taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
2A35ED1E0664500F8461A6EFFFC6C926.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621142	http://doi.org/10.3897/zookeys.1072.69025.figure7	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
2A35ED1E0664500F8461A6EFFFC6C926.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621144	http://doi.org/10.3897/zookeys.1072.69025.figure9	Figure 9. External and internal morphology of Edwardsianthus amethystus sp. nov. (CMNH-ZG 09763) A oral view of living specimen in the habitat B outer view of preserved specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of retractor muscle F transverse section of testis G transverse section of trace of nemathybome. Abbreviations: fi, filament; me, mesoglea; ne, nemathybome-like structure; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacle longitudinal muscle; ts, testis. Scale bars: 1 cm (A, B); 500 µm (E, F); 100 µm (C, D, G).	Figure 9. External and internal morphology of Edwardsianthus amethystus sp. nov. (CMNH-ZG 09763) A oral view of living specimen in the habitat B outer view of preserved specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of retractor muscle F transverse section of testis G transverse section of trace of nemathybome. Abbreviations: fi, filament; me, mesoglea; ne, nemathybome-like structure; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacle longitudinal muscle; ts, testis. Scale bars: 1 cm (A, B); 500 µm (E, F); 100 µm (C, D, G).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
2A35ED1E0664500F8461A6EFFFC6C926.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621145	http://doi.org/10.3897/zookeys.1072.69025.figure10	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
EA704A69959E541C9C0F1550DE68CC12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621138	http://doi.org/10.3897/zookeys.1072.69025.figure3	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
EA704A69959E541C9C0F1550DE68CC12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621140	http://doi.org/10.3897/zookeys.1072.69025.figure5	Figure 5. External and internal morphology of Edwardsianthus carbunculus sp. nov. (CMNH-ZG 05954) A outer view of preserved specimen (aboral end is damaged) B oral view of living specimen in the habitat (photograph Kensuke Yanagi) C transverse section of column in upper part D transverse section of lower column in lower part E transverse section of the macrocneme F transverse section of testis G transverse section of nemathybomes. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; me, mesoglea; ne, nemathybomes; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A, B); 500 µm (C-F); 100 µm (G). Picture B taken by Kensuke Yanagi.	Figure 5. External and internal morphology of Edwardsianthus carbunculus sp. nov. (CMNH-ZG 05954) A outer view of preserved specimen (aboral end is damaged) B oral view of living specimen in the habitat (photograph Kensuke Yanagi) C transverse section of column in upper part D transverse section of lower column in lower part E transverse section of the macrocneme F transverse section of testis G transverse section of nemathybomes. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; me, mesoglea; ne, nemathybomes; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A, B); 500 µm (C-F); 100 µm (G). Picture B taken by Kensuke Yanagi.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
EA704A69959E541C9C0F1550DE68CC12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621145	http://doi.org/10.3897/zookeys.1072.69025.figure10	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
EA704A69959E541C9C0F1550DE68CC12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621136	http://doi.org/10.3897/zookeys.1072.69025.figure1	Figure 1. Localities of the specimens of the Edwardsianthus species collected in this study. Purple star marked by p indicates collection locality of Edwardsianthus pudicus; orange stars are those of E. gilbertensis; red star marked by c is E. carbunculus sp. nov.; blue star marked by a is E. sapphirus sp. nov.; green star marked by s is E. smaragdus sp. nov.	Figure 1. Localities of the specimens of the Edwardsianthus species collected in this study. Purple star marked by p indicates collection locality of Edwardsianthus pudicus; orange stars are those of E. gilbertensis; red star marked by c is E. carbunculus sp. nov.; blue star marked by a is E. sapphirus sp. nov.; green star marked by s is E. smaragdus sp. nov.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
5B726EEA8CCF56B8B680BA201207CCF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621136	http://doi.org/10.3897/zookeys.1072.69025.figure1	Figure 1. Localities of the specimens of the Edwardsianthus species collected in this study. Purple star marked by p indicates collection locality of Edwardsianthus pudicus; orange stars are those of E. gilbertensis; red star marked by c is E. carbunculus sp. nov.; blue star marked by a is E. sapphirus sp. nov.; green star marked by s is E. smaragdus sp. nov.	Figure 1. Localities of the specimens of the Edwardsianthus species collected in this study. Purple star marked by p indicates collection locality of Edwardsianthus pudicus; orange stars are those of E. gilbertensis; red star marked by c is E. carbunculus sp. nov.; blue star marked by a is E. sapphirus sp. nov.; green star marked by s is E. smaragdus sp. nov.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
5B726EEA8CCF56B8B680BA201207CCF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621145	http://doi.org/10.3897/zookeys.1072.69025.figure10	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
3736F24343895DFFBA57229A0452E87E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621138	http://doi.org/10.3897/zookeys.1072.69025.figure3	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
3736F24343895DFFBA57229A0452E87E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621139	http://doi.org/10.3897/zookeys.1072.69025.figure4	Figure 4. External and internal morphology of Edwardsianthus gilbertensis (CMNH-ZG 06527 except for B) A outer view of preserved specimen B oral view of living specimen with 18 tentacles in the habitat C longitudinal section of oral end D transverse section of nemathybome E transverse section of column in lower part; F. Enlarged view of transverse section of mesenteries. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; ne, nemathybomes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A); 1 mm (B); 500 µm (C, E, F); 100 µm (D). Photograph B by Kensuke Yanagi.	Figure 4. External and internal morphology of Edwardsianthus gilbertensis (CMNH-ZG 06527 except for B) A outer view of preserved specimen B oral view of living specimen with 18 tentacles in the habitat C longitudinal section of oral end D transverse section of nemathybome E transverse section of column in lower part; F. Enlarged view of transverse section of mesenteries. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; ne, nemathybomes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A); 1 mm (B); 500 µm (C, E, F); 100 µm (D). Photograph B by Kensuke Yanagi.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
11CE36E72E175D6AA1F958A3E40FC11F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621137	http://doi.org/10.3897/zookeys.1072.69025.figure2	Figure 2. External and internal morphology of Edwardsianthus pudicus (NSMT-Co 1702) A outer view of living specimen B oral view of living specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of parietal muscle F transverse section of macrocnemes G longitudinal section of physa. Abbreviations: cap, capitulum; fi, filament; me, mesoglea; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 5 mm (A, B); 1 mm (C); 500 µm (D, F, G); 100 µm (E).	Figure 2. External and internal morphology of Edwardsianthus pudicus (NSMT-Co 1702) A outer view of living specimen B oral view of living specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of parietal muscle F transverse section of macrocnemes G longitudinal section of physa. Abbreviations: cap, capitulum; fi, filament; me, mesoglea; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 5 mm (A, B); 1 mm (C); 500 µm (D, F, G); 100 µm (E).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
11CE36E72E175D6AA1F958A3E40FC11F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621138	http://doi.org/10.3897/zookeys.1072.69025.figure3	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	Figure 3. Cnidae of Edwardsianthus species A-E E. pudicus (NSMT-Co 1702) A 1 spirocyst in tentacle; A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx B 3 microbasic p - mastigophore in actinopharynx C basitrich in nemathybome D basitrich in column E 1 small basitrich in filament E 2 large basitrich in filament E 3 microbasic p - mastigophore in filament F-J E. gilbertensis (CMNH-ZG 06527) F 1 spirocyst in tentacle F 2 basitrich in tentacle G 1 small basitrich in actinopharynx G 2 large basitrich in actinopharynx G 3 microbasic p - mastigophore in actinopharynx H basitrich in nemathybome I basitrich in nemathybome J 1 basitrich in filament J 2 microbasic p - mastigophore in filament K-O E. carbunculus sp. nov. (CMNH-ZG 05954) K 1 spirocyst in tentacle K 2 basitrich in tentacle L 1 small basitrich in actinopharynx L 2 large basitrich in actinopharynx M 1 small basitrich in nemathybome M 2 large basitrich in nemathybome N basitrich in column O 1 small basitrich in filament O 2 large basitrich in filament O 3 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
D4431F22BDB7546DA6FC73888125A467.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621141	http://doi.org/10.3897/zookeys.1072.69025.figure6	Figure 6. External and internal morphology of Edwardsianthus sapphirus sp. nov. (CMNH-ZG 09761). A oral view of living specimen in the habitat B outer view of preserved specimen; C. Transverse section of column in upper part D enlarged view of transverse section of parietal muscle E enlarged view of transverse section of retractor muscle F transverse section of ovary G longitudinal section of tentacle H transverse section of tentacle. Abbreviations: a, actinopharynx; fi, filament; me, mesoglea; ne, nemathybome; oo, oocytes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm (C-E); 100 µm (F-H).	Figure 6. External and internal morphology of Edwardsianthus sapphirus sp. nov. (CMNH-ZG 09761). A oral view of living specimen in the habitat B outer view of preserved specimen; C. Transverse section of column in upper part D enlarged view of transverse section of parietal muscle E enlarged view of transverse section of retractor muscle F transverse section of ovary G longitudinal section of tentacle H transverse section of tentacle. Abbreviations: a, actinopharynx; fi, filament; me, mesoglea; ne, nemathybome; oo, oocytes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm (C-E); 100 µm (F-H).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
D4431F22BDB7546DA6FC73888125A467.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621142	http://doi.org/10.3897/zookeys.1072.69025.figure7	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621142	http://doi.org/10.3897/zookeys.1072.69025.figure7	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	Figure 7. Cnidae of Edwardsianthus species A-D E. sapphirus sp. nov. (CMNH-ZG 09761) A 1 spirocyst in tentacle A 2 basitrich in tentacle B 1 small basitrich in actinopharynx B 2 large basitrich in actinopharynx C 1 small basitrich in nemathybome C 2 large basitrich in nemathybome D 1 small basitrich in filament D 2 large basitrich in filament D 3 microbasic p - mastigophore in filament E-H E. smaragdus sp. nov. (CMNH-ZG 09762) E 1 spirocyst in tentacle E 2 basitrich in tentacle F 1 small basitrich in actinopharynx F 2 large basitrich in actinopharynx F 3 microbasic p - mastigophore in actinopharynx G 1 small basitrich in nemathybome; G 2 large basitrich in nemathybome; H 1 spirocyst in filament; H 2 small basitrich in filament H 3 large basitrich in filament H 4 microbasic p - mastigophore in filament I-K E. amethystus sp. nov. (CMNH-ZG 09763) I 1 spirocyst in tentacle I 2 basitrich in tentacle J 1 small basitrich in actinopharynx J 2 large basitrich in actinopharynx K 1 spirocyst in filament K 2 small basitrich in filament K 3 large basitrich in filament K 4 microbasic p - mastigophore in filament.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621143	http://doi.org/10.3897/zookeys.1072.69025.figure8	Figure 8. External and internal morphology of Edwardsianthus smaragdus sp. nov. (CMNH-ZG 09762). A outer view of living specimen B outer view of preserved specimen C transverse section of retractor muscle D transverse section of the tentacle E longitudinal section of the tentacle F transverse section of column G enlarged view of transverse section of ovary H transverse section of a nemathybome. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; me, mesoglea; ne, nemathybomes; ov, ovary; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tcm, tentacular circular muscle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm in (C - H).	Figure 8. External and internal morphology of Edwardsianthus smaragdus sp. nov. (CMNH-ZG 09762). A outer view of living specimen B outer view of preserved specimen C transverse section of retractor muscle D transverse section of the tentacle E longitudinal section of the tentacle F transverse section of column G enlarged view of transverse section of ovary H transverse section of a nemathybome. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; me, mesoglea; ne, nemathybomes; ov, ovary; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tcm, tentacular circular muscle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm in (C - H).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621137	http://doi.org/10.3897/zookeys.1072.69025.figure2	Figure 2. External and internal morphology of Edwardsianthus pudicus (NSMT-Co 1702) A outer view of living specimen B oral view of living specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of parietal muscle F transverse section of macrocnemes G longitudinal section of physa. Abbreviations: cap, capitulum; fi, filament; me, mesoglea; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 5 mm (A, B); 1 mm (C); 500 µm (D, F, G); 100 µm (E).	Figure 2. External and internal morphology of Edwardsianthus pudicus (NSMT-Co 1702) A outer view of living specimen B oral view of living specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of parietal muscle F transverse section of macrocnemes G longitudinal section of physa. Abbreviations: cap, capitulum; fi, filament; me, mesoglea; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 5 mm (A, B); 1 mm (C); 500 µm (D, F, G); 100 µm (E).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621139	http://doi.org/10.3897/zookeys.1072.69025.figure4	Figure 4. External and internal morphology of Edwardsianthus gilbertensis (CMNH-ZG 06527 except for B) A outer view of preserved specimen B oral view of living specimen with 18 tentacles in the habitat C longitudinal section of oral end D transverse section of nemathybome E transverse section of column in lower part; F. Enlarged view of transverse section of mesenteries. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; ne, nemathybomes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A); 1 mm (B); 500 µm (C, E, F); 100 µm (D). Photograph B by Kensuke Yanagi.	Figure 4. External and internal morphology of Edwardsianthus gilbertensis (CMNH-ZG 06527 except for B) A outer view of preserved specimen B oral view of living specimen with 18 tentacles in the habitat C longitudinal section of oral end D transverse section of nemathybome E transverse section of column in lower part; F. Enlarged view of transverse section of mesenteries. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; ma, macrocneme; ne, nemathybomes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A); 1 mm (B); 500 µm (C, E, F); 100 µm (D). Photograph B by Kensuke Yanagi.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621140	http://doi.org/10.3897/zookeys.1072.69025.figure5	Figure 5. External and internal morphology of Edwardsianthus carbunculus sp. nov. (CMNH-ZG 05954) A outer view of preserved specimen (aboral end is damaged) B oral view of living specimen in the habitat (photograph Kensuke Yanagi) C transverse section of column in upper part D transverse section of lower column in lower part E transverse section of the macrocneme F transverse section of testis G transverse section of nemathybomes. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; me, mesoglea; ne, nemathybomes; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A, B); 500 µm (C-F); 100 µm (G). Picture B taken by Kensuke Yanagi.	Figure 5. External and internal morphology of Edwardsianthus carbunculus sp. nov. (CMNH-ZG 05954) A outer view of preserved specimen (aboral end is damaged) B oral view of living specimen in the habitat (photograph Kensuke Yanagi) C transverse section of column in upper part D transverse section of lower column in lower part E transverse section of the macrocneme F transverse section of testis G transverse section of nemathybomes. Abbreviations: a, actinopharynx; cap, capitulum; fi, filament; me, mesoglea; ne, nemathybomes; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle. Scale bars: 5 mm (A, B); 500 µm (C-F); 100 µm (G). Picture B taken by Kensuke Yanagi.	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621141	http://doi.org/10.3897/zookeys.1072.69025.figure6	Figure 6. External and internal morphology of Edwardsianthus sapphirus sp. nov. (CMNH-ZG 09761). A oral view of living specimen in the habitat B outer view of preserved specimen; C. Transverse section of column in upper part D enlarged view of transverse section of parietal muscle E enlarged view of transverse section of retractor muscle F transverse section of ovary G longitudinal section of tentacle H transverse section of tentacle. Abbreviations: a, actinopharynx; fi, filament; me, mesoglea; ne, nemathybome; oo, oocytes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm (C-E); 100 µm (F-H).	Figure 6. External and internal morphology of Edwardsianthus sapphirus sp. nov. (CMNH-ZG 09761). A oral view of living specimen in the habitat B outer view of preserved specimen; C. Transverse section of column in upper part D enlarged view of transverse section of parietal muscle E enlarged view of transverse section of retractor muscle F transverse section of ovary G longitudinal section of tentacle H transverse section of tentacle. Abbreviations: a, actinopharynx; fi, filament; me, mesoglea; ne, nemathybome; oo, oocytes; pa, parietal muscle; ph, physa; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacular longitudinal muscle. Scale bars: 1 cm (A, B); 500 µm (C-E); 100 µm (F-H).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621144	http://doi.org/10.3897/zookeys.1072.69025.figure9	Figure 9. External and internal morphology of Edwardsianthus amethystus sp. nov. (CMNH-ZG 09763) A oral view of living specimen in the habitat B outer view of preserved specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of retractor muscle F transverse section of testis G transverse section of trace of nemathybome. Abbreviations: fi, filament; me, mesoglea; ne, nemathybome-like structure; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacle longitudinal muscle; ts, testis. Scale bars: 1 cm (A, B); 500 µm (E, F); 100 µm (C, D, G).	Figure 9. External and internal morphology of Edwardsianthus amethystus sp. nov. (CMNH-ZG 09763) A oral view of living specimen in the habitat B outer view of preserved specimen C longitudinal section of tentacle D transverse section of tentacle E transverse section of retractor muscle F transverse section of testis G transverse section of trace of nemathybome. Abbreviations: fi, filament; me, mesoglea; ne, nemathybome-like structure; pa, parietal muscle; rm, retractor muscle; scs, scapus; te, tentacle; tlm, tentacle longitudinal muscle; ts, testis. Scale bars: 1 cm (A, B); 500 µm (E, F); 100 µm (C, D, G).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
E7110EE880295AC792592D11B062ADC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/621145	http://doi.org/10.3897/zookeys.1072.69025.figure10	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	Figure 10. Maximum-likelihood tree of the order Actiniaria based on the combined dataset of mitochondrial 12 S and 16 S and nuclear 18 S rDNA (total 2866 bp). The clade of the genus Edwardsianthus is colored in a red box. Red bar at node indicates the position at which nemathybomes would be obtained. Numbers indicate ML bootstrap support values followed by BI posterior probabilities of the nodes (bootstrap values of ≥ 50 % and posterior probabilities ≥ 0.5 are shown).	2021-12-10	Izumi, Takato;Fujii, Takuma		Zenodo	biologists	Izumi, Takato;Fujii, Takuma			
