taxonID	type	description	language	source
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	description	(Fig. 1, 3).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	materials_examined	Typus: BRAZIL. Roraima: SEMA Ecological Reserve, Ilha de Maracá, riverine vegetation bordering the Igarapé Pedra Sentada, 3 ° 30 ’ N 61 ° 43 ’ W, 30. III. 1987, Milliken & Lima 35 (holo-: NY!; iso-: E, K!).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	diagnosis	Arbor parva, ramulis terminalis puberulentis vel pilosis. Nonnulli pili longior quam 0,8 mm. Lamina subcoriacea, elliptica, rarius ovato-elliptica, (2,8 -) 4,8 - 14,1 Χ (1,1 -) 2,2 - 4,6 cm. Inflorescentia masculina 11 - 32 - flora, hermaphrodita 5 - 11 - flora. Fructus 0,6 - 0,9 Χ 0,55 - 0,8 cm.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	description	Small trees or shrubs to 12 m tall. Terminal internodes laterally compressed or subterete, longitudinally striate, 0.8 - 2 mm in diameter, densely puberulent to pilose, with trichomes 0.1 - 0.4 mm long, erect, 0.2 - 1.5 mm long (at least some hairs> 0.8 mm long), curved, multicellular. Leaves petiolate; stipules caducous, elliptical, subulate, or narrowly ovate, (0.45 -) 0.9 - 3.2 Χ (0.06 -) 0.1 - 0.6 cm, apex acute, abaxially pilose, the adjacent stipules 0.1 - 0.4 mm distant at base, scars transversely elliptical to orbicular, 0.3 - 0.6 Χ 0.4 - 1 mm; petiole subterete, inconspicuously canaliculate or even, ± equally thickened, 0.1 - 0.5 cm long, 0.8 - 2 mm in diameter; lamina subcoriaceous, elliptical (or ovate-elliptical), (2.8 -) 4.8 - 14.1 Χ (1.1 -) 2.2 - 4.6 cm, base attenuate, cuneate, or acute, apex (sub-) acuminate, adaxially glabrous or pilose at base, abaxially pilose along midvein (or covering the whole surface when juvenile); margin inconspicuously revolute, minutely serrulate or entire; secondary veins 7 - 14 per side, adaxially slightly impressed or prominulous, abaxially prominulous, 0.25 - 1.7 cm distant at middle of lamina, interspersed by 0 - 2 (- 4) conspicuous intersecondary veins. Male inflorescence thyrsoid, 11 - 32 - flowered, with flowers in fascicles of (1 -) 3, hermaphroditic inflorescence botryoid, 5 - 11 - flowered, with flowers solitary, not in fascicles; rachis 1 per axile, laterally compressed (or subterete), longitudinally furrowed, inconspicuously thickened in fruit, 0.8 - 1.7 (- 2.7) cm long, 0.5 - 1 mm in diameter, densely pilose; bracts (sub-) opposite, ovate to elliptical, (0.9 -) 1.4 - 2.5 Χ 0.7 - 1.3 mm, apex acute, glabrous or sparsely hairy; bracteoles subulate to ovate, 0.4 - 1.3 0.2 - 0.4 mm, apex acuminate; pedicel below articulation to (0.3 -) 0.7 - 2.4 mm long, above articulation (sub-) terete, (1.5 -) 3 - 5 mm long, 0.3 - 0.7 mm in diameter, inconspicuously puberulent or glabrous. Flowers 2.5 - 3 mm in diameter; sepals 4, coriaceous, elliptical to ovate (to suborbicular), 1.4 - 1.7 (- 2.3) Χ 1.1 - 1.6 mm, apex rounded or obtuse; petals 4, yellow, elliptical to obovate, c. 2 Χ 1.6 mm, apex rounded, glabrous, rarely ciliolate; stamens in male flowers 21 - 26, in hermaphroditic flowers c. 12, filaments to 1.8 mm long, free or in hermaphroditic flowers sometimes adnate to petals; ovary 2 - locular, styles 2, 1.8 - 1.9 mm long. Fruit subglobose, ellipsoid or obovoid, longitudinally striate, 0.6 - 0.9 Χ 0.55 - 0.8 cm, apex rounded, glabrous; seeds 1 per fruit, ellipsoid to subglobose, 0.55 Χ 0.5 cm, densely villous (trichomes 0.3 - 0.7 mm long).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	distribution	Distribution. – Known from southern Venezuela and northern Brazil. Occurs in terra firme and riverine forests, in campina or capoeira, on sandy or bauxitic (lateritic) soils; up to 800 m.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	biology_ecology	Phenology. – Flowering in February and August (few data available), fruiting from February to August.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	etymology	Etymology. – The species epithet refers to the type locality, the Ilha de Maracá.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	discussion	Notes. – This species is characterized by the combination of small ellipsoid to subglobose fruits, small ± lanceolate stipules, pedicels with basal articulation, and especially the long multicellular trichomes (at least in some parts> 0.8 mm long) at terminal internodes, leaves and inflorescences. Quiina florida Tul. is similar in leaf shape, but differs in larger fruits and inflorescences, the usually broader stipules, and shorter trichomes. Quiina wurdackii Pires differs in shorter inflorescences and few, rather obscure secondary veins, while Q. parvifolia Lanj. & Heerdt is distinguished by broader stipules, broader leaf blades, and longer pedicels.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFEFF8ED13CF9C26B7BF984.taxon	materials_examined	Additional material examined. – VENEZUELA. Bolívar: Municipio Cedeño, Cabeceras del río Túriba y Caño La Miel, 45 km E de Los Pijiguaos, VIII. 1989, Sanoja & Fernández 2904 (MO); Municipio Cedeño, headwaters of río Túriba and Caño La Miel, 45 km E of Pijiguaos, 6 ° 34 ’ N 66 ° 23 ’ W, VIII. 1989, Fernández & Sanoja 5892 (MO, NY, PORT); Municipio Raúl Leoni, headwaters of río Túriba and Caño La Miel, 45 km E of Pijiguaos, 6 ° 34 ’ N 66 ° 23 ’ W, VI. 1989, Fernández & Delgado 5813 (MO, NY, PORT). Amazonas: Alto Orinoco, Salto Salas, 18. VIII. 1951, Croizat 536 (NY); Dept. Atabapo, Caño Jayuwapuey, tributary of the Ocamo River, 3 ° 4 ’ N 64 ° 40 ’ W, I. 1990, Fernández 6789 (PORT). BRAZIL. Amazonas: Vicinity of Pico Rondon, Perimetral Norte, Highway Km 211, 3 km from Km 211, lower slopes of Pico Rondon, 1 ° 32 ’ N 62 ° 48 ’ W, 2. II. 1984, Prance & al. 28761 (F, GH, K, NY, UB). Pará: Sete Varas airstrip on Rio Curua, 0 ° 95 ’ S 54 ° 92 ’ W, 4. VIII. 1981, Strudwick & al. 4083 (F, K, NY, UEC). Roraima: Município Alto Alegre, Ilha de Maracá, SEMA Estação, forest trails close to Estação, 3 ° 22 ’ N 61 ° 20 ’ W, 6. VI. 1986, Hopkins & al. 515 (FR, NY); Município Alto Alegre, Ilha de Maracá, SEMA Estação, Furo Pananá de Firmino of Rio Uraricuera on S side of island, forest near Casa Maracá, 3 ° 24 ’ N 61 ° 26 ’ W, 10. VI. 1986, Hopkins & al. 634 (FR, NY); Município de Boa Vista, Estação Ecológica de Maracá, 21. V. 1987, Lima 804 (K); SEMA Ecological Reserve, Ilha de Maracá, 3 ° 22 ’ N 61 ° 26 ’ W, 21. IV. 1987, Milliken 77 (E, K, NY); Serrinha, Rio Mucajaí, 31. I. 1967, Prance & al. 4198 (COL, F, GH, IAN, K, NY, R, S, U); Posto Mucajaí, Rio Mucajaí, Vicinity of Mucajaí airstrip, 13. III. 1971, Prance & al. 10928 (F, IAN, K, M, NY, P, R, S, U); SEMA Ecological Station, Ilha de Maracá, in forest at Santa Rosa, 3 ° 22 ’ N 61 ° 25 ’ W, 7. III. 1987, Ratter & al. 5587 (E, K); SEMA Ecological Station, Ilha de Maracá, 3 ° 22 ’ N 61 ° 25 ’ W, 11. III. 1987 (st), Ratter & al. 5679 (K); SEMA Ecological Sation, Ilha de Maracá, 3 ° 22 ’ N 61 ° 25 ’ W, 18. III. 1987, Ratter & al. 5748 (E, K); Surroundings of Ecological Station of Maracá, 3 ° 22 ’ N 61 ° 25 ’ W, 9. II. 1979, Rosa 3121 (F, NY).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	description	(Fig. 2, 3).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	materials_examined	Typus: ECUADOR. Zamora-Chinchipe: road Loja-Zamora, c. 35 km from Loja, Estación Científica San Francisco, 3 ° 58 ’ S 79 ° 04 ’ W, 1800 m, 16. III. 2000, Homeier 333 (holo-: BIEL!; iso-: QCNE, MO, LOJA).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	description	Folia rigida. Inflorescentia hermaphrodita solum visa racemiforma, 1 - 11 - flora, rhachidi 0,3 - 0,7 cm longa. Trees. Terminal internodes terete, longitudinally furrowed, 2 - 3 mm in diameter, glabrous. Leaves petiolate; stipules caducous, (narrowly) triangular-ovate, 0.3 - 0.9 Χ 0.08 - 0.15 cm, apex acute, glabrous or adaxially puberulent, the adjacent stipules joint at base or to 0.3 mm distant; petiole canaliculate, 0.3 - 0.65 cm long, 1.5 - 3 mm in diameter; lamina coriaceous, rigid, elliptical, 7 - 16 Χ 3.2 - 6.9 cm, base shortly attenuate to broadly cuneate, apex shortly (sub-) acuminate, the very apex rounded or obtuse, adaxially glabrous, abaxially sparsely hairy along midvein, the hairs red-brown, 0.3 - 1.1 mm long; margin not or minutely revolute, entire or very inconspicuously serrulate; secondary veins 8 - 16 per side, adaxially impressed to prominulous, abaxially prominent, 0.7 - 1.7 cm distant at middle of lamina, interspersed by 0 - 1 (- 4) intersecondary veins. Hermaphroditic inflorescence botryoid, 1 - 11 - flowered, male inflorescence not seen; rachis 1 (- 3) per axile, subterete to inconspicuously quadrangular, longitudinally furrowed, 0.3 - 0.7 cm long, c. 1 mm in diameter, ± glabrous or sparsely puberulent with brownish hairs to 0.3 mm long; bracts subopposite, ovate, 0.6 - 1.3 Χ 0.6 - 1 mm, apex acute, puberulent, bracteoles not seen; pedicel below articulation to 2 mm long, above articulation subterete, apically inconspicuously widened, 5 - 10 mm long (in male flowers to 12 mm long), 0.7 - 0.8 mm in diameter, glabrous. Flowers subglobose in bud; sepals 4, coriaceous, elliptical to suborbicular, strongly concave, 2 - 3 Χ 1.3 - 4 mm, rounded, abaxially glabrous, margin ciliate; petals 5, obovate, reflexed, 4 - 5 Χ 2 - 4 mm, rounded, margin ciliolate; stamens in hermaphroditic flowers c. 17, in male flowers c. 65 (see remarks), filaments c. 5 mm long, anthers c. 0.4 - 0.7 Χ 0.3 - 0.6 mm, in male flowers larger; ovary 2 - locular, styles 2, 2.5 - 3 mm long, stigma inconspicuously broadened. Fruit ellipsoid, longitudinally striate, 2 - 3 Χ 1.2 - 1.5 cm, apically obtuse to subacute, glabrous; seeds not seen.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	distribution	Distribution. – Only known from Ecuador. Occurs in tropical montane rain forest at about 1800 m.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	biology_ecology	Phenology. – Flowers in March.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	etymology	Etymology. – The species epithet refers to the type locality near Zamora.	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	materials_examined	Additional material examined. – ECUADOR. Zamora-Chinchipe: road Loja-Zamora, c. 35 km from Loja, Estación Científica San Francisco, 1830 m, 7 Apr 2004, Homeier 1375 (BIEL, FR).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
A2708787FFFBFF8ED12DF9CC6E68F959.taxon	discussion	Notes. – This species is similar in leaf shape to Q. florida, but contrasts in a rigid-coriaceous leaf texture. The leaf texture is similar to Q. integrifolia Pulle, but this species has longer inflorescences with more flowers. The short inflorescences, in turn, are similar to Q. blackii Pires. However, Q. blackii differs in chartaceous to subcoriaceous leaves and subulate stipules. Male flowers were provided by J. Homeier (no specimen seen).	en	Schneider, Julio V., Zizka, Georg (2012): Quiina maracaensis J. V. Schneid. & Zizka and Quiina zamorensis J. V. Schneid. & Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67 (2): 261-267, DOI: 10.15553/c2012v672a7
