taxonID	type	description	language	source
075A8FD8A3B45743840D9C7DBB0D8B38.taxon	description	http: // zoobank. org / 32557822 - 9245 - 4 CE 3 - 9 E 5 B- 3 E 8 ABF 5 DEF 12 Figures 2, 3	en	Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B., Rađa 5, Tonci (2021): A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus. Subterranean Biology 41: 69-85, DOI: http://dx.doi.org/10.3897/subtbiol.41.75613, URL: http://dx.doi.org/10.3897/subtbiol.41.75613
075A8FD8A3B45743840D9C7DBB0D8B38.taxon	etymology	Etymology. The new species is named after Sinisa Solak, a geographer and naturalist, who was our guide during cave investigations conducted near the town of Glamoc.	en	Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B., Rađa 5, Tonci (2021): A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus. Subterranean Biology 41: 69-85, DOI: http://dx.doi.org/10.3897/subtbiol.41.75613, URL: http://dx.doi.org/10.3897/subtbiol.41.75613
075A8FD8A3B45743840D9C7DBB0D8B38.taxon	diagnosis	Diagnosis. The new species is closely related to some other species of Leonhardia by its large size and by the presence of a high mesosternal carina. Those other species are L. hilfi, L. reitteri, L. delminiumica, L. jajcensis, and L. sebesicensis (Figs 1, 4) (Jeannel 1924; Nonveiller et al. 2002; Curcic et al. 2014, 2018). Leonhardia solaki sp. nov. differs from L. hilfi in regard to AL in males (antennae exceeding the middle of elytra vs. antennae reaching the middle of elytra); A 10 L / A 10 W (less than 2.375 vs. more than 2.50); pronotum form (bell-shaped, lateral margins well-rounded anteriorly vs. subquadrate, lateral margins obtuse anteriorly); PL / PW (pronotum wider than long vs. pronotum as long as wide); shape of the mesosternal carina (higher, almost right-angled, posterior margin concave vs. lower, obtuse-angled, posterior margin somewhat elevated); shape of elytra (inversely ovate vs. widely oval); shape of the median lobe in dorsal view (apically flattened vs. apically narrowed); length of the median lobe (barely longer than parameres vs. markedly longer than parameres); shape and size of the basal bulb in dorsal view (larger, rounded vs. smaller, not rounded); and shape of the aedeagus in lateral view (median lobe more curved, basal bulb larger vs. median lobe less curved, basal bulb smaller) (Figs 2 - 4) (Reitter 1901; Jeannel 1924; Knirsch 1928; Perreau 1999; Nonveiller et al. 2002). Leonhardia solaki sp. nov. is easily distinguished from L. reitteri, from which it differs with respect to AL (antennae exceeding the middle of elytra in males and reaching the middle of elytra in females vs. antennae barely reaching the middle of body); A 2 L / A 3 L + A 4 L (antennomere II shorter than the following two antennomeres combined vs. antennomere II as long as the following two antennomeres combined); A 8 L / A 8 W (M in males 2.06, in females 1.58 vs. 1.50 in both genders); A 10 L / A 10 W (R in females 1.54 - 1.78 vs. more than 2.00 in both genders); shape of the hind pronotal angles (obtuse vs. right or weakly acute); shape of the mesosternal carina (almost right-angled, anterior margin convex, posterior margin regularly concave vs. obtuse-angled, anterior margin obtuse, posterior margin deeply incised); shape of elytra (inversely ovate vs. oval); shape of the median lobe in dorsal (wider in apical half, apically flattened vs. thinner in apical half, apically narrowed) and lateral (more elongate vs. less elongate) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs 2 - 4) (Breit 1902; Jeannel 1924; Knirsch 1928; Mueller 1937; Perreau 1999; Nonveiller et al. 2002). Leonhardia solaki sp. nov. differs from L. delminiumica in regard to TL (R 3.40 - 3.65 mm in males, 3.55 - 3.93 mm in females vs. R 3.10 - 3.20 mm in males, 3.40 mm in a female); A 8 L / A 8 W (M 2.06 in males, 1.58 in females vs. antennomere VIII barely longer than broad); shape of the mesosternal carina (almost right-angled, posterior margin less concave vs. obtuse-angled, posterior margin more concave); shape of the median lobe in dorsal (apically flattened, with a rounded apex vs. apically narrowed, with a pointed apex) and lateral (more curved, wider vs. less curved, narrower) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta closer to the two remaining setae vs. inner pre-apical seta farther away from the two remaining setae) (Figs 2 - 4) (Nonveiller et al. 2002). Leonhardia solaki sp. nov. is easily distinguished from L. jajcensis, from which it differs with respect to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse vs. almost right); shape of the mesosternal carina (higher, anterior margin more convex, posterior margin more concave vs. lower, anterior margin less convex, posterior margin less concave); form of the elytra (more elongate vs. less elongate); shape of the median lobe in dorsal (apically flattened vs. apically narrowed) and lateral (more thickened in its basal half, less convex ventrally vs. less thickened in its basal half, more convex ventrally) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); size of the basal bulb in dorsal view (larger vs. smaller); and position of parameral setae in dorsal view (inner pre-apical seta farther away from the two remaining setae vs. inner pre-apical seta closer to the two remaining setae) (Figs 2 - 4) (Curcic et al. 2014). Leonhardia solaki sp. nov. differs from L. sebesicensis in regard to AL in females (antennae reaching the middle of elytra vs. antennae ending before the middle of elytra); shape of the hind pronotal angles (obtuse-angled in both genders vs. almost right-angled in males); shape of the pronotum and elytra (less elongate vs. more elongate); shape of the mesosternal carina (apically rounded vs. apically toothed); shape of the median lobe in dorsal (apically flattened vs. apically rounded) and lateral (more curved, narrower in apical half vs. less curved, wider in apical half) views; length of the median lobe (barely longer than parameres vs. markedly longer than parameres); and position of parameral setae in dorsal view (inner pre-apical seta somewhat separated from two remaining setae vs. all setae equidistant) (Figs 2 - 4) (Curcic et al. 2018).	en	Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B., Rađa 5, Tonci (2021): A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus. Subterranean Biology 41: 69-85, DOI: http://dx.doi.org/10.3897/subtbiol.41.75613, URL: http://dx.doi.org/10.3897/subtbiol.41.75613
075A8FD8A3B45743840D9C7DBB0D8B38.taxon	description	Description. Medium-sized leptodirine. TL M 3.62 mm (3.55 mm in males, 3.69 mm in females), R 3.40 - 3.93 mm (3.40 - 3.65 mm in males, 3.55 - 3.93 mm in females). Habitus: Body shape pholeuonoid, colour reddish-brown (Figs 2 A, 3 A, B). Integument: Shiny, microsculptured (Fig. 3 D, G, L). Body covered with a number of densely distributed deep punctures and yellow pubescence of medium length (erect on head, recumbent on both pronotum and elytra). Head: Longer than wide (HL / HW M 1.11, R 1.06 - 1.19), anophthalmous, with no occipital carina (Figs 2 A, 3 C). Labrum slightly emarginate, with several long setae. Penultimate maxillary palpomere widened apically. Ultimate maxillary palpomere short, thin, gradually narrowing apically. Vertex with a longitudinal impression. Antennae inserted in middle third of head, slender, narrow proximally, slightly dilated distally, longer in males, AL M 2.15 mm, R 1.93 - 2.33 mm (2.165 - 2.33 mm in males, 1.93 - 2.165 mm in females), exceeding middle of elytra in males and reaching middle of elytra in females (Figs 2 A, 3 E). Antennomere I short and wide. A 1 L / A 2 L M 0.54, R 0.52 - 0.58. Antennomere II narrow, elongate, shorter than III and IV combined (A 2 L / A 3 + A 4 L M 0.86, R 0.79 - 0.90). Antennomeres III-VI of similar shape and size, narrow, among which V is the longest. Antennomeres VII and IX-XI widened (VII, IX, and X dilated distally), large, among which VII is the shortest in males, VIII the shortest in females, and XI the longest in both sexes. Antennomere VIII small, elongate in males (A 8 L / A 8 W M 2.06, R 1.82 - 2.40), wide in females (A 8 L / A 8 W M 1.58, R 1.33 - 1.90). A 9 L / A 9 W in males M 2.35 (R 2.25 - 2.50), in females M 1.68 (R 1.60 - 1.78). A 10 L / A 10 W in males M 2.27 (R 2.21 - 2.375), in females M 1.65 (R 1.54 - 1.78). Ultimate antennomere slender, narrowing apically, A 11 L / A 11 W in males M 3.10 (R 2.54 - 3.50), in females M 2.82 (R 2.67 - 3.00). Prothorax: Pronotum bell-shaped, transverse (PL / PW M 0.87, R 0.84 - 0.91), widest slightly after anterior third, markedly broader than head (Figs 2 A, 3 F). Lateral margins well-rounded anteriorly, somewhat concave posteriorly, sub-parallel basally. Pronotal base straight, markedly shorter than elytral base (PB / EB M 0.95, R 0.90 - 0.99). PB / AM M 1.40, R 1.18 - 1.55. Anterior margin somewhat convex medially. Fore pronotal angles small, rounded, obtuse. Hind pronotal angles obtuse, prominent, not protruding backwards. Pronotal disc moderately convex (Fig. 3 B). Mesothorax: Mesosternal carina high anteriorly, low posteriorly (Figs 2 B, 3 H). Anterior margin convex, posterior margin concave, with a few setae. Tooth obtuse, rounded. No mesoventral processus on mesoventrite (Fig. 3 I). Scutellum large, triangular (Figs 2 A, 3 J). Metathorax: Metasternum without carina. Elytra: Wide, obovoid, slightly wider in females (EL / EW in males M 1.47, R 1.40 - 1.57; in females M 1.44, R 1.41 - 1.48), markedly wider than pronotum (Figs 2 A, 3 K). Maximum width slightly before middle. Lateral margins arcuate. Marginal furrows visible on nearly entire elytra. Shoulders weakly expressed, almost rounded. Elytral disc markedly convex, gently declining basally and steeply declining apically in lateral view (Fig. 3 B). Sutural striae developed. Elytral apex attenuated, rounded. Pygidium covered by elytra. Legs: Elongate (Fig. 2 A). Femora widened basally, constricted sub-apically. Tibiae thin, very gently curved, gradually widening distally, each with several spines. Each fore tibia with an outer distal brush-like structure. Male fore tarsi five-segmented, weakly dilated. Tarsal claws elongate, narrow, curved, pointed apically. Male genitalia: Aedeagus elongate, straight, thin, sclerotized (Figs 2 C, D). Basal bulb small and rounded in dorsal view (Fig. 2 C), moderately large in lateral view (Fig. 2 D). Median lobe in dorsal view sub-parallel, rounded anteriorly, with a flattened apex, longer than parameres (Fig. 2 C). Median lobe in lateral view curved, straight in basal half, sub-apically concave dorsally, almost straight ventrally in basal part, with an acute apex curved downwards (Fig. 2 D). Parameres slender, narrow, arcuate, sub-apically curved towards exterior, each with a dilated rounded apex in dorsal view (Figs 2 C, E), while gently narrowing distally in lateral view (Fig. 2 D). Paramere bearing three setae: two strong, long, in apical and pre-apical position, respectively, and one inner, thin, short, in pre-apical position (Fig. 2 E). Two long parameral setae close-set, short one somewhat separated, positioned somewhat below level of other two parameral setae (Fig. 2 E). Endophallus elongate, tubular. Copulatory piece having a Y-shaped phanera and two lateral basal sclerifications, in front of which a weakly chitinized structure resembling teeth is present. In addition, two lateral bands are present in apical half of median lobe (Fig. 2 C). Female genitalia: Gonostyli slender, thin, gradually narrowing distally, straight, pointed apically (Fig. 2 F). Each gonostylus carrying one apical seta, three inner setae, and one outer seta (Fig. 2 F). Spermatheca elongate, chitinized, curved, constricted posteriorly in sub-apical part, spherical apically (Fig. 2 G). Female abdominal sternite VIII: Large, transverse, with a thin anterior process, carrying pubescence (Fig. 2 H).	en	Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B., Rađa 5, Tonci (2021): A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus. Subterranean Biology 41: 69-85, DOI: http://dx.doi.org/10.3897/subtbiol.41.75613, URL: http://dx.doi.org/10.3897/subtbiol.41.75613
075A8FD8A3B45743840D9C7DBB0D8B38.taxon	distribution	Geographic distribution. Thus far, the species is known only from its type locality - the Golubnjaca kod Skucana Pit, situated in the village of Skucani, close to the town of Glamoc in western Bosnia and Herzegovina. This site represents the westernmost location of a Leonhardia species. We assume that the new species probably inhabits other subterranean sites in the surrounding areas of western Bosnia and Herzegovina, although one of us (TR) visited the Ledenjaca Cave situated in the same village (Skucani), but found no specimens of the new species there.	en	Curcic 1, Srecko, Vesovic 1, Nikola, Vrbica 1, Maja, Popovic 1,2, Slađana, Radovanovic 3, Zeljko, Curcic 4, Nina B., Rađa 5, Tonci (2021): A new species of Leonhardia Reitter, 1901 (Coleoptera, Leiodidae, Leptodirini) from Bosnia and Herzegovina, with a key to species of the genus. Subterranean Biology 41: 69-85, DOI: http://dx.doi.org/10.3897/subtbiol.41.75613, URL: http://dx.doi.org/10.3897/subtbiol.41.75613
