identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
397C5D1DFFC8FFDCFF6FB545FEEDFBA0.text	397C5D1DFFC8FFDCFF6FB545FEEDFBA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum ahrensi Shavrin 2021	<div><p>Amphichroum ahrensi Shavrin, sp. n.</p> <p>(Figs. 1, 8–9, 15)</p> <p>Type material examined: Holotype, ♂: ‘ INDIA, West Bengal | Darjeeling, Tonglu | 19.-23.V.1998 | 2600-3000m NN | leg. Fabrizi &amp; Ahrens’ &lt;printed&gt;, ‘HOLOTYPE | Amphichroum | ahrensi sp. n. | Shavrin A.V. des. 2021’ &lt;red, printed&gt; (NME).</p> <p>Paratypes: 7 ♂♂ (two specimens dissected), 11 ♀♀: same data as the holotype. (5 ♂♂, 7 ♀♀: NME; 2 ♂♂, 4 ♀♀: cSh). All paratypes with red label ‘ PARATYPE | Amphichroum | ahrensi sp. n. | Shavrin A.V. des. 2021’.</p> <p>Description. Measurements (n=19): HL: 0.30–0.43; HW: 0.55–0.70; AL (holotype): 1.54; OL: 0.19–0.25; PL: 0.53–0.72; PW: 0.88–1.00; ESL: 1.05–1.25; EW: 1.18–1.32; AW: 1.16–1.30; MTbL (holotype): 0.57; MTrL (holotype): 0.52 (MTrL 1–4: 0.30; MTrL 5: 0.22); AedL: 0.70–0.75; TL: 2.65–3.75 (holotype: 3.70).</p> <p>Habitus as in Fig. 1. Head, mouthparts, antennomeres 4–11, disc of pronotum, scutellum, legs and abdomen yellow-brown; antennomeres 1–3, lateral and basal portion of pronotum, elytra and paratergites yellow. Forebody glossy; frontal portion of head with dense transverse microsculpture, middle portion of head without or with very fine transverse microreticulation, infraorbital ridges with fine, rounded or longitudinal meshes; neck with dense transverse microsculpture; pronotum with distinct isodiametric microreticulation, without meshes in mediobasal third; exposed part of scutellum with dense, transverse sculpture; abdominal tergites with very dense, fine, isodiametric microreticulation. Head with very fine and sparse punctation, slightly denser in middle portion and with several moderately large punctures between anteocellar foveae and eyes; punctation of pronotum distinctly larger, denser and deeper than that in middle portion of head, sometimes finer and sparser in medioapical portion or without punctures in middle and/or mediobasal portions; punctation of elytra large, distinctly denser, coarser and deeper in basal portion, finer and sparser in middle along suture; abdominal tergites with dense, fine punctures, coarser and denser on abdominal tergite VII. Forebody without setation except for head with several long setae on frontal portion and two to four moderately long and erect setae between anteocellar foveae and eyes, and lateral margins of pronotum with four moderately long setae; abdominal tergites with regular, short, moderately dense setation.</p> <p>Head subtrapezoidal, transverse, 1.6–1.8 times as broad as long, flattened in middle and slightly convex on posterior portions of infraorbital ridges; anteocellar foveae narrow, deep and long, diagonally stretching toward antennal bases about to level of anterior margins of eyes. Eyes very large, convex. Ocelli large, located at level of posterior margins of eyes near deep occipital furrow, distance between ocelli as long as or slightly shorter than distance between ocellus and posterior margin of eye. Apical maxillary palpomere 1.3 times as long as preceding segment, gradually narrowed from basal third toward subacute apex. Antenna moderately long, reaching basal part of elytra when reflexed, from antennomere 5 progressively widened apicad; basal antennomere oblong, about three times as long as broad, antennomere 2 distinctly shorter than and about as broad as basal antennomere, 3 slightly longer than 2, 4 slightly shorter than 3, 5–6 slightly broader than 4, 7–8 slightly broader than 6, 9–10 slightly broader than 8, apical antennomere about 1.3 times as long as 10, sharply narrowed from apical third toward subacute apex.</p> <p>Pronotum 1.3–1.6 times as broad as long, 1.4–1.6 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; anterior angles widely rounded, slightly protruded anteriad; posterior angles widely rounded; laterobasal margins slightly sinuate in some specimens; lateral portions flattened, moderately wide, slightly explanate, explanate area sometimes gradually widened posteriad.</p> <p>Elytra slightly broader than long and slightly widened apicad, reaching apical margin of abdominal tergite IV or V, slightly less than twice as long as pronotum, with widely rounded or straight apical margins. Hind wings fully developed.</p> <p>Legs moderately long; metatibia slightly longer than metatarsus, with long strong thorns on inner and long setation on outer surfaces.</p> <p>Abdomen slightly narrower than elytra, with a pair of very large transverse tomentose spots (wing-folding patches) in middle of tergite IV, with narrow palisade fringe on apical margin of tergite VII.</p> <p>Male. Protarsomeres 1–4 very wide. Medial margin of inner side of protibia with two parallel rows of eight to twelve short peg setae; mesotibia strongly curved mediad, with dense row of numerous very short and wide thorns beginning from medial side of bend and stretching to apex of mesotibia; lateral margins of mesotibia with very long and strong, sparse thorns and apical margins with several strong and long thorns. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII widely rounded. Aedeagus (Fig. 8) with median lobe widest at basal bulb, gradually narrowed apicad; parameres moderately narrow, distinctly exceeding apex of median lobe, each with two moderately short preapical setae; internal sac long, with two fields of long thorns in apical and two complicated sclerotized structures in basal portions. Lateral aspect of aedeagus as in Fig. 9.</p> <p>Female. Protarsomeres 1–4 narrow. Medial margin of protibia without peg setae; mesotibia without modifications, slender, with long strong thorn along entire length of lateral margin. Abdominal tergite VIII and sternite VIII with small rounded or straight apical margin.</p> <p>Comparative notes. Amphichroum ahrensi sp. n. differs from the remaining species of the Himalayan Region by the absence of pubescence on the disc of pronotum and elytra. Based on the shape of the median lobe, it somewhat similar to the Chinese A. maculosum Shavrin &amp; Smetana, 2018 and A. subelongatum Shavrin &amp; Smetana, 2018, but differs from them by the paler coloration of the body and completely different morphology of the internal sac.</p> <p>Distribution. The new species is at present known only from the type locality (Fig. 15) in West Bengal, India.</p> <p>Bionomics. Specimens were collected at elevations from 2600 to 3000 m a.s.l.</p> <p>Etymology. Patronymic, the species is named to honor Dirk Ahrens (Bonn, Germany), one of the collectors of the type specimens.</p></div> 	http://treatment.plazi.org/id/397C5D1DFFC8FFDCFF6FB545FEEDFBA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFCAFFDBFF6FB5FDFC0CFCC8.text	397C5D1DFFCAFFDBFF6FB5FDFC0CFCC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum altivagans Cameron 1941	<div><p>Amphichroum altivagans Cameron, 1941</p> <p>(Figs. 2, 10–11, 15)</p> <p>Amphichroum altivagans Cameron, 1941: 143</p> <p>Type material examined: Holotype by monotypy (“Type in my collection”), ♂ (Fig. 2): ‘Type’ &lt;rounded label with red margin, printed&gt;, ‘Kashmir | Khelanmarg | 10,000 ft. | vi-vii-1931 | Dr.Cameron’ &lt;printed&gt;, ‘A. | altivagans | TYPE [handwritten in red] Cam’ &lt;handwritten in black&gt;, ‘M.Cameron. | Bequest. | B.M.1955-147’. &lt;printed&gt;, ‘ Amphichroum | altivagans Cameron, 1941 | Shavrin A.V. det. 2021’ &lt;printed&gt; (BMNH).</p> <p>Additional material studied: 2 ♂♂, 2 ♀♀: ‘ Kashmir Khelanmarg 10,000 ft. vi-vii-1931 Dr.Cameron’, ‘ M.Cameron. Bequest. B.M.1955-147.’ (BMNH).</p> <p>Redescription. Measurements (n=5): HL: 0.30–0.32; HW: 0.50–0.55; AL (holotype): 0.93; OL: 0.17–0.19; PL: 0.48–0.52; PW: 0.71–0.81; ESL: 1.00–1.07; EW: 1.00–1.05; AW: 0.87–0.97; MTbL (holotype): 0.57; MTrL (holotype): 0.41 (MTrL 1–4: 0.21; MTrL 5: 0.20); AedL: 0.60; TL: 2.42–3.01 (holotype: 2.55).</p> <p>Habitus as in Fig. 2. Forebody, mouthparts, antennomeres 1–3 and legs yellow (middle portion of head and neck slightly darker); abdomen yellow to dark-brown; two apical maxillary palpomeres and antennomeres 4–11 brown. Head with distinct, dense microsculpture, transverse on clypeus and isodiametric in middle portion and neck; pronotum with distinct isodiametric meshes, finer than in middle portion of head, without microreticulation in middle and/or mediobasal third; abdominal tergites with dense transverse microreticulation. Middle portion of head with fine and sparse punctation; punctation of pronotum dense, moderately large and deep, finer in mediobasal portion, sparser and finer in middle and/or mediobasal third (some specimens without punctures in mediobasal portion); punctation of elytra about as that on pronotum, but deeper, coarser in basal portion and sometimes finer in parascutellar portion and along suture; abdominal tergites without or with very fine and sparse punctation. Frontal portion of head with several long setae; pronotum and elytra covered by dense, long, semierect setation.</p> <p>Head 1.6–1.7 times as broad as long, with basal part of clypeus slightly impressed; anteocellar foveae deep and long, reaching level of anterior margins of eyes. Distance between ocelli about one and a half times distance between ocellus and posterior margin of eyes. Antennomere 3 slightly narrower and about as long as 2, 4 slightly broader and shorter than 3, 5–6 distinctly broader than 4, 7–9 slightly shorter than 6, 10 slightly broader than 9, about as long as previous antennomere.</p> <p>Pronotum 1.4–1.5 times as broad as long, 1.4 times as broad as head, widest in middle, slightly more narrowed anteriad than posteriad.</p> <p>Elytra about as long as broad, twice as long as pronotum.</p> <p>Male. Apical margin of abdominal tergite VIII more or less straight. Aedeagus (Fig. 10) with wide basal portion, significantly narrowed apicad toward subacute apex; parameres narrow, slightly exceeding apex of median lobe, with one pair of short preapical seta; internal sac very long, with several large sclerotized teeth in preapical and middle portions. Lateral aspect of aedeagus as in Fig. 11.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Regarding the general shape of the narrow and elongate apical portion of the median lobe and the length of the narrow parameres, A. altivagans is similar to A. anthobioides and A. telnovi sp. n. From A. anthobioides it differs by the paler pronotum and elytra, and from A. telnovi sp. n. by the smaller body and longer elytra. It can be distinguished from both species by the shape of pronotum, from middle slightly more narrowed anteriad than posteriad, and by details of the structure of the internal sac.</p> <p>Distribution. The new species is at present known only from the type locality (Fig. 15) in Kashmir, India.</p> <p>Bionomics. Specimens were collected at elevation 3048 m a.s.l.</p></div> 	http://treatment.plazi.org/id/397C5D1DFFCAFFDBFF6FB5FDFC0CFCC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFCDFFD8FF6FB2D5FBAAFBCC.text	397C5D1DFFCDFFD8FF6FB2D5FBAAFBCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum anthobioides Champion 1925	<div><p>Amphichroum anthobioides Champion, 1925</p> <p>(Figs. 3, 12–15)</p> <p>Amphichroum anthobioides Champion, 1925: 104; Cameron 1930: 158</p> <p>Type material examined: Lectotype (here designated) of Amphichroum anthobioides Champion, 1925, ♂ (Fig. 3; left specimen glued on the rectangular card with the paralectotype; under it I put a black dot): ‘ ♂ ’ &lt;printed&gt;, ‘Type | H.T.’ &lt;round label with red margin, printed&gt;, ‘Burphu, | Gori V., 11500ft. | India. H.G.C.’ &lt;printed&gt;, Amphichroum | anthobioides | 1925 Ch. types’ &lt;handwritten in black Indian ink&gt;, ‘ Amphichroum | anthobioides, | Champ.’ &lt;printed&gt;, ‘E.M.M. 1925. | det. G.C.C.’ &lt;printed&gt;, ‘G.C.Champion. | Brit.Mus. | 1925–42.’ &lt;printed&gt;, ‘On umbels | (..[unreadable]...)’ &lt;handwritten&gt;, ‘ LECTOTYPE | Amphichroum | anthobioides Champion, 1925 | Shavrin A.V. des. 2021’ &lt;red, printed&gt; (BMNH).</p> <p>Paralectotypes of A. anthobioides Champion, 1925: 1 ♂ (specimen is glued on the same card with the lectotype): same labels as the lectotype (BMNH); 4 ♂♂ (one specimen dissected): ‘ ♂ ’ &lt;printed&gt;, ‘SYN- | TYPE’ &lt;round label with blue margin, printed&gt;, ‘Burphu, | Gori V., 11500ft. | India. H.G.C.’ &lt;printed&gt;, ‘ Amphichroum | anthobioides, | Champ.’ &lt;printed&gt;, ‘H.G.Champion Coll. | B.M. 1953-156.’ &lt;printed&gt;, ‘ Amphichroum | anthobioides Champion, 1925 | Shavrin A.V. des. 2021’ &lt;printed&gt; (BMNH).</p> <p>Additional material studied: 2 ♂♂, 5 ♀♀: ‘ NEPAL Prov. Seti Distr. Bajura 19km W Simikot, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=81.64444&amp;materialsCitation.latitude=29.887222" title="Search Plazi for locations around (long 81.64444/lat 29.887222)">Kuwadi Khola</a>, 3500 m 29°53’14’’N, 81°38’40’’E 04- 05.07.2001. leg. A. Kopetz riverbank, coniferous-birch forest’ (1 ♂, 3 ♀♀: NME; 1 ♂, 2 ♀♀: cSh).</p> <p>Redescription. Measurements (n=13): HL: 0.30–0.33; HW: 0.47–0.50; AL (lectotype): 0.98; OL: 0.12; PL: 0.47–0.51; PW: 0.67–0.77; ESL: 0.82–1.00; EW: 0.98–1.07; AW: 0.76–0.95; MTbL (lectotype): 0.55; MTrL (lectotype): 0.44 (MTrL 1–4: 0.22; MTrL 5: 0.20); AedL: 0.50; TL: 1.96 (lectotype)–2.54.</p> <p>Habitus as in Fig. 3. Head, disc of pronotum and elytra yellow to yellow-brown; two apical maxillary palpomeres, antennomeres 4–11 and abdomen brown to reddish-brown; antennomeres 1–3, lateral and sometimes basal portion of pronotum, legs and apical segments of abdomen yellow. Frontal portion of head with fine transverse microsculpture, middle portion of head with variable, fine and very indistinct to distinct, transverse microreticulation, slightly coarser on infraorbital ridges; neck with dense and transverse meshes; pronotum without visible sculpture; scutellum without or with very fine transverse microreticulation; abdominal tergites with dense, isodiametric microsculpture. Head with sparse and irregular punctation, fine or moderately large, denser and deeper in middle portion between posterior portions of eyes; punctation of pronotum dense, moderately large and deep, finer in middle portion and sometimes missing in mediobasal third; scutellum without or with several fine punctures in middle; punctation of elytra dense, significantly larger, coarser and deeper than that on pronotum, finer in parascutellar portion and sometimes finer along suture; abdominal tergites without or with very fine and sparse punctation. Forebody with moderately dense and long setation, distinctly denser on pronotum and elytra.</p> <p>Head 1.5 times as broad as long, flattened or slightly elevated in middle; anteocellar foveae deep, long and almost reaching level of apical third of eyes. Distance between ocelli distinctly longer than distance between ocellus and posterior margin of eye. Apical maxillary palpomere about as long as or slightly longer than preapical segment. Antenna reaching basal third of elytra when reflexed; antennomere 3 distinctly narrower and about as long as 2, 4 shorter and slightly broader than 3, 5–6 slightly broader and longer than 4, 7–8 slightly broader than 6, 9–10 slightly shorter and broader than 8, apical antennomere about 1.3–1.5 times as long as 10, from about middle gradually narrowed toward rounded or subacute apex.</p> <p>Pronotum 1.4–1.5 times as broad as long, 1.4–1.5 times as broad as head, from widest middle evenly narrowed both anteriad and posteriad.</p> <p>Male. Apical margins of abdominal tergite VIII and sternite VIII more or less straight. Median lobe moderately wide (Fig. 12), sharply narrowed from preapical part toward subacute apex; parameres slightly exceeding apex of median lobe, with moderately short preapical and apical setae (Figs. 12–13); internal sac long and narrow, with six moderately large sclerotized structures in basal portion. Lateral aspect of aedeagus as in Fig. 14.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Regarding the general shape of the median lobe and parameres, A. anthobioides is similar to A. altivagans and A. telnovi sp. n. From A. altivagans it differs by the shape of pronotum, evenly narrowed middle both anteriad and posteriad, and from A. telnovi sp. n. by the slightly shorter elytra. From both species it can be distinguished by somewhat smaller body and details of the structure of the internal sac.</p> <p>Distribution. The species is known from three locations in India (Uttarakhand) and Nepal (Fig. 15).</p> <p>Bionomics. Specimens were collected at elevation 3500 m a.s.l. (type specimens were collected about from the same altitude (“ 11,000 ft. ”). According to Champion (1925), type specimens were found “…in thistle heads”. The specimens from Nepal were collected along a riverbank in coniferous-birch forest.</p> </div>	http://treatment.plazi.org/id/397C5D1DFFCDFFD8FF6FB2D5FBAAFBCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFCEFFD7FF6FB5D1FDDFFB50.text	397C5D1DFFCEFFD7FF6FB5D1FDDFFB50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum monticola Cameron 1928	<div><p>Amphichroum monticola Cameron, 1928</p> <p>(Figs. 4, 16, 18–20)</p> <p>Amphichroum monticola Cameron, 1928: 558; Cameron 1930: 157</p> <p>&lt;?&gt; Amphichroum monticola: Coiffait 1978: 114</p> <p>Type material examined: Holotype, ♂ (Fig. 4; dissected): ‘Type’ &lt;round label with red margins&gt;, ‘ Tibet: [underlined in yellow] Tropde. 11.000ft. 23.vi.1924. Maj.R.W.G.Hingston’ &lt;printed&gt;, ‘Everest Exp. | Brit.Mus. | 1924– 386.’ &lt;printed&gt;, ‘ Amphichroum alticola [sic!] Cam TYPE [handwritten by red]’ &lt;handwritten in black&gt;, ‘ Amphichroum | monticola Cameron, 1928 | teste Shavrin A.V. det. 2021’ &lt;printed&gt; (BMNH).</p> <p>Additional material studied: CHINA: 1 ♀: ‘ Tibet: Tropde. 11,000ft. 23.vi.1924. Maj.R.W.G.Hingston’, ‘ Everest Exp. Brit.Mus. 1924-386.’, ‘ A. monticola Cam’, ‘ M.Cameron Bequest. B.M.1955-147.’ (BMNH); NEPAL: 1 ♂, 1 ♀: ‘NEPAL | Deorali – Bhandar env. 7.6.– 12.6.2012 lgt. E. Kučera’ (1 ♂: cSch; 1 ♀: cSh); 1 ♂: ‘NEPAL, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.48333&amp;materialsCitation.latitude=28.366667" title="Search Plazi for locations around (long 84.48333/lat 28.366667)">Manaslu Mts.</a> 28°22’N 84°29’E E slope of Ngali Khola Vall. 2000-2300 m, leg. Schmidt 15.V.2005 ’ (NME); 13 ♂♂, 5 ♀♀: ‘ Nepal Manaslu Mts. E-slope, E Gupchi 2500-2700 m Danda, 22.V.2006 leg. J. Schmidt’ (9 ♂♂, 3 ♀♀: NME; 4 ♂♂, 2 ♀♀: cSh); 2 ♂♂: ‘NEPAL Manaslu Mts. E-slope E Gupchi <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.81917&amp;materialsCitation.latitude=28.185833" title="Search Plazi for locations around (long 84.81917/lat 28.185833)">Danda</a> 2500-2700m 28°11’09N 84°49’09E 22.5.2006 leg. J. Schmidt’ (1 ♂: NME; 1 ♂: cSh); 1 ♂, 2 ♀♀: ‘NEPAL, Prov. Janakpur distr. Dolakha, upp. Simigau village, 2700-2800 m NN, 01.VI.2000 leg. J. Schmidt’ (1 ♀: NME; 1 ♂, 1 ♀: cSh).</p> <p>Redescription. Measurements (n=25): HL: 0.29–0.37; HW: 0.52–0.58; AL (holotype): 1.25; OL: 0.13–0.17; PL: 0.54–0.62; PW: 0.80–0.88; ESL: 0.86–1.02; EW: 0.92–1.23; AW: 0.88–1.25; MTbL (holotype): 0.52; MTrL (holotype): 0.48 (MTrL 1–4: 0.26; MTrL 5: 0.22); AedL: 0.62–0.68; TL: 2.65–4.00 (holotype: 3.00).</p> <p>Habitus as in Fig. 4. Head and pronotum yellow-brown to reddish-brown; elytra yellowish to brown; abdomen and antennomeres 4–11 brown to reddish-brown (some specimens with paler abdomen); mouthparts, antennomeres 1–3, legs and sometimes paratergites yellow. Clypeus with dense transverse microsculpture, middle portion of head with fine transverse and infraorbital portions with dense isodiametric or transverse meshes; pronotum without or with very fine, isodiametric microreticulation in medioapical and/or middle portion; scutellum with fine, transverse sculpture; abdomen with dense, transverse microsculpture on tergites IV–VI and dense isodiametric meshes on tergites VII–VIII. Head with fine and irregular punctation, denser in middle portion and sometimes slightly deeper between anteocellar foveae and eyes; punctation of pronotum dense, moderately large and deep, sometimes finer in middle and/or mediobasal portions or without punctures in mediobasal third; punctation of elytra dense, distinctly larger and deeper than that on pronotum; abdominal tergites with very indistinct, fine, irregular punctation. Forebody with moderately dense pubescence.</p> <p>Head 1.5–1.7 times as broad as long. Distance between ocelli about one and a half times to twice as long as distance between ocellus and posterior margin of eyes. Antennomere 2 about as long as and broad as basal antennomere, 3 slightly shorter than 2, 4 distinctly shorter and slightly broader than 3, 5–7 disitinctly longer and broader than 4, 8–10 slightly broader and sometimes longer than 7.</p> <p>Pronotum 1.4 times as broad as long, 1.4–1.5 times as broad as head, from the widest middle narrowed evenly anteriad and posteriad.</p> <p>Elytra moderately short, 1.5–1.6 times as long as pronotum, slightly widened posteriad or somewhat parallelsided.</p> <p>Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded. Median lobe moderately narrow (Fig. 18–19), abruptly narrowed in preapical portion toward narrowly rounded (Fig. 18) or subacute (Fig. 19) apex; parameres slightly broadened apically, exceeding apex of median lobe, with two preapical setae; internal sac long, with oval ring-shaped structure in basal portion. Lateral aspect of aedeagus as in Fig. 20.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Based on the coloration, length and proportions of the body, and the general shape of the aedeagus, A. monticola is similar to A. nepalicum and A. pindarense, from which it can be distinguished by the denser punctation of the pronotum, and details of the structure of the internal sac.</p> <p>Distribution. The species is known from China (Tibet) and Nepal (Fig. 16).</p> <p>Bionomics. Specimens were collected at elevations from 2000 to 3352 m a.s.l.</p> <p>Remarks. Coiffait (1978) recorded A. monticola from Bhutan and provided figures of the aedeagus, with narrow apex of the median lobe (Coiffait, 1978: figures 4E, 4F), but these specimens requires re-examination. It is here recorded from Nepal for the first time.</p> </div>	http://treatment.plazi.org/id/397C5D1DFFCEFFD7FF6FB5D1FDDFFB50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFC1FFD6FF6FB46DFCD8FB50.text	397C5D1DFFC1FFD6FF6FB46DFCD8FB50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum nepalicum (Coiffait 1982) Shavrin 2021	<div><p>Amphichroum nepalicum (Coiffait, 1982) comb. nov.</p> <p>(Figs. 5, 17, 21–22)</p> <p>Arpedium nepalicum Coiffait, 1982: 145</p> <p>Type material examined: Holotype of Arpedium nepalicum Coiffait, 1982, ♀: ‘Dzunda Khola-Tal | b. Talphi 3000- 3500 m’ &lt;printed&gt;, ‘Gebiet von Jumla | Westnepal, lg.H.Franz’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘ Arpedium | nepalicum | H. COIFFAIT det.19[printed]79’ &lt;handwritten&gt;, ‘ Amphichroum [handwritten] | sp. [handwritten] | det. M. K. Thayer 198 9[handwritten]’ &lt;printed&gt;, ‘ Amphichroum | nepalicum Coiffait, 1982 | Shavrin A.V. det. 2015’ &lt;printed&gt; (NHMW).</p> <p>Additional material studied: 4 ♂♂, 5 ♀♀: ‘ NEPAL, W-Manaslu Himal Ngali Khola-Gebiet unterh Bhara Pokh. Lekh. 2200mNN N 28°22’34’’ E 84°29’52’’ 15./ 16.V.2005, leg. O. Jäger’ (2 ♂♂, 3 ♀♀: SNSD; 2 ♂♂, 2 ♀♀: cSh); 2 ♂♂, 2 ♀♀: ‘W Nepal, NW Pokhara, Modi Khola, 1600-2000 m. Landrung – Gandrung. 8.V.1984. C. Holzschuh leg.’ (1 ♂, 2 ♀♀: cZan; 1 ♂: cSh); 12 ♂♂, 10 ♀♀: ‘ NEPAL <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.48333&amp;materialsCitation.latitude=28.366667" title="Search Plazi for locations around (long 84.48333/lat 28.366667)">Manaslu Mts.</a> 28°22’N 84°29’E E slope of N gali Khola Vall. 2000-2300 m. leg. Schmidt 15. V. 2005 ’ (3 ♂♂, 2 ♀♀: cSh; 9 ♂♂, 8 ♀♀: NME); 5 ♀♀: ‘ Nepal Manaslu Mts. | E-slope, E Gupchi | 2500-2700 m | Danda, 22.V.2006 | leg. J. Schmidt’ (2 ♀♀: cSh; 3 ♀♀: NME); 1 ♀: ‘ NEPAL, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.910835&amp;materialsCitation.latitude=27.472778" title="Search Plazi for locations around (long 87.910835/lat 27.472778)">Prov.</a> Mechi | 27°28’22’’N, 087°54’39’’E | Omje Khola bis | Yamphudin; 17.IV.2003 | 1700-2550 m NN | leg.: J. Weipert’ (cSh).</p> <p>Redescription. Measurements (n=30): HL: 0.29–0.41; HW: 0.48–0.61; AL (averaged): 1.28; OL: 0.15–0.17; PL: 0.54–0.66; PW: 0.73–0.91; ESL: 0.88–1.08; EW: 0.96–1.29; AW: 0.88–1.36;MTbL (averaged): 0.58; MTrL (averaged): 0.42 (MTrL 1–4: 0.20; MTrL 5: 0.22); AedL: 0.61–0.74; TL: 2.50 (holotype)–3.85.</p> <p>Habitus as in Fig. 5. Head and disc of pronotum yellow-brown; antennomeres 4–11 and abdomen reddishbrown; mouthparts, antennomeres 1–3, lateral portions of pronotum, elytra, legs and sometimes paratergites yellow. Head with dense transverse microsculpture, sparser and finer in middle portion; pronotum with very fine and sometimes indistinct transverse microreticulation, usually without meshes in middle and/or mediobasal third. Head with moderately fine, irregular punctation, denser in middle portion; punctation of pronotum about as that in middle portion of head, but slightly larger and deeper, sparser in middle and mediobasal portion; punctation of elytra larger and deeeper than that on pronotum, finer in parascutellar portion and along suture; punctation of abdomen invisible or very fine and sparse. Forebody with dense and short pubescence.</p> <p>Head 1.4–1.6 times as broad as long, with deep and long anteocellar foveae, almost reaching level of anterior margins of eyes. Distance between ocelli about one and a half times distance between ocellus and posterior margin of eyes. Antennomere 3 slightly narrower than 2, 4 distinctly shorter and broader than 3, 5–6 slightly broader than 4, 7–10 slightly broader than 6.</p> <p>Pronotum 1.3 times as broad as long, 1.4–1.5 times as broad as head, evenly narrowed both anteriad and posteriad.</p> <p>Elytra 1.6 times as long as pronotum.</p> <p>Male. Apical margin of abdominal tergite VIII and sternite VIII rounded. Aedeagus (Figs. 21) with wide basal bulb of median lobe, narrowed toward middle; median lobe from preapical portion sharply narrowed toward subacute apex; parameres narrow, widened apically, significantly exceeding apex of median lobe, with two pairs of short apical and preapical setae; internal sac very long, with lateral fields of small and oval sclerites in middle and apical portions. Lateral aspect of aedeagus as in Fig. 22.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Based on the coloration, length and general proportions of the body and aedeagus, A. nepalicum is similar to A. monticola and A. pindarense. From A. monticola it differs by the sparser punctation of the pronotum. From both species it can be distinguished by the details of the structure of the internal sac.</p> <p>Distribution. Amphichroum nepalicum is known from several localities in Nepal (Fig. 17).</p> <p>Bionomics. Specimens were collected at elevations from 1600 to 3500 m a.s.l.</p> <p>Remarks. Arpedium nepalicum was originally described based on a male from “ Népal (occidental), Dzunda, Khola Tal, près Talphi…”. I studied the holotype from NHMW, and based on the external morphology it is clearly belongs to the genus Amphichroum (see morphological features of the genus in Newton et al. 2020, Shavrin &amp; Smetana (2018)). Thus, a new combination is proposed here.</p> </div>	http://treatment.plazi.org/id/397C5D1DFFC1FFD6FF6FB46DFCD8FB50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFC0FFD5FF6FB46DFBA0FC78.text	397C5D1DFFC0FFD5FF6FB46DFBA0FC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum pindarense Champion 1920	<div><p>Amphichroum pindarense Champion, 1920</p> <p>(Figs. 6, 16, 23–24)</p> <p>Amphichroum pindarense Champion, 1920: 243; Cameron 1928: 558, 1930: 157, Coiffait 1978: 114</p> <p>Type material examined: Syntype, ♂ (Fig. 6): ‘ ♂ ’ &lt;printed&gt;, ‘Type | H.T.’ &lt;round label with red margin, printed&gt;, ‘Sunderhunga V., | 8-12,000 ft. | W. Almora. H.G.C.’ &lt;printed&gt;, ‘H.G. Champion | 1920-69’ &lt;printed&gt;, ‘Amphichroum | pindarense | Champ.’ &lt;handwritten&gt;, ‘ Amphichroum | pindarense Champion, 1920 | Shavrin A.V. det. 2021’ (BMNH); Syntype, ♀: ‘ ♀ ’ &lt;printed&gt;, other labels as the previous syntype (BMNH).</p> <p>Additional material studied: INDIA: 1 ♂, 1 ♀: same data as in syntypes, but without type labels (BMNH); CHINA: 5 ♂♂, 7 ♀♀: ‘ Tibet: Rongshar Valley 11,000ft 27.vi.1924. Maj.R.W.G.Hingston’, ‘ Everest Exp. Brit. Mus. 1924-386.’, ‘ Amphichroum pindarense Champ. ’, ‘ M.Cameron. Bequest. B.M.1955-147.’ (BMNH).</p> <p>Redescription. Measurements (n=16): HL: 0.27–0.36; HW: 0.57–0.65; AL (averaged): 1.45; OL: 0.13–0.16; PL: 0.57–0.68; PW: 0.90–1.09; ESL: 1.07–1.32; EW: 1.20–1.36; AW: 1.18–1.39; MTbL (averaged): 0.65; MTrL (averaged): 0.57 (MTrL 1–4: 0.30; MTrL 5: 0.27); AedL: 0.55–0.75; TL: 2.70–4.20 (holotype: 3.25).</p> <p>Habitus as in Fig. 6. Body yellow-brown to reddish brown, usually with slightly darker head, pronotum and abdomen, and paler elytra; apical maxillary segment and antennomeres 4–11 brown; mouthparts, antennomeres 1–3, legs and paratergites yellow. Head with distinct, dense, transverse microsculpture, finer and sparser between anteocellar foveae, and denser and coarser on infraorbital ridges; pronotum with very fine and sometimes indistinct microreticulation, without meshes in middle and mediobasal portion. Head with fine, irregular punctation, usually denser and larger in middle; pronotum with dense punctation, slightly larger and deeper than that on head, sparser and finer in mediobasal portion; abdomen with very sparse and fine punctation. Forebody with dense and short pubescence.</p> <p>Head about twice as broad as long or slightly narrower. Distance between ocelli about one and a half times distance between ocellus and posterior margin of eyes. Antennomere 3 slightly shorter and narrower than 2, 4 slightly broader than 3, 5–7 slightly longer and broader than 4, 8–10 slightly broader and distinctly shorter than 7.</p> <p>Pronotum 1.5–1.6 times as broad as long and 1.5–1.6 times as broad as head, widest about middle, slightly more narrowed posteriad than anteriad.</p> <p>Elytra 1.7–1.8 times as long as pronotum.</p> <p>Abdomen slightly narrower or broader than elytra.</p> <p>Male. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII straight or slightly concave. Aedeagus (Fig. 23) with moderately narrow basal portion, gradually narrowed toward subacute apex; parameres moderately narrow, slightly broadened apically, significantly exceeding apex of median lobe, with short apical and preapical setae; internal sac narrow and long, with two pairs of relatively large subtriangular teeth in basal and preapical portions. Lateral aspect of aedeagus as in Fig. 24.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Regarding the coloration and length of the body, A. pindarense is similar to A. monticola sp. n. and A. nepalicum, from which it can be distinguished by the wider body, longer elytra, narrower preapical part of the median lobe and the presence of paired large sclerotized structures in preapical and basal portions of the internal sac. The length of the elytra is similar to that in A. milkensis, but the apical portions of the parameres of A. pindarense are somewhat wider.</p> <p>Distribution. Amphichroum pindarense is known from several locations in India (Uttarakhand), China (Tibet, Xizang) and Bhutan (Fig. 16).</p> <p>Bionomics. Specimens were collected at elevations from 2400 to 3650 m a.s.l.</p></div> 	http://treatment.plazi.org/id/397C5D1DFFC0FFD5FF6FB46DFBA0FC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
397C5D1DFFC3FFD3FF6FB545FB6FFD3F.text	397C5D1DFFC3FFD3FF6FB545FB6FFD3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphichroum telnovi Shavrin 2021	<div><p>Amphichroum telnovi Shavrin, sp. n.</p> <p>(Figs. 7, 16, 25–26)</p> <p>Type material examined: Holotype, ♂ (dissected): ‘ NEPAL P: Seti D: Bajura | 15km S Simikot, N pass | slope W Malikasthan’ &lt;printed&gt;, ‘ 41-4200m, 29°50’42’’N | 81°47’25’’E 07.VII.2001 | leg. A.Weigel stone-deb.’ &lt;printed&gt;, ‘HOLOTYPE | Amphichroum | telnovi sp. n. | Shavrin A.V. des. 2021’ &lt;red, printed&gt; (NME).</p> <p>Paratypes: 1 ♂, 2 ♀♀ (apical and preapical antennomeres of both females are missing): same data as the holotype, with additional red label ‘ PARATYPE | Amphichroum | telnovi sp. n. | Shavrin A.V. des. 2021’ (1 ♂, 1 ♀: cSh, 1 ♀: NME).</p> <p>Description. Measurements (n=4): HL: 0.37–0.41; HW: 0.59–0.62; AL (holotype): 1.44; OL: 0.17–0.21; PL: 0.61–0.65; PW: 0.81–0.92; ESL: 1.12–1.20; EW: 1.16–1.26; AW: 0.90–1.12; MTbL (holotype): 0.67; MTrL (holotype): 0.54 (MTrL 1–4: 0.29; MTrL 5: 0.25); AedL: 0.65; TL: 3.56–3.95 (holotype: 3.70).</p> <p>Habitus as in Fig. 7. Body, antennomeres 4–11 and legs yellow-brown (head and disc of pronotum slightly darker); mouthparts, antennomeres 1–3 and tarsi yellow. Head with distinct and dense microsculpture, transverse on clypeus, finer in middle and isodiametric on infraorbital ridges; pronotum without visible microreticulation except for medioapical portion with very fine transverse meshes. Head with irregular, fine and sparse punctation, denser between anteocellar foveae; punctation of pronotum moderately dense, slightly larger and deeper than that in middle of head, sparser in middle and/or mediobasal portion; punctation of elytra larger and deeper than that on pronotum, coarser in basal portion and finer along suture; abdominal tergites with indistinct, sparse and fine punctation. Pronotum and elytra with dense and short pubescence.</p> <p>Head 1.5 times as broad as long, with narrow or moderately wide and deep anteocellar foveae, reaching level of middle or anterior margins of eyes. Ocelli very large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Antennomere 3 slightly longer and narrower than 2, 4 distinctly shorter than 3, 5–6 slightly longer and broader than 4, 7 slightly longer than 6, 8 slightly shorter than 7, 9–10 slightly longer than 8.</p> <p>Pronotum 1.3–1.4 times as broad as long and head, widest in about middle, evenly narrowed both anteriad and posteriad.</p> <p>Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded. Aedeagus (Fig. 25) with wide basal portion, gradually narrowed toward subacute apex; parameres very narrow, slightly exceeding apex of median lobe, with one short apical and preapical setae; internal sac narrow, long, with a pair of subapical and median and four basal sclerotized, elongate structures. Lateral aspect of aedeagus as in Fig. 26.</p> <p>Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</p> <p>Comparative notes. Based on the general shapes of the apical portion of the median lobe and the parameres, A. telnovi sp. n. is similar to A. altivagans and A. anthobioides. From A. altivagans it differs by the larger body, the shape of the pronotum, from middle evenly narrowed both anteriad and posteriad, and shorter elytra, and from A. anthobioides by the paler pronotum and elytra. From both species it can be distinguished by details of the structure of the internal sac.</p> <p>Distribution. The new species is known only from the type locality in western Nepal (Fig. 16).</p> <p>Bionomics. Specimens were collected at elevations from 4100 to 4200 m a.s.l.</p> <p>Etymology. Patronymic, the species is named to honor my colleague, Dmitry Telnov (London).</p></div> 	http://treatment.plazi.org/id/397C5D1DFFC3FFD3FF6FB545FB6FFD3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2021): New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5082 (1): 1-14, DOI: https://doi.org/10.11646/zootaxa.5082.1.1
