taxonID	type	description	language	source
517F87A6FFC789321E55F90380985970.taxon	discussion	The genus comprises six species, described from the Chinese Province of Yunnan (NOMURA 2002, 2004). It is distinguished by pronotal characters, notably by presence of a mesal impression and carinate lateral margins. Two species have been collected in Nepal, one described below, with a large number of specimens, and thesecond with a single female. This species is probably also new. We decline to describe it in the absence of at least one available male.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	materials_examined	Holotype ♂: NEPAL Patan Distr. Phulchoki 2400 - 2600 m 28 - 30. IV. 1984 I. LÖbl A. Smetana (MHNG). Paratypes: 44 ex., NEPAL, Bagmati Zone, Patan District, Phulchoki, 2500 - 2600 m, 28. and 29. IV. 1984, leg. I. LÖbl (MHNG); 35 ex., Bagmati Zone, Patan District, Phulchoki, 2500 m, 10. V. 1981, leg. I. LÖbl & A. Smetana (MHNG, NMNS); 16 ex., Bagmati Zone, Patan District, Phulchoki, 2550 and 2600 m, 20. and 21. IV. 1982, leg A. & Z. Smetana (PCAM, MHNG); 54 ex., Bagmati Zone, Patan District, Phulchoki, 2300 - 2700 m, 13. - 17. X. 1983, leg. I. LÖbl & A. Smetana (MHNG); 1 ex., Bagmati Zone, Patan District, Phulchoki, leg. H. Franz (NHMW); 2 ex., Bagmati Zone, Patan District, Godawari, 1700 m, 19. X. 1983, leg. I. LÖbl (MHNG); 10 ex., Bagmati Zone, Sindhupalchok District, Pokhare NE Barahbise, 3000 m, 7. IV. 1981, leg. I. LÖbl & A. Smetana (MHNG); 16 ex., Bagmati Zone, Sindhupalchok District, Malemchi, 2900 m, 14. IV. 1981, leg. I. LÖbl & A. Smetana (MHNG, PCVB); 4 ex., Bagmati Zone, Sindhupalchok District, Tarke Ghyang, 2650 m, 19. IV. 1981, leg. I. LÖbl & A. Smetana (MHNG); 1 ex., Bagmati Zone, Sindhupalchok District, above Shermathang, 2900 m, 26. IV. 1981, leg. I. LÖbl & A. Smetana (MHNG); 1 ex., Bagmati Zone, Sindhupalchok District, Dobate Ridge NE Barahbise, 2800 m, 2. V. 1981, leg. I. LÖbl & A. Smetana (MHNG); 3 ex., Bagmati Zone, Helambu, Gul Bhanjyang, 2600 m, 6. IV. 1981, leg. I. LÖbl & A. Smetana (MHNG); 1 ♀, Bagmati Zone, Rasuwa District, Ramche, 1800 - 3350 m, 18. IV. 1978, leg. B. C. Bhakta (NHMB); 1 ex., Central Nepal, forest below “ Fulung ”, IX. - X. 1971, leg. H. Franz (NHMW); 1 ex., Gandaki Zone, between Mulkharka and Tare Pati, IX. - X. 1971, leg. H. Franz (NHMW); 1 ex., Gandaki Zone, ridge E Goropani, 3100 m, 7. X. 1983, leg. I. LÖbl & A. Smetana (MHNG); 1 ex., Koshi Zone, Dhankuta District, Maharabhat, 2350 m, 20. X. 1977, leg. L. Deharveng [Nep. 174] (MHNG); 1 ex., Koshi Zone, Namche Bazar, ridge between Lamjura and Sete, 3000 m, 21. X. 1984, L. Deharveng (MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	etymology	Etymology. The species epithet refers to Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	description	Description. Length 1.55 - 1.62 mm. Head, body and appendages light brown, venter of thorax often darkened. Pubescence sparse, except for patches of white squamous setae covering venter of head posterior gular mound, most of proventrite, mesoventrite anterior mesocoxae, apices of elytra, apical margin of metaventrite and base of ventrite II. Head elongate, narrow, about 1.5 long as wide, with frontal rostrum prominent, clypeus and mandibles not visible in dorsal view, antennal tubercles large, each with ridge extending to nearly asetose vertexal foveae; with narrow, parallel-sided mesal sulcus. Mesal sulcus with setose patch between antennal tubercles. Margins of frontal sulcus angulate, lateral borders vertical. Eyes across head mid-length, prominent, with five to eight facets. Vertex raised, shallowly impressed mesally, inflexed almost vertically to level of upper eye margins. Vertexal foveae slightly anterior level of hind eyes margins. Temple each with carina extending from lower eye margins to neck constriction. Contours of temples rounded in dorsal view. Lateral parts of frontal rostrum, genae and antennomeres Iand III reticulate, remaining dorsal and lateral surface of head smooth. Pubescence short, rather appressed. Gular-mandibular carinae defining laterodorsally gular mound. Antennae 11 - segmented, with 3 - segmented club. Length / width ratio of antennomeres as: I 20 / 8: II 10 / 7: III 8 / 6: IV 7 / 6: V 7 / 6: VI 8 / 6: VII 8 / 6: VIII 8 / 6: IX 12 / 8: X 12 / 9: XI 24 / 13. Maxillary palpi elongate, segment I narrow, cylindrical, slightly bent near base, segment II pedunculate, about 3 times as long as segment I, longer thanhalf of head length, its peduncle about twice as long as thickened apical part, slightly swollen at base, segment III as long as wide, segment IV longer than head, with enlarged apical part lacking tubercles, shorter than peduncle (ratio 15 / 22), sharply delimited sensory area in apical triangular notch extending by carinae. Gular mound delimited anterolaterally by rounded impressions, posteriorly by flattened longitudinal impression. Gular foveae obscured by setose patch. Pronotum shorter than head, longer than wide, widest anterior mid-point, with lateral contours nearly evenly rounded, lateral margins appearing in dorsal view angulate. Pronotal disc raised dorsally above level of vertex, with mesal impression delimited by longitudinal carinae, antebasal sulcus interrupted in middle by two short carinae. Antebasal sulcus reaching distinct antebasal lateral foveae, mesal antebasal fovea absent. Sides nearly vertical below carinae upto level of lateral antebasal foveae, explanate at level of lateral antebasal foveae, explanate surface delimited by stria. Pronotal dorsum smooth, not microsculptured, punctation as on head hardly visible (x 160 x magnification), pubescence sparse, short, rather appressed. Lateral procoxal foveae obscured by patch of squamous setae. Elytra somewhat longer than head, their combined width about 1.5 times length, basal ridge raised. Each elytron with three basal foveae. Inner two basal foveae in common deep impression; outer fovea in similar but somewhat smaller impression, separated by median longitudinal carina reaching elytral apex; marginal carinae conspicuous, oblique along basal fourth of elytral lateral length, curved to form inconspicuous outer discal carina. Antebasal fovea present. Adsutural stria close to suture, adsutural carina raised. Outer apical angles prominent. Apical area of disc with striagulate microsculpture visible on cleared specimens. Pubescence short and appressed on discal carinae, absent from intervals, present and longer near apical squamous rim. Few short appressed setae present along lateral margins. Hind wings atrophied. Proventrite with setose patch reaching nearly lateral margins, obscuring foveae. Mesoventrite with patch of squamous setae overlapping mesal ridge. Metaventrite lacking foveae, with deep, glabrous mesal impression delimited by ridges bearing short setae. Apical margins of tergites broadly rounded. Tergite I broadened apically, about 1.5 times as wide at long. Ventrite II with two inconspicuous admesal setal rows. Femora reticulate, clavate, tibiae straight, gradually thickened apically. Tibiae straight, graduallythickened apically, weakly curved. Tarsomeres III cylindrical, longer and narrower than tarsomeres II. Male characters. Obvious secondary sexual characters absent. Aedeagus (Figs 7, 8) 0.26 - 0.28 mm long, with narrow, flat, and bent ventral process, middle of apical side prominent. Parameres bearing apical and subapical setae. Internal sac with sclerotized rod acute at apex. Measurements of the holotype (in mm). Total length: 1.58, length / width: head 0.32 / 0.21, pronotum 0.30 / 0.26, elytra 0.34 / 0.53 (0.17), tergite I 0.40 / 0.62; length of maxillary palpomere IV 0.39, length of aedeagus 0.26.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	diagnosis	Diagnosis. Frontal sulcus parallel-sided, approaching vertexal foveae, with setal patch between antennal tubercles. Vertex sulcate mesally. Pronotum strongly raised dorsally, with deep mesal impression delimited by carinae, sides nearly vertical below carinae. Lateral margins of elytra carinate anteriorly. Aedeagus with setose parameres, internal sac lacking denticulate or spinose structures, with single rod acute at tip.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	distribution	Distribution. Western, Central and Eastern Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC789311D32FF6384F455D0.taxon	discussion	Comments. The new species differs markedlyfrom the Chinese members of Nabepselaphus by its deep pronotal impression delimited by admesal carinae and by the carinate basolateral margins of the elytra. The aedeagus of the new species is also distinctive in possessing comparativelyshort setose parameres. The internal sac has a single long, undivided sclerite, similar to that of N. jizushanus Nomura, 2004. With respectto the pronotal, elytral and aedeagal characters, the species is tentatively placed in Nabepselaphus. It occurs mainly in mountain forests with oak, rhododendron, and bamboo species but was also found in subtropical broadleaved Schima and Castanopsis forest. The available data give an altitudinal range from 1700 to 3100 m abovesea level. The distribution, ranging from Western to Eastern Nepal, is quite large for an apterous species. Most specimenscome fromthe Mahabharat Range south of the Kathmandu Valley while only two specimens were collected in East Nepal and two in Western Nepal. The absence of this species from extensive collections made in the Indian Darjeeling District, including on Tonglu at the Indian / Nepalese border, is also notable.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC489301D18F983855151B7.taxon	type_taxon	Type species Himallaphus pahilo n. sp. Gender masculine.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC489301D18F983855151B7.taxon	etymology	Etymology. The name is derived from Himalaya, combined with “ laphus ”, a meaningless part of the generic name Pselaphus.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC489301D18F983855151B7.taxon	description	Description. Length 1.60 - 1.95 mm. Pubescence sparse, except for patches of white squamous setae covering anterior parts of frontal sulcus, venter of head posterior gular mound, most of proventrite, mesoventrite anterior mesocoxae, apices of elytra, apical margin of metaventrite and base of ventrite II. Head narrow, elongate, with frontal rostrum prominent and with setose patch between prominent antennal tubercles. Frontal ridges extending to near anterior eyes margins, impressed posteriad and from level of posterior eye margins replaced by sulci joined on posterior inclined part of vertex. Mesal area of frons raised to form low ridge. Eyes small, situated at about mid-length of head. Vertex with mesal sulcus. Vertexal foveae obscured, situated behind level of posterior eye margin. Temple with impressed stria extending from upper eye margins to neck constriction. Contours of temples rounded or oblique in dorsal view. Gular-mandibular carinae conspicuous, forming lateroventral head margin. Antennae 11 - segmented, antennomeres Iand II reticulate, club 3 - segmented, weekly formed, segment XI asymmetrical. Maxillary palpi elongate, segment Inarrow, cylindrical, bent near base, segment II long, pedunculate, about 3 times as long as segment I, exceeding half of head length, with peduncle much longer than enlarged apical part, slightly swollen at base, segment III short, about long as wide, segment IV glabrous, pedunculate, bent, sensory area in apical triangular notch with sharply delimited margins extending bycarinae on lateral side of palpomere. Gula strongly swollen, gular mound longitudinal, narrowed posteriad, delimited laterally and posteriad by impression completely covered by setose patch. Gular foveae obscured. Pronotum rounded laterally, convex dorsally, widest about at mid-point, anterior antebasal sulcus with dense row of longitudinal striae, with antebasal lateral foveae and antebasal median fovea joined with antebasal sulcus, narrow basal area uneven. Elytra short, strongly widened apically, impressed basally, with rounded or oblique lateral margins, three basal and one antebasal foveae, discal carinae close to adsutural carinae. Hind wings atrophied. Proventrite withsetose patch reaching nearly lateral margins, obscuring foveae. Foveae of mesoventrite obscured by setose patch. Metaventrite afoveate. Legs without spines. Femora clavate, reticulate. Tibiae straight, thickened apically. Tarsomeres III cylindrical, longer than tarsomeres II. Abdomen with tergite Imuch longer than rest of tergites combined, convex, widened apically, with rounded lateral margins, deep basal sulcus and large foveae near lateral margin obscured by rows of flat setae; apical margins of tergites broadly rounded. Ventrite II with deep basal sulcus obscured by rows of flat setae. Aedeagus symmetrical, median lobe sub-oval, with strongly sclerotized ventral process, parameres weakly sclerotized, bearing apical setae.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC489301D18F983855151B7.taxon	diagnosis	Diagnosis. Head with frontal sulcus approaching vertexal foveae. Vertex transversely carinate, with mesal sulcus. Maxillary palpomere IV pedunculate, bent, about long as head, lacking tubercles, with sensory area in apical notch extended by carinae. Pronotum rounded, with deep longitudinal striae separated by carinae, except on area between antebasal sulcus and basal margin, antebasal sulcus not interrupted in middle, antebasal mesal and lateral foveae present. Elytra short, strongly widened apically, with three basal foveae, sutural and discal carinae.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC489301D18F983855151B7.taxon	discussion	Comments. Members of Himallaphus resemble Nabepselaphus by their body-shape, the pattern of the frontal carinae and sulci, the small eyes situated about at head mid-length, the long maxillary palpi with margins of the sensory organs V-shaped, the presence of antebasal pronotal sulcus joined with foveae, the short and apically strongly widened elytra, and the pattern of patches or stripes of widened setae. Himallaphus differs markedly from Nabepselaphus, and from all other Pselaphini, by its densely striate pronotum.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC589371D32FEC380FB5310.taxon	materials_examined	Holotype ♂: NEPAL Patan Distr. Phulchoki 2500 m 10. V. 1981 LÖbl (MHNG). Paratypes: 1 ♂, 1 ♀, NEPAL, Bagmati Zone, with the same locality data as the holotype (MHNG); 2 ♂, 3 ♀, same data but 2600 - 2700 m (MHNG); 2 ♀, same data but 2650 m, 14. X. 1993 (MHNG); 1 ♂, same data but 2500 m, 28 - 29. IV. 1984 (MHNG); 2 ♂, same data but 2550 m, 15. X. 1983, Smetana & LÖbl (MHNG); 1 ♂, 1 ♀, Bagmati Zone, Sindhupalchok Distr ,, Dobate ridge NE Barahbise, 2800 m, 2. V. 1981, I. LÖbl & A. Smetana (MHNG); 1 ♂, 3 ♀, Bagmati Zone, Sindhupalchok District, Pokhare NE Barahbise, 3000 m, 2. V. 1981, I. LÖbl & A. Smetana (MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC589371D32FEC380FB5310.taxon	etymology	Etymology. The species epithet means first in Nepali.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC589371D32FEC380FB5310.taxon	description	Description. Length 1.70 - 1.80 mm. Head about 1.5 times as long as wide. Frontal ridges extended to near level of posterior eye margins, narrowed posteriad. Frontal sulcus parallel-sided, raised from level of anterior eye margins to form gradually widened longitudinal mesal ridge. Mesal ridge delimited laterally by striae, apically by short carina, followed by deep mesal sulcus extended to neck. Setose patch in frontal sulcus about 0.06 mm long. Eyes in level of head mid-length, prominent, with eight to ten facets. Temples explanate below level of eyes, with rounded contours in dorsal view and transverse striae extended onto vertex. Neck and area near neck lacking striae. Glabrous genal area extended behind eyes and gradually narrowed. Temples and vertex anterior neck rounded, with conspicuous appressed pubescence. Segment IV of maxillary palpus shorter than head, its apical enlarged area about as long as peduncle. Length / width ratio of antennomeres as: I 28 / 9: II 10 / 8: III 9 / 5.5: IV 9 / 5.5: V 9 / 5.5: VI 9 / 5.5: VII 9 / 5.5: VIII 9 / 5.5: IX 11 / 8: X 11 / 9: XI 27 / 13. Gular mound subtriangular. Pronotum slightly longer than wide, strongly convex, shorter than head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal carinae dense, reaching almost anterior pronotal margin, striae between carinae narrower than carinae. Pronotal pubescence moderately short,. Elytron gradually impressed toward base, with sutural and discal carinae parallel, discal carina slightly wider than sutural carina; basal third of lateral margin carinate and oblique, pubescence short, similar to pronotal pubescence. Male characters. Metaventrite with mesal impression delimited by large admesal ridges overlapping metacoxae, their inner sides sharply delimited, diverging apically. Surface of ridges oval, flat, completely covered by patches of short flat setae. Middle of ventrite II throughout shallowly impressed, impression about as long as wide. Aedeagus (Figs 9, 10) 0.34 - 0.35 mm long. Ventral process short, bent, narrow, at apex about as sixth of maximal aedeagal width. Parameres widened apically (lateral view). Internal sac broad, lacking distinct sclerotized pieces, at base variably sclerotized, densely spinose. Measurements of the holotype (in mm). Total length 1.75; length / width of head 0.40 / 0.26, pronotum 0.37 / 0.32, elytra 0.50 / 0.66 (0.19), tergite I 0.54 / 0.78; length of maxillary palpomere IV 0.34, length of aedeagus 0.34.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC589371D32FEC380FB5310.taxon	diagnosis	Diagnosis. Vertex and temples transversallystriate. Mesal vertexal sulcus extending to neck constriction. Eyes prominent. Maxillary palpomere IV with enlarged apical part about as long as peduncle. Base of elytra impressed. Protuberances of male metaventrite overlapping metatrochanters. Apicoventral process of aedeagus narrow. Parameres wide in dorsal view. Internal sac broad, densely spinose.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC589371D32FEC380FB5310.taxon	distribution	Distribution. Central Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289371EB3FD6385675570.taxon	materials_examined	Holotype ♂: NEPAL: Kosi, ridge S Mangming 2800 m 7. IV. 84 LÖbl-Smetana (MHNG). Paratypes: 9 ♀, NEPAL, Koshi Zone, Sankhuawasabha District, with the same data as the holotype (MHNG); 1 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, Induwa Kola valley. 2800 m, 15. IV. 1984, I. LÖbl & A. Smetana (MHNG); 1 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, “ Bakan ” Wof Tashigaon, 3200 m, 5. IV. 1982, A. & Z. Smetana (MHNG, PCPH); 1 ♂, 1 ♀, Koshi Zone, Sankhuawasabha District, above Tashigaon, 3600 m, 6. IV. 1982, A. & Z. Smetana (MHNG); 2 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, above Sheduwa, 3000 m, 31. III- 1. IV. 1982, A. & Z. Smetana (MHNG); 3 ♂, 3 ♀, Meshi Zone, Panchtar District, above Pahakhola, 2600 - 2800 m, 31. V. 1988, J. Martens & W. Schawaller (SMNS, MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289371EB3FD6385675570.taxon	etymology	Etymology. The species epithet is a Nepali word meaning next.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289371EB3FD6385675570.taxon	description	Description. Length 1.60 - 1.75 mm. Headabout 1.4 times as long as wide. Frontal ridges extended to level of eye midlength, narrowed posteriad. Frontal sulcus widened posterior antennal tubercles, parallel-sided, raised at level of eye midlength to form shallow mesal vertexal sulcus reaching level of highest point of vertex. Setose patch in frontal sulcus about 0.07 to 0.09 mm long. Eyes at head mid-length, not prominent, with eight or nine facets. Contours of temples rounded in dorsal view. Lower part of temples explanate, delimited by longitudinal carina, glabrous, very finely pubescent. Upper anterior part of temples with short striae extended onto vertex. Neck and area near neck without striae. Glabrous genal area extended behind eyes and gradually narrowed. Vertex anterior neck convex, with inconspicuous appressed pubescence. Segment IV of maxillary palpus shorter than head, with peduncle much longer than enlarged apical area (ratio 29 / 16). Length / width ratio of antennomeres as: I 25 / 10: II 12 / 8: III 9 / 6.5: IV 9 / 6.5: V 9 / 6.5: VI 9 / 6.5: VII 9 / 6.5: VIII 9 / 6.5: IX 11 / 8: X 11 / 9: XI 30 / 14. Pronotum hardly longer than wide, strongly convex, somewhat shorter than head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal carinae dense, not reaching anterior pronotal margin, striae between carinae narrower than carinae. Anterior 0.02 to 0.03 mm of pronotal disc smooth. Pronotal pubescence moderately short. Elytron gradually impressed toward base, with discal and sutural carinae parallel or converging in anterior two thirds; discal carina wide as orwider than sutural carina; basal elytral margin carinate; anterior third of lateral margin carinate and hardly rounded, pubescence short, similar to pronotal pubescence. Male characters. Metaventrite with mesal impression delimited by large admesal ridges with inner sides well delimited, diverging apically. Surface of ridges oval, flat and completely covered by patches of short flat setae. Ventrite II with shallow mesal impression. Aedeagus (Figs XY) 0.34 - 0.35 mm long. Ventral process rather long, slightly bent, broad, at apex nearly as maximal aedeagal width. Parameres gradually widened apically (lateral view). Internal sac with two tufts of long spines. Measurements of the holotype (in mm). Total length 1.73; length / width of head 0.37 / 0.26, pronotum 0.34 / 0.32, elytra 0.40 / 0.63 (0.22), tergite I 0.55 / 0.75; length of maxillary palpomere IV 0.48, length of aedeagus 0.27.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289371EB3FD6385675570.taxon	diagnosis	Diagnosis. Upper part of temples vertex striate. Contours of temples rounded. Eyes not prominent. Mesal vertexal sulcus extended to level of highest point of vertex. Maxillary palpomere IV with peduncle much longerthan enlarged apical area. Baseof elytra impressed. Ventral process of aedeagus nearly as broad as median lobe. Parameres gradually widened in lateral view. Internal sac with two dense spinose tufts.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289371EB3FD6385675570.taxon	distribution	Distribution. East Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289361D18FB83838655F0.taxon	materials_examined	Holotype ♂: INDIA Darjeeling Dist., Tonglu 3100 m 16. X. 1978, LÖbl & Besuchet # 16 b under shrubs (MHNG). Paratypes: 3 ♂, 7 ♀, INDIA, with the same locality data as the holotype (MHNG, PCPH).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289361D18FB83838655F0.taxon	etymology	Etymology. The species epithet means sister in Hindi.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289361D18FB83838655F0.taxon	description	Description. Length 1.60 - 1.80 mm. Head about 1.6 times as long as wide. Frontal ridges extended to near level of posterior eye margins, narrowed posteriad. Frontal sulcus parallel-sided, widened posterior antennal tubercles, reaching mesal ridge about at level of eye mid-length. Mesal ridge delimited by narrow sulci joined posteriad to form vertexal sulcus. Vertexal sulcus reaching neck constriction. Setose patch in frontal sulcus about 0.08 - 0.10 mm long. Eyes in level of head mid-length, not prominent, with eight or nine facets. Temples with margins oblique or slightly concave in dorsal view, nearly angulate below eyes in dorsal view, lower and upper parts of temples separated by stria. Temples, vertex, and neck without vertical striae. Glabrous genal area not extended behind eyes. Temples and vertex anterior neck rounded, with inconspicuous, oblique and appressed pubescence. Segment IV of maxillary palpus with peduncle longer than enlarged apical area (ratio 21 / 17 to 22 / 15). Length / widthratio of antennomeres as: I 26 / 10: II 13 / 8: III 10 / 6: IV 8 / 6: V 8 / 6: VI 8 / 6: VII 10 / 6: VIII 9 / 6: IX 13 / 8: X 13 / 9: XI 28 / 13. Pronotum slightly longer than wide, strongly convex, nearlylong as head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal carinae dense, reaching near anterior pronotal margin, striae between carinae narrower than carinae. Pubescence short. Elytron impressed toward base, with sutural and discal carinae parallel, discal carina slightly wider than sutural carina and slightly bent; basal third to half of lateral margins oblique, carinate, pubescence similar to pronotal pubescence. Male characters. Metaventrite with mesal impression delimited by admesal ridges diverging apically, not overlapping metacoxae, their inner sides not clearly delimited, bearing long setae not forming compact patches. Middle of ventrite II throughout with narrow shallow impression. Aedeagus (Figs 13, 14) 0.48 - 0.53 mm long. Ventral process large, strongly bent, wider than half of median lobe. Parameres narrow in dorsal and lateral views. Internal sac lacking sclerotized pieces, with admesal spinose stripes and large, densely spinose tuft in apical half. Measurements of the holotype (in mm). Total length 1.80; length / width of head 0.43 / 0.27, pronotum 0.38 / 0.34, elytra 0.38 / 0.67 (0.21), tergite I 0.54 / 0.81; length of maxillary palpomere IV 0.41, length of aedeagus 0.45.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289361D18FB83838655F0.taxon	diagnosis	Diagnosis. Upper part of temples and vertex withouttransverse striae. Contours of temples oblique. Eyes not prominent. Vertexal sulcus reaching neck constriction. Maxillary palpomere IV with peduncle longer than enlarged apical area. Base of elytra impressed. Ventral process of aedeagus wider than half of median lobe. Parameres narrow in dorsal and lateral views Internal sac with admesal spinose stripes and large, densely spinose tuft in apical half.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC289361D18FB83838655F0.taxon	distribution	Distribution. India, West Bengal: Darjeeling District.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389361E55FB03846154D0.taxon	materials_examined	Holotype ♂: INDIA Darjeeling Distr., Tiger Hill 2200 - 2300 m 16. X. 78, Besuchet LÖbl # 13 (MHNG). Paratypes: 5 ♂, 6 ♀, INDIA, with the same locality data as the holotype (MHNG); 1 ♂, with the same data but 2500 - 2600 m, 18. X. (MHNG); 1 ♀, with the same locality data but 2200 m, 7. X. 1967, Gy. TopÁl (HNMB).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389361E55FB03846154D0.taxon	etymology	Etymology. The species epithet means face in Hindi.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389361E55FB03846154D0.taxon	description	Description. Length 1.60 - 1.85 mm. Head about 1.4 times as long as wide. Frontal ridges extended to near level of posterior eye margins, narrowed posteriad. Frontal sulcus parallel-sided, not widened posterior antennal tubercles, reaching mesal ridge about at level of anterior eye margin. Mesal ridge delimited by narrow striae converging posteriad, converging posteriad and forming sulcus extended to neck constriction. Setose patch in frontal sulcus about 0.08 - 0.09 mm long. Eyes in level of head mid-length, not prominent, with nine or ten facets. Temples appearing smooth, very finely punctate, with margins rounded in dorsal view, lower parts explanate, separated from upper parts by striae. Neck and area near neck with inconspicuous mesal carina. Temples and vertex anterior neck with inconspicuous, oblique and appressed pubescence. Segment IV of maxillary palpus with peduncle longer than enlarged apical area (ratio 21 / 15). Length / width ratio of antennomeres as: I 28 / 9: II 13 / 8: III 9 / 5: IV 9 / 5: V 9 / 5: VI 9 / 5: VII 9 / 5: VIII 9 / 5: IX 12 / 7: X 14 / 8: XI 26 / 12. Pronotum slightly longer than wide, strongly convex, nearly long as head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal carinae dense, reachingnearly anterior pronotal margin, striae between carinae narrower than carinae. Pubescence rather long. Elytron impressed toward base, withbasal margin carinate, sutural and discal carinae parallel, discal carina slightly wider than sutural carina and slightly bent; lateral margins throughout convex, carinate in basal third, pubescence rather long, to part longer than pronotal pubescence. Male characters. Metaventrite with mesal impression delimited by short admesal ridges diverging apically, not reaching metacoxae, sharp marginate, bearing at inner sides setae not forming compact patches. Middle of ventrite II with narrow, shallow impression delimited by dense oblique setae. Aedeagus (Figs 15, 16) 0.37 - 0.40 mm long. Ventral process narrow, at apex about as wide as fifth of median lobe. Parameres narrow and nearly straight in dorsal and lateral views. Internal sac without sclerotized pieces, with two narrow tufts of short spines overlapping denticulate lobe. Measurements of the holotype (in mm). Total length 1.83; length / width of head 0.38 / 0.27, pronotum 0.34 / 0.32, elytra 0.32 / 0.65 (0.22), tergite I 0.54 / 0.77; length of segment IV of maxillary palpus 0.38, length of aedeagus 0.40.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389361E55FB03846154D0.taxon	diagnosis	Diagnosis. Upper part of temples and vertex smooth. Contours of temples rounded. Eyes not prominent. Vertexal sulcus reaching neck constriction. Maxillary palpomere IV with peduncle longer than enlarged apical area. Base of elytra impressed, lateral elytral margins carinate. Ventral process of aedeagus narrow, about as fifth of median lobe. Parameres narrow. Internal sac without sclerites, with two tufts of short spines.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389361E55FB03846154D0.taxon	distribution	Distribution. India, West Bengal: Darjeeling District.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	materials_examined	Holotype ♂: INDIA Darjeeling Distr., Tonglu 3100 m, 16. X. 1978, LÖbl & Besuchet # 16 b under shrubs (MHNG). Paratypes: 15 ♂, 5 ♀, INDIA, with the same locality data as the holotype (MHNG, PCPH); 4 ♂, 2 ♀, with the same data but 2700 m (MHNG); 10 ♂, 4 ♀, NEPAL, Koshi Zone, Panchthar District, Dorpar Kharka, mature Rhododendron - Lithocarpus forest, 2700 m, 13. - 16. IV. 1988, J. Martens & W. Schawaller (SMNS, MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	etymology	Etymology. The species epithet means younger brother in Nepali.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	description	Description. Length 1.60 - 1.78 mm. Head about 1.5 times as long as wide. Frontal ridges extended to near level of posterior eye margins, narrowed posteriad. Frontal sulcus widened posterior antennal tubercles, parallel-sided, reaching minute tubercle about at level of eye mid-length and delimited by narrow striae joined posteriad to form vertexal sulcus. Vertexal sulcus nearly reaching neck constriction, followed by short mesal carina. Setose patch in frontal sulcus about 0.05 to 0.06 mm long. Eyes in level of head mid-length, hardly prominent, with eight or nine facets. Temples not explanate, with contours rounded in dorsal view. Temples and vertex above temples with oblique striae. Neck and area near neck without striae. Glabrous genal area not extended behind eyes. Temples and vertex anterior neck with conspicuous oblique appressed pubescence. Segment IV of maxillary palpus long as head, its apical thickened area somewhat longer than peduncle. Length / width ratio of antennomeres as: I 24 / 9: II 10 / 7: III 9 / 6: IV 7 / 6: V 8 / 6: VI 8 / 6: VII 8 / 6: VIII 7 / 6: IX 10 / 7: X 12 / 8: XI 26 / 12. Gular mound rounded. Gular setose patch reaching level of narrowest neck area. Pronotum slightly longer than wide, strongly convex, long nearly as head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal striae dense, reaching almost anterior pronotal margin, carinae between striae broader than carinae. Pubescence rather long apically. Elytron not impressed toward base, with sutural and discal carinae converging apically, discalcarina slightly wider than sutural carina; basal third of lateral margin oblique, not carinate, pubescence similar to pronotal pubescence. Male characters. Metaventrite with mesal impression delimited by admesal ridges diverging apically, not overlapping metacoxae, their inner sides not clearly delimited, bearing fine setae not forming compact patches. Middle of ventrite II throughout narrowly and shallowly impressed. Aedeagus (Figs 17 - 19) 0.48 - 0.53 mm long. Ventral process narrow, at apex broad as fifth to tenth of median lobe, bent. Parameres widened apically in dorsal and lateral views. Internal sac with two admesal sclerites to part overlapping very finely denticulate membranes. Measurements of the holotype (in mm). Total length 1.63; length / width of head 0.37 / 0.25, pronotum 0.36 / 0.31, elytra 0.45 / 0.63 (0.22), tergite I 0.50 / 0.78; length of maxillary palpomere IV 0.37, length of aedeagus 0.50.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	diagnosis	Diagnosis. Temples and vertex with oblique striae. Contours of temples rounded. Eyes hardly prominent. Vertexal sulcus nearly reaching neck constriction, followed by short carina. Maxillary palpomere IV with peduncle slightly shorter than thickened apical area. Base of elytra not impressed, lateral elytral margins not carinate. Ventral process of aedeagus narrow, wide as fifth to tenth of median lobe. Parameres widened apically in dorsal and lateral views. Internal sac with admesal sclerites and finely denticulate membranes.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	distribution	Distribution. India, West Bengal: Darjeeling District: Eastern Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC389351D32FA23808D56B0.taxon	discussion	Comments. The sclerites in the internal sac are expanded and definitively larger in specimens from Dorpar Kharka (Fig. 19) than in specimens from Tonglu (Fig. 17). Thus, it seems uncertain the whither the populations from the two sampled localities are comnspecific.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089351D18FCE385F25710.taxon	discussion	This genus comprises over 60 species and is widely distributed in the Old World and in Australia. An overview and key to its Asian members are lacking, and the available information on most included species is limited to their original descriptions. Threeare three species in the Himalayan collections examined. Two of them have maxillary palpomeres IV apically truncate with a sensory area in a circular impression. This character is not congruent with the definition of the genus as given by CHANDLER (2001). In addition, they lack carinate outer margins of the pronotal sulci, situated between the basal margin and the lateral antebasal foveae. The third Himalayan species has the maxillary palpomere IV with a sharply defined Vshaped sensory area but unlike other examined species lacks lateral pronotal sulci. The three Himalayan species have the gular moundweakly narrowed posteriad, with the lateral margins oblique or slightly convex and the apical margin nearly transverse. As only a single specimen is available for the two new species described below, we did not clear the elytra to verify the number of the basal foveae and the state of the hind wings. However, their short elytra indicate atrophy of hind wings, a characterstate correlated with reducedeyes. The third unnamed species has the hind wings atrophied and the elytra each with three basal foveae, the outer foveae being oblique, and the antebasal foveae are lacking.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	materials_examined	Holotype ♂: Zentral-Nepal Sept. - Okt. 1971 lg. H. Franz / Umg. Goropani w. Pokhara (NHMW).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	etymology	Etymology. The species epithet means alone in Nepali.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	description	Description. Length about 1.70 mm. Headabout 1.3 longer than wide. Anterior margin of frontal rostrum truncate, setose patch between antennal tubercles absent. Frontal sulcus deep, slightly widened posteriad, glabrous, step-like raised to continue as mesal impression of vertex. Frontal carinae oblique posterior antennal tubercles, with sharply delimited inner and lower outer margins, punctate, bearing short setae oriental posteriad. Vertex strongly raised, convex, impressed mesally, punctate and with short setae. Vertexal foveae setose. Mesal vertexal carina absent. Eyes prominent, with diameter clearly shorter than interval to neck, consisting of twelve facets. Genae smooth. Temples in lateral view with rounded contours. Temples and vertex with shortappressed setae. Maxillarypalpomere IVlong as head, with thickened apical part about 1.5 times as long as peduncle, apex truncate, margins of sensory area circular and carinate, lateral longitudinal sulcus very shallow and narrow, appearing as fine stria, withclearly defined margins onlyat apex. Scape reticulate. Length / width ratioof antennomeresas: I 26 / 9: II 12 / 8: III 8 / 6: IV 8 / 6: V 8 / 6: VI 9 / 6: VII 9 / 6: VIII 8 / 6: IX 12 / 8: X 13 / 9: XI 25 / 13. Gular-mandibular carina branched at level of anterior margin of gular mound, inferior branch defining laterally gular mound. Gular mound nearly globose, hardly narrowed posteriad, not ridged, gular foveae obscured by setose patch. Pronotum about 1.2 timesas long as wide, hardly longer than head, median antebasal fovea indistinct, joined to short impression, antebasal sulcus narrow, distinct, lateralantebasalfoveaedistinctand setose, lateral sulcus between lateral antebasal fovea and basalmargin narrow, not carinate. Discal setae short, appressed. Elytra slightly longer than pronotum, strongly widened apically, at apices well three times as wide as at bases, not microsculptured, flat near bases, with three distinct basal foveae, basal margins not raised, lateral margins rounded, adsuturalandmedian longitudinalcarinae fine, median longitudinal carina clearly visibly in basal two thirds of elytral length, setae covering entire surface, short and appressed as on pronotum. Abdomen with tergite I widened apically, slightly wider than elytra, with broad paratergites, rounded lateral margins, pubescence short and appressed. Male characters. Mesal area of metaventrite impressed and glabrous, delimited laterally by ridges. Top of ridges truncate, forming oval setose pockets. Aedeagus (Figs. 20, 21) 0.37 mm long, with median lobe oval, apical process abruptly bent and gradually narrowed in lateral view. Parameres weakly sclerotized, moderately bent, with apical and subapical setae, in lateral view widest near apices. Internal sac with narrow, sinuate rod and membranous vesicle bearing extremely short spines. Measurements of the holotype (in mm). Total length 1.72; length / width of head 0.34 / 0.27, pronotum 0.35 / 0.30, elytra 0.38 / 0.68 (0.21), tergite I 0.50 / 0.70; length of maxillary palpomere IV 0.34, length of aedeagus 0.37.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	diagnosis	Diagnosis. Length about 1.7 mm. Gular-mandibular carina branched. Eyes small, with diameter as length of temples. Antennal club three-segmented. Maxillarypalpomere IV truncate at apex, with very narrow lateral stria, densely and very finely punctate and bearing short erect setae. Thickened part of maxillary palpomere IV longer than peduncle. Pronotum hardly longer than head, with short lateral sulci extending from lateral antebasal foveae to basal margin. Elytra slightly longer than pronotum. Internal sac of aedeagus with single narrow sinuate rod.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	distribution	Distribution. Western Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC089341D18F96384AA5090.taxon	discussion	Comments. This new species can easily be distinguished from the type species of Pselaphaulax, P. dresdensis (Herbst, 1791), and from the other six Palaearctic congeners, by its small eyes, the pronotum lacking carinae between the lateral antebasal foveae and the basal margin, and the elytra much shorter than the ventrite II. The few other Asian species of Pselaphaulax, P. articularis (Schaufuss, 1877), P. bifoveolatum (Schaufuss, 1877), P. ceylanicus Jeannel, 1961 and P. vestitus (Raffray, 1904), are from the tropics. According to their respective descriptions, they may be distinguishedfrom P. eklai bythe long elytra. As the type material held in the Muséum national d’Histoire naturelle in Paris is at present unavailable, we were not able to study types of these species.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC189341D32FDE382C85677.taxon	materials_examined	Holotype ♂: NEPAL Kosi ELamjura Pass, 2950 m, 21. X. 1984, L. Deharveng (MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC189341D32FDE382C85677.taxon	etymology	Etymology. The species epithet refers to the ethnic group living mainly in Eastern Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC189341D32FDE382C85677.taxon	description	Description. Length about 1.70 mm. Most characters as in S. eklai. Head longer, 1.45 times as long as wide. Eyes about as long as temples, with 10 facets. Maxillary palpomere IV somewhat shorter than head. Length / width ratio of antennomeres as: I 26 / 11: II 11 / 8: III 6 / 6: IV 6 / 6: V 6 / 6: VI 6 / 6: VII 7 / 6: VIII 7 / 6: IX 1208: X 10 / 9: XI 24 / 13. Pronotum distinctly shorter than head, long as wide. Elytral carinae longer and more distinct, reaching apical fourth. Tergite Ishorter than head, with nearly oblique lateral margins. Male characters. Mesal area of metaventrite impressed, glabrous, delimited by high lateral ridges gradually narrowed to acute tip. Aedeagus (Figs. 22, 23) 0.28 mm long, with median lobe oval, apical process short, weakly bent and gradually narrowed in lateral view. Parameres weakly sclerotized, in axis with median lobe, strongly widened, in lateral view widest posterior apical third, with single apical seta. Internal sac lacking sclerotized structures or spinose membranes. Measurements of the holotype (in mm). Total length 1.45; length / width of head 0.38 / 0.27, pronotum 0.32 / 0.32, elytra 0.36 / 0.61 (0.23), tergite I 0.35 / 0.75; length of maxillary palpomere IV 0.32, length of aedeagus 0.28.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC189341D32FDE382C85677.taxon	distribution	Distribution. Eastern Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFC189341D32FDE382C85677.taxon	discussion	Comments. This species may be distinguished from P. eklai by the longer head, the eyes long as temples, the antennomeres III to VI long as wide, the pronotum long as wide, the discal carinae of elytra longer, extending onto apical fourth of the disc, the aedeagus much smaller, with broader parameres bearing each a single apical seta, and the internal sac lacking sclerotized structures.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	materials_examined	Holotype ♂: NEPAL Parbat Distr. Punhill at Goropani Pass 3050 - 3100 m 8. X. 83 Smetana & LÖbl (MHNG). Paratypes: 5 ♂, 3 ♀, NEPAL, with the same locality data as the holotype (MHNG); 3 ♂, 3 ♀, Parbat District, ridge E Goropani, 3100 m, 7. X. 1983, I. LÖbl & A. Smetana (MHNG); 6 ♂, 10 ♀, Parbat District, Goropani, north slope, 2750 m, 5. X. 1983, I. LÖbl & A. Smetana (MHNG, NMNT, PCPH); 3 ♂, 5 ♀, Parbat District, Goropani Pass, 2700 m, 6. X. 1983, I. LÖbl & A. Smetana (MHNG); 1 ♂, Parbat District, near Goropani, IX-X. 1971, H. Franz (NHMW); 3 ♂, Kali-Gandaki Valley, forest above Nawronkot near Larjung, 2900 - 3000 m, 3. X. 1978, H. Franz (NHMW); 2 ♂, 3 ♀, Kali Gandaki Valley, 2 km N Kalopani, 2550 m, 1. X. 1983, I. LÖbl & A. Smetana (MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	etymology	Etymology. The species epithet means strong in Nepali.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	description	Description. Length 1.55 - 1.70 mm. Head about 1.6 longer than wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Frontal sulcus parallel-sided, with carinate inner margins, reaching nearly level of posterior eye margins, frontal carinae delimited laterally by oblique impressions. Setose patch between antennal tubercle lacking in some specimens. Vertex convex, lacking distinct mesal impression, mesal carina near neck absent. Eyes prominent, with diameter as half of interval to neck, consisting of eight facets. Genae rugose. Temples weakly rounded, in dorsal view with nearly oblique contours. Margins of frontal carinae bearing each row of short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV long as or somewhat longer than head, with peduncle about 1.4 times as long as thickened apical part. Length / width ratio of antennomeres as: I 28 / 9: II 12 / 8: III 8 / 6: IV 8 / 6: V 8 / 6: VI 8 / 6: VII 8 / 6: VIII 8 / 6: IX 11 / 8: X 13 / 9: XI 25 / 15. Gular mound circular, not ridged, gular foveae obscured by setose patch. Pronotum about 1.4 times as longas wide, long as or slightly longer than head, with punctiform median antebasal fovea, distinct lateral antebasal foveae, hardly visible antebasal transverse stria. Elytra hardly longer than pronotum, strongly widened apically, at apices nearly three times as wide as at bases, not microsculptured, with distinct three basal foveae, outer margin of median longitudinal carina clearly delimited, inner margin indistinct, adsutural setose row not reduced, median setose row double. Hind wings atrophied. Abdomen with tergite I wider than elytra, broad paratergites, rounded lateral margins. Male characters. Mesal area of metaventrite flat, glabrous, delimited laterally by slightly raised, punctate ridges bearing rather long setae. Ventrite II without sexual characters. Aedeagus (Figs 24, 25) 0.27 - 0.30 mm long. Median lobe oval, prominent apically to form short lamina covering ostium. Ventral process apical, inflexed, slightly bent and gradually narrowed in lateral view. Internal sac with single, weakly sclerotized rod. Parameres bent and widened apically, with four rather long setae, very weakly sclerotized inner sides. Measurements of the holotype (in mm). Total length 1.60; length / width of head 0.36 / 0.22, pronotum 0.38 / 0.28, elytra 0.40 / 0.61 (0.21), tergite I 0.43 / 0.68; length of maxillary palpomere IV 0.44, length of aedeagus 0.28.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	diagnosis	Diagnosis. Frontal sulcus parallel-sided. Eyes small, with diameters about half that of temples. Pronotum with median and lateral antebasal foveae and hardly visible transverse antebasal stria. Ventrite II without obvious male characters. Aedeagus with median lobe oval, nearly symmetrical, with short lamina covering ostium and obliquely inflexed ventral process appearing nearly pentagonal in dorsal view. Parameres widened apically and bearing long setae.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	distribution	Distribution. Western Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FC2385E55717.taxon	discussion	Comments. This species shares with P. emeishanus Nomura, 2003 and P. huqin Nomura, 2003 a lamina above the ostium, the base of the ventral process of the aedeagus situated near aedeagal apex, and the internal sac lacking a brush-like lobe. In addition, it possesses like these two Chinese species an antebasal pronotal stria. It differs markedly in the distinct median antebasal fovea, the tip of the ventral process of the aedeagus not abruptly bent and the parameres widened apically in lateral view and bearing long setae.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCE893B1EB3FD83822952D0.taxon	discussion	This genus comprises 76 species (NEWTON 2020), many of them described from the temperate part of the Euro - Asian continent. The group was redefined by NOMURA (1998) and its east Asian members were keyed in NOMURA (2003). Two species have been collected in Nepal, one is described below, the second represented by two females, is probably also new. We decline to describe it in absence of at least one available male.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF893A1E55FD0382F55490.taxon	type_taxon	Type species Nepallaphus ekam n. sp. Gender masculine.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF893A1E55FD0382F55490.taxon	etymology	Etymology. The epithet is derived from Nepal, combined with “ laphus ”, a meaningless part of the generic name Pselaphus.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF893A1E55FD0382F55490.taxon	description	Description. Length 1.70 - 1.90 mm. Pubescence short, appressed and sparse, excepted on middle of metaventrite, and for patches of white squamous setae covering venter of head posterior gular mound, most of proventrite and mesoventrite, apices of elytra, apical margin of metaventrite and bases of tergite Iand ventrite II. Head narrow, elongate, with frontal rostrum prominent, antennaltubercles smooth, postantennal notches absent. Frontal ridges extended to level of eye mid-length. Mesal sulcus glabrous, narrow, parallel-sided or somewhat widened apically, extended onto vertical part of frontoclypeus, lacking setose patch between antennal tubercles. Anterior margin of frontal rostrum notched in dorsal view. Vertex abruptly raised. Vertexal foveae setose, in globous pocket, oblique, situated at level of posterior eye margin. Eyes small, prominent, anterior eyes margins about at head-mid-length. Temple rounded. Genae reticulate, explanate. Gular-mandibular carinae conspicuous, forming prominent lateroventral margin of anterior part of head. Antennae 11 - segmented, antennomeres Iand II reticulate, club 3 - segmented, weekly formed, segment XI asymmetrical. Maxillary palpi elongate, segment Inarrow, cylindrical, bent, segment II long, pedunculate, about 3 times as long as segment I, longer than half of head length, with peduncle much longer than enlarged apical part, slightly swollen at base, segment III short, subtriangular, long about as wide, segment IV pedunculate, bent, lacking sensory area and without apical notch and lateral sulcus, basal part of peduncle glabrous, apical part of peduncle and enlarged apical part appearing reticulate, densely, very finely studded, with short, bent setae. Gula strongly swollen, gular lacking striae. Gular foveae narrowly separated, obscured by setose patch. Pronotum evenly convex, rounded laterally, widest about at mid-point, with distinct antebasal lateral foveae, punctiform antebasal median fovea, without antebasal sulcus, basal area glabrous. Elytra short, strongly widened apically, not impressed basally, basal margin not raised, with three basal foveae, middle basal fovea oblique, antebasal fovea lacking, discal carinae close to adsutural carinae or absent. Hind wings atrophied. Proventrite with setose patch reaching nearly lateral margins, obscuring foveae. Mesoventrite covered by setose patch; foveae obscured. Metaventrite asetose. Legs without spines Femora clavate, reticulate. Tibiae thickened apically. Third tarsomeres subcylindrical, narrower and longer than second tarsomeres. Abdomen with tergite Imuch longer than rest of tergites combined, convex, widened apically, with rounded lateral margins, deep basal sulcus and large basolateral and mediobasal foveae obscured by rows of flat setae; apical margins of tergites broadly rounded. Ventrite II with deep basal sulcus obscured by rows of flat setae. Aedeagus symmetrical, median lobe sub-oval, with strongly sclerotized ventral process, parameres asetose, weakly sclerotized. Basal bulb delimited apicodorsally by transverse sclerotizedband, with conspicuous dorsal vesicle below window in bulb’s dorsal sclerotization, attached by muscles to ventral side of bulb; internal sac with two narrow admesal rods.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF893A1E55FD0382F55490.taxon	discussion	Comments. Members of this genus resembles Pselaphus Herbst, 1791, as redefined by BESUCHET (1999). Adults of Nepallaphus have maxillary palpomeres IV pedunculate, densely setose, lacking a sensory area in the apical notch and lateral sulcus. They share these characters with the type species of Pselaphus, P. heisei Herbst, 1791, and with numerous other species currently placed in Pselaphus. They also share with Pselaphus the frontal sulcus lackinga setose patch between the antennal tubercles. Nevertheless, some members of Pselaphus, suchas P. kubischteki (Reitter, 1891) and P. lomnickii (Reitter, 1901), differ markedly in palpal characters, P. kubischteki also differs in possessing extended setose patches covering the temples, basolateral areas of pronotum and bases of elytra (pers. observation). Thus, there is an obvious need to revisit the genus. However, Nepallaphus is clearly distinguished from Pselaphus by the pronotum and elytra possessing antebasal and basal foveae, respectively. In addition, the new genus also differs in possessing microtuberculate palpomeres IV, while the palpomeres IV bear prominent tubercles in Pselaphus heisei and in many other species currently considered members of Pselaphus. The basal bulb of the aedeagus clearly separates Nepallaphus from Pselaphus based on the presence a dorsal vesicle.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF893A1E55FF6383AC50B0.taxon	discussion	The genus comprises seven species. One of them, P. indicus (Jeannel, 1960), has been reported from the Indian subcontinent. It was described from Dehra Dun and placed in Trichopselaphus Jeannel, 1949, a junior homonym replaced by Pselaphotrichus. The newly examined collections comprise a single female that clearly differs from P. indicus by the much longer scape and long erect setae situated on the pronotal and elytral discs. This species is probably new. We decline to describe it in absence of at least one available male.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	materials_examined	Holotype ♂: NEPAL (Prov. Bagmati) near Mere Dara, 3000 m, 7. IV. 1981 LÖbl & Smetana (MHNG). Paratypes: 5 ♂, 5 ♀, NEPAL, with the same data as the holotype (MHNG, PCPH); 1 ♂, 1 ♀, Bagmati Zone, Mere Dara, 3200 m, north slope, 8. IV. 1981, I. LÖbl & A. Smetana (MHNG); 1 ♂, 7 ♀, Bagmati Zone, below Thare Pati, 3300 m, 9. IV. 1981, I. LÖbl & A. Smetana (MHNG, PCVB).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	etymology	Etymology. The species epithet is the number one in Sanskrit.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	description	Description. Length 1.70 - 1.90 mm. Head about 1.6 times as long as wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Anterior margin of frontal rostrum prominent, notched in middle, bidentate. Frontal sulcus narrow, with carinate inner marginsextended to level of eye mid-length. Frontal carinae horizontal, reticulate. Antennal tubercles prominent. Vertex convex, lacking distinct mesal impression, mesal carina near neck absent. Eyes prominent, with diameter as half of interval to neck, consisting of seven to eight facets. Genae reticulate. Temples weakly rounded, in dorsal view with nearly oblique contours. Frontal carinae bearing rows of short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV longer than head, with peduncle about as long as thickened apical part. Length / width ratio of antennomeres as: I 24 / 10: II 12 / 9: III 10 / 6: IV 9 / 6: V 9 / 6: VI 9 / 6: VII 10 / 6: VIII 10 / 6: IX 13 / 8: X 14 / 9: XI 28 / 14. Pronotum slightly longer than wide and slightly shorter than head, with punctiform median antebasal fovea, distinct lateral antebasal foveae, punctation sparser than on head. Elytra about 1.4 times as long as pronotum, with lateral margins rounded, at apices well 4 times as wide as at bases, not microsculptured, with three basal foveae, median longitudinal carina with margins well delimited, extending to or nearly to apical third of elytral disc, adsutural and median setose rows simple, disc with outer double row od setae, intervals between rows of setae glabrous. Hind wings atrophied. Abdomen with tergite Islightly wider than elytra, withbroad paratergites and roundedlateral margins. Male characters. Middle of metaventrite raised to form a flat, circular area inflexed apically and completely covered by short, appressed setae. Aedeagus (Figs. 26, 27) 0.49 - 0.53 mm long, with basal bulb not widened apically, its sclerotized apicodorsal band not notched, dorsal capsule cordiform. Measurements of the holotype (in mm). Total length 1.76; length / width of head 0.39 / 0.25, pronotum 0.35 / 0.32, elytra 0.49 / 0.76 (0.17), tergite I 0.50 / 0.83; length of maxillary palpomere IV 0.43, length of aedeagus 0.50.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	diagnosis	Diagnosis. Length 1.70 - 1.90 mm. Anterior margin of frontal rostrum prominent andnotched in middle. Frontal sulcus narrow, parallel-sided. Maxillary palpomere IV longer than head, with peduncle about as long as thickened apical part. Elytron with discal carina extending to or nearly to apical third of disc. Interval between carinae and laterally discal carina glabrous. Aedeagus with basal bulb not widened apically, its apicodorsal band not notched, dorsal capsule cordiform.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	distribution	Distribution. Central Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCF89391D32F9E3801C57D0.taxon	discussion	Comments. See key below to distinguish this species from its congeners.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	materials_examined	Holotype ♂: NEPAL (Prov. Bagmati) Pokhare NE Barahbise 2800 m 2. V. 81 LÖbl & Smetana (MHNG). Paratypes: 1 ♂, 2 ♀, NEPAL, Bagmati Zone, Yardang Ridge NE Barahbise, 3250 m, 5. IV. 1981, I. LÖbl & A. Smetana (MHNG); 1 ♂, Bagmati Zone, Kalingchok NE Barahbise, Amatal Khola, Tingsang La, 3100 m, 27. X .. 1981, P. Cassagnau (MHNG); 1 ♀, Bagmati Zone, Kalingchok NE Barahbise, Yardang Ridge, 3250 m, 28. X. 1981, P. Cassagnou (MHNG).	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	etymology	Etymology. The species epithet is the numbertwo in Sanskrit.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	description	Description. Length 1.60 - 1.82 mm. Head about 1.7 times as long as wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Anterior marginof frontal rostrum rounded. Frontal sulcus between antennal tubercles narrow, widened posteriad, with carinate inner margins, extended to level of eye mid-length. Frontal carinae oblique, reticulate. Antennal tubercles prominent. Vertex convex, with mesal impression very shallow, narrowed posteriad, mesal carina near neck absent. Eyes prominent, with diameter about as half of interval to neck, consisting of seven facets. Genae reticulate. Temples weakly rounded, contours in dorsal view nearly oblique. Frontal carinae bearing rowsof short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV long about as head, with peduncle somewhat shorter than thickened apical part. Length / width ratio of antennomeres as: I 22 / 10: II 12 / 8: III 9 / 6: IV 7 / 6: V 7 / 6: VI 8 / 6: VII 8 / 6: VIII 8 / 6: IX 10 / 7: X 12 / 8: XI 24 / 13. Gular mound longer than wide, narrowed posteriad. Pronotum similar to that in N. ekam. Elytra nearly 1.4 times as long as pronotum, with lateral margins weakly rounded, at apices about 3.5 times as wide asat bases, not microsculptured, with three basal foveae, median longitudinal carina with margins well delimited, reaching nearly elytral mid-length, bearing double rows of setae, adsutural carina with single row of setae; disc with additional, outer double row of setae, intervals between rows of setae glabrous. Hind wings atrophied. Abdomen with tergite I slightly wider than elytra, broad paratergites, rounded lateral margins. Male characters. Metaventrite as in N. ekam. Aedeagus (Figs 28, 29) 0.35 - 0.37 mm long, with basal bulb not widened apically, its apicodorsal band narrowly notched in middle, dorsal capsule oval. Measurements of the holotype (in mm). Total length 1.82; length / width of head 0.41 / 0.24, pronotum 0.33 / 0.30, elytra 0.45 / 0.70 (0.20), tergite I 0.51 / 0.75; length of maxillary palpomere IV 0.40, length of aedeagus 0.37.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	diagnosis	Diagnosis. Length 1.60 - 1.82 mm. Anterior margin of frontal rostrum rounded. Frontal sulcus rather wide, distinctly widened posteriad. Maxillary palpomere IV long about as head, with peduncle shorter than thickened apical part. Elytron with discal carina short, extended to mid-length of disc, interval between carinae and laterally discal carina glabrous. Aedeagus with basal bulb not widened apically, oval apicodorsal capsule, and minute notch in middle of apicodorsal band.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	distribution	Distribution. Central Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89391EB3F92382F85670.taxon	discussion	Comments. See key below to distinguish this species from its congeners.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	materials_examined	Holotype ♂: Népal 15. VII. 1981 Langtang N. Park près Gosaikunda Lake 4400 m Beron [handwritten] (NMNS)	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	etymology	Etymology. The species epithet is the number three in Sanskrit.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	description	Description. Length 2.10 mm. Headabout 1.6 longer than wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Middle of anterior margin of frontal rostrum shallowly notched, bidentate. Frontal sulcus between antennal tubercles narrow, parallel to level of anterior eye margins, widened between eyes, with carinate inner margins, extended to level of eye mid-length. Frontal carinae oblique, reticulate. Antennal tubercles prominent. Vertex convex, with mesal impression very shallow, narrowed posteriad, mesal carina near neck absent. Eyes prominent, with diameter about as half of interval to neck, consisting of eight facets. Genae reticulate. Temples weakly rounded, in dorsal view with nearly oblique contours. Frontal carinae bearing rows of short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV long as head, with peduncle somewhat shorter than thickened apical part. Length / width ratio of antennomeres as: I 26 / 11: II 12 / 9: III 8 / 6: IV 8 / 6: V 8 / 6: VI 8 / 6: VII 8 / 6: VIII 8 / 6: IX 10 / 7: X 12 / 9: XI 26 / 15. Gular mound wider than long, nearly circular. Pronotum similar to that of N. ekam. Elytra about 1.3 times as long as pronotum, with lateral margins oblique in basal halves, weakly rounded posteriad mid-length, at apices nearly 4 times as wide as at bases, not microsculptured, with three basal foveae, inner two foveae in elongate impression, median longitudinalcarina absent, adsutural carina with single row of setae; entire disc setose. Abdomen with tergite I slightly wider than elytra, broad paratergites, rounded lateral margins. Male characters. Metaventrite as in N. ekam. Aedeagus (Figs 30, 31) 0.62 mm long, with basal bulb widened apically, its apicodorsal band deeply notched in middle, dorsal capsule oval. Measurements of the holotype (in mm). Total length 2.10; length / width of head 0.41 / 0.26, pronotum 0.36 / 0.31, elytra 0.48 / 0.75 (0.20), tergite I 0.64 / 0.84; length of maxillary palpomere IV 0.40, length of aedeagus 0.62.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	diagnosis	Diagnosis. Length 2.10 mm. Anterior margin of frontal rostrum bidentate. Frontal sulcus rather wide, distinctly widened posteriad. Maxillary palpomere IV long as head, with peduncle shorter than thickened apical part. Elytron without discal carina, pubescence covering entire elytral disc. Basal bulb of aedeagus widened apically, its apicodorsalband deeply notched.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	distribution	Distribution. Central Nepal.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
517F87A6FFCC89381D18F88383CE57D0.taxon	discussion	Comments. The state of the hind wings was not examined. The narrow elytral bases and the small eyes suggest reduced hind wings. See key to distinguish the species from its congeners.	en	Löbl, Ivan, Kodada, Ján (2021): On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae). In: Biodiversität und Naturausstattung im Himalaya VII. Erfurt: Naturkundemuseum: 349-368, DOI: http://doi.org/10.5281/zenodo.5793818
