geniculate seta with subapical flagellum). Labrum (Fig. 3 A) subtriangular, with denticles along apical margin and large distolateral denticles plus 2 patches of minute denticles on posterior face. Mandible (Fig. 3 B) composed of coxa and 2 - segmented palp. Coxa with inner subapical process, numerous unicuspidate teeth along distal margin, and unilaterally denticulate seta on inner distal angle. Proximal segment of palp unarmed, but furnished with 2 proximomedial spinules, 1 medial process, and row of apical spinules; distal segment armed with 5 apical naked setae. Maxillule (Fig. 3 C) composed of praecoxa and 3 - segmented palp. Praecoxal arthrite bears proximal crescentic row of spinules, 2 chitinized naked setae on inner subapical margin, 2 long naked setae on anterior surface, and 8 apical elements (3 highly chitinized, of which 2 each furnished with minute apical teeth; 1 unipinnate; 1 with bristled tip; 3 naked). Coxal endite elongated, with subapical row of spinules and 1 geniculate and 2 naked setae at distal end. Basis as long as coxa, bears 5 apical naked setae. Endopod 1 - segmented, vestigial, armed with 2 short naked setae. Exopod absent. Maxilla (Fig. 3 D) 3 - segmented, composed of syncoxa, allobasis, and 1 - segmented endopod. Syncoxa large, with 3 longitudinal rows of spinules on anterior surface and 1 pectinate and 2 naked apical setae on distal endite. Allobasis drawn out into long claw furnished with spinules along distal half of inner margin and bears 1 pectinate seta. Endopod 1 - segmented, armed with 2 long apical setae. Maxilliped (Fig. 3 E) 3 - segmented, comprising syncoxa, basis, and 1 - segmented endopod. Syncoxa stout, with 2 rows of long spinules and 1 distal pinnate seta. Basis naked and unarmed, about 1.5 times as long as syncoxa. Endopod drawn out into long claw, with 1 proximal naked seta and denticles along distal half of inner margin. Legs 1 - 4 biramous (Figs 3 F, G, 4 A, B); leg 1 with trimerous rami; legs 2 - 4 with trimerous exopod and bimerous endopod. Armature on rami of legs 1 to 4 as follows (Roman numerals = spines; Arabic numerals = setae): Leg 1 (Fig. 3 F) intercoxal sclerite naked and about twice as wide as long. Coxa with 2 rows of minute spinules and 1 row of large spinules on anterior surface; outer margin with 1 row of large spinules and 2 rows of fine spinules; inner distal corner with row of fine spinules. Basis with row of large spinules at insertion of each ramus and row of fine spinules along inner margin and on posterior surface; 1 additional large spinule present near base of inner spine; both spines with subapical flagellum. Outer spine on proximal exopodal segment with subapical flagellum. First two exopodal segments with large spinules along outer margin and on outer distal corner, as well as fine spinules along inner margin; distal segment with large spinules along outer margin and 1 spinule on apical margin. Both setae on terminal exopodal segment geniculate. Proximal endopodal segment long, extending almost to mid-point of distal exopodal segment, with fine spinules along inner margin; middle segment with fine spinules along outer and inner margins; distal segment with fine spinules along outer margin. Two (of 3) setae on distal endopodal segment geniculate. Both rami with minute surface pits (not drawn). Leg 2 (Fig. 3 G) intercoxal sclerite posteriorly bilobate, with row of spinules on each lobe. Coxa with 4 rows of minute spinules on anterior surface; outer margin with 2 rows of fine spinules; inner distal corner with row of minute spinules. Basis ornamented as in leg 1, except lacks large spinule near inner margin and with additional row of minute spinules on anterior surface. Exopod ornamented as in leg 1, except with additional spinulated frill on inner distal corner of proximal and middle segments. First two exopodal segments protruded on outer distal corner. Proximal endopodal segment with spinules along outer and inner margins and short spinulated frill on inner distal corner. Distal endopodal segment about 1.5 times longer than proximal endopodal segment and furnished with spinules along outer and inner margins. Both rami with minute surface pits (not drawn) as in leg 1. Leg 3 (Fig. 4 A) similar to leg 2, except with naked intercoxal sclerite, outer seta (instead of spine) on basis, longer inner spine on middle exopodal segment, and inner distal seta (instead of spine) and longer inner proximal spine on distal endopodal segment. Leg 4 (Fig. 4 B) similar to leg 3, except with smaller intercoxal sclerite, row of spinules absent on posterior surface of basis, less protruded outer distal corner on first two exopodal segments, shorter inner spine on middle exopodal segment, 2 more elements on distal exopodal segment, shorter distal endopodal segment, and only 3 elements on distal endopodal segment. Leg 5 (Fig. 4 C) biramous. Basoendopod with long outer basal seta and 3 spinulated setae on endopodal lobe. Exopod 1 - segmented, slightly longer than wide, with few spinules along inner margin and 4 unequal naked setae. Leg 6 (Fig. 2 B) represented by simple operculum covering genital pore, armed with 1 minute naked seta on outer distal corners. Male. Unknown.	en	Tang, Danny, Eberhard, Stefan M. (2016): Two new species of Nitocrella (Crustacea, Copepoda, Harpacticoida) from groundwaters of northwestern Australia expand the geographic range of the genus in a global hotspot of subterranean biodiversity. Subterranean Biology 20: 51-76, DOI: http://dx.doi.org/10.3897/subtbiol.20.10389, URL: http://dx.doi.org/10.3897/subtbiol.20.10389
4701EEAA8A72A7234E62335499BA7090.taxon	etymology	Etymology. This species is named in honour of the late Professor Brenton Knott (The University of Western Australia) who made significant contributions to research on groundwater fauna in Western Australia.	en	Tang, Danny, Eberhard, Stefan M. (2016): Two new species of Nitocrella (Crustacea, Copepoda, Harpacticoida) from groundwaters of northwestern Australia expand the geographic range of the genus in a global hotspot of subterranean biodiversity. Subterranean Biology 20: 51-76, DOI: http://dx.doi.org/10.3897/subtbiol.20.10389, URL: http://dx.doi.org/10.3897/subtbiol.20.10389
5ADCDF2CF5D22E8A148C28D8179F1B15.taxon	materials_examined	Other material examined. All material collected from boreholes in the Ethel Gorge aquifer, approximately 15 km ENE of Newman, Western Australia (Fig. 1). 3 females and 1 male (WAM C 51833) in absolute alcohol, borehole EEX 917 (23 ° 19 ' 47 " S; 119 ° 52 ' 13 " E), 8 February 2012, S. Catomore and N. Coen leg.; 7 females (WAM C 51834) in absolute alcohol, borehole EEX 917 (23 ° 19 ' 47 " S; 119 ° 52 ' 13 " E), 12 April 2012, P. Bell and S. Catomore leg.; 1 male (WAM C 51835) in absolute alcohol, borehole P 13 S (23 ° 18 ' 56 " S; 119 ° 50 ' 58 " E), 21 November 2009, P. Bell and G. Perina leg.; 1 female (WAM C 51836) in absolute alcohol, borehole T 399 (23 ° 17 ' 03 " S; 119 ° 52 ' 07 " E), 5 November 2010, P. Bell and S. Catomore leg.; 13 females and 3 males (WAM C 51846) in absolute alcohol, borehole UNK 02 (23 ° 15 ' 00 " S; 119 ° 53 ' 41 " E), 9 February 2011, P. Bell and S. Catomore leg.; 2 females (WAM C 51847) in absolute alcohol, borehole W 105 (23 ° 19 ' 37 " S; 119 ° 51 ' 50 " E), 9 February 2011, P. Bell and S. Catomore leg.; 7 females (WAM C 51848) in absolute alcohol, borehole W 105 (23 ° 19 ' 37 " S; 119 ° 51 ' 50 " E), 13 April 2012, P. Bell and S. Catomore leg.; 3 females (WAM C 51849) in absolute alcohol, borehole W 105 (23 ° 19 ' 37 " S; 119 ° 51 ' 50 " E), 10 February 2012, S. Catomore and N. Coen leg.; 2 females (WAM C 51850) in absolute alcohol, borehole W 116 (23 ° 14 ' 47 " S; 119 ° 54 ' 25 " E), 8 February 2011, P. Bell and S. Catomore leg.; 3 females (WAM C 51851) in absolute alcohol, borehole W 116 (23 ° 14 ' 47 " S; 119 ° 54 ' 25 " E), 22 November 2009, P. Bell and G. Perina leg.; 5 females (WAM C 51852) in absolute alcohol, borehole W 116 (23 ° 14 ' 47 " S; 119 ° 54 ' 25 " E), 21 April 2010, P. Bell and G. Perina leg.; 8 females and 1 male (WAM C 51853) in absolute alcohol, borehole W 116 (23 ° 14 ' 47 " S; 119 ° 54 ' 25 " E), 8 February 2012, S. Catomore and N. Coen leg.; 6 females and 3 males (WAM C 51854) in absolute alcohol, borehole W 116 (23 ° 14 ' 47 " S; 119 ° 54 ' 25 " E), 11 April 2012, P. Bell and S. Catomore leg.; 1 female (WAM C 51855) in absolute alcohol, borehole W 79 D (23 ° 19 ' 42 " S; 119 ° 50 ' 39 " E), 22 November 2009, P. Bell and G. Perina leg.; 3 females (WAM C 51856) in absolute alcohol, borehole W 79 D (23 ° 19 ' 42 " S; 119 ° 50 ' 39 " E), 12 April 2012, P. Bell and S. Catomore leg.; 2 males (WAM C 51857) in absolute alcohol, borehole WP 126 NRE (23 ° 15 ' 01 " S; 119 ° 53 ' 42 " E), 21 November 2009, P. Bell and G. Perina leg.	en	Tang, Danny, Eberhard, Stefan M. (2016): Two new species of Nitocrella (Crustacea, Copepoda, Harpacticoida) from groundwaters of northwestern Australia expand the geographic range of the genus in a global hotspot of subterranean biodiversity. Subterranean Biology 20: 51-76, DOI: http://dx.doi.org/10.3897/subtbiol.20.10389, URL: http://dx.doi.org/10.3897/subtbiol.20.10389
5ADCDF2CF5D22E8A148C28D8179F1B15.taxon	description	Description. Female. Body (Fig. 5 A) cylindrical, 450 - 495 µm (mean 471 µm; n = 6) long (measured from tip of rostrum to posterior margin of caudal rami) and 95 - 105 µm (mean 103 µm; n = 6) wide (at first free pedigerous somite). Prosome composed of cephalothorax and 3 free pedigerous somites. Urosome comprised of fifth pedigerous somite, genital double-somite, and 3 free abdominal somites. Components of genital double-somite (Fig. 5 A, B, C) not fused dorsally but completely fused ventrally, with elliptical integumental window laterally, row of small spinules immediately posterior to each integumental window, and row of large spinules and frill of minute spinules encircling posterior margin; genital field with large median copulatory pore, chitinized copulatory duct leading anteriorly to pair of lobate seminal receptacles, and median genital pore covered by operculiform leg 6. First free abdominal somite with anteroventral pair of oval integumental windows and row of unequal spinules and frill of minute spinules ringing posterior border. Second free abdominal somite with row of subequal spinules and frill of minute spinules encircling posterior edge. Anal somite (Fig. 5 B, D, E) with anterior and posterior row of spinules on ventral surface, several rows of spinules on lateral surface, and spinules along posterior margin of anal operculum. Caudal ramus (Fig. 5 B, D, E) about 1.5 times longer than wide, bearing 7 setae. Spinules present at insertion point of setae III, VI and VII. Setae IV and V spinulate, with proximal breaking planes; other setae naked. Seta VII basally tri-articulate. Rostrum (Fig. 5 F) about 1.25 times longer than wide, not demarcated at base, with rounded apex and 2 dorsal sensilla. Antennule (Fig. 6 A) similar to that of Nitocrella knotti sp. n. Antenna (Fig. 6 B), comprising coxa, basis, 1 - segmented exopod, and 2 - segmented endopod. Coxa naked and unarmed. Basis with 2 long spinules on inner margin and short row of minute spinules on anterior surface. Exopod armed with 3 pinnate setae (2 highly chitinized). Proximal endopodal segment naked and unarmed. Distal endopodal segment slightly longer than proximal endopodal segment; ornamented with row of spinules on proximal half of inner margin, 2 distolateral hyaline frills, and 2 distomedial spinules; armed distomedially with 2 naked unequal spines plus 1 long naked seta and apically with 1 pilose and 5 geniculate setae (lateralmost geniculate seta with 2 spinules at mid-point and fused basally with pilose seta). Labrum (Fig. 6 C) as in Nitocrella knotti sp. n., except shorter and with smaller denticles along distal margin. Mandible (Fig. 6 D) similar to that of Nitocrella knotti sp. n. Maxillule (Fig. 6 E) similar to that of Nitocrella knotti sp. n. Maxilla (Fig. 6 F) as in Nitocrella knotti sp. n., except with naked syncoxa. Maxilliped (Fig. 6 G) similar to that of Nitocrella knotti sp. n., except with only 1 row of spinules on syncoxa and no proximal seta on endopod. Legs 1 - 4 biramous (Fig. 7 A, B, C, D); leg 1 with trimerous rami; legs 2 - 4 with trimerous exopod and bimerous endopod. Armature on rami of legs 1 to 4 as follows (Roman numerals = spines; Arabic numerals = setae): Leg 1 (Fig. 7 A) intercoxal sclerite naked and concave on posterior margin. Coxa with 1 row of spinules on anterior surface and another row of spinules on posterolateral surface. Basis with row of long spinules at insertion of each ramus and 3 additional large spinules proximal to inner spine; both spines with subapical flagellum. Exopodal segments with large spinules along outer margin and on outer distal corner; middle segment also with fine spinules along inner margin. Endopodal segments with large spinules on outer margin and fine spinules along inner margin. Both setae on terminal exopodal segment and 1 of 3 setae on distal endopodal segment geniculate. Leg 2 (Fig. 7 B) intercoxal sclerite naked and posteriorly bilobate. Coxa with 1 row of minute spinules on posterolateral surface. Basis with row of spinules at insertion of each ramus and several fine spinules (only 1 depicted) on inner margin; outer spine with subapical flagellum. Exopod ornamented as in leg 1, except with additional spinulated frill on inner distal corner of proximal and middle segments. First two exopodal segments protruded on outer distal corner. Both endopodal segments with spinules along outer margin. Leg 3 (Fig. 7 C) similar to leg 2, except with outer seta (instead of spine) on basis and 3 elements on distal endopodal segment. Leg 4 (Fig. 7 D) similar to leg 3, except with much smaller spinules at insertion of endopod, 6 elements on distal exopodal segment (of which inner distal seta is longer and ornamented with tightly packed spinules on inner margin of apex), and 2 elements on distal endopodal segment. Leg 5 (Fig. 7 E) biramous. Basoendopod with long outer basal seta plus median pore, short row of spinules laterally and 4 distal spinulated setae on endopodal lobe. Exopod 1 - segmented, about twice as long as wide, with spinules along inner margin and 4 setae (3 naked; 1 spinulated). Leg 6 (Fig. 5 B) represented by genital operculum covering genital pore, and armed with 1 minute naked seta on distolateral borders. Male. Body length (measured from tip of rostrum to posterior margin of caudal rami) 400 - 440 µm (mean 417 µm; n = 7); body width 90 - 95 µm (mean 91 µm; n = 7). Prosome composed of cephalothorax and 3 free pedigerous somites. Urosome comprised of fifth pedigerous somite, genital somite and 4 free postgenital somites. Genital somite (Fig. 8 A) wider than long, with elliptical integumental window laterally and row of small spinules immediately posterior to each integumental window. First postgenital somite (Fig. 8 A) with ventral pair of oval integumental windows, posterior row of large spinules, and frill of minute spinules along posterior border. Second and third postgenital somites (Fig. 8 A) posteriorly with row of spinules and frill of minute spinules. Anal somite (Fig. 8 A) ornamented as in female. Caudal ramus (Fig. 8 A) about 2 times as long as wide; armed and ornamented as in female. Antennule (Fig. 8 B) 10 - segmented, haplocerate, with geniculation between segments 7 and 8. Armature as follows: 1, 10, 8, 2, 7 + ae, 2, 3, 4, 4, and 8. Short spinulate seta (e) with flagellate tip present on segments 5 - 7. Aesthetasc and adjacent apical seta on segment 5 basally fused forming acrothek. One (of 3) and 3 (of 4) elements on segments 7 and 8, respectively, modified as digitate spines. Two apical setae on segment 10 basally fused. Inner spine on basis of leg 1 (Fig. 8 C) modified as is typical for members of Ameiridae. Leg 5 (Fig. 8 D) biramous, with basoendopods fused medially. Basoendopod with outer basal seta and median pore and 2 apical spinulated elements on endopodal lobe. Exopod 1 - segmented, about 1.3 times as long as wide, with 5 setae (3 naked; 2 spinulate). Leg 6 (Fig. 8 A) asymmetrical, with right side modified as operculum and left side basally fused to somite; each side armed with 2 unequal distolateral setae.	en	Tang, Danny, Eberhard, Stefan M. (2016): Two new species of Nitocrella (Crustacea, Copepoda, Harpacticoida) from groundwaters of northwestern Australia expand the geographic range of the genus in a global hotspot of subterranean biodiversity. Subterranean Biology 20: 51-76, DOI: http://dx.doi.org/10.3897/subtbiol.20.10389, URL: http://dx.doi.org/10.3897/subtbiol.20.10389
5ADCDF2CF5D22E8A148C28D8179F1B15.taxon	etymology	Etymology. This species is named for Dr. Tomislav Karanovic, in recognition of his extensive taxonomic research on subterranean copepods of Australia.	en	Tang, Danny, Eberhard, Stefan M. (2016): Two new species of Nitocrella (Crustacea, Copepoda, Harpacticoida) from groundwaters of northwestern Australia expand the geographic range of the genus in a global hotspot of subterranean biodiversity. Subterranean Biology 20: 51-76, DOI: http://dx.doi.org/10.3897/subtbiol.20.10389, URL: http://dx.doi.org/10.3897/subtbiol.20.10389
