identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
489497C318E25994AEC98FDF11379162.text	489497C318E25994AEC98FDF11379162.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thymus richardii subsp. aureopunctatus (Beck) (Beck) L. Sáez, Bogunić & Bogdanović, comb. & 2021	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 2.  
Thymus richardii subsp. aureopunctatus (Beck) L. 
Saez
, 
Bogunic
&amp; 
Bogdanovic
, comb. &amp;
 stat. nov.</p>
            <p> Thymus humifusus aureopunctatus ≡  Thymus humifusus var. aureopunctatus Beck, Ann. Naturhist. Mus. Wien 2: 142. 1887, basionym. </p>
            <p> Thymus aureopunctatus ≡  Th. aureopunctatus (Beck) K.  Malý , Prilozi za floru Bosne i Herzegovine: 557. 1908. </p>
            <p>Type.</p>
            <p>  Herc. [Herzegovina],  Nächst Konjica, 8 July 1885, G. Beck (lectotype: PRC 455886! designated here, Fig. 5)  . </p>
            <p>Description.</p>
            <p> Stems up to 45 cm long, procumbent to reptant. Leaf blade up to 7.7  × 5.3 mm, suborbicular to elliptical, entire. Inflorescence 8-21 mm long, capitate; bracts up to 6.5  × 5 mm, similar to leaves, entire, usually hairy at margin (eglandular hairs up to 1 mm long). Calyx 3-5 mm long, glabrescent to sparsely hairy, with eglandular hairs up to 0.5 mm long, without stipitate glandular hairs; calyx tube 1.3-2.2 mm long, sparsely hairy, with eglandular hairs up to 0.5 mm long on the ventral surface; central tooth of upper lip 0.7-1.6 mm long, lower teeth 2-3 mm long, with hairs pectinate up to 0.5 mm long. Corolla 6-9 mm long, rose (Fig. 4, A, B, D). </p>
            <p>Chromosome number.</p>
            <p>2 n = 28 (Kaleva 1969).</p>
            <p>Distribution.</p>
            <p> Endemic to surroundings of Konjic (  Podorašac , Koznik,  Dužani , Dudle,  Džepi , Zlatar, Borci, Spiljani,  Glavatičevo , Pribilja, Repovica), northern Herzegovina. The taxon covers an area of c. 280 km2. </p>
            <p>Habitat.</p>
            <p>Sandy dolomites and dolomitic rocky places, 400-1040 m a.s.l.</p>
            <p>Remarks.</p>
            <p> Beck (1887) described  Th. humifusus var. aureopunctatus from "In saxosis prope Konjicam" [Bosnia and Herzegovina] and related this new variety to  Thymus humifusus Bernh. ex Link, which is currently regarded a synonym of the tetraploid  Th. praecox Opiz (Jalas 1971; Euro+Med 2006; Plant List 2021; WFO 2021).  Günther Beck (1856-1931) was a Bohemian botanist, and his herbarium is currently kept at PRC and W (Stafleu and Cowan 1976). We have been able to locate original material of  Th. humifusus var. aureopunctatus at PRC. This is a well-prepared specimen; it matches the description and the provenance indicated in the protologue. Therefore, we designate the specimen with barcode PRC 455886 as the lectotype of the name  Th. humifusus var. aureopunctatus (Fig. 5). The taxon occurs in fragmented subpopulations in Bosnia and Herzegovina. Their habitats are threatened by forest succession and canopy closure, but frequent fire incidences represent the most serious threat to its subpopulations. However, the overall population trend of  Th. richardii subsp. aureopunctatus is inferred to be generally stable (F.  Bogunić , pers. observ.). </p>
            <p>Specimens examined.</p>
            <p>
                  Bosnia and Herzegovina.  
                <a title="Search Plazi for locations around (long 18.121262/lat 43.540565)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.121262&amp;materialsCitation.latitude=43.540565">Konjic</a>
                 , 8 July 1885, G. Beck (PRC 455886!, lectoptype); Konjic,  Dužani , 43.509894N 18.152114E, 830 m, 10 July 2020, F.  Bogunić (SARA, ZAGR, L.  Sáez herb. pers.); Konjic,  Džepi , 43.675506N 18.011992E 757 m, 10 July 2020, F.  Bogunić (SARA, ZAGR, L.  Sáez herb. pers.); Bosnia and Herzegovina, Dudle, 43.540567N 18.121261E, 1034 m, 10 July 2020, F.  Bogunić (SARA, ZAGR, L.  Sáez herb. pers.); Flora Herzegovinae. In pineti (Pinus nigra) inter  Pričepa-Bigolje ; solo dolomitico, 720 m, 9 August 1908, K. Maly (ZA); Flora  Hercegovinae . In saxosis dolomiticis ad Repovica prope Konjic, 12 July 1931, V. Loschingg (ZA)  . 
            </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/489497C318E25994AEC98FDF11379162	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Saez, Llorenc;Bogunic, Faruk;Cambria, Salvatore;Riera, Jesus;Bogdanovic, Sandro	Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus, Bogdanovic, Sandro (2021): On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex. PhytoKeys 186: 139-158, DOI: http://dx.doi.org/10.3897/phytokeys.186.75412, URL: http://dx.doi.org/10.3897/phytokeys.186.75412
2A0D0D5311135CC8A0B31705A7C52B0C.text	2A0D0D5311135CC8A0B31705A7C52B0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thymus richardii subsp. ebusitanus (Font Quer) (Font Quer) Jalas, Bot. J. Linn. Soc. 64 (3): 264. 1971	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 3.  Thymus richardii subsp. ebusitanus (Font Quer) Jalas, Bot. J. Linn. Soc. 64(3): 264. 1971</p>
            <p> Thymus richardii ebusitanus ≡  Thymus richardii var. ebusitanus Font Quer, Cavanillesia 7: 77. 1935; </p>
            <p> Thymus ebusitanus ≡  Th. ebusitanus (Font Quer) Romo, Fl. Silvestres Baleares: 266. 1994. </p>
            <p>Type.</p>
            <p>Eivissa, cala de les Torretes, 29 May 1918, Font Quer &amp; Gros (lectotype: BC 50117! designated by Jalas 1971: 264).</p>
            <p>Description.</p>
            <p> Stems up to 54 cm long, more or less reptant to suberect. Leaf blade up to 11  × 8 mm, suborbicular to elliptical, entire. Inflorescence 19-62 mm long, oblong; bracts up to 8.3  × 7.3 mm, similar to leaves, entire; glabrous to hairy at margin and midrib (eglandular hairs up to 1 mm long). Calyx 4.5-6.9 mm long, densely hairy, sometimes hirsute, with eglandular hairs up to 1 mm long, usually with stipitate glandular hairs; calyx tube 1.8-2.7 mm long, densely hairy, with eglandular hairs up to 1 mm long on the ventral surface; central tooth of upper lip 0.9-1.9 mm long, lower teeth 1.9-3.1 mm long, with pectinate hairs up to 1 mm long. Corolla 6-8.5 mm long, pale rose. </p>
            <p>Chromosome number.</p>
            <p>2 n = 30 (Morales 1990).</p>
            <p>Distribution.</p>
            <p>Endemic to northern Eivissa, Western Balearic Islands (Spain).</p>
            <p>Habitat.</p>
            <p>Limestone rocky places, 5-370 m a.s.l.</p>
            <p>Specimens examined.</p>
            <p>  Spain. Balearic Islands, Eivissa: cala de les Torretes, 29 May 1918, Font Quer &amp; Gros (BC 50117, lectotype); Santa  Agnès , a la Cala de les Torretes, 15 June 1918,  Gros (BC 50116);  Cala de’n Damià , 10 July 1920,  Gros (BC 858975, P 04438273); cala de les Torretes, 8 July 1920,  Gros (BC 859210, P 04438274); cala de Santa  Agnès , 5 m, July 1935, Gros (BC 87078, BC87079); Cala Aubarca, 1 Aug 1974, J.Y.  Lesouëf (MA 620032)  ;   vicum Sant Mateu, cala  d’Aubarca , 31SCD 52, 50 m, 23 June 1979,  Fernández Casas 2883 (BC 633215)  ;   Cala den  Sardina , 2 June 1981, Cardona &amp; al. (BC 644574)  ;   Cala den  Sardina , 20 June 1983, L. Llorens (Herb. Univ. Illes Balears)  ;   cingles  d’en Recó , 8 June 1997, N. Torres, M. Mayol &amp; L.  Sáez (MA 592780)  ;   Ses Balandres , 31SCD 5523, 131 m, 3 June 2010, C.  Benedí &amp; L.  Sáez (L.  Sáez , herb. pers.)  . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/2A0D0D5311135CC8A0B31705A7C52B0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Saez, Llorenc;Bogunic, Faruk;Cambria, Salvatore;Riera, Jesus;Bogdanovic, Sandro	Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus, Bogdanovic, Sandro (2021): On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex. PhytoKeys 186: 139-158, DOI: http://dx.doi.org/10.3897/phytokeys.186.75412, URL: http://dx.doi.org/10.3897/phytokeys.186.75412
8108DAB1BEB25D36B6F72521E8F08340.text	8108DAB1BEB25D36B6F72521E8F08340.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thymus richardii subsp. nitidus (Guss.) (Guss.) Jalas, Bot. J. Linn. Soc. 64: 264. 1971	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 4.  Thymus richardii subsp. nitidus (Guss.) Jalas, Bot. J. Linn. Soc. 64: 264. 1971</p>
            <p> Thymus nitidus ≡  Thymus nitidus Guss., Fl. Sicul. Syn. 2(1): 97. 1844; </p>
            <p> Thymus serpyllum nitidus ≡  Th. serpyllum var. nitidus (Guss.)  Bég . in Fiori &amp;  Béguinot , Fl. Italia 3: 66. 1903. -  Th. sensu lucidus Guss., Fl. Sicul. Prodr., Suppl.: 198. 1843 </p>
            <p>Type.</p>
            <p>Marettimo, 10 May 1829, Herb. Gussone Sicilia s.c., bottom-right specimen (Lectotype: NAP-Gussone!, designated by Bartolucci et al. 2013: 1310).</p>
            <p>Description.</p>
            <p> Stems up to 25 cm long, procumbent or suberect. Leaf blade up to 10  × 4.5 mm, elliptical, entire. Inflorescence 8-30 mm long, subcapitate to oblong; bracts up to 7  × 4 mm, similar to leaves, entire, glabrous to hairy at margin and midrib (eglandular hairs up to 0.4 mm long, mixed with stipitate glandular hairs). Calyx 4.5-6.3 mm long, densely covered by stipitate glandular hairs and sparse eglandular hairs up to 0.5 mm long; calyx tube 1.9-2.5 mm long, with eglandular hairs up to 0.5 mm long on the ventral surface; central tooth of upper lip 0.8-1.5 mm long, lower teeth 2-3 mm long, with pectinate hairs up to 0.5 mm long. Corolla 6.5-9.5 mm long, pale rose. </p>
            <p>Chromosome number.</p>
            <p>2 n = 28 (Morales 1997)</p>
            <p>Distribution.</p>
            <p>Endemic to Island of Marettimo, Sicily (Italy).</p>
            <p>Habitat.</p>
            <p>Limestone rocky places, 10-600 m a.s.l.</p>
            <p>Specimens examined.</p>
            <p>  Italy. Sicily, Marettimo, sine leg. (PAL);  Isola di Marettimo , rupi di P. Anzine, 21 July 2007,  Scuderi (VAL 184304)  . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/8108DAB1BEB25D36B6F72521E8F08340	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Saez, Llorenc;Bogunic, Faruk;Cambria, Salvatore;Riera, Jesus;Bogdanovic, Sandro	Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus, Bogdanovic, Sandro (2021): On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex. PhytoKeys 186: 139-158, DOI: http://dx.doi.org/10.3897/phytokeys.186.75412, URL: http://dx.doi.org/10.3897/phytokeys.186.75412
8DECCD33118D5BEB9A360BC03E884E09.text	8DECCD33118D5BEB9A360BC03E884E09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thymus richardii subsp. richardii Pers., Syn. Pl. 2: 130. 1806	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 1.  Thymus richardii subsp. richardii Pers., Syn. Pl. 2: 130. 1806</p>
            <p> Thymus serpyllum richardii ≡  Thymus serpyllum var. richardii (Pers.) Knoche, Fl. Balear. 2: 354. 1922. </p>
            <p> Thymus serpyllum richardii ≡  Th. serpyllum subsp. richardii (Pers.) Malag., H. Bianor, Educador  Botánico Baleares: 150. 1971. </p>
            <p>Type.</p>
            <p> Holotype (see  Rosselló and  Sáez 2001: 109): P-Lamarck. </p>
            <p>Description.</p>
            <p> Stems up to 47 cm long, procumbent to reptant. Leaf blade up to 13  × 7.7 mm, broadly ovate to elliptical, entire. Inflorescence 15-30 mm long, capitate to oblong; bracts up to 11  × 7.8 mm, similar to leaves, entire, glabrous. Calyx 6-8 mm long, glabrescent (sometimes glabrous), with eglandular hairs up to 0.3 mm long, occasionally with scattered stipitate glandular hairs; calyx tube 2.2-3.2 mm long, glabrescent (sometimes glabrous on the dorsal surface), with eglandular hairs up to 0.3 mm long on the ventral surface; central tooth of upper lip 1.3-2.2 mm long, lower teeth 2.8-3.8 mm long, with pectinate hairs up to 0.3 mm long. Corolla 7-11 mm long, rose to pinkish-purple (Fig. 4, C, E). </p>
            <p>Chromosome number.</p>
            <p>2 n = 30 (Morales 1995).</p>
            <p>Distribution.</p>
            <p>Endemic to Mallorca, Eastern Balearic Islands (Spain).</p>
            <p>Habitat.</p>
            <p>Cliffs, on humid and north-facing limestone rocks, 250-1430 m a.s.l.</p>
            <p>Remarks.</p>
            <p> This is a rare plant, documented from three localities in the north of Mallorca (Ternelles mountain, Formentor peninsula and Puig Major) of which we have only been able to verify its presence in the last locality, growing on cliffs with very difficult access. This taxon could be facing a population decline. Bianor (1917) at the beginning of the 20th century, considered it as abundant in the Puig Major ["Abondant dans les endroits peu accessibles"]. In fact, there are dozens of specimens from this mountain and which are preserved in various herbaria; mostly collected in the late 19th and early 20th centuries. Currently,  Th. richardii subsp. richardii is very scarce at the same locality where it was reported by Bianor (1917) and the plants are practically inaccessible if climbing techniques are not used. Another population located on a different slope of the same massif is also scarce and very difficult to access. This possible population decline could be due to a loss of potential habitat and intense predation by feral goats (  Capra hircus ). </p>
            <p>Specimens examined.</p>
            <p>  Spain. Balearic Islands, Mallorca: Comma de  n’Arbona ,  Puig Major , 12 June 1852, G. Vigineix (P 04436032, P 04436034);  Majorque , 19 June 1869,  Bourgeau (P 04436046); rochers des  Arbonas [  n’Arbona ], 17 Apr 1870, F.  Barceló (COI 00045051, P 03389631);  
Coma de 
n'Arbona et  Puig Major de Son Torrella , 1000-1300 m, 24 June 1885 and 29 July 1885, Porta &amp; Rigo (P 04436045);  Mallorca :  
Coma de 
n'Arbona ,  Sóller , 30 June 1879, A.  Crespí (BC 651145, P 04407218); Puig Major, 1000-1400 m, 12 July 1917, F. Bianor (BC 50119);  Féntes des rochers, Puig Major, 1000-1450 m, 12 July 1918, F. Bianor (BC 50118); Puig Major,  Féntes des rochers, 1000-1500 m, 7 July 1919, F. Bianor, Pl. Espagne F. Sennen 3768 (BC 50123);  
Coma de 
n'Arbona , 18 June 1920, Gros (BC 859198, P 04407218);  
Coma de 
n'Arbona , 4 July, 1936, Kennedy 48 (BC 103732);  Sóller , escletxes dels espadats de la  
Coma de 
n'Arbona , July, 1958, L.  Garcías Font (BC 145169); Puig Major,  
Coma de 
n'Arbona , 27 June 1985, T. Rabassa (HBJS 5700); Puig Major, Escorca, 10 July 1986, L.  Sáez (MA 592837);  Puig Major de Son Torrella , c. via des Bufador, Escorca, 31SDE8207, 1200 m, 14 June 2006, L.G. Valle &amp; L.  Sáez LS-6445 (L.  Sáez , herb. pers.); Escorca, Puig Major,  Penyal des Bufador , 31DE 8206, 1340 m, 30 June 2020, L.  Sáez (L.  Sáez , herb. pers.); Fornalutx, Coma  N’Arbona , 31SDE8105, 1100 m, 17 June 2021, L.  Sáez (L.  Sáez , herb. pers.)  . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/8DECCD33118D5BEB9A360BC03E884E09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Saez, Llorenc;Bogunic, Faruk;Cambria, Salvatore;Riera, Jesus;Bogdanovic, Sandro	Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus, Bogdanovic, Sandro (2021): On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex. PhytoKeys 186: 139-158, DOI: http://dx.doi.org/10.3897/phytokeys.186.75412, URL: http://dx.doi.org/10.3897/phytokeys.186.75412
0D988EF069CF5B1896B2A0B427C1A8C1.text	0D988EF069CF5B1896B2A0B427C1A8C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thymus richardii subsp. vigoi Riera, Güemes & Rosselló, Fl. Montiber. 37: 78. 2007	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 5.  
Thymus richardii subsp. vigoi Riera, 
Gueemes
&amp; 
Rossello
, Fl. Montiber. 37: 78. 2007
</p>
            <p> Thymus richardii vigoi Type. Spain, Valencia, Villalonga, La Safor, ad  l’Orxa , 30SYJ3706, 600 m, 4 July 2000, J. Riera &amp; J.  Güemes (holotype: VAL 185406!; isotype: MA 757804!; Riera et al. 2007). </p>
            <p> Thymus richardii valentinus =  Thymus richardii var. valentinus O.  Bolòs &amp; Vigo, Collect. Bot. (Barcelona) 14: 95. 1983 </p>
            <p>Type.</p>
            <p>  Spain, Valencia province, Valentia, c.  Gandia , 15 Sept 1950, P.  Cañigueral (holotype: BC 119858!;  Bolòs and Vigo 1983)  . </p>
            <p>Description.</p>
            <p> Stems up to 16 cm long, suberect to erect. Leaf blade up to 11  × 8.1 mm, ovate-triangular, denticulate. Inflorescence 15-34 mm long, usually oblong; bracts up to 8  × 7 mm, similar to leaves, denticulate, usually glabrous. Calyx 5.8-7 mm long, sparsely hairy, with eglandular hairs up to 0.7 mm long, sometimes with sparse stipitate glandular hairs; calyx tube 2.1-2.9 mm long, with eglandular hairs up to 0.5 mm long on the ventral surface; central tooth of upper lip 1.8-2.6 mm long, lower teeth 3-3.5 mm long, with pectinate hairs up to 0.7 mm long. Corolla 7-10 mm long, whitish to pale rose, sometimes cream. </p>
            <p>Chromosome number.</p>
            <p>Unknown.</p>
            <p>Distribution.</p>
            <p>Endemic to Alicante and Valencia provinces (Spain).</p>
            <p>Habitat.</p>
            <p>Open scrub, on limestone soil, 130-600 m a.s.l.</p>
            <p>Remarks.</p>
            <p> Plants which were considered to be hybrids between  Th. richardii subsp. vigoi and  T. piperella L. have been called  
T. 
x
bolosii
 . The hybrid has been reported from a small area of Serra de la Safor, eastern Spain (Riera et al. 2020). </p>
            <p>Specimens examined.</p>
            <p>  Spain. Alicante province: La Vilallonga,  La Safor , 136 m, 22 June 1984, J.B. Peris &amp; G.  Stübing (BC 674556); Valencia province, Valentia, c. Gandia, 15 Sept 1950, P.  Cañigueral (BC 119858); Villalonga, La Safor, ad  l’Orxa , 30SYJ 3706, 600 m, 4 July 2000, J. Riera &amp; J.  Güemes (VAL 185406!)  . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/0D988EF069CF5B1896B2A0B427C1A8C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Saez, Llorenc;Bogunic, Faruk;Cambria, Salvatore;Riera, Jesus;Bogdanovic, Sandro	Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus, Bogdanovic, Sandro (2021): On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex. PhytoKeys 186: 139-158, DOI: http://dx.doi.org/10.3897/phytokeys.186.75412, URL: http://dx.doi.org/10.3897/phytokeys.186.75412
