taxonID	type	description	language	source
20E1B7B63D6E5803A70E990EA4551A91.taxon	diagnosis	Diagnosis. Only one species of Alesiella is known and can be recognized by characters in the above key to genera. The specimen from Thailand does not differ from the type material (previously studied by the author), though the aedeagus was lost during mounting (Y. Hayashi, pers. comm.).	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
20E1B7B63D6E5803A70E990EA4551A91.taxon	distribution	Distribution. Myanmar and Thailand (new country record).	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
2EA08D86F9085E09BFA318FAF72F74D5.taxon	diagnosis	Diagnosis. Cyrtoquediini (as recently redefined by Brunke et al. (2021)) can be recognized among other Staphylininae based on the following combination of characters: microsculpture on disc of head and pronotum absent; obvious presence of both posterior frontal and basal punctures (Brunke et al., 2019: fig. 1); profemora with apical row of lateroventral spines (near joint with protibia) (Brunke et al. 2021: fig. 8 B); protibia without subapical notch (Brunke et al. 2021: fig. 8 C); metatarsomeres 1 - 4 flattened and trapezoidal, not elongate and cylindrical. Most genera in Cyrtoquediini can also be recognized by the unique row of coarse, impressed setose punctures on the elytral epipleuron (Brunke et al. 2016: fig. 4).	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
E50B322C5347585B901BF70A83A4E7CE.taxon	description	Fig. 3 A-G	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
E50B322C5347585B901BF70A83A4E7CE.taxon	etymology	Etymology. The generic name is a combination of the Latin word fluvium (river, stream) and Raphirus (a subgenus of Quedius), where the only species of Fluviphirus was previously classified and to which it bears a superficial resemblance. Noun in apposition.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
E50B322C5347585B901BF70A83A4E7CE.taxon	diagnosis	Diagnosis. Among other Indoquediini, Fluviphirus is easily recognized by the combination of meshed microsculpture on the forebody and the absence of interocular punctures on the head. It is also the only genus of Nearctic Indoquediini.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
E50B322C5347585B901BF70A83A4E7CE.taxon	description	Description. With the character states of Indoquediini (see Brunke et al. 2021) and the following: disc of head and pronotum with meshed microsculpture; eyes moderately convex, not strongly bulging, large, distinctly larger than temples (Fig. 3 A); head with single basal puncture, interocular punctures absent, temples with numerous smaller punctures, with single parocular puncture; antennomere 3 with dense but not tomentose pubescence; apical maxillary palpomere glabrous; penultimate labial palpomere with brush of dense setae (but sparser than that of Indoquedius); pronotum with two punctures in dorsal row, ' second' puncture present (Fig. 3 A); postcoxal process fused across inferior marginal line; elytra with sub-basal ridge reduced to horizontal fragment, with evidence of mesoscutellar collar; humeral spines absent; protibia without lateral spines (Fig. 3 A); metatibia with only two thin spines on outer face (Fig. 3 A); pretarsi of all legs with one pair of empodial setae; abdominal sternite III with basal transverse carina produced posteriad at a sharp angle.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
E50B322C5347585B901BF70A83A4E7CE.taxon	distribution	Distribution. Western North America, broadly distributed along the western cordilleras at a variety of elevations.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
EB0F25A806945B9B8CD30A0AB2C937C6.taxon	description	Fig. 3 A-G	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
EB0F25A806945B9B8CD30A0AB2C937C6.taxon	diagnosis	Diagnosis. As given above for the genus.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
EB0F25A806945B9B8CD30A0AB2C937C6.taxon	distribution	Distribution. Canada: BC. United States: CA, ID, NV, OR, WA	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B96953A861FA5081ADC10AABFFC4BB27.taxon	diagnosis	Diagnosis. Indoquediini (as recently redefined by Brunke et al. (2021)) can be recognized among other Staphylininae by the combination of: head with obvious presence of both posterior frontal and basal punctures (Brunke et al. 2019: fig. 1); protibiae subapically with distinct and unique notch (Brunke et al. 2021: fig. 8 C); all antennomeres longer than wide.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B03D9F4FA9915D55B14AA316150212E5.taxon	description	Figs 1 D-F, 2 A-F	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B03D9F4FA9915D55B14AA316150212E5.taxon	etymology	Etymology. The species epithet means ' shortened' or ' reduced', and refers to the shorter dorsal rows of punctures on the pronotum compared to S. nigropolitus.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B03D9F4FA9915D55B14AA316150212E5.taxon	diagnosis	Diagnosis. Sundaquedius abbreviatus can be distinguished by the presence of only two or three punctures in the dorsal row of the pronotum.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B03D9F4FA9915D55B14AA316150212E5.taxon	description	Description. Measurements. Male (n = 2): HW / HL 1.30 - 1.35; PW / PL 1.06 - 1.08; EW / EL 1.22 - 1.24; PW / HW 1.14 - 1.19; forebody length 4.9 - 5.4 mm. Female (n = 4): HW / HL 1.25 - 1.29; PW / PL 1.03 - 1.10; EW / EL 1.13 - 1.15; PW / HW 1.21 - 1.23; forebody length 4.7 - 5.0 mm. Similar to S. nigropolitus and differing only in the following: antennomeres dark except apical three segments paler, becoming successively paler to antennal apex; maxillary and labial palpi paler, entirely medium reddish brown; head, without additional punctures between named punctures, distinctly transverse, more so in males, head also broader relative to pronotum in males; antennae overall more robust, with apical segments less strongly transverse; pronotum slightly to distinctly transverse, with two or three punctures in the dorsal row, third puncture, when present, smaller, sometimes rudimentary and without seta; elytra more transverse than in S. nigropolitus, and even more so in males, with two discal rows and without scattered additional punctures; abdominal tergites III and IV with distinct impressions, V with only vague impression; abdominal punctation slightly denser but punctures generally still well separated. Male with sternite VII broadly but shallowly emarginate; sternite VIII with slightly deeper emargination and distinct, triangular impressed and glabrous area; tergite X elongate, with distinct shallow emargination, with many long setae at apex (Fig. 2 D); sternite IX with bulky, asymmetrical base, apex deeply emarginate (Fig. 2 E); median lobe of aedeagus in ventral view subparallel sided, narrowing to rounded, acute apex, paramere with broad base, becoming slender to strongly acute apex (Fig. 2 A); median lobe in lateral view with nearly straight ventral face, with short, rounded apical part (Fig. 2 C); apex of paramere with short, sparse paired row of marginal peg setae (Fig. 2 B); aedeagus with ventral paired copulatory sclerites broadest at base and strongly narrowed to sharp apex. Female tergite X similar to that of S. nigropolitus but with slightly narrower apex (Fig. 2 F).	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
B03D9F4FA9915D55B14AA316150212E5.taxon	distribution	Distribution. Known only from the type locality in the central highlands of Vietnam.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
496DB27496DD55CD98997BC808D87444.taxon	description	Figs 1 A-F, 2 A-G	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
496DB27496DD55CD98997BC808D87444.taxon	etymology	Etymology. The generic name refers to the Sunda Plate and Quedius, with which members of this genus and closely related genus Cyrtoquedius were associated with for a long time. Much of the Sunda Plate is currently below sea level but had connected terrestrial species on Borneo, Sumatra, Java and the present southeast Asian mainland in multiple episodes, from about the Eocene to as recently as the Pleistocene (e. g., Inger and Voris 2001). Noun in apposition.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
496DB27496DD55CD98997BC808D87444.taxon	diagnosis	Diagnosis. Among other Oriental Cyrtoquediini, Sundaquedius is easily recognized by a combination of the large eyes (more than 3 x as long as temples) (Fig. 1 B, E), incomplete infraorbital ridge and elytra with rows of setose punctures. It can be distinguished from its putative close relatives Cyrtoquedius and Parisanopus by any one of: more than one puncture in the dorsal row of the pronotum (Fig. 1 C, F), two or more parocular punctures on the head (Fig. 1 B, E), the incomplete infraorbital ridge and presence of peg setae on the paramere (Fig. 2 B).	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
496DB27496DD55CD98997BC808D87444.taxon	description	Description. With the character states of Cyrtoquediini (see Brunke et al. 2021) and the following: head with basal puncture present but not doubled; two or three parocular punctures present; antennae non-geniculate, antennomeres 1 - 3 sparsely pubescent and without tomentose pubescence, antennomere 4 with some tomentose pubescence but much sparser than 5; labrum with two usual lobes and moderately incised median emargination; apical maxillary and labial palpi fusiform, apical labial palpomere with sparse, short setae; mandibles slender in apical half and markedly broad in basal half, bearing a single proximal tooth; gular sutures convergent, separate but running extremely close in basal half; infraorbital ridge / nuchal ridge incomplete, reaching ~ 1 / 3 to 1 / 2 the distance to mandible base; pronotum strongly convex, non-explanate and slightly elongate, with 2 - 8 punctures in the dorsal row, ' second' puncture present; basisternum with pair of macrosetae at middle; mesoscutellum glabrous and without micropunctures; disc of elytra without microsculpture and glabrous, except for three rows of coarse setose macropunctures (one sutural, two discal), rows slightly disorganized due to extra punctures in S. nigropolitus; elytra with epipleuron bearing row of coarse, setose macropunctures, epipleuron with additional rows and clusters of coarse setae; epipleural margin not thickened; mesocoxae contiguous; metatibia spinose, with three spines on outer face, inner face without spines; pro- and metatarsomeres with setae on disc, setae not restricted to margins; metatarsomere 4 with ventral setae distinctly interrupted medially and removed from apical margin; abdominal tergite IV with impression but punctures only slightly more impressed, not markedly coarser in impression (as in Bolitogyrus); abdominal sternite III with basal transverse line sharply produced posteriad forming an acute angle at middle; abdominal sternite IV with basal transverse line not produced; aedeagus with single fused paramere bearing well-developed peg setae, internal sac with ventral, paired copulatory sclerites, with an additional sclerotized structure similar to dorsal copulatory piece, but singular, and more weakly sclerotized compared to Cyrtoquedius or Parisanopus, and held within spinose internal sac.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
496DB27496DD55CD98997BC808D87444.taxon	distribution	Distribution. Sundaquedius is presently known only from central Vietnam and East Java but likely occurs at medium elevations across southeast Asia, west of Wallace's line.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
35316AE5F20359258E78B9EF35BD318C.taxon	description	Figs 1 A-C, 2 G	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
35316AE5F20359258E78B9EF35BD318C.taxon	diagnosis	Diagnosis. Sundaquedius nigropolitus can be easily recognized by the dorsal rows of the pronotum, which have seven or eight punctures in each row. The only other known species is allopatric.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
35316AE5F20359258E78B9EF35BD318C.taxon	distribution	Distribution. Known only from the type locality in East Java, which is at the northern edge of the plateau.	en	Brunke, Adam J. (2021): New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae). ZooKeys 1076: 109-124, DOI: http://dx.doi.org/10.3897/zookeys.1076.73103, URL: http://dx.doi.org/10.3897/zookeys.1076.73103
