taxonID	type	description	language	source
03A4A269793961161B11F9728A6D659A.taxon	diagnosis	Diagnosis (after Hutchings et al. 2021 a; most important diagnostic characters highlighted in bold) Transverse prostomium attached to dorsal surface of upper lip; basal part usually as thick horse-shoe shaped crest, eye spots absent; distal part either as another thick crest, with flaring distal lobes, with or without mid-dorsal process, or extending along upper lip until near anterior margin of lip; prostomium Fig. 1. Schematic illustrations of different uncini morphologies, in lateral view (following Nogueira et al. 2010). A. Terebellides sp., SG IX (SMA-BR-Terebellides-KER 1). B. Polycirrus catalanensis Lavesque, Hutchings, Daffe & Londoño-Mesa, 2020, SG XX (holotype MNHN-IA-TYPE 2007). C. Pista sauriaui Lavesque, Daffe, Londoño-Mesa & Hutchings, 2021, SG V (paratype MNHN-IA- TYPE 2036). D. Lomia ramzega Lavesque, Bonifácio, Londoño-Mesa, Le Garrec & Grall, 2017, SG XII (paratype MNHN-IA-TYPE 1791). E. Streblosoma cabiochi Lavesque, Londoño-Mesa, Daffe & Hutchings, 2020, SG VI (holotype MNHN-IA-TYPE 2000). F. Thelepus japonicus Marenzeller, 1884, SG XVII (MNHN-IA-PNT 117). Abbreviations: Af = anterior filament; Ba = base; Ca = capitium; Cext = coma-shape extension; Cr = crest; Db = dorsal button; He = heel; Lh = long handle; Mf = main fang; Oc = occipitium; Pr = prow. frequently extending ventrally, terminating laterally to mouth (Fig. 2 A – D). Buccal tentacles of two types at least, short ones thin, uniformly cylindrical, long tentacles stouter, expanded at tips to variable degrees, distally spatulate (Fig. 2 B, D) or more specialised. Peristomium forming lips; lips expanded, upper lip large, frequently circular and convoluted, folded into three lobes; swollen lower lip, only midventral or cushion-like across ventrum, sometimes extending posteriorly for a few segments (Fig. 2 A – D). Segment I reduced, frequently only visible ventrally, sometimes completely hidden. Segment II distinctly narrower than following segments, constricting body posteriorly to “ lips head ”; SG II usually with rectangular or pentagonal mid-ventral shield at beginning of mid-ventral groove, sometimes extending anteriorly through SG I until near posterior margin of lower lip (Fig. 2 C). Anterior segments highly glandular ventrally, frequently papillose or tessellated, with paired ventro-lateral pads separated from each other within pairs by mid-ventral groove extending from SG II – IV to posterior body (Fig. 2 A – D). Branchiae absent. Notopodia, if present, from SG III (Fig. 2 A – D), extending for variable number of segments, usually few; bilobed, elongate notopodia, post-chaetal lobes sometimes longer, notochaetae originating between lobes along all extension of notopodia, separating lobes from base on ventral side of notopodia (Fig. 2 A – D); notochaetae winged (Fig. 2 E) and / or pinnate, wings of variable width. Neuropodia, if present, located posteriorly to notopodia, frequently from posterior thoracic segments or only on abdomen; neurochaetae as acicular spines or avicular uncini, of two types, and arranged in a single row (Figs 1 C, 2 F – G). Nephridial and genital papillae usually present, at anterior bases of all notopodia, or only at anteriormost notopodia (Fig. 2 A). Pygidium smooth or with rounded ventral papilla.	en	Lavesque, Nicolas, Hutchings, Pat, Londoño-Mesa, Mario H., Nogueira, João M. M., Daffe, Guillemine, Nygren, Arne, Blanchet, Hugues, Bonifácio, Paulo, Broudin, Caroline, Dauvin, Jean-Claude, Droual, Gabin, Gouillieux, Benoit, Grall, Jacques, Guyonnet, Benjamin, Houbin, Céline, Humbert, Suzie, Janson, Anne-Laure, Jourde, Jérôme, Labrune, Céline, Lamarque, Bastien, Latry, Lise, Garrec, Vincent Le, Pelaprat, Corine, Pezy, Jean-Philippe, Sauriau, Pierre-Guy, Montaudouin, Xavier De (2021): The “ Spaghetti Project ”: the final identification guide to European Terebellidae (sensu lato) (Annelida, Terebelliformia). European Journal of Taxonomy 782 (1): 108-156, DOI: https://doi.org/10.5852/ejt.2021.782.1593, URL: http://dx.doi.org/10.5852/ejt.2021.782.1593
03A4A269792D610918B2FEA98C98652D.taxon	diagnosis	Diagnosis (after Nogueira et al. 2018; Hutchings et al. 2021 a, most important diagnostic characters highlighted in bold) Transverse prostomium attached to dorsal surface of upper lip; basal part as thick crest, eyespots frequently present in one pair of dorso-lateral clusters, each with several rows of eyespots (Fig. 3 A); distal part at base of upper lip, frequently with low or erect mid-dorsal tongue-like process, fused to upper lip at variable degrees, with free distal lobe (s), or free from the base. Buccal tentacles of two types, short ones thin, uniformly cylindrical, long tentacles stouter and expanded at tips, slightly spatulate (Figs 3 A – B, F, 4 A). Peristomium forming lips and continuing dorsally at least for short extension, with dorso-lateral nuchal organs at margin with prostomium; lips expanded, upper lip distinctly elongate and narrow, undulated to convoluted; swollen lower lip extending across ventrum, cushion-like or segment-like, frequently deeply grooved (Figs 3 A – B, 4 A). Either SG I or SG II reduced, not forming complete ring in many species. Anterior segments glandular ventrally, smooth, discrete shields absent and frequently with glandular regions poorly developed in comparison to other families of Terebellidae s. l.; mid-ventral groove frequently extending from anterior segments. Two pairs of cirriform branchiae on SG II – III, each pair with simple thin, curled and relatively short filaments progressively tapering to tips (Figs 3 A, 4 A), originating from raised crests on anterior margins of SG II and III, or from specialised, apparently glandular, dorso-lateral cushion-like pads occupying from anterior margins to level of posterior bases of notopodia of those segments. Notopodia beginning from SG II or III, usually SG III, extending for at least 15 segments; notopodia as short cones, notochaetae originating from central core on top, distal lobes absent; notochaetae winged, sometimes with bulbous head and alimbate tips (bayonet-like chaetae), at least in anterior row of anterior thoracic segments. Neuropodia beginning posteriorly to notopodia, usually around SG VIII – XII; neuropodia in conjunction with notopodia as sessile tori, as distinctly low pinnules after notopodia terminate; neurochaetae in single row, as avicular uncini about as long as high, with short triangular heel directed posteriorly, wide and slightly curved base, and dorsal button near midlength of uncini, but closer to anterior margin (Fig. 4 E). Nephridial and genital papillae, if conspicuous, on SG V – VII, posterior to bases of notopodia.	en	Lavesque, Nicolas, Hutchings, Pat, Londoño-Mesa, Mario H., Nogueira, João M. M., Daffe, Guillemine, Nygren, Arne, Blanchet, Hugues, Bonifácio, Paulo, Broudin, Caroline, Dauvin, Jean-Claude, Droual, Gabin, Gouillieux, Benoit, Grall, Jacques, Guyonnet, Benjamin, Houbin, Céline, Humbert, Suzie, Janson, Anne-Laure, Jourde, Jérôme, Labrune, Céline, Lamarque, Bastien, Latry, Lise, Garrec, Vincent Le, Pelaprat, Corine, Pezy, Jean-Philippe, Sauriau, Pierre-Guy, Montaudouin, Xavier De (2021): The “ Spaghetti Project ”: the final identification guide to European Terebellidae (sensu lato) (Annelida, Terebelliformia). European Journal of Taxonomy 782 (1): 108-156, DOI: https://doi.org/10.5852/ejt.2021.782.1593, URL: http://dx.doi.org/10.5852/ejt.2021.782.1593
03A4A269792D610C1B63F9248A6D6064.taxon	diagnosis	Diagnosis (after Hutchings et al. 2021 a, most important diagnostic characters highlighted in bold) Transverse prostomium attached to dorsal surface of upper lip; basal part as thick crest, eyespots frequently present, in short lateral rows, or extending transversely across basal part of prostomium, usually progressively more spaced towards dorsal mid-line, with mid-dorsal gap or not; distal part of base of upper lip short, from nearly indistinct to shelf-like. Buccal tentacles all uniformly thin and cylindrical, to slightly spatulate distally (Figs 3 D, F, 4 B). Peristomium forming lips, sometimes also complete annulation, with dorso-lateral nuchal organs as ciliated grooves; lips expanded, relatively short upper lip, hood-like, about as long as wide; swollen, button-like, mid-ventral lower lip (Figs 3 D, F, 4 B – C). Segment 1 usually present all around, frequently with ventral lobe marginal to mouth (Figs 3 D, F, 4 B – C); SG II typically with anterior margin as protruding crest, at least ventrally (Figs 3 D – E, 4 B – C); lobes on following anterior segments sometimes present. Anterior segments highly glandular ventrally, smooth to highly corrugated between neuropodia within pairs, discrete shields absent (Figs 3 D F, 4 B); mid-ventral groove frequently extending from anterior segments with notopodia. Two to three pairs of branchiae, on SG II – III or II – IV, each pair with simple thin, curled and relatively short filaments progressively tapering to tips (Figs 3 C, E, 4 C), leaving mid-dorsal gap or not between filaments within pairs; branchial filaments originating directly from the body wall or from specialised dorsolateral cushion-like pads. Notopodia beginning on SG II – III, usually extending to mid-body, at least, sometimes until near posterior end; cylindrical to rectangular, distally bilobed notopodia, notochaetae originating between lobes; most taxa with winged notochaetae only, with wings of variable width (Fig. 4 D), distally serrated notochaetae sometimes also present; bayonet-like and pinnate chaetae both absent. Neuropodia beginning posteriorly to notopodia, on SG IV – VI, typically on SG V; neuropodia in conjunction with notopodia as fleshy, swollen ridges, as raised rectangular to cylindrical pinnules after notopodia terminate; neurochaetae as avicular uncini frequently longer than high, with short triangular heel directed posteriorly, distinctly curved and wide base, and dorsal button near anterior margin of uncini, or within anterior third of distance between anterior margin of uncini and base of main fang (Fig. 4 F). Nephridial and genital papillae usually present, on SG IV – VII, posterior to bases of notopodia or between parapodial lobes (Fig. 3 C).	en	Lavesque, Nicolas, Hutchings, Pat, Londoño-Mesa, Mario H., Nogueira, João M. M., Daffe, Guillemine, Nygren, Arne, Blanchet, Hugues, Bonifácio, Paulo, Broudin, Caroline, Dauvin, Jean-Claude, Droual, Gabin, Gouillieux, Benoit, Grall, Jacques, Guyonnet, Benjamin, Houbin, Céline, Humbert, Suzie, Janson, Anne-Laure, Jourde, Jérôme, Labrune, Céline, Lamarque, Bastien, Latry, Lise, Garrec, Vincent Le, Pelaprat, Corine, Pezy, Jean-Philippe, Sauriau, Pierre-Guy, Montaudouin, Xavier De (2021): The “ Spaghetti Project ”: the final identification guide to European Terebellidae (sensu lato) (Annelida, Terebelliformia). European Journal of Taxonomy 782 (1): 108-156, DOI: https://doi.org/10.5852/ejt.2021.782.1593, URL: http://dx.doi.org/10.5852/ejt.2021.782.1593
03A4A2697928610518C6FCE68C1A660A.taxon	diagnosis	Diagnosis (after Hutchings et al. 2021 a, most important diagnostic characters highlighted in bold) Transverse prostomium attached to dorsal surface of upper lip; basal part as thick crest, eyespots frequently present (Fig. 5 B), in short rows at each lateral sides, or extending transversely across basal part of prostomium. Buccal tentacles all usually uniformly cylindrical. Peristomium usually forming lips only; lips expanded, relatively short upper lip, hood-like, about as long as wide; swollen, usually button-like and mid-ventral lower lip. Segment I terminating laterally to ventro-laterally, partially fused to expanded lower lip, or developed, forming lobes of variable extension and position. Lobes on anterior segments frequently present, of variable length, sometimes extending to SGV – VII (Figs 5 B – D, 6 A – D). Anterior segments highly glandular ventrally, with discrete, smooth to corrugated, rectangular to trapezoidal mid-ventral shields extending from anterior segments until termination of notopodia, or near it; mid-ventral groove extending from termination of mid-ventral shields. Two to three pairs of branchiae usually present (Figs 5 A – D, 6 A – D), but three genera have a single pair and several are abranchiate; branchial filaments originating all together from single point on body wall, on either side of branchiferous segments, unbranched, or, more frequently, originating from conspicuous main stalk on either side of pair, branching from one to several levels, in plumose (spiraled), dichotomous, pectinate or arborescent arrangement. Notopodia beginning on SGII – V, SGIV in most genera, usually extending to mid-body, around SGXX, but sometimes present on fewer segments or extending more posteriorly for variable extension, rarely until near posterior end; first pairs of notopodia inserted dorso-laterally, progressively more laterally, then vertically aligned; cylindrical to rectangular notopodia. Notochaetae originating from central core on top, distal lobes absent; notochaetae distally winged, wings of variable length and width, or serrate, sometimes with wings at midlength, basally to a serrated blade; some more specialised types of notochaetae may be present (Fig. 5 E – G). Neuropodia beginning posteriorly to notopodia, on SGV – IX, usually on SGV; neuropodia in conjunction with notopodia as low, sessile ridges, sometimes continuing posteriorly until pygidium, but most taxa with rectangular to cylindrical or foliaceous neuropodial pinnules after notopodia terminate; neurochaetae as avicular uncini usually as long as high, with short triangular heel directed posteriorly, slightly curved and wide base, and dorsal button (Figs 1 C – D, 6 E – F); uncini arranged in double rows (Fig. 6 E) from around SGXI usually until termination of notopodia, but several genera with double rows.	en	Lavesque, Nicolas, Hutchings, Pat, Londoño-Mesa, Mario H., Nogueira, João M. M., Daffe, Guillemine, Nygren, Arne, Blanchet, Hugues, Bonifácio, Paulo, Broudin, Caroline, Dauvin, Jean-Claude, Droual, Gabin, Gouillieux, Benoit, Grall, Jacques, Guyonnet, Benjamin, Houbin, Céline, Humbert, Suzie, Janson, Anne-Laure, Jourde, Jérôme, Labrune, Céline, Lamarque, Bastien, Latry, Lise, Garrec, Vincent Le, Pelaprat, Corine, Pezy, Jean-Philippe, Sauriau, Pierre-Guy, Montaudouin, Xavier De (2021): The “ Spaghetti Project ”: the final identification guide to European Terebellidae (sensu lato) (Annelida, Terebelliformia). European Journal of Taxonomy 782 (1): 108-156, DOI: https://doi.org/10.5852/ejt.2021.782.1593, URL: http://dx.doi.org/10.5852/ejt.2021.782.1593
03A4A269792161381B3CFA388A6C6409.taxon	diagnosis	Diagnosis (after Hutchings et al. 2021 a, most important diagnostic characters highlighted in bold) Transverse prostomium attached to dorsal surface of upper lip; basal part as thick crest, eyespots sometimes present; distal part at base of upper lip or extending along lip. Buccal tentacles of two types, uniformly cylindrical and expanded at tips, spatulate. Peristomium forming lips, sometimes also a ventral lobe, as an extension of the lower lip; lips expanded, circular upper lip, distal margin folded or convoluted; lower lip button-like, usually continuing by ventral lobe, or expanded, forming large scoop-shaped process (Figs 7 A – C, 8 A, C – D). Segment I usually short, frequently only visible ventrally; anterior margin of anterior segments with lobes as low, even-length collars covering posterior margins of preceding segments, at least ventrally; ventro-lateral or lateral lobes on anterior segments sometimes present. Anterior segments poorly glandular ventrally, smooth, discrete shields absent; midventral groove extending from posterior segments with notopodia. Two to four pairs of branchiae, beginning from SGII, each pair with single, thick and elongate, tapered or foliaceous filament, or two pairs fused in single four lobed structure originating mid-dorsally between SGII – III or II – IV (Figs 7 C, 8 C – D). Notopodia beginning from SGIII – VI, typically terminating at SGXX; short, conical notopodia, chaetae emerging from central core on top, distal lobes absent; narrowly-winged notochaetae in both rows throughout. Neuropodia beginning on same segment as notopodia or slightly posteriorly, rarely beginning before notopodia; sessile neuropodia until termination of notopodia, neurochaetae emerging directly from body wall, as rectangular to foliaceous pinnules after termination of notopodia; thoracic neurochaetae as acicular uncini (Figs 1 A, 7 D, 8 F), sometimes with small hood or beard below main fang; avicular abdominal uncini, with secondary teeth in rows on top and laterally to main fang. Nephridial papillae on SGIII usually present, other papillae sometimes present on SGVI and SGVII, but reduced to inconspicuous in most taxa. Pygidium smooth to slightly crenulate, sometimes bilobed.	en	Lavesque, Nicolas, Hutchings, Pat, Londoño-Mesa, Mario H., Nogueira, João M. M., Daffe, Guillemine, Nygren, Arne, Blanchet, Hugues, Bonifácio, Paulo, Broudin, Caroline, Dauvin, Jean-Claude, Droual, Gabin, Gouillieux, Benoit, Grall, Jacques, Guyonnet, Benjamin, Houbin, Céline, Humbert, Suzie, Janson, Anne-Laure, Jourde, Jérôme, Labrune, Céline, Lamarque, Bastien, Latry, Lise, Garrec, Vincent Le, Pelaprat, Corine, Pezy, Jean-Philippe, Sauriau, Pierre-Guy, Montaudouin, Xavier De (2021): The “ Spaghetti Project ”: the final identification guide to European Terebellidae (sensu lato) (Annelida, Terebelliformia). European Journal of Taxonomy 782 (1): 108-156, DOI: https://doi.org/10.5852/ejt.2021.782.1593, URL: http://dx.doi.org/10.5852/ejt.2021.782.1593
