taxonID	type	description	language	source
370987F0FFADFFA3A2F5FD85FDF8FB6B.taxon	description	Figs 2 – 3	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFADFFA3A2F5FD85FDF8FB6B.taxon	diagnosis	Diagnosis Russula adwanitekae sp. nov. is mainly separated from similarly looking species with reddish lilac, orchid purple or greyish to deep magenta colored pileus in subsect. Laricinae by its sequence data (nrITS) and geographic distribution.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFADFFA3A2F5FD85FDF8FB6B.taxon	etymology	Etymology After the name of the forest locality, Adwani-Teka.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFADFFA3A2F5FD85FDF8FB6B.taxon	materials_examined	Material examined Holotype INDIA • Uttarakhand, Pauri Garhwal, Adwani-Teka forest; 30 ° 05.681 ′ N, 78 ° 43.890 ′ E; alt. 1989 m a. s. l.; in temperate mixed forest; 3 Oct. 2016; A. Ghosh, AG 16 - 1430; GenBank: MN 263242 (ITS); CAL [1821]. Paratype INDIA • Uttarakhand, Pauri Garhwal, Adwani-Teka forest; 30 ° 05.675 ′ N, 78 ° 43.880 ′ E; alt. 1953 m a. s. l.; in temperate mixed forest; 5 Oct. 2016, A. Ghosh, AG 16 - 1435; GenBank: MN 263243 (ITS); CAL [1822]. MycoBank: MB 838571; Index Fungorum number: IF 558126; Facesoffungi number: FoF 09581	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFADFFA3A2F5FD85FDF8FB6B.taxon	description	Description Pileus small to medium-sized, 36 – 53 mm diam., convex when young, becoming plano-convex, applanate to uplifted with maturity, broadly depressed centre, near the margin becoming tuberculately striate and more or less straight with age, entire; pileipellis peeling to mid-radius; surface dry, shiny and viscid when moist, glabrous, reddish lilac (14 C 3 – 5), orchid purple (14 C 8) or greyish magenta (14 D 5 – 7) to deep magenta (14 D 8) and centrally dark magenta (13 – 14 F 4 – 8). Pileus context firm, brittle, yellowish white (1 – 2 A 2), unchanging after bruising, on exposure or with age. Lamellae adnexed to subdecurrent, equal but mixed with some dispersed lamellulae of different length, close (10 – 12 / cm at pileus margin), cream to pale yellow or light yellow (4 A 2 – 4), up to 6 mm broad, occasionally forked near the stipe apex; lamella edge even and concolorous. Stipe 35 – 51 × 7 – 14 mm, cylindrical to subclavate, tapered at the apex, central, solid but not firm; surface dry, finely longitudinally venose, chalky white (1 – 2 A 1) but with yellowish white to pale yellow (4 A 2 – 3) flush near its middle portion, changing greyish white (1 – 2 B 1) after bruising. Stipe context solid, chalky white (1 – 2 A 1), changing dirty white or greyish white (1 – 2 B 1) after bruising or on exposure; turning reddish brown (8 E 7 – 8) to dark brown (8 F 7 – 8) with application of guaiacol. Odour indistinctive. Taste mild. Spore print not obtained. Basidiospores subglobose to broadly ellipsoid, rarely ellipsoid, (7 –) 7.73 – 8.23 – 8.72 (– 9.5) × (6 –) 6.6 – 6.93 – 7.3 (– 8) μm, Q = (1.07 –) 1.13 – 1.19 – 1.25 (– 1.33), ornamentation amyloid (up to 1.7 µm high), consisting of thick ridges and warts forming incomplete reticulum, with some isolated intermediate warts; suprahilar plage amyloid; apiculi up to 2 µm high. Basidia (29 –) 32 – 35 – 38 (– 40) × (10 –) 10 – 11 – 13 (– 15) µm, 4 - spored, subclavate to clavate; sterigmata up to 8 µm long. Subhymenium layer up to 37 µm thick, pseudoparenchymatous. Hymenial cystidia on lamellar sides (55 –) 61.5 – 71.5 – 81 (– 92) × (7 –) 8 – 9.5 – 10.5 (– 11) µm, cylindrical, subclavate to fusiform with frequent lageniform (up to 23 µm long) or appendiculate or few rounded-obtuse apices, emergent up to 32 µm beyond the basidiole tips; contents crystalline, without reaction in sulfovanillin. Lamellae edges fertile with basidia and cystidia. Hymenial cystidia on lamellar edges (37 –) 42.5 – 52.5 – 62.5 (– 66) × (6 –) 6 – 7 – 8 (– 9) µm, cylindrical to subclavate with lageniform (up to 20 µm long) or appendiculate or rounded apex; contents crystalline, without reaction in sulfovanillin. Hymenophoral trama mainly composed of large nests of sphaerocytes and few hyphal elements. Pileipellis orthochromatic in Cresyl Blue, sharply delimited from the underlying sphaerocytes of the context, 100 – 120 μm thick, two-layered, distinctly divided in 40 – 50 μm deep suprapellis composed of erect or ascending hyphal terminations, arranged in a trichodermal structure and dispersed pileocystidia, and subpellis 60 – 70 μm deep, composed of more or less dense, horizontally oriented hyphae. Acid-resistant incrustations absent. Hyphal terminations near the pileus margin usually branched at the subterminal cells or the cells just below, thin-walled; terminal cells (11 –) 19 – 28.5 – 37.5 (– 50) × 3 – 4 – 4.5 (– 6) μm, mainly subulate, sometimes cylindrical to subcylindrical, apically obtuse or slightly narrowed towards tips and wider near base or sometimes attenuated; subterminal cells usually equal in size, sometimes with lateral branches; near the pileus centre with slightly shorter and less wide terminal cells, measuring (15 –) 18.5 – 25 – 32 (– 44) × (2 –) 2.5 – 3 – 3.5 (– 4) μm, but equally wide subterminal cells. Pileocystidia near the pileus margin 1 – 4 - celled, cylindrical, usually originating deep in subpellis and often originating from branched subterminal cells, thin-walled; terminal cells (23 –) 40.5 – 62 – 83 (– 110) × (4 –) 4.5 – 5 – 6 (– 7) μm, cylindrical or sometimes slightly tapered towards tips, rounded-obtuse apex, without any incrustations; contents crystalline, without reaction in sulfovanillin; those near the pileus centre with 0 – 3 septa and shorter terminal cells (15 –) 25.5 – 38 – 51 (– 70) × (3.5 –) 4 – 5 – 5.5 (– 7) μm. Clamp connections absent from all tissues.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFAEFFAFA28EFAA4FD5FFE02.taxon	description	Figs 4 – 5	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFAEFFAFA28EFAA4FD5FFE02.taxon	diagnosis	Diagnosis Russula purpureozonata sp. nov. is a blackening species with a dark purplish concentric zonation on the pileus surface and otherwise typical features of subsect. Decolorantes; it differs from the microscopically similar, North American R. burkei Burl. in its mild taste and ectomycorrhizal association with Abies densa, and from North American R. californiensis and R. magna in its distinctly smaller spores.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFAEFFAFA28EFAA4FD5FFE02.taxon	etymology	Etymology Refers to the dark purplish concentric zones on the pileus surface.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFAEFFAFA28EFAA4FD5FFE02.taxon	materials_examined	Material examined Holotype INDIA • Sikkim, East district, Memeinchu; 27 ° 21.108 ′ N, 88 ° 49.660 ′ E; alt. 3539 m a. s. l.; on the soil under Abies densa; 2 Aug. 2018; K. Das, KD 18 - 003; GenBank: MN 267570 (ITS); CAL [1817]. Additional material INDIA • Sikkim, East district, opposite to Gnathang firing range forest; 27 ° 18.605 ′ N, 88 ° 48.794 ′ E; alt. 3885 m a. s. l.; on the soil under Abies densa; 5 Aug. 2018; K. Das, KD 18 - 15 (CAL 1818); GenBank: MN 269951 (ITS); CAL [1818]. MycoBank: MB 838572; Index Fungorum number: IF 558127; Facesoffungi number: FoF 09580	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
370987F0FFAEFFAFA28EFAA4FD5FFE02.taxon	description	Description Pileus large-sized, 85 – 105 mm diam., hemispherical when young, then convex, plano-convex to applanate, broadly but shallowly depressed in the centre when mature; margin decurved to plane with maturity, entire; surface viscid when moist, then dry, finely areolate, peeling to ½ of the radius, greyish yellow (2 C 3 – 4) or olive to linden green (2 D 5 – 6), centrally dark brown (7 F 4 – 5) to dark purple or purple-black with pastel yellow to light yellow patches (3 A 4 – 5), fading towards mid in distinctive concentric zones. Pileus context up to 8 mm thick, thinning towards margin, firm, brittle, chalky white (1 – 2 A 2), changing first orange-red, then black when cut or bruised; turning greyish red to reddish brown (8 C – D 5) and dull green (26 E 3 – 4) with guaiacol and FeSO 4, respectively. Lamellae adnexed to free, subdistant (6 – 7 / cm at pileus margin), yellowish white (2 – 3 A 2), forked near the stipe apex; edge even and concolorous. Stipe 70 – 100 × 20 – 30 mm, subclavate to clavate, tapered at the apex, central, solid; surface dry, finely longitudinally venose, chalky white (1 – 2 A 1) with yellowish white to pale yellow (4 A 2 – 3) flush at one side of the centre, changing first orange-red, then black when cut or bruised. Stipe context solid, chalky white (1 – 2 A 1), changing first orange-red, then black when cut or bruised; turning greyish red to reddish brown (8 C – D 5) and dull green (26 E 3 – 4) with guaiacol and FeSO 4, respectively. Odour indistinctive. Taste mild. Spore print yellowish white (3 A 2). Basidiospores globose, subglobose to broadly ellipsoid, (7.4 –) 7.6 – 8.02 – 8.4 (– 8.9) × (6.5 –) 6.8 – 7.2 – 7.7 (– 8) μm, Q = (1.04 –) 1.07 – 1.11 – 1.15 (– 1.23), ornamentation amyloid, composed of somewhat cylindric (mostly with rounded or obtuse apices) isolated very small (0.3 μm) to high (up to 1.2 μm) spines and most of spines fused laterally and connected by thin to thick ridges (like small crests) forming partial reticulum; suprahilar plage amyloid; apiculi up to 2 μm high. Basidia (35 –) 41 – 49 – 57 (– 68) × 10 – 11 – 12 (– 14) μm, 4 - spored, subclavate to clavate. Subhymenium layer 20 – 25 μm thick, made up of pseudoparenchymatous cells. Hymenial cystidia on lamellar sides (63 –) 71.5 – 86 – 101 (– 134) × (9 –) 9.5 – 11 – 12.5 (– 14) μm, subcylindrical, cylindrical to ventricose with capitate, mucronate, moniliform or appendiculate (up to 7 μm long appendage) apex, emergent up to 55 μm beyond the basidiole tips; contents dense, heteromorphous and partly or completely filled with fibrous to somewhat crystalloid components, hardly staining in sulfovanillin. Lamellae edges fertile with frequent basidia. Hymenial cystidia on lamellar edges (72 –) 75 – 84 – 92.5 (– 98) × (9 –) 9.6 – 10.5 – 11.5 (– 12) μm, subcylindrical, cylindrical to ventricose with obtuse-rounded, capitate, mucronate or appendiculate apex; contents dense, heteromorphous and partly or completely filled with fibrous to somewhat crystalloid components, hardly staining in sulfovanillin. Hymenophoral trama composed of numerous sphaerocytes and connecting hyphae; sphaerocytes globose to elliptical. Pileipellis orthochromatic in Cresyl Blue, sharply delimited from the underlying sphaerocytes of the context, 120 – 180 μm thick, two-layered, distinctly divided in 50 – 80 μm deep suprapellis composed of erect or ascending hyphal terminations, arranged in a densely turf of trichodermal structure and dispersed pileocystidia, and subpellis 70 – 100 μm deep, composed of more or less horizontally irregularly oriented, moderately dense, 2.5 – 3.5 μm wide pilear hyphae. Acid-resistant incrustations absent. Hyphal terminations near the pileus margin usually branched at the subterminal cells or the cells just below, occasionally slightly flexuous, thin-walled; terminal cells (13 –) 15.5 – 24.5 – 34 (– 53) × 3 – 3.5 – 4.5 (– 5) μm, mainly subulate to tapering towards tips or cylindrical to subcylindrical, apically obtuse or slightly narrowed towards tips and wider near base; subterminal cells usually equal in size, rarely with lateral branches or nodulose, equally wide or more or slightly wider. Hyphal terminations near the pileus centre with shorter but slightly wider terminal cells measuring (9 –) 17 – 22.5 – 28.5 (– 35) × (2.5 –) 3 – 4 – 4.5 (– 6) μm, mainly tapering to subulate towards tips or cylindrical or ventricose or occasionally lageniform, apically obtuse or slightly narrowed towards tips and wider near base; subterminal cells equally wide, sometimes slightly wider or ventricose, rarely with nodulose or with lateral branches. Pileocystidia near the pileus margin 1 – 4 - celled, numerous, cylindrical, usually originating deep in subpellis and often originating from branched subterminal cells, thin-walled; terminal cells (30 –) 32.5 – 52 – 71 (– 108) × (4 –) 4.5 – 5 – 6 (– 6.5) μm, cylindrical or sometimes slightly tapered towards tips, rounded-obtuse, without any incrustations; contents heteromorphous-crystalline, without reaction in sulfovanillin. Pileocystidia near the pileus centre with often more septa (1 – 7); terminal cells (14 –) 22.5 – 38 – 54 (– 73) × (4 –) 4.5 – 5 – 6 (– 7) μm, cylindrical or slightly tapered towards tips, rounded-obtuse apex. Clamp connections absent from all tissues.	en	Ghosh, Aniket, Das, Kanad, Buyck, Bart (2021): Two new species in the Russula (Russulaceae, Basidiomycota) crown clade from Indian Himalaya. European Journal of Taxonomy 782: 157-172, DOI: https://doi.org/10.5852/ejt.2021.782.1595, URL: http://dx.doi.org/10.5852/ejt.2021.782.1595
